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<mods:title id="73530D351581C4BFE3FE4B73021DBBE5">Taxonomic reassessment of the Herpetoreas xenura (Wall, 1907) (Squamata: Serpentes: Natricidae) from Myanmar with description of a new species</mods:title>
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<mods:namePart id="CFB7FF85386B2414D0929D84598169CE">Nguyen, Tan Van</mods:namePart>
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<taxonomicName id="4C554D32FFE97E550E17FD5E1354FCBE" authority="Lalremsanga &amp; Biakzuala, 2024" authorityName="Lalremsanga &amp; Biakzuala" authorityYear="2024" box="[630,859,742,769]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="172" phylum="Chordata" rank="species" species="davidi" status="sp. nov.">
<emphasis id="B921EAA3FFE97E550E17FD5E1354FCBE" bold="true" box="[630,859,742,769]" italics="true" pageId="14" pageNumber="172">Herpetoreas davidi</emphasis>
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<figureCitation id="136E2A34FFE97E550ED1FC9412F8FCF7" box="[688,759,813,840]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[203,1384,265,1847]" captionTargetId="figure-11@16.[203,1384,265,1847]" captionTargetPageId="16" captionText="Fig. 5. Herpetoreas davidi sp. nov. AF. Holotype, ♂ (CAS 222969) in preservation. A. Lateral view of the head, right side. B. Lateral view of the head, left side. C. Ventral view of the head. D. Dorsal view of the head. E. General dorsal view. F. General ventral view. GH. Paratype, ♀ (CAS 220256). G. General dorsal view. H. General ventral view. Photos by G. Vogel." figureDoi="http://doi.org/10.5281/zenodo.11112831" httpUri="https://zenodo.org/record/11112831/files/figure.png" pageId="14" pageNumber="172">Fig. 5</figureCitation>
;
<tableCitation id="C6D7030AFFE97E550F65FC94138CFCF8" box="[772,899,813,839]" captionStart-0="Table 1" captionStart-1="Table 2" captionStartId-0="4.[181,250,551,579]" captionStartId-1="11.[180,249,593,621]" captionTargetPageId-0="4" captionTargetPageId-1="11" captionText-0="Table 1 (continued on next page). Main measurements and meristic characters of the Herpetoreas davidi sp. nov., H. xenura (Wall, 1907), and Hebius khasiensis (Boulenger, 1890). Abbreviations: F = adult female; Heb. = Hebius; Her. = Herpetoreas; M= adult male; N/a = not available; SF = subadult female; SM = subadult male." captionText-1="Table 2(continued on next two pages). Comparison of morphological characteristics ofHerpetoreas davidi sp. nov. with those of the genus Herpetoreas Günther, 1860. Abbreviations: Heb. = Hebius; Her. = Herpetoreas; N/a = not available." pageId="14" pageNumber="172">Tables 12</tableCitation>
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<emphasis id="B921EAA3FFE97E550CDCFCD61192FC36" box="[189,413,879,905]" italics="true" pageId="14" pageNumber="172">Amphiesma xenura</emphasis>
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<emphasis id="B921EAA3FFE97E550D80FCC912C5FC36" box="[481,714,880,905]" italics="true" pageId="14" pageNumber="172">Herpetoreas xenura</emphasis>
<bibRefCitation id="EFC44B40FFE97E550EB1FCD6135EFC36" accessDate="2024-04-04" author="Wall F." box="[720,849,879,905]" firstAuthor="Wall" journalOrPublisher="The Journal of the Bombay Natural History Society" pageId="14" pageNumber="172" pagination="337 - 357" part="19" publicationUrl="https://www.biodiversitylibrary.org/page/5449715" refId="ref14630" refString="Wall F. 1907. Notes on snakes from the neighbourhood of Darjeeling. The Journal of the Bombay Natural History Society 19: 337 - 357. Available from https: // www. biodiversitylibrary. org / page / 5449715 [accessed 4 Apr. 2024]." title="Notes on snakes from the neighbourhood of Darjeeling" type="journal article" year="1907">Wall, 1907</bibRefCitation>
)
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<bibRefCitation id="EFC44B40FFE97E550F14FCD61478FC36" author="Wogan G. O. U. &amp; Vindum J. V. &amp; Wilkinson J. A. &amp; Koo M. S. &amp; Slowinski J. B. &amp; Win H. &amp; Thin T. &amp; Kyi S. W. &amp; Oo S. L. &amp; Lwin K. S." box="[885,1143,879,905]" etAl="et al." firstAuthor="Wogan" journalOrPublisher="Hamadryad" page="87" pageId="14" pageNumber="172" pagination="83 - 96" part="33" refId="ref14764" refString="Wogan G. O. U., Vindum J. V., Wilkinson J. A., Koo M. S., Slowinski J. B., Win H., Thin T., Kyi S. W., Oo S. L. &amp; Lwin K. S. 2008. New country records and range extensions for Myanmar amphibians and reptiles. Hamadryad 33: 83 - 96." title="New country records and range extensions for Myanmar amphibians and reptiles" type="journal article" year="2008">
Wogan
<emphasis id="B921EAA3FFE97E550FADFCC91409FC36" box="[972,1030,879,905]" italics="true" pageId="14" pageNumber="172">et al.</emphasis>
2008: 87
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, in part. —
<treatmentCitation id="0AF410A0FFE97E55089EFCD61116FC12" author="Das I." page="335" pageId="14" pageNumber="172" year="2010">
<bibRefCitation id="EFC44B40FFE97E55089EFCD61116FC12" author="Das I." firstAuthor="Das" journalOrPublisher="New Holland Publishers Ltd, London" page="335" pageId="14" pageNumber="172" refId="ref11888" refString="Das I. 2010. A Field Guide to the Reptiles of South-east Asia. New Holland Publishers Ltd, London." title="A Field Guide to the Reptiles of South-east Asia" type="book" year="2010">Das 2010: 335</bibRefCitation>
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, in part. —
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<bibRefCitation id="EFC44B40FFE97E550DC2FC2B1240FC13" author="Reza A." box="[419,591,914,940]" firstAuthor="Reza" journalOrPublisher="Hamadryad" page="64" pageId="14" pageNumber="172" pagination="64 - 72" part="35" refId="ref14308" refString="Reza A. 2010. First record of Amphiesma venningi (Wall, 1910) (Serpentes, Colubridae, Natricinae) from Bangladesh, with notes on its taxonomy, natural history, biogeography and other sympatric species. Hamadryad 35: 64 - 72." title="First record of Amphiesma venningi (Wall, 1910) (Serpentes, Colubridae, Natricinae) from Bangladesh, with notes on its taxonomy, natural history, biogeography and other sympatric species" type="journal article" year="2010">Reza 2010: 64</bibRefCitation>
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, in part. —
<treatmentCitation id="0AF410A0FFE97E550EBBFC2B13E1FC12" author="Wallach V. &amp; Williams K. L. &amp; Boundy J." box="[730,1006,914,941]" page="35" pageId="14" pageNumber="172" year="2014">
<bibRefCitation id="EFC44B40FFE97E550EBBFC2B13E1FC12" author="Wallach V. &amp; Williams K. L. &amp; Boundy J." box="[730,1006,914,941]" etAl="et al." firstAuthor="Wallach" journalOrPublisher="CRC Press, Boca Raton, FL, USA" page="35" pageId="14" pageNumber="172" refId="ref14726" refString="Wallach V., Williams K. L. &amp; Boundy J. 2014. Snakes of the World. A Catalogue of Living and Extinct Species. CRC Press, Boca Raton, FL, USA." title="Snakes of the World. A Catalogue of Living and Extinct Species" type="book" year="2014">
Wallach
<emphasis id="B921EAA3FFE97E550F20FC2A1373FC13" box="[833,892,914,940]" italics="true" pageId="14" pageNumber="172">et al.</emphasis>
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<emphasis id="B921EAA3FFE97E550CDCFC0F11A7FC70" box="[189,424,950,975]" italics="true" pageId="14" pageNumber="172">Herpetoreas xenura</emphasis>
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<bibRefCitation id="EFC44B40FFE97E550DA6FC0F12A0FC70" author="Ren J. L. &amp; Jiang K. &amp; Huang J. J. &amp; David P. &amp; Li J. T." box="[455,687,949,976]" etAl="et al." firstAuthor="Ren" journalOrPublisher="Diversity" page="79" pageId="14" pageNumber="172" pagination="79" part="14" refId="ref14241" refString="Ren J. L., Jiang K., Huang J. J., David P. &amp; Li J. T. 2022. Taxonomic review of the genus Herpetoreas (Serpentes: Natricidae), with the description of a new species from Tibet, China. Diversity 14: 79. https: // doi. org / 10.3390 / d 14020079" title="Taxonomic review of the genus Herpetoreas (Serpentes: Natricidae), with the description of a new species from Tibet, China" type="journal article" year="2022">
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<treatmentCitation id="0AF410A0FFE97E550F5CFC0F140FFC70" author="Uetz et al." box="[829,1024,949,976]" pageId="14" pageNumber="172" year="2023">
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Uetz
<emphasis id="B921EAA3FFE97E550F1CFC0F13B6FC70" box="[893,953,949,975]" italics="true" pageId="14" pageNumber="172">et al.</emphasis>
2023
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: page “
<emphasis id="B921EAA3FFE97E550839FC0F154CFC70" box="[1112,1347,950,975]" italics="true" pageId="14" pageNumber="172">Herpetoreas xenura</emphasis>
”, in part.
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sp.
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<treatmentCitation id="0AF410A0FFE97E550DF2FC4212A3FBA9" author="Deepak V. &amp; Cooper N. &amp; Poyarkov N. A. &amp; Kraus F. &amp; Burin G. &amp; Das A. &amp; Narayanan S. &amp; Streicher J. W. &amp; Smith S. - J. &amp; Gower D. J." box="[403,684,1019,1046]" page="285" pageId="14" pageNumber="172" year="2022">
<bibRefCitation id="EFC44B40FFE97E550DF2FC4212A3FBA9" author="Deepak V. &amp; Cooper N. &amp; Poyarkov N. A. &amp; Kraus F. &amp; Burin G. &amp; Das A. &amp; Narayanan S. &amp; Streicher J. W. &amp; Smith S. - J. &amp; Gower D. J." box="[403,684,1019,1046]" etAl="et al." firstAuthor="Deepak" journalOrPublisher="Zoological Journal of the Linnean Society" page="285" pageId="14" pageNumber="172" pagination="279 - 298" part="1" refId="ref12062" refString="Deepak V., Cooper N., Poyarkov N. A., Kraus F., Burin G., Das A., Narayanan S., Streicher J. W., Smith S. - J. &amp; Gower D. J. 2022. Multilocus phylogeny, natural history traits and classification of natricine snakes (Serpentes: Natricinae). Zoological Journal of the Linnean Society 1: 279 - 298. https: // doi. org / 10.1093 / zoolinnean / zlab 099" title="Multilocus phylogeny, natural history traits and classification of natricine snakes (Serpentes: Natricinae)" type="journal article" year="2022">
Deepak
<emphasis id="B921EAA3FFE97E550D93FC451223FBAA" box="[498,556,1019,1045]" italics="true" pageId="14" pageNumber="172">et al.</emphasis>
2022: 285
</bibRefCitation>
</treatmentCitation>
, in part. —
<treatmentCitation id="0AF410A0FFE97E550F55FC421424FBA9" author="Patel H. &amp; Thackeray T. &amp; Campbell P. D. &amp; Mirza Z. A." box="[820,1067,1019,1046]" page="431" pageId="14" pageNumber="172" year="2023">
<bibRefCitation id="EFC44B40FFE97E550F55FC421424FBA9" author="Patel H. &amp; Thackeray T. &amp; Campbell P. D. &amp; Mirza Z. A." box="[820,1067,1019,1046]" etAl="et al." firstAuthor="Patel" journalOrPublisher="Taxonomy" page="431" pageId="14" pageNumber="172" pagination="415 - 434" part="3" refId="ref13773" refString="Patel H., Thackeray T., Campbell P. D. &amp; Mirza Z. A. 2023. Systematic assessment of Hebius beddomei (Gunther, 1864) (Serpentes: Colubridae: Natricinae) with description of a new genus and a new allied species from the Western Ghats, India. Taxonomy 3: 415 - 434. https: // doi. org / 10.3390 / taxonomy 3030024" title="Systematic assessment of Hebius beddomei (Gunther, 1864) (Serpentes: Colubridae: Natricinae) with description of a new genus and a new allied species from the Western Ghats, India" type="journal article" year="2023">
Patel
<emphasis id="B921EAA3FFE97E550F14FC4513A0FBAA" box="[885,943,1019,1045]" italics="true" pageId="14" pageNumber="172">et al.</emphasis>
2023:
<quantity id="4CAD9B54FFE97E550F9EFC451424FBA9" box="[1023,1067,1020,1046]" metricMagnitude="1" metricUnit="m" metricValue="1.09474" pageId="14" pageNumber="172" unit="in" value="431.0">431</quantity>
</bibRefCitation>
</treatmentCitation>
in supplementary materials.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34F653AFFE97E550CDCFB841480FAE6" pageId="14" pageNumber="172" type="diagnosis">
<paragraph id="8BEA36B1FFE97E550CDCFB841139FBE8" blockId="14.[189,310,1085,1111]" box="[189,310,1085,1111]" pageId="14" pageNumber="172">
<heading id="D0A281DDFFE97E550CDCFB841139FBE8" bold="true" box="[189,310,1085,1111]" fontSize="11" level="3" pageId="14" pageNumber="172" reason="3">
<emphasis id="B921EAA3FFE97E550CDCFB841139FBE8" bold="true" box="[189,310,1085,1111]" pageId="14" pageNumber="172">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA36B1FFE97E550CDCFBD51480FAE6" blockId="14.[188,1400,1131,1369]" pageId="14" pageNumber="172">
<taxonomicName id="4C554D32FFE97E550CDCFBD51191FB3A" authority="Lalremsanga &amp; Biakzuala, 2024" authorityName="Lalremsanga &amp; Biakzuala" authorityYear="2024" box="[189,414,1131,1157]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="172" phylum="Chordata" rank="species" species="davidi" status="sp. nov.">
<emphasis id="B921EAA3FFE97E550CDCFBD51191FB3A" box="[189,414,1131,1157]" italics="true" pageId="14" pageNumber="172">Herpetoreas davidi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257D8FFE97E550DC2FBD511F3FB39" box="[419,508,1132,1158]" pageId="14" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
differs from all other members of the genus
<taxonomicName id="4C554D32FFE97E550F9FFBD5149FFB3A" authorityName="Gunther" authorityYear="1860" box="[1022,1168,1132,1157]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="172" phylum="Chordata" rank="genus">
<emphasis id="B921EAA3FFE97E550F9FFBD5149FFB3A" box="[1022,1168,1132,1157]" italics="true" pageId="14" pageNumber="172">Herpetoreas</emphasis>
</taxonomicName>
by the combination of the following morphological characters: (1) body cylindrical, small size, max. TL
<quantity id="4CAD9B54FFE97E5508D0FB371518FB16" box="[1201,1303,1166,1193]" metricMagnitude="-1" metricUnit="m" metricValue="5.52" pageId="14" pageNumber="172" unit="mm" value="552.0">552 mm</quantity>
; (2) tail relatively long in both sexes, TaL/TL
<geoCoordinate id="EE615076FFE97E550E16FB0B12B6FB73" box="[631,697,1202,1228]" degrees="0.301" direction="north" orientation="latitude" pageId="14" pageNumber="172" precision="55" value="0.301">0.301</geoCoordinate>
<geoCoordinate id="EE615076FFE97E550ED8FB0B1301FB73" box="[697,782,1202,1228]" degrees="0.310" direction="west" orientation="longitude" pageId="14" pageNumber="172" precision="55" value="-0.31">0.310</geoCoordinate>
; (3) dorsal scale rows 191917; all strongly keeled; (4) ventrals 151156; cloacal plate single, subcaudals single; subcaudals 97100; (5) supralabials 9, 4
<superScript id="7C209BF9FFE97E550CADFB4110D6FAB9" attach="none" box="[204,217,1272,1286]" fontSize="6" pageId="14" pageNumber="172">th</superScript>
6th entering orbit; (6) head with a faint off-white sagittal line just behind the parietal; (7) dorsum dark olive-brown or dark brown with indistinct series of pale spots; (8) venter off-whitish, each ventral decorated with a pair of indistinct and faint dark-brown spots on their lateral edges.
</paragraph>
</subSubSection>
<subSubSection id="C34F653AFFE97E550CDCFA3914BCF9AF" pageId="14" pageNumber="172" type="etymology">
<paragraph id="8BEA36B1FFE97E550CDCFA39114BFA25" blockId="14.[189,324,1408,1434]" box="[189,324,1408,1434]" pageId="14" pageNumber="172">
<heading id="D0A281DDFFE97E550CDCFA39114BFA25" bold="true" box="[189,324,1408,1434]" fontSize="11" level="3" pageId="14" pageNumber="172" reason="3">
<emphasis id="B921EAA3FFE97E550CDCFA39114BFA25" bold="true" box="[189,324,1408,1434]" pageId="14" pageNumber="172">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA36B1FFE97E550CDCFA1614BCF9AF" blockId="14.[189,1399,1455,1552]" pageId="14" pageNumber="172">
The species epithet is an eponym in honour of Patrick David (MNHN, Paris,
<collectingCountry id="F3427621FFE97E550832FA1614A8FA76" box="[1107,1191,1455,1481]" name="France" pageId="14" pageNumber="172">France</collectingCountry>
), a French reptile taxonomist, for his contribution to the systematics of snakes of the family
<taxonomicName id="4C554D32FFE97E550852FA6B14A2FA53" authorityName="Bonaparte" authorityYear="1838" box="[1075,1197,1490,1516]" family="Natricidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="172" phylum="Chordata" rank="family">Natricidae</taxonomicName>
and to the Asian snake fauna in general. We suggest “Rakhine Keelback” as its English common name.
</paragraph>
</subSubSection>
<subSubSection id="C34F653AFFE97E540CDCF98E1578FEF8" lastPageId="15" lastPageNumber="173" pageId="14" pageNumber="172" type="materials_examined">
<paragraph id="8BEA36B1FFE97E550CDCF98E1163F9EF" blockId="14.[189,364,1590,1617]" box="[189,364,1590,1617]" pageId="14" pageNumber="172">
<heading id="D0A281DDFFE97E550CDCF98E1163F9EF" bold="true" box="[189,364,1590,1617]" fontSize="11" level="3" pageId="14" pageNumber="172" reason="3">
<emphasis id="B921EAA3FFE97E550CDCF98E1163F9EF" bold="true" box="[189,364,1590,1617]" pageId="14" pageNumber="172">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA36B1FFE97E550C8BF9DC1299F956" blockId="14.[189,1398,1637,1769]" pageId="14" pageNumber="172">
<materialsCitation id="3B3D3CECFFE97E550C8BF9DC1299F956" collectingDate="2002-01-23" collectionCode="CAS" collectorName="G. O. U. Wogan &amp; S. W. Kyi" country="Myanmar" county="Daung Stream" elevation="120" latitude="17.584417" location="Rakhine Yoma Elephant Sanctuary" longLatPrecision="1" longitude="94.681274" municipality="Yoma Elephant Sanctuary" pageId="14" pageNumber="172" specimenCode="CAS 222969" specimenCount="1" specimenCount-male="1" stateProvince="Rakhine State" typeStatus="holotype">
<emphasis id="B921EAA3FFE97E550C8BF9DC1153F9C0" bold="true" box="[234,348,1637,1663]" pageId="14" pageNumber="172">
<typeStatus id="54EE8813FFE97E550C8BF9DC1153F9C0" box="[234,348,1637,1663]" pageId="14" pageNumber="172" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3427621FFE97E550CDCF9311159F91D" box="[189,342,1672,1698]" name="Myanmar" pageId="14" pageNumber="172">MYANMAR</collectingCountry>
<specimenCount id="9D53FD38FFE97E550D19F9301181F91C" box="[376,398,1673,1699]" count="1" pageId="14" pageNumber="172" type="male"></specimenCount>
adult;
<collectingCounty id="628B4E3DFFE97E550D88F9301296F91D" box="[489,665,1672,1699]" pageId="14" pageNumber="172">Daung Stream</collectingCounty>
, within
<location id="8E8A606AFFE97E550F60F93114A7F91D" LSID="urn:lsid:plazi:treatment:03FC87A7FFE97E480E17FD5E1106FB5E:8E8A606AFFE97E550F60F93114A7F91D" box="[769,1192,1672,1699]" country="Myanmar" county="Daung Stream" latitude="17.584417" longLatPrecision="1" longitude="94.681274" municipality="Yoma Elephant Sanctuary" name="Rakhine Yoma Elephant Sanctuary" pageId="14" pageNumber="172" stateProvince="Rakhine State">Rakhine Yoma Elephant Sanctuary</location>
,
<location id="8E8A606AFFE97E5508D8F931157DF91D" LSID="urn:lsid:plazi:treatment:03FC87A7FFE97E480E17FD5E1106FB5E:8E8A606AFFE97E5508D8F931157DF91D" box="[1209,1394,1672,1698]" country="Myanmar" county="Daung Stream" latitude="17.584417" longLatPrecision="1" longitude="94.681274" municipality="Yoma Elephant Sanctuary" name="Gwa Township" pageId="14" pageNumber="172" stateProvince="Rakhine State">Gwa Township</location>
,
<collectingRegion id="4991F853FFE97E550CDCF912116BF979" box="[189,356,1707,1734]" country="Myanmar" name="Rakhine" pageId="14" pageNumber="172">Rakhine State</collectingRegion>
;
<geoCoordinate id="EE615076FFE97E550D10F912125BF979" box="[369,596,1707,1734]" degrees="17.5844166667" direction="north" orientation="latitude" pageId="14" pageNumber="172" precision="1" value="17.584417">17.5844166667° N</geoCoordinate>
,
<geoCoordinate id="EE615076FFE97E550E01F9121330F979" box="[608,831,1707,1734]" degrees="94.6812777778" direction="east" orientation="longitude" pageId="14" pageNumber="172" precision="1" value="94.681274">94.6812777778° E</geoCoordinate>
, WGS 84; ca
<elevation id="0078D182FFE97E550F82F9151461F97A" box="[995,1134,1707,1734]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="172" unit="m" value="120.0">
<quantity id="4CAD9B54FFE97E550F82F9151422F979" box="[995,1069,1708,1734]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="172" unit="m" value="120.0">120 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEB1071FFE97E55081AF915151BF979" box="[1147,1300,1708,1734]" pageId="14" pageNumber="172" value="2002-01-23">
<collectingDate id="EFAFE999FFE97E55081AF915151BF979" box="[1147,1300,1708,1734]" pageId="14" pageNumber="172" value="2002-01-23">23 Jan. 2002</collectingDate>
</date>
;
<collectorName id="26A05367FFE97E550940F9151100F956" pageId="14" pageNumber="172">G.O.U. Wogan</collectorName>
and
<collectorName id="26A05367FFE97E550D28F97611BBF956" box="[329,436,1743,1769]" pageId="14" pageNumber="172">S.W. Kyi</collectorName>
leg.;
<specimenCode id="DBF39ECAFFE97E550D97F976129DF956" box="[502,658,1743,1769]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="172" type="Museum">CAS 222969</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BEA36B1FFE97E540C8BF8A91578FEF8" blockId="14.[188,1401,1808,1976]" lastBlockId="15.[189,1399,265,327]" lastPageId="15" lastPageNumber="173" pageId="14" pageNumber="172">
<materialsCitation id="3B3D3CECFFE97E550C8BF8A91353F808" collectingDate="1926-04-27" collectingDateMax="1926-04-29" collectingDateMin="1926-04-27" collectionCode="CAS" collectorName="J. B. Slowinski &amp; G. O. U. Wogan &amp; H. Win &amp; T. Thin &amp; K. S. Lwin" country="Myanmar" county="Kyat Stream" elevation="175" latitude="17.703888" location="Rakhine Yoma Elephant Sanctuary" longLatPrecision="1" longitude="94.648415" municipality="Yoma Elephant Sanctuary" pageId="14" pageNumber="172" specimenCode="CAS 220256, CAS 220332, CAS 220378" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Rakhine State" typeStatus="paratype">
<emphasis id="B921EAA3FFE97E550C8BF8A91168F895" bold="true" box="[234,359,1808,1834]" pageId="14" pageNumber="172">
<typeStatus id="54EE8813FFE97E550C8BF8A91168F895" box="[234,359,1808,1834]" pageId="14" pageNumber="172" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F3427621FFE97E550CDCF88A1159F8F2" box="[189,342,1843,1869]" name="Myanmar" pageId="14" pageNumber="172">MYANMAR</collectingCountry>
<specimenCount id="9D53FD38FFE97E550D0EF88A1194F8F1" box="[367,411,1843,1870]" count="1" pageId="14" pageNumber="172" type="female">1 ♀</specimenCount>
adult,
<specimenCount id="9D53FD38FFE97E550D8AF88A1218F8F1" box="[491,535,1843,1870]" count="1" pageId="14" pageNumber="172" type="female">1 ♀</specimenCount>
subadult,
<specimenCount id="9D53FD38FFE97E550EF1F88A12B3F8F1" box="[656,700,1843,1870]" count="1" pageId="14" pageNumber="172" type="male">1 ♂</specimenCount>
subadult;
<collectingCounty id="628B4E3DFFE97E550F57F88D13C2F8F2" box="[822,973,1843,1870]" pageId="14" pageNumber="172">Kyat Stream</collectingCounty>
,
<location id="8E8A606AFFE97E550FB9F88A157DF8F2" LSID="urn:lsid:plazi:treatment:03FC87A7FFE97E480E17FD5E1106FB5E:8E8A606AFFE97E550FB9F88A157DF8F2" box="[984,1394,1843,1870]" country="Myanmar" county="Kyat Stream" latitude="17.703888" longLatPrecision="1" longitude="94.648415" municipality="Yoma Elephant Sanctuary" name="Rakhine Yoma Elephant Sanctuary" pageId="14" pageNumber="172" stateProvince="Rakhine State">Rakhine Yoma Elephant Sanctuary</location>
,
<location id="8E8A606AFFE97E550CDCF8EE117FF8CF" LSID="urn:lsid:plazi:treatment:03FC87A7FFE97E480E17FD5E1106FB5E:8E8A606AFFE97E550CDCF8EE117FF8CF" box="[189,368,1878,1905]" country="Myanmar" county="Kyat Stream" latitude="17.703888" longLatPrecision="1" longitude="94.648415" municipality="Yoma Elephant Sanctuary" name="Gwa Township" pageId="14" pageNumber="172" stateProvince="Rakhine State">Gwa Township</location>
,
<collectingRegion id="4991F853FFE97E550D1DF8EF122DF8CE" box="[380,546,1878,1905]" country="Myanmar" name="Rakhine" pageId="14" pageNumber="172">Rakhine State</collectingRegion>
;
<geoCoordinate id="EE615076FFE97E550E4CF8EE1300F8CE" box="[557,783,1879,1905]" degrees="17.7038888889" direction="north" orientation="latitude" pageId="14" pageNumber="172" precision="1" value="17.703888">17.7038888889° N</geoCoordinate>
,
<geoCoordinate id="EE615076FFE97E550F7BF8EF13F7F8CE" box="[794,1016,1878,1905]" degrees="94.6484166667" direction="east" orientation="longitude" pageId="14" pageNumber="172" precision="1" value="94.648415">94.6484166667° E</geoCoordinate>
, WGS 84; ca
<elevation id="0078D182FFE97E5508F9F8EF152FF8CF" box="[1176,1312,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="14" pageNumber="172" unit="m" value="175.0">
<quantity id="4CAD9B54FFE97E5508F9F8EF14EEF8CE" box="[1176,1249,1878,1905]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="14" pageNumber="172" unit="m" value="175.0">175 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEB1071FFE97E55094CF8EF1146F82B" pageId="14" pageNumber="172" value="1926-04-27" valueMax="1926-04-29" valueMin="1926-04-27">
<collectingDate id="EFAFE999FFE97E55094CF8EF1146F82B" pageId="14" pageNumber="172" value="1926-04-27" valueMax="1926-04-29" valueMin="1926-04-27">2627 and 29 Apr.</collectingDate>
</date>
2021, respectively;
<collectorName id="26A05367FFE97E550E5DF8C312E4F82B" box="[572,747,1914,1940]" pageId="14" pageNumber="172">J.B. Slowinski</collectorName>
,
<collectorName id="26A05367FFE97E550E98F8C313A2F82B" box="[761,941,1914,1940]" pageId="14" pageNumber="172">G.O.U. Wogan</collectorName>
,
<collectorName id="26A05367FFE97E550FDAF8C3141BF82B" box="[955,1044,1914,1940]" pageId="14" pageNumber="172">H. Win</collectorName>
,
<collectorName id="26A05367FFE97E550843F8C31473F82B" box="[1058,1148,1914,1940]" pageId="14" pageNumber="172">T. Thin</collectorName>
, and
<collectorName id="26A05367FFE97E5508DFF8C31536F82B" box="[1214,1337,1914,1940]" pageId="14" pageNumber="172">K.S. Lwin</collectorName>
leg.;
<specimenCode id="DBF39ECAFFE97E550CDEF8241150F807" box="[191,351,1949,1976]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="172" type="Museum">CAS 220256</specimenCode>
,
<specimenCode id="DBF39ECAFFE97E550D0EF8241201F808" box="[367,526,1949,1975]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="172" type="Museum">CAS 220332</specimenCode>
,
<specimenCode id="DBF39ECAFFE97E550E7FF82412B1F808" box="[542,702,1949,1975]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="172" type="Museum">CAS 220378</specimenCode>
, respectively
</materialsCitation>
<materialsCitation id="3B3D3CECFFE97E540F1CF824157CFEF8" collectingDate="2021-05-01" collectionCode="CAS" collectorName="J. B. Slowinski &amp; G. O. U. Wogan &amp; H. Win &amp; T. Thin &amp; K. S. Lwin" country="Myanmar" county="Daung Stream" elevation="125" lastPageId="15" lastPageNumber="173" latitude="17.584183" location="Rakhine Yoma Elephant Sanctuary" longLatPrecision="1" longitude="94.67815" municipality="Yoma Elephant Sanctuary" pageId="14" pageNumber="172" specimenCode="CAS 220550" specimenCount="1" specimenCount-male="1" stateProvince="Rakhine State" typeStatus="paratype">
<specimenCount id="9D53FD38FFE97E550F1CF82413A1F808" box="[893,942,1949,1975]" count="1" pageId="14" pageNumber="172" type="male">1 ♂</specimenCount>
adult;
<collectingCounty id="628B4E3DFFE97E550869F82414B8F808" box="[1032,1207,1949,1975]" pageId="14" pageNumber="172">Daung Stream</collectingCounty>
,
<location id="8E8A606AFFE97E5408A6F82411ACFE9C" LSID="urn:lsid:plazi:treatment:03FC87A7FFE97E480E17FD5E1106FB5E:8E8A606AFFE97E5408A6F82411ACFE9C" country="Myanmar" county="Daung Stream" lastPageId="15" lastPageNumber="173" latitude="17.584183" longLatPrecision="1" longitude="94.67815" municipality="Yoma Elephant Sanctuary" name="Rakhine Yoma Elephant Sanctuary" pageId="14" pageNumber="172" stateProvince="Rakhine State">Rakhine Yoma Elephant Sanctuary</location>
,
<location id="8E8A606AFFE87E540DCEFEB0126CFE9C" LSID="urn:lsid:plazi:treatment:03FC87A7FFE97E480E17FD5E1106FB5E:8E8A606AFFE87E540DCEFEB0126CFE9C" box="[431,611,265,291]" country="Myanmar" county="Daung Stream" latitude="17.584183" longLatPrecision="1" longitude="94.67815" municipality="Yoma Elephant Sanctuary" name="Gwa Township" pageId="15" pageNumber="173" stateProvince="Rakhine State">Gwa Township</location>
,
<collectingRegion id="4991F853FFE87E540E0EFEB01318FE9C" box="[623,791,265,291]" country="Myanmar" name="Rakhine" pageId="15" pageNumber="173">Rakhine State</collectingRegion>
;
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, WGS 84; ca
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a.s.l.
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;
<collectorName id="26A05367FFE87E540DDBFE941269FEF9" box="[442,614,300,327]" pageId="15" pageNumber="173">J.B. Slowinski</collectorName>
,
<collectorName id="26A05367FFE87E540E10FE95132EFEF8" box="[625,801,300,327]" pageId="15" pageNumber="173">G.O.U. Wogan</collectorName>
,
<collectorName id="26A05367FFE87E540F4DFE94138DFEF9" box="[812,898,300,327]" pageId="15" pageNumber="173">H. Win</collectorName>
,
<collectorName id="26A05367FFE87E540FECFE9413EBFEF9" box="[909,996,300,327]" pageId="15" pageNumber="173">T. Thin</collectorName>
, and
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leg.;
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.
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<emphasis id="B921EAA3FFE87E540CDCFED4121FFE37" bold="true" box="[189,528,365,392]" pageId="15" pageNumber="173">
Description of the
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</emphasis>
(
<figureCitation id="136E2A34FFE87E540E7EFED7126AFE37" box="[543,613,365,392]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[203,1384,265,1847]" captionTargetId="figure-11@16.[203,1384,265,1847]" captionTargetPageId="16" captionText="Fig. 5. Herpetoreas davidi sp. nov. AF. Holotype, ♂ (CAS 222969) in preservation. A. Lateral view of the head, right side. B. Lateral view of the head, left side. C. Ventral view of the head. D. Dorsal view of the head. E. General dorsal view. F. General ventral view. GH. Paratype, ♀ (CAS 220256). G. General dorsal view. H. General ventral view. Photos by G. Vogel." figureDoi="http://doi.org/10.5281/zenodo.11112831" httpUri="https://zenodo.org/record/11112831/files/figure.png" pageId="15" pageNumber="173">Fig. 5</figureCitation>
)
</paragraph>
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. Body slender, cylindrical, and elongate (
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<quantity id="4CAD9B54FFE87E540FE8FE2513E1FE08" box="[905,1006,412,439]" metricMagnitude="-1" metricUnit="m" metricValue="3.84" pageId="15" pageNumber="173" unit="mm" value="384.0">384 mm</quantity>
, TaL
<quantity id="4CAD9B54FFE87E540850FE251498FE08" box="[1073,1175,412,439]" metricMagnitude="-1" metricUnit="m" metricValue="1.68" pageId="15" pageNumber="173" unit="mm" value="168.0">168 mm</quantity>
, TL
<quantity id="4CAD9B54FFE87E5408AFFE25153BFE08" box="[1230,1332,412,439]" metricMagnitude="-1" metricUnit="m" metricValue="5.52" pageId="15" pageNumber="173" unit="mm" value="552.0">552 mm</quantity>
, ratio TaL/TL 0.304). Head moderately large, moderately distinct from neck (ratio HL/
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0.048, ratio HW/ HL 0.512), moderately distinct from neck (
<figureCitation id="136E2A34FFE87E540ECCFE5B1328FE42" box="[685,807,482,509]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[203,1384,265,1847]" captionTargetId="figure-11@16.[203,1384,265,1847]" captionTargetPageId="16" captionText="Fig. 5. Herpetoreas davidi sp. nov. AF. Holotype, ♂ (CAS 222969) in preservation. A. Lateral view of the head, right side. B. Lateral view of the head, left side. C. Ventral view of the head. D. Dorsal view of the head. E. General dorsal view. F. General ventral view. GH. Paratype, ♀ (CAS 220256). G. General dorsal view. H. General ventral view. Photos by G. Vogel." figureDoi="http://doi.org/10.5281/zenodo.11112831" httpUri="https://zenodo.org/record/11112831/files/figure.png" pageId="15" pageNumber="173">Fig. 5EF</figureCitation>
); snout broad, obtuse as seen from above; eye large (
<collectionCode id="ED44AE74FFE87E540CA5FDBF10E2FD9F" box="[196,237,518,544]" pageId="15" pageNumber="173">ED</collectionCode>
<quantity id="4CAD9B54FFE87E540C92FDBF1151FD9F" box="[243,350,518,544]" metricMagnitude="-3" metricUnit="m" metricValue="2.86" pageId="15" pageNumber="173" unit="mm" value="2.86">2.86 mm</quantity>
, ratio ED/ESn 1.036, ratio ED/ENs 2.000); pupils round; nostrils crescentic, piercing in the middle of the nasal, oriented laterally (
<figureCitation id="136E2A34FFE87E540ECFFD901320FDFB" box="[686,815,553,580]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[203,1384,265,1847]" captionTargetId="figure-11@16.[203,1384,265,1847]" captionTargetPageId="16" captionText="Fig. 5. Herpetoreas davidi sp. nov. AF. Holotype, ♂ (CAS 222969) in preservation. A. Lateral view of the head, right side. B. Lateral view of the head, left side. C. Ventral view of the head. D. Dorsal view of the head. E. General dorsal view. F. General ventral view. GH. Paratype, ♀ (CAS 220256). G. General dorsal view. H. General ventral view. Photos by G. Vogel." figureDoi="http://doi.org/10.5281/zenodo.11112831" httpUri="https://zenodo.org/record/11112831/files/figure.png" pageId="15" pageNumber="173">Fig. 5AB</figureCitation>
).
</paragraph>
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<collectionCode id="ED44AE74FFE87E540CDCFDD210F1FD3C" box="[189,254,619,645]" pageId="15" pageNumber="173">BODY</collectionCode>
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. Dorsal scales in 191917 rows; all scales strongly keeled, notched at apex; 154 ventrals (plus 2 preventrals); cloacal plate single; 99 subcaudals, all single.
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<collectionCode id="ED44AE74FFE87E540D62FD6C1179FD57" box="[259,374,725,744]" pageId="15" pageNumber="173">SCALATION</collectionCode>
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. Rostral wider than high, width approximately twice as long as high, visible from above (
<figureCitation id="136E2A34FFE87E540CA4FD4B112EFCB2" box="[197,289,754,781]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[203,1384,265,1847]" captionTargetId="figure-11@16.[203,1384,265,1847]" captionTargetPageId="16" captionText="Fig. 5. Herpetoreas davidi sp. nov. AF. Holotype, ♂ (CAS 222969) in preservation. A. Lateral view of the head, right side. B. Lateral view of the head, left side. C. Ventral view of the head. D. Dorsal view of the head. E. General dorsal view. F. General ventral view. GH. Paratype, ♀ (CAS 220256). G. General dorsal view. H. General ventral view. Photos by G. Vogel." figureDoi="http://doi.org/10.5281/zenodo.11112831" httpUri="https://zenodo.org/record/11112831/files/figure.png" pageId="15" pageNumber="173">Fig. 5D</figureCitation>
); nasals pentagonal, slightly elongate, completely divided, anterior part slightly smaller than the posterior one; nostrils located on the middle of nasals (
<figureCitation id="136E2A34FFE87E540F2BFCAC13C2FC8F" box="[842,973,789,816]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[203,1384,265,1847]" captionTargetId="figure-11@16.[203,1384,265,1847]" captionTargetPageId="16" captionText="Fig. 5. Herpetoreas davidi sp. nov. AF. Holotype, ♂ (CAS 222969) in preservation. A. Lateral view of the head, right side. B. Lateral view of the head, left side. C. Ventral view of the head. D. Dorsal view of the head. E. General dorsal view. F. General ventral view. GH. Paratype, ♀ (CAS 220256). G. General dorsal view. H. General ventral view. Photos by G. Vogel." figureDoi="http://doi.org/10.5281/zenodo.11112831" httpUri="https://zenodo.org/record/11112831/files/figure.png" pageId="15" pageNumber="173">Fig. 5AB</figureCitation>
); internasals subtriangular, in broad contact with each other, not in contact with loreal, about equal length to width, slightly, but distinctly narrowing anteriorly, the width of the posterior margin approximately twice as long as the anterior margin; 2 prefrontals, hexagonal, slightly broader than long, about 1.3 times as long as internasals; prefrontal sutures slightly longer than length internasal sutures; frontal pentagonal, elongate, length 1.4 times as long as width, about twice as long as prefrontal; supraocular 1/1, rectangular, elongate, narrowed anteriorly, broadly in contact with prefrontal, with supraocular two-thirds as broad as the frontal; parietals in broad contact with each other, parietal suture subequal to length of frontal (
<figureCitation id="136E2A34FFE87E54096AFBB51566FB98" box="[1291,1385,1036,1063]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[203,1384,265,1847]" captionTargetId="figure-11@16.[203,1384,265,1847]" captionTargetPageId="16" captionText="Fig. 5. Herpetoreas davidi sp. nov. AF. Holotype, ♂ (CAS 222969) in preservation. A. Lateral view of the head, right side. B. Lateral view of the head, left side. C. Ventral view of the head. D. Dorsal view of the head. E. General dorsal view. F. General ventral view. GH. Paratype, ♀ (CAS 220256). G. General dorsal view. H. General ventral view. Photos by G. Vogel." figureDoi="http://doi.org/10.5281/zenodo.11112831" httpUri="https://zenodo.org/record/11112831/files/figure.png" pageId="15" pageNumber="173">Fig. 5D</figureCitation>
). Loreal 1/1, subrectangular, small, about equal in length to depth, in contact with 2
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3
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supralabials, not entering orbit; preoculars 1/1; postoculars 3/3, size decreasing from top to bottom; supralabials 9/9, 2
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3
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in contact with the loreal, 4
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6
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entering orbit, 8
<superScript id="7C209BF9FFE87E540F35FBCC136EFB3C" attach="left" box="[852,865,1141,1155]" fontSize="6" pageId="15" pageNumber="173">th</superScript>
largest; temporals 2+3 on both sides; mental subtriangular, wide, 2.5 times as long as high (
<figureCitation id="136E2A34FFE87E540E8CFB20137FFB0C" box="[749,880,1176,1203]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[203,1384,265,1847]" captionTargetId="figure-11@16.[203,1384,265,1847]" captionTargetPageId="16" captionText="Fig. 5. Herpetoreas davidi sp. nov. AF. Holotype, ♂ (CAS 222969) in preservation. A. Lateral view of the head, right side. B. Lateral view of the head, left side. C. Ventral view of the head. D. Dorsal view of the head. E. General dorsal view. F. General ventral view. GH. Paratype, ♀ (CAS 220256). G. General dorsal view. H. General ventral view. Photos by G. Vogel." figureDoi="http://doi.org/10.5281/zenodo.11112831" httpUri="https://zenodo.org/record/11112831/files/figure.png" pageId="15" pageNumber="173">Fig. 5AB</figureCitation>
); 9/10 infralabials, first pair in contact with each other behind mental, 1
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5
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in contact with anterior chin shields; two pars of chin shields; posterior chin shields 1.3 times as long as anterior ones, separated from each other by small scales; mental groove apparent (
<figureCitation id="136E2A34FFE87E540D50FABB1181FAA2" box="[305,398,1282,1309]" captionStart="Fig" captionStartId="16.[189,232,1892,1918]" captionTargetBox="[203,1384,265,1847]" captionTargetId="figure-11@16.[203,1384,265,1847]" captionTargetPageId="16" captionText="Fig. 5. Herpetoreas davidi sp. nov. AF. Holotype, ♂ (CAS 222969) in preservation. A. Lateral view of the head, right side. B. Lateral view of the head, left side. C. Ventral view of the head. D. Dorsal view of the head. E. General dorsal view. F. General ventral view. GH. Paratype, ♀ (CAS 220256). G. General dorsal view. H. General ventral view. Photos by G. Vogel." figureDoi="http://doi.org/10.5281/zenodo.11112831" httpUri="https://zenodo.org/record/11112831/files/figure.png" pageId="15" pageNumber="173">Fig. 5C</figureCitation>
).
</paragraph>
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<heading id="D0A281DDFFE87E540CDCFAFA120AFAE2" bold="true" box="[189,517,1347,1373]" fontSize="11" level="3" pageId="15" pageNumber="173" reason="3">
<emphasis id="B921EAA3FFE87E540CDCFAFA120AFAE2" bold="true" box="[189,517,1347,1373]" pageId="15" pageNumber="173">Coloration in preservative</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA36B1FFE87E540CDCFACB1443F9A6" blockId="15.[188,1399,1394,1562]" pageId="15" pageNumber="173">
Dorsal surface of head dark brown, with two small and white stripes run from the posterior edge of the 8
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supralabials to the nape. Ventral surface of head uniform whitish. Dorsal surface of body and tail pale dark brown with irregular brown speckling on either side. Ventral surface of body and tail whitish with scales bordered on either side by a blackish-brown stripe. According to
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staff, no photographs of living specimens of the
<typeStatus id="54EE8813FFE87E540DB6F9B91205F9A5" box="[471,522,1536,1562]" pageId="15" pageNumber="173">type</typeStatus>
series of
<taxonomicName id="4C554D32FFE87E540E18F9B91354F9A6" authority="Lalremsanga &amp; Biakzuala, 2024" authorityName="Lalremsanga &amp; Biakzuala" authorityYear="2024" box="[633,859,1535,1561]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="173" phylum="Chordata" rank="species" species="davidi" status="sp. nov.">
<emphasis id="B921EAA3FFE87E540E18F9B91354F9A6" box="[633,859,1535,1561]" italics="true" pageId="15" pageNumber="173">Herpetoreas davidi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257D8FFE87E540F03F9B913B2F9A5" box="[866,957,1536,1562]" pageId="15" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
were taken.
</paragraph>
<paragraph id="8BEA36B1FFE87E540CDCF9F9139FF9E4" blockId="15.[189,912,1600,1627]" box="[189,912,1600,1627]" pageId="15" pageNumber="173">
<emphasis id="B921EAA3FFE87E540CDCF9F9113CF9E5" bold="true" box="[189,307,1600,1626]" pageId="15" pageNumber="173">Variation</emphasis>
(for details of informations see
<tableCitation id="C6D7030AFFE87E540ED2F9F91305F9E4" box="[691,778,1600,1627]" captionStart="Table 1" captionStartId="4.[181,250,551,579]" captionTargetPageId="4" captionText="Table 1 (continued on next page). Main measurements and meristic characters of the Herpetoreas davidi sp. nov., H. xenura (Wall, 1907), and Hebius khasiensis (Boulenger, 1890). Abbreviations: F = adult female; Heb. = Hebius; Her. = Herpetoreas; M= adult male; N/a = not available; SF = subadult female; SM = subadult male." pageId="15" pageNumber="173">Table 1</tableCitation>
and
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)
</paragraph>
<paragraph id="8BEA36B1FFE87E540CDCF9D61531F935" blockId="15.[189,1342,1647,1674]" box="[189,1342,1647,1674]" pageId="15" pageNumber="173">
The longest known specimen is
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long (
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, TaL
<quantity id="4CAD9B54FFE87E540FA7F9D61425F936" box="[966,1066,1647,1673]" metricMagnitude="-1" metricUnit="m" metricValue="1.78" pageId="15" pageNumber="173" unit="mm" value="178.0">178 mm</quantity>
, female;
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220550).
</paragraph>
<paragraph id="8BEA36B1FFE87E540CDCF9081500F951" blockId="15.[189,1398,1712,1774]" pageId="15" pageNumber="173">
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<collectionCode id="ED44AE74FFE87E540CDCF90810F1F976" box="[189,254,1713,1739]" pageId="15" pageNumber="173">BODY</collectionCode>
</smallCapsWord>
<smallCapsWord id="8D0CA06DFFE87E540D66F90F1175F976" baselines="1732" box="[263,378,1718,1737]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="scalation" pageId="15" pageNumber="173">
<collectionCode id="ED44AE74FFE87E540D66F90F1175F976" box="[263,378,1718,1737]" pageId="15" pageNumber="173">SCALATION</collectionCode>
</smallCapsWord>
. 191917 DSR; 151156
<collectingRegion id="4991F853FFE87E540EA5F908130CF974" box="[708,771,1713,1739]" country="Austria" name="Wien" pageId="15" pageNumber="173">VEN</collectingRegion>
, 97100 SC, single in all examined specimens, total number of
<collectingRegion id="4991F853FFE87E540D5EF96D1172F951" box="[319,381,1748,1774]" country="Austria" name="Wien" pageId="15" pageNumber="173">VEN</collectingRegion>
+SC 248253, without sexual dimorphism. SL:
<quantity id="4CAD9B54FFE87E540FCFF96D13D4F952" box="[942,987,1747,1774]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="15" pageNumber="173" unit="in" value="9.0">9 in</quantity>
all occurrences, IL: 910.
</paragraph>
<paragraph id="8BEA36B1FFE87E540CDCF8AF13F1F826" blockId="15.[189,1400,1813,1945]" pageId="15" pageNumber="173">
<smallCapsWord id="8D0CA06DFFE87E540CDCF8AF10F3F891" baselines="1833,1833" box="[189,252,1814,1840]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Main" pageId="15" pageNumber="173">
<collectionCode id="ED44AE74FFE87E540CDCF8AF10F3F891" box="[189,252,1814,1840]" pageId="15" pageNumber="173">MAIN</collectionCode>
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<smallCapsWord id="8D0CA06DFFE87E540D64F8A21181F891" baselines="1833" box="[261,398,1819,1838]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="characters" pageId="15" pageNumber="173">
<collectionCode id="ED44AE74FFE87E540D64F8A21181F891" box="[261,398,1819,1838]" pageId="15" pageNumber="173">CHARACTERS</collectionCode>
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<collectionCode id="ED44AE74FFE87E540DF9F8A211BDF891" box="[408,434,1819,1838]" pageId="15" pageNumber="173">OF</collectionCode>
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<smallCapsWord id="8D0CA06DFFE87E540DDAF8A2121BF891" baselines="1833" box="[443,532,1819,1838]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="pattern" pageId="15" pageNumber="173">
<collectionCode id="ED44AE74FFE87E540DDAF8A2121BF891" box="[443,532,1819,1838]" pageId="15" pageNumber="173">PATTERN</collectionCode>
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. Head olive-brown with two small, faint off-white spots on the parietals and a faint off-white sagittal line just behind the parietal. Dorsum dark olive-brown or dark brown with indistinct narrow blackish cross-bars or series of spots; venter off-whitish or cream, each ventral decorated with a pair of dark brown square spots on their lateral edges.
</paragraph>
</subSubSection>
<caption id="DF2A6639FFF77E4B0CDCF8DD136AF857" ID-DOI="http://doi.org/10.5281/zenodo.11112831" ID-Zenodo-Dep="11112831" httpUri="https://zenodo.org/record/11112831/files/figure.png" pageId="16" pageNumber="174" startId="16.[189,232,1892,1918]" targetBox="[203,1384,265,1847]" targetPageId="16">
<paragraph id="8BEA36B1FFF77E4B0CDCF8DD136AF857" blockId="16.[188,1399,1892,2025]" pageId="16" pageNumber="174">
<emphasis id="B921EAA3FFF77E4B0CDCF8DD1105F8C0" bold="true" box="[189,266,1892,1919]" pageId="16" pageNumber="174">Fig. 5.</emphasis>
<taxonomicName id="4C554D32FFF77E4B0D70F8DF11FDF8C0" authority="Lalremsanga &amp; Biakzuala, 2024" authorityName="Lalremsanga &amp; Biakzuala" authorityYear="2024" box="[273,498,1893,1919]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="174" phylum="Chordata" rank="species" species="davidi" status="sp. nov.">
<emphasis id="B921EAA3FFF77E4B0D70F8DF11FDF8C0" box="[273,498,1893,1919]" italics="true" pageId="16" pageNumber="174">Herpetoreas davidi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257D8FFF77E4B0D98F8DC125CF8C0" box="[505,595,1893,1919]" pageId="16" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B921EAA3FFF77E4B0E39F8DD1282F8C1" bold="true" box="[600,653,1892,1918]" pageId="16" pageNumber="174">AF</emphasis>
. Holotype, ♂ (CAS 222969) in preservation.
<emphasis id="B921EAA3FFF77E4B08C0F8DD14B9F8C1" bold="true" box="[1185,1206,1892,1918]" pageId="16" pageNumber="174">A</emphasis>
. Lateral view of the head, right side.
<emphasis id="B921EAA3FFF77E4B0DC6F83111B4F81D" bold="true" box="[423,443,1928,1954]" pageId="16" pageNumber="174">B</emphasis>
. Lateral view of the head, left side.
<emphasis id="B921EAA3FFF77E4B0F3BF8311360F81D" bold="true" box="[858,879,1928,1954]" pageId="16" pageNumber="174">C</emphasis>
. Ventral view of the head.
<emphasis id="B921EAA3FFF77E4B08C7F83114B4F81D" bold="true" box="[1190,1211,1928,1954]" pageId="16" pageNumber="174">D</emphasis>
. Dorsal view of the head.
<emphasis id="B921EAA3FFF77E4B0D4AF8121130F87A" bold="true" box="[299,319,1963,1989]" pageId="16" pageNumber="174">E</emphasis>
. General dorsal view.
<emphasis id="B921EAA3FFF77E4B0E23F812125BF87A" bold="true" box="[578,596,1963,1989]" pageId="16" pageNumber="174">F</emphasis>
. General ventral view.
<emphasis id="B921EAA3FFF77E4B0F00F8121391F87A" bold="true" box="[865,926,1963,1989]" pageId="16" pageNumber="174">GH</emphasis>
. Paratype, ♀ (CAS 220256).
<emphasis id="B921EAA3FFF77E4B0894F812151BF87A" bold="true" box="[1269,1300,1963,1989]" pageId="16" pageNumber="174">G.</emphasis>
General dorsal view.
<emphasis id="B921EAA3FFF77E4B0D33F8771166F857" bold="true" box="[338,361,1998,2024]" pageId="16" pageNumber="174">H</emphasis>
. General ventral view. Photos by G. Vogel.
</paragraph>
</caption>
<caption id="DF2A6639FFF67E4A0CD5F9CD1524F969" ID-DOI="http://doi.org/10.5281/zenodo.11112833" ID-Zenodo-Dep="11112833" httpUri="https://zenodo.org/record/11112833/files/figure.png" pageId="17" pageNumber="175" startId="17.[180,223,1652,1680]" targetBox="[215,1899,532,1603]" targetPageId="17">
<paragraph id="8BEA36B1FFF67E4A0CD5F9CD1524F969" blockId="17.[180,1934,1652,1750]" pageId="17" pageNumber="175">
<emphasis id="B921EAA3FFF67E4A0CD5F9CD1148F92F" bold="true" box="[180,327,1652,1680]" pageId="17" pageNumber="175">Fig. 6. AF</emphasis>
.
<taxonomicName id="4C554D32FFF67E4A0D3BF9CD12E4F92F" authority="(Wall, 1907)" baseAuthorityName="Wall" baseAuthorityYear="1907" box="[346,747,1652,1680]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="175" phylum="Chordata" rank="species" species="xenura">
<emphasis id="B921EAA3FFF67E4A0D3BF9CD1247F92F" box="[346,584,1652,1680]" italics="true" pageId="17" pageNumber="175">Herpetoreas xenura</emphasis>
(
<bibRefCitation id="EFC44B40FFF67E4A0E3DF9CD12ECF92F" accessDate="2024-04-04" author="Wall F." box="[604,739,1652,1680]" firstAuthor="Wall" journalOrPublisher="The Journal of the Bombay Natural History Society" pageId="17" pageNumber="175" pagination="337 - 357" part="19" publicationUrl="https://www.biodiversitylibrary.org/page/5449715" refId="ref14630" refString="Wall F. 1907. Notes on snakes from the neighbourhood of Darjeeling. The Journal of the Bombay Natural History Society 19: 337 - 357. Available from https: // www. biodiversitylibrary. org / page / 5449715 [accessed 4 Apr. 2024]." title="Notes on snakes from the neighbourhood of Darjeeling" type="journal article" year="1907">Wall, 1907</bibRefCitation>
)
</taxonomicName>
in life.
<emphasis id="B921EAA3FFF67E4A0F32F9CD1383F92F" bold="true" box="[851,908,1652,1680]" pageId="17" pageNumber="175">AC</emphasis>
. Mizoram, India.
<emphasis id="B921EAA3FFF67E4A080DF9CD14AEF92F" bold="true" box="[1132,1185,1652,1680]" pageId="17" pageNumber="175">DF</emphasis>
. Kaptai National Park, Chittagong, Bangladesh.
<emphasis id="B921EAA3FFF67E4A0A96F9CD1727F92F" bold="true" box="[1783,1832,1652,1680]" pageId="17" pageNumber="175">GI</emphasis>
.
<taxonomicName id="4C554D32FFF67E4A0B5AF9CD1201F90C" authority="(Boulenger, 1890)" baseAuthorityName="Boulenger" baseAuthorityYear="1890" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="17" pageNumber="175" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="B921EAA3FFF67E4A0B5AF9CD1122F90C" italics="true" pageId="17" pageNumber="175">Hebius khasiensis</emphasis>
(Boulenger, 1890)
</taxonomicName>
in life.
<emphasis id="B921EAA3FFF67E4A0E12F92E1287F90C" bold="true" box="[627,648,1687,1715]" pageId="17" pageNumber="175">G</emphasis>
. Tamdil, Mizoram, India.
<emphasis id="B921EAA3FFF67E4A0FA6F92E13D3F90C" bold="true" box="[967,988,1687,1715]" pageId="17" pageNumber="175">H</emphasis>
. Bhamo, Kachin, Myanmar.
<emphasis id="B921EAA3FFF67E4A095DF92E154AF90C" bold="true" box="[1340,1349,1687,1715]" pageId="17" pageNumber="175">I</emphasis>
. Nakhon Thai, Phitsanulok, Thailand. Photos by G. Vogel (AC),
<bibRefCitation id="EFC44B40FFF67E4A0D1AF903121DF969" author="Reza A." box="[379,530,1722,1750]" firstAuthor="Reza" journalOrPublisher="Hamadryad" pageId="17" pageNumber="175" pagination="64 - 72" part="35" refId="ref14308" refString="Reza A. 2010. First record of Amphiesma venningi (Wall, 1910) (Serpentes, Colubridae, Natricinae) from Bangladesh, with notes on its taxonomy, natural history, biogeography and other sympatric species. Hamadryad 35: 64 - 72." title="First record of Amphiesma venningi (Wall, 1910) (Serpentes, Colubridae, Natricinae) from Bangladesh, with notes on its taxonomy, natural history, biogeography and other sympatric species" type="journal article" year="2010">Reza (2010)</bibRefCitation>
(DE), P. Shinde (G), K.C. Ouyang (H), and P. Pawangkhanant (I).
</paragraph>
</caption>
<subSubSection id="C34F653AFFF57E480CDCFEB114D9FD87" lastPageId="19" lastPageNumber="177" pageId="18" pageNumber="176" type="diagnosis">
<paragraph id="8BEA36B1FFF57E490CDCFEB11157FE9D" blockId="18.[189,344,264,290]" box="[189,344,264,290]" pageId="18" pageNumber="176">
<heading id="D0A281DDFFF57E490CDCFEB11157FE9D" bold="true" box="[189,344,264,290]" fontSize="11" level="3" pageId="18" pageNumber="176" reason="3">
<emphasis id="B921EAA3FFF57E490CDCFEB11157FE9D" bold="true" box="[189,344,264,290]" pageId="18" pageNumber="176">Comparison</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA36B1FFF57E490CDCFE811578FECB" blockId="18.[189,1399,311,373]" pageId="18" pageNumber="176">
We summarize the main characters separating
<taxonomicName id="4C554D32FFF57E490EBFFE8113CFFEEE" authority="Lalremsanga &amp; Biakzuala, 2024" authorityName="Lalremsanga &amp; Biakzuala" authorityYear="2024" box="[734,960,311,337]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="davidi" status="sp. nov.">
<emphasis id="B921EAA3FFF57E490EBFFE8113CFFEEE" box="[734,960,311,337]" italics="true" pageId="18" pageNumber="176">Herpetoreas davidi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257D8FFF57E490FA6FE81142DFEED" box="[967,1058,312,338]" pageId="18" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
from the other seven species of the genus
<taxonomicName id="4C554D32FFF57E490D2DFEE211D1FECB" authorityName="Gunther" authorityYear="1860" box="[332,478,347,372]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="genus">
<emphasis id="B921EAA3FFF57E490D2DFEE211D1FECB" box="[332,478,347,372]" italics="true" pageId="18" pageNumber="176">Herpetoreas</emphasis>
</taxonomicName>
in
<tableCitation id="C6D7030AFFF57E490E61FEE31257FECB" box="[512,600,346,372]" captionStart="Table 2" captionStartId="11.[180,249,593,621]" captionTargetPageId="11" captionText="Table 2(continued on next two pages). Comparison of morphological characteristics ofHerpetoreas davidi sp. nov. with those of the genus Herpetoreas Günther, 1860. Abbreviations: Heb. = Hebius; Her. = Herpetoreas; N/a = not available." pageId="18" pageNumber="176">Table 2</tableCitation>
. More specifically, the comparisons with these species are as follows:
</paragraph>
<paragraph id="8BEA36B1FFF57E480CDCFE2514D5FED5" blockId="18.[189,1399,412,544]" lastBlockId="19.[189,1399,265,362]" lastPageId="19" lastPageNumber="177" pageId="18" pageNumber="176">
<taxonomicName id="4C554D32FFF57E490CDCFE251191FE09" authority="Lalremsanga &amp; Biakzuala, 2024" authorityName="Lalremsanga &amp; Biakzuala" authorityYear="2024" box="[189,414,412,438]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="davidi" status="sp. nov.">
<emphasis id="B921EAA3FFF57E490CDCFE251191FE09" box="[189,414,412,438]" italics="true" pageId="18" pageNumber="176">Herpetoreas davidi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257D8FFF57E490DC5FE2511F2FE09" box="[420,509,412,438]" pageId="18" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
is easily distinguished from
<taxonomicName id="4C554D32FFF57E490F29FE2513D6FE09" authorityName="Guo, Zhu, Liu, Zhang, Li, Huang &amp; Pyron" authorityYear="2014" box="[840,985,412,438]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="burbrinki">
<emphasis id="B921EAA3FFF57E490F29FE2513D6FE09" box="[840,985,412,438]" italics="true" pageId="18" pageNumber="176">H. burbrinki</emphasis>
</taxonomicName>
,
<taxonomicName id="4C554D32FFF57E490F89FE251452FE09" authorityName="Lalremsanga, Bal, Vogel &amp; Biakzuala" authorityYear="2022" box="[1000,1117,412,438]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="murlen">
<emphasis id="B921EAA3FFF57E490F89FE251452FE09" box="[1000,1117,412,438]" italics="true" pageId="18" pageNumber="176">H. murlen</emphasis>
</taxonomicName>
,
<taxonomicName id="4C554D32FFF57E49080AFE2514DFFE09" baseAuthorityName="Sclater" baseAuthorityYear="1891" box="[1131,1232,412,438]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="pealii">
<emphasis id="B921EAA3FFF57E49080AFE2514DFFE09" box="[1131,1232,412,438]" italics="true" pageId="18" pageNumber="176">H. pealii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C554D32FFF57E4908BFFE251560FE09" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[1246,1391,412,438]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="platyceps">
<emphasis id="B921EAA3FFF57E4908BFFE251560FE09" box="[1246,1391,412,438]" italics="true" pageId="18" pageNumber="176">H. platyceps</emphasis>
</taxonomicName>
,
<taxonomicName id="4C554D32FFF57E490CDCFE791149FE66" authorityName="Gunther" authorityYear="1860" box="[189,326,447,473]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="sieboldii">
<emphasis id="B921EAA3FFF57E490CDCFE791149FE66" box="[189,326,447,473]" italics="true" pageId="18" pageNumber="176">H. sieboldii</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C554D32FFF57E490DE6FE7911E9FE66" authorityName="Ren, Jiang, Huang, David &amp; Li" authorityYear="2022" box="[391,486,448,473]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="tpser">
<emphasis id="B921EAA3FFF57E490DE6FE7911E9FE66" box="[391,486,448,473]" italics="true" pageId="18" pageNumber="176">H. tpser</emphasis>
</taxonomicName>
by having single subcaudals vs divided subcaudals. Particularly,
<taxonomicName id="4C554D32FFF57E490884FE791109FE43" authorityName="Lalremsanga &amp; Biakzuala" authorityYear="2024" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="davidi">
<emphasis id="B921EAA3FFF57E490884FE791109FE43" italics="true" pageId="18" pageNumber="176">Herpetoreas davidi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C554D32FFF57E490DC2FE5A123AFE43" authorityName="Guo, Zhu, Liu, Zhang, Li, Huang &amp; Pyron" authorityYear="2014" box="[419,565,482,508]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="burbrinki">
<emphasis id="B921EAA3FFF57E490DC2FE5A123AFE43" box="[419,565,482,508]" italics="true" pageId="18" pageNumber="176">H. burbrinki</emphasis>
</taxonomicName>
,
<taxonomicName id="4C554D32FFF57E490E24FE5A12B3FE43" authorityName="Lalremsanga, Bal, Vogel &amp; Biakzuala" authorityYear="2022" box="[581,700,482,508]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="murlen">
<emphasis id="B921EAA3FFF57E490E24FE5A12B3FE43" box="[581,700,482,508]" italics="true" pageId="18" pageNumber="176">H. murlen</emphasis>
</taxonomicName>
,
<taxonomicName id="4C554D32FFF57E490EAAFE5A1350FE43" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[715,863,482,508]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="platyceps">
<emphasis id="B921EAA3FFF57E490EAAFE5A1350FE43" box="[715,863,482,508]" italics="true" pageId="18" pageNumber="176">H. platyceps</emphasis>
</taxonomicName>
,
<taxonomicName id="4C554D32FFF57E490F0FFE5A13F8FE43" authorityName="Gunther" authorityYear="1860" box="[878,1015,482,508]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="sieboldii">
<emphasis id="B921EAA3FFF57E490F0FFE5A13F8FE43" box="[878,1015,482,508]" italics="true" pageId="18" pageNumber="176">H. sieboldii</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C554D32FFF57E490858FE5A1497FE43" authorityName="Ren, Jiang, Huang, David &amp; Li" authorityYear="2022" box="[1081,1176,483,508]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="tpser">
<emphasis id="B921EAA3FFF57E490858FE5A1497FE43" box="[1081,1176,483,508]" italics="true" pageId="18" pageNumber="176">H. tpser</emphasis>
</taxonomicName>
by having a single cloacal plate (vs divided) and fewer ventrals
<collectingRegion id="4991F853FFF57E490E83FDBF1311FD9F" box="[738,798,518,544]" country="Austria" name="Wien" pageId="18" pageNumber="176">VEN</collectingRegion>
151156 (vs
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<taxonomicName id="4C554D32FFF57E49083BFDBF14FFFDA0" authorityName="Guo, Zhu, Liu, Zhang, Li, Huang &amp; Pyron" authorityYear="2014" box="[1114,1264,517,543]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="burbrinki">
<emphasis id="B921EAA3FFF57E49083BFDBF14FFFDA0" box="[1114,1264,517,543]" italics="true" pageId="18" pageNumber="176">H. burbrinki</emphasis>
</taxonomicName>
,
<quantity id="4CAD9B54FFF57E490963FDBC155FFDA0" box="[1282,1360,517,543]" metricMagnitude="0" metricUnit="m" metricValue="4.5466" pageId="18" pageNumber="176" unit="in" value="179.0">179 in</quantity>
<taxonomicName id="4C554D32FFF57E48093BFDBF111FFE9C" authorityName="Lalremsanga, Bal, Vogel &amp; Biakzuala" authorityYear="2022" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" lastPageId="19" lastPageNumber="177" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="murlen">
<emphasis id="B921EAA3FFF57E48093BFDBF111FFE9C" italics="true" lastPageId="19" lastPageNumber="177" pageId="18" pageNumber="176">H. murlen</emphasis>
</taxonomicName>
,
<quantity id="4CAD9B54FFF47E480D41FEB011A9FE9C" box="[288,422,265,291]" metricMagnitude="0" metricUnit="m" metricValue="5.3975" metricValueMax="5.9436" metricValueMin="4.8514" pageId="19" pageNumber="177" unit="in" value="212.5" valueMax="234.0" valueMin="191.0">191234 in</quantity>
<taxonomicName id="4C554D32FFF47E480DCEFEB3124DFE9C" baseAuthorityName="Blyth" baseAuthorityYear="1854" box="[431,578,265,291]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="177" phylum="Chordata" rank="species" species="platyceps">
<emphasis id="B921EAA3FFF47E480DCEFEB3124DFE9C" box="[431,578,265,291]" italics="true" pageId="19" pageNumber="177">H. platyceps</emphasis>
</taxonomicName>
, and
<quantity id="4CAD9B54FFF47E480EE5FEB01305FE9C" box="[644,778,265,291]" metricMagnitude="0" metricUnit="m" metricValue="4.8387" metricValueMax="5.2578" metricValueMin="4.4196" pageId="19" pageNumber="177" unit="in" value="190.5" valueMax="207.0" valueMin="174.0">174207 in</quantity>
<taxonomicName id="4C554D32FFF47E480F72FEB31393FE9C" authorityName="Gunther" authorityYear="1860" box="[787,924,265,291]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="177" phylum="Chordata" rank="species" species="sieboldii">
<emphasis id="B921EAA3FFF47E480F72FEB31393FE9C" box="[787,924,265,291]" italics="true" pageId="19" pageNumber="177">H. sieboldii</emphasis>
</taxonomicName>
); differs from
<taxonomicName id="4C554D32FFF47E48082CFEB314BAFE9C" baseAuthorityName="Sclater" baseAuthorityYear="1891" box="[1101,1205,265,291]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="177" phylum="Chordata" rank="species" species="pealii">
<emphasis id="B921EAA3FFF47E48082CFEB314BAFE9C" box="[1101,1205,265,291]" italics="true" pageId="19" pageNumber="177">H. pealii</emphasis>
</taxonomicName>
by having more ventrals
<collectingRegion id="4991F853FFF47E480D42FE941150FEF8" box="[291,351,301,327]" country="Austria" name="Wien" pageId="19" pageNumber="177">VEN</collectingRegion>
151156 (vs 136144); and differs from
<taxonomicName id="4C554D32FFF47E480F2DFE9413A3FEF9" authorityName="Ren, Jiang, Huang, David &amp; Li" authorityYear="2022" box="[844,940,301,326]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="177" phylum="Chordata" rank="species" species="tpser">
<emphasis id="B921EAA3FFF47E480F2DFE9413A3FEF9" box="[844,940,301,326]" italics="true" pageId="19" pageNumber="177">H. tpser</emphasis>
</taxonomicName>
by having a smaller size in both sexes (max TL
<quantity id="4CAD9B54FFF47E480D4AFEF61182FED5" box="[299,397,335,362]" metricMagnitude="-1" metricUnit="m" metricValue="5.52" pageId="19" pageNumber="177" unit="mm" value="552.0">552 mm</quantity>
in males,
<quantity id="4CAD9B54FFF47E480E64FEF61269FED5" box="[517,614,335,362]" metricMagnitude="-1" metricUnit="m" metricValue="5.75" pageId="19" pageNumber="177" unit="mm" value="575.0">575 mm</quantity>
in females vs
<quantity id="4CAD9B54FFF47E480F6EFEF6137EFED5" box="[783,881,335,362]" metricMagnitude="-1" metricUnit="m" metricValue="5.84" pageId="19" pageNumber="177" unit="mm" value="584.0">584 mm</quantity>
in males,
<quantity id="4CAD9B54FFF47E480F88FEF61445FED5" box="[1001,1098,335,362]" metricMagnitude="-1" metricUnit="m" metricValue="6.79" pageId="19" pageNumber="177" unit="mm" value="679.0">679 mm</quantity>
in females).
</paragraph>
<caption id="DF2A6639FFF57E490CDCF8A711B7F856" ID-DOI="http://doi.org/10.5281/zenodo.11112835" ID-Zenodo-Dep="11112835" httpUri="https://zenodo.org/record/11112835/files/figure.png" pageId="18" pageNumber="176" startId="18.[189,232,1822,1848]" targetBox="[201,1390,605,1765]" targetPageId="18">
<paragraph id="8BEA36B1FFF57E490CDCF8A711B7F856" blockId="18.[189,1399,1822,2025]" pageId="18" pageNumber="176">
<emphasis id="B921EAA3FFF57E490CDCF8A7111DF887" bold="true" box="[189,274,1822,1848]" pageId="18" pageNumber="176">Fig. 7.</emphasis>
Map showing the distribution of
<taxonomicName id="4C554D32FFF57E490EA2F8A613A4F887" authority="Lalremsanga &amp; Biakzuala, 2024" authorityName="Lalremsanga &amp; Biakzuala" authorityYear="2024" box="[707,939,1822,1848]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="davidi" status="sp. nov.">
<emphasis id="B921EAA3FFF57E490EA2F8A613A4F887" box="[707,939,1822,1848]" italics="true" pageId="18" pageNumber="176">Herpetoreas davidi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257D8FFF57E490FD8F8A61414F886" box="[953,1051,1823,1849]" pageId="18" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
in Rakhine Yoma Elephant Sanctuary, Rakhine, Myanmar (red stars).
<emphasis id="B921EAA3FFF57E490EC8F8F812B7F8E4" bold="true" box="[681,696,1857,1883]" pageId="18" pageNumber="176">1</emphasis>
. Daung Stream (type locality).
<emphasis id="B921EAA3FFF57E490846F8F81439F8E4" bold="true" box="[1063,1078,1857,1883]" pageId="18" pageNumber="176">2</emphasis>
. Kyat Stream.
<taxonomicName id="4C554D32FFF57E490885F8FB11A6F8C0" authority="(Wall, 1907)" baseAuthorityName="Wall" baseAuthorityYear="1907" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" sensu="stricto" species="xenura">
<emphasis id="B921EAA3FFF57E490885F8FB1101F8C0" italics="true" pageId="18" pageNumber="176">Herpetoreas xenura</emphasis>
(
<bibRefCitation id="EFC44B40FFF57E490D7FF8DC11AEF8C0" accessDate="2024-04-04" author="Wall F." box="[286,417,1893,1919]" firstAuthor="Wall" journalOrPublisher="The Journal of the Bombay Natural History Society" pageId="18" pageNumber="176" pagination="337 - 357" part="19" publicationUrl="https://www.biodiversitylibrary.org/page/5449715" refId="ref14630" refString="Wall F. 1907. Notes on snakes from the neighbourhood of Darjeeling. The Journal of the Bombay Natural History Society 19: 337 - 357. Available from https: // www. biodiversitylibrary. org / page / 5449715 [accessed 4 Apr. 2024]." title="Notes on snakes from the neighbourhood of Darjeeling" type="journal article" year="1907">Wall, 1907</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A21257D8FFF57E490DD1F8DC124EF8C0" box="[432,577,1893,1919]" pageId="18" pageNumber="176" sensu="stricto">sensu stricto</taxonomicNameLabel>
in red circles,
<taxonomicName id="4C554D32FFF57E490E8FF8DC1491F8C0" authority="(Boulenger, 1890)" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[750,1182,1893,1919]" class="Squamata" family="Colubridae" genus="Hebius" kingdom="Animalia" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="B921EAA3FFF57E490E8FF8DC13CDF8C0" box="[750,962,1893,1919]" italics="true" pageId="18" pageNumber="176">Hebius khasiensis</emphasis>
(Boulenger, 1890)
</taxonomicName>
in blue diamonds, and
<taxonomicName id="4C554D32FFF57E490C93F83111FCF81D" baseAuthorityName="Boulenger" baseAuthorityYear="1890" box="[242,499,1928,1954]" class="Squamata" family="Colubridae" genus="Hebius" isUncertain="true" kingdom="Animalia" pageId="18" pageNumber="176" phylum="Chordata" rank="species" species="khasiensis">
<emphasis id="B921EAA3FFF57E490C93F831114AF81D" box="[242,325,1928,1954]" italics="true" pageId="18" pageNumber="176">Hebius</emphasis>
cf.
<emphasis id="B921EAA3FFF57E490D1BF83111FCF81D" box="[378,499,1928,1954]" italics="true" pageId="18" pageNumber="176">khasiensis</emphasis>
</taxonomicName>
in blue triangle. Our estimated distribution range for
<taxonomicName id="4C554D32FFF57E4908E9F8301579F81D" baseAuthorityName="Wall" baseAuthorityYear="1907" box="[1160,1398,1929,1954]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="176" phylum="Chordata" rank="species" sensu="stricto" species="xenura">
<emphasis id="B921EAA3FFF57E4908E9F8301579F81D" box="[1160,1398,1929,1954]" italics="true" pageId="18" pageNumber="176">Herpetoreas xenura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257D8FFF57E490CDCF8151141F87A" box="[189,334,1963,1990]" pageId="18" pageNumber="176" sensu="stricto">sensu stricto</taxonomicNameLabel>
is shaded in red. See Appendix IV for the corresponding locality numbers of the other taxa presented in the map.
</paragraph>
</caption>
<paragraph id="8BEA36B1FFF47E480CDCFE2814D9FD87" blockId="19.[187,1400,401,568]" pageId="19" pageNumber="177">
Finally,
<taxonomicName id="4C554D32FFF47E480D7AFE2B11F3FE14" authority="Lalremsanga &amp; Biakzuala, 2024" authorityName="Lalremsanga &amp; Biakzuala" authorityYear="2024" box="[283,508,401,427]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="177" phylum="Chordata" rank="species" species="davidi" status="sp. nov.">
<emphasis id="B921EAA3FFF47E480D7AFE2B11F3FE14" box="[283,508,401,427]" italics="true" pageId="19" pageNumber="177">Herpetoreas davidi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257D8FFF47E480E63FE281254FE14" box="[514,603,401,427]" pageId="19" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from the superficially similar species
<taxonomicName id="4C554D32FFF47E480962FE2B1579FE14" baseAuthorityName="Wall" baseAuthorityYear="1907" box="[1283,1398,402,427]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="177" phylum="Chordata" rank="species" species="xenura">
<emphasis id="B921EAA3FFF47E480962FE2B1579FE14" box="[1283,1398,402,427]" italics="true" pageId="19" pageNumber="177">H. xenura</emphasis>
</taxonomicName>
by having a smaller size in both sexes (max TL
<quantity id="4CAD9B54FFF47E480E9FFE0D136EFE70" box="[766,865,436,463]" metricMagnitude="-1" metricUnit="m" metricValue="5.52" pageId="19" pageNumber="177" unit="mm" value="552.0">552 mm</quantity>
in males,
<quantity id="4CAD9B54FFF47E480FBEFE0D144DFE70" box="[991,1090,436,463]" metricMagnitude="-1" metricUnit="m" metricValue="5.75" pageId="19" pageNumber="177" unit="mm" value="575.0">575 mm</quantity>
in females vs
<quantity id="4CAD9B54FFF47E480893FE0D155AFE70" box="[1266,1365,436,463]" metricMagnitude="-1" metricUnit="m" metricValue="8.7" pageId="19" pageNumber="177" unit="mm" value="870.0">870 mm</quantity>
in males,
<quantity id="4CAD9B54FFF47E480D72FE6E1179FE4D" box="[275,374,471,498]" metricMagnitude="-1" metricUnit="m" metricValue="7.54" pageId="19" pageNumber="177" unit="mm" value="754.0">754 mm</quantity>
in females), higher ratio TaL/TL in both sexes (
<quantity id="4CAD9B54FFF47E480FD5FE6E1466FE4E" box="[948,1129,471,497]" metricMagnitude="-3" metricUnit="m" metricValue="7.6834999999999996" metricValueMax="7.7216" metricValueMin="7.6454" pageId="19" pageNumber="177" unit="in" value="0.3025" valueMax="0.304" valueMin="0.301">
<geoCoordinate id="EE615076FFF47E480FD5FE6E13FAFE4E" box="[948,1013,471,497]" degrees="0.301" direction="north" orientation="latitude" pageId="19" pageNumber="177" precision="55" value="0.301">0.301</geoCoordinate>
<geoCoordinate id="EE615076FFF47E480F97FE6E1446FE4E" box="[1014,1097,471,497]" degrees="0.304" direction="west" orientation="longitude" pageId="19" pageNumber="177" precision="55" value="-0.304">0.304</geoCoordinate>
in
</quantity>
males,
<quantity id="4CAD9B54FFF47E4808A7FE6E1577FE4E" box="[1222,1400,471,497]" metricMagnitude="-3" metricUnit="m" metricValue="7.8359" metricValueMax="7.874" metricValueMin="7.7978" pageId="19" pageNumber="177" unit="in" value="0.3085" valueMax="0.31" valueMin="0.307">
<geoCoordinate id="EE615076FFF47E4808A7FE6E1508FE4E" box="[1222,1287,471,497]" degrees="0.307" direction="north" orientation="latitude" pageId="19" pageNumber="177" precision="55" value="0.307">0.307</geoCoordinate>
<geoCoordinate id="EE615076FFF47E480966FE6E1557FE4E" box="[1287,1368,471,497]" degrees="0.310" direction="west" orientation="longitude" pageId="19" pageNumber="177" precision="55" value="-0.31">0.310</geoCoordinate>
in
</quantity>
females vs
<quantity id="4CAD9B54FFF47E480D25FE4211F9FDAB" box="[324,502,506,533]" metricMagnitude="-3" metricUnit="m" metricValue="6.3881" metricValueMax="7.188199999999999" metricValueMin="5.587999999999999" pageId="19" pageNumber="177" unit="in" value="0.2515" valueMax="0.283" valueMin="0.22">
<geoCoordinate id="EE615076FFF47E480D25FE421189FDAA" box="[324,390,507,533]" degrees="0.220" direction="north" orientation="latitude" pageId="19" pageNumber="177" precision="55" value="0.22">0.220</geoCoordinate>
<geoCoordinate id="EE615076FFF47E480DE6FE4211D8FDAA" box="[391,471,507,533]" degrees="0.283" direction="west" orientation="longitude" pageId="19" pageNumber="177" precision="55" value="-0.283">0.283</geoCoordinate>
in
</quantity>
males,
<quantity id="4CAD9B54FFF47E480E33FE43130BFDAB" box="[594,772,506,532]" metricMagnitude="-3" metricUnit="m" metricValue="6.4135" metricValueMax="7.0866" metricValueMin="5.7404" pageId="19" pageNumber="177" unit="in" value="0.2525" valueMax="0.279" valueMin="0.226">
<geoCoordinate id="EE615076FFF47E480E33FE43129BFDAB" box="[594,660,506,532]" degrees="0.226" direction="north" orientation="latitude" pageId="19" pageNumber="177" precision="55" value="0.226">0.226</geoCoordinate>
<geoCoordinate id="EE615076FFF47E480EF5FE4312EAFDAB" box="[660,741,506,532]" degrees="0.279" direction="west" orientation="longitude" pageId="19" pageNumber="177" precision="55" value="-0.279">0.279</geoCoordinate>
in
</quantity>
females), and slightly lower number of
<collectingRegion id="4991F853FFF47E480881FE421513FDAA" box="[1248,1308,507,533]" country="Austria" name="Wien" pageId="19" pageNumber="177">VEN</collectingRegion>
in both sexes (
<quantity id="4CAD9B54FFF47E480D6CFDA4119CFD87" box="[269,403,541,568]" metricMagnitude="0" metricUnit="m" metricValue="3.937" metricValueMax="3.9624" metricValueMin="3.9116" pageId="19" pageNumber="177" unit="in" value="155.0" valueMax="156.0" valueMin="154.0">154156 in</quantity>
males,
<quantity id="4CAD9B54FFF47E480D8FFDA4127CFD87" box="[494,627,541,568]" metricMagnitude="0" metricUnit="m" metricValue="3.8862" metricValueMax="3.937" metricValueMin="3.8354" pageId="19" pageNumber="177" unit="in" value="153.0" valueMax="155.0" valueMin="151.0">151155 in</quantity>
females vs
<quantity id="4CAD9B54FFF47E480E9FFDA4138CFD87" box="[766,899,541,568]" metricMagnitude="0" metricUnit="m" metricValue="4.1021" metricValueMax="4.191" metricValueMin="4.013199999999999" pageId="19" pageNumber="177" unit="in" value="161.5" valueMax="165.0" valueMin="158.0">158165 in</quantity>
males,
<quantity id="4CAD9B54FFF47E480FBCFDA4146DFD87" box="[989,1122,541,568]" metricMagnitude="0" metricUnit="m" metricValue="4.064" metricValueMax="4.191" metricValueMin="3.937" pageId="19" pageNumber="177" unit="in" value="160.0" valueMax="165.0" valueMin="155.0">155165 in</quantity>
females).
</paragraph>
</subSubSection>
<subSubSection id="C34F653AFFF47E480CDCFDE6110EFC5B" pageId="19" pageNumber="177" type="distribution">
<paragraph id="8BEA36B1FFF47E480CDCFDE61296FDC6" blockId="19.[189,665,607,633]" box="[189,665,607,633]" pageId="19" pageNumber="177">
<heading id="D0A281DDFFF47E480CDCFDE61296FDC6" bold="true" box="[189,665,607,633]" fontSize="11" level="3" pageId="19" pageNumber="177" reason="3">
<emphasis id="B921EAA3FFF47E480CDCFDE61296FDC6" bold="true" box="[189,665,607,633]" pageId="19" pageNumber="177">Distribution and natural history notes</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA36B1FFF47E480CDCFD37110EFC5B" blockId="19.[189,1399,654,996]" pageId="19" pageNumber="177">
<taxonomicName id="4C554D32FFF47E480CDCFD3711A4FD17" authority="Lalremsanga &amp; Biakzuala, 2024" authorityName="Lalremsanga &amp; Biakzuala" authorityYear="2024" box="[189,427,654,680]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="177" phylum="Chordata" rank="species" species="davidi" status="sp. nov.">
<emphasis id="B921EAA3FFF47E480CDCFD3711A4FD17" box="[189,427,654,680]" italics="true" pageId="19" pageNumber="177">Herpetoreas davidi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257D8FFF47E480DA1FD371228FD17" box="[448,551,654,680]" pageId="19" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
is so far known only from mountain forests in two locations within
<collectingRegion id="4991F853FFF47E480D7AFD081172FD74" box="[283,381,689,715]" country="Myanmar" name="Rakhine" pageId="19" pageNumber="177">Rakhine</collectingRegion>
Yoma Elephant Sanctuary, Gwa Township,
<collectingRegion id="4991F853FFF47E480FA7FD081476FD74" box="[966,1145,689,715]" country="Myanmar" name="Rakhine" pageId="19" pageNumber="177">Rakhine State</collectingRegion>
,
<collectingCountry id="F3427621FFF47E4808F1FD08150DFD74" box="[1168,1282,689,715]" name="Myanmar" pageId="19" pageNumber="177">Myanmar</collectingCountry>
(
<figureCitation id="136E2A34FFF47E48097CFD08157EFD74" box="[1309,1393,689,715]" captionStart="Fig" captionStartId="18.[189,232,1822,1848]" captionTargetBox="[201,1390,605,1765]" captionTargetId="figure-119@18.[189,1398,600,1779]" captionTargetPageId="18" captionText="Fig. 7. Map showing the distribution of Herpetoreas davidi sp. nov. in Rakhine Yoma Elephant Sanctuary, Rakhine, Myanmar (red stars). 1. Daung Stream (type locality). 2. Kyat Stream. Herpetoreas xenura (Wall, 1907) sensu stricto in red circles, Hebius khasiensis (Boulenger, 1890) in blue diamonds, and Hebius cf. khasiensis in blue triangle. Our estimated distribution range for Herpetoreas xenura sensu stricto is shaded in red. See Appendix IV for the corresponding locality numbers of the other taxa presented in the map." figureDoi="http://doi.org/10.5281/zenodo.11112835" httpUri="https://zenodo.org/record/11112835/files/figure.png" pageId="19" pageNumber="177">Fig. 7</figureCitation>
;
<figureCitation id="136E2A34FFF47E480CDCFD6D115BFD51" box="[189,340,724,750]" captionStart="Appendix 4" captionStartId="33.[181,304,550,578]" captionTargetPageId="33" captionText="Appendix 4 (continued on next seven pages). Uncorrected p-distance estimated using Cytb gene sequence among the genera Hebius Thompson, 1913,Herpetoreas Günther, 1860, Amphiesma Duméril, Bibron &amp; Duméril, 1854, and Sahyadriophis Patel, Thackeray, Campbell &amp; Mirza, 2023, with Amphiesmoides Malnate, 1961 as outgroup." pageId="19" pageNumber="177">Appendix 4</figureCitation>
). Further studies in neighbouring areas, especially in the northern
<collectingRegion id="4991F853FFF47E4808A2FD6D152AFD51" box="[1219,1317,724,750]" country="Myanmar" name="Rakhine" pageId="19" pageNumber="177">Rakhine</collectingRegion>
Hills,
<collectingCountry id="F3427621FFF47E480CDCFD41113EFCAD" box="[189,305,760,786]" name="Myanmar" pageId="19" pageNumber="177">Myanmar</collectingCountry>
, are required to determine the actual distribution and population size and trend of this new species. According to data recorded for specimens deposited in the collections of CAS (http://portal.vertnet.org/search?q=
<taxonomicName id="4C554D32FFF47E480E36FC87134DFCE8" baseAuthorityName="Wall" baseAuthorityYear="1907" box="[599,834,830,856]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="177" phylum="Chordata" rank="species" species="xenura">
<emphasis id="B921EAA3FFF47E480E36FC8712EFFCE7" box="[599,736,830,856]" italics="true" pageId="19" pageNumber="177">Amphiesma</emphasis>
+
<emphasis id="B921EAA3FFF47E480E90FC87134DFCE8" box="[753,834,830,855]" italics="true" pageId="19" pageNumber="177">xenura</emphasis>
</taxonomicName>
+CAS+
<collectingCountry id="F3427621FFF47E480FF8FC871401FCE7" box="[921,1038,830,856]" name="Myanmar" pageId="19" pageNumber="177">Myanmar</collectingCountry>
), specimens of
<taxonomicName id="4C554D32FFF47E480884FC871109FCC4" authorityName="Lalremsanga &amp; Biakzuala" authorityYear="2024" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="177" phylum="Chordata" rank="species" species="davidi">
<emphasis id="B921EAA3FFF47E480884FC871109FCC4" italics="true" pageId="19" pageNumber="177">Herpetoreas davidi</emphasis>
</taxonomicName>
were collected both during the day and at night on the ground near streams. The air temperature varied from 28°C to 31°C, and relative humidity was 94%. This species seems to be distributed at low elevations. The stomach of the
<typeStatus id="54EE8813FFF47E480E55FC1E1294FC7E" box="[564,667,935,961]" pageId="19" pageNumber="177" type="holotype">holotype</typeStatus>
CAS 222969 contained a frog of the family
<taxonomicName id="4C554D32FFF47E4808D7FC1E10F2FC5B" authority="Batsch, 1796" authorityName="Batsch" authorityYear="1796" class="Amphibia" family="Ranidae" kingdom="Animalia" order="Anura" pageId="19" pageNumber="177" phylum="Chordata" rank="family">Ranidae Batsch, 1796</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F653AFFF47E480CDCFBB51106FB5E" pageId="19" pageNumber="177" type="conservation">
<paragraph id="8BEA36B1FFF47E480CDCFBB511BAFB99" blockId="19.[189,437,1036,1062]" box="[189,437,1036,1062]" pageId="19" pageNumber="177">
<heading id="D0A281DDFFF47E480CDCFBB511BAFB99" bold="true" box="[189,437,1036,1062]" fontSize="11" level="3" pageId="19" pageNumber="177" reason="3">
<emphasis id="B921EAA3FFF47E480CDCFBB511BAFB99" bold="true" box="[189,437,1036,1062]" pageId="19" pageNumber="177">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA36B1FFF47E480CDCFB821106FB5E" blockId="19.[189,1401,1082,1249]" pageId="19" pageNumber="177">
<taxonomicName id="4C554D32FFF47E480CDCFB8211ACFBEB" authority="Lalremsanga &amp; Biakzuala, 2024" authorityName="Lalremsanga &amp; Biakzuala" authorityYear="2024" box="[189,419,1082,1108]" class="Reptilia" family="Colubridae" genus="Herpetoreas" kingdom="Animalia" order="Squamata" pageId="19" pageNumber="177" phylum="Chordata" rank="species" species="davidi" status="sp. nov.">
<emphasis id="B921EAA3FFF47E480CDCFB8211ACFBEB" box="[189,419,1082,1108]" italics="true" pageId="19" pageNumber="177">Herpetoreas davidi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257D8FFF47E480DCFFB821202FBEA" box="[430,525,1083,1109]" pageId="19" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
is currently known only from
<collectingRegion id="4991F853FFF47E480FEBFB8313E3FBEB" box="[906,1004,1082,1108]" country="Myanmar" name="Rakhine" pageId="19" pageNumber="177">Rakhine</collectingRegion>
Yoma Elephant Sanctuary, Gwa Township,
<collectingRegion id="4991F853FFF47E480D20FBE411E6FBC7" box="[321,489,1117,1144]" country="Myanmar" name="Rakhine" pageId="19" pageNumber="177">Rakhine State</collectingRegion>
,
<collectingCountry id="F3427621FFF47E480D97FBE71265FBC7" box="[502,618,1118,1144]" name="Myanmar" pageId="19" pageNumber="177">Myanmar</collectingCountry>
, a protected area. In absence of other biological data to assess its population status and potential threat, we suggest that the species should be considered a Data Deficient (DD) species, following the IUCNs Red List categories (
<bibRefCitation id="EFC44B40FFF47E480F1AFB1D10F2FB5E" accessDate="2024-04-04" author="IUCN Standards and Petitions Committee" firstAuthor="IUCN Standards" journalOrPublisher="Standards and Petitions Committee" pageId="19" pageNumber="177" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="ref12733" refString="IUCN Standards and Petitions Committee 2019. Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Committee. Available from http: // www. iucnredlist. org / documents / RedListGuidelines. pdf [accessed 4 Apr. 2024]." title="Guidelines for Using the IUCN Red List Categories and Criteria. Version 14" type="book" year="2019">IUCN Standards and Petitions Committee 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>