548 lines
88 KiB
XML
548 lines
88 KiB
XML
<document id="18D05F32C800B872AA7B477434219502" ID-DOI="10.5281/zenodo.184238" ID-GBIF-Dataset="e24bcebe-de5c-4ff9-b005-dd9523f718e8" ID-ISSN="1175-5326" ID-Zenodo-Dep="184238" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459963415029" checkinUser="plazi" docAuthor="Komai, Tomoyuki & Eletskaya, Maria" docDate="2008" docId="03FC8790FF8DFF94FF23875187E7BAE1" docLanguage="en" docName="zt01882p056.pdf" docOrigin="Zootaxa 1882" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Pandalus ivanovi Komai & Eletskaya, 2008, n. sp." docType="treatment" docVersion="8" lastPageNumber="55" masterDocId="FFC5FFE8FF8CFF9DFFB482548441BD42" masterDocTitle="A new species of the pandalid shrimp genus Pandalus (Crustacea: Decapoda: Caridea) from the Sea of Okhotsk off eastern Sakhalin, Russian Far East" masterLastPageNumber="56" masterPageNumber="46" pageNumber="47" updateTime="1698230947729" updateUser="plazi">
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<mods:mods id="5F46FEABB4B91F50BE20321DC6C817AE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5BBE2124FBB6D6E576B6EC2171F8E59B">
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<mods:title id="8FA580DEF82ED0217A981A094380DC7A">A new species of the pandalid shrimp genus Pandalus (Crustacea: Decapoda: Caridea) from the Sea of Okhotsk off eastern Sakhalin, Russian Far East</mods:title>
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</mods:titleInfo>
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<mods:name id="A0871765383F47BCC34AE38641FF2A33" type="personal">
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<mods:roleTerm id="3A91548D1BD7BA9BBE875E8473CF954F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BFDAD6FA36561B32EA9B692E15224D1E">Komai, Tomoyuki</mods:namePart>
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</mods:name>
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<mods:name id="0E37D8D8C6DCF1745A25EAA0161D7050" type="personal">
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<mods:roleTerm id="29D165CCF885D3007A0AD5235FFEC091">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="05945E1D3C14476E33A23B357276EF1B">Eletskaya, Maria</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="F294BB1815955221A02AC7B1AF5AABA4">text</mods:typeOfResource>
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<mods:titleInfo id="824015C17E032B22826A076B519BABB9">
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<mods:title id="9CF465F3DF683FD8476D0A5452B33B64">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="12A40A8CD68FB7E037ACFF9B0B86E524">
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<mods:date id="17B8A4619496837BA97D4B8BC32EC59E">2008</mods:date>
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<mods:detail id="93FB14B02B545BD4E8E100D695542E0B" type="volume">
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<mods:number id="88F82D6694F40BA608B575F6B63672D0">1882</mods:number>
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<mods:start id="89C0B60856D0F3CB218B8412F2AE0FFD">46</mods:start>
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<mods:end id="9458241224A99ECFA8364DDE387B94F7">56</mods:end>
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<mods:classification id="A293ACB2A5A963A8F1E65BE94CC8E799">journal article</mods:classification>
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<mods:identifier id="289733E09E7823B2195D410C1EEFEF60" type="DOI">10.5281/zenodo.184238</mods:identifier>
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<mods:identifier id="80BA8966C2C41AD59B3889F4E50B505D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="3CC5D078816EA0A3A0D7729608E82FE5" type="Zenodo-Dep">184238</mods:identifier>
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<treatment id="03FC8790FF8DFF94FF23875187E7BAE1" ID-DOI="http://doi.org/10.5281/zenodo.6228294" ID-GBIF-Taxon="119377076" ID-Zenodo-Dep="6228294" LSID="urn:lsid:plazi:treatment:03FC8790FF8DFF94FF23875187E7BAE1" httpUri="http://treatment.plazi.org/id/03FC8790FF8DFF94FF23875187E7BAE1" lastPageId="9" lastPageNumber="55" pageId="1" pageNumber="47">
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<subSubSection id="C34F650DFF8DFF9CFF2387518619B85D" box="[151,600,1285,1311]" pageId="1" pageNumber="47" type="nomenclature">
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<paragraph id="8BEA3686FF8DFF9CFF2387518619B85D" blockId="1.[151,600,1285,1311]" box="[151,600,1285,1311]" pageId="1" pageNumber="47">
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<heading id="D0A281EAFF8DFF9CFF2387518619B85D" bold="true" box="[151,600,1285,1311]" fontSize="11" level="1" pageId="1" pageNumber="47" reason="1">
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<emphasis id="B921EA94FF8DFF9CFF2387518619B85D" bold="true" box="[151,600,1285,1311]" pageId="1" pageNumber="47">
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<taxonomicName id="4C554D05FF8DFF9CFF2387518529B85D" ID-CoL="9CZ4B" box="[151,360,1285,1311]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="47" phylum="Arthropoda" rank="species" species="ivanovi" status="sp. nov.">
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<emphasis id="B921EA94FF8DFF9CFF2387518529B85D" bold="true" box="[151,360,1285,1311]" italics="true" pageId="1" pageNumber="47">Pandalus ivanovi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21257EFFF8DFF9CFEDB875185F3B85D" box="[367,434,1285,1311]" pageId="1" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
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(
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<figureCitation id="136E2A03FF8DFF9CFE7C87518673B85D" box="[456,562,1285,1311]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,255,871,895]" captionStartId-1="3.[151,255,1735,1759]" captionStartId-2="5.[151,255,1827,1851]" captionStartId-3="6.[151,255,1364,1388]" captionStartId-4="7.[151,255,1245,1269]" captionStartId-5="8.[151,255,1508,1532]" captionTargetBox-0="[169,1423,195,843]" captionTargetBox-1="[225,1357,197,1702]" captionTargetBox-2="[227,1382,201,1797]" captionTargetBox-3="[239,1343,428,1343]" captionTargetBox-4="[157,1376,211,1219]" captionTargetBox-5="[169,1423,199,1483]" captionTargetId-0="figure@2.[169,1425,195,844]" captionTargetId-1="figure@3.[219,1370,194,1712]" captionTargetId-2="figure@5.[205,1391,194,1806]" captionTargetId-3="figure@6.[231,1355,422,1353]" captionTargetId-4="figure@7.[151,1436,194,1222]" captionTargetId-5="figure@8.[169,1423,194,1484]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionText-0="FIGURE 1. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221), habitus in lateral view, in preservative." captionText-1="FIGURE 2. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view (setae omitted); C, posterior four pleonal somites and telson, lateral view (setae omitted); D, telson, dorsal view; E, left ocular appendage, antennule and antenna, dorsal view. Scale bars: 10 mm for A, C; 5 mm for B, D, E." captionText-2="FIGURE 3. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221), thoracic appendages in lateral view. A, left third maxilliped; B, left first pereopod; C, left second pereopod; D, right second pereopod; E, left third pereopod; F, left fourth pereopod; G, left fifth pereopod. Scale bar: 5 mm." captionText-3="FIGURE 4. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, ultimate segment of left third maxilliped, lateral view (setae partially omitted); B, ischium of left first pereopod, lateral view (setae omitted); C, dactylus and distal part of propodus of left third pereopod, lateral view; D, distal part of propodus of left third pereopod, ventral (flexor) view; E, proximal part of propodus and carpus of left third pereopod, mesial view; F, dactylus and distal part of propodus of left fourth pereopod, lateral view; G, dactylus and distal part of propodus of left fifth pereopod, lateral view. Scale bars: 2 mm." captionText-4="FIGURE 5. Pandalus ivanovi n. sp. A – C, paratype, male (CL 27.8 mm; VNIRO); D, E, paratype, male (CL 29.7 mm; same lot). A, carapace, lateral view; B, D, endopod of first pleopod; C, E, appendix masculina and appendix interna of second pleopod. Scale bars: 10 mm for A; 1 mm for B – E." captionText-5="FIGURE 6. Photographs showing coloration in fresh. A, Pandalus ivanovi n. sp., paratype, female (CL 31.4 mm; CBM- ZC 9222); B, Pandalus hypsinotus Brandt, 1851, female (CL 33.8 mm; CBM-ZC 9473), Canada." httpUri-0="https://zenodo.org/record/184239/files/figure.png" httpUri-1="https://zenodo.org/record/184240/files/figure.png" httpUri-2="https://zenodo.org/record/184241/files/figure.png" httpUri-3="https://zenodo.org/record/184242/files/figure.png" httpUri-4="https://zenodo.org/record/184243/files/figure.png" httpUri-5="https://zenodo.org/record/184244/files/figure.png" pageId="1" pageNumber="47">Figs 1–6</figureCitation>
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,
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<figureCitation id="136E2A03FF8DFF9CFDF48751860EB85D" box="[576,591,1285,1311]" captionStart="FIGURE 8" captionStartId="9.[151,255,1739,1763]" captionTargetBox="[240,1359,1077,1713]" captionTargetId="figure@9.[223,1367,1068,1716]" captionTargetPageId="9" captionText="FIGURE 8. Scatter plot of the relative length of the rostrum (indicated by rostral length / carapace length against carapace length) between Pandalus ivanovi n. sp. and P. hypsinotus Brandt, 1851." httpUri="https://zenodo.org/record/184246/files/figure.png" pageId="1" pageNumber="47">8</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34F650DFF8DFF9CFF23871E853ABB3E" pageId="1" pageNumber="47" type="materials_examined">
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<paragraph id="8BEA3686FF8DFF9CFF23871E853ABB3E" blockId="1.[151,1437,1354,2036]" pageId="1" pageNumber="47">
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<emphasis id="B921EA94FF8DFF9CFF23871E85C5B826" bold="true" box="[151,388,1354,1380]" pageId="1" pageNumber="47">Material examined</emphasis>
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.
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<typeStatus id="54EE8824FF8DFF9CFE20871E8647B826" box="[404,518,1354,1380]" pageId="1" pageNumber="47" type="holotype">Holotype</typeStatus>
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: CBM-ZC 9221, female (CL
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<quantity id="4CAD9B63FF8DFF9CFCD6871E8791B826" box="[866,976,1354,1380]" metricMagnitude="-2" metricUnit="m" metricValue="2.84" pageId="1" pageNumber="47" unit="mm" value="28.4">28.4 mm</quantity>
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), off eastern Sakhalin, Sea of Okhotsk, 46°N, 145°E, 2005, 150–
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<quantity id="4CAD9B63FF8DFF9CFE6487268662B8CE" box="[464,547,1394,1420]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="47" unit="m" value="200.0">200 m</quantity>
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, rocky bottom, commercial fishing using shrimp basket, S. Miyaki leg.
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<typeStatus id="54EE8824FF8DFF9CFF2387CE854FB8F6" box="[151,270,1434,1460]" pageId="1" pageNumber="47" type="paratype">Paratypes</typeStatus>
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: CBM-ZC 9222,
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<specimenCount id="9D53FD0FFF8DFF9CFE6D87CE860BB8F6" box="[473,586,1434,1460]" pageId="1" pageNumber="47" type="female">2 females</specimenCount>
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(CL 31.0,
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<quantity id="4CAD9B63FF8DFF9CFD7D87CE8777B8F6" box="[713,822,1434,1460]" metricMagnitude="-2" metricUnit="m" metricValue="3.1399999999999997" pageId="1" pageNumber="47" unit="mm" value="31.4">31.4 mm</quantity>
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), same data as
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<typeStatus id="54EE8824FF8DFF9CFC5187CE800EB8F6" box="[997,1103,1434,1460]" pageId="1" pageNumber="47" type="holotype">holotype</typeStatus>
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; CBM-ZC 9223,
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<specimenCount id="9D53FD0FFF8DFF9CFAA887CE81DAB8F6" box="[1308,1435,1434,1460]" pageId="1" pageNumber="47" type="female">20 females</specimenCount>
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(CL
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<quantity id="4CAD9B63FF8DFF9CFF65879685C5B89E" box="[209,388,1474,1500]" metricMagnitude="-2" metricUnit="m" metricValue="3.29" metricValueMax="3.56" metricValueMin="3.02" pageId="1" pageNumber="47" unit="mm" value="32.9" valueMax="35.6" valueMin="30.2">30.2–35.6 mm</quantity>
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), same data as
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<typeStatus id="54EE8824FF8DFF9CFDF7879686ECB89E" box="[579,685,1474,1500]" pageId="1" pageNumber="47" type="holotype">holotype</typeStatus>
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; MNHN-Na 16873, same data as
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<typeStatus id="54EE8824FF8DFF9CFBE4879680FBB89E" box="[1104,1210,1474,1500]" pageId="1" pageNumber="47" type="holotype">holotype</typeStatus>
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; NSMT-Cr 18306, same data as
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<typeStatus id="54EE8824FF8DFF9CFE8E87BE85E5BB46" box="[314,420,1514,1540]" pageId="1" pageNumber="47" type="holotype">holotype</typeStatus>
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; CBM-ZC 9224,
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<specimenCount id="9D53FD0FFF8DFF9CFDCE87BE86AFBB46" box="[634,750,1514,1540]" pageId="1" pageNumber="47" type="female">4 females</specimenCount>
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(CL
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<quantity id="4CAD9B63FF8DFF9CFC8587BE87A2BB46" box="[817,995,1514,1540]" metricMagnitude="-2" metricUnit="m" metricValue="3.185" metricValueMax="3.31" metricValueMin="3.06" pageId="1" pageNumber="47" unit="mm" value="31.85" valueMax="33.1" valueMin="30.6">30.6–33.1 mm</quantity>
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), similar locality, 2005, M. Fujiwara leg.;
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<collectionCode id="ED44AE43FF8DFF9CFF6684468573BB6E" box="[210,306,1554,1580]" httpUri="http://grbio.org/cool/kxi4-27ep" name="Institute of Oceanography" pageId="1" pageNumber="47">VNIRO</collectionCode>
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, 19 transitional males (CL 27.6–30.0 mm),
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<specimenCount id="9D53FD0FFF8DFF9CFC8D844687EBBB6E" box="[825,938,1554,1580]" pageId="1" pageNumber="47" type="female">5 females</specimenCount>
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(CL 29.0–
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<quantity id="4CAD9B63FF8DFF9CFB99844680DBBB6E" box="[1069,1178,1554,1580]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" pageId="1" pageNumber="47" unit="mm" value="31.5">31.5 mm</quantity>
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), 3 ovigerous females (CL
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<quantity id="4CAD9B63FF8DFF9CFF79846E853ABB16" box="[205,379,1594,1620]" metricMagnitude="-2" metricUnit="m" metricValue="3.06" metricValueMax="3.3899999999999997" metricValueMin="2.73" pageId="1" pageNumber="47" unit="mm" value="30.6" valueMax="33.9" valueMin="27.3">27.3–33.9 mm</quantity>
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), off eastern Sakhalin,
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<geoCoordinate id="EE615041FF8DFF9CFD31846E86ABBB16" box="[645,746,1594,1620]" direction="north" orientation="latitude" pageId="1" pageNumber="47" precision="925" value="49.25">49°15’N</geoCoordinate>
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<geoCoordinate id="EE615041FF8DFF9CFD45846E8720BB16" box="[753,865,1594,1620]" direction="east" orientation="longitude" pageId="1" pageNumber="47" precision="925" value="144.58333">144°35’E</geoCoordinate>
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to
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<geoCoordinate id="EE615041FF8DFF9CFC31846E87ABBB16" box="[901,1002,1594,1620]" direction="north" orientation="latitude" pageId="1" pageNumber="47" precision="925" value="49.25">49°15’N</geoCoordinate>
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<geoCoordinate id="EE615041FF8DFF9CFC45846E8025BB16" box="[1009,1124,1594,1620]" direction="east" orientation="longitude" pageId="1" pageNumber="47" precision="925" value="144.65">144°39’E</geoCoordinate>
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,
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<quantity id="4CAD9B63FF8DFF9CFBC4846E80A9BB16" box="[1136,1256,1594,1620]" metricMagnitude="2" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="1" pageNumber="47" unit="m" value="105.0" valueMax="120.0" valueMin="90.0">90–120 m</quantity>
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, FRV
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<emphasis id="B921EA94FF8DFF9CFA85846F81DBBB16" box="[1329,1434,1595,1620]" italics="true" pageId="1" pageNumber="47">Viktoriya</emphasis>
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2,
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<date id="FFEB1046FF8DFF9CFF0084368536BB3E" box="[180,375,1634,1660]" pageId="1" pageNumber="47" value="2001-09">September 2001</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34F650DFF8DFF99FF7184DE81D1BBE0" lastPageId="4" lastPageNumber="50" pageId="1" pageNumber="47" type="description">
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<paragraph id="8BEA3686FF8DFF9CFF7184DE85A5BAB6" blockId="1.[151,1437,1354,2036]" pageId="1" pageNumber="47">
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<emphasis id="B921EA94FF8DFF9CFF7184DE851CBBE6" bold="true" box="[197,349,1674,1700]" pageId="1" pageNumber="47">Description.</emphasis>
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<emphasis id="B921EA94FF8DFF9CFED284DF8588BBE6" box="[358,457,1675,1700]" italics="true" pageId="1" pageNumber="47">Females</emphasis>
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. Body (
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<figureCitation id="136E2A03FF8DFF9CFD9E84DE862FBBE6" box="[554,622,1674,1700]" captionStart="FIGURE 1" captionStartId="2.[151,255,871,895]" captionTargetBox="[169,1423,195,843]" captionTargetId="figure@2.[169,1425,195,844]" captionTargetPageId="2" captionText="FIGURE 1. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221), habitus in lateral view, in preservative." httpUri="https://zenodo.org/record/184239/files/figure.png" pageId="1" pageNumber="47">Fig. 1</figureCitation>
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) moderately robust. Rostrum (
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<figureCitation id="136E2A03FF8DFF9CFC6984DE8065BBE6" box="[989,1060,1674,1700]" captionStart="FIGURE 2" captionStartId="3.[151,255,1735,1759]" captionTargetBox="[225,1357,197,1702]" captionTargetId="figure@3.[219,1370,194,1712]" captionTargetPageId="3" captionText="FIGURE 2. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view (setae omitted); C, posterior four pleonal somites and telson, lateral view (setae omitted); D, telson, dorsal view; E, left ocular appendage, antennule and antenna, dorsal view. Scale bars: 10 mm for A, C; 5 mm for B, D, E." httpUri="https://zenodo.org/record/184240/files/figure.png" pageId="1" pageNumber="47">Fig. 2</figureCitation>
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A, B) strongly curving dorsally, overreaching distal margin of antennal scale, 1.02–1.25 times as long as carapace; dorsal margin armed with 16–24 movable spines, including 9–14 on rostrum proper and 6–9 on carapace posterior to orbital margin, and 1 or 2 small fixed teeth near apex of rostrum, subdistal 0.50–0.60 leaving unarmed, posteriormost (first) spine arising at posterior to midlength of carapace (0.55–0.59 of carapace length); ventral margin armed with 7–9 fixed teeth increasing in size posteriorly, posteriormost tooth distinctly stronger than others; lateral carina conspicuous in posterior half. Carapace (
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<figureCitation id="136E2A03FF8DFF9CFDE0853E86DABAC6" box="[596,667,1898,1924]" captionStart="FIGURE 2" captionStartId="3.[151,255,1735,1759]" captionTargetBox="[225,1357,197,1702]" captionTargetId="figure@3.[219,1370,194,1712]" captionTargetPageId="3" captionText="FIGURE 2. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view (setae omitted); C, posterior four pleonal somites and telson, lateral view (setae omitted); D, telson, dorsal view; E, left ocular appendage, antennule and antenna, dorsal view. Scale bars: 10 mm for A, C; 5 mm for B, D, E." httpUri="https://zenodo.org/record/184240/files/figure.png" pageId="1" pageNumber="47">Fig. 2</figureCitation>
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A) with postrostral ridge moderately high, somewhat crested but non-arched, extending nearly to posterodorsal margin; dorsal margin gently convex in lateral view, posteriormost 1–4 spines on peak; antennal tooth moderately strong; branchiostegal tooth small; no conspicuous ridges or carinae on lateral surface.
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</paragraph>
|
||
<caption id="DF2A660EFF8EFF9FFF238133849EBEE3" httpUri="https://zenodo.org/record/184239/files/figure.png" pageId="2" pageNumber="48" targetBox="[169,1423,195,843]" targetPageId="2">
|
||
<paragraph id="8BEA3686FF8EFF9FFF238133849EBEE3" blockId="2.[151,1436,871,929]" pageId="2" pageNumber="48">
|
||
<emphasis id="B921EA94FF8EFF9FFF2381338558BE3D" bold="true" box="[151,281,871,895]" pageId="2" pageNumber="48">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C554D05FF8EFF9FFE95813C859ABE3D" box="[289,475,872,895]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="48" phylum="Arthropoda" rank="species" species="ivanovi" status="sp. nov.">
|
||
<emphasis id="B921EA94FF8EFF9FFE95813C859ABE3D" box="[289,475,872,895]" italics="true" pageId="2" pageNumber="48">Pandalus ivanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257EFFF8EFF9FFE56813C865DBE3D" box="[482,540,872,895]" pageId="2" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
|
||
Holotype, female (CL 28.4 mm; CBM-ZC 9221), habitus in lateral view, in preservative.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEA3686FF8EFF9FFF7181818055B9CD" blockId="2.[151,1437,981,1927]" pageId="2" pageNumber="48">
|
||
Pleon (
|
||
<figureCitation id="136E2A03FF8EFF9FFEAD81818527BEAD" box="[281,358,981,1007]" captionStart="FIGURE 1" captionStartId="2.[151,255,871,895]" captionTargetBox="[169,1423,195,843]" captionTargetId="figure@2.[169,1425,195,844]" captionTargetPageId="2" captionText="FIGURE 1. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221), habitus in lateral view, in preservative." httpUri="https://zenodo.org/record/184239/files/figure.png" pageId="2" pageNumber="48">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="136E2A03FF8EFF9FFEC2818185C4BEAD" box="[374,389,981,1007]" captionStart="FIGURE 2" captionStartId="3.[151,255,1735,1759]" captionTargetBox="[225,1357,197,1702]" captionTargetId="figure@3.[219,1370,194,1712]" captionTargetPageId="3" captionText="FIGURE 2. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view (setae omitted); C, posterior four pleonal somites and telson, lateral view (setae omitted); D, telson, dorsal view; E, left ocular appendage, antennule and antenna, dorsal view. Scale bars: 10 mm for A, C; 5 mm for B, D, E." httpUri="https://zenodo.org/record/184240/files/figure.png" pageId="2" pageNumber="48">2</figureCitation>
|
||
C) dorsally smooth. Third somite without middorsal projection or posterodorsal median tooth. Fourth and fifth somites each with posterolateral tooth on pleuron. Sixth somite about 0.40 times as long as carapace and 1.70–1.80 times longer than high; posterolateral process terminating in acute tooth. Telson (
|
||
<figureCitation id="136E2A03FF8EFF9FFF7B86198554B925" box="[207,277,1101,1127]" captionStart="FIGURE 2" captionStartId="3.[151,255,1735,1759]" captionTargetBox="[225,1357,197,1702]" captionTargetId="figure@3.[219,1370,194,1712]" captionTargetPageId="3" captionText="FIGURE 2. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view (setae omitted); C, posterior four pleonal somites and telson, lateral view (setae omitted); D, telson, dorsal view; E, left ocular appendage, antennule and antenna, dorsal view. Scale bars: 10 mm for A, C; 5 mm for B, D, E." httpUri="https://zenodo.org/record/184240/files/figure.png" pageId="2" pageNumber="48">Fig. 2</figureCitation>
|
||
D) moderately broad, armed normally with 5 or 6 pairs of dorsolateral spines; posterior margin terminating in small blunt triangular projection, with 3 pairs of unequal spines.
|
||
</paragraph>
|
||
<paragraph id="8BEA3686FF8EFF9FFF7186C984B5B99D" blockId="2.[151,1437,981,1927]" pageId="2" pageNumber="48">
|
||
Eye (
|
||
<figureCitation id="136E2A03FF8EFF9FFEB786C9850BB9F5" box="[259,330,1181,1207]" captionStart="FIGURE 2" captionStartId="3.[151,255,1735,1759]" captionTargetBox="[225,1357,197,1702]" captionTargetId="figure@3.[219,1370,194,1712]" captionTargetPageId="3" captionText="FIGURE 2. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view (setae omitted); C, posterior four pleonal somites and telson, lateral view (setae omitted); D, telson, dorsal view; E, left ocular appendage, antennule and antenna, dorsal view. Scale bars: 10 mm for A, C; 5 mm for B, D, E." httpUri="https://zenodo.org/record/184240/files/figure.png" pageId="2" pageNumber="48">Fig. 2</figureCitation>
|
||
A, E) broadly subpyriform, maximal diameter of cornea 0.17–0.20 of carapace length; ocellus present.
|
||
</paragraph>
|
||
<paragraph id="8BEA3686FF8EFF9FFF7186B9857EB83D" blockId="2.[151,1437,981,1927]" pageId="2" pageNumber="48">
|
||
Antennular peduncle (
|
||
<figureCitation id="136E2A03FF8EFF9FFE7F86B98650B845" box="[459,529,1261,1287]" captionStart="FIGURE 2" captionStartId="3.[151,255,1735,1759]" captionTargetBox="[225,1357,197,1702]" captionTargetId="figure@3.[219,1370,194,1712]" captionTargetPageId="3" captionText="FIGURE 2. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view (setae omitted); C, posterior four pleonal somites and telson, lateral view (setae omitted); D, telson, dorsal view; E, left ocular appendage, antennule and antenna, dorsal view. Scale bars: 10 mm for A, C; 5 mm for B, D, E." httpUri="https://zenodo.org/record/184240/files/figure.png" pageId="2" pageNumber="48">Fig. 2</figureCitation>
|
||
A, E) reaching midlength of antennal scale; first segment with short, rounded stylocerite; second segment unarmed on dorsal surface, but with few spinules on dorsodistal margin; distal 2 segments slightly shorter than first segment; aesthetasc-bearing portion of outer flagellum distinctly shorter than carapace.
|
||
</paragraph>
|
||
<paragraph id="8BEA3686FF8EFF9FFF7187D985BBBB5D" blockId="2.[151,1437,981,1927]" pageId="2" pageNumber="48">
|
||
Antennal (
|
||
<figureCitation id="136E2A03FF8EFF9FFEF187D985D1B8E5" box="[325,400,1421,1447]" captionStart="FIGURE 2" captionStartId="3.[151,255,1735,1759]" captionTargetBox="[225,1357,197,1702]" captionTargetId="figure@3.[219,1370,194,1712]" captionTargetPageId="3" captionText="FIGURE 2. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view (setae omitted); C, posterior four pleonal somites and telson, lateral view (setae omitted); D, telson, dorsal view; E, left ocular appendage, antennule and antenna, dorsal view. Scale bars: 10 mm for A, C; 5 mm for B, D, E." httpUri="https://zenodo.org/record/184240/files/figure.png" pageId="2" pageNumber="48">Fig. 2</figureCitation>
|
||
A, E) with stout basicerite bearing moderately strong ventrolateral tooth; carpocerite moderately stout, far falling short of midlength of antennal scale; antennal scale 0.82–0.88 times as long as carapace and 4.20–4.50 times longer than broad, lateral margin nearly straight, distolateral tooth reaching distal margin of rounded lamella.
|
||
</paragraph>
|
||
<paragraph id="8BEA3686FF8EFF9FFF71847985D0BAC5" blockId="2.[151,1437,981,1927]" pageId="2" pageNumber="48">
|
||
Mouthparts usual of genus (see
|
||
<bibRefCitation id="EFC44B77FF8EFF9FFDF3847986A6BB05" author="Komai" box="[583,743,1581,1607]" pageId="2" pageNumber="56" refString="Komai, T. (1999) A revision of the genus Pandalus (Crustacea: Decapoda: Caridea: Pandalidae). Journal of Natural History, 33, 1265 - 1372." type="journal article" year="1999">Komai, 1999</bibRefCitation>
|
||
). Third maxilliped (
|
||
<figureCitation id="136E2A03FF8EFF9FFC6384798065BB05" box="[983,1060,1581,1607]" captionStart="FIGURE 1" captionStartId="2.[151,255,871,895]" captionTargetBox="[169,1423,195,843]" captionTargetId="figure@2.[169,1425,195,844]" captionTargetPageId="2" captionText="FIGURE 1. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221), habitus in lateral view, in preservative." httpUri="https://zenodo.org/record/184239/files/figure.png" pageId="2" pageNumber="48">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="136E2A03FF8EFF9FFB8184798002BB05" box="[1077,1091,1581,1607]" captionStart="FIGURE 3" captionStartId="5.[151,255,1827,1851]" captionTargetBox="[227,1382,201,1797]" captionTargetId="figure@5.[205,1391,194,1806]" captionTargetPageId="5" captionText="FIGURE 3. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221), thoracic appendages in lateral view. A, left third maxilliped; B, left first pereopod; C, left second pereopod; D, right second pereopod; E, left third pereopod; F, left fourth pereopod; G, left fifth pereopod. Scale bar: 5 mm." httpUri="https://zenodo.org/record/184241/files/figure.png" pageId="2" pageNumber="48">3</figureCitation>
|
||
A, 4A) slightly overreaching distal margin of antennal scale; ultimate segment about 1.80 as long as penultimate segment (= carpus), terminating in small corneous spine, with numerous scattered spinules and several tufts of short setae on lateral surface and 2 or 3 tufts of longer spinules on dorsal margin subdistally; mesial face of ultimate segment with numerous transverse tracts of stiff setae and few long spines; carpus also with several spinules on lateral surface and numerous stiff setae on mesial surface; antepenultimate segment (merus-ischium-basis fused segment) slightly shorter than distal 2 segments combined, with small rounded tubercle basally on dorsal surface; dorsal surface elevated at midlength; ventral margin forming blunt edge fringed with row of numerous short setae; exopod absent.
|
||
</paragraph>
|
||
<caption id="DF2A660EFF8FFF9EFF23849385FFBA04" httpUri="https://zenodo.org/record/184240/files/figure.png" pageId="3" pageNumber="49" targetBox="[225,1357,197,1702]" targetPageId="3">
|
||
<paragraph id="8BEA3686FF8FFF9EFF23849385FFBA04" blockId="3.[151,1436,1735,1862]" pageId="3" pageNumber="49">
|
||
<emphasis id="B921EA94FF8FFF9EFF2384938558BB9D" bold="true" box="[151,281,1735,1759]" pageId="3" pageNumber="49">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C554D05FF8FFF9EFE94849C859BBB9D" box="[288,474,1736,1759]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="49" phylum="Arthropoda" rank="species" species="ivanovi" status="sp. nov.">
|
||
<emphasis id="B921EA94FF8FFF9EFE94849C859BBB9D" box="[288,474,1736,1759]" italics="true" pageId="3" pageNumber="49">Pandalus ivanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257EFFF8FFF9EFE5484938658BB9C" box="[480,537,1735,1758]" pageId="3" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
|
||
Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view (setae omitted); C, posterior four pleonal somites and telson, lateral view (setae omitted); D, telson, dorsal view; E, left ocular appendage, antennule and antenna, dorsal view. Scale bars: 10 mm for A, C; 5 mm for B, D, E.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEA3686FF8FFF99FF71852E85C7B908" blockId="3.[151,1436,1914,2020]" lastBlockId="4.[151,1437,152,2018]" lastPageId="4" lastPageNumber="50" pageId="3" pageNumber="49">
|
||
First pereopod (
|
||
<figureCitation id="136E2A03FF8FFF9EFECB852E8582BAD6" box="[383,451,1914,1940]" captionStart="FIGURE 3" captionStartId="5.[151,255,1827,1851]" captionTargetBox="[227,1382,201,1797]" captionTargetId="figure@5.[205,1391,194,1806]" captionTargetPageId="5" captionText="FIGURE 3. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221), thoracic appendages in lateral view. A, left third maxilliped; B, left first pereopod; C, left second pereopod; D, right second pereopod; E, left third pereopod; F, left fourth pereopod; G, left fifth pereopod. Scale bar: 5 mm." httpUri="https://zenodo.org/record/184241/files/figure.png" pageId="3" pageNumber="49">Fig. 3</figureCitation>
|
||
B) minutely chelate, slender, slightly falling short of distal margin of antennal scale; propodus about 0.60 times as long as carpus, tapering distally, with distinct longitudinal groove lined with tufts of short setae; ischium with narrow ventral lamina terminating acutely or subacutely at ventrodistal angle; row of spinules partially obscured by stiff setae present on ventral margin of ischium. Second pereopods greatly unequal with left much longer and slender than right. Left second pereopod (
|
||
<figureCitation id="136E2A03FF88FF99FB6F82948163BD98" box="[1243,1314,192,218]" captionStart="FIGURE 3" captionStartId="5.[151,255,1827,1851]" captionTargetBox="[227,1382,201,1797]" captionTargetId="figure@5.[205,1391,194,1806]" captionTargetPageId="5" captionText="FIGURE 3. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221), thoracic appendages in lateral view. A, left third maxilliped; B, left first pereopod; C, left second pereopod; D, right second pereopod; E, left third pereopod; F, left fourth pereopod; G, left fifth pereopod. Scale bar: 5 mm." httpUri="https://zenodo.org/record/184241/files/figure.png" pageId="4" pageNumber="50">Fig. 3</figureCitation>
|
||
C) greatly elongate, overreaching antennal scale by 0.80–0.90 length of carpus; chela small, dactylus slightly shorter than palm; carpus elongate, divided in 54–61 articles; merus entirely annulated; ischium distally annulated, ventral margin slightly expanded in proximal half to accommodate chela. Right second pereopod (
|
||
<figureCitation id="136E2A03FF88FF99FA81836C813DBC10" box="[1333,1404,312,338]" captionStart="FIGURE 3" captionStartId="5.[151,255,1827,1851]" captionTargetBox="[227,1382,201,1797]" captionTargetId="figure@5.[205,1391,194,1806]" captionTargetPageId="5" captionText="FIGURE 3. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221), thoracic appendages in lateral view. A, left third maxilliped; B, left first pereopod; C, left second pereopod; D, right second pereopod; E, left third pereopod; F, left fourth pereopod; G, left fifth pereopod. Scale bar: 5 mm." httpUri="https://zenodo.org/record/184241/files/figure.png" pageId="4" pageNumber="50">Fig. 3</figureCitation>
|
||
D) overreaching antennal scale by 0.3 length of carpus; chela about 0.25 times as long as carpus, dactylus shorter than palm; carpus divided in 19–24 articles; merus with few annulations distally; ischium entire, similar to left in structure. Third to fifth pereopods decreasing in length toward posterior. Third pereopod (
|
||
<figureCitation id="136E2A03FF88FF99FB4583E48176BC88" box="[1265,1335,432,458]" captionStart="FIGURE 3" captionStartId="5.[151,255,1827,1851]" captionTargetBox="[227,1382,201,1797]" captionTargetId="figure@5.[205,1391,194,1806]" captionTargetPageId="5" captionText="FIGURE 3. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221), thoracic appendages in lateral view. A, left third maxilliped; B, left first pereopod; C, left second pereopod; D, right second pereopod; E, left third pereopod; F, left fourth pereopod; G, left fifth pereopod. Scale bar: 5 mm." httpUri="https://zenodo.org/record/184241/files/figure.png" pageId="4" pageNumber="50">Fig. 3</figureCitation>
|
||
E) overreaching antennal scale by 0.5–0.6 length of propodus, subprehensile; dactylus (
|
||
<figureCitation id="136E2A03FF88FF99FBF5838C80C6BCB0" box="[1089,1159,472,498]" captionStart="FIGURE 4" captionStartId="6.[151,255,1364,1388]" captionTargetBox="[239,1343,428,1343]" captionTargetId="figure@6.[231,1355,422,1353]" captionTargetPageId="6" captionText="FIGURE 4. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, ultimate segment of left third maxilliped, lateral view (setae partially omitted); B, ischium of left first pereopod, lateral view (setae omitted); C, dactylus and distal part of propodus of left third pereopod, lateral view; D, distal part of propodus of left third pereopod, ventral (flexor) view; E, proximal part of propodus and carpus of left third pereopod, mesial view; F, dactylus and distal part of propodus of left fourth pereopod, lateral view; G, dactylus and distal part of propodus of left fifth pereopod, lateral view. Scale bars: 2 mm." httpUri="https://zenodo.org/record/184242/files/figure.png" pageId="4" pageNumber="50">Fig. 4</figureCitation>
|
||
C) sickle-shaped, 0.32– 0.47 times as long as propodus, with 22–27 accessory spinules becoming longer and more widely spaced distally and 1 subterminal spinule appressed to slender unguis; propodus slightly recurved, laterally compressed, all surfaces with scattered spinules, particularly flexor surface with numerous scattered spinules flanked by rows of long spinules (spinules becoming fewer proximally) (
|
||
<figureCitation id="136E2A03FF88FF99FCDC802C87EFBFD0" box="[872,942,632,658]" captionStart="FIGURE 4" captionStartId="6.[151,255,1364,1388]" captionTargetBox="[239,1343,428,1343]" captionTargetId="figure@6.[231,1355,422,1353]" captionTargetPageId="6" captionText="FIGURE 4. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, ultimate segment of left third maxilliped, lateral view (setae partially omitted); B, ischium of left first pereopod, lateral view (setae omitted); C, dactylus and distal part of propodus of left third pereopod, lateral view; D, distal part of propodus of left third pereopod, ventral (flexor) view; E, proximal part of propodus and carpus of left third pereopod, mesial view; F, dactylus and distal part of propodus of left fourth pereopod, lateral view; G, dactylus and distal part of propodus of left fifth pereopod, lateral view. Scale bars: 2 mm." httpUri="https://zenodo.org/record/184242/files/figure.png" pageId="4" pageNumber="50">Fig. 4</figureCitation>
|
||
D); carpus 0.50–0.60 times as long as propodus, with 2 or 3 lateral and 1 ventromesial spines and with scattered spinules on lateral to mesial faces (
|
||
<figureCitation id="136E2A03FF88FF99FAD980F484E6BFA0" captionStart="FIGURE 4" captionStartId="6.[151,255,1364,1388]" captionTargetBox="[239,1343,428,1343]" captionTargetId="figure@6.[231,1355,422,1353]" captionTargetPageId="6" captionText="FIGURE 4. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, ultimate segment of left third maxilliped, lateral view (setae partially omitted); B, ischium of left first pereopod, lateral view (setae omitted); C, dactylus and distal part of propodus of left third pereopod, lateral view; D, distal part of propodus of left third pereopod, ventral (flexor) view; E, proximal part of propodus and carpus of left third pereopod, mesial view; F, dactylus and distal part of propodus of left fourth pereopod, lateral view; G, dactylus and distal part of propodus of left fifth pereopod, lateral view. Scale bars: 2 mm." httpUri="https://zenodo.org/record/184242/files/figure.png" pageId="4" pageNumber="50">Fig. 4</figureCitation>
|
||
E); merus 0.86–0.94 times as long as carapace, armed with 7–9 lateral and 6–9 ventral spines; ischium with 1 ventrolateral spine. Fourth pereopod (
|
||
<figureCitation id="136E2A03FF88FF99FDC480A486F7BE48" box="[624,694,752,778]" captionStart="FIGURE 3" captionStartId="5.[151,255,1827,1851]" captionTargetBox="[227,1382,201,1797]" captionTargetId="figure@5.[205,1391,194,1806]" captionTargetPageId="5" captionText="FIGURE 3. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221), thoracic appendages in lateral view. A, left third maxilliped; B, left first pereopod; C, left second pereopod; D, right second pereopod; E, left third pereopod; F, left fourth pereopod; G, left fifth pereopod. Scale bar: 5 mm." httpUri="https://zenodo.org/record/184241/files/figure.png" pageId="4" pageNumber="50">Fig. 3</figureCitation>
|
||
F) overreaching antennal scale by length of dactylus; dactylus (
|
||
<figureCitation id="136E2A03FF88FF99FF2B814C84A7BE70" box="[159,230,792,818]" captionStart="FIGURE 4" captionStartId="6.[151,255,1364,1388]" captionTargetBox="[239,1343,428,1343]" captionTargetId="figure@6.[231,1355,422,1353]" captionTargetPageId="6" captionText="FIGURE 4. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, ultimate segment of left third maxilliped, lateral view (setae partially omitted); B, ischium of left first pereopod, lateral view (setae omitted); C, dactylus and distal part of propodus of left third pereopod, lateral view; D, distal part of propodus of left third pereopod, ventral (flexor) view; E, proximal part of propodus and carpus of left third pereopod, mesial view; F, dactylus and distal part of propodus of left fourth pereopod, lateral view; G, dactylus and distal part of propodus of left fifth pereopod, lateral view. Scale bars: 2 mm." httpUri="https://zenodo.org/record/184242/files/figure.png" pageId="4" pageNumber="50">Fig. 4</figureCitation>
|
||
F) 0.22–0.34 times as long as propodus, armed with 17–21 accessory and 1 subterminal spinules; propodus nearly straight, less spinose than third; carpus with 3 lateral and 1 ventral spines and scattered spinules on lateral to mesial faces; merus 0.72–0.79 times as long as carapace, with 5–8 lateral and 5–8 ventral spines; ischium with 1 ventral spine. Fifth pereopod (
|
||
<figureCitation id="136E2A03FF88FF99FD1E81C486AFBEE8" box="[682,750,912,938]" captionStart="FIGURE 3" captionStartId="5.[151,255,1827,1851]" captionTargetBox="[227,1382,201,1797]" captionTargetId="figure@5.[205,1391,194,1806]" captionTargetPageId="5" captionText="FIGURE 3. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221), thoracic appendages in lateral view. A, left third maxilliped; B, left first pereopod; C, left second pereopod; D, right second pereopod; E, left third pereopod; F, left fourth pereopod; G, left fifth pereopod. Scale bar: 5 mm." httpUri="https://zenodo.org/record/184241/files/figure.png" pageId="4" pageNumber="50">Fig. 3</figureCitation>
|
||
G) falling short of distal margin of antennal scale; dactylus (
|
||
<figureCitation id="136E2A03FF88FF99FF2B81EC84A4BE90" box="[159,229,952,978]" captionStart="FIGURE 4" captionStartId="6.[151,255,1364,1388]" captionTargetBox="[239,1343,428,1343]" captionTargetId="figure@6.[231,1355,422,1353]" captionTargetPageId="6" captionText="FIGURE 4. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, ultimate segment of left third maxilliped, lateral view (setae partially omitted); B, ischium of left first pereopod, lateral view (setae omitted); C, dactylus and distal part of propodus of left third pereopod, lateral view; D, distal part of propodus of left third pereopod, ventral (flexor) view; E, proximal part of propodus and carpus of left third pereopod, mesial view; F, dactylus and distal part of propodus of left fourth pereopod, lateral view; G, dactylus and distal part of propodus of left fifth pereopod, lateral view. Scale bars: 2 mm." httpUri="https://zenodo.org/record/184242/files/figure.png" pageId="4" pageNumber="50">Fig. 4</figureCitation>
|
||
G) short, somewhat curved, 0.15–0.22 times as long as propodus, with 9–12 accessory and 1 subterminal spinules; propodus with 2 rows of long spinules on flexor surface and with tufts of grooming setae distally (
|
||
<figureCitation id="136E2A03FF88FF99FF2B865C84A8B960" box="[159,233,1032,1058]" captionStart="FIGURE 4" captionStartId="6.[151,255,1364,1388]" captionTargetBox="[239,1343,428,1343]" captionTargetId="figure@6.[231,1355,422,1353]" captionTargetPageId="6" captionText="FIGURE 4. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, ultimate segment of left third maxilliped, lateral view (setae partially omitted); B, ischium of left first pereopod, lateral view (setae omitted); C, dactylus and distal part of propodus of left third pereopod, lateral view; D, distal part of propodus of left third pereopod, ventral (flexor) view; E, proximal part of propodus and carpus of left third pereopod, mesial view; F, dactylus and distal part of propodus of left fourth pereopod, lateral view; G, dactylus and distal part of propodus of left fifth pereopod, lateral view. Scale bars: 2 mm." httpUri="https://zenodo.org/record/184242/files/figure.png" pageId="4" pageNumber="50">Fig. 4</figureCitation>
|
||
G); carpus with scattered spinules on dorsal surface and with 2 lateral and 1 ventral spines; ischium with 1 ventral spine.
|
||
</paragraph>
|
||
<paragraph id="8BEA3686FF88FF99FF71860C8077B9D8" blockId="4.[151,1437,152,2018]" pageId="4" pageNumber="50">
|
||
First pleopod with simple endopod. Second pleopod completely lacking trace of appendix masculina. Uropod (
|
||
<figureCitation id="136E2A03FF88FF99FEB486D48507B9D8" box="[256,326,1152,1178]" captionStart="FIGURE 2" captionStartId="3.[151,255,1735,1759]" captionTargetBox="[225,1357,197,1702]" captionTargetId="figure@3.[219,1370,194,1712]" captionTargetPageId="3" captionText="FIGURE 2. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view (setae omitted); C, posterior four pleonal somites and telson, lateral view (setae omitted); D, telson, dorsal view; E, left ocular appendage, antennule and antenna, dorsal view. Scale bars: 10 mm for A, C; 5 mm for B, D, E." httpUri="https://zenodo.org/record/184240/files/figure.png" pageId="4" pageNumber="50">Fig. 2</figureCitation>
|
||
C) with rami subequal in length, reaching nearly to tip of telson.
|
||
</paragraph>
|
||
<paragraph id="8BEA3686FF88FF99FF7186FC808EB980" blockId="4.[151,1437,152,2018]" box="[197,1231,1192,1218]" pageId="4" pageNumber="50">
|
||
Eggs numerous; size 0.83–1.00 mm along short axis and 1.00–
|
||
<quantity id="4CAD9B63FF88FF99FC1286FC804CB980" box="[934,1037,1192,1218]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="4" pageNumber="50" unit="mm" value="1.17">1.17 mm</quantity>
|
||
along long axis.
|
||
</paragraph>
|
||
<paragraph id="8BEA3686FF88FF99FF7186848771B820" blockId="4.[151,1437,152,2018]" pageId="4" pageNumber="50">
|
||
<emphasis id="B921EA94FF88FF99FF71868485E2B9AB" box="[197,419,1232,1257]" italics="true" pageId="4" pageNumber="50">Transitional males</emphasis>
|
||
. Generally similar to females (
|
||
<figureCitation id="136E2A03FF88FF99FCAA86848727B9A8" box="[798,870,1232,1258]" captionStart="FIGURE 5" captionStartId="7.[151,255,1245,1269]" captionTargetBox="[157,1376,211,1219]" captionTargetId="figure@7.[151,1436,194,1222]" captionTargetPageId="7" captionText="FIGURE 5. Pandalus ivanovi n. sp. A – C, paratype, male (CL 27.8 mm; VNIRO); D, E, paratype, male (CL 29.7 mm; same lot). A, carapace, lateral view; B, D, endopod of first pleopod; C, E, appendix masculina and appendix interna of second pleopod. Scale bars: 10 mm for A; 1 mm for B – E." httpUri="https://zenodo.org/record/184243/files/figure.png" pageId="4" pageNumber="50">Fig. 5</figureCitation>
|
||
A). Rostrum 1.08–1.33 times longer than carapace. Endopod of first pleopod (
|
||
<figureCitation id="136E2A03FF88FF99FD9186AC8631B850" box="[549,624,1272,1298]" captionStart="FIGURE 5" captionStartId="7.[151,255,1245,1269]" captionTargetBox="[157,1376,211,1219]" captionTargetId="figure@7.[151,1436,194,1222]" captionTargetPageId="7" captionText="FIGURE 5. Pandalus ivanovi n. sp. A – C, paratype, male (CL 27.8 mm; VNIRO); D, E, paratype, male (CL 29.7 mm; same lot). A, carapace, lateral view; B, D, endopod of first pleopod; C, E, appendix masculina and appendix interna of second pleopod. Scale bars: 10 mm for A; 1 mm for B – E." httpUri="https://zenodo.org/record/184243/files/figure.png" pageId="4" pageNumber="50">Fig. 5</figureCitation>
|
||
B, D) with subterminal appendix interna; proximomesial lobe well delineated. Appendix masculina on second pleopod (
|
||
<figureCitation id="136E2A03FF88FF99FCBD87748711B878" box="[777,848,1312,1338]" captionStart="FIGURE 5" captionStartId="7.[151,255,1245,1269]" captionTargetBox="[157,1376,211,1219]" captionTargetId="figure@7.[151,1436,194,1222]" captionTargetPageId="7" captionText="FIGURE 5. Pandalus ivanovi n. sp. A – C, paratype, male (CL 27.8 mm; VNIRO); D, E, paratype, male (CL 29.7 mm; same lot). A, carapace, lateral view; B, D, endopod of first pleopod; C, E, appendix masculina and appendix interna of second pleopod. Scale bars: 10 mm for A; 1 mm for B – E." httpUri="https://zenodo.org/record/184243/files/figure.png" pageId="4" pageNumber="50">Fig. 5</figureCitation>
|
||
C, E) slender, 1.8–2.0 times longer than appendix interna, bearing single or few spiniform setae terminally.
|
||
</paragraph>
|
||
<paragraph id="8BEA3686FF88FF99FF71873B81D1BBE0" blockId="4.[151,1437,152,2018]" pageId="4" pageNumber="50">
|
||
<emphasis id="B921EA94FF88FF99FF71873B85FAB8CB" bold="true" box="[197,443,1391,1417]" pageId="4" pageNumber="50">Coloration in fresh</emphasis>
|
||
. See
|
||
<figureCitation id="136E2A03FF88FF99FDB18724860FB8C8" box="[517,590,1392,1418]" captionStart="FIGURE 6" captionStartId="8.[151,255,1508,1532]" captionTargetBox="[169,1423,199,1483]" captionTargetId="figure@8.[169,1423,194,1484]" captionTargetPageId="8" captionText="FIGURE 6. Photographs showing coloration in fresh. A, Pandalus ivanovi n. sp., paratype, female (CL 31.4 mm; CBM- ZC 9222); B, Pandalus hypsinotus Brandt, 1851, female (CL 33.8 mm; CBM-ZC 9473), Canada." httpUri="https://zenodo.org/record/184244/files/figure.png" pageId="4" pageNumber="50">Fig. 6</figureCitation>
|
||
A. Background reddish brown. Rostrum pale, with red band distal to midlength, tip also reddish. Carapace with obscure transverse band along anterolateral margin, otherwise mottled with red or reddish brown. Red or reddish brown blotches on dorsal and lateral surfaces of first to sixth pleonal somites, often forming broad transverse band. Antennal flagellum with alternating red and white. Third maxilliped red, with dark reddish brown band around joint between ultimate and penultimate segments. First pereopod also red, with banding on propodus and carpus. Second pereopod each with carpus fading distally, chela white; merus and ischium red. Third to fifth pereopods generally pink; propodi each with reddish brown band subdistally, and often with second subproximal band; banding also present on meri. Uropod red.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F650DFF88FF99FF7184FB811FBB88" box="[197,1374,1711,1738]" pageId="4" pageNumber="50" type="distribution">
|
||
<paragraph id="8BEA3686FF88FF99FF7184FB811FBB88" blockId="4.[151,1437,152,2018]" box="[197,1374,1711,1738]" pageId="4" pageNumber="50">
|
||
<emphasis id="B921EA94FF88FF99FF7184FB851FBB8B" bold="true" box="[197,350,1711,1737]" pageId="4" pageNumber="50">Distribution</emphasis>
|
||
. So far Known only from off eastern Sakhalin, Sea of Okhotsk, at depths of
|
||
<quantity id="4CAD9B63FF88FF99FB5484E48118BB88" box="[1248,1369,1712,1738]" metricMagnitude="2" metricUnit="m" metricValue="1.45" metricValueMax="2.0" metricValueMin="0.9" pageId="4" pageNumber="50" unit="m" value="145.0" valueMax="200.0" valueMin="90.0">
|
||
90–
|
||
<specimenCount id="9D53FD0FFF88FF99FAB884E48118BB88" box="[1292,1369,1712,1738]" pageId="4" pageNumber="50" type="generic">200 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F650DFF88FF9BFF7184838010BC68" lastPageId="6" lastPageNumber="52" pageId="4" pageNumber="50" type="biology_ecology">
|
||
<paragraph id="8BEA3686FF88FF99FF7184838546BAA0" blockId="4.[151,1437,152,2018]" pageId="4" pageNumber="50">
|
||
<emphasis id="B921EA94FF88FF99FF7184838560BBB3" bold="true" box="[197,289,1751,1777]" pageId="4" pageNumber="50">Biology</emphasis>
|
||
. In the Sea of Okhotsk, shrimp of the genus
|
||
<taxonomicName id="4C554D05FF88FF99FCE0848C8782BBB3" box="[852,963,1752,1777]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B921EA94FF88FF99FCE0848C8782BBB3" box="[852,963,1752,1777]" italics="true" pageId="4" pageNumber="50">Pandalus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C554D05FF88FF99FBB7848C80D5BBB3" box="[1027,1172,1752,1777]" class="Malacostraca" family="Pandalidae" genus="Pandalopsis" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B921EA94FF88FF99FBB7848C80D5BBB3" box="[1027,1172,1752,1777]" italics="true" pageId="4" pageNumber="50">Pandalopsis</emphasis>
|
||
</taxonomicName>
|
||
form a major fishery resource, and it has been generally considered that the local shrimp fauna is well documented. Thus, the discovery of a new species having a high potential of commercial value from a rather well explored area is surprising. Perhaps, the habitat preference has prevented the discovery of this new species. The captain of the fishing boat capturing the new shrimp reported that the shrimp inhabited hard bottom, and that
|
||
<taxonomicName id="4C554D05FF88FF99FA99852C8555BAFB" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF88FF99FA99852C8555BAFB" italics="true" pageId="4" pageNumber="50">Pandalus hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
did not occur in the fishing ground where
|
||
<taxonomicName id="4C554D05FF88FF99FCB085F48737BAFB" box="[772,886,1952,1977]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="50" phylum="Arthropoda" rank="species" species="ivanovi" status="sp. nov.">
|
||
<emphasis id="B921EA94FF88FF99FCB085F48737BAFB" box="[772,886,1952,1977]" italics="true" pageId="4" pageNumber="50">P. ivanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257EFFF88FF99FCC985F487FDBAF8" box="[893,956,1952,1978]" pageId="4" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
|
||
was fished (S. Miyaki, personal communication).
|
||
</paragraph>
|
||
<caption id="DF2A660EFF89FF98FF2385778738BA3D" httpUri="https://zenodo.org/record/184241/files/figure.png" pageId="5" pageNumber="51" targetBox="[227,1382,201,1797]" targetPageId="5">
|
||
<paragraph id="8BEA3686FF89FF98FF2385778738BA3D" blockId="5.[151,1436,1827,1919]" pageId="5" pageNumber="51">
|
||
<emphasis id="B921EA94FF89FF98FF238577855ABA79" bold="true" box="[151,283,1827,1851]" pageId="5" pageNumber="51">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C554D05FF89FF98FE91857785A0BA78" box="[293,481,1827,1850]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="ivanovi" status="sp. nov.">
|
||
<emphasis id="B921EA94FF89FF98FE91857785A0BA78" box="[293,481,1827,1850]" italics="true" pageId="5" pageNumber="51">Pandalus ivanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257EFFF89FF98FE5E85778667BA78" box="[490,550,1827,1850]" pageId="5" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
|
||
Holotype, female (CL 28.4 mm; CBM-ZC 9221), thoracic appendages in lateral view. A, left third maxilliped; B, left first pereopod; C, left second pereopod; D, right second pereopod; E, left third pereopod; F, left fourth pereopod; G, left fifth pereopod. Scale bar: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEA3686FF89FF9BFF7185E48010BC68" blockId="5.[151,1436,1968,2034]" lastBlockId="6.[151,1436,152,378]" lastPageId="6" lastPageNumber="52" pageId="5" pageNumber="51">
|
||
The available material suggests that
|
||
<taxonomicName id="4C554D05FF89FF98FD3585E58711BA88" box="[641,848,1969,1994]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="51" phylum="Arthropoda" rank="species" species="ivanovi" status="sp. nov.">
|
||
<emphasis id="B921EA94FF89FF98FD3585E58711BA88" box="[641,848,1969,1994]" italics="true" pageId="5" pageNumber="51">Pandalus ivanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257EFFF89FF98FCEF85E487DCBA88" box="[859,925,1968,1994]" pageId="5" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
|
||
is a protandrous hermaphrodite like other congeneric species (
|
||
<bibRefCitation id="EFC44B77FF89FF98FE31858C8664BAB0" author="Komai" box="[389,549,2008,2034]" pageId="5" pageNumber="56" refString="Komai, T. (1999) A revision of the genus Pandalus (Crustacea: Decapoda: Caridea: Pandalidae). Journal of Natural History, 33, 1265 - 1372." type="journal article" year="1999">Komai, 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC44B77FF89FF98FD86858C86BCBAB0" author="Bergstrom" box="[562,765,2008,2034]" pageId="5" pageNumber="56" refString="Bergstrom, B. I. (2000) The biology of Pandalus. In: Southward, A. J., Tyler, P. A., Young, C. M. & Fuiman, L. (Eds.), Advances in Marine Biology. Vol. 38. Academic Press, London, pp. 57 - 228." type="book" year="2000">Bergstrom, 2000</bibRefCitation>
|
||
), although no specimens of functional males were available for study. Nineteen specimens (CL 27.6–30.0 mm) have subterminal appendices internae on the endopod of the first pleopod and slender, nearly unarmed appendix masculina on the second pleopod. These features suggest that the specimens are in an early stage of transitional phase (
|
||
<bibRefCitation id="EFC44B77FF8AFF9BFC0982BC8012BC40" author="Butler" box="[957,1107,232,258]" pageId="6" pageNumber="56" refString="Butler, T. H. (1980) Shrimps of the Pacific coast of Canada. Canadian Bulletin of Fisheries and Aquatic Science, 202, 1 - 280, pls 1 - 6." type="journal article" year="1980">Butler, 1980</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC44B77FF8AFF9BFBEB82BC80BDBC40" author="Komai" box="[1119,1276,232,258]" pageId="6" pageNumber="56" refString="Komai, T. (1999) A revision of the genus Pandalus (Crustacea: Decapoda: Caridea: Pandalidae). Journal of Natural History, 33, 1265 - 1372." type="journal article" year="1999">Komai, 1999</bibRefCitation>
|
||
). Females are generally larger than those of transitional males, measuring 29.0–
|
||
<quantity id="4CAD9B63FF8AFF9BFC23834487BEBC68" box="[919,1023,272,298]" metricMagnitude="-2" metricUnit="m" metricValue="3.56" pageId="6" pageNumber="52" unit="mm" value="35.6">35.6 mm</quantity>
|
||
in CL.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34F650DFF8AFF9BFF71836380D3BC38" pageId="6" pageNumber="52" type="etymology">
|
||
<paragraph id="8BEA3686FF8AFF9BFF71836380D3BC38" blockId="6.[151,1436,152,378]" pageId="6" pageNumber="52">
|
||
<emphasis id="B921EA94FF8AFF9BFF718363850BBC13" bold="true" box="[197,330,311,337]" pageId="6" pageNumber="52">Etymology</emphasis>
|
||
. This new species is dedicated to the late Boris G. Ivanov, in recognition of his contributions to the taxonomy of the decapod Crustacea. He was the supervisor of the second author.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2A660EFF8AFF9BFF23870085D7BB49" httpUri="https://zenodo.org/record/184242/files/figure.png" pageId="6" pageNumber="52" targetBox="[239,1343,428,1343]" targetPageId="6">
|
||
<paragraph id="8BEA3686FF8AFF9BFF23870085D7BB49" blockId="6.[151,1436,1364,1547]" pageId="6" pageNumber="52">
|
||
<emphasis id="B921EA94FF8AFF9BFF238700855BB82E" bold="true" box="[151,282,1364,1388]" pageId="6" pageNumber="52">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C554D05FF8AFF9BFE958701859DB82E" box="[289,476,1365,1388]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="ivanovi" status="sp. nov.">
|
||
<emphasis id="B921EA94FF8AFF9BFE958701859DB82E" box="[289,476,1365,1388]" italics="true" pageId="6" pageNumber="52">Pandalus ivanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257EFFF8AFF9BFE578700865CB829" box="[483,541,1364,1387]" pageId="6" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
|
||
Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, ultimate segment of left third maxilliped, lateral view (setae partially omitted); B, ischium of left first pereopod, lateral view (setae omitted); C, dactylus and distal part of propodus of left third pereopod, lateral view; D, distal part of propodus of left third pereopod, ventral (flexor) view; E, proximal part of propodus and carpus of left third pereopod, mesial view; F, dactylus and distal part of propodus of left fourth pereopod, lateral view; G, dactylus and distal part of propodus of left fifth pereopod, lateral view. Scale bars: 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34F650DFF8AFF94FF71846D87E7BAE1" lastPageId="9" lastPageNumber="55" pageId="6" pageNumber="52" type="discussion">
|
||
<paragraph id="8BEA3686FF8AFF9BFF71846D803BBAAD" blockId="6.[151,1437,1593,2031]" pageId="6" pageNumber="52">
|
||
<emphasis id="B921EA94FF8AFF9BFF71846D8576BB11" bold="true" box="[197,311,1593,1619]" pageId="6" pageNumber="52">Remarks</emphasis>
|
||
. The new species is referred to the
|
||
<taxonomicName id="4C554D05FF8AFF9BFD60846E8786BB11" box="[724,967,1594,1619]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8AFF9BFD60846E8786BB11" box="[724,967,1594,1619]" italics="true" pageId="6" pageNumber="52">Pandalus hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
group of
|
||
<bibRefCitation id="EFC44B77FF8AFF9BFB8D846E80A1BB16" author="Komai" box="[1081,1248,1594,1620]" pageId="6" pageNumber="56" refString="Komai, T. (1999) A revision of the genus Pandalus (Crustacea: Decapoda: Caridea: Pandalidae). Journal of Natural History, 33, 1265 - 1372." type="journal article" year="1999">Komai (1999)</bibRefCitation>
|
||
, including eight species, viz.,
|
||
<taxonomicName id="4C554D05FF8AFF9BFE8084348677BB3B" authority="Komai, 1999" authorityName="Komai" authorityYear="1999" box="[308,566,1631,1657]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="chani">
|
||
<emphasis id="B921EA94FF8AFF9BFE80843485D3BB3B" box="[308,402,1632,1657]" italics="true" pageId="6" pageNumber="52">P. chani</emphasis>
|
||
<bibRefCitation id="EFC44B77FF8AFF9BFE2E840B8677BB3B" author="Komai" box="[410,566,1631,1657]" pageId="6" pageNumber="56" refString="Komai, T. (1999) A revision of the genus Pandalus (Crustacea: Decapoda: Caridea: Pandalidae). Journal of Natural History, 33, 1265 - 1372." type="journal article" year="1999">Komai, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C554D05FF8AFF9BFDF68434869DBB3B" box="[578,732,1632,1657]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8AFF9BFDF68434869DBB3B" box="[578,732,1632,1657]" italics="true" pageId="6" pageNumber="52">P. hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C554D05FF8AFF9BFD5884348074BB3B" authority="Komai, 1999" authorityName="Komai" authorityYear="1999" box="[748,1077,1631,1657]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="formosanus">
|
||
<emphasis id="B921EA94FF8AFF9BFD58843487D3BB3B" box="[748,914,1632,1657]" italics="true" pageId="6" pageNumber="52">P. formosanus</emphasis>
|
||
<bibRefCitation id="EFC44B77FF8AFF9BFC2D840B8074BB3B" author="Komai" box="[921,1077,1631,1657]" pageId="6" pageNumber="56" refString="Komai, T. (1999) A revision of the genus Pandalus (Crustacea: Decapoda: Caridea: Pandalidae). Journal of Natural History, 33, 1265 - 1372." type="journal article" year="1999">Komai, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C554D05FF8AFF9BFBF58434813BBB3B" authority="Stimpson, 1860" authorityName="Stimpson" authorityYear="1860" box="[1089,1402,1631,1657]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B921EA94FF8AFF9BFBF584348011BB3B" box="[1089,1104,1632,1657]" italics="true" pageId="6" pageNumber="52">P</emphasis>
|
||
.
|
||
<emphasis id="B921EA94FF8AFF9BFBEB843480F8BB3B" box="[1119,1209,1632,1657]" italics="true" pageId="6" pageNumber="52">gracilis</emphasis>
|
||
Stimpson, 1860
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C554D05FF8AFF9BFA32843485F2BBDD" authority="Stimpson, 1871" authorityName="Stimpson" authorityYear="1871" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="gurneyi">
|
||
<emphasis id="B921EA94FF8AFF9BFA32843484B0BBDC" italics="true" pageId="6" pageNumber="52">P. gurneyi</emphasis>
|
||
Stimpson, 1871
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C554D05FF8AFF9BFE7484D18759BBDD" authority="Yokoya, 1933" authorityName="Yokoya" authorityYear="1933" box="[448,792,1669,1695]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="nipponensis">
|
||
<emphasis id="B921EA94FF8AFF9BFE7484D18628BBDC" box="[448,617,1669,1694]" italics="true" pageId="6" pageNumber="52">P. nipponensis</emphasis>
|
||
Yokoya, 1933
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B921EA94FF8AFF9BFC9084D187D8BBDC" box="[804,921,1669,1694]" italics="true" pageId="6" pageNumber="52">P. p re n s o r</emphasis>
|
||
Stimpson, 1869, and
|
||
<taxonomicName id="4C554D05FF8AFF9BFB2A84D181D9BBDD" authority="Kubo, 1937" authorityName="Kubo" authorityYear="1937" box="[1182,1432,1669,1695]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="teraoi">
|
||
<emphasis id="B921EA94FF8AFF9BFB2A84D18140BBDC" box="[1182,1281,1669,1694]" italics="true" pageId="6" pageNumber="52">P. teraoi</emphasis>
|
||
<collectingCountry id="F3427616FF8AFF9BFABD84D1810CBBDD" box="[1289,1357,1669,1695]" name="Cuba" pageId="6" pageNumber="52">Kubo</collectingCountry>
|
||
, 1937
|
||
</taxonomicName>
|
||
. It is very similar to
|
||
<taxonomicName id="4C554D05FF8AFF9BFE3584FE865ABB81" box="[385,539,1706,1731]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8AFF9BFE3584FE865ABB81" box="[385,539,1706,1731]" italics="true" pageId="6" pageNumber="52">P. hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
in the morphology and the general coloration in life. Shared characters are: the number of spines of the dorsal rostral series could be within a range of 18–24, of which seven to nine spines are on the carapace; the posteriormost spine of the dorsal rostral series arises distinctly posterior to the midlength of the carapace; the postrostral ridge is markedly elevated, thus the dorsal margin of the carapace is convex; the pleon is banded with dark reddish brown. Counts of spines or spinules on segments of the pereopods also agree for each other. Because of the strong similarity between the two taxa, we have examined the
|
||
<typeStatus id="54EE8824FF8AFF9BFF2385DE84BFBAE6" box="[151,254,1930,1956]" pageId="6" pageNumber="52" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C554D05FF8AFF9BFE9085DE8656BAE1" box="[292,535,1930,1955]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="52" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8AFF9BFE9085DE8656BAE1" box="[292,535,1930,1955]" italics="true" pageId="6" pageNumber="52">Pandalus hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54EE8824FF8AFF9BFD9285DE861BBAE6" box="[550,602,1930,1956]" pageId="6" pageNumber="52">type</typeStatus>
|
||
locality: Unalaska;
|
||
<bibRefCitation id="EFC44B77FF8AFF9BFCF085DE879EBAE6" author="Brandt" box="[836,991,1930,1956]" pageId="6" pageNumber="56" refString="Brandt, F. (1851) Krebse. In: Middendorff, A. Th. v. (Ed.) Reise in den aussersten Norden und Osten Sibiriens wahrend der Jahre 1843 und 1844 mit allerhochster Genehmigung auf veranstaltung der Kaiserlichen Akademie der Wissenchaften zu St. Petersburg ausgefuhrt und in Verbindung mit vielen Gelehten herausgegeben, 2 (1), 79 - 148, pls. 5, 6." type="journal article" year="1851">Brandt, 1851</bibRefCitation>
|
||
) in order to confirm the specific identity. The condition of the
|
||
<typeStatus id="54EE8824FF8AFF9BFE7785FB8668BA8B" box="[451,553,1967,1993]" pageId="6" pageNumber="52" type="syntype">syntypes</typeStatus>
|
||
is poor, but characters necessary for species recognition are still preserved (
|
||
<figureCitation id="136E2A03FF8AFF9BFF2B858184A6BAAD" box="[159,231,2005,2031]" captionStart="FIGURE 7" captionStartId="9.[151,255,973,997]" captionTargetBox="[449,1137,184,948]" captionTargetId="figure@9.[449,1138,184,949]" captionTargetPageId="9" captionText="FIGURE 7. Pandalus hypsinotus Brandt, 1851. Syntypes (ZISP 1453). A, four specimens removed from jar; B, one of the syntypes; C, largest syntype (CL about 38.0 mm)." httpUri="https://zenodo.org/record/184245/files/figure.png" pageId="6" pageNumber="52">Fig. 7</figureCitation>
|
||
). It was confirmed that the eastern Sakhalin shrimp is an undescribed species.
|
||
</paragraph>
|
||
<caption id="DF2A660EFF8BFF9AFF23868986BEB878" httpUri="https://zenodo.org/record/184243/files/figure.png" pageId="7" pageNumber="53" targetBox="[157,1376,211,1219]" targetPageId="7">
|
||
<paragraph id="8BEA3686FF8BFF9AFF23868986BEB878" blockId="7.[151,1436,1245,1338]" pageId="7" pageNumber="53">
|
||
<emphasis id="B921EA94FF8BFF9AFF238689855AB9B7" bold="true" box="[151,283,1245,1269]" pageId="7" pageNumber="53">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C554D05FF8BFF9AFE96868A859CB9B7" box="[290,477,1246,1269]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="ivanovi" status="sp. nov.">
|
||
<emphasis id="B921EA94FF8BFF9AFE96868A859CB9B7" box="[290,477,1246,1269]" italics="true" pageId="7" pageNumber="53">Pandalus ivanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257EFFF8BFF9AFE51868A865EB9B7" box="[485,543,1246,1269]" pageId="7" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
|
||
A–C, paratype, male (CL 27.8 mm; VNIRO); D, E, paratype, male (CL 29.7 mm; same lot). A, carapace, lateral view; B, D, endopod of first pleopod; C, E, appendix masculina and appendix interna of second pleopod. Scale bars: 10 mm for A; 1 mm for B–E.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEA3686FF8BFF9AFF71873F8556BA99" blockId="7.[151,1436,1387,2011]" pageId="7" pageNumber="53">
|
||
The new species differs from
|
||
<taxonomicName id="4C554D05FF8BFF9AFD98873F8689B8C6" box="[556,712,1387,1412]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8BFF9AFD98873F8689B8C6" box="[556,712,1387,1412]" italics="true" pageId="7" pageNumber="53">P. hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
in variable but apparently significant morphological characters. The rostrum is more strongly curved (cf.
|
||
<figureCitation id="136E2A03FF8BFF9AFD0B87C58751B8E9" box="[703,784,1425,1451]" captionStart="FIGURE 2" captionStartId="3.[151,255,1735,1759]" captionTargetBox="[225,1357,197,1702]" captionTargetId="figure@3.[219,1370,194,1712]" captionTargetPageId="3" captionText="FIGURE 2. Pandalus ivanovi n. sp. Holotype, female (CL 28.4 mm; CBM-ZC 9221). A, carapace and cephalic appendages, lateral view; B, rostrum, lateral view (setae omitted); C, posterior four pleonal somites and telson, lateral view (setae omitted); D, telson, dorsal view; E, left ocular appendage, antennule and antenna, dorsal view. Scale bars: 10 mm for A, C; 5 mm for B, D, E." httpUri="https://zenodo.org/record/184240/files/figure.png" pageId="7" pageNumber="53">Figs. 2</figureCitation>
|
||
A and 6B), and tends to be proportionally shorter in
|
||
<taxonomicName id="4C554D05FF8BFF9AFA3287C584ACB88D" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="ivanovi">
|
||
<emphasis id="B921EA94FF8BFF9AFA3287C584ACB88D" italics="true" pageId="7" pageNumber="53">P. ivanovi</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C554D05FF8BFF9AFEE687E285AFB88D" box="[338,494,1462,1487]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8BFF9AFEE687E285AFB88D" box="[338,494,1462,1487]" italics="true" pageId="7" pageNumber="53">P. hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
although the values partly overlap (
|
||
<figureCitation id="136E2A03FF8BFF9AFC1487E287A8B892" box="[928,1001,1462,1488]" captionStart="FIGURE 8" captionStartId="9.[151,255,1739,1763]" captionTargetBox="[240,1359,1077,1713]" captionTargetId="figure@9.[223,1367,1068,1716]" captionTargetPageId="9" captionText="FIGURE 8. Scatter plot of the relative length of the rostrum (indicated by rostral length / carapace length against carapace length) between Pandalus ivanovi n. sp. and P. hypsinotus Brandt, 1851." httpUri="https://zenodo.org/record/184246/files/figure.png" pageId="7" pageNumber="53">Fig. 8</figureCitation>
|
||
). The proportional value against the carapace length ranges 1.02–1.29 (1.12 on average, n=55) in
|
||
<taxonomicName id="4C554D05FF8BFF9AFCCB878F87B4B8B6" box="[895,1013,1499,1524]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="ivanovi">
|
||
<emphasis id="B921EA94FF8BFF9AFCCB878F87B4B8B6" box="[895,1013,1499,1524]" italics="true" pageId="7" pageNumber="53">P. ivanovi</emphasis>
|
||
</taxonomicName>
|
||
, and 1.19–1.58 (1.332 on average, n=43) in
|
||
<taxonomicName id="4C554D05FF8BFF9AFEB7845585DCBB58" box="[259,413,1537,1562]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8BFF9AFEB7845585DCBB58" box="[259,413,1537,1562]" italics="true" pageId="7" pageNumber="53">P. hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
. The postrostral ridge in transitional males and females is distinctly lower in
|
||
<taxonomicName id="4C554D05FF8BFF9AFA9D845581DDBB58" box="[1321,1436,1537,1562]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="ivanovi">
|
||
<emphasis id="B921EA94FF8BFF9AFA9D845581DDBB58" box="[1321,1436,1537,1562]" italics="true" pageId="7" pageNumber="53">P. ivanovi</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C554D05FF8BFF9AFF44847285CABB7D" box="[240,395,1574,1599]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8BFF9AFF44847285CABB7D" box="[240,395,1574,1599]" italics="true" pageId="7" pageNumber="53">P. hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
. The development of the postrostral ridge shows an ontogenetic variation in
|
||
<taxonomicName id="4C554D05FF8BFF9AFAAD847284F8BB26" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8BFF9AFAAD847284F8BB26" italics="true" pageId="7" pageNumber="53">P. hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<bibRefCitation id="EFC44B77FF8BFF9AFF44841F85CFBB27" author="Komai" box="[240,398,1611,1637]" pageId="7" pageNumber="56" refString="Komai, T. (1999) A revision of the genus Pandalus (Crustacea: Decapoda: Caridea: Pandalidae). Journal of Natural History, 33, 1265 - 1372." type="journal article" year="1999">Komai, 1999</bibRefCitation>
|
||
), and thus perhaps it will be not reliable in identifying functional males. The carpal articles of the left second pereopods are constantly fewer in
|
||
<taxonomicName id="4C554D05FF8BFF9AFC98842587B9BBC8" box="[812,1016,1649,1674]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="ivanovi">
|
||
<emphasis id="B921EA94FF8BFF9AFC98842587B9BBC8" box="[812,1016,1649,1674]" italics="true" pageId="7" pageNumber="53">Pandalus ivanovi</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C554D05FF8BFF9AFBE2842580B1BBC8" box="[1110,1264,1649,1674]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8BFF9AFBE2842580B1BBC8" box="[1110,1264,1649,1674]" italics="true" pageId="7" pageNumber="53">P. hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
(54–63 versus 72–85). The dactylus of the fifth pereopod seems to be more strongly curved in
|
||
<taxonomicName id="4C554D05FF8BFF9AFBF284C280F8BBED" box="[1094,1209,1686,1711]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="ivanovi">
|
||
<emphasis id="B921EA94FF8BFF9AFBF284C280F8BBED" box="[1094,1209,1686,1711]" italics="true" pageId="7" pageNumber="53">P. ivanovi</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C554D05FF8BFF9AFAAE84C284F8BB96" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8BFF9AFAAE84C284F8BB96" italics="true" pageId="7" pageNumber="53">P. hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
. The coloration in life is also different between the two species (cf.
|
||
<figureCitation id="136E2A03FF8BFF9AFC6E84EF805EBB97" box="[986,1055,1723,1749]" captionStart="FIGURE 7" captionStartId="9.[151,255,973,997]" captionTargetBox="[449,1137,184,948]" captionTargetId="figure@9.[449,1138,184,949]" captionTargetPageId="9" captionText="FIGURE 7. Pandalus hypsinotus Brandt, 1851. Syntypes (ZISP 1453). A, four specimens removed from jar; B, one of the syntypes; C, largest syntype (CL about 38.0 mm)." httpUri="https://zenodo.org/record/184245/files/figure.png" pageId="7" pageNumber="53">Fig. 7</figureCitation>
|
||
A, B). The general body color is more reddish in the new species than in
|
||
<taxonomicName id="4C554D05FF8BFF9AFDD884B58747BBB8" box="[620,774,1761,1786]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8BFF9AFDD884B58747BBB8" box="[620,774,1761,1786]" italics="true" pageId="7" pageNumber="53">P. hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
; bands on the pleon are wider in
|
||
<taxonomicName id="4C554D05FF8BFF9AFB3884B580BFBBB8" box="[1164,1278,1761,1786]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="ivanovi" status="sp. nov.">
|
||
<emphasis id="B921EA94FF8BFF9AFB3884B580BFBBB8" box="[1164,1278,1761,1786]" italics="true" pageId="7" pageNumber="53">P. ivanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257EFFF8BFF9AFAB284B58105BBB9" box="[1286,1348,1761,1787]" pageId="7" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
|
||
than in
|
||
<taxonomicName id="4C554D05FF8BFF9AFF2385528572BA5D" box="[151,307,1798,1823]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8BFF9AFF2385528572BA5D" box="[151,307,1798,1823]" italics="true" pageId="7" pageNumber="53">P. hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
. Furthermore,
|
||
<taxonomicName id="4C554D05FF8BFF9AFE5D855286F9BA5D" box="[489,696,1798,1823]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="ivanovi" status="sp. nov.">
|
||
<emphasis id="B921EA94FF8BFF9AFE5D855286F9BA5D" box="[489,696,1798,1823]" italics="true" pageId="7" pageNumber="53">Pandalus ivanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257EFFF8BFF9AFD7685528742BA62" box="[706,771,1798,1824]" pageId="7" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
|
||
is characterized by its relatively small size; the largest specimen of
|
||
<taxonomicName id="4C554D05FF8BFF9AFE86857F85E9BA06" box="[306,424,1835,1860]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="ivanovi">
|
||
<emphasis id="B921EA94FF8BFF9AFE86857F85E9BA06" box="[306,424,1835,1860]" italics="true" pageId="7" pageNumber="53">P. ivanovi</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<quantity id="4CAD9B63FF8BFF9AFE64857F867ABA07" box="[464,571,1835,1861]" metricMagnitude="-2" metricUnit="m" metricValue="3.56" pageId="7" pageNumber="53" unit="mm" value="35.6">35.6 mm</quantity>
|
||
in CL; female specimens, including ovigerous, are
|
||
<quantity id="4CAD9B63FF8BFF9AFB1B857F8120BA07" box="[1199,1377,1835,1861]" metricMagnitude="-2" metricUnit="m" metricValue="3.145" metricValueMax="3.56" metricValueMin="2.73" pageId="7" pageNumber="53" unit="mm" value="31.45" valueMax="35.6" valueMin="27.3">27.3–35.6 mm</quantity>
|
||
. The maximum size appears considerably different in
|
||
<taxonomicName id="4C554D05FF8BFF9AFD59850587CABA28" box="[749,907,1873,1898]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="53" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF8BFF9AFD59850587CABA28" box="[749,907,1873,1898]" italics="true" pageId="7" pageNumber="53">P. hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
according to different populations. Female specimens from
|
||
<collectingCountry id="F3427616FF8BFF9AFEED85228589BAD2" box="[345,456,1910,1936]" name="China" pageId="7" pageNumber="53">East Asia</collectingCountry>
|
||
are 36.1–45.0 mm in CL, and the specimens of less than
|
||
<quantity id="4CAD9B63FF8BFF9AFBDD852280FDBAD2" box="[1129,1212,1910,1936]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="7" pageNumber="53" unit="mm" value="36.0">36 mm</quantity>
|
||
in CL are all functional males or transitional individuals. On the other hand, the female specimens from
|
||
<collectingCountry id="F3427616FF8BFF9AFB2E85CF80B3BAF7" box="[1178,1266,1947,1973]" name="Canada" pageId="7" pageNumber="53">Canada</collectingCountry>
|
||
are 32.5–40.0 mm in CL.
|
||
</paragraph>
|
||
<caption id="DF2A660EFF84FF95FF2387B080E4BB5C" httpUri="https://zenodo.org/record/184244/files/figure.png" pageId="8" pageNumber="54" targetBox="[169,1423,199,1483]" targetPageId="8">
|
||
<paragraph id="8BEA3686FF84FF95FF2387B080E4BB5C" blockId="8.[151,1436,1508,1566]" pageId="8" pageNumber="54">
|
||
<emphasis id="B921EA94FF84FF95FF2387B08558B8BE" bold="true" box="[151,281,1508,1532]" pageId="8" pageNumber="54">FIGURE 6.</emphasis>
|
||
Photographs showing coloration in fresh. A,
|
||
<taxonomicName id="4C554D05FF84FF95FD4887B087F7B8B9" box="[764,950,1508,1531]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="54" phylum="Arthropoda" rank="species" species="ivanovi" status="sp. nov.">
|
||
<emphasis id="B921EA94FF84FF95FD4887B087F7B8B9" box="[764,950,1508,1531]" italics="true" pageId="8" pageNumber="54">Pandalus ivanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257EFFF84FF95FC0887B087B6B8B9" box="[956,1015,1508,1531]" pageId="8" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
|
||
, paratype, female (CL 31.4 mm; CBM- ZC 9222); B,
|
||
<taxonomicName id="4C554D05FF84FF95FE9F845386DDBB5C" authority="Brandt, 1851" authorityName="Brandt" authorityYear="1851" box="[299,668,1543,1566]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="54" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF84FF95FE9F84538649BB5C" box="[299,520,1543,1566]" italics="true" pageId="8" pageNumber="54">Pandalus hypsinotus</emphasis>
|
||
Brandt, 1851
|
||
</taxonomicName>
|
||
, female (CL 33.8 mm; CBM-ZC 9473), Canada.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEA3686FF84FF95FF71841B8611BAEB" blockId="8.[151,1437,1615,1961]" pageId="8" pageNumber="54">
|
||
It is interesting to note that the two separate populations of
|
||
<taxonomicName id="4C554D05FF84FF95FC3F841B803EBB2A" box="[907,1151,1615,1640]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="54" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF84FF95FC3F841B803EBB2A" box="[907,1151,1615,1640]" italics="true" pageId="8" pageNumber="54">Pandalus hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
show differences in the relative length of the rostrum and overall body size. The rostrum seems to be proportionally longer in the Asian population than in the Canadian population (1.29–1.57 times as long as the carapace versus 1.19–1.42), although the values partially overlap. The body size is larger in the Asian population than in the Canadian population (transitional males CL
|
||
<quantity id="4CAD9B63FF84FF95FD8284BB86A6BA4B" box="[566,743,1775,1801]" metricMagnitude="-2" metricUnit="m" metricValue="3.945" metricValueMax="4.2299999999999995" metricValueMin="3.66" pageId="8" pageNumber="54" unit="mm" value="39.45" valueMax="42.3" valueMin="36.6">36.6–42.3 mm</quantity>
|
||
, females CL
|
||
<quantity id="4CAD9B63FF84FF95FC3884BB807BBA4B" box="[908,1082,1775,1801]" metricMagnitude="-2" metricUnit="m" metricValue="4.18" metricValueMax="4.75" metricValueMin="3.61" pageId="8" pageNumber="54" unit="mm" value="41.8" valueMax="47.5" valueMin="36.1">36.1–47.5 mm</quantity>
|
||
versus transitional males CL 31.0–33.0 mm, females CL 33.6–40.0 mm).
|
||
<bibRefCitation id="EFC44B77FF84FF95FD0F8543872EBA73" author="Haynes" box="[699,879,1815,1841]" pageId="8" pageNumber="56" refString="Haynes, E. (1976) Descriptions of zoeae of coonstripe shrimp, Pandalus hypsinotus, reared in the laboratory. Fishery Bulletin, 74 (2), 323 - 342." type="journal article" year="1976">Haynes (1976)</bibRefCitation>
|
||
, who described the larval stages of
|
||
<taxonomicName id="4C554D05FF84FF95FA9985438555BA1A" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="54" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF84FF95FA9985438555BA1A" italics="true" pageId="8" pageNumber="54">Pandalus hypsinotus</emphasis>
|
||
</taxonomicName>
|
||
from Alaska, noted several morphological differences between his larvae and those described by
|
||
<bibRefCitation id="EFC44B77FF84FF95FF238533857CBAC3" author="Kurata" box="[151,317,1895,1921]" pageId="8" pageNumber="56" refString="Kurata, H. (1964) Larvae of decapod Crustacea of Hokkaido. 3. Pandalidae. Bulletin of the Hokkaido Regional Fisheries Research Laboratory, 28, 23 - 34." type="journal article" year="1964">Kurata (1964)</bibRefCitation>
|
||
from Hokkaido,
|
||
<collectingCountry id="F3427616FF84FF95FDBF85338610BAC3" box="[523,593,1895,1921]" name="Japan" pageId="8" pageNumber="54">Japan</collectingCountry>
|
||
. Detailed comparison will be necessary if the geographically separated populations belong to a same species.
|
||
</paragraph>
|
||
<caption id="DF2A660EFF85FF94FF2381998693B945" httpUri="https://zenodo.org/record/184245/files/figure.png" pageId="9" pageNumber="55" targetBox="[449,1137,184,948]" targetPageId="9">
|
||
<paragraph id="8BEA3686FF85FF94FF2381998693B945" blockId="9.[151,1436,973,1031]" pageId="9" pageNumber="55">
|
||
<emphasis id="B921EA94FF85FF94FF238199855BBEA7" bold="true" box="[151,282,973,997]" pageId="9" pageNumber="55">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C554D05FF85FF94FE96819A86D6BEA7" authority="Brandt, 1851" authorityName="Brandt" authorityYear="1851" box="[290,663,974,997]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF85FF94FE96819A8641BEA7" box="[290,512,974,997]" italics="true" pageId="9" pageNumber="55">Pandalus hypsinotus</emphasis>
|
||
Brandt, 1851
|
||
</taxonomicName>
|
||
. Syntypes (ZISP 1453). A, four specimens removed from jar; B, one of the syntypes; C, largest syntype (CL about 38.0 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2A660EFF85FF94FF23849F87DABA47" httpUri="https://zenodo.org/record/184246/files/figure.png" pageId="9" pageNumber="55" targetBox="[240,1359,1077,1713]" targetPageId="9">
|
||
<paragraph id="8BEA3686FF85FF94FF23849F87DABA47" blockId="9.[151,1435,1739,1798]" pageId="9" pageNumber="55">
|
||
<emphasis id="B921EA94FF85FF94FF23849F8558BBA1" bold="true" box="[151,281,1739,1763]" pageId="9" pageNumber="55">FIGURE 8.</emphasis>
|
||
Scatter plot of the relative length of the rostrum (indicated by rostral length/carapace length against carapace length) between
|
||
<taxonomicName id="4C554D05FF85FF94FEFC84BB8642BA44" box="[328,515,1775,1798]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="ivanovi" status="sp. nov.">
|
||
<emphasis id="B921EA94FF85FF94FEFC84BB8642BA44" box="[328,515,1775,1798]" italics="true" pageId="9" pageNumber="55">Pandalus ivanovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21257EFFF85FF94FDBD84BA8603BA47" box="[521,578,1774,1797]" pageId="9" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C554D05FF85FF94FDC284BB87D6BA47" authority="Brandt, 1851" authorityName="Brandt" authorityYear="1851" box="[630,919,1774,1798]" class="Malacostraca" family="Pandalidae" genus="Pandalus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="55" phylum="Arthropoda" rank="species" species="hypsinotus">
|
||
<emphasis id="B921EA94FF85FF94FDC284BB8743BA44" box="[630,770,1775,1798]" italics="true" pageId="9" pageNumber="55">P. hypsinotus</emphasis>
|
||
Brandt, 1851
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEA3686FF85FF94FF71856D87E7BAE1" blockId="9.[151,1436,1849,1955]" pageId="9" pageNumber="55">It remains unclear whether the new species is endemic to the Sea of Okhotsk. Nevertheless, in spite of our collecting efforts, no specimens of the new species have been encountered in the collections made during recent fishery or scientific surveys or those in museum collections.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |