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<mods:title id="F55578E26E2CAA059BC43B9493429C60">Two new species of Aegopsis Burmeister, 1847 (Coleoptera: Scarabaeidae: Dynastinae) from the central Brazilian Cerrado</mods:title>
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<emphasis id="B921EAEDFF86063AFF4CC8D7FAAEC1B7" bold="true" box="[151,421,151,177]" italics="true" pageId="2" pageNumber="177">Aegopsis bolboceridus</emphasis>
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<figureCitation id="136E2A7AFF86063AFF44C8FAFA00C1D5" box="[159,267,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="3.[151,250,1977,1999]" captionStartId-1="4.[151,250,1882,1904]" captionStartId-2="5.[151,250,1876,1898]" captionStartId-3="6.[151,250,1693,1715]" captionStartId-4="7.[151,250,1292,1314]" captionStartId-5="9.[151,250,1775,1797]" captionStartId-6="10.[151,250,1892,1914]" captionStartId-7="11.[151,250,1322,1344]" captionTargetBox-0="[184,1388,203,1935]" captionTargetBox-1="[200,1377,193,1853]" captionTargetBox-2="[231,1354,193,1844]" captionTargetBox-3="[151,1415,201,1637]" captionTargetBox-4="[151,1436,193,1270]" captionTargetBox-5="[152,1421,205,1743]" captionTargetBox-6="[185,1389,202,1867]" captionTargetBox-7="[151,1436,193,1283]" captionTargetId-0="figure@3.[184,1388,203,1935]" captionTargetId-1="figure@4.[200,1377,193,1853]" captionTargetId-2="figure@5.[231,1354,193,1844]" captionTargetId-3="figure@6.[151,1436,193,1672]" captionTargetId-4="figure@7.[151,1436,193,1270]" captionTargetId-5="figure@9.[151,1436,193,1754]" captionTargetId-6="figure@10.[181,1389,193,1871]" captionTargetId-7="figure@11.[151,1436,193,1283]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionText-0="FIGURE 1. Habitus of Aegopsis bolboceridus. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." captionText-1="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." captionText-2="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." captionText-3="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." captionText-4="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." captionText-5="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." captionText-6="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." captionText-7="FIGURE 8. A, Mandible of Aegopsis bolboceridus female in ventral view; B, mandible of Aegopsis diceratops female in ventral view; C, mandible of Aegopsis vazdemelloi female in ventral view; D, mentum of Aegopsis bolboceridus female in ventral view; E, mentum of Aegopsis diceratops female in ventral view; F, mentum of Aegopsis vazdemelloi female in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; arrow indicating middle part of mesal brushes in B; arrow indicating internal lobe in C. Scale bars: 0.5 mm." httpUri-0="https://zenodo.org/record/2611532/files/figure.png" httpUri-1="https://zenodo.org/record/2611534/files/figure.png" httpUri-2="https://zenodo.org/record/2611536/files/figure.png" httpUri-3="https://zenodo.org/record/2611538/files/figure.png" httpUri-4="https://zenodo.org/record/2611541/files/figure.png" httpUri-5="https://zenodo.org/record/2611544/files/figure.png" httpUri-6="https://zenodo.org/record/2611546/files/figure.png" httpUri-7="https://zenodo.org/record/2611548/files/figure.png" pageId="2" pageNumber="177">Figs. 1–8</figureCitation>
, 
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<emphasis id="B921EAEDFF86063AFF4CC942FA9BC01E" box="[151,400,258,280]" italics="true" pageId="2" pageNumber="177">Agacephala bolbocerida</emphasis>
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<bibRefCitation id="EFC44B0EFF86063AFC6BC900FF6AC050" author="Dechambre, R. - P." box="[944,1121,320,342]" pageId="2" pageNumber="177" pagination="173 - 174" refId="ref8804" refString="Dechambre, R. - P. (1999) Une nouvelle espece d' Aegopsis Burmeister, 1847. Revue Francaise d'Entomologie, Nouvelle-Serie, 21, 173 - 174." type="journal article" year="1999">Dechambre 1999</bibRefCitation>
: 174
</treatmentCitation>
; 
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<bibRefCitation id="EFC44B0EFF86063AFB46C900FE6EC050" author="Oliveira, C. M. &amp; Moron, M. A. &amp; Frizzas, M. R." box="[1181,1381,320,342]" pageId="2" pageNumber="177" pagination="324 - 327" refId="ref9705" refString="Oliveira, C. M., Moron, M. A. &amp; Frizzas, M. R. (2008) Aegopsis bolboceridus (Coleoptera: Melolonthidae): an important pest on vegetables and corn in Central Brazil. Florida Entomologist, 91, 324 - 327. https: // doi. org / 10.1653 / 0015 - 4040 (2008) 91 [324: ABCMAI] 2.0. CO; 2" type="journal article" year="2008">
Oliveira 
<emphasis id="B921EAEDFF86063AFB2DC901FE28C050" box="[1270,1315,320,342]" italics="true" pageId="2" pageNumber="177">et al</emphasis>
. 2008
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: 324
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; 
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<bibRefCitation id="EFC44B0EFF86063AFF1CC91EFACBC072" author="Oliveira, C. M. &amp; Frizzas, M. R." box="[199,448,350,372]" pageId="2" pageNumber="177" pagination="1 - 15" refId="ref9651" refString="Oliveira, C. M. &amp; Frizzas, M. R. (2013) Field biology of the beetle Aegopsis bolboceridus in Brazil, with a list of host plants. Journal of Insect Science, 13, 1 - 15. https: // doi. org / 10.1673 / 031.013.4801" type="journal article" year="2013">Oliveira &amp; Frizzas 2013</bibRefCitation>
: 1
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; 
<treatmentCitation id="0AF410EEFF86063AFE38C91EF811C072" author="Neita-Moreno, J. C. &amp; Oliveira, C. M. &amp; Moron, M. A." box="[483,794,350,373]" page="114" pageId="2" pageNumber="177" year="2014">
<bibRefCitation id="EFC44B0EFF86063AFE38C91EF9E3C072" author="Neita-Moreno, J. C. &amp; Oliveira, C. M. &amp; Moron, M. A." box="[483,744,350,373]" pageId="2" pageNumber="177" pagination="114 - 125" refId="ref9453" refString="Neita-Moreno, J. C., Oliveira, C. M. &amp; Moron, M. A. (2014) Larval and pupal descriptions of two Aegopsis species (Coleoptera: Melolonthidae: Dynastinae). Florida Entomologist, 97, 114 - 125. https: // doi. org / 10.1653 / 024.097.0116" type="journal article" year="2014">
Neita-Moreno 
<emphasis id="B921EAEDFF86063AFDA2C920F9ADC072" box="[633,678,350,373]" italics="true" pageId="2" pageNumber="177">et al</emphasis>
. 2014
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: 114
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.
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</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF86063AFF4CC9E1FE30C368" pageId="2" pageNumber="177" type="diagnosis">
<paragraph id="8BEA36FFFF86063AFF4CC9E1FE30C368" blockId="2.[151,1437,417,2026]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFF4CC9E1FA1AC0BC" bold="true" box="[151,273,417,442]" pageId="2" pageNumber="177">Diagnosis.</emphasis>
Males: clypeus subrectangular, apex emarginate (
<figureCitation id="136E2A7AFF86063AFCAEC9E1F8DCC0BF" box="[885,983,417,442]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="2" pageNumber="177">Fig. 2A</figureCitation>
); internal lobe of mandibles apically protruding, mesal brush covering internal lobe laterally (
<figureCitation id="136E2A7AFF86063AFCE4C985F8ABC0D8" box="[831,928,453,478]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="2" pageNumber="177">Fig. 5A</figureCitation>
); mandibles with apical protuberance near external carina (
<figureCitation id="136E2A7AFF86063AFE95C9A9FAA1C304" box="[334,426,489,514]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="2" pageNumber="177">Fig. 5A</figureCitation>
); mentum subrectangular, carina on mentum absent (
<figureCitation id="136E2A7AFF86063AFBDEC9A9FF6BC304" box="[1029,1120,489,514]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="2" pageNumber="177">Fig. 5D</figureCitation>
); labrum concave at middle (
<figureCitation id="136E2A7AFF86063AFF44CA4DFBF7C320" box="[159,252,525,550]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="2" pageNumber="177">Fig. 4A</figureCitation>
); parameres, in lateral view, thinner than apex of phallobase, lateral carina slightly curved basally (
<figureCitation id="136E2A7AFF86063AFAB4CA4DFBB7C34C" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="2" pageNumber="177">Fig. 6A</figureCitation>
). Females: clypeus round, punctures shallow and annular on vertex (
<figureCitation id="136E2A7AFF86063AFC08CA71FF24C34F" box="[979,1071,561,586]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="2" pageNumber="177">Fig. 7A</figureCitation>
); distance from canthi carina to canthi apex almost equal to canthi thickness (
<figureCitation id="136E2A7AFF86063AFD48CA15F9E0C368" box="[659,747,597,622]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="2" pageNumber="177">Fig. 3B</figureCitation>
); sensorial area of maxillary palpus striate (
<figureCitation id="136E2A7AFF86063AFB0ECA15FE25C368" box="[1237,1326,597,622]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="2" pageNumber="177">Fig. 4C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF86063FFF1CCA39FA51C7BC" lastPageId="7" lastPageNumber="182" pageId="2" pageNumber="177" type="description">
<paragraph id="8BEA36FFFF86063AFF1CCA39FF2EC4AC" blockId="2.[151,1437,417,2026]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFF1CCA39F90BC394" bold="true" box="[199,512,633,658]" pageId="2" pageNumber="177">
Male redescription. 
<emphasis id="B921EAEDFF86063AFE6DCA39FAFCC394" bold="true" box="[438,503,633,658]" italics="true" pageId="2" pageNumber="177">Color</emphasis>
:
</emphasis>
Surface lustrous; dorsum, venter, and appendages brown with weak metallic shine (
<figureCitation id="136E2A7AFF86063AFF44CADDFBF0C3B0" box="[159,251,669,694]" captionStart="FIGURE 1" captionStartId="3.[151,250,1977,1999]" captionTargetBox="[184,1388,203,1935]" captionTargetId="figure@3.[184,1388,203,1935]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Aegopsis bolboceridus. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611532/files/figure.png" pageId="2" pageNumber="177">Fig. 1A</figureCitation>
). 
<emphasis id="B921EAEDFF86063AFEC8CADCFA51C3B0" bold="true" box="[275,346,668,694]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFEC8CADCFA5AC3B3" bold="true" box="[275,337,668,693]" italics="true" pageId="2" pageNumber="177">Head</emphasis>
:
</emphasis>
Frons moderately punctate; punctures moderate and small, mixed. Cephalic horns moderately punctate, weakly rugopunctate dorsally on outer side; punctures sparse and small. Clypeus moderately punctate, punctures coalescent basally, becoming sparser and smaller to apex; shape subrectangular, apex truncate (
<figureCitation id="136E2A7AFF86063AFAEECAA5FE84C3F8" box="[1333,1423,741,766]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="2" pageNumber="177">Fig. 2A</figureCitation>
); clypeus strongly concave from frontoclypeal suture to apex. Ocular canthi straight anteriorly, posteriorly projecting, tip round; anterior canthi carina sinuous, apex inconspicuous near external edge of canthi (
<figureCitation id="136E2A7AFF86063AFAE9CB6DFE84C240" box="[1330,1423,813,838]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="2" pageNumber="177">Fig. 3A</figureCitation>
). 
<emphasis id="B921EAEDFF86063AFF4CCB11FA2CC26C" bold="true" box="[151,295,849,874]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFF4CCB11FA15C26C" bold="true" box="[151,286,849,874]" italics="true" pageId="2" pageNumber="177">Mouthparts</emphasis>
:
</emphasis>
Labrum setose, concave at middle, sides round (
<figureCitation id="136E2A7AFF86063AFC50CB11F8E6C26C" box="[907,1005,849,874]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="2" pageNumber="177">Fig. 4A</figureCitation>
). Mandibles subtriangular with two prominent teeth apically and two ventral carinae; in ventral view, apical protuberance near external carina, internal lobe apically protruding, mesal brush covering internal lobe laterally (
<figureCitation id="136E2A7AFF86063AFC6FCBD9FF1BC2B4" box="[948,1040,921,946]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="2" pageNumber="177">Fig. 5A</figureCitation>
). Maxilla with galea subtriangular, teeth absent; maxillary palpomere IV 3.6 times longer than wide; sensorial area densely striated (
<figureCitation id="136E2A7AFF86063AFB09CBFDFE21C2D0" box="[1234,1322,957,982]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="2" pageNumber="177">Fig. 4B</figureCitation>
). Mentum subrectangular, sides strongly convex, carina absent (
<figureCitation id="136E2A7AFF86063AFD33CBA1F84AC2FC" box="[744,833,993,1018]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="2" pageNumber="177">Fig. 5D</figureCitation>
). 
<emphasis id="B921EAEDFF86063AFC8FCBA1F8DFC2FC" bold="true" box="[852,980,993,1018]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFC8FCBA1F8C1C2FC" bold="true" box="[852,970,993,1018]" italics="true" pageId="2" pageNumber="177">Prothorax</emphasis>
:
</emphasis>
Pronotum bordered, disc weakly convex and distinctly punctate. Thoracic horn strong, apex slightly downturned. Puncture size almost uniform on disc, becoming smaller towards thoracic horn apex and denser on sides (
<figureCitation id="136E2A7AFF86063AFC4DCC69F8FBC544" box="[918,1008,1065,1090]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="2" pageNumber="177">Fig. 2B</figureCitation>
). Anterior angles and lateral edges of horn completely wrinkled. 
<taxonomicName id="4C554D7CFF86063AFE17CC0CF9E7C560" authority=": Subtriangular" authorityName="Subtriangular" box="[460,748,1100,1126]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="2" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAEDFF86063AFE17CC0CF941C560" bold="true" box="[460,586,1100,1126]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFE17CC0CF94AC563" bold="true" box="[460,577,1100,1125]" italics="true" pageId="2" pageNumber="177">Scutellum</emphasis>
:
</emphasis>
Subtriangular
</taxonomicName>
in shape, apex weakly projected; punctures large and dense medially and apically, small basally. 
<emphasis id="B921EAEDFF86063AFDE1CC31F980C58C" bold="true" box="[570,651,1137,1162]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFDE1CC31F989C58C" bold="true" box="[570,642,1137,1162]" italics="true" pageId="2" pageNumber="177">Elytra</emphasis>
:
</emphasis>
Form 1.7 times longer than wide. Surface glabrous, densely covered by large ocellate punctures and fine annular punctures, microsculpture slightly wrinkled. Elytral punctures generally irregular, with distinct sutural striae; striae not impressed, punctures small. Humeral umbone weak, apical umbone distinct not crossing elytral external edges in dorsal view. 
<emphasis id="B921EAEDFF86063AFC14CC9EFF04C5F0" bold="true" box="[975,1039,1245,1270]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFC14CC9EFF0EC5F3" bold="true" box="[975,1029,1246,1269]" italics="true" pageId="2" pageNumber="177">Legs</emphasis>
:
</emphasis>
Protibial surface punctate dorsally, with two setose striae. Protibia with three well-developed external teeth posteriorly decreasing in size. 
<emphasis id="B921EAEDFF86063AFAFFCD41FE97C41C" bold="true" box="[1316,1436,1281,1306]" pageId="2" pageNumber="177">
<taxonomicName id="4C554D7CFF86063AFAFFCD41FE99C41C" box="[1316,1426,1281,1306]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="177" phylum="Chordata" rank="genus">
<emphasis id="B921EAEDFF86063AFAFFCD41FE99C41C" bold="true" box="[1316,1426,1281,1306]" italics="true" pageId="2" pageNumber="177">Pygidium</emphasis>
</taxonomicName>
:
</emphasis>
Form 2.6 times wider than long, strongly convex. Disc and posterior edge glabrous, hirsute laterally, densely wrinkled on edges, larger and denser punctures medially. 
<emphasis id="B921EAEDFF86063AFCF7CD09F8ABC464" bold="true" box="[812,928,1353,1378]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFCF7CD09F89CC464" bold="true" box="[812,919,1353,1378]" italics="true" pageId="2" pageNumber="177">Genitalia</emphasis>
:
</emphasis>
Parameres, in dorsal view, as in 
<figureCitation id="136E2A7AFF86063AFAC4CD09FE72C464" box="[1311,1401,1353,1378]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="2" pageNumber="177">Fig. 6C</figureCitation>
; in lateral view, thinner than apex of phallobase, lateral carina slightly curved at base (
<figureCitation id="136E2A7AFF86063AFB9ACD2DFF90C480" box="[1089,1179,1389,1414]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="2" pageNumber="177">Fig. 6A</figureCitation>
); in ventral view, apex round, ventral carina basally acute, inferior edge of parameres concave (
<figureCitation id="136E2A7AFF86063AFC1BCDD1FF13C4AC" box="[960,1048,1425,1450]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="2" pageNumber="177">Fig. 6B</figureCitation>
).
</paragraph>
<paragraph id="8BEA36FFFF86063AFF1CCDF5FE6DC784" blockId="2.[151,1437,417,2026]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFF1CCDF5FA9AC4C8" bold="true" box="[199,401,1461,1486]" pageId="2" pageNumber="177">Variation, males.</emphasis>
Major males with cephalic horns about 2–4 times longer than vertex; body length 
<quantity id="4CAD9B1AFF86063AFAF7CDF5FBC8C4F7" metricMagnitude="-2" metricUnit="m" metricValue="3.01" metricValueMax="3.4799999999999995" metricValueMin="2.54" pageId="2" pageNumber="177" unit="mm" value="30.099999999999998" valueMax="34.8" valueMin="25.4">25.4–34.8 mm</quantity>
(
<emphasis id="B921EAEDFF86063AFF0FCD9AFBE9C4F7" box="[212,226,1498,1521]" italics="true" pageId="2" pageNumber="177">n</emphasis>
= 53) and pronotal width 
<quantity id="4CAD9B1AFF86063AFDCBCD99F9BFC4F7" box="[528,692,1497,1522]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" metricValueMax="1.55" metricValueMin="1.21" pageId="2" pageNumber="177" unit="mm" value="13.8" valueMax="15.5" valueMin="12.1">12.1–15.5 mm</quantity>
(
<emphasis id="B921EAEDFF86063AFD1DCD9AF9DFC4F7" box="[710,724,1498,1521]" italics="true" pageId="2" pageNumber="177">n</emphasis>
= 53), as measured from the apex of the cephalic horns to the apices of the elytra; ocular canthi sometimes posteriorly shorter. Minor males (
<figureCitation id="136E2A7AFF86063AFBDFCDBDFF52C710" box="[1028,1113,1533,1558]" captionStart="FIGURE 1" captionStartId="3.[151,250,1977,1999]" captionTargetBox="[184,1388,203,1935]" captionTargetId="figure@3.[184,1388,203,1935]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Aegopsis bolboceridus. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611532/files/figure.png" pageId="2" pageNumber="177">Fig. 1B</figureCitation>
) with cephalic horns smaller or as long as vertex and frons together; body length 
<quantity id="4CAD9B1AFF86063AFD3DCE61F882C73F" box="[742,905,1569,1594]" metricMagnitude="-2" metricUnit="m" metricValue="2.025" metricValueMax="2.26" metricValueMin="1.7899999999999998" pageId="2" pageNumber="177" unit="mm" value="20.25" valueMax="22.6" valueMin="17.9">17.9–22.6 mm</quantity>
(
<emphasis id="B921EAEDFF86063AFC41CE62F8A3C73F" box="[922,936,1570,1593]" italics="true" pageId="2" pageNumber="177">n</emphasis>
= 14) and pronotal width 
<quantity id="4CAD9B1AFF86063AFB0ACE61FE6EC73F" box="[1233,1381,1569,1594]" metricMagnitude="-2" metricUnit="m" metricValue="1.0399999999999998" metricValueMax="1.16" metricValueMin="0.9199999999999999" pageId="2" pageNumber="177" unit="mm" value="10.399999999999999" valueMax="11.6" valueMin="9.2">9.2–11.6 mm</quantity>
(
<emphasis id="B921EAEDFF86063AFAADCE62FE8FC73F" box="[1398,1412,1570,1593]" italics="true" pageId="2" pageNumber="177">n</emphasis>
= 14); thoracic horn short, tuberculate, not crossing anterior margin of pronotum nor crossing vertex; ocular canthi subtriangular, carina more notable; punctures of pronotum larger and stronger; parameres shorter and broader.
</paragraph>
<paragraph id="8BEA36FFFF86063AFF1CCECDFFBDC7CF" blockId="2.[151,1437,417,2026]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFF1CCECDFA76C7A0" bold="true" box="[199,381,1677,1702]" pageId="2" pageNumber="177">Measurements.</emphasis>
Body length: 
<quantity id="4CAD9B1AFF86063AFDFFCECEF9C7C7A3" box="[548,716,1678,1702]" metricMagnitude="-2" metricUnit="m" metricValue="2.635" metricValueMax="3.4799999999999995" metricValueMin="1.7899999999999998" pageId="2" pageNumber="177" unit="mm" value="26.349999999999998" valueMax="34.8" valueMin="17.9">17.9–34.8 mm</quantity>
. Cephalic horns length: 
<quantity id="4CAD9B1AFF86063AFC3DCECEFF79C7A3" box="[998,1138,1678,1702]" metricMagnitude="-3" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="8.1" metricValueMin="1.2" pageId="2" pageNumber="177" unit="mm" value="4.6499999999999995" valueMax="8.1" valueMin="1.2">1.2–8.1 mm</quantity>
. Elytral length: 
<quantity id="4CAD9B1AFF86063AFAF6CECDFBCDC7CF" metricMagnitude="-2" metricUnit="m" metricValue="1.4949999999999999" metricValueMax="1.85" metricValueMin="1.14" pageId="2" pageNumber="177" unit="mm" value="14.95" valueMax="18.5" valueMin="11.4">11.4–18.5 mm</quantity>
. Elytral width: 
<quantity id="4CAD9B1AFF86063AFEA9CEF1F901C7CF" box="[370,522,1713,1738]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="10.6" metricValueMin="6.8" pageId="2" pageNumber="177" unit="mm" value="8.7" valueMax="10.6" valueMin="6.8">6.8–10.6 mm</quantity>
. Pronotal width: 
<quantity id="4CAD9B1AFF86063AFD12CEF1F86AC7CF" box="[713,865,1713,1738]" metricMagnitude="-2" metricUnit="m" metricValue="1.2349999999999999" metricValueMax="1.55" metricValueMin="0.9199999999999999" pageId="2" pageNumber="177" unit="mm" value="12.35" valueMax="15.5" valueMin="9.2">9.2–15.5 mm</quantity>
. Protibial length: 
<quantity id="4CAD9B1AFF86063AFBF3CEF2FFB9C7CF" box="[1064,1202,1714,1738]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" metricValueMax="9.8" metricValueMin="4.8" pageId="2" pageNumber="177" unit="mm" value="7.3" valueMax="9.8" valueMin="4.8">4.8–9.8 mm</quantity>
.
</paragraph>
<paragraph id="8BEA36FFFF86063AFF1CCE95FE7DC6EC" blockId="2.[151,1437,417,2026]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFF1CCE95FA3BC7E8" bold="true" box="[199,304,1749,1774]" pageId="2" pageNumber="177">Females.</emphasis>
Females are as males in general aspect, except the following (
<figureCitation id="136E2A7AFF86063AFC3ECE95FF37C7E8" box="[997,1084,1749,1774]" captionStart="FIGURE 1" captionStartId="3.[151,250,1977,1999]" captionTargetBox="[184,1388,203,1935]" captionTargetId="figure@3.[184,1388,203,1935]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Aegopsis bolboceridus. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611532/files/figure.png" pageId="2" pageNumber="177">Fig. 1C</figureCitation>
). 
<emphasis id="B921EAEDFF86063AFB8BCE95FF9CC7E8" bold="true" box="[1104,1175,1749,1774]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFB8BCE95FF85C7E8" bold="true" box="[1104,1166,1749,1774]" italics="true" pageId="2" pageNumber="177">Head</emphasis>
:
</emphasis>
Cephalic horns absent; frons with two inconspicuous tubercles (
<figureCitation id="136E2A7AFF86063AFDB6CEB9F9C2C614" box="[621,713,1785,1810]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="2" pageNumber="177">Fig. 7B</figureCitation>
); clypeus round, slightly concave from frontoclypeal suture to apex, punctures larger and denser, especially on clypeus, wrinkles in front of the tubercles (
<figureCitation id="136E2A7AFF86063AFB60CF5DFE13C630" box="[1211,1304,1821,1846]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="2" pageNumber="177">Fig. 7A</figureCitation>
); punctures shallow and annular on vertex. Ocular canthi convex anteriorly, concave posteriorly, tip round; distance from canthi carina to canthi apex almost equal to canthi thickness (
<figureCitation id="136E2A7AFF86063AFCB9CF25F8B7C678" box="[866,956,1893,1918]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="2" pageNumber="177">Fig. 3B</figureCitation>
). 
<emphasis id="B921EAEDFF86063AFC08CF25FF68C678" bold="true" box="[979,1123,1893,1918]" pageId="2" pageNumber="177">
<emphasis id="B921EAEDFF86063AFC08CF25FF51C678" bold="true" box="[979,1114,1893,1918]" italics="true" pageId="2" pageNumber="177">Mouthparts</emphasis>
:
</emphasis>
Labrum convex anteriorly, sides more round. Mandibles and teeth shorter, internal lobe more protruding, mesal brush sparsely covering the internal lobe (
<figureCitation id="136E2A7AFF86063AFEE1CFEDFA9CC6C0" box="[314,407,1965,1990]" captionStart="FIGURE 8" captionStartId="11.[151,250,1322,1344]" captionTargetBox="[151,1436,193,1283]" captionTargetId="figure@11.[151,1436,193,1283]" captionTargetPageId="11" captionText="FIGURE 8. A, Mandible of Aegopsis bolboceridus female in ventral view; B, mandible of Aegopsis diceratops female in ventral view; C, mandible of Aegopsis vazdemelloi female in ventral view; D, mentum of Aegopsis bolboceridus female in ventral view; E, mentum of Aegopsis diceratops female in ventral view; F, mentum of Aegopsis vazdemelloi female in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; arrow indicating middle part of mesal brushes in B; arrow indicating internal lobe in C. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611548/files/figure.png" pageId="2" pageNumber="177">Fig. 8A</figureCitation>
). Maxilla with stipes less hirsute, maxillary palpomere IV three times longer than wide, sensorial area shorter, striae sparser and distinct (
<figureCitation id="136E2A7AFF86063AFD3CCF91F84BC6EC" box="[743,832,2001,2026]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="2" pageNumber="177">Fig. 4C</figureCitation>
). Mentum shorter, more hirsute on apex (
<figureCitation id="136E2A7AFF86063AFAD4CF91FE62C6EC" box="[1295,1385,2001,2026]" captionStart="FIGURE 8" captionStartId="11.[151,250,1322,1344]" captionTargetBox="[151,1436,193,1283]" captionTargetId="figure@11.[151,1436,193,1283]" captionTargetPageId="11" captionText="FIGURE 8. A, Mandible of Aegopsis bolboceridus female in ventral view; B, mandible of Aegopsis diceratops female in ventral view; C, mandible of Aegopsis vazdemelloi female in ventral view; D, mentum of Aegopsis bolboceridus female in ventral view; E, mentum of Aegopsis diceratops female in ventral view; F, mentum of Aegopsis vazdemelloi female in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; arrow indicating middle part of mesal brushes in B; arrow indicating internal lobe in C. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611548/files/figure.png" pageId="2" pageNumber="177">Fig. 8D</figureCitation>
).
</paragraph>
<caption id="DF2A6677FF87063BFF4CCFF9FAC6C6E8" httpUri="https://zenodo.org/record/2611532/files/figure.png" pageId="3" pageNumber="178" startId="3.[151,250,1977,1999]" targetBox="[184,1388,203,1935]" targetPageId="3">
<paragraph id="8BEA36FFFF87063BFF4CCFF9FAC6C6E8" blockId="3.[151,1436,1977,2030]" pageId="3" pageNumber="178">
<emphasis id="B921EAEDFF87063BFF4CCFF9FA1EC6C8" bold="true" box="[151,277,1977,1999]" pageId="3" pageNumber="178">FIGURE 1.</emphasis>
Habitus of 
<taxonomicName id="4C554D7CFF87063BFE4FCFFAF976C6C9" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[404,637,1977,1999]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF87063BFE4FCFFAF976C6C9" box="[404,637,1977,1999]" italics="true" pageId="3" pageNumber="178">Aegopsis bolboceridus</emphasis>
</taxonomicName>
. 
<emphasis id="B921EAEDFF87063BFD57CFF9F995C6C8" bold="true" box="[652,670,1977,1998]" pageId="3" pageNumber="178">A</emphasis>
, Major male in dorsal view; 
<emphasis id="B921EAEDFF87063BFC0FCFFAF8EEC6C9" bold="true" box="[980,997,1978,1999]" pageId="3" pageNumber="178">B</emphasis>
, minor male in dorsal view; 
<emphasis id="B921EAEDFF87063BFAC2CFF9FE20C6C9" bold="true" box="[1305,1323,1977,1999]" pageId="3" pageNumber="178">C</emphasis>
, female in dorsal view. Scale bars: 3 mm.
</paragraph>
</caption>
<caption id="DF2A6677FF80063CFF4CCF1AFA6CC6EC" httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="4" pageNumber="179" startId="4.[151,250,1882,1904]" targetBox="[200,1377,193,1853]" targetPageId="4">
<paragraph id="8BEA36FFFF80063CFF4CCF1AFA6CC6EC" blockId="4.[151,1436,1882,2026]" pageId="4" pageNumber="179">
<emphasis id="B921EAEDFF80063CFF4CCF1AFA24C669" bold="true" box="[151,303,1882,1904]" pageId="4" pageNumber="179">FIGURE 2. A</emphasis>
, Head of 
<taxonomicName id="4C554D7CFF80063CFE43CF1AF98BC676" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[408,640,1882,1904]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF80063CFE43CF1AF98BC676" box="[408,640,1882,1904]" italics="true" pageId="4" pageNumber="179">Aegopsis bolboceridus</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF80063CFCB0CF1AF877C669" bold="true" box="[875,892,1882,1903]" pageId="4" pageNumber="179">B</emphasis>
, pronotum of 
<taxonomicName id="4C554D7CFF80063CFBC9CF1AFFF0C676" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[1042,1275,1882,1904]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF80063CFBC9CF1AFFF0C676" box="[1042,1275,1882,1904]" italics="true" pageId="4" pageNumber="179">Aegopsis bolboceridus</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF80063CFF0DCF38FBE3C688" bold="true" box="[214,232,1912,1934]" pageId="4" pageNumber="179">C</emphasis>
, head of 
<taxonomicName id="4C554D7CFF80063CFE93CF39F91EC688" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[328,533,1912,1934]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF80063CFE93CF39F91EC688" box="[328,533,1912,1934]" italics="true" pageId="4" pageNumber="179">Aegopsis diceratops</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF80063CFD23CF39F800C688" bold="true" box="[760,779,1913,1934]" pageId="4" pageNumber="179">D</emphasis>
, pronotum of 
<taxonomicName id="4C554D7CFF80063CFC46CF39FF60C688" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[925,1131,1912,1934]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF80063CFC46CF39FF60C688" box="[925,1131,1912,1934]" italics="true" pageId="4" pageNumber="179">Aegopsis diceratops</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF80063CFA95CF39FE54C688" bold="true" box="[1358,1375,1913,1934]" pageId="4" pageNumber="179">E</emphasis>
, head of 
<taxonomicName id="4C554D7CFF80063CFF6CCFD8FA9CC6AB" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[183,407,1943,1965]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF80063CFF6CCFD8FA9CC6AB" box="[183,407,1943,1965]" italics="true" pageId="4" pageNumber="179">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF80063CFD48CFD7F9ABC6AA" bold="true" box="[659,672,1943,1964]" pageId="4" pageNumber="179">F</emphasis>
, pronotum of 
<taxonomicName id="4C554D7CFF80063CFC9ACFD8FF2AC6AB" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[833,1057,1943,1965]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="179" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF80063CFC9ACFD8FF2AC6AB" box="[833,1057,1943,1965]" italics="true" pageId="4" pageNumber="179">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
male, in dorsal view. Black arrow indicating emarginate apex of clypeus in 
<emphasis id="B921EAEDFF80063CFDE2CFF6F962C6CD" bold="true" box="[569,617,1974,1995]" pageId="4" pageNumber="179">A, E</emphasis>
; white arrow indicating micropunctures on pronotum in F. Scale bars: 
<emphasis id="B921EAEDFF80063CFAECCFF6FE42C6CD" bold="true" box="[1335,1353,1974,1995]" pageId="4" pageNumber="179">A</emphasis>
, 
<emphasis id="B921EAEDFF80063CFA8DCFF5FE63C6CD" bold="true" box="[1366,1384,1973,1995]" pageId="4" pageNumber="179">C</emphasis>
, 
<emphasis id="B921EAEDFF80063CFAAECFF6FE89C6CD" bold="true" box="[1397,1410,1974,1995]" pageId="4" pageNumber="179">F</emphasis>
, 2 mm; 
<emphasis id="B921EAEDFF80063CFF17CF95FBD6C6EC" bold="true" box="[204,221,2005,2026]" pageId="4" pageNumber="179">B</emphasis>
, 
<emphasis id="B921EAEDFF80063CFF32CF95FBF0C6EC" bold="true" box="[233,251,2005,2026]" pageId="4" pageNumber="179">D</emphasis>
, 
<emphasis id="B921EAEDFF80063CFED3CF95FA12C6EC" bold="true" box="[264,281,2005,2026]" pageId="4" pageNumber="179">E</emphasis>
, 3 mm.
</paragraph>
</caption>
<caption id="DF2A6677FF81063DFF4CCF14FE4CC6E3" httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="5" pageNumber="180" startId="5.[151,250,1876,1898]" targetBox="[231,1354,193,1844]" targetPageId="5">
<paragraph id="8BEA36FFFF81063DFF4CCF14FE4CC6E3" blockId="5.[151,1436,1876,2021]" pageId="5" pageNumber="180">
<emphasis id="B921EAEDFF81063DFF4CCF14FA27C66F" bold="true" box="[151,300,1876,1898]" pageId="5" pageNumber="180">FIGURE 3. A</emphasis>
, Canthus of 
<taxonomicName id="4C554D7CFF81063DFE75CF15F99FC66C" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[430,660,1876,1898]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF81063DFE75CF15F99FC66C" box="[430,660,1876,1898]" italics="true" pageId="5" pageNumber="180">Aegopsis bolboceridus</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF81063DFCACCF14F883C66F" bold="true" box="[887,904,1876,1897]" pageId="5" pageNumber="180">B</emphasis>
, canthus of 
<taxonomicName id="4C554D7CFF81063DFBDECF15FFE0C66C" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[1029,1259,1876,1898]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF81063DFBDECF15FFE0C66C" box="[1029,1259,1876,1898]" italics="true" pageId="5" pageNumber="180">Aegopsis bolboceridus</emphasis>
</taxonomicName>
female, in dorsal view; 
<emphasis id="B921EAEDFF81063DFF0CCF33FBE2C68F" bold="true" box="[215,233,1907,1929]" pageId="5" pageNumber="180">C</emphasis>
, canthus of 
<taxonomicName id="4C554D7CFF81063DFEB0CF33F931C68F" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[363,570,1907,1929]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF81063DFEB0CF33F931C68F" box="[363,570,1907,1929]" italics="true" pageId="5" pageNumber="180">Aegopsis diceratops</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF81063DFCFFCF33F83DC68E" bold="true" box="[804,822,1907,1928]" pageId="5" pageNumber="180">D</emphasis>
, canthus of 
<taxonomicName id="4C554D7CFF81063DFC63CF33FF8CC68F" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[952,1159,1907,1929]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF81063DFC63CF33FF8CC68F" box="[952,1159,1907,1929]" italics="true" pageId="5" pageNumber="180">Aegopsis diceratops</emphasis>
</taxonomicName>
female, in dorsal view; 
<emphasis id="B921EAEDFF81063DFA5ECF33FE9DC68E" bold="true" box="[1413,1430,1907,1928]" pageId="5" pageNumber="180">E</emphasis>
, canthus of 
<taxonomicName id="4C554D7CFF81063DFEDDCFD2FAEAC6A1" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[262,481,1937,1959]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF81063DFEDDCFD2FAEAC6A1" box="[262,481,1937,1959]" italics="true" pageId="5" pageNumber="180">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF81063DFD1FCFD2F9DAC6A1" bold="true" box="[708,721,1938,1959]" pageId="5" pageNumber="180">F</emphasis>
, canthus of 
<taxonomicName id="4C554D7CFF81063DFC95CFD2FF22C6A1" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[846,1065,1937,1959]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="180" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF81063DFC95CFD2FF22C6A1" box="[846,1065,1937,1959]" italics="true" pageId="5" pageNumber="180">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in 
<emphasis id="B921EAEDFF81063DFD40CFF0F9A6C6C3" bold="true" box="[667,685,1968,1989]" pageId="5" pageNumber="180">A</emphasis>
; black arrows indicating canthi carina split in 
<emphasis id="B921EAEDFF81063DFB48CFF0FFAEC6C0" bold="true" box="[1171,1189,1968,1990]" pageId="5" pageNumber="180">C</emphasis>
; black arrow indicating apex conspicuous of canthi carina in 
<emphasis id="B921EAEDFF81063DFDD5CF8FF914C6E2" bold="true" box="[526,543,1999,2020]" pageId="5" pageNumber="180">E</emphasis>
; black arrows indicating external edge of canthi in 
<emphasis id="B921EAEDFF81063DFBF3CF8FFF32C6E2" bold="true" box="[1064,1081,1999,2020]" pageId="5" pageNumber="180">B</emphasis>
, 
<emphasis id="B921EAEDFF81063DFB9ECF8FFF5CC6E2" bold="true" box="[1093,1111,1999,2020]" pageId="5" pageNumber="180">D</emphasis>
, 
<emphasis id="B921EAEDFF81063DFBBFCF8FFF7AC6E2" bold="true" box="[1124,1137,1999,2020]" pageId="5" pageNumber="180">F</emphasis>
. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="DF2A6677FF82063EFF4CCEDDFBE0C66D" httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="6" pageNumber="181" startId="6.[151,250,1693,1715]" targetBox="[151,1415,201,1637]" targetPageId="6">
<paragraph id="8BEA36FFFF82063EFF4CCEDDFBE0C66D" blockId="6.[151,1437,1693,1899]" pageId="6" pageNumber="181">
<emphasis id="B921EAEDFF82063EFF4CCEDDFA26C7B5" bold="true" box="[151,301,1693,1715]" pageId="6" pageNumber="181">FIGURE 4. A</emphasis>
, Labrum of 
<taxonomicName id="4C554D7CFF82063EFE74CEDEF99DC7B5" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[431,662,1693,1715]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF82063EFE74CEDEF99DC7B5" box="[431,662,1693,1715]" italics="true" pageId="6" pageNumber="181">Aegopsis bolboceridus</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF82063EFCA6CEDEF885C7B5" bold="true" box="[893,910,1694,1715]" pageId="6" pageNumber="181">B</emphasis>
, maxillary palpomere IV of 
<taxonomicName id="4C554D7CFF82063EFB6ECEDEFE97C7B5" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[1205,1436,1693,1715]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF82063EFB6ECEDEFE97C7B5" box="[1205,1436,1693,1715]" italics="true" pageId="6" pageNumber="181">Aegopsis bolboceridus</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF82063EFEA5CEFCFA9BC7D4" bold="true" box="[382,400,1724,1746]" pageId="6" pageNumber="181">C</emphasis>
, maxillary palpomere IV of 
<taxonomicName id="4C554D7CFF82063EFD1ACEFDF8A1C7D4" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[705,938,1724,1746]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF82063EFD1ACEFDF8A1C7D4" box="[705,938,1724,1746]" italics="true" pageId="6" pageNumber="181">Aegopsis bolboceridus</emphasis>
</taxonomicName>
female, in dorsal view; 
<emphasis id="B921EAEDFF82063EFB74CEFDFFCAC7D4" bold="true" box="[1199,1217,1725,1746]" pageId="6" pageNumber="181">D</emphasis>
, labrum of 
<taxonomicName id="4C554D7CFF82063EFA9BCEFDFA09C7F7" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF82063EFA9BCEFDFA09C7F7" italics="true" pageId="6" pageNumber="181">Aegopsis diceratops</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF82063EFE25CE9BF904C7F6" bold="true" box="[510,527,1755,1776]" pageId="6" pageNumber="181">E</emphasis>
, maxillary palpomere IV of 
<taxonomicName id="4C554D7CFF82063EFC97CE9BFF14C7F7" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[844,1055,1755,1777]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF82063EFC97CE9BFF14C7F7" box="[844,1055,1755,1777]" italics="true" pageId="6" pageNumber="181">Aegopsis diceratops</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF82063EFAC0CE9BFE23C7F6" bold="true" box="[1307,1320,1755,1776]" pageId="6" pageNumber="181">F</emphasis>
, maxillary palpomere IV of 
<taxonomicName id="4C554D7CFF82063EFE95CEBAF915C609" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[334,542,1785,1807]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF82063EFE95CEBAF915C609" box="[334,542,1785,1807]" italics="true" pageId="6" pageNumber="181">Aegopsis diceratops</emphasis>
</taxonomicName>
female, in dorsal view; 
<emphasis id="B921EAEDFF82063EFCF8CEB9F83FC609" bold="true" box="[803,820,1785,1807]" pageId="6" pageNumber="181">G</emphasis>
, labrum of 
<taxonomicName id="4C554D7CFF82063EFC68CEBAFF9AC609" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[947,1169,1785,1807]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF82063EFC68CEBAFF9AC609" box="[947,1169,1785,1807]" italics="true" pageId="6" pageNumber="181">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF82063EFA59CEBAFE9DC609" bold="true" box="[1410,1430,1786,1807]" pageId="6" pageNumber="181">H</emphasis>
, maxillary palpomere IV of 
<taxonomicName id="4C554D7CFF82063EFE6CCF59F99EC628" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[439,661,1816,1838]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF82063EFE6CCF59F99EC628" box="[439,661,1816,1838]" italics="true" pageId="6" pageNumber="181">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
male, in dorsal view; 
<emphasis id="B921EAEDFF82063EFC5FCF58F885C62B" bold="true" box="[900,910,1816,1837]" pageId="6" pageNumber="181">I</emphasis>
, maxillary palpomere IV of 
<taxonomicName id="4C554D7CFF82063EFB65CF59FE97C628" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1214,1436,1816,1838]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="181" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF82063EFB65CF59FE97C628" box="[1214,1436,1816,1838]" italics="true" pageId="6" pageNumber="181">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
female, in dorsal view. Arrows indicating the middle of the labrum in 
<emphasis id="B921EAEDFF82063EFC82CF77F860C64A" bold="true" box="[857,875,1847,1868]" pageId="6" pageNumber="181">A</emphasis>
, 
<emphasis id="B921EAEDFF82063EFCACCF77F880C64B" bold="true" box="[887,907,1847,1869]" pageId="6" pageNumber="181">G</emphasis>
; arrows indicating sensorial area in 
<emphasis id="B921EAEDFF82063EFB22CF77FE00C64B" bold="true" box="[1273,1291,1847,1869]" pageId="6" pageNumber="181">C</emphasis>
, 
<emphasis id="B921EAEDFF82063EFACCCF77FE2AC64A" bold="true" box="[1303,1313,1847,1868]" pageId="6" pageNumber="181">I</emphasis>
. Scale bars: 0.5 mm.
</paragraph>
</caption>
<caption id="DF2A6677FF83063FFF4CCD4CF80CC49A" httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="7" pageNumber="182" startId="7.[151,250,1292,1314]" targetBox="[151,1436,193,1270]" targetPageId="7">
<paragraph id="8BEA36FFFF83063FFF4CCD4CF80CC49A" blockId="7.[151,1436,1292,1436]" pageId="7" pageNumber="182">
<emphasis id="B921EAEDFF83063FFF4CCD4CFA20C427" bold="true" box="[151,299,1292,1314]" pageId="7" pageNumber="182">FIGURE 5. A</emphasis>
, Mandible of 
<taxonomicName id="4C554D7CFF83063FFE67CD4DF9AAC424" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[444,673,1292,1314]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF83063FFE67CD4DF9AAC424" box="[444,673,1292,1314]" italics="true" pageId="7" pageNumber="182">Aegopsis bolboceridus</emphasis>
</taxonomicName>
male, in ventral view; 
<emphasis id="B921EAEDFF83063FFC57CD4CF896C427" bold="true" box="[908,925,1292,1313]" pageId="7" pageNumber="182">B</emphasis>
, mandible of 
<taxonomicName id="4C554D7CFF83063FFBF1CD4DFFFCC424" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1066,1271,1292,1314]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF83063FFBF1CD4DFFFCC424" box="[1066,1271,1292,1314]" italics="true" pageId="7" pageNumber="182">Aegopsis diceratops</emphasis>
</taxonomicName>
male, in ventral view; 
<emphasis id="B921EAEDFF83063FFF0DCD6AFBE3C446" bold="true" box="[214,232,1322,1344]" pageId="7" pageNumber="182">C</emphasis>
, mandible of 
<taxonomicName id="4C554D7CFF83063FFEADCD6BF95AC446" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[374,593,1322,1344]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF83063FFEADCD6BF95AC446" box="[374,593,1322,1344]" italics="true" pageId="7" pageNumber="182">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
male, in ventral view; 
<emphasis id="B921EAEDFF83063FFCE7CD6BF845C446" bold="true" box="[828,846,1323,1344]" pageId="7" pageNumber="182">D</emphasis>
, mentum of 
<taxonomicName id="4C554D7CFF83063FFC0ACD6BFFBCC446" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[977,1207,1322,1344]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF83063FFC0ACD6BFFBCC446" box="[977,1207,1322,1344]" italics="true" pageId="7" pageNumber="182">Aegopsis bolboceridus</emphasis>
</taxonomicName>
male, in ventral view; 
<emphasis id="B921EAEDFF83063FFF4CCD09FBA3C458" bold="true" box="[151,168,1353,1374]" pageId="7" pageNumber="182">E</emphasis>
, mentum of 
<taxonomicName id="4C554D7CFF83063FFEF5CD0AFAF6C459" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[302,509,1353,1375]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF83063FFEF5CD0AFAF6C459" box="[302,509,1353,1375]" italics="true" pageId="7" pageNumber="182">Aegopsis diceratops</emphasis>
</taxonomicName>
male, in ventral view; 
<emphasis id="B921EAEDFF83063FFD34CD09F9F7C458" bold="true" box="[751,764,1353,1374]" pageId="7" pageNumber="182">F</emphasis>
, mentum of 
<taxonomicName id="4C554D7CFF83063FFC5FCD0AFF6BC459" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[900,1120,1353,1375]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="182" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF83063FFC5FCD0AFF6BC459" box="[900,1120,1353,1375]" italics="true" pageId="7" pageNumber="182">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in 
<emphasis id="B921EAEDFF83063FFC1FCD28F8DDC47B" bold="true" box="[964,982,1384,1405]" pageId="7" pageNumber="182">A</emphasis>
; Arrows indicating the internal lobe in 
<emphasis id="B921EAEDFF83063FFABECD28FE7DC47B" bold="true" box="[1381,1398,1384,1405]" pageId="7" pageNumber="182">B</emphasis>
, 
<emphasis id="B921EAEDFF83063FFA58CD28FE9EC478" bold="true" box="[1411,1429,1384,1406]" pageId="7" pageNumber="182">C</emphasis>
; arrow indicating the mentum carina in 
<emphasis id="B921EAEDFF83063FFDFBCDC7F93AC49A" bold="true" box="[544,561,1415,1436]" pageId="7" pageNumber="182">E</emphasis>
. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BEA36FFFF83063FFF4CCD8AFE08C72C" blockId="7.[151,1436,1481,2010]" pageId="7" pageNumber="182">
<emphasis id="B921EAEDFF83063FFF4CCD8AFA1EC4E4" bold="true" box="[151,277,1481,1506]" pageId="7" pageNumber="182">
<emphasis id="B921EAEDFF83063FFF4CCD8AFA07C4E7" bold="true" box="[151,268,1482,1505]" italics="true" pageId="7" pageNumber="182">Pronotum</emphasis>
:
</emphasis>
Weakly convex; prothoracic horn absent; punctures large and dense, wrinkles absent. 
<taxonomicName id="4C554D7CFF83063FFAC5CD89FA0EC703" authority=": Punctures" authorityName="Punctures" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="7" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAEDFF83063FFAC5CD89FE97C4E4" bold="true" box="[1310,1436,1481,1506]" pageId="7" pageNumber="182">
<emphasis id="B921EAEDFF83063FFAC5CD89FE98C4E4" bold="true" box="[1310,1427,1481,1506]" italics="true" pageId="7" pageNumber="182">Scutellum</emphasis>
:
</emphasis>
Punctures
</taxonomicName>
small on base, absent on apex. 
<taxonomicName id="4C554D7CFF83063FFDB7CDACF84DC700" authority=": Convex" authorityName="Convex" box="[620,838,1516,1542]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" pageNumber="182" phylum="Chordata" rank="genus">
<emphasis id="B921EAEDFF83063FFDB7CDACF9E8C700" bold="true" box="[620,739,1516,1542]" pageId="7" pageNumber="182">
<emphasis id="B921EAEDFF83063FFDB7CDACF9D1C703" bold="true" box="[620,730,1516,1541]" italics="true" pageId="7" pageNumber="182">Pygidium</emphasis>
:
</emphasis>
Convex
</taxonomicName>
, straight near apex; hirsute on sides, short bristles on middle; punctures coalescent and dense, merging with wrinkles laterally, punctures annular medially.
</paragraph>
<paragraph id="8BEA36FFFF83063FFF1CCE75FFDEC777" blockId="7.[151,1436,1481,2010]" pageId="7" pageNumber="182">
<emphasis id="B921EAEDFF83063FFF1CCE75F909C748" bold="true" box="[199,514,1589,1614]" pageId="7" pageNumber="182">Measurements of females.</emphasis>
Body length: 
<quantity id="4CAD9B1AFF83063FFD71CE76F858C74B" box="[682,851,1590,1614]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.8899999999999997" metricValueMin="1.91" pageId="7" pageNumber="182" unit="mm" value="24.0" valueMax="28.9" valueMin="19.1">19.1–28.9 mm</quantity>
. Elytral length: 
<quantity id="4CAD9B1AFF83063FFBD5CE76FFBDC74B" box="[1038,1206,1589,1614]" metricMagnitude="-2" metricUnit="m" metricValue="1.545" metricValueMax="1.8399999999999999" metricValueMin="1.25" pageId="7" pageNumber="182" unit="mm" value="15.45" valueMax="18.4" valueMin="12.5">12.5–18.4 mm</quantity>
. Elytral width: 7.0– 
<quantity id="4CAD9B1AFF83063FFF4CCE19FBF5C777" box="[151,254,1625,1650]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="7" pageNumber="182" unit="mm" value="10.6">10.6 mm</quantity>
. Head length: 
<quantity id="4CAD9B1AFF83063FFE44CE19F922C777" box="[415,553,1625,1650]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="4.1" metricValueMin="2.5" pageId="7" pageNumber="182" unit="mm" value="3.3" valueMax="4.1" valueMin="2.5">2.5–4.1 mm</quantity>
. Pronotal width: 
<quantity id="4CAD9B1AFF83063FFD32CE19F88AC777" box="[745,897,1625,1650]" metricMagnitude="-2" metricUnit="m" metricValue="1.145" metricValueMax="1.35" metricValueMin="0.94" pageId="7" pageNumber="182" unit="mm" value="11.45" valueMax="13.5" valueMin="9.4">9.4–13.5 mm</quantity>
. Protibial length: 
<quantity id="4CAD9B1AFF83063FFB9CCE19FFDAC777" box="[1095,1233,1625,1650]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" metricValueMax="7.7" metricValueMin="5.1" pageId="7" pageNumber="182" unit="mm" value="6.4" valueMax="7.7" valueMin="5.1">5.1–7.7 mm</quantity>
.
</paragraph>
<paragraph id="8BEA36FFFF83063FFF1CCE3DFA51C7BC" blockId="7.[151,1436,1481,2010]" pageId="7" pageNumber="182">
<emphasis id="B921EAEDFF83063FFF1CCE3DFAF9C790" bold="true" box="[199,498,1661,1686]" pageId="7" pageNumber="182">Geographic distribution.</emphasis>
<materialsCitation id="3B3D3CA2FF83063FFE27CE3DF80EC790" ID-GBIF-Occurrence="2029162033" box="[508,773,1661,1686]" country="Brazil" location="Goias" pageId="7" pageNumber="182" specimenCount="1" stateProvince="Goias">
<collectingCountry id="F342766FFF83063FFE27CE3DF94EC790" box="[508,581,1661,1686]" name="Brazil" pageId="7" pageNumber="182">Brazil</collectingCountry>
: 
<collectingRegion id="4991F81DFF83063FFD89CE3DF99DC790" box="[594,662,1661,1686]" country="Brazil" name="Goias" pageId="7" pageNumber="182">Goiás</collectingRegion>
, southern
</materialsCitation>
<collectingRegion id="4991F81DFF83063FFCCBCE3DF8A7C790" box="[784,940,1661,1686]" country="Brazil" name="Minas Gerais" pageId="7" pageNumber="182">Minas Gerais</collectingRegion>
, 
<collectingRegion id="4991F81DFF83063FFC61CE3EFFA3C790" box="[954,1192,1661,1686]" country="Brazil" name="Mato Grosso do Sul" pageId="7" pageNumber="182">Mato Grosso do Sul</collectingRegion>
, 
<collectingRegion id="4991F81DFF83063FFB6DCE3EFE0EC790" box="[1206,1285,1662,1686]" country="Brazil" name="Parana" pageId="7" pageNumber="182">Paraná</collectingRegion>
, and Distrito Federal (
<figureCitation id="136E2A7AFF83063FFF21CEE1FA47C7BC" box="[250,332,1697,1722]" captionStart="FIGURE 11" captionStartId="14.[151,250,1333,1355]" captionTargetBox="[150,1437,193,1311]" captionTargetId="figure@14.[150,1437,193,1311]" captionTargetPageId="14" captionText="FIGURE 11. Distribution map of the species in the Aegopsis bolboceridus species group." httpUri="https://zenodo.org/record/2611555/files/figure.png" pageId="7" pageNumber="182">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF830630FF1CCE85FF76C01A" lastPageId="8" lastPageNumber="183" pageId="7" pageNumber="182" type="materials_examined">
<paragraph id="8BEA36FFFF830630FF1CCE85FF76C01A" blockId="7.[151,1436,1481,2010]" lastBlockId="8.[151,1437,151,536]" lastPageId="8" lastPageNumber="183" pageId="7" pageNumber="182">
<emphasis id="B921EAEDFF83063FFF1CCE85F92CC7D8" bold="true" box="[199,551,1733,1758]" pageId="7" pageNumber="182">
Material examined. 
<materialsCitation id="3B3D3CA2FF83063FFE61CE85F92CC7D8" ID-GBIF-Occurrence="2029161553" box="[442,551,1733,1758]" collectionCode="MNHN" pageId="7" pageNumber="182" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EE885DFF83063FFE61CE85F92CC7D8" box="[442,551,1733,1758]" pageId="7" pageNumber="182" type="holotype">Holotype</typeStatus>
</materialsCitation>
</emphasis>
male (MNHN) labeled: a) “Bias”; b) “Ex-Musaeo / JAMES THOMSON”; c) “Bolbocerida / Brés int Thoms / Type”; d) MUSÉUM PARIS / 1952 / COLL R. OBERTHUR”; e) “ 
<typeStatus id="54EE885DFF83063FFAD6CEAAFE87C604" box="[1293,1420,1770,1794]" pageId="7" pageNumber="182" type="lectotype">Lectotypus</typeStatus>
/ Agaocephala bolbocerida / Thomson”; f) “ 
<materialsCitation id="3B3D3CA2FF830630FDA0CF4EFF76C01A" ID-GBIF-Occurrence="2029162537" collectingDate="1942-12" collectingDateMax="2015-10" collectingDateMin="1942-12" collectionCode="CERPE, EPGC, UFRN" collectorName="Luz &amp; Mortos &amp; Arroz', P. &amp; Distrito &amp; Federal &amp; Planaltina &amp; Nucleo Rural Taquara &amp; Chacara &amp; C. M. Oliveira &amp; Brasilia &amp; Asa Norte &amp; Embrapa &amp; M. O. Breda &amp; Distrito Federal &amp; Formosa &amp; J. Ampessar &amp; Ponta Grossa &amp; Lageado &amp; Dourados &amp; L. R. Moraes &amp; Lavras &amp; Silvio Gusmao &amp; Guara" country="Brazil" elevation="1100" lastPageId="8" lastPageNumber="183" location="Brasilia" pageId="7" pageNumber="182" specimenCount="165" specimenCount-female="53" specimenCount-male="112" stateProvince="Goias" typeStatus="lectotype">
<typeStatus id="54EE885DFF83063FFDA0CF4EF828C620" box="[635,803,1806,1830]" pageId="7" pageNumber="182" type="lectotype">LECTOTYPE</typeStatus>
”; g) “MNHN / EC2464”. 
<emphasis id="B921EAEDFF83063FFB95CF4DFE1CC620" bold="true" box="[1102,1303,1805,1830]" pageId="7" pageNumber="182">Other specimens</emphasis>
: 
<specimenCount id="9D53FD76FF83063FFAF3CF4EFE97C620" box="[1320,1436,1805,1830]" pageId="7" pageNumber="182" type="male">100 males</specimenCount>
and 
<specimenCount id="9D53FD76FF83063FFF1CCF71FA49C64C" box="[199,322,1841,1866]" pageId="7" pageNumber="182" type="female">50 females</specimenCount>
(
<collectionCode id="ED44AE3AFF83063FFE89CF72FAA5C64C" box="[338,430,1842,1866]" pageId="7" pageNumber="182">CERPE</collectionCode>
, 
<collectionCode id="ED44AE3AFF83063FFE62CF72F909C64C" box="[441,514,1842,1866]" pageId="7" pageNumber="182">EPGC</collectionCode>
) labeled: “ 
<collectingCountry id="F342766FFF83063FFDA6CF72F9EDC64C" box="[637,742,1842,1866]" name="Brazil" pageId="7" pageNumber="182">BRASIL</collectingCountry>
, DF, 
<location id="8E8A6024FF83063FFCF9CF71F877C64C" LSID="urn:lsid:plazi:treatment:03FC87E9FF860630FF4CC8D7FE6BC31E:8E8A6024FF83063FFCF9CF71F877C64C" box="[802,892,1841,1866]" country="Brazil" name="Brasilia" pageId="7" pageNumber="182" stateProvince="Goias">Brasília</location>
, / 
<quantity id="4CAD9B1AFF83063FFC4CCF72F8E5C64F" box="[919,1006,1842,1866]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="182" unit="m" value="1100.0">
<elevation id="0078D1CCFF83063FFC4CCF72F8E5C64F" box="[919,1006,1842,1866]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="182" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, 
<date id="FFEB103FFF83063FFC20CF71FF76C64C" box="[1019,1149,1841,1866]" pageId="7" pageNumber="182" value="1988-12-23">
<collectingDate id="EFAFE9D7FF83063FFC20CF71FF76C64C" box="[1019,1149,1841,1866]" pageId="7" pageNumber="182" value="1988-12-23">23.xii.1988</collectingDate>
</date>
, / 
<collectorName id="26A05329FF83063FFB42CF72FFCCC64F" box="[1177,1223,1842,1865]" pageId="7" pageNumber="182">Luz</collectorName>
, ‘ 
<collectorName id="26A05329FF83063FFB03CF72FE25C64C" box="[1240,1326,1842,1866]" pageId="7" pageNumber="182">Mortos</collectorName>
em palha de / 
<collectorName id="26A05329FF83063FFF17CF16FA31C66B" box="[204,314,1878,1902]" pageId="7" pageNumber="182">Arroz’, P.</collectorName>
&amp; E. Grossi Legs.”; 
<specimenCount id="9D53FD76FF83063FFDEBCF16F98BC668" box="[560,640,1877,1902]" pageId="7" pageNumber="182" type="male">1 male</specimenCount>
, 
<specimenCount id="9D53FD76FF83063FFD55CF16F9FAC668" box="[654,753,1877,1902]" pageId="7" pageNumber="182" type="female">1 female</specimenCount>
(
<collectionCode id="ED44AE3AFF83063FFCD8CF16F859C668" box="[771,850,1878,1902]" pageId="7" pageNumber="182">EPGC</collectionCode>
): “ 
<collectingCountry id="F342766FFF83063FFCABCF16F8D2C668" box="[880,985,1878,1902]" name="Brazil" pageId="7" pageNumber="182">BRASIL</collectingCountry>
, 
<collectorName id="26A05329FF83063FFC3DCF15FF30C668" box="[998,1083,1877,1902]" pageId="7" pageNumber="182">Distrito</collectorName>
/ 
<collectorName id="26A05329FF83063FFB8DCF15FFA6C668" box="[1110,1197,1877,1902]" pageId="7" pageNumber="182">Federal</collectorName>
, 
<collectorName id="26A05329FF83063FFB61CF15FE26C668" box="[1210,1325,1877,1902]" pageId="7" pageNumber="182">Planaltina</collectorName>
, / 
<collectorName id="26A05329FF83063FFA90CF15FA31C697" pageId="7" pageNumber="182">Núcleo Rural Taquara</collectorName>
, / 
<collectorName id="26A05329FF83063FFE8ECF39FABAC694" box="[341,433,1913,1938]" pageId="7" pageNumber="182">Chácara</collectorName>
70, 
<date id="FFEB103FFF83063FFE38CF39F96EC694" box="[483,613,1913,1938]" pageId="7" pageNumber="182" value="2005-07-10">
<collectingDate id="EFAFE9D7FF83063FFE38CF39F96EC694" box="[483,613,1913,1938]" pageId="7" pageNumber="182" value="2005-07-10">10.vii.2005</collectingDate>
</date>
, / ‘coleta manual em raiz de hortaliças’, / 
<collectorName id="26A05329FF83063FFB9ACF3AFFEDC694" box="[1089,1254,1913,1938]" pageId="7" pageNumber="182">C. M. Oliveira</collectorName>
col.”; 
<specimenCount id="9D53FD76FF83063FFAE8CF3AFE89C694" box="[1331,1410,1913,1938]" pageId="7" pageNumber="182" type="male">1 male</specimenCount>
, 
<specimenCount id="9D53FD76FF83063FFA55CF3AFBE8C6B0" pageId="7" pageNumber="182" type="female">1 female</specimenCount>
(
<collectionCode id="ED44AE3AFF83063FFF2CCFDEFA4DC6B0" box="[247,326,1950,1974]" pageId="7" pageNumber="182">EPGC</collectionCode>
): “ 
<collectingCountry id="F342766FFF83063FFEBCCFDEFADBC6B0" box="[359,464,1950,1974]" name="Brazil" pageId="7" pageNumber="182">BRASIL</collectingCountry>
, DF, 
<collectorName id="26A05329FF83063FFDCECFDDF967C6B0" box="[533,620,1949,1974]" pageId="7" pageNumber="182">Brasília</collectorName>
/ 
<collectorName id="26A05329FF83063FFD57CFDEF80CC6B3" box="[652,775,1950,1974]" pageId="7" pageNumber="182">Asa Norte</collectorName>
, 
<date id="FFEB103FFF83063FFCCCCFDDF86FC6B0" box="[791,868,1949,1974]" pageId="7" pageNumber="182" value="2015-10">
<collectingDate id="EFAFE9D7FF83063FFCCCCFDDF86FC6B0" box="[791,868,1949,1974]" pageId="7" pageNumber="182" value="2015-10">x.2015</collectingDate>
</date>
/ 
<collectorName id="26A05329FF83063FFC5FCFDDF8E2C6B0" box="[900,1001,1949,1974]" pageId="7" pageNumber="182">Embrapa</collectorName>
CENARGEN, 
<collectorName id="26A05329FF83063FFB7ACFDEFE3BC6B0" box="[1185,1328,1949,1974]" pageId="7" pageNumber="182">M.O. Breda</collectorName>
”; 
<specimenCount id="9D53FD76FF83063FFA90CFDEFE97C6B0" box="[1355,1436,1949,1974]" pageId="7" pageNumber="182" type="male">1 male</specimenCount>
(
<collectionCode id="ED44AE3AFF83063FFF44CF82FBE5C6DC" box="[159,238,1986,2010]" pageId="7" pageNumber="182">EPGC</collectionCode>
): “ 
<collectingCountry id="F342766FFF83063FFECBCF82FA72C6DC" box="[272,377,1986,2010]" name="Brazil" pageId="7" pageNumber="182">BRASIL</collectingCountry>
, 
<collectorName id="26A05329FF83063FFE51CF81F948C6DC" box="[394,579,1985,2010]" pageId="7" pageNumber="182">Distrito Federal</collectorName>
, / 
<taxonomicName id="4C554D7CFF83063FFDB2CF81F8CDC6DC" authority=", Embrapa Cerrados" authorityName="Embrapa Cerrados" box="[617,966,1985,2010]" class="Actinopterygii" family="Characidae" genus="Planaltina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Characiformes" pageId="7" pageNumber="182" phylum="Chordata" rank="genus">Planaltina, Embrapa Cerrados</taxonomicName>
, / 
<date id="FFEB103FFF83063FFC37CF81FF6CC6DC" box="[1004,1127,1985,2010]" pageId="7" pageNumber="182" value="2012-11-19">
<collectingDate id="EFAFE9D7FF83063FFC37CF81FF6CC6DC" box="[1004,1127,1985,2010]" pageId="7" pageNumber="182" value="2012-11-19">19.xi.2012</collectingDate>
</date>
, ‘coleta manual’, / 
<collectorName id="26A05329FF830630FA8DCF82FBF3C1B6" lastPageId="8" lastPageNumber="183" pageId="7" pageNumber="182">C. M. Oliveira</collectorName>
”; 
<specimenCount id="9D53FD76FF8C0630FEC8C8D8FA79C1B6" box="[275,370,151,176]" pageId="8" pageNumber="183" type="male">4 males</specimenCount>
, 
<specimenCount id="9D53FD76FF8C0630FE58C8D8FAE2C1B6" box="[387,489,151,176]" pageId="8" pageNumber="183" type="female">1 female</specimenCount>
(
<collectionCode id="ED44AE3AFF8C0630FE25C8D8F946C1B6" box="[510,589,152,176]" pageId="8" pageNumber="183">EPGC</collectionCode>
): “ 
<collectingCountry id="F342766FFF8C0630FDB4C8D8F9DCC1B6" box="[623,727,152,176]" name="Brazil" pageId="8" pageNumber="183">BRASIL</collectingCountry>
, 
<collectingRegion id="4991F81DFF8C0630FD33C8D7F827C1B6" box="[744,812,151,176]" country="Brazil" name="Goias" pageId="8" pageNumber="183">Goiás</collectingRegion>
, / 
<collectorName id="26A05329FF8C0630FC8AC8D8F8BDC1A9" box="[849,950,152,175]" pageId="8" pageNumber="183">Formosa</collectorName>
, / 
<date id="FFEB103FFF8C0630FC00C8D7FF5EC1B6" box="[987,1109,151,176]" pageId="8" pageNumber="183" value="1996-11-01">
<collectingDate id="EFAFE9D7FF8C0630FC00C8D7FF5EC1B6" box="[987,1109,151,176]" pageId="8" pageNumber="183" value="1996-11-01">01.xi.1996</collectingDate>
</date>
, 
<collectorName id="26A05329FF8C0630FBBDC8D8FFFEC1B6" box="[1126,1269,152,176]" pageId="8" pageNumber="183">J. Ampessar</collectorName>
leg.”; 
<specimenCount id="9D53FD76FF8C0630FA90C8D8FE97C1B6" box="[1355,1436,151,176]" pageId="8" pageNumber="183" type="male">1 male</specimenCount>
(
<collectionCode id="ED44AE3AFF8C0630FF44C8FCFBF6C1D2" box="[159,253,188,212]" pageId="8" pageNumber="183">CERPE</collectionCode>
): “ 
<collectingCountry id="F342766FFF8C0630FEC0C8FCFA8FC1D2" box="[283,388,188,212]" name="Brazil" pageId="8" pageNumber="183">BRASIL</collectingCountry>
, 
<collectingRegion id="4991F81DFF8C0630FE4BC8FCFAD4C1D2" box="[400,479,188,212]" country="Brazil" name="Parana" pageId="8" pageNumber="183">Paraná</collectingRegion>
, / 
<collectorName id="26A05329FF8C0630FE20C8FDF99FC1D2" box="[507,660,188,212]" pageId="8" pageNumber="183">Ponta Grossa</collectorName>
, / 
<collectorName id="26A05329FF8C0630FD74C8FCF819C1D3" box="[687,786,188,213]" pageId="8" pageNumber="183">Lageado</collectorName>
, 
<date id="FFEB103FFF8C0630FCC4C8FCF88BC1D3" box="[799,896,188,213]" pageId="8" pageNumber="183" value="1942-12">
<collectingDate id="EFAFE9D7FF8C0630FCC4C8FCF88BC1D3" box="[799,896,188,213]" pageId="8" pageNumber="183" value="1942-12">xii.1942</collectingDate>
</date>
”; 
<specimenCount id="9D53FD76FF8C0630FC43C8FCF8EEC1D3" box="[920,997,188,213]" pageId="8" pageNumber="183" type="male">1 male</specimenCount>
(
<collectionCode id="ED44AE3AFF8C0630FC2FC8FCFF59C1D2" box="[1012,1106,188,212]" pageId="8" pageNumber="183">CERPE</collectionCode>
): “ 
<collectingCountry id="F342766FFF8C0630FBABC8FCFFD2C1D2" box="[1136,1241,188,212]" name="Brazil" pageId="8" pageNumber="183">BRASIL</collectingCountry>
, 
<collectingRegion id="4991F81DFF8C0630FB3EC8FDFBB4C1FE" country="Brazil" name="Mato Grosso do Sul" pageId="8" pageNumber="183">Mato Grosso do Sul</collectingRegion>
, / 
<collectorName id="26A05329FF8C0630FF39C89FFA59C1FE" box="[226,338,223,248]" pageId="8" pageNumber="183">Dourados</collectorName>
, 
<date id="FFEB103FFF8C0630FEB9C89FFAB2C1FE" box="[354,441,223,248]" pageId="8" pageNumber="183" value="2000-11">
<collectingDate id="EFAFE9D7FF8C0630FEB9C89FFAB2C1FE" box="[354,441,223,248]" pageId="8" pageNumber="183" value="2000-11">xi.2000</collectingDate>
</date>
, / 
<collectorName id="26A05329FF8C0630FE07C8A0F972C1F1" box="[476,633,224,247]" pageId="8" pageNumber="183">L. R. Moraes</collectorName>
leg.”; 
<specimenCount id="9D53FD76FF8C0630FD15C8A0F816C1FE" box="[718,797,223,248]" pageId="8" pageNumber="183" type="male">1 male</specimenCount>
(
<collectionCode id="ED44AE3AFF8C0630FCEAC8A0F88BC1FE" box="[817,896,224,248]" pageId="8" pageNumber="183">EPGC</collectionCode>
): “ 
<collectingCountry id="F342766FFF8C0630FC7AC8A0FF01C1FE" box="[929,1034,224,248]" name="Brazil" pageId="8" pageNumber="183">BRASIL</collectingCountry>
, 
<collectingRegion id="4991F81DFF8C0630FBC2C89FFFBCC1FE" box="[1049,1207,223,248]" country="Brazil" name="Minas Gerais" pageId="8" pageNumber="183">Minas Gerais</collectingRegion>
, / 
<collectorName id="26A05329FF8C0630FB01C8A0FE22C1F1" box="[1242,1321,224,247]" pageId="8" pageNumber="183">Lavras</collectorName>
, UFLA, / 
<date id="FFEB103FFF8C0630FF4CC944FA1AC01B" box="[151,273,260,285]" pageId="8" pageNumber="183" value="1992-11-01">
<collectingDate id="EFAFE9D7FF8C0630FF4CC944FA1AC01B" box="[151,273,260,285]" pageId="8" pageNumber="183" value="1992-11-01">01.xi.1992</collectingDate>
</date>
, 
<collectorName id="26A05329FF8C0630FEC7C944FAC8C01A" box="[284,451,260,285]" pageId="8" pageNumber="183">Silvio Gusmão</collectorName>
leg.”; 
<specimenCount id="9D53FD76FF8C0630FDD4C944F96EC01B" box="[527,613,260,285]" pageId="8" pageNumber="183" type="male">2 males</specimenCount>
(
<collectionCode id="ED44AE3AFF8C0630FDAEC944F9CCC01A" box="[629,711,260,284]" country="Brazil" lsid="urn:lsid:biocol.org:col:15771" name="Universidade Federal do Rio Grande do Norte" pageId="8" pageNumber="183" type="Herbarium">UFRN</collectionCode>
): “ 
<collectingCountry id="F342766FFF8C0630FD38C944F847C01A" box="[739,844,260,284]" name="Brazil" pageId="8" pageNumber="183">BRASIL</collectingCountry>
, 
<collectorName id="26A05329FF8C0630FC8CC944FF01C01B" box="[855,1034,260,285]" pageId="8" pageNumber="183">Distrito Federal</collectorName>
, / 
<collectorName id="26A05329FF8C0630FBFFC944FF65C01A" box="[1060,1134,260,284]" pageId="8" pageNumber="183">Guará</collectorName>
”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF8C0630FF1CC967FE6BC31E" pageId="8" pageNumber="183" type="discussion">
<paragraph id="8BEA36FFFF8C0630FF1CC967FE6BC31E" blockId="8.[151,1437,151,536]" pageId="8" pageNumber="183">
<emphasis id="B921EAEDFF8C0630FF1CC967FA30C046" bold="true" box="[199,315,295,320]" pageId="8" pageNumber="183">Remarks.</emphasis>
The redescription of 
<taxonomicName id="4C554D7CFF8C0630FDE7C968F9F9C046" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[572,754,296,320]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8C0630FDE7C968F9F9C046" box="[572,754,296,320]" italics="true" pageId="8" pageNumber="183">A. bolboceridus</emphasis>
</taxonomicName>
is based on specimens collected by P.C.G. and his father, Everardo J. Grossi, in Brasília and additional material from other states in 
<collectingCountry id="F342766FFF8C0630FBCDC90CFF56C063" box="[1046,1117,332,357]" name="Brazil" pageId="8" pageNumber="183">Brazil</collectingCountry>
, after comparison with the 
<typeStatus id="54EE885DFF8C0630FF4CC92FFA09C08E" box="[151,258,367,392]" pageId="8" pageNumber="183" type="lectotype">lectotype</typeStatus>
. The 
<typeStatus id="54EE885DFF8C0630FE97C92FFAB8C08E" box="[332,435,367,392]" pageId="8" pageNumber="183" type="lectotype">lectotype</typeStatus>
is a very small specimen with the genitalia missing (
<bibRefCitation id="EFC44B0EFF8C0630FB93C92FFE9BC08E" author="Dechambre, R. - P. &amp; Grossi, E. J." box="[1096,1424,367,392]" pageId="8" pageNumber="183" pagination="237 - 240" refId="ref8835" refString="Dechambre, R. - P. &amp; Grossi, E. J. (1990) Agaocephala bolbocerida Thomson: nouvelles captures et mise au point systematique. Bulletin de la Societe entomologique de France, 95, 237 - 240." type="journal article" year="1990">Dechambre &amp; Grossi 1990</bibRefCitation>
). However, the other external traits of the 
<typeStatus id="54EE885DFF8C0630FDB8C9D4F9C1C0AB" box="[611,714,404,429]" pageId="8" pageNumber="183" type="lectotype">lectotype</typeStatus>
are shared with small-horned males from Brasília municipality, including body color, pronotal punctation, and ocular canthi shape. Within the species group, 
<taxonomicName id="4C554D7CFF8C0630FB67C9F8FE65C0D6" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[1212,1390,440,464]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF8C0630FB67C9F8FE65C0D6" box="[1212,1390,440,464]" italics="true" pageId="8" pageNumber="183">A. bolboceridus</emphasis>
</taxonomicName>
has the broadest geographic distribution in the Cerrado, despite not occurring in localities where 
<taxonomicName id="4C554D7CFF8C0630FB6EC99CFE40C0F2" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1205,1355,476,500]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF8C0630FB6EC99CFE40C0F2" box="[1205,1355,476,500]" italics="true" pageId="8" pageNumber="183">A. diceratops</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C554D7CFF8C0630FA5FC99CFA17C31E" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="183" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF8C0630FA5FC99CFA17C31E" italics="true" pageId="8" pageNumber="183">A. vazdemelloi</emphasis>
</taxonomicName>
were found. For comparisons among these species, see the Remarks section of both new species.
</paragraph>
</subSubSection>
</treatment>
</document>