155 lines
16 KiB
XML
155 lines
16 KiB
XML
<document id="DE1A7E04486628762E0949B6CF5E4965" ID-CLB-Dataset="20903" ID-DOI="10.11646/zootaxa.4980.1.9" ID-GBIF-Dataset="c6aedffd-0196-45d9-acc3-071c4e886e66" ID-ISSN="1175-5326" ID-Zenodo-Dep="4883181" ID-ZooBank="E4D00067-F4F3-4BC5-9CDA-6532773B56B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622464354611" checkinUser="plazi" docAuthor="Makarkin, Vladimir N., Archibald, S. Bruce & Mathewes, Rolf W." docDate="2021" docId="03F07D0C043AFFF6F0957F64FDE8049B" docLanguage="en" docName="zootaxa.4980.1.9.pdf" docOrigin="Zootaxa 4980 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Osmylidia glastrai Makarkin & Archibald & Mathewes 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="145" masterDocId="FFC905740438FFF5F0027825FF9D0753" masterDocTitle="New Protosmylinae (Neuroptera: Osmylidae) from the early Eocene of western North America, with taxonomic remarks" masterLastPageNumber="156" masterPageNumber="142" pageNumber="144" updateTime="1698995111136" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="313DE68B66F922D23F7BA08A336526DE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="3E6FB64F1A71119C3C0741CF61973692">
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<mods:title id="B2DD51F38E2A964824B8CFA447C3C498">New Protosmylinae (Neuroptera: Osmylidae) from the early Eocene of western North America, with taxonomic remarks</mods:title>
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<mods:name id="B160C38D1332F50650AFE8A765E801F2" type="personal">
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<mods:namePart id="5CE9972BBFE41962983F7911F29DEE65">Makarkin, Vladimir N.</mods:namePart>
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<mods:nameIdentifier id="96C8BA52BF7EC59A854F812CAC71F882" type="ORCID">0000-0002-1304-0461</mods:nameIdentifier>
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<mods:affiliation id="8FC5B2A52E24780D1E0386644E1299F8">Federal Scientific Centre of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivostok 690022, Russia.</mods:affiliation>
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<mods:nameIdentifier id="9A666C83660142F8D68D1CCA02CF52A6" type="email">vnmakarkin@mail.ru</mods:nameIdentifier>
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<mods:name id="EF57DC3D6725C852F5F78ADBCE05BC3B" type="personal">
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<mods:roleTerm id="2DD339E02E5A981C65661E9708171767">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9D874CC3B9A934CFC8131813DD23AA47">Archibald, S. Bruce</mods:namePart>
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<mods:affiliation id="FFA1D728F217BFA602747EE47CFF9B53">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, BC, V 5 A 1 S 6, Canada. & Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America; and Royal British Columbia Museum, 675 Belleville Street, Victoria, BC, V 8 W 9 W 2, Canada.</mods:affiliation>
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</mods:name>
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<mods:name id="0337829E516987DD623F94823E1955E2" type="personal">
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<mods:roleTerm id="5B69349836B6388BB76072765136B546">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AC259DD96B891AF9AF9058BB8C67AF1A">Mathewes, Rolf W.</mods:namePart>
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<mods:affiliation id="365A4F9AE5244E3BB3DBC24ECC508065">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, BC, V 5 A 1 S 6, Canada.</mods:affiliation>
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<mods:typeOfResource id="A51BA12F6635D78534738E1A7F4994D3">text</mods:typeOfResource>
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<mods:title id="6CEBEA3E47C94D22005EC96B812909B8">Zootaxa</mods:title>
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<mods:part id="86A6DD71E0297F82FA12242DD976A87A">
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<mods:date id="F587CF6FAAD986ECD15A575CE05D3E44">2021</mods:date>
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<mods:number id="32A8BC5ADF8B57E08BD7894494BF5730">2021-05-31</mods:number>
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<mods:number id="F27D0660D39BEE37CF1235914ED408BB">4980</mods:number>
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<mods:identifier id="62BF6745954CCF36CAD34CDE6FF22217" type="DOI">10.11646/zootaxa.4980.1.9</mods:identifier>
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<treatment id="03F07D0C043AFFF6F0957F64FDE8049B" ID-DOI="http://doi.org/10.5281/zenodo.4883191" ID-GBIF-Taxon="192720198" ID-Zenodo-Dep="4883191" LSID="urn:lsid:plazi:treatment:03F07D0C043AFFF6F0957F64FDE8049B" httpUri="http://treatment.plazi.org/id/03F07D0C043AFFF6F0957F64FDE8049B" lastPageId="3" lastPageNumber="145" pageId="2" pageNumber="144">
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<subSubSection id="C3439F91043AFFF7F0957F64FE440008" box="[151,473,1857,1884]" pageId="2" pageNumber="144" type="nomenclature">
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<paragraph id="8BE6CC1A043AFFF7F0957F64FE440008" blockId="2.[151,473,1857,1919]" box="[151,473,1857,1884]" pageId="2" pageNumber="144">
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<heading id="D0AE7B76043AFFF7F0957F64FE440008" bold="true" box="[151,473,1857,1884]" fontSize="11" level="1" pageId="2" pageNumber="144" reason="1">
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<emphasis id="B92D1008043AFFF7F0957F64FE440008" bold="true" box="[151,473,1857,1884]" pageId="2" pageNumber="144">
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<taxonomicName id="4C59B799043AFFF7F0957F64FEE9000F" authority="Makarkin & Archibald & Mathewes, 2021" authorityName="Makarkin & Archibald & Mathewes" authorityYear="2021" box="[151,372,1857,1884]" class="Insecta" family="Osmylidae" genus="Osmylidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="144" phylum="Arthropoda" rank="species" species="glastrai" status="sp. nov.">
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<emphasis id="B92D1008043AFFF7F0957F64FEE9000F" bold="true" box="[151,372,1857,1884]" italics="true" pageId="2" pageNumber="144">Osmylidia glastrai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21EAD73043AFFF7F1797F64FE440008" box="[379,473,1857,1883]" pageId="2" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3439F91043AFFF7F0957F40FF44002C" box="[151,217,1893,1919]" pageId="2" pageNumber="144" type="description">
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<paragraph id="8BE6CC1A043AFFF7F0957F40FF44002C" blockId="2.[151,473,1857,1919]" box="[151,217,1893,1919]" pageId="2" pageNumber="144">
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<figureCitation id="1362D09F043AFFF7F0957F40FF44002C" box="[151,217,1893,1919]" captionStart="FIGURE 1" captionStartId="3.[151,250,1939,1963]" captionTargetBox="[151,1436,990,1902]" captionTargetId="figure-368@3.[151,1436,990,1913]" captionTargetPageId="3" captionText="FIGURE 1. Osmylidia glastrai sp. nov., holotype forewing SR 02-22-15. A, photograph; B, drawing. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4883183" httpUri="https://zenodo.org/record/4883183/files/figure.png" pageId="2" pageNumber="144">Fig. 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3439F91043AFFF6F0957F88FC3C0786" lastPageId="3" lastPageNumber="145" pageId="2" pageNumber="144" type="materials_examined">
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<paragraph id="8BE6CC1A043AFFF7F0957F88FDAA00B8" blockId="2.[151,1437,1965,2027]" pageId="2" pageNumber="144">
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<emphasis id="B92D1008043AFFF7F0957F88FED90094" bold="true" box="[151,324,1965,1991]" pageId="2" pageNumber="144">Type material.</emphasis>
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<typeStatus id="54E272B8043AFFF7F14B7F88FE2C0094" box="[329,433,1965,1991]" pageId="2" pageNumber="144" type="holotype">Holotype</typeStatus>
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SR 02-22-15 (part) collected by Joshua Glastra,
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<date id="FFE7EADA043AFFF7F3C97F8BFB1B0094" box="[971,1158,1965,1992]" pageId="2" pageNumber="144" value="2002-08-24">August 24, 2002</date>
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; deposited in SR. A wellpreserved almost complete forewing.
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</paragraph>
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<paragraph id="8BE6CC1A043BFFF6F0C578B3FC3C0786" blockId="3.[151,1437,150,969]" pageId="3" pageNumber="145">
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<materialsCitation id="3B31C647043BFFF6F0C578B3FC000786" ID-GBIF-Occurrence="3459552302" collectorName="Exposure B" country="United States of America" location="Klondike Mountain" municipality="Mountain Formation" pageId="3" pageNumber="145" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
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<emphasis id="B92D1008043BFFF6F0C578B3FE6107E3" bold="true" box="[199,508,150,176]" pageId="3" pageNumber="145">
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<typeStatus id="54E272B8043BFFF6F0C578B3FE9F07E3" box="[199,258,150,176]" pageId="3" pageNumber="145">Type</typeStatus>
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locality and horizon.
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</emphasis>
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<collectorName id="26ACA9CC043BFFF6F20178B2FD1907E3" box="[515,644,150,177]" pageId="3" pageNumber="145">Exposure B</collectorName>
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4131 of the Tom Thumb Member of the
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<location id="8E869AC1043BFFF6F44478B3FABC07E3" LSID="urn:lsid:plazi:treatment:03F07D0C043AFFF6F0957F64FDE8049B:8E869AC1043BFFF6F44478B3FABC07E3" box="[1094,1313,150,176]" country="United States of America" municipality="Mountain Formation" name="Klondike Mountain" pageId="3" pageNumber="145" stateProvince="Washington">
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Klondike
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<collectingMunicipality id="6B825660043BFFF6F4B678B3FABC07E3" box="[1204,1313,150,176]" pageId="3" pageNumber="145">Mountain</collectingMunicipality>
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</location>
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Formation in the town of Republic,
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<collectingRegion id="499D02F8043BFFF6F1AF789FFDA80787" box="[429,565,186,212]" country="United States of America" name="Washington" pageId="3" pageNumber="145">Washington</collectingRegion>
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,
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<collectingCountry id="F34E8C8A043BFFF6F242789FFD0D0787" box="[576,656,186,212]" name="United States of America" pageId="3" pageNumber="145">U.S.A.</collectingCountry>
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; Eocene (late Ypresian)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3439F91043BFFF6F0C578FBFC6807AB" box="[199,1013,222,249]" pageId="3" pageNumber="145" type="etymology">
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<paragraph id="8BE6CC1A043BFFF6F0C578FBFC6807AB" blockId="3.[151,1437,150,969]" box="[199,1013,222,249]" pageId="3" pageNumber="145">
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<emphasis id="B92D1008043BFFF6F0C578FBFED507AB" bold="true" box="[199,328,222,248]" pageId="3" pageNumber="145">Etymology</emphasis>
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. From the surname of Joshua Glastra, collector of this fossil.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3439F91043BFFF6F0C57927FC9D0612" pageId="3" pageNumber="145" type="diagnosis">
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<paragraph id="8BE6CC1A043BFFF6F0C57927FC9D0612" blockId="3.[151,1437,150,969]" pageId="3" pageNumber="145">
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<emphasis id="B92D1008043BFFF6F0C57927FEA7064F" bold="true" box="[199,314,258,284]" pageId="3" pageNumber="145">Diagnosis</emphasis>
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. Forewings may be distinguished from those of other species of the genus by faint colour patterning (
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<taxonomicName id="4C59B799043BFFF6F0A37902FF660613" box="[161,251,295,321]" class="Insecta" family="Osmylidae" genus="Osmylidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="145" phylum="Arthropoda" rank="species" species="picta">
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<emphasis id="B92D1008043BFFF6F0A37902FF660613" box="[161,251,295,321]" italics="true" pageId="3" pageNumber="145">O. picta</emphasis>
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</taxonomicName>
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: strongly spotted;
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<taxonomicName id="4C59B799043BFFF6F1CE7902FDD70613" baseAuthorityName="Scudder" baseAuthorityYear="1890" box="[460,586,295,321]" class="Insecta" family="Osmylidae" genus="Osmylidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="145" phylum="Arthropoda" rank="species" species="requieta">
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<emphasis id="B92D1008043BFFF6F1CE7902FDD70613" box="[460,586,295,321]" italics="true" pageId="3" pageNumber="145">O. requieta</emphasis>
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</taxonomicName>
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: without spots).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3439F91043BFFF6F0C5796FFF5C040E" pageId="3" pageNumber="145" type="description">
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<paragraph id="8BE6CC1A043BFFF6F0C5796FFF5C040E" blockId="3.[151,1437,150,969]" pageId="3" pageNumber="145">
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<emphasis id="B92D1008043BFFF6F0C5796FFECC0637" bold="true" box="[199,337,330,356]" pageId="3" pageNumber="145">Description</emphasis>
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. Forewing
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<emphasis id="B92D1008043BFFF6F1D1796EFE700637" box="[467,493,331,356]" italics="true" pageId="3" pageNumber="145">ca</emphasis>
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.
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<quantity id="4CA161FF043BFFF6F1FE796FFDD10636" box="[508,588,330,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="145" unit="mm" value="17.0">17 mm</quantity>
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long,
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<emphasis id="B92D1008043BFFF6F291796EFD300637" box="[659,685,331,356]" italics="true" pageId="3" pageNumber="145">ca</emphasis>
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.
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<quantity id="4CA161FF043BFFF6F2BE796FFC8E0636" box="[700,787,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="3" pageNumber="145" unit="mm" value="5.7">5.7 mm</quantity>
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wide. Costal space broad, with subcostal veinlets simple, closely spaced. Sc fused with RA far from wing apex. Sc+RA veinlets simple, closely spaced. Subcostal space narrow; crossveins not detected. RA space relatively narrow, with five preserved crossveins, rather regularly spaced; one distal crossvein probably not preserved. RP originates rather close to wing base, with 14 pectinate branches, one of these (RP10) deeply forked. M forked distad origin of RP1, proximad origin of RP2. All crossveins (beside one proximal) in radial and radio-medial spaces arranged in three gradate series: proximal series with five crossveins; intermediate series with eight crossveins; outer series with seven preserved crossveins (partly fragmentarily preserved). Basal crossvein in radio-medial space connects R, M. Both MA, MP pectinately branched distally, with three branches. Four crossveins between MA, MP; five preserved crossveins between MP, CuA. CuA pectinately branched, with three branches. CuP pectinately branched, with five preserved branches (probably six or seven in life). One preserved crossvein detected between CuA, CuP. One distal crossvein between CuP, A1. A1 relatively long, with five pectinate, simple branches. A2 relatively long, with four preserved pectinate, simple branches. Wing colouration: irregular spots throughout wing (surrounding some crossveins, in subcostal space and along margin), most prominently three between MP, CuA, one along margin near termination of CuP, A1, and two on pterostigma.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3439F91043BFFF6F0C57B43FDE8049B" pageId="3" pageNumber="145" type="discussion">
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<paragraph id="8BE6CC1A043BFFF6F0C57B43FDE8049B" blockId="3.[151,1437,150,969]" pageId="3" pageNumber="145">
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<emphasis id="B92D1008043BFFF6F0C57B43FEA604D3" bold="true" box="[199,315,870,896]" pageId="3" pageNumber="145">Remarks.</emphasis>
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The species has a five-branched CuP and, therefore, is assigned to
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<taxonomicName id="4C59B799043BFFF6F4337B43FB3904D3" authorityName="Cockerell" authorityYear="1908" box="[1073,1188,870,896]" class="Insecta" family="Osmylidae" genus="Osmylidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis id="B92D1008043BFFF6F4337B43FB3904D3" box="[1073,1188,870,896]" italics="true" pageId="3" pageNumber="145">Osmylidia</emphasis>
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</taxonomicName>
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(see genus diagnosis) but only tentatively, as this character is unknown in one fossil genus, and
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<taxonomicName id="4C59B799043BFFF6F3E17BAFFBCB04F7" authorityName="Cockerell" authorityYear="1908" box="[995,1110,906,932]" class="Insecta" family="Osmylidae" genus="Osmylidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="145" phylum="Arthropoda" rank="genus">
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<emphasis id="B92D1008043BFFF6F3E17BAFFBCB04F7" box="[995,1110,906,932]" italics="true" pageId="3" pageNumber="145">Osmylidia</emphasis>
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</taxonomicName>
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is barely distinguished from extant genera solely by forewing venation.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |