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<mods:title id="24340288BB2A710D4F707E3F5ABEE06A">Taxonomy of the genus Protocypus J. Müller, 1923 (Coleoptera: Staphylinidae: Staphylinini) and two new species from China</mods:title>
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Key to species of the genus
<taxonomicName id="4C554D615B63856FFE3C6917881F7895" ID-CoL="8W2QF" authority="J. Muller, 1923" authorityName="J. Muller" authorityYear="1923" box="[491,820,1644,1670]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="459" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAF05B63856FFE3C691789597895" bold="true" box="[491,626,1644,1670]" italics="true" pageId="1" pageNumber="459">Protocypus</emphasis>
<bibRefCitation id="EFC44B135B63856FFDAE6917881F7895" author="Muller" box="[633,820,1644,1670]" pageId="1" pageNumber="459" refString="Muller, J. (1923) Contributo alla conoscenze del genere Staphylinus L. Bollettino della Societa Entomologica Italiana, 55, 135 - 144." type="journal article" year="1923">J. Müller, 1923</bibRefCitation>
</taxonomicName>
from China
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(modified from
<bibRefCitation id="EFC44B135B63856FFE9D69F48AD978B4" author="Smetana" box="[330,498,1678,1703]" pageId="1" pageNumber="459" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
)
</paragraph>
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<keyLead id="30A495D75B63856FFF4069AE8EB7791A" pageId="1" pageNumber="459">1. Antenna short, when reclined, tip of third segment distinctly not reaching posterior margin of eye, segments 3 and 4 only slightly longer than wide................................................................................2</keyLead>
</paragraph>
<paragraph id="8BEA36E25B63856FFF40686A8EB07950" blockId="1.[151,1436,1749,2035]" pageId="1" pageNumber="459">
<keyLead id="30A495D75B63856FFF40686A8EB07950" pageId="1" pageNumber="459">- Antenna long, when reclined, tip of third segment reaching variably beyond posterior margin of eye, segments 3 and 4 distinctly longer than wide................................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E475B63856FFF4068308EB079C5" pageId="1" pageNumber="459">
<paragraph id="8BEA36E25B63856FFF4068308EB77988" blockId="1.[151,1436,1749,2035]" pageId="1" pageNumber="459">
<keyLead id="30A495D75B63856FFF4068308EB77988" pageId="1" pageNumber="459">
2. Legs at least partially darkened, dark brunneous to brunneopiceous. Each elytron with vague, slightly elevated area in middle behind scutellum. Pronotal hypomeron extensively and relatively densely setose. Aedeagus as in
<bibRefCitation id="EFC44B135B63856FFB5768138EB3796D" author="Smetana" box="[1152,1432,1896,1918]" pageId="1" pageNumber="459" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana (2005: Figs. 1517)</bibRefCitation>
. Length
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. Chonqing........................................................
<taxonomicName id="4C554D615B63856FFB5F68FD8EB77988" ID-CoL="4ZJGZ" authority="Bernhauer, 1933" authorityName="Bernhauer" authorityYear="1933" box="[1160,1436,1925,1947]" class="Insecta" family="Staphylinidae" genus="Staphylinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="459" phylum="Arthropoda" rank="species" species="beckeri">
<emphasis id="B921EAF05B63856FFB5F68FD8FCD7988" box="[1160,1254,1925,1947]" italics="true" pageId="1" pageNumber="459">P. beckeri</emphasis>
(
<bibRefCitation id="EFC44B135B63856FFB2468FE8EBE7988" author="Bernhauer" box="[1267,1429,1925,1947]" pageId="1" pageNumber="459" refString="Bernhauer, M. (1933) Neuheiten der chinesischen Staphylinidenfauna. Wiener Entomologische Zeitung, 50, 25 - 48." type="journal article" year="1933">Bernhauer, 1933</bibRefCitation>
)
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<keyLead id="30A495D75B63856FFF4068DF8EB079C5" pageId="1" pageNumber="459">- Legs entirely reddish. Elytron without elevated area in middle behind scutellum. Pronotal hypomeron not extensively and densely setose. Aedeagus different........................................................................ 3</keyLead>
</paragraph>
</keyStep>
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<keyLead id="30A495D75B63856FFF4068A68EB779E0" box="[151,1436,2013,2035]" pageId="1" pageNumber="459">3. Apical portion of median lobe of aedeagus relatively narrow (Fig. 3-2. F), paramere smaller with apical portion shorter and</keyLead>
</paragraph>
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with apex wider (Figs. 3-1. D, 3-2. G). Aedeagus (Figs. 3-1. B &amp; C, 3-2. B, C &amp; D) in general only moderately large (on average
<quantity id="4CAD9B075B60856CFEDF6FCF8A7A7ED9" box="[264,337,180,202]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="460" unit="mm" value="2.3">2.3 mm</quantity>
long). Smaller, less robust species. Length
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.
<collectingRegion id="4991F8005B60856CFC5E6FCF88F97ED9" box="[905,978,180,202]" country="China" name="Beijing" pageId="2" pageNumber="460">Beijing</collectingRegion>
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,
<materialsCitation id="3B3D3CBF5B60856CFC0B6FCF8F4C7ED9" ID-GBIF-Occurrence="1572323554" box="[988,1127,180,202]" location="Gansu" pageId="2" pageNumber="460" specimenCount="1" stateProvince="Gansu">
<collectingRegion id="4991F8005B60856CFC0B6FCF8F377ED9" box="[988,1052,180,202]" country="China" name="Gansu" pageId="2" pageNumber="460">Gansu</collectingRegion>
, Heibei
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,
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<collectingRegion id="4991F8005B60856CFBA66FCF8FEA7ED9" box="[1137,1217,180,202]" country="China" name="Ningxia" pageId="2" pageNumber="460">Ningxia</collectingRegion>
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,
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<collectingRegion id="4991F8005B60856CFB1D6FCF8E3B7ED9" box="[1226,1296,180,202]" country="China" name="Shanxi" pageId="2" pageNumber="460">Shanxi</collectingRegion>
..................................................................................
<taxonomicName id="4C554D615B60856CFBF66FA98EB07EF4" ID-CoL="9PTKZ" authority="Eppelsheim, 1889" authorityName="Eppelsheim" authorityYear="1889" box="[1057,1435,209,231]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="460" phylum="Arthropoda" rank="species" species="fulvotomentosus">
<emphasis id="B921EAF05B60856CFBF66FA98FF37EF4" box="[1057,1240,209,231]" italics="true" pageId="2" pageNumber="460">P. fulvotomentosus</emphasis>
(
<bibRefCitation id="EFC44B135B60856CFB326FAA8EBF7EF4" author="Eppelsheim" box="[1253,1428,209,231]" pageId="2" pageNumber="460" refString="Eppelsheim, E. (1889) Insecta, A Cl. G. N. Potanin in China et in Mongolia novissime lecta. V. Neue Staphylinen. Horae Societatis Entomologicae Rossicae, 23, 169 - 184." type="journal article" year="1889">Eppelsheim, 1889</bibRefCitation>
)
</taxonomicName>
<keyLead id="30A495D75B60856CFF406F948F147F2C" pageId="2" pageNumber="460">
- Apical portion of median lobe of aedeagus wider and of different shape (
<bibRefCitation id="EFC44B135B60856CFC7D6F948FA47F16" author="Smetana" box="[938,1167,239,261]" pageId="2" pageNumber="460" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana, 2005, Fig. 11</bibRefCitation>
), paramere distinctly larger with apical portion longer and with apex narrower (
<bibRefCitation id="EFC44B135B60856CFD196E77889A7F31" author="Smetana" box="[718,945,268,290]" pageId="2" pageNumber="460" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana, 2005: Fig. 12</bibRefCitation>
). Aedeagus (
<bibRefCitation id="EFC44B135B60856CFBFA6E778E047F31" author="Smetana" box="[1069,1327,268,290]" pageId="2" pageNumber="460" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana, 2005: Figs. 913</bibRefCitation>
) in general markedly larger (on average 3.0 mm). Larger, more robust species. Length 18.020.0 mm
</keyLead>
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.
<materialsCitation id="3B3D3CBF5B60856CFB9F6E518FA27F2C" ID-GBIF-Occurrence="1572323543" box="[1096,1161,298,319]" location="Henan" pageId="2" pageNumber="460" specimenCount="1" stateProvince="Henan">
<collectingRegion id="4991F8005B60856CFB9F6E518FA27F2C" box="[1096,1161,298,319]" country="China" name="Henan" pageId="2" pageNumber="460">Henan</collectingRegion>
</materialsCitation>
,
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<collectingRegion id="4991F8005B60856CFB456E528FC97F2C" box="[1170,1250,297,319]" country="China" name="Shaanxi" pageId="2" pageNumber="460">Shaanxi</collectingRegion>
</materialsCitation>
,
<materialsCitation id="3B3D3CBF5B60856CFB3B6E528EB77F4E" ID-GBIF-Occurrence="1572323551" location="Shanxi" pageId="2" pageNumber="460" specimenCount="1" stateProvince="Shanxi">
<collectingRegion id="4991F8005B60856CFB3B6E528E1F7F2C" box="[1260,1332,297,319]" country="China" name="Shanxi" pageId="2" pageNumber="460">Shanxi</collectingRegion>
..........................................................................................
<taxonomicName id="4C554D615B60856CFB656E338EB77F4E" ID-CoL="4N535" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[1202,1436,327,349]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="460" phylum="Arthropoda" rank="species" species="vulpes">
<emphasis id="B921EAF05B60856CFB656E338E2C7F4E" box="[1202,1287,327,349]" italics="true" pageId="2" pageNumber="460">P. vulpes</emphasis>
<bibRefCitation id="EFC44B135B60856CFADA6E3C8EB77F4E" author="Smetana" box="[1293,1436,327,349]" pageId="2" pageNumber="460" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana, 2005</bibRefCitation>
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</paragraph>
</keyStep>
<keyStep id="30A12E475B60856CFF406E1F8EB07F84" pageId="2" pageNumber="460">
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<keyLead id="30A495D75B60856CFF406E1F8EB07F69" box="[151,1435,356,378]" pageId="2" pageNumber="460">4. Visible abdominal tergites 4 and 5 each with a patch of golden-yellow tomentose pubescence at middle.................5</keyLead>
</paragraph>
<paragraph id="8BEA36E25B60856CFF406EF98EB07F84" blockId="2.[151,1436,151,1698]" box="[151,1435,385,407]" pageId="2" pageNumber="460">
<keyLead id="30A495D75B60856CFF406EF98EB07F84" box="[151,1435,385,407]" pageId="2" pageNumber="460">- Visible abdominal tergites 4 and 5 with uniform, dark pubescence, without patches of golden-yellow pubescence at middle..8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E475B60856CFF406EE48EBE7FFC" pageId="2" pageNumber="460">
<paragraph id="8BEA36E25B60856CFF406EE48EB77FA6" blockId="2.[151,1436,151,1698]" box="[151,1436,415,437]" pageId="2" pageNumber="460">
<keyLead id="30A495D75B60856CFF406EE48EB77FA6" box="[151,1436,415,437]" pageId="2" pageNumber="460">5. Apex of median lobe of aedeagus in ventral view acute; apex of paramere narrowly arcuate to subacute.................6</keyLead>
</paragraph>
<paragraph id="8BEA36E25B60856CFF406EC68EBE7FFC" blockId="2.[151,1436,151,1698]" pageId="2" pageNumber="460">
<keyLead id="30A495D75B60856CFF406EC68EBE7FFC" pageId="2" pageNumber="460">- Apex of median lobe of aedeagus in ventral view obtuse, widely rounded to subtruncate; apical portion of paramere wide, with apex obtuse to widely rounded.......................................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E475B60856CFF406E8C8EB77C91" pageId="2" pageNumber="460">
<paragraph id="8BEA36E25B60856CFF406E8C8EB77C54" blockId="2.[151,1436,151,1698]" pageId="2" pageNumber="460">
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<keyLead id="30A495D75B60856CFF406E8C8EB77C54" pageId="2" pageNumber="460">
6. Aedeagus moderately robust (Figs. 5-1 C, 5-2 C). Apex of paramere widely rounded, shaped as in Figs. 5-1 D, 5-2
<collectorName id="26A053345B60856CFA996E8C8EB07C1E" box="[1358,1435,503,525]" pageId="2" pageNumber="460">G. Male</collectorName>
sternite 9 markedly elongate (Fig. 5-2 H). Length 19.0
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...................................................................................................
<taxonomicName id="4C554D615B60856CFBA96D498E757C54" ID-CoL="4N52S" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[1150,1374,561,583]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="460" phylum="Arthropoda" rank="species" species="lupus">
<emphasis id="B921EAF05B60856CFBA96D498FE27C54" box="[1150,1225,561,583]" italics="true" pageId="2" pageNumber="460">P. lupus</emphasis>
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- Aedeagus markedly slender (
<figureCitation id="136E2A675B60856CFE2A6D34897E7C76" box="[509,597,591,613]" captionStart="FIGURE 1 - 1" captionStartId="3.[151,250,1946,1968]" captionTargetBox="[180,1408,424,1915]" captionTargetId="figure@3.[161,1434,409,1933]" captionTargetPageId="3" captionText="FIGURE 1 - 1. Protocypus taibaiensis sp. nov.: A. habitus of species (2.0 mm); B. aedeagus, lateral view; (0.5 mm); C. aedeagus, ventral view (0.5 mm); D. underside of apical portion of paramere (0.5 mm); E. second gonocoxite of female genital segment (0.5 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/827608/files/figure.png" pageId="2" pageNumber="460">Figs. 1-1</figureCitation>
C, 1-2 C). Apex of paramere narrow and knife-like, shaped as in
<figureCitation id="136E2A675B60856CFB366D348E137C76" box="[1249,1336,591,613]" captionStart="FIGURE 1 - 1" captionStartId="3.[151,250,1946,1968]" captionTargetBox="[180,1408,424,1915]" captionTargetId="figure@3.[161,1434,409,1933]" captionTargetPageId="3" captionText="FIGURE 1 - 1. Protocypus taibaiensis sp. nov.: A. habitus of species (2.0 mm); B. aedeagus, lateral view; (0.5 mm); C. aedeagus, ventral view (0.5 mm); D. underside of apical portion of paramere (0.5 mm); E. second gonocoxite of female genital segment (0.5 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/827608/files/figure.png" pageId="2" pageNumber="460">Figs. 1-1</figureCitation>
D, 1-2
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sternite 9 not distinctly elongate (
<figureCitation id="136E2A675B60856CFD8E6D17898D7C91" box="[601,678,620,642]" captionStart-0="FIGURE 1 - 1" captionStart-1="FIGURE 1 2" captionStartId-0="3.[151,250,1946,1968]" captionStartId-1="4.[151,250,1945,1967]" captionTargetBox-0="[180,1408,424,1915]" captionTargetBox-1="[165,1429,193,1925]" captionTargetId-0="figure@3.[161,1434,409,1933]" captionTargetId-1="figure@4.[165,1429,193,1925]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1 - 1. Protocypus taibaiensis sp. nov.: A. habitus of species (2.0 mm); B. aedeagus, lateral view; (0.5 mm); C. aedeagus, ventral view (0.5 mm); D. underside of apical portion of paramere (0.5 mm); E. second gonocoxite of female genital segment (0.5 mm). (Scale in parentheses)." captionText-1="FIGURE 1 2. Protocypus taibaiensis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm)." httpUri-0="https://zenodo.org/record/827608/files/figure.png" httpUri-1="https://zenodo.org/record/827612/files/figure.png" pageId="2" pageNumber="460">Fig. 1-2</figureCitation>
H). Length
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................
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<emphasis id="B921EAF05B60856CFA986D178EB77C91" bold="true" box="[1359,1436,620,642]" pageId="2" pageNumber="460">sp. nov.</emphasis>
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7. Median lobe only slightly curved, with almost parallel lateral sides (
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). Length 18.023.0 mm. Shaanxi......................................................................
<taxonomicName id="4C554D615B60856CFBA26DD38E757CAE" ID-CoL="4N52T" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[1141,1374,679,701]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="460" phylum="Arthropoda" rank="species" species="meles">
<emphasis id="B921EAF05B60856CFBA26DD38FE27CAE" box="[1141,1225,679,701]" italics="true" pageId="2" pageNumber="460">P. meles</emphasis>
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(pars)
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</paragraph>
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<keyLead id="30A495D75B60856CFF406DBE8EB07CE4" pageId="2" pageNumber="460">
- Median lobe distinctly curved, with apex pointed towards left sides in ventral view (
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). Length 20.025.0 mm. Gansu........................................................................
<taxonomicName id="4C554D615B60856CFB6B6D998EB07CE4" ID-CoL="4N534" authority="Smetana, 2011" authorityName="Smetana" authorityYear="2011" box="[1212,1435,737,759]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="460" phylum="Arthropoda" rank="species" species="ursus">
<emphasis id="B921EAF05B60856CFB6B6D998E237CE4" box="[1212,1288,738,759]" italics="true" pageId="2" pageNumber="460">P. ursus</emphasis>
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<paragraph id="8BEA36E25B60856CFF406D848EB77D06" blockId="2.[151,1436,151,1698]" box="[151,1436,767,789]" pageId="2" pageNumber="460">
<keyLead id="30A495D75B60856CFF406D848EB77D06" box="[151,1436,767,789]" pageId="2" pageNumber="460">8. Pronotal hypomeron with numerous setiferous punctures distributed over major portion of hypomeron..................9</keyLead>
</paragraph>
<paragraph id="8BEA36E25B60856CFF406C668EB07D21" blockId="2.[151,1436,151,1698]" box="[151,1435,796,818]" pageId="2" pageNumber="460">
<keyLead id="30A495D75B60856CFF406C668EB07D21" box="[151,1435,796,818]" pageId="2" pageNumber="460">- Pronotal hypomeron with only a few setiferous punctures.....................................................14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E475B60856CFF406C428EB07DB4" pageId="2" pageNumber="460">
<paragraph id="8BEA36E25B60856CFF406C428EB77D99" blockId="2.[151,1436,151,1698]" pageId="2" pageNumber="460">
<keyLead id="30A495D75B60856CFF406C428EB77D99" pageId="2" pageNumber="460">
9. Aedeagus in ventral view with apical portion of paramere conspicuously wrapped around left side of median lobe, with prominent longitudinal carina; apical portion of median lobe subacute, right side of median lobe below apex conspicuously, arcuately dilated (
<bibRefCitation id="EFC44B135B60856CFEB46C0F896C7D99" author="Smetana" box="[355,583,884,906]" pageId="2" pageNumber="460" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana, 2005: Fig. 46</bibRefCitation>
). Length 18.020.0 mm. Shaanxi..........................
<taxonomicName id="4C554D615B60856CFB106C0E8EB77D99" ID-CoL="4N52D" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[1223,1436,884,906]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="460" phylum="Arthropoda" rank="species" species="felis">
<emphasis id="B921EAF05B60856CFB106C0E8E2C7D99" box="[1223,1287,884,906]" italics="true" pageId="2" pageNumber="460">P. felis</emphasis>
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<keyLead id="30A495D75B60856CFF406CE98EB07DB4" box="[151,1435,913,935]" pageId="2" pageNumber="460">- Aedeagus in ventral view with apical portions of both paramere and median lobe markedly different in shape............10</keyLead>
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</keyStep>
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<paragraph id="8BEA36E25B60856CFF406CD48EB07DF1" blockId="2.[151,1436,151,1698]" pageId="2" pageNumber="460">
<keyLead id="30A495D75B60856CFF406CD48EB07DF1" pageId="2" pageNumber="460">
10. Apical portion of median lobe of aedeagus in ventral view with distinct, more or less acute triangular apex (
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)................................................................................ 11
</keyLead>
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<keyLead id="30A495D75B60856CFF406C918EB07A0E" pageId="2" pageNumber="460">
- Apical portion of median lobe of aedeagus in ventral view with apex of different shape, with no more than minute, sharp dent at middle (
<bibRefCitation id="EFC44B135B60856CFE906B7C89CD7A0E" author="Smetana" box="[327,742,1031,1053]" pageId="2" pageNumber="460" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana, 2005: Figs. 26, 28, 40, 42, 46, 48</bibRefCitation>
).......................................................12
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<paragraph id="8BEA36E25B60856CFF406B5F8EB07A44" blockId="2.[151,1436,151,1698]" pageId="2" pageNumber="460">
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11. Apical portion of paramere of aedeagus markedly asymmetrical, with subacute apex (Figs. 7-1 C, 7-2 C). Aedeagus as in Figs. 7-1 B &amp; C, 7-2
<collectorName id="26A053345B60856CFEA06B3989017A44" box="[375,554,1089,1111]" pageId="2" pageNumber="460">B, C &amp; D. Smaller</collectorName>
species of length l
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....................
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- Apical portion of paramere of aedeagus not appreciably asymmetrical (Figs. 5-1 D, 5-2 G). Aedeagus as in Figs. 5-1 B &amp; C, 5- 2
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species of length
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........
<taxonomicName id="4C554D615B60856CFBA96B068E757A81" ID-CoL="4N52S" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[1150,1374,1148,1170]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="460" phylum="Arthropoda" rank="species" species="lupus">
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12. Apex of median lobe of aedeagus in ventral view with minute, sharp projection in middle (
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). Smaller species of length 16.0
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. Hubei............................................
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- Apex of median lobe of aedeagus in ventral view without minute, sharp projection in middle, either simple (
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), or with oblique double carina splitting apical portion (
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). Larger species of length 18.0
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.......................................................................................13
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<paragraph id="8BEA36E25B60856CFF406A578EB77B4C" blockId="2.[151,1436,151,1698]" pageId="2" pageNumber="460">
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13. Apical portion of median lobe in ventral view straight and simple (
<bibRefCitation id="EFC44B135B60856CFCB86A578F7C7B51" author="Smetana" box="[879,1111,1324,1346]" pageId="2" pageNumber="460" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana, 2005: Fig. 40</bibRefCitation>
). Aedeagus as in
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. Length 18.023.0 mm. Shaanxi........................................
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<emphasis id="B921EAF05B60856CFBA26A318FE37B4C" box="[1141,1224,1353,1375]" italics="true" pageId="2" pageNumber="460">P. meles</emphasis>
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(pars)
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- Median lobe with right side distinctly curved in ventral view (Figs. 6-1 C, 6-2 C &amp; F). Aedeagus as in Figs. 6-1 B &amp; C, 6-2
<collectorName id="26A053345B60856CFA526A1C8A5A7B89" pageId="2" pageNumber="460">B, C &amp; D. Length</collectorName>
18.0
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.
<collectingRegion id="4991F8005B60856CFDD96AFF898A7B89" box="[526,673,1412,1434]" isEnumeration="true" pageId="2" pageNumber="460">Hubei, Shaanxi</collectingRegion>
............................................
<taxonomicName id="4C554D615B60856CFB676AFE8EB07B89" ID-CoL="4N52W" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[1200,1435,1412,1434]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="460" phylum="Arthropoda" rank="species" species="pilifer">
<emphasis id="B921EAF05B60856CFB676AFE8E2C7B89" box="[1200,1287,1412,1434]" italics="true" pageId="2" pageNumber="460">P. pilifer</emphasis>
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14. Apical portion of median lobe in ventral view distinctly expanded, more robust than middle portion (
<bibRefCitation id="EFC44B135B60856CFB7B6ADA8EA47BA4" author="Smetana" box="[1196,1423,1441,1463]" pageId="2" pageNumber="460" refString="Smetana, A. (2005) Contributions to the knowledge of the genera of the &quot; Staphylinus - complex &quot; (Coleoptera: Staphylinidae) of China Part. 5. The genus Protocypus J. Muller, 1923. Elytra, 33 (1), 269 - 301." type="journal article" year="2005">Smetana, 2005: Fig. 65</bibRefCitation>
). Large, robust species with wide abdomen, abdomen markedly dilated to posterior margin of third visible tergite. Length 19.023.0 mm. Shaanxi..........................................................
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<emphasis id="B921EAF05B60856CFB5A6AA68E2C7BE1" box="[1165,1287,1500,1522]" italics="true" pageId="2" pageNumber="460">P. lativentris</emphasis>
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<keyLead id="30A495D75B60856CFF406A818EB0783E" pageId="2" pageNumber="460">- Apical portion of median lobe in ventral view not distinctly expanded. Abdomen only moderately dilated to posterior margin of third visible tergite.................................................................................15</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E475B60856CFF40694F8EB778B1" pageId="2" pageNumber="460">
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15. Apical portion of median lobe shaped as in
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; paramere with semicircular apical portion widely rounded (
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). Length 17.0
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. Hubei......................................
<taxonomicName id="4C554D615B60856CFB6A69298EB07874" ID-CoL="4N52R" authority="Smetana, 2005" authorityName="Smetana" authorityYear="2005" box="[1213,1435,1617,1639]" class="Insecta" family="Staphylinidae" genus="Protocypus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="460" phylum="Arthropoda" rank="species" species="latro">
<emphasis id="B921EAF05B60856CFB6A69298E2C7874" box="[1213,1287,1617,1639]" italics="true" pageId="2" pageNumber="460">P. latro</emphasis>
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<paragraph id="8BEA36E25B60856CFF4069148EB778B1" blockId="2.[151,1436,151,1698]" pageId="2" pageNumber="460">
<keyLead id="30A495D75B60856CFF4069148EB778B1" pageId="2" pageNumber="460">
- Apical portion of median lobe distinctly different, shaped as in
<figureCitation id="136E2A675B60856CFCEC691488A27896" box="[827,905,1647,1669]" captionStart="FIGURE 1 2" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[165,1429,193,1925]" captionTargetId="figure@4.[165,1429,193,1925]" captionTargetPageId="4" captionText="FIGURE 1 2. Protocypus taibaiensis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm)." httpUri="https://zenodo.org/record/827612/files/figure.png" pageId="2" pageNumber="460">Fig. 2-2</figureCitation>
. F; paramere with apical portion lingulate (
<figureCitation id="136E2A675B60856CFAF369148E597896" box="[1316,1394,1647,1669]" captionStart="FIGURE 1 2" captionStartId="4.[151,250,1945,1967]" captionTargetBox="[165,1429,193,1925]" captionTargetId="figure@4.[165,1429,193,1925]" captionTargetPageId="4" captionText="FIGURE 1 2. Protocypus taibaiensis sp. nov.: A. male tergite 10; B and D. aedeagus, lateral view; C. aedeagus, ventral view; E. female tergite 10; F. apical portion of median lobe, ventral view, paramere removed; G. underside of apical portion of paramere; H. male sternite 9; I. second gonocoxite of female genital segment; J. male sternite 8 (Scale bars = 0.50 mm)." httpUri="https://zenodo.org/record/827612/files/figure.png" pageId="2" pageNumber="460">Fig. 2-1</figureCitation>
. D). Length
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. Hubei...........................................................
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<emphasis id="B921EAF05B60856CFB6B69F68E6278B1" box="[1212,1353,1676,1698]" italics="true" pageId="2" pageNumber="460">P. admirabilis</emphasis>
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<emphasis id="B921EAF05B60856CFA9869F78EB778B1" bold="true" box="[1359,1436,1676,1698]" pageId="2" pageNumber="460">sp. nov.</emphasis>
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