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<document id="2594FC6496E47D4E112E36F693660813" ID-CLB-Dataset="31106" ID-DOI="10.11646/zootaxa.4365.1.1" ID-GBIF-Dataset="8e2d27e3-dec0-4ebb-bcd3-dd16e56c8092" ID-ISSN="1175-5326" ID-Zenodo-Dep="1125219" ID-ZooBank="5F132840-6E2F-42E6-8DF2-9EF191DE243C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1513931221510" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Smetana, Aleš" docDate="2017" docId="03FC87DCFFA19D39FF0D3942FD22FF28" docLanguage="en" docName="zootaxa.4365.1.1.pdf.imf" docOrigin="Zootaxa 4365 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthobium conjunctum Shavrin &amp; Smetana 2017, sp.n." docType="treatment" docVersion="8" lastPageNumber="32" masterDocId="FFC5FFA4FFBB9D26FF9A3F56FFB8FFFD" masterDocTitle="A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). I. Gracilipalpe, morchella and nigrum groups" masterLastPageNumber="39" masterPageNumber="1" pageNumber="27" updateTime="1698511183827" updateUser="ExternalLinkService">
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<mods:title id="E9CC9255B5F7B62ADCD369602BBCC577">A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). I. Gracilipalpe, morchella and nigrum groups</mods:title>
</mods:titleInfo>
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<mods:namePart id="0D0C25D8933939189BC9604B89E33D47">Shavrin, Alexey V.</mods:namePart>
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<mods:namePart id="AC9D82FD47D8C4021FB65BD9473F2732">Smetana, Aleš</mods:namePart>
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<mods:typeOfResource id="E64A075ACEE02267A74B43BB883D0C8F">text</mods:typeOfResource>
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<mods:title id="56110EE9797C07E1F243AA0744AF513B">Zootaxa</mods:title>
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<mods:part id="62E426D77468BCB09867614856FA36B0">
<mods:date id="AB12E26D2C39AFC2DE9F634DD57E8690">2017</mods:date>
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<mods:number id="5AFC5B43DD8E8C7820AB08D29D12FD76">2017-12-15</mods:number>
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<mods:number id="D6CD5DAF6D1DAD6FAC11650F98491BB3">4365</mods:number>
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<mods:identifier id="1864459E194DA1E813B1A3846C1ABC1C" type="DOI">10.11646/zootaxa.4365.1.1</mods:identifier>
<mods:identifier id="621891B337E73B65D0AE51F955EA58C9" type="GBIF-Dataset">8e2d27e3-dec0-4ebb-bcd3-dd16e56c8092</mods:identifier>
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<treatment id="03FC87DCFFA19D39FF0D3942FD22FF28" ID-DOI="http://doi.org/10.5281/zenodo.6017660" ID-GBIF-Taxon="139444208" ID-Zenodo-Dep="6017660" LSID="urn:lsid:plazi:treatment:03FC87DCFFA19D39FF0D3942FD22FF28" httpUri="http://treatment.plazi.org/id/03FC87DCFFA19D39FF0D3942FD22FF28" lastPageId="31" lastPageNumber="32" pageId="26" pageNumber="27">
<subSubSection id="C34F6541FFA19D3CFF0D3942FDE1F9D3" box="[151,601,1556,1582]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BEA36CAFFA19D3CFF0D3942FDE1F9D3" blockId="26.[151,601,1556,1615]" box="[151,601,1556,1582]" pageId="26" pageNumber="27">
<heading id="D0A281A6FFA19D3CFF0D3942FDE1F9D3" bold="true" box="[151,601,1556,1582]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B921EAD8FFA19D3CFF0D3942FDE1F9D3" bold="true" box="[151,601,1556,1582]" pageId="26" pageNumber="27">
3.3.2.3.
<taxonomicName id="4C554D49FFA19D3CFF6C3942FDA0F9D3" ID-CoL="67CD9" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[246,536,1556,1582]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="conjunctum" status="sp. nov.">
<emphasis id="B921EAD8FFA19D3CFF6C3942FDA0F9D3" bold="true" box="[246,536,1556,1582]" italics="true" pageId="26" pageNumber="27">Anthobium conjunctum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257A3FFA19D3CFD843942FDE1F9D3" box="[542,601,1556,1582]" pageId="26" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FFA19D3CFF0D3960FE3AF9B2" box="[151,386,1590,1615]" pageId="26" pageNumber="27" type="description">
<paragraph id="8BEA36CAFFA19D3CFF0D3960FE3AF9B2" blockId="26.[151,601,1556,1615]" box="[151,386,1590,1615]" pageId="26" pageNumber="27">
(
<figureCitation id="136E2A4FFFA19D3CFF053960FF44F9B2" box="[159,252,1590,1615]" captionStart="FIGURES 65–69" captionStartId="24.[151,264,1459,1482]" captionTargetBox="[182,1395,210,1407]" captionTargetId="figure@24.[151,1436,193,1438]" captionTargetPageId="24" captionText="FIGURES 65–69. Habitus of Anthobium: 65—A. nigrum (INDIA: Uttarkashi), 66—A. anishchenkoi (paratype), 67—A. conjunctum (holotype), 68—A. densepunctatum (holotype), 69—A. ivani (paratype).Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1125247/files/figure.png" pageId="26" pageNumber="27">Figs. 67</figureCitation>
,
<figureCitation id="136E2A4FFFA19D3CFE923960FE9EF9B2" box="[264,294,1590,1615]" captionStart="FIGURE 76" captionStartId="29.[151,250,906,928]" captionTargetBox="[179,1408,193,884]" captionTargetId="figure@29.[179,1408,193,884]" captionTargetPageId="29" captionText="FIGURE 76. Distribution of Anthobium anishchenkoi (black circles), A. conjunctum (black square) and A. densepunctatum (black triangle) in China." httpUri="https://zenodo.org/record/1125255/files/figure.png" pageId="26" pageNumber="27">76</figureCitation>
,
<figureCitation id="136E2A4FFFA19D3CFEA83961FEC1F9B2" box="[306,377,1591,1615]" captionStart-0="FIGURES 77–78" captionStart-1="FIGURE 79" captionStartId-0="30.[151,264,1431,1454]" captionStartId-1="32.[151,250,1885,1907]" captionTargetBox-0="[194,1401,190,1412]" captionTargetBox-1="[167,1420,193,1864]" captionTargetId-0="figure@30.[151,1435,173,1432]" captionTargetId-1="figure@32.[167,1420,193,1864]" captionTargetPageId-0="30" captionTargetPageId-1="32" captionText-0="FIGURES 77–78. Aedeagusof Anthobium conjunctum (paratype):77—parameral view, 78—lateral view. Scale bar: 0.1mm." captionText-1="FIGURE 79.Type locality of Anthobium conjunctum in the Diancang Shan, Yunnan, China (photograph:M.Schülke)." httpUri-0="https://zenodo.org/record/1125257/files/figure.png" httpUri-1="https://zenodo.org/record/1125259/files/figure.png" pageId="26" pageNumber="27">77–79</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FFA19D3CFF0D3928FD74F84A" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8BEA36CAFFA19D3CFF0D3928FC07F8DA" blockId="26.[151,1436,1662,1975]" pageId="26" pageNumber="27">
<emphasis id="B921EAD8FFA19D3CFF0D3928FE78F96A" bold="true" box="[151,448,1662,1687]" pageId="26" pageNumber="27">
<typeStatus id="54EE8868FFA19D3CFF0D3928FF6AF96A" box="[151,210,1662,1687]" pageId="26" pageNumber="27">Type</typeStatus>
material examined:
</emphasis>
<materialsCitation id="3B3D3C97FFA19D3CFE5C3928FC07F8DA" ID-GBIF-Occurrence="1799096747" collectedFrom="small creek valley, litter and" collectingDate="2007-05-28" collectingMethod="debris sifted" collectionCode="CSC" collectorName=". Schülke’" country="China" elevation="3160" latitude="25.688889" location="Diancang Shan W | Dali" longLatPrecision="21" longitude="100.10333" municipality="Dali Bai Auton. Pref" pageId="26" pageNumber="27" specimenCode="CH07-02 A" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54EE8868FFA19D3CFE5C3928FD96F96A" box="[454,558,1662,1687]" pageId="26" pageNumber="27" type="holotype">Holotype</typeStatus>
<specimenCount id="9D53FD43FFA19D3CFDAF3929FDF2F96B" box="[565,586,1663,1686]" pageId="26" pageNumber="27" type="male">♂</specimenCount>
[plastic plate with aedeagus, abdominal tergite VIII, sternite VIII and apical segment in Canadian balsam was pinned under the plate with beetle]: ‘
<collectingCountry id="F342765AFFA19D3CFC2139F5FBA5F946" box="[955,1053,1699,1723]" name="China" pageId="26" pageNumber="27">CHINA</collectingCountry>
:
<collectingRegion id="4991F828FFA19D3CFBB339F5FBC6F947" box="[1065,1150,1699,1722]" country="China" name="Yunnan" pageId="26" pageNumber="27">Yunnan</collectingRegion>
[
<specimenCode id="DBF39EB1FFA19D3CFB1439F5FAA1F946" box="[1166,1305,1699,1723]" pageId="26" pageNumber="27">CH07-02 A</specimenCode>
], |
<collectingMunicipality id="6B8EACB0FFA19D3CFAA639F4FE9AF922" pageId="26" pageNumber="27">Dali Bai Auton. Pref</collectingMunicipality>
.,
<location id="8E8A6011FFA19D3CFEA93990FDF6F922" LSID="urn:lsid:plazi:treatment:03FC87DCFFA19D39FF0D3942FD22FF28:8E8A6011FFA19D3CFEA93990FDF6F922" box="[307,590,1734,1759]" country="China" latitude="25.688889" longLatPrecision="21" longitude="100.10333" municipality="Dali Bai Auton. Pref" name="Diancang Shan W | Dali" pageId="26" pageNumber="27" stateProvince="Yunnan">Diancang Shan W | Dali</location>
,
<geoCoordinate id="EE61500DFFA19D3CFDC13990FD59F922" box="[603,737,1734,1759]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="25.688889">25°41'20''N</geoCoordinate>
,
<geoCoordinate id="EE61500DFFA19D3CFD743990FCC4F922" box="[750,892,1734,1759]" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="100.10333">100°06'12''E</geoCoordinate>
|
<elevation id="0078D1F9FFA19D3CFC0E3990FC55F923" box="[916,1005,1734,1759]" metricMagnitude="3" metricUnit="m" metricValue="3.16" pageId="26" pageNumber="27" unit="m" value="3160.0">
<quantity id="4CAD9B2FFFA19D3CFC0E3990FC55F923" box="[916,1005,1734,1759]" metricMagnitude="3" metricUnit="m" metricValue="3.16" pageId="26" pageNumber="27" unit="m" value="3160.0">3160 m</quantity>
</elevation>
,
<collectedFrom id="38497537FFA19D3CFC613990FA87F922" box="[1019,1343,1734,1759]" pageId="26" pageNumber="27">small creek valley, litter and</collectedFrom>
|
<collectingMethod id="52144EDDFFA19D3CFAC23990FF60F8FE" pageId="26" pageNumber="27">debris sifted</collectingMethod>
,
<collectingDate id="EFAFE9E2FFA19D3CFF7339BDFEE6F8FE" box="[233,350,1771,1795]" pageId="26" pageNumber="27" value="2007-05-28">
<date id="FFEB100AFFA19D3CFF7339BDFEE6F8FE" box="[233,350,1771,1795]" pageId="26" pageNumber="27" value="2007-05-28">28.V.2007</date>
</collectingDate>
, M
<collectorName id="26A0531CFFA19D3CFE1739BDFDBAF8FE" box="[397,514,1770,1795]" pageId="26" pageNumber="27">. Schülke’</collectorName>
&lt;rectangular label, printed&gt;, ‘HOLOTYPE |
<taxonomicName id="4C554D49FFA19D3CFBBF39BDFB18F8FE" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1061,1184,1771,1795]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B921EAD8FFA19D3CFBBF39BDFB18F8FE" box="[1061,1184,1771,1795]" italics="true" pageId="26" pageNumber="27">Anthobium</emphasis>
</taxonomicName>
|
<taxonomicName id="4C554D49FFA19D3CFB5B39BDFAFCF8FF" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1217,1348,1771,1794]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="conjunctum" status="sp. nov.">
<emphasis id="B921EAD8FFA19D3CFB5B39BDFAFCF8FF" box="[1217,1348,1771,1794]" italics="true" pageId="26" pageNumber="27">conjunctum</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FFA19D3CFACB39BCFA30F8FE" bold="true" box="[1361,1416,1770,1795]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="A21257A3FFA19D3CFACB39BCFA30F8FE" box="[1361,1416,1770,1795]" pageId="26" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A. &amp; Smetana A. 2017’ &lt;red rectangular label, printed&gt; (
<collectionCode id="ED44AE0FFFA19D3CFCE23859FC0BF8DA" box="[888,947,1807,1831]" country="Cuba" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12869" name="Colegio del Sagrado Coraz<61>n" pageId="26" pageNumber="27">CSC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BEA36CAFFA19D3CFF5D3865FD74F84A" blockId="26.[151,1436,1662,1975]" pageId="26" pageNumber="27">
Paratypes (2 specimens): 2 ♂♂: ‘CHINA (Yunnan) | Dali Bai Auton. Pref., | Diancang Shan W Dali | 25°41'20''N/100°06'12''E |
<quantity id="4CAD9B2FFFA19D3CFE563800FD9FF893" box="[460,551,1878,1903]" metricMagnitude="3" metricUnit="m" metricValue="3.16" pageId="26" pageNumber="27" unit="m" value="3160.0">3160 m</quantity>
(small creek valley, | litter and debris sifted |
<date id="FFEB100AFFA19D3CFBC23801FB73F892" box="[1112,1227,1879,1903]" pageId="26" pageNumber="27" value="2007-05-27">27.V.2007</date>
D.W.Wrase [02]’ &lt;rectangular label, printed&gt;, ‘PARATYPE |
<taxonomicName id="4C554D49FFA19D3CFD10382DFCBDF86E" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[650,773,1915,1939]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B921EAD8FFA19D3CFD10382DFCBDF86E" box="[650,773,1915,1939]" italics="true" pageId="26" pageNumber="27">Anthobium</emphasis>
</taxonomicName>
|
<taxonomicName id="4C554D49FFA19D3CFC81382DFC26F86F" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[795,926,1915,1938]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="conjunctum" status="sp. nov.">
<emphasis id="B921EAD8FFA19D3CFC81382DFC26F86F" box="[795,926,1915,1938]" italics="true" pageId="26" pageNumber="27">conjunctum</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FFA19D3CFC3F382CFC65F86E" bold="true" box="[933,989,1914,1939]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="A21257A3FFA19D3CFC3F382CFC65F86E" box="[933,989,1914,1939]" pageId="26" pageNumber="27" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A. &amp; Smetana A. 2017’ &lt;red rectangular label, printed&gt; (1 ♂: CAS; 1 ♂: CSC).
</paragraph>
</subSubSection>
<caption id="DF2A6642FFA09D3DFF0D38F8FCACF819" httpUri="https://zenodo.org/record/1125249/files/figure.png" pageId="27" pageNumber="28" targetBox="[314,1276,204,1934]" targetPageId="27">
<paragraph id="8BEA36CAFFA09D3DFF0D38F8FCACF819" blockId="27.[151,1436,1966,2020]" pageId="27" pageNumber="28">
<emphasis id="B921EAD8FFA09D3DFF0D38F8FEEDF839" bold="true" box="[151,341,1966,1989]" pageId="27" pageNumber="28">FIGURES 70–73.</emphasis>
Aedeagus of
<taxonomicName id="4C554D49FFA09D3DFE7838F9FD19F838" authorityName=": Smetana" authorityYear="2004" box="[482,673,1967,1989]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis id="B921EAD8FFA09D3DFE7838F9FD19F838" box="[482,673,1967,1989]" italics="true" pageId="27" pageNumber="28">Anthobium nigrum</emphasis>
</taxonomicName>
(70–71—INDIA: Uttarkashi) and
<taxonomicName id="4C554D49FFA09D3DFB9E38E6FB1EF838" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1028,1190,1967,1989]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="anishchenkoi">
<emphasis id="B921EAD8FFA09D3DFB9E38E6FB1EF838" box="[1028,1190,1967,1989]" italics="true" pageId="27" pageNumber="28">A. anishchenkoi</emphasis>
</taxonomicName>
(72–73—paratype): 70, 72—parameral view, 71, 73—lateral view. Scale bars: 0.1 mm.
</paragraph>
</caption>
<caption id="DF2A6642FFA79D3AFF0D3C80FA82FC11" box="[151,1338,982,1005]" httpUri="https://zenodo.org/record/1125251/files/figure.png" pageId="28" pageNumber="29" targetBox="[179,1408,193,961]" targetPageId="28">
<paragraph id="8BEA36CAFFA79D3AFF0D3C80FA82FC11" blockId="28.[151,1338,982,1005]" box="[151,1338,982,1005]" pageId="28" pageNumber="29">
<emphasis id="B921EAD8FFA79D3AFF0D3C80FE98FC11" bold="true" box="[151,288,982,1004]" pageId="28" pageNumber="29">FIGURE 74.</emphasis>
Distribution of
<taxonomicName id="4C554D49FFA79D3AFE583C80FD38FC10" authorityName=": Smetana" authorityYear="2004" box="[450,640,982,1005]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis id="B921EAD8FFA79D3AFE583C80FD38FC10" box="[450,640,982,1005]" italics="true" pageId="28" pageNumber="29">Anthobium nigrum</emphasis>
</taxonomicName>
(black circles) and
<taxonomicName id="4C554D49FFA79D3AFCD23C81FC2FFC10" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[840,919,983,1005]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="ivani">
<emphasis id="B921EAD8FFA79D3AFCD23C81FC2FFC10" box="[840,919,983,1005]" italics="true" pageId="28" pageNumber="29">A. ivani</emphasis>
</taxonomicName>
(white circles) in the Himalayan Region.
</paragraph>
</caption>
<caption id="DF2A6642FFA79D3AFF0D38D2FEF3F845" httpUri="https://zenodo.org/record/1125253/files/figure.png" pageId="28" pageNumber="29" targetBox="[157,1429,1054,1902]" targetPageId="28">
<paragraph id="8BEA36CAFFA79D3AFF0D38D2FEF3F845" blockId="28.[151,1437,1924,1976]" pageId="28" pageNumber="29">
<emphasis id="B921EAD8FFA79D3AFF0D38D2FE98F864" bold="true" box="[151,288,1924,1946]" pageId="28" pageNumber="29">FIGURE 75.</emphasis>
Habitat of
<taxonomicName id="4C554D49FFA79D3AFE0F38D2FDECF867" authorityName=": Smetana" authorityYear="2004" box="[405,596,1924,1946]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis id="B921EAD8FFA79D3AFE0F38D2FDECF867" box="[405,596,1924,1946]" italics="true" pageId="28" pageNumber="29">Anthobium nigrum</emphasis>
</taxonomicName>
in the northern India, left tributary of Bhagirathi River, Uttarakhand (photograph: A. Anishchenko).
</paragraph>
</caption>
<caption id="DF2A6642FFA69D3BFF0D3CDCFE2EFC43" httpUri="https://zenodo.org/record/1125255/files/figure.png" pageId="29" pageNumber="30" targetBox="[179,1408,193,884]" targetPageId="29">
<paragraph id="8BEA36CAFFA69D3BFF0D3CDCFE2EFC43" blockId="29.[151,1435,906,958]" pageId="29" pageNumber="30">
<emphasis id="B921EAD8FFA69D3BFF0D3CDCFE9AFC62" bold="true" box="[151,290,906,928]" pageId="29" pageNumber="30">FIGURE 76.</emphasis>
Distribution of
<taxonomicName id="4C554D49FFA69D3BFE513CDCFD70FC5D" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[459,712,906,928]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="anishchenkoi">
<emphasis id="B921EAD8FFA69D3BFE513CDCFD70FC5D" box="[459,712,906,928]" italics="true" pageId="29" pageNumber="30">Anthobium anishchenkoi</emphasis>
</taxonomicName>
(black circles),
<taxonomicName id="4C554D49FFA69D3BFCE93CDCFBB0FC5D" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[883,1032,906,928]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="conjunctum">
<emphasis id="B921EAD8FFA69D3BFCE93CDCFBB0FC5D" box="[883,1032,906,928]" italics="true" pageId="29" pageNumber="30">A. conjunctum</emphasis>
</taxonomicName>
(black square) and
<taxonomicName id="4C554D49FFA69D3BFB433CDCFA23FC5D" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1241,1435,906,928]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="densepunctatum">
<emphasis id="B921EAD8FFA69D3BFB433CDCFA23FC5D" box="[1241,1435,906,928]" italics="true" pageId="29" pageNumber="30">A. densepunctatum</emphasis>
</taxonomicName>
(black triangle) in China.
</paragraph>
</caption>
<subSubSection id="C34F6541FFA69D38FF5D3CBDFE28F8F3" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
<paragraph id="8BEA36CAFFA69D3BFF5D3CBDFB79FBB1" blockId="29.[151,1437,1003,2036]" pageId="29" pageNumber="30">
<emphasis id="B921EAD8FFA69D3BFF5D3CBDFEE9FBF9" bold="true" box="[199,337,1003,1028]" pageId="29" pageNumber="30">Description</emphasis>
. Measurements (n=3): HW: 0.83–0.84, HL: 0.52–0.55; AL (
<typeStatus id="54EE8868FFA69D3BFB9B3CBDFBD2FBF9" box="[1025,1130,1003,1028]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
): 1.66; OL: 0.22–0.23; PL: 0.68–0.71; PW: 1.35–1.38; ESL: 1.63–1.66; EW: 1.93–1.99; AW: 1.65–1.81; MTbL (
<typeStatus id="54EE8868FFA69D3BFB0B3B59FB42FBD5" box="[1169,1274,1039,1064]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
): 0.91; MTrL (
<typeStatus id="54EE8868FFA69D3BFF053B65FEB0FBB1" box="[159,264,1075,1100]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
): 0.42 (MTrL 1–4: 0.30; MTrL 5: 0.12); AedL: 1.01; TL: 3.90–4.20 (
<typeStatus id="54EE8868FFA69D3BFB973B65FBCDFBB1" box="[1037,1141,1075,1100]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
: 4.15).
</paragraph>
<paragraph id="8BEA36CAFFA69D3BFF5D3B01FCA0F9DD" blockId="29.[151,1437,1003,2036]" pageId="29" pageNumber="30">
Body moderately elongated, wide, distinctly convex. Body and antennomeres 4–11 reddish-brown; lateral and basal parts of pronotum and lateral sides of elytra yellow-brown; mouthparts, antennomeres 1–3 and legs yellow. Body glossy; anterior portion of head between antennal insertion and anterior margin of eye with traces of transverse microsculpture, posterio-lateral parts of head behind postocular ridges and neck with fine rugose sculpture; abdomen with distinct isodiametric microsculpture. Head with irregular sparse punctation, markedly denser and deeper on medio-basal portion; middle part of neck with deep and small punctation; pronotum with punctation as that on medio-basal portion of head, sparser and more irregular on medio-basal and latero-apical portions; scutellum with irregular small punctation; punctation of elytra as that in
<taxonomicName id="4C554D49FFA69D3BFBC13A05FAA9FA96" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1115,1297,1363,1387]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="anishchenkoi" status="sp. nov.">
<emphasis id="B921EAD8FFA69D3BFBC13A05FAA9FA96" box="[1115,1297,1363,1387]" italics="true" pageId="29" pageNumber="30">A. anishchenkoi</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FFA69D3BFA863A05FAECFA91" bold="true" box="[1308,1364,1363,1388]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A21257A3FFA69D3BFA863A05FAECFA91" box="[1308,1364,1363,1388]" pageId="29" pageNumber="30" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
Body glabrous; clypeus with irregular long setation; frons and posterior portions of infraorbital ridges with very small setae, middle portion of head between middle length of eyes with a pair of elongated setae; anterior portions of middle elevation of pronotum with two pairs of elongated setae, latero-apical and lateral portions of pronotum with six to nine elongated setae very irregularly arranged; each elytron with very irregular and sparse setation indistinctly longitudinaly arranged. Habitus as in
<figureCitation id="136E2A4FFFA69D3BFD5B3951FCACF9DD" box="[705,788,1543,1568]" captionStart="FIGURES 65–69" captionStartId="24.[151,264,1459,1482]" captionTargetBox="[182,1395,210,1407]" captionTargetId="figure@24.[151,1436,193,1438]" captionTargetPageId="24" captionText="FIGURES 65–69. Habitus of Anthobium: 65—A. nigrum (INDIA: Uttarkashi), 66—A. anishchenkoi (paratype), 67—A. conjunctum (holotype), 68—A. densepunctatum (holotype), 69—A. ivani (paratype).Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1125247/files/figure.png" pageId="29" pageNumber="30">Fig. 67</figureCitation>
.
</paragraph>
<paragraph id="8BEA36CAFFA69D3BFF5D397DFAFBF8E6" blockId="29.[151,1437,1003,2036]" pageId="29" pageNumber="30">
Head 1.5 times as wide as long; middle portion of head slightly elevated or somewhat flattened, with indistinct medio-basal transverse impression; occipital furrow between ocelli moderately wide and deep; grooves in front of ocelli moderately short, reaching posterior third of length of eye; postocular ridges markedly acute, distance between postocular ridge and posterior margin of eye as long as two nearest facets; anterior portion between antennal insertion and anterior margin of eye with wide semicircular notch. Eyes and ocelli as in
<taxonomicName id="4C554D49FFA69D3BFABA39EDFA2DF92E" authorityName=": Smetana" authorityYear="2004" box="[1312,1429,1723,1747]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis id="B921EAD8FFA69D3BFABA39EDFA2DF92E" box="[1312,1429,1723,1747]" italics="true" pageId="29" pageNumber="30">A. nigrum</emphasis>
</taxonomicName>
. Antenna with elongated antennomeres 3–8; measurements of antennomeres (
<typeStatus id="54EE8868FFA69D3BFB9E3989FBD5F905" box="[1028,1133,1759,1784]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
): 1: 0.18 × 0.07; 2: 0.13 × 0.05; 3: 0.16 × 0.05; 4–5: 0.15 × 0.05; 6–7: 0.16 × 0.06; 8: 0.15 × 0.06; 9–10: 0.12 × 0.07; 11: 0.18 × 0.07.
</paragraph>
<paragraph id="8BEA36CAFFA69D3BFF5D387EFEC9F809" blockId="29.[151,1437,1003,2036]" pageId="29" pageNumber="30">
Pronotum about twice as wide as long, 1.6 times as wide as head, widest in middle, more or less gradually narrowed posteriad and anteriad; anterior angles rounded, strongly protruded anteriad; posterior angles obtuse; latero-apical margins with irregular smoothed crenulation; disc of pronotum with very wide middle elevation with indistinct and irregular pair of transverse impressions in medio-basal third, or with moderately wide medio-basal impression uninterrupted in middle (one
<typeStatus id="54EE8868FFA69D3BFDF238E1FD76F832" box="[616,718,1975,1999]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
); lateral portions wide and markedly explanate, each with deep pit at about middle.
</paragraph>
<caption id="DF2A6642FFA59D38FF0D3AC1FA2CFA50" box="[151,1428,1431,1454]" httpUri="https://zenodo.org/record/1125257/files/figure.png" pageId="30" pageNumber="31" targetBox="[194,1401,190,1412]" targetPageId="30">
<paragraph id="8BEA36CAFFA59D38FF0D3AC1FA2CFA50" blockId="30.[151,1428,1431,1454]" box="[151,1428,1431,1454]" pageId="30" pageNumber="31">
<emphasis id="B921EAD8FFA59D38FF0D3AC1FEECFA51" bold="true" box="[151,340,1431,1454]" pageId="30" pageNumber="31">FIGURES 77–78.</emphasis>
Aedeagus of
<taxonomicName id="4C554D49FFA59D38FE453AC1FD72FA53" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[479,714,1431,1454]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="conjunctum">
<emphasis id="B921EAD8FFA59D38FE453AC1FD72FA53" box="[479,714,1431,1454]" italics="true" pageId="30" pageNumber="31">Anthobium conjunctum</emphasis>
</taxonomicName>
(paratype): 77—parameral view, 78—lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BEA36CAFFA59D38FF5D3A82FE71F9A7" blockId="30.[151,1437,1492,2022]" pageId="30" pageNumber="31">Elytra about 1.2 times as wide as long, reaching apical margin of abdominal tergite IV or basal margin of tergite V, more than twice as long as pronotum, with widely rounded shoulders, with widely rounded apical margins truncated at suture; lateral portions moderately wide, markedly explanated; latero-apical margins with small flattened crenulation.</paragraph>
<paragraph id="8BEA36CAFFA59D38FF5D3933FF02F914" blockId="30.[151,1437,1492,2022]" pageId="30" pageNumber="31">
Male. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII slightly sinuate. Aedeagus (
<figureCitation id="136E2A4FFFA59D38FEEE39DFFE7EF95C" box="[372,454,1673,1698]" captionStart="FIGURES 77–78" captionStartId="30.[151,264,1431,1454]" captionTargetBox="[194,1401,190,1412]" captionTargetId="figure@30.[151,1435,173,1432]" captionTargetPageId="30" captionText="FIGURES 77–78. Aedeagusof Anthobium conjunctum (paratype):77—parameral view, 78—lateral view. Scale bar: 0.1mm." httpUri="https://zenodo.org/record/1125257/files/figure.png" pageId="30" pageNumber="31">Fig. 77</figureCitation>
) very narrow, with widely rounded apex of median lobe; parameres slightly exceeding apex of median lobe, with short apical and two preapical setae; internal sac very long. Aedeagus laterally as in
<figureCitation id="136E2A4FFFA59D38FAEA39FBFF0EF914" captionStart="FIGURES 77–78" captionStartId="30.[151,264,1431,1454]" captionTargetBox="[194,1401,190,1412]" captionTargetId="figure@30.[151,1435,173,1432]" captionTargetPageId="30" captionText="FIGURES 77–78. Aedeagusof Anthobium conjunctum (paratype):77—parameral view, 78—lateral view. Scale bar: 0.1mm." httpUri="https://zenodo.org/record/1125257/files/figure.png" pageId="30" pageNumber="31">Fig. 78</figureCitation>
.
</paragraph>
<paragraph id="8BEA36CAFFA59D38FF5D39A3FE28F8F3" blockId="30.[151,1437,1492,2022]" box="[199,400,1781,1806]" pageId="30" pageNumber="31">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C34F6541FFA59D38FF5D384FFC24F887" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="8BEA36CAFFA59D38FF5D384FFC24F887" blockId="30.[151,1437,1492,2022]" pageId="30" pageNumber="31">
<emphasis id="B921EAD8FFA59D38FF5D384FFE0CF8CF" bold="true" box="[199,436,1817,1842]" pageId="30" pageNumber="31">Comparative notes.</emphasis>
<taxonomicName id="4C554D49FFA59D38FE25384FFD70F8CC" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[447,712,1817,1841]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="conjunctum" status="sp. nov.">
<emphasis id="B921EAD8FFA59D38FE25384FFD70F8CC" box="[447,712,1817,1841]" italics="true" pageId="30" pageNumber="31">Anthobium conjunctum</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FFA59D38FD49384FFCB2F8CF" bold="true" box="[723,778,1817,1842]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="A21257A3FFA59D38FD49384FFCB2F8CF" box="[723,778,1817,1842]" pageId="30" pageNumber="31" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
may be distinguished from all remaining species of the
<taxonomicName id="4C554D49FFA59D38FF0D3868FF50F8A8" authorityName=": Smetana" authorityYear="2004" box="[151,232,1854,1877]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis id="B921EAD8FFA59D38FF0D3868FF50F8A8" box="[151,232,1854,1877]" italics="true" pageId="30" pageNumber="31">nigrum</emphasis>
</taxonomicName>
group by anterior angles of the pronotum markedly protruded anteriad, by the elytra markedly wider than long and by details of external and internal structure of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FFA59D38FF5D38D3FA2FF863" box="[199,1431,1925,1950]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BEA36CAFFA59D38FF5D38D3FA2FF863" blockId="30.[151,1437,1492,2022]" box="[199,1431,1925,1950]" pageId="30" pageNumber="31">
<emphasis id="B921EAD8FFA59D38FF5D38D3FED8F863" bold="true" box="[199,352,1925,1950]" pageId="30" pageNumber="31">Distribution.</emphasis>
The new species is known only from one locality (
<figureCitation id="136E2A4FFFA59D38FC0738D3FC56F863" box="[925,1006,1925,1950]" captionStart="FIGURE 76" captionStartId="29.[151,250,906,928]" captionTargetBox="[179,1408,193,884]" captionTargetId="figure@29.[179,1408,193,884]" captionTargetPageId="29" captionText="FIGURE 76. Distribution of Anthobium anishchenkoi (black circles), A. conjunctum (black square) and A. densepunctatum (black triangle) in China." httpUri="https://zenodo.org/record/1125255/files/figure.png" pageId="30" pageNumber="31">Fig. 76</figureCitation>
) in Diancang
<collectingRegion id="4991F828FFA59D38FB1738D3FB7DF863" box="[1165,1221,1925,1950]" country="Myanmar" name="Shan" pageId="30" pageNumber="31">Shan</collectingRegion>
in
<collectingRegion id="4991F828FFA59D38FB7338D0FAFAF860" box="[1257,1346,1926,1949]" country="China" name="Yunnan" pageId="30" pageNumber="31">Yunnan</collectingRegion>
,
<collectingCountry id="F342765AFFA59D38FAD638D3FA2BF863" box="[1356,1427,1925,1950]" name="China" pageId="30" pageNumber="31">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FFA59D38FF5D38FFFD95F818" pageId="30" pageNumber="31" type="biology_ecology">
<paragraph id="8BEA36CAFFA59D38FF5D38FFFD95F818" blockId="30.[151,1437,1492,2022]" pageId="30" pageNumber="31">
<emphasis id="B921EAD8FFA59D38FF5D38FFFEF2F83F" bold="true" box="[199,330,1961,1986]" pageId="30" pageNumber="31">Bionomics.</emphasis>
Specimens were sifted from litter and debris in a small creek valley at elevation
<quantity id="4CAD9B2FFFA59D38FA9C38FFFAE6F83C" box="[1286,1374,1961,1986]" metricMagnitude="3" metricUnit="m" metricValue="3.16" pageId="30" pageNumber="31" unit="m" value="3160.0">3160 m</quantity>
a.s.l. (localities:
<collectingCountry id="F342765AFFA59D38FE8E389BFE84F818" box="[276,316,1997,2021]" name="Switzerland" pageId="30" pageNumber="31">CH</collectingCountry>
07-02A (
<figureCitation id="136E2A4FFFA59D38FE3A389BFE4AF818" box="[416,498,1997,2022]" captionStart="FIGURES 80–83" captionStartId="33.[151,264,1971,1994]" captionTargetBox="[274,1291,206,1932]" captionTargetId="figure@33.[251,1339,193,1957]" captionTargetPageId="33" captionText="FIGURES 80–83. Aedeagus of Anthobium densepunctatum (80–81—paratype), A. ivani (82–83—paratype): 80, 82— parameral view, 81, 83—ateral view. Scale bars:0.1 mm." httpUri="https://zenodo.org/record/1125261/files/figure.png" pageId="30" pageNumber="31">Fig. 81</figureCitation>
), 02)
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FFA49D39FF5D3FC1FD22FF28" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8BEA36CAFFA49D39FF5D3FC1FD22FF28" blockId="31.[151,1436,151,213]" pageId="31" pageNumber="32">
<emphasis id="B921EAD8FFA49D39FF5D3FC1FEF6FF4D" bold="true" box="[199,334,151,176]" pageId="31" pageNumber="32">Etymology.</emphasis>
The specific epithet is the Latin adjective
<emphasis id="B921EAD8FFA49D39FCB13FCEFC1CFF52" box="[811,932,152,175]" italics="true" pageId="31" pageNumber="32">conjunctus</emphasis>
, -
<emphasis id="B921EAD8FFA49D39FC213FCFFC71FF4D" box="[955,969,153,176]" italics="true" pageId="31" pageNumber="32">a</emphasis>
, -
<emphasis id="B921EAD8FFA49D39FC7B3FCFFBBBFF4D" box="[993,1027,153,176]" italics="true" pageId="31" pageNumber="32">um</emphasis>
(connected, related). It confirms the belonging of this species to the
<taxonomicName id="4C554D49FFA49D39FE633FEBFDF2FF29" authorityName=": Smetana" authorityYear="2004" box="[505,586,189,212]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis id="B921EAD8FFA49D39FE633FEBFDF2FF29" box="[505,586,189,212]" italics="true" pageId="31" pageNumber="32">nigrum</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
</treatment>
</document>
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