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<document id="99EF7CEF34635B1A00D712151797D5A6" ID-DOI="10.5281/zenodo.172764" ID-GBIF-Dataset="57f77567-9f74-4e9f-a79a-a19a41e960c8" ID-ISSN="1175­5326" ID-Zenodo-Dep="172764" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459761781846" checkinUser="plazi" docAuthor="Blahnik, Roger J. &amp; Holzenthal, Ralph W." docDate="2006" docId="03FC8793FFC16F79846AF993C2421E05" docLanguage="en" docName="zt01233p052.pdf" docOrigin="Zootaxa 1233" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Culoptila bidentata Blahnik &amp; Holzenthal, 2006, new species" docType="treatment" docVersion="10" lastPageNumber="19" masterDocId="FFC5FFEBFFD16F6B8562FFAFC6051D3B" masterDocTitle="Revision of the genus Culoptila (Trichoptera: Glossosomatidae)" masterLastPageNumber="52" masterPageNumber="1" pageNumber="17" updateTime="1698215326134" updateUser="plazi">
<mods:mods id="12EC17595CABA760FA22BD5CC69B059A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4BFAB77E1C1A0D1E093CD9A86A8B9161">
<mods:title id="052551D6D53DCD945D1DFB2A423E3596">Revision of the genus Culoptila (Trichoptera: Glossosomatidae)</mods:title>
</mods:titleInfo>
<mods:name id="3DEB056E507F73658D2D9C75898C9B1B" type="personal">
<mods:role id="36AECEC3FE9E5623326BDEB934D82454">
<mods:roleTerm id="CAC0A52AE265692CFFF83472ECB2430E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F7EB7F97F31ABBF763862E6A879CBD2D">Blahnik, Roger J.</mods:namePart>
</mods:name>
<mods:name id="698FC5DACC4BF4328B5BEF66373C0CBE" type="personal">
<mods:role id="C6B583A3AAE2F455ECF15EC13D7C5624">
<mods:roleTerm id="539E4AD30AA9D3870C5A0ED2456E4560">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="30C41306B2AF45FF51F7EFF06AF681C2">Holzenthal, Ralph W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="22508C48A22D973A3129555C938C4D6F">text</mods:typeOfResource>
<mods:relatedItem id="54654B651AE061B0C000856BF993549B" type="host">
<mods:titleInfo id="01E6EE49F8F641E925260AC3B31EA65C">
<mods:title id="3C7ED763A408DD06EE7540768C1F4F72">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="561466FA27B9B53D77F31F53C358156A">
<mods:date id="94F7A9E26DF370C7084126C0188B5F02">2006</mods:date>
<mods:detail id="ED6B4D1527E16A412C9557E121BE6CA7" type="volume">
<mods:number id="74644603AE1A9B8F2BDE9DFD0C90A271">1233</mods:number>
</mods:detail>
<mods:extent id="209107B1039E36A51DAEAB46C7FCCB0D" unit="page">
<mods:start id="38104CEF329289E44EF8458711FCE95B">1</mods:start>
<mods:end id="7A5A78476CEC1FDC23CF3A600BEE64C6">52</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="AF584A60B66A64A792A8B701C7B51A8E">journal article</mods:classification>
<mods:identifier id="C5A43801608E7D29AAD68B9CB6E12495" type="DOI">10.5281/zenodo.172764</mods:identifier>
<mods:identifier id="135AF8F04C7D66F0553CF6FD0474F461" type="GBIF-Dataset">57f77567-9f74-4e9f-a79a-a19a41e960c8</mods:identifier>
<mods:identifier id="9E16014BB9F5E983A367F2F9A901B761" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="87F4772B65B95C4A8E9A9EA97831D7B2" type="Zenodo-Dep">172764</mods:identifier>
</mods:mods>
<treatment id="03FC8793FFC16F79846AF993C2421E05" ID-DOI="http://doi.org/10.5281/zenodo.6256629" ID-GBIF-Taxon="119345086" ID-Zenodo-Dep="6256629" LSID="urn:lsid:plazi:treatment:03FC8793FFC16F79846AF993C2421E05" httpUri="http://treatment.plazi.org/id/03FC8793FFC16F79846AF993C2421E05" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<subSubSection id="C34F650EFFC16F7B846AF993C4041B42" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BEA3685FFC16F7B846AF993C48B1B6D" blockId="16.[264,654,1596,1657]" box="[264,654,1596,1622]" pageId="16" pageNumber="17">
<heading id="D0A281E9FFC16F7B846AF993C48B1B6D" bold="true" box="[264,654,1596,1622]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B921EA97FFC16F7B846AF993C48B1B6D" bold="true" box="[264,654,1596,1622]" pageId="16" pageNumber="17">
<taxonomicName id="4C554D06FFC16F7B846AF993C7EA1B6D" box="[264,495,1596,1622]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bidentata" status="sp. nov.">
<emphasis id="B921EA97FFC16F7B846AF993C7EA1B6D" bold="true" box="[264,495,1596,1622]" italics="true" pageId="16" pageNumber="17">Culoptila bidentata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21257ECFFC16F7B849CF993C48B1B6D" box="[510,654,1596,1622]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA3685FFC16F7B846AF9F0C4041B42" blockId="16.[264,654,1596,1657]" box="[264,513,1631,1657]" pageId="16" pageNumber="17">
<figureCitation id="136E2A00FFC16F7B846AF9F0C7631B42" box="[264,358,1631,1657]" captionStart="FIGURES 15 – 16" captionStartId="17.[264,383,1393,1417]" captionTargetBox="[358,1233,398,1379]" captionTargetId="figure@17.[358,1235,398,1379]" captionTargetPageId="17" captionText="FIGURES 15 – 16. Culoptila bidentata, new species. 15. Male genitalia: 15 A—lateral; 15 Bventral. 16. Female genitalia: 16 A—lateral; 16 B—ventral." httpUri="https://zenodo.org/record/172771/files/figure.png" pageId="16" pageNumber="17">Figs. 15</figureCitation>
A, B, 16A, B
</paragraph>
</subSubSection>
<subSubSection id="C34F650EFFC16F79846AF900C5441CC5" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="description">
<paragraph id="8BEA3685FFC16F7A846AF900C5B51C65" blockId="16.[264,1323,1711,1897]" lastBlockId="17.[264,1322,284,350]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
This species is probably closely related to
<taxonomicName id="4C554D06FFC16F7B8664F900C56E1BF3" box="[774,875,1711,1736]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="aluca">
<emphasis id="B921EA97FFC16F7B8664F900C56E1BF3" box="[774,875,1711,1736]" italics="true" pageId="16" pageNumber="17">C. aluca</emphasis>
</taxonomicName>
and
<taxonomicName id="4C554D06FFC16F7B86C5F900C2121BF3" box="[935,1047,1711,1736]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="azulae">
<emphasis id="B921EA97FFC16F7B86C5F900C2121BF3" box="[935,1047,1711,1736]" italics="true" pageId="16" pageNumber="17">C. azulae</emphasis>
</taxonomicName>
, agreeing in having an inflated phallobase with 2 prominent spines, one strongly curved and with a greatly enlarged base. In general form it probably most closely resembles
<taxonomicName id="4C554D06FFC16F7B8150F950C2A11A23" box="[1074,1188,1791,1816]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="azulae">
<emphasis id="B921EA97FFC16F7B8150F950C2A11A23" box="[1074,1188,1791,1816]" italics="true" pageId="16" pageNumber="17">C. azulae</emphasis>
</taxonomicName>
in that the longer spine is the one with the enlarged base and the inferior appendages are nearly as elongate. It differs from either of those species in being smaller overall, and with a shorter tergum X. The apicolateral process of tergum X bears a prominent apical spine, somewhat like
<taxonomicName id="4C554D06FFC06F7A845EFEEBC79A1C66" box="[316,415,324,349]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="aluca">
<emphasis id="B921EA97FFC06F7A845EFEEBC79A1C66" box="[316,415,324,349]" italics="true" pageId="17" pageNumber="18">C. aluca</emphasis>
</taxonomicName>
, but not so enlarged and differently oriented.
</paragraph>
<caption id="DF2A660DFFC06F7A846AFADEC57D1896" httpUri="https://zenodo.org/record/172771/files/figure.png" pageId="17" pageNumber="18" targetBox="[358,1233,398,1379]" targetPageId="17">
<paragraph id="8BEA3685FFC06F7A846AFADEC57D1896" blockId="17.[264,1322,1393,1453]" pageId="17" pageNumber="18">
<emphasis id="B921EA97FFC06F7A846AFADEC7D618B2" bold="true" box="[264,467,1393,1417]" pageId="17" pageNumber="18">FIGURES 15–16.</emphasis>
<taxonomicName id="4C554D06FFC06F7A8482FADEC4B318B3" box="[480,694,1393,1416]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="bidentata" status="sp. nov.">
<emphasis id="B921EA97FFC06F7A8482FADEC4B318B3" box="[480,694,1393,1416]" italics="true" pageId="17" pageNumber="18">Culoptila bidentata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21257ECFFC06F7A87ABFADEC55718B3" box="[713,850,1393,1416]" pageId="17" pageNumber="18" rank="species">new species</taxonomicNameLabel>
. 15. Male genitalia: 15A—lateral; 15Bventral. 16. Female genitalia: 16A—lateral; 16B—ventral.
</paragraph>
</caption>
<paragraph id="8BEA3685FFC06F7A845AFA4BC73E1BA5" blockId="17.[264,1324,1508,1894]" pageId="17" pageNumber="18">
Adult. Length of forewing: male
<quantity id="4CAD9B60FFC06F7A87B7FA4BC55D18C5" box="[725,856,1508,1534]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="17" pageNumber="18" unit="mm" value="2.1" valueMax="2.2" valueMin="2.0">2–2.2 mm</quantity>
.; female
<quantity id="4CAD9B60FFC06F7A86ABFA4BC25B18C5" box="[969,1118,1508,1534]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="17" pageNumber="18" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.2–2.5 mm</quantity>
. Color brown in alcohol. Mesothorax of male not noticeably modified; mesoscutal wart prominent, subtriangular, wide at anterior margin. Mesothoracic tegulae of male flattened and weakly cupped, distinctly enlarged; tegular setae short; tegular gland short, pleated, concertinalike.
</paragraph>
<paragraph id="8BEA3685FFC06F79845AF903C5441CC5" blockId="17.[264,1324,1508,1894]" lastBlockId="18.[264,1324,284,510]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">Male genitalia. Sternum VI process short, subtruncate. Tergum IX ventral margin rounded, tapered from anterolateral margin, slightly produced posteroventrally. Inferior appendages moderately long, narrow apically, divergent from base. Tergum X very short, length much less than width, apex in ventral view truncate to subtruncate; ventrolateral processes with apices incurved, slightly narrowed and anteriorly bent, and with diverging fork or branch from posterior margin, forming acute spine­like projection. Phallobase large, broadly rounded in lateral view, apicodorsal projection short, with slight inflection at base, apex subacute. Phallic apparatus with 2 phallic spines, one greatly elongate, overall length exceeding length of phallobase, narrow, strongly recurved at base, basally greatly enlarged, sclerotized, the other spine elongate, narrow, nearly straight, about 1/2 length of phallobase, without basal enlargement.</paragraph>
</subSubSection>
<subSubSection id="C34F650EFFC36F79846AFD9BC2421E05" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BEA3685FFC36F79846AFD9BC7E31F76" blockId="18.[264,1325,564,830]" box="[264,486,564,589]" pageId="18" pageNumber="19">
<emphasis id="B921EA97FFC36F79846AFD9BC7E31F76" box="[264,486,564,589]" italics="true" pageId="18" pageNumber="19">Material examined</emphasis>
</paragraph>
<paragraph id="8BEA3685FFC36F79845AFDF4C7761FFD" blockId="18.[264,1325,564,830]" pageId="18" pageNumber="19">
<emphasis id="B921EA97FFC36F79845AFDF4C55E1F4E" bold="true" box="[312,859,603,629]" pageId="18" pageNumber="19">
<typeStatus id="54EE8827FFC36F79845AFDF4C7AF1F4E" box="[312,426,603,629]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F3427615FFC36F798772FDF4C4D71F4E" box="[528,722,603,629]" name="Costa Rica" pageId="18" pageNumber="19">COSTA RICA</collectingCountry>
: Alajuela:
</emphasis>
Río Pizote, ca.
<quantity id="4CAD9B60FFC36F798154FDF3C27E1F4D" box="[1078,1147,604,630]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="km" value="5.0">5 km</quantity>
N Dos Rios,
<geoCoordinate id="EE615042FFC36F79846AFD2BC79C1FA5" box="[264,409,644,670]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="15" value="10.948055">10°56’53&quot;N</geoCoordinate>
,
<geoCoordinate id="EE615042FFC36F7984CBFD2BC4481FA5" box="[425,589,644,670]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="15" value="-85.29111">085°17’28&quot;W</geoCoordinate>
,
<quantity id="4CAD9B60FFC36F79873FFD2BC4AB1FA5" box="[605,686,644,670]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="18" pageNumber="19" unit="m" value="470.0">470 m</quantity>
,
<date id="FFEB1045FFC36F7987DFFD2BC5281FA5" box="[701,813,644,670]" pageId="18" pageNumber="19" value="1986-03-09">9.iii.1986</date>
(Holzenthal &amp; Fasth) (
<collectionCode id="ED44AE40FFC36F79812EFD2BC2981FA5" box="[1100,1181,644,670]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="18" pageNumber="19">UMSP</collectionCode>
000018803) (
<collectionCode id="ED44AE40FFC36F798472FD03C7631FFD" box="[272,358,684,710]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="18" pageNumber="19">UMSP</collectionCode>
).
</paragraph>
<paragraph id="8BEA3685FFC36F79845AFD7BC2421E05" blockId="18.[264,1325,564,830]" pageId="18" pageNumber="19">
<emphasis id="B921EA97FFC36F79845AFD7BC50C1FD5" bold="true" box="[312,777,724,750]" pageId="18" pageNumber="19">
<typeStatus id="54EE8827FFC36F79845AFD7BC7BF1FD5" box="[312,442,724,750]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F3427615FFC36F7984ABFD7BC4831FD5" box="[457,646,724,750]" name="Costa Rica" pageId="18" pageNumber="19">COSTA RICA</collectingCountry>
: Alajuela:
</emphasis>
same data as
<typeStatus id="54EE8827FFC36F7986DBFD7BC2251FD5" box="[953,1056,724,750]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
—
<specimenCount id="9D53FD0CFFC36F798130FD7BC2A21FD5" box="[1106,1191,724,750]" pageId="18" pageNumber="19" type="male">1 male</specimenCount>
,
<specimenCount id="9D53FD0CFFC36F7981D4FD7BC32E1FD5" box="[1206,1323,724,750]" pageId="18" pageNumber="19" type="female">6 females</specimenCount>
(
<collectionCode id="ED44AE40FFC36F798472FD53C7631E2D" box="[272,358,764,790]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="18" pageNumber="19">UMSP</collectionCode>
); Río Pizote, ca.
<quantity id="4CAD9B60FFC36F79872CFD53C4971E2D" box="[590,658,764,790]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="km" value="5.0">5 km</quantity>
(air) S
<collectingCountry id="F3427615FFC36F798661FD53C5641E2D" box="[771,865,764,790]" name="Brazil" pageId="18" pageNumber="19">Brasilia</collectingCountry>
,
<geoCoordinate id="EE615042FFC36F798617FD53C2031E2D" box="[885,1030,764,790]" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="15" value="10.971944">10°58’19&quot;N</geoCoordinate>
,
<geoCoordinate id="EE615042FFC36F798179FD53C2BA1E2D" box="[1051,1215,764,790]" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="15" value="-85.345">085°20’42&quot;W</geoCoordinate>
,
<quantity id="4CAD9B60FFC36F7981B1FD53C32D1E2D" box="[1235,1320,764,790]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="18" pageNumber="19" unit="m" value="390.0">390 m</quantity>
,
<date id="FFEB1045FFC36F79846AFC8BC7821E05" box="[264,391,804,830]" pageId="18" pageNumber="19" value="1986-03-12">12.iii.1986</date>
(Holzenthal &amp; Fasth) —
<specimenCount id="9D53FD0CFFC36F7987D6FC8BC50A1E05" box="[692,783,804,830]" pageId="18" pageNumber="19" type="male">7 males</specimenCount>
(
<collectionCode id="ED44AE40FFC36F79867CFC8BC56A1E05" box="[798,879,804,830]" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="18" pageNumber="19">UMSP</collectionCode>
,
<collectionCode id="ED44AE40FFC36F798619FC8BC5DD1E05" box="[891,984,804,830]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19">NMNH</collectionCode>
,
<collectionCode id="ED44AE40FFC36F798686FC8BC23E1E05" box="[996,1083,804,830]" pageId="18" pageNumber="19">INBIO</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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