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<document id="C1BA8FF0B98441C819FA79576BC7F03D" ID-DOI="10.11646/zootaxa.5383.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10350971" ID-ZooBank="1D2AD0FF-CE98-462C-A238-CC3DBF2906C9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1702283818271" checkinUser="plazi" docAuthor="Lin, Chen, Zhang, Chunmin &amp; Yang, Ding" docDate="2023" docId="03F03213FFA919609AD6FE07636D4444" docLanguage="en" docName="zootaxa.5383.2.3.pdf" docOrigin="Zootaxa 5383 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5383.2.3/52446" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systenus flavilateralis Lin &amp; Zhang &amp; Yang 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="160" masterDocId="FFC94A6BFFAE19679A41FF95672F4308" masterDocTitle="The genus Systenus Meigen, 1803 (Diptera: Dolichopodidae) from Yunnan, China, with descriptions of six new species" masterLastPageNumber="166" masterPageNumber="153" pageNumber="160" updateTime="1702284527099" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="058F74CDDEBC8A4F368782F3B83A9EE2">The genus Systenus Meigen, 1803 (Diptera: Dolichopodidae) from Yunnan, China, with descriptions of six new species</mods:title>
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<mods:namePart id="777FCD4EA2027C5304CF29AD51A62655">Lin, Chen</mods:namePart>
<mods:affiliation id="B1CA7BC5CB6F040B171402131BE19815">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100193, China &amp; Institute of Life Science and Technology, Inner Mongolia Normal University, Huhhot 010022, China</mods:affiliation>
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<mods:namePart id="E97F0B52AAE80BEF6911B8BF2295C3DD">Zhang, Chunmin</mods:namePart>
<mods:affiliation id="58191BA86AB588B42E8AEEAF829ADD4F">Tongliao Agricultural Technology Extension Center, Tongliao 028000, China</mods:affiliation>
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<mods:namePart id="B3D596C8507B071C40348ABCEE0AAA20">Yang, Ding</mods:namePart>
<mods:affiliation id="3551FC88848430550F16481D8B0CBA28">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing 100193, China</mods:affiliation>
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<mods:title id="30B8B6085B3CBD8AAFE47A29E2F6E337">Zootaxa</mods:title>
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<mods:date id="0C2936028B4F9059DD75AD571EE377CA">2023</mods:date>
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<treatment id="03F03213FFA919609AD6FE07636D4444" ID-DOI="http://doi.org/10.5281/zenodo.10350937" ID-Zenodo-Dep="10350937" LSID="urn:lsid:plazi:treatment:03F03213FFA919609AD6FE07636D4444" httpUri="http://treatment.plazi.org/id/03F03213FFA919609AD6FE07636D4444" lastPageNumber="160" pageId="7" pageNumber="160">
<subSubSection id="C343D08EFFA919609AD6FE07651A42A4" box="[151,565,402,429]" pageId="7" pageNumber="160" type="nomenclature">
<paragraph id="8BE68305FFA919609AD6FE07651A42A4" blockId="7.[151,565,402,464]" box="[151,565,402,429]" pageId="7" pageNumber="160">
<heading id="D0AE3469FFA919609AD6FE07651A42A4" bold="true" box="[151,565,402,429]" fontSize="11" level="1" pageId="7" pageNumber="160" reason="1">
<emphasis id="B92D5F17FFA919609AD6FE07651A42A4" bold="true" box="[151,565,402,429]" pageId="7" pageNumber="160">
<emphasis id="B92D5F17FFA919609AD6FE0766FF42A5" bold="true" box="[151,464,402,429]" italics="true" pageId="7" pageNumber="160">
4.
<taxonomicName id="4C59F886FFA919609A86FE0766FF42A5" authority="Lin &amp; Zhang &amp; Yang, 2023" authorityName="Lin &amp; Zhang &amp; Yang" authorityYear="2023" box="[199,464,402,429]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="160" phylum="Arthropoda" rank="species" species="flavilateralis" status="sp. nov.">Systenus flavilateralis</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A21EE26CFFA919609B96FE07651A42A4" box="[471,565,402,428]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA919609AD6FE23666B42D8" box="[151,324,438,464]" pageId="7" pageNumber="160" type="description">
<paragraph id="8BE68305FFA919609AD6FE23666B42D8" blockId="7.[151,565,402,464]" box="[151,324,438,464]" pageId="7" pageNumber="160">
(
<figureCitation id="13629F80FFA919609ADEFE2367C742D8" box="[159,232,438,464]" captionStart="FIGURES 16. 1" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[203,1384,189,1907]" captionTargetId="figure-21@2.[193,1394,179,1917]" captionTargetPageId="2" captionText="FIGURES 16. 1. Systenus apicidigitatus sp. nov.; 2. Systenus basiflavus sp. nov.; 3. Systenus digitatus sp. nov.; 4. Systenus flavilateralis sp. nov.; 5. Systenus huanglianus sp. nov.; 6. Systenus latimaculatus sp. nov. All male. Scale bar =1.0mm." figureDoi="http://doi.org/10.5281/zenodo.10350973" httpUri="https://zenodo.org/record/10350973/files/figure.png" pageId="7" pageNumber="160">Figs 4</figureCitation>
, 13, 14)
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA919609AD6FE6B67D941A0" pageId="7" pageNumber="160" type="diagnosis">
<paragraph id="8BE68305FFA919609AD6FE6B67D941A0" blockId="7.[151,1437,510,1869]" pageId="7" pageNumber="160">
<emphasis id="B92D5F17FFA919609AD6FE6B663D4110" bold="true" box="[151,274,510,536]" pageId="7" pageNumber="160">Diagnosis.</emphasis>
Thoracic pleura mostly dark yellow. Abdomen mostly dark yellow. Antennal scape and pedicel and base of postpedicel yellow to dark yellow. Postpedicel rather short, subconical, 2.0 times longer than wide; arista subapical. All coxae yellow. Hind femur dark brown at apical 1/3; tibia brown to dark brown except extreme base and tip; tarsus dark brown. Surstylus short and thick with two small outer apical processes. Male cercus bifurcated apically.
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA919609A86FD2764FD4558" pageId="7" pageNumber="160" type="description">
<paragraph id="8BE68305FFA919609A86FD2764C141C5" blockId="7.[151,1437,510,1869]" box="[199,1006,690,717]" pageId="7" pageNumber="160">
<emphasis id="B92D5F17FFA919609A86FD27667641C4" bold="true" box="[199,345,690,716]" pageId="7" pageNumber="160">Description.</emphasis>
Male (
<figureCitation id="13629F80FFA919609BE9FD2766C141C4" box="[424,494,690,716]" captionStart="FIGURES 16. 1" captionStartId="2.[152,270,1943,1967]" captionTargetBox="[203,1384,189,1907]" captionTargetId="figure-21@2.[193,1394,179,1917]" captionTargetPageId="2" captionText="FIGURES 16. 1. Systenus apicidigitatus sp. nov.; 2. Systenus basiflavus sp. nov.; 3. Systenus digitatus sp. nov.; 4. Systenus flavilateralis sp. nov.; 5. Systenus huanglianus sp. nov.; 6. Systenus latimaculatus sp. nov. All male. Scale bar =1.0mm." figureDoi="http://doi.org/10.5281/zenodo.10350973" httpUri="https://zenodo.org/record/10350973/files/figure.png" pageId="7" pageNumber="160">Fig. 4</figureCitation>
). Body length
<quantity id="4CA12EE0FFA9196098D2FD2765C441C5" box="[659,747,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="160" unit="mm" value="2.6">2.6 mm</quantity>
, wing length
<quantity id="4CA12EE0FFA9196099C3FD2764C641C5" box="[898,1001,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="7" pageNumber="160" unit="mm" value="2.25">2.25 mm</quantity>
.
</paragraph>
<paragraph id="8BE68305FFA919609A86FD43630F40AC" blockId="7.[151,1437,510,1869]" pageId="7" pageNumber="160">Head metallic green with pale gray pollinosity. Frons metallic green with white gray pollinosity; face dark metallic green with dense silvery white pollinosity, nearly as wide as postpedicel. Hairs and bristles on head black, but postocular bristles yellowish. 2 oc, 2 vt, 2 pvt. Antenna [aristal apical segment broken] dark brown, but scape and pedicel yellow and postpedicel dark yellow at base; pedicel with black apical bristles; postpedicel rather short, subconical, 2.0 times longer than wide, with dense pale pubescence (Fig. 13). Proboscis short, dark brown with blackish hairs; palpus small, blackish, with black hairs and 1 black apical bristle.</paragraph>
<paragraph id="8BE68305FFA919609A86FC3B6450473C" blockId="7.[151,1437,510,1869]" pageId="7" pageNumber="160">Thorax metallic green with pale gray pollinosity, but humerus brownish yellow; pleura mostly dark yellow except mesopleuron mostly blackish, pteropleuron with black spot, sternopleuron blackish at posterior margin and hypopleuron blackish anteriorly. Hairs and bristles on thorax black; 6 dc; 8 to 9 irregular pairs of acr; 2 strong npl, 1 strong sa, 2 strong pa; sc long and strong, lsc short and hair-like.</paragraph>
<paragraph id="8BE68305FFA919609A86FBAA642C4604" blockId="7.[151,1437,510,1869]" pageId="7" pageNumber="160">Legs [hind tarsomeres 45 broken] mainly yellow; all coxae yellow; hind femur dark brown at apical 1/3; hind tibia brown to dark brown except extreme base and tip; mid tarsus brownish yellow, hind tarsus dark brown. Hairs and bristles on legs black. Hind coxa with 1 strong outer bristle basally. Mid tibia with 2 ad, 2 pd, 3 pv and 4 apical bristles; hind tibia with 2 ad and 4 apical bristles. Mid and hind tarsomeres 14 each with 4 short apical bristles. Relative length of tibiae and five tarsomeres of legs LI 3.0: 1.8: 1.0: 0.9: 0.6: 0.5; LII 4.4: 2.3: 1.3: 1.0: 0.7: 0.5; LIII 5.4: 1.4: 1.7: 1.0: distal tarsomeres missing.</paragraph>
<paragraph id="8BE68305FFA919609A86FA836331465C" blockId="7.[151,1437,510,1869]" pageId="7" pageNumber="160">
Wing hyaline, weakly tinged brownish; veins dark brown; M
<subScript id="17DD8140FFA919609933FABC6455463F" attach="left" box="[882,890,1321,1335]" fontSize="6" pageId="7" pageNumber="160">1</subScript>
bent medially, R
<subScript id="17DD8140FFA919609E78FABD637C463E" attach="left" box="[1081,1107,1320,1334]" fontSize="6" pageId="7" pageNumber="160">4+5</subScript>
and M
<subScript id="17DD8140FFA919609EE3FABC6385463F" attach="left" box="[1186,1194,1321,1335]" fontSize="6" pageId="7" pageNumber="160">1</subScript>
distinctly convergent apically; CuAx ratio 0.6. Squama yellow with blackish hairs. Halter pale yellow.
</paragraph>
<paragraph id="8BE68305FFA919609A86FACB65DA46C8" blockId="7.[151,1437,510,1869]" pageId="7" pageNumber="160">Abdomen mostly dark yellow with pale gray pollinosity. Terga 13 dark yellow except tergum 3 pale metallic green at middle; terga 46 dark metallic green; venter dark yellow. Hairs and bristles on abdomen black. First tergum with very long black hairs at posterior margin.</paragraph>
<paragraph id="8BE68305FFA919609A86FA5F625D4524" blockId="7.[151,1437,510,1869]" pageId="7" pageNumber="160">Male genitalia (Fig. 14): epandrium distinctly longer than wide; epandrial lobe long and thick, apically distinctly narrowed toward tip; surstylus short and thick with two small outer apical processes. Cercus long and thick, apically bifurcated. Hypandrium long, somewhat thick, apically narrow and slightly curved. Phallus long and very thin.</paragraph>
<paragraph id="8BE68305FFA919609A86F9A364FD4558" blockId="7.[151,1437,510,1869]" box="[199,978,1590,1617]" pageId="7" pageNumber="160">
<emphasis id="B92D5F17FFA919609A86F9A3660A4558" bold="true" box="[199,293,1590,1616]" pageId="7" pageNumber="160">Female.</emphasis>
Body length
<quantity id="4CA12EE0FFA919609BFCF9A3650C4559" box="[445,547,1590,1617]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" pageId="7" pageNumber="160" unit="mm" value="2.45">2.45 mm</quantity>
, wing length
<quantity id="4CA12EE0FFA9196098FBF9A3643C4559" box="[698,787,1590,1617]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="160" unit="mm" value="2.4">2.4 mm</quantity>
. Similar to male.
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA919609A86F9CF668645B4" pageId="7" pageNumber="160" type="materials_examined">
<paragraph id="8BE68305FFA919609A86F9CF668645B4" blockId="7.[151,1437,510,1869]" pageId="7" pageNumber="160">
<emphasis id="B92D5F17FFA919609A86F9CF665E457C" bold="true" box="[199,369,1626,1652]" pageId="7" pageNumber="160">Type material.</emphasis>
<materialsCitation id="3B318958FFA919609B32F9CF63424590" collectingDate="2019-04-19" collectingDateMax="2019-05-19" collectingDateMin="2019-04-19" collectionCode="CAU" collectorName="Malaise &amp; Liang Wang &amp; Xin Li" country="China" county="Lvchun" elevation="1779" latitude="23.057" location="Shuikuxiafang" longLatPrecision="1" longitude="102.777145" municipality="Yakou" pageId="7" pageNumber="160" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
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</emphasis>
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,
<collectingCountry id="F34EC395FFA91960987AF9CF65AD457C" box="[571,642,1626,1652]" name="China" pageId="7" pageNumber="160">China</collectingCountry>
:
<collectingRegion id="499D4DE7FFA9196098C6F9CE65CF457D" box="[647,736,1627,1653]" country="China" name="Yunnan" pageId="7" pageNumber="160">Yunnan</collectingRegion>
,
<collectingCounty id="6287FB89FFA9196098A7F9CF6411457C" box="[742,830,1626,1652]" pageId="7" pageNumber="160">Lvchun</collectingCounty>
,
<collectingMunicipality id="6B82197FFFA919609905F9CF64A0457C" box="[836,911,1626,1652]" pageId="7" pageNumber="160">Yakou</collectingMunicipality>
,
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[
<geoCoordinate id="EE6DE5C2FFA919609E03F9CF63F8457D" box="[1090,1239,1626,1653]" degrees="23" direction="north" minutes="3" orientation="latitude" pageId="7" pageNumber="160" precision="1" seconds="25.2" value="23.057">23°325.2”N</geoCoordinate>
,
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,
<quantity id="4CA12EE0FFA919609AD6F9EB67DD4591" box="[151,242,1662,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.779" pageId="7" pageNumber="160" unit="m" value="1779.0">
<elevation id="00746436FFA919609AD6F9EB67DD4591" box="[151,242,1662,1689]" metricMagnitude="3" metricUnit="m" metricValue="1.779" pageId="7" pageNumber="160" unit="m" value="1779.0">1779 m</elevation>
</quantity>
], collected by
<collectorName id="26ACE6D3FFA919609BD8F9EB66DD4590" box="[409,498,1662,1688]" pageId="7" pageNumber="160">Malaise</collectorName>
trap,
<date id="FFE7A5C5FFA919609877F9EB65D84590" box="[566,759,1662,1688]" pageId="7" pageNumber="160" value="2019-04-19" valueMax="2019-05-19" valueMin="2019-04-19">
<collectingDate id="EFA35C2DFFA919609877F9EB65D84590" box="[566,759,1662,1688]" pageId="7" pageNumber="160" value="2019-04-19" valueMax="2019-05-19" valueMin="2019-04-19">2019.IV.19V.19</collectingDate>
</date>
,
<collectorName id="26ACE6D3FFA919609942F9EB64A14591" box="[771,910,1662,1689]" pageId="7" pageNumber="160">Liang Wang</collectorName>
and
<collectorName id="26ACE6D3FFA919609986F9EB633C4590" box="[967,1043,1662,1688]" pageId="7" pageNumber="160">Xin Li</collectorName>
(
<collectionCode id="ED481BC0FFA919609E65F9EB634A4590" box="[1060,1125,1662,1688]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="7" pageNumber="160">CAU</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B318958FFA919609E3BF9EB668A45B4" collectingDate="2019-04-19" collectingDateMax="2019-05-19" collectingDateMin="2019-04-19" collectionCode="CAU" collectorName="Malaise &amp; Liang Wang &amp; Xin Li" country="China" county="Lvchun" elevation="1779" latitude="23.057" location="Shuikuxiafang" longLatPrecision="1" longitude="102.777145" municipality="Yakou" pageId="7" pageNumber="160" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B92D5F17FFA919609E3BF9EB62314590" bold="true" box="[1146,1310,1662,1688]" pageId="7" pageNumber="160">
<typeStatus id="54E23DA7FFA919609E3BF9EB623A4590" box="[1146,1301,1662,1688]" pageId="7" pageNumber="160" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
<specimenCount id="9D5F488CFFA919609F66F9EB627C4591" box="[1319,1363,1662,1689]" count="1" pageId="7" pageNumber="160" type="female">1 ♀</specimenCount>
, same data as holotype (
<collectionCode id="ED481BC0FFA919609B1DF93766B245B4" box="[348,413,1698,1724]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="7" pageNumber="160">CAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA919609A86F953650E45E8" box="[199,545,1734,1760]" pageId="7" pageNumber="160" type="distribution">
<paragraph id="8BE68305FFA919609A86F953650E45E8" blockId="7.[151,1437,510,1869]" box="[199,545,1734,1760]" pageId="7" pageNumber="160">
<emphasis id="B92D5F17FFA919609A86F953664E45E8" bold="true" box="[199,353,1734,1760]" pageId="7" pageNumber="160">Distribution.</emphasis>
<collectingCountry id="F34EC395FFA919609B26F953668545E8" box="[359,426,1734,1760]" name="China" pageId="7" pageNumber="160">China</collectingCountry>
(
<collectingRegion id="499D4DE7FFA919609BF8F953653A45E8" box="[441,533,1734,1760]" country="China" name="Yunnan" pageId="7" pageNumber="160">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA919609A86F97F64364420" pageId="7" pageNumber="160" type="discussion">
<paragraph id="8BE68305FFA919609A86F97F64364420" blockId="7.[151,1437,510,1869]" pageId="7" pageNumber="160">
<emphasis id="B92D5F17FFA919609A86F97F6614440C" bold="true" box="[199,315,1770,1796]" pageId="7" pageNumber="160">Remarks.</emphasis>
<taxonomicName id="4C59F886FFA919609B03F97E6517440C" authority="Lin &amp; Zhang &amp; Yang, 2023" authorityName="Lin &amp; Zhang &amp; Yang" authorityYear="2023" box="[322,568,1770,1797]" class="Insecta" family="Dolichopodidae" genus="Systenus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="160" phylum="Arthropoda" rank="species" species="flavilateralis" status="sp. nov.">
<emphasis id="B92D5F17FFA919609B03F97E6517440C" box="[322,568,1770,1797]" italics="true" pageId="7" pageNumber="160">Systenus flavilateralis</emphasis>
</taxonomicName>
<emphasis id="B92D5F17FFA91960987EF97F65B7440C" bold="true" box="[575,664,1770,1796]" pageId="7" pageNumber="160">
<taxonomicNameLabel id="A21EE26CFFA91960987EF97F65B7440C" box="[575,664,1770,1796]" pageId="7" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
here is somewhat similar to
<emphasis id="B92D5F17FFA919609999F97E6366440C" box="[984,1097,1771,1797]" italics="true" pageId="7" pageNumber="160">S. Tenorio</emphasis>
<bibRefCitation id="EFC8FEF4FFA919609E11F97F63F0440D" author="Bickel, D. J." box="[1104,1247,1770,1797]" pageId="7" pageNumber="160" pagination="169 - 182" refId="ref7927" refString="Bickel, D. J. (2015) The Costa Rican Systenus Loew (Diptera: Dolichopodidae): rich local sympatry in an otherwise rare genus. Zootaxa, 4020 (1), 169 - 182. https: // doi. org / 10.11646 / zootaxa. 4020.1.7" type="journal article" year="2015">Bickel, 2015</bibRefCitation>
from
<collectingCountry id="F34EC395FFA919609F63F97F62B3440C" box="[1314,1436,1770,1796]" name="Costa Rica" pageId="7" pageNumber="160">Costa Rica</collectingCountry>
in having the short thick postpedicel and subapical arista.
</paragraph>
</subSubSection>
<subSubSection id="C343D08EFFA919609A86F8A7636D4444" box="[199,1090,1842,1869]" pageId="7" pageNumber="160" type="etymology">
<paragraph id="8BE68305FFA919609A86F8A7636D4444" blockId="7.[151,1437,510,1869]" box="[199,1090,1842,1869]" pageId="7" pageNumber="160">
<emphasis id="B92D5F17FFA919609A86F8A766614444" bold="true" box="[199,334,1842,1868]" pageId="7" pageNumber="160">Etymology.</emphasis>
The specific name refers to the thoracic pleura mostly dark yellow.
</paragraph>
</subSubSection>
</treatment>
</document>