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<document id="C3D55DA61BF7E5E3DE62B5BE4DAD759B" ID-CLB-Dataset="26265" ID-DOI="10.11646/zootaxa.4629.4.1" ID-GBIF-Dataset="4d76ea5b-ee7f-4361-8f18-595631a770e6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3274050" ID-ZooBank="0DB18F90-AAFD-4FB7-B1ED-09A86A707C9F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1562653840885" checkinUser="plazi" docAuthor="Rodrigues, Higor D. D. &amp; Sites, Robert W." docDate="2019" docId="03E58815E920FFA9FF61F7E7FAE75BE0" docLanguage="en" docName="zootaxa.4629.4.1.pdf" docOrigin="Zootaxa 4629 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Limnocoris panamensis : La Rivers 1971" docType="treatment" docVersion="13" lastPageNumber="482" masterDocId="FFDCF06DE93CFFB6FFF6F34AFFB45F60" masterDocTitle="Revision of Limnocoris (Heteroptera: Nepomorpha: Naucoridae) of North America" masterLastPageNumber="497" masterPageNumber="451" pageNumber="479" updateTime="1698738605784" updateUser="ExternalLinkService">
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<mods:title id="F0FEE3A6E3DA181C5FAD1420E0B64A6C">Revision of Limnocoris (Heteroptera: Nepomorpha: Naucoridae) of North America</mods:title>
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<mods:namePart id="E971EE818B6DD91C848DE3D161B9C1E7">Rodrigues, Higor D. D.</mods:namePart>
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<subSubSection id="C3566A88E920FFAAFF61F7E7FD8E5BA6" box="[151,570,1196,1223]" pageId="28" pageNumber="479" type="nomenclature">
<paragraph id="8BF33903E920FFAAFF61F7E7FD8E5BA6" blockId="28.[151,570,1196,1259]" box="[151,570,1196,1223]" pageId="28" pageNumber="479">
<heading id="D0BB8E6FE920FFAAFF61F7E7FD8E5BA6" bold="true" box="[151,570,1196,1223]" fontSize="11" level="1" pageId="28" pageNumber="479" reason="1">
<taxonomicName id="4C4C4280E920FFAAFF61F7E7FD8E5BA6" ID-CoL="6QBMN" authority="La Rivers" authorityName=": La Rivers" authorityYear="1971" box="[151,570,1196,1223]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B938E511E920FFAAFF61F7E7FD8E5BA6" bold="true" box="[151,570,1196,1223]" pageId="28" pageNumber="479">
<emphasis id="B938E511E920FFAAFF61F7E7FE0C5BA7" bold="true" box="[151,440,1197,1223]" italics="true" pageId="28" pageNumber="479">Limnocoris panamensis</emphasis>
La Rivers
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3566A88E920FFAAFF61F79AFE765B8B" box="[151,450,1232,1259]" pageId="28" pageNumber="479" type="description">
<paragraph id="8BF33903E920FFAAFF61F79AFE765B8B" blockId="28.[151,570,1196,1259]" box="[151,450,1232,1259]" pageId="28" pageNumber="479">
(
<figureCitation id="13772586E920FFAAFF69F79AFF485B8B" box="[159,252,1232,1259]" captionStart="FIGURE 2" captionStartId="7.[151,250,1461,1486]" captionTargetBox="[156,1431,221,1424]" captionTargetId="figure@7.[151,1435,211,1426]" captionTargetPageId="7" captionText="FIGURE 2. (AB) Lateral view of meso- and metasternal carinae, (A) L. pygmaeus, white arrow indicates anterior projection, black arrow indicates excavation posterior to fossa, (B) L. zacki n. sp., black arrow indicates median projection of mesosternal carina. (CD) Ventral view of meso- and metasternal carinae, (C) L. inornatus, white arrow indicates row of longitudinal golden setae between mesobasisternum and mesoepisternum, (D) L. zacki n. sp., white arrow indicates fossa partially open anteriorly, dotted lines indicate anterior and posterior limits of fossa. (EF) Ventral view of abdominal segments IIIV, (E) L. virescens, black arrows indicate posterolateral corners, (F) L. panamensis. (GH) Ventral view of female abdominal segments IVIX, (G) L. moapensis, sgp = subgenital plate, (H) L. major n. sp., white arrows indicate elongate golden setae, gs= posterolateral elongate golden setae on lateral margin of subgenital plate." httpUri="https://zenodo.org/record/3274057/files/figure.png" pageId="28" pageNumber="479">Figs. 2F</figureCitation>
,
<figureCitation id="13772586E920FFAAFEFEF79BFE925B8B" box="[264,294,1233,1259]" captionStart="FIGURE 13" captionStartId="29.[151,250,1924,1949]" captionTargetBox="[192,1398,172,1898]" captionTargetId="figure@29.[184,1404,166,1904]" captionTargetPageId="29" captionText="FIGURE 13. Limnocoris panamensis La Rivers. (A) Dorsal habitus and (B) ventral habitus of female paratype, hindwing brachypterous (USNM), (C) ventral view of part of head and thorax, (D) terminal abdominal sterna of female, and (E) lateral view of meso- and metasternal carinae. Size bar = 1.00 mm and applies only to Figs. AB." httpUri="https://zenodo.org/record/3274079/files/figure.png" pageId="28" pageNumber="479">13</figureCitation>
,
<figureCitation id="13772586E920FFAAFEC4F79BFEF85B8B" box="[306,332,1233,1259]" captionStart="FIGURE 14" captionStartId="30.[151,250,1653,1678]" captionTargetBox="[151,1435,273,1634]" captionTargetId="figure@30.[151,1435,273,1634]" captionTargetPageId="30" captionText="FIGURE 14. Structures of male terminalia. (A, C, E) 6th and 7th abdominal terga, (B, D, F) 8th abdominal tergum. (AB) Limnocoris moapensis, (CD) L. nanus n. sp., and (EF) L. panamensis." figureDoi="http://doi.org/10.5281/zenodo.3446039" httpUri="https://zenodo.org/record/3446039/files/figure.png" pageId="28" pageNumber="479">14</figureCitation>
EF, 20A)
</paragraph>
</subSubSection>
<subSubSection id="C3566A88E920FFAAFF61F65FFA265A2C" pageId="28" pageNumber="479" type="reference_group">
<paragraph id="8BF33903E920FFAAFF61F65FFCFD5A4D" blockId="28.[151,1426,1300,1356]" box="[151,841,1300,1325]" pageId="28" pageNumber="479">
<taxonomicName id="4C4C4280E920FFAAFF61F65FFDD15A4D" ID-CoL="6QBMN" authority="La Rivers, 1970: 6 - 8" authorityName=": La Rivers" authorityPageNumber="6" authorityYear="1971" box="[151,613,1300,1325]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B938E511E920FFAAFF61F65FFE3D5A4C" box="[151,393,1301,1324]" italics="true" pageId="28" pageNumber="479">Limnocoris panamensis</emphasis>
<bibRefCitation id="EFDD44F2E920FFAAFE79F65FFDD15A4D" author="La Rivers, I." box="[399,613,1300,1325]" pageId="28" pageNumber="479" pagination="1 - 10" refId="ref32890" refString="La Rivers, I. (1970) New species of Limnocoris from Latin America (Hemiptea, Naucoridae). Biological Society of Nevada Occasional Papers, 23, 1 - 10." type="journal article" year="1970">La Rivers, 1970: 68</bibRefCitation>
</taxonomicName>
(original description).
</paragraph>
<paragraph id="8BF33903E920FFAAFF61F67EFA265A2C" blockId="28.[151,1426,1300,1356]" box="[151,1426,1331,1356]" pageId="28" pageNumber="479">
<treatmentCitationGroup id="AB5C1E2DE920FFAAFF61F67EFA265A2C" box="[151,1426,1331,1356]" pageId="28" pageNumber="479">
<taxonomicName id="4C4C4280E920FFAAFF61F67EFDED5A2C" ID-CoL="6QBMN" authority=": La Rivers 1971: 76" authorityName=": La Rivers" authorityPageNumber="76" authorityYear="1971" box="[151,601,1331,1356]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="479" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B938E511E920FFAAFF61F67EFE3D5A2B" box="[151,393,1332,1355]" italics="true" pageId="28" pageNumber="479">Limnocoris panamensis</emphasis>
:
<treatmentCitation id="0AED1F12E920FFAAFE60F67EFDED5A2C" author="La Rivers, I." box="[406,601,1331,1356]" page="76" pageId="28" pageNumber="479" year="1971">
<bibRefCitation id="EFDD44F2E920FFAAFE60F67EFDED5A2C" author="La Rivers, I." box="[406,601,1331,1356]" pageId="28" pageNumber="479" pagination="1 - 120" refId="ref32924" refString="La Rivers, I. (1971) Studies of Naucoridae (Hemiptera). Biological Society of Nevada Memoirs, 2, 1 - 120." type="journal article" year="1971">La Rivers 1971: 76</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(catalog);
<treatmentCitation id="0AED1F12E920FFAAFD31F679FBB75A2B" author="Nieser, N. &amp; Lopez-Ruf, M." box="[711,1027,1331,1356]" page="319" pageId="28" pageNumber="479" year="2001">
<bibRefCitation id="EFDD44F2E920FFAAFD31F679FBB75A2B" author="Nieser, N. &amp; Lopez-Ruf, M." box="[711,1027,1331,1356]" pageId="28" pageNumber="479" pagination="261 - 328" refId="ref33464" refString="Nieser, N. &amp; Lopez-Ruf, M. (2001) A review of Limnocoris Stal (Heteroptera: Naucoridae) in southern South America east of the Andes. Tijdschrift voor Entomologie, 144, 261 - 328. https: // doi. org / 10.1163 / 22119434 - 900000091" type="journal article" year="2001">Nieser &amp; López-Ruf 2001: 319</bibRefCitation>
</treatmentCitation>
(catalog);
<treatmentCitation id="0AED1F12E920FFAAFB87F67EFA9F5A2B" author="Herrera, F." box="[1137,1323,1331,1355]" page="227" pageId="28" pageNumber="479" year="2013">
<bibRefCitation id="EFDD44F2E920FFAAFB87F67EFA9F5A2B" author="Herrera, F." box="[1137,1323,1331,1355]" pageId="28" pageNumber="479" pagination="221 - 232" refId="ref32671" refString="Herrera, F. (2013) Estado del conocimiento y catalogo de la familia Naucoridae (Insecta: Heteroptera) en Centroamerica. Dugesiana, 20 (2), 221 - 232." type="journal article" year="2013">
<collectingRegion id="4988F7E1E920FFAAFB87F67EFB0A5A2B" box="[1137,1214,1332,1355]" country="Panama" name="Herrera" pageId="28" pageNumber="479">Herrera</collectingRegion>
2013: 227
</bibRefCitation>
</treatmentCitation>
(catalog).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3566A88E920FFAAFF61F633FB505960" pageId="28" pageNumber="479" type="diagnosis">
<paragraph id="8BF33903E920FFAAFF61F633FB505960" blockId="28.[151,1437,1401,2040]" pageId="28" pageNumber="479">
<emphasis id="B938E511E920FFAAFF61F633FEA65AF4" bold="true" box="[151,274,1401,1428]" pageId="28" pageNumber="479">Diagnosis.</emphasis>
The pubescent area of the propleuron is not extended posteriorly on the lateral margin (as in
<figureCitation id="13772586E920FFAAFAC3F630FA3A5AF4" box="[1333,1422,1402,1428]" captionStart="FIGURE 1" captionStartId="6.[151,250,1505,1530]" captionTargetBox="[155,1429,494,1482]" captionTargetId="figure@6.[151,1432,489,1485]" captionTargetPageId="6" captionText="FIGURE 1. Structures of Limnocoris. (AD) Propleuron, white arrows indicate posterior end of pubescent area, yellow arrows indicate mid-lateral region of posterior margin, (A) L. zacki n. sp., (B) L. inornatus, (C) L. virescens, (D) L. chaetocarinatus n. sp. (EH) Part of abdominal segments, showing pubescence patterns of sternum II (circled), (E) L. moapensis, (F) L. submontandoni, (G) L. nanus n. sp., and (H) L. stali. pa = pubescent area, pra = pruinose area." httpUri="https://zenodo.org/record/3274054/files/figure.png" pageId="28" pageNumber="479">Fig. 1D</figureCitation>
). The fossa of the mesosternal carina is distinctly deep and elliptical to ovate (
<figureCitation id="13772586E920FFAAFBF4F6D4FBDE5AD8" box="[1026,1130,1438,1464]" captionStart="FIGURE 13" captionStartId="29.[151,250,1924,1949]" captionTargetBox="[192,1398,172,1898]" captionTargetId="figure@29.[184,1404,166,1904]" captionTargetPageId="29" captionText="FIGURE 13. Limnocoris panamensis La Rivers. (A) Dorsal habitus and (B) ventral habitus of female paratype, hindwing brachypterous (USNM), (C) ventral view of part of head and thorax, (D) terminal abdominal sterna of female, and (E) lateral view of meso- and metasternal carinae. Size bar = 1.00 mm and applies only to Figs. AB." httpUri="https://zenodo.org/record/3274079/files/figure.png" pageId="28" pageNumber="479">Fig. 13C</figureCitation>
). The metasternal carina is depressed medially, and acuminate posteriorly (
<figureCitation id="13772586E920FFAAFD45F688FCAF5ABC" box="[691,795,1474,1500]" captionStart="FIGURE 13" captionStartId="29.[151,250,1924,1949]" captionTargetBox="[192,1398,172,1898]" captionTargetId="figure@29.[184,1404,166,1904]" captionTargetPageId="29" captionText="FIGURE 13. Limnocoris panamensis La Rivers. (A) Dorsal habitus and (B) ventral habitus of female paratype, hindwing brachypterous (USNM), (C) ventral view of part of head and thorax, (D) terminal abdominal sterna of female, and (E) lateral view of meso- and metasternal carinae. Size bar = 1.00 mm and applies only to Figs. AB." httpUri="https://zenodo.org/record/3274079/files/figure.png" pageId="28" pageNumber="479">Fig. 13C</figureCitation>
). The lateral margin of the female subgenital plate has a tuft of elongate golden setae at mid-length, and the posterior margin is evenly rounded (
<figureCitation id="13772586E920FFAAFB99F6ACFB635960" box="[1135,1239,1510,1536]" captionStart="FIGURE 13" captionStartId="29.[151,250,1924,1949]" captionTargetBox="[192,1398,172,1898]" captionTargetId="figure@29.[184,1404,166,1904]" captionTargetPageId="29" captionText="FIGURE 13. Limnocoris panamensis La Rivers. (A) Dorsal habitus and (B) ventral habitus of female paratype, hindwing brachypterous (USNM), (C) ventral view of part of head and thorax, (D) terminal abdominal sterna of female, and (E) lateral view of meso- and metasternal carinae. Size bar = 1.00 mm and applies only to Figs. AB." httpUri="https://zenodo.org/record/3274079/files/figure.png" pageId="28" pageNumber="479">Fig. 13D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3566A88E920FFA9FF31F540FAAA5E7F" lastPageId="31" lastPageNumber="482" pageId="28" pageNumber="479" type="description">
<paragraph id="8BF33903E920FFAAFF31F540FC2159F0" blockId="28.[151,1437,1401,2040]" pageId="28" pageNumber="479">
<emphasis id="B938E511E920FFAAFF31F540FEC25944" bold="true" box="[199,374,1546,1572]" pageId="28" pageNumber="479">Measurements</emphasis>
(n =
<specimenCount id="9D4AF28AE920FFAAFE47F540FDB25944" box="[433,518,1546,1572]" pageId="28" pageNumber="479" type="male">3 males</specimenCount>
/
<specimenCount id="9D4AF28AE920FFAAFDE3F540FD315945" box="[533,645,1546,1573]" pageId="28" pageNumber="479" type="female">3 females</specimenCount>
). Body length 8.208.23/ 8.608.74, body width 5.655.80/ 5.866.00, synthlipsis 1.491.52/ 1.501.51, head length 1.211.40/ 1.40, head width 2.752.80/ 2.802.82, pronotum length on midline 1.50/ 1.501.60, pronotum width 5.373.40/ 5.655.68, scutellum length 1.201.40/ 1.40, scutellum width 2.512.70/ 2.612.80, hemelytra length 5.906.12/ 6.206.40.
</paragraph>
<paragraph id="8BF33903E920FFA8FF31F5D3FDDC5F9B" blockId="28.[151,1437,1401,2040]" lastBlockId="30.[151,1437,153,251]" lastPageId="30" lastPageNumber="481" pageId="28" pageNumber="479">
<emphasis id="B938E511E920FFAAFF31F5D3FDB659D4" bold="true" box="[199,514,1689,1716]" pageId="28" pageNumber="479">Supplemental description.</emphasis>
Maxillary plate tumescent anteriorly. Distolateral margins of labrum tapering to broadly rounded point. Antenna not exceeding lateral margin of eye; pedicel quadrate; flagellomeres slender, not partially fused, with long setae. Posterior margin of propleuron convex at mid-length (
<figureCitation id="13772586E920FFAAFB70F5A8FB44599C" box="[1158,1264,1762,1788]" captionStart="FIGURE 13" captionStartId="29.[151,250,1924,1949]" captionTargetBox="[192,1398,172,1898]" captionTargetId="figure@29.[184,1404,166,1904]" captionTargetPageId="29" captionText="FIGURE 13. Limnocoris panamensis La Rivers. (A) Dorsal habitus and (B) ventral habitus of female paratype, hindwing brachypterous (USNM), (C) ventral view of part of head and thorax, (D) terminal abdominal sterna of female, and (E) lateral view of meso- and metasternal carinae. Size bar = 1.00 mm and applies only to Figs. AB." httpUri="https://zenodo.org/record/3274079/files/figure.png" pageId="28" pageNumber="479">Fig. 13C</figureCitation>
); posteromesal corner near prosternellum flat. Region between mesobasisternum and mesoepisternum with longitudinal row of elongate golden setae (partially visible in
<figureCitation id="13772586E920FFAAFD99F460FD615824" box="[623,725,1834,1860]" captionStart="FIGURE 13" captionStartId="29.[151,250,1924,1949]" captionTargetBox="[192,1398,172,1898]" captionTargetId="figure@29.[184,1404,166,1904]" captionTargetPageId="29" captionText="FIGURE 13. Limnocoris panamensis La Rivers. (A) Dorsal habitus and (B) ventral habitus of female paratype, hindwing brachypterous (USNM), (C) ventral view of part of head and thorax, (D) terminal abdominal sterna of female, and (E) lateral view of meso- and metasternal carinae. Size bar = 1.00 mm and applies only to Figs. AB." httpUri="https://zenodo.org/record/3274079/files/figure.png" pageId="28" pageNumber="479">Fig. 13C</figureCitation>
). Lateral margin of abdomen with minute serration; posterolateral corners of IIVI narrowly rounded to right angled, not spinose; sterna with elongate golden setae generally dispersed, concentrated at midline of segments IIIV; sternum II with sinuous row of golden setae.
<emphasis id="B938E511E920FFAAFB31F438FAB558EC" box="[1223,1281,1906,1932]" italics="true" pageId="28" pageNumber="479">Male</emphasis>
: mediotergite VI with accessory genitalic process poorly developed; posterior margin of mediotergite VII convex medially, with small rounded lobe on each side; laterotergite VII with lateral and mesal margins subparallel, and posterior margin narrowly rounded (
<figureCitation id="13772586E920FFAAFE87F494FE6D5898" box="[369,473,2014,2040]" captionStart="FIGURE 14" captionStartId="30.[151,250,1653,1678]" captionTargetBox="[151,1435,273,1634]" captionTargetId="figure@30.[151,1435,273,1634]" captionTargetPageId="30" captionText="FIGURE 14. Structures of male terminalia. (A, C, E) 6th and 7th abdominal terga, (B, D, F) 8th abdominal tergum. (AB) Limnocoris moapensis, (CD) L. nanus n. sp., and (EF) L. panamensis." figureDoi="http://doi.org/10.5281/zenodo.3446039" httpUri="https://zenodo.org/record/3446039/files/figure.png" pageId="28" pageNumber="479">Fig. 14E</figureCitation>
). Lateral lobe of tergum VIII with lateral margin almost straight in anterior half; left medial lobe slightly angled laterally at apex, distal margin rounded; right medial lobe twisted in distal third (
<figureCitation id="13772586E922FFA8FA99F3D3FF725FB7" captionStart="FIGURE 14" captionStartId="30.[151,250,1653,1678]" captionTargetBox="[151,1435,273,1634]" captionTargetId="figure@30.[151,1435,273,1634]" captionTargetPageId="30" captionText="FIGURE 14. Structures of male terminalia. (A, C, E) 6th and 7th abdominal terga, (B, D, F) 8th abdominal tergum. (AB) Limnocoris moapensis, (CD) L. nanus n. sp., and (EF) L. panamensis." figureDoi="http://doi.org/10.5281/zenodo.3446039" httpUri="https://zenodo.org/record/3446039/files/figure.png" pageId="30" pageNumber="481">Fig. 14F</figureCitation>
).
<emphasis id="B938E511E922FFA8FF2CF3F7FE9A5FB7" box="[218,302,189,215]" italics="true" pageId="30" pageNumber="481">Female</emphasis>
: subgenital plate with posterior margin broadly convex; laterosternite VII with posterior margin shal- lowly angled to rounded apex (
<figureCitation id="13772586E922FFA8FE05F3ABFDEF5F9B" box="[499,603,225,251]" captionStart="FIGURE 13" captionStartId="29.[151,250,1924,1949]" captionTargetBox="[192,1398,172,1898]" captionTargetId="figure@29.[184,1404,166,1904]" captionTargetPageId="29" captionText="FIGURE 13. Limnocoris panamensis La Rivers. (A) Dorsal habitus and (B) ventral habitus of female paratype, hindwing brachypterous (USNM), (C) ventral view of part of head and thorax, (D) terminal abdominal sterna of female, and (E) lateral view of meso- and metasternal carinae. Size bar = 1.00 mm and applies only to Figs. AB." httpUri="https://zenodo.org/record/3274079/files/figure.png" pageId="30" pageNumber="481">Fig. 13D</figureCitation>
).
</paragraph>
<caption id="DF33698BE921FFABFF61F4CEFC6D5885" httpUri="https://zenodo.org/record/3274079/files/figure.png" pageId="29" pageNumber="480" startId="29.[151,250,1924,1949]" targetBox="[192,1398,172,1898]" targetPageId="29">
<paragraph id="8BF33903E921FFABFF61F4CEFC6D5885" blockId="29.[151,1437,1924,2021]" pageId="29" pageNumber="480">
<emphasis id="B938E511E921FFABFF61F4CEFEAA58FD" bold="true" box="[151,286,1924,1949]" pageId="29" pageNumber="480">FIGURE 13.</emphasis>
<taxonomicName id="4C4C4280E921FFABFED2F4CCFD3658FD" authority="La Rivers." authorityName=": La Rivers" authorityYear="1971" box="[292,642,1924,1949]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B938E511E921FFABFED2F4CCFDA058FD" box="[292,532,1926,1949]" italics="true" pageId="29" pageNumber="480">Limnocoris panamensis</emphasis>
La Rivers.
</taxonomicName>
(A) Dorsal habitus and (B) ventral habitus of female paratype, hindwing brachypterous (USNM), (C) ventral view of part of head and thorax, (D) terminal abdominal sterna of female, and (E) lateral view of meso- and metasternal carinae. Size bar = 1.00 mm and applies only to Figs. AB.
</paragraph>
</caption>
<caption id="DF33698BE922FFA8FF61F53FFCCC59D1" ID-DOI="http://doi.org/10.5281/zenodo.3446039" ID-Zenodo-Dep="3446039" httpUri="https://zenodo.org/record/3446039/files/figure.png" pageId="30" pageNumber="481" startId="30.[151,250,1653,1678]" targetBox="[151,1435,273,1634]" targetPageId="30">
<paragraph id="8BF33903E922FFA8FF61F53FFCCC59D1" blockId="30.[151,1436,1653,1714]" pageId="30" pageNumber="481">
<emphasis id="B938E511E922FFA8FF61F53FFE9659EE" bold="true" box="[151,290,1653,1678]" pageId="30" pageNumber="481">FIGURE 14.</emphasis>
Structures of male terminalia. (A, C, E) 6th and 7th abdominal terga, (B, D, F) 8th abdominal tergum. (AB)
<emphasis id="B938E511E922FFA8FF61F5D1FEC859D2" box="[151,380,1691,1714]" italics="true" pageId="30" pageNumber="481">Limnocoris moapensis</emphasis>
, (CD)
<emphasis id="B938E511E922FFA8FE26F5D1FD9359D2" box="[464,551,1691,1714]" italics="true" pageId="30" pageNumber="481">L. nanus</emphasis>
<emphasis id="B938E511E922FFA8FDDBF5D0FDD259D2" bold="true" box="[557,614,1690,1714]" pageId="30" pageNumber="481">n. sp.</emphasis>
, and (EF)
<emphasis id="B938E511E922FFA8FD29F5D1FCC659D2" box="[735,882,1691,1714]" italics="true" pageId="30" pageNumber="481">L. panamensis</emphasis>
.
</paragraph>
</caption>
<paragraph id="8BF33903E922FFA9FF31F5AAFAAA5E7F" blockId="30.[151,1437,1760,2039]" lastBlockId="31.[151,1437,153,1152]" lastPageId="31" lastPageNumber="482" pageId="30" pageNumber="481">
<emphasis id="B938E511E922FFA8FF31F5AAFE1B599B" bold="true" box="[199,431,1760,1787]" pageId="30" pageNumber="481">Comparative notes.</emphasis>
This species has a characteristic mesosternal carina with a distinctly deep fossa. Five other species of
<taxonomicName id="4C4C4280E922FFA8FEFAF44FFE38587E" box="[268,396,1797,1822]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="genus">
<emphasis id="B938E511E922FFA8FEFAF44FFE38587E" box="[268,396,1797,1822]" italics="true" pageId="30" pageNumber="481">Limnocoris</emphasis>
</taxonomicName>
share a deep fossa of the mesosternal carina:
<taxonomicName id="4C4C4280E922FFA8FC72F44FFACF587E" authority="Nieser, Gonzalez &amp; Eichelkraut" authorityName="Nieser, Gonzalez &amp; Eichelkraut" authorityYear="1993" box="[900,1403,1796,1823]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="B938E511E922FFA8FC72F44FFBBB587E" box="[900,1039,1796,1822]" italics="true" pageId="30" pageNumber="481">L. angulatus</emphasis>
Nieser, González &amp; Eichelkraut
</taxonomicName>
,
<taxonomicName id="4C4C4280E922FFA8FA70F44FFECB5822" authority="Montandon" authorityName="Montandon" authorityYear="1898" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="bouvieri">
<emphasis id="B938E511E922FFA8FA70F44FFF405822" italics="true" pageId="30" pageNumber="481">L. bouvieri</emphasis>
Montandon
</taxonomicName>
,
<taxonomicName id="4C4C4280E922FFA8FE7CF463FCFC5822" authority="Nieser, Gonzalez &amp; Eichelkraut" authorityName="Nieser, Gonzalez &amp; Eichelkraut" authorityYear="1993" box="[394,840,1832,1859]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="calii">
<emphasis id="B938E511E922FFA8FE7CF463FE6E5822" box="[394,474,1832,1858]" italics="true" pageId="30" pageNumber="481">L. calii</emphasis>
Nieser, González &amp; Eichelkraut
</taxonomicName>
,
<taxonomicName id="4C4C4280E922FFA8FCA5F463FBDF5822" authority="Montandon" authorityName="Montandon" authorityYear="1897" box="[851,1131,1832,1858]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="pectoralis">
<emphasis id="B938E511E922FFA8FCA5F463FC545822" box="[851,992,1832,1858]" italics="true" pageId="30" pageNumber="481">L. pectoralis</emphasis>
Montandon
</taxonomicName>
, and
<taxonomicName id="4C4C4280E922FFA8FB53F463FA9D5822" box="[1189,1321,1833,1858]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="virescens">
<emphasis id="B938E511E922FFA8FB53F463FA9D5822" box="[1189,1321,1833,1858]" italics="true" pageId="30" pageNumber="481">L. virescens</emphasis>
</taxonomicName>
. Of these, only
<taxonomicName id="4C4C4280E922FFA8FF27F407FEE25806" box="[209,342,1869,1894]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="virescens">
<emphasis id="B938E511E922FFA8FF27F407FEE25806" box="[209,342,1869,1894]" italics="true" pageId="30" pageNumber="481">L. virescens</emphasis>
</taxonomicName>
occurs in North America. In
<taxonomicName id="4C4C4280E922FFA8FD54F407FCA95806" authorityName="Montandon" authorityYear="1898" box="[674,797,1868,1894]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="bouvieri">
<emphasis id="B938E511E922FFA8FD54F407FCA95806" box="[674,797,1868,1894]" italics="true" pageId="30" pageNumber="481">L. bouvieri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4280E922FFA8FCA0F407FC6F5806" box="[854,987,1869,1894]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="virescens">
<emphasis id="B938E511E922FFA8FCA0F407FC6F5806" box="[854,987,1869,1894]" italics="true" pageId="30" pageNumber="481">L. virescens</emphasis>
</taxonomicName>
the posterolateral corners of abdominal segments IIIV are spinose, whereas in
<taxonomicName id="4C4C4280E922FFA8FD97F43BFCB058EA" authorityName=": La Rivers" authorityYear="1971" box="[609,772,1905,1930]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B938E511E922FFA8FD97F43BFCB058EA" box="[609,772,1905,1930]" italics="true" pageId="30" pageNumber="481">L. panamensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4280E922FFA8FCE2F43BFC1658EA" authorityName="Nieser, Gonzalez &amp; Eichelkraut" authorityYear="1993" box="[788,930,1904,1930]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="B938E511E922FFA8FCE2F43BFC1658EA" box="[788,930,1904,1930]" italics="true" pageId="30" pageNumber="481">L. angulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4280E922FFA8FC44F43BFBB758EA" authorityName="Nieser, Gonzalez &amp; Eichelkraut" authorityYear="1993" box="[946,1027,1904,1930]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="calii">
<emphasis id="B938E511E922FFA8FC44F43BFBB758EA" box="[946,1027,1904,1930]" italics="true" pageId="30" pageNumber="481">L. calii</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C4C4280E922FFA8FBB2F43BFB6058EA" authorityName="Montandon" authorityYear="1897" box="[1092,1236,1904,1930]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="pectoralis">
<emphasis id="B938E511E922FFA8FBB2F43BFB6058EA" box="[1092,1236,1904,1930]" italics="true" pageId="30" pageNumber="481">L. pectoralis</emphasis>
</taxonomicName>
these corners are slightly acuminate to rounded, without forming spines.
<emphasis id="B938E511E922FFA8FD0BF4DFFBC258CE" box="[765,1142,1940,1966]" italics="true" pageId="30" pageNumber="481">
<taxonomicName id="4C4C4280E922FFA8FD0BF4DFFC0058CE" authorityName="Nieser, Gonzalez &amp; Eichelkraut" authorityYear="1993" box="[765,948,1940,1966]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="calii">Limnocoris calii</taxonomicName>
and
<taxonomicName id="4C4C4280E922FFA8FC1FF4DFFBC258CE" authorityName="Montandon" authorityYear="1897" box="[1001,1142,1940,1966]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="pectoralis">L. pectoralis</taxonomicName>
</emphasis>
differ from
<taxonomicName id="4C4C4280E922FFA8FB0AF4DFFA2858CE" authorityName=": La Rivers" authorityYear="1971" box="[1276,1436,1941,1966]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B938E511E922FFA8FB0AF4DFFA2858CE" box="[1276,1436,1941,1966]" italics="true" pageId="30" pageNumber="481">L. panamensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4280E922FFA8FF30F4F3FEE658B2" authorityName="Nieser, Gonzalez &amp; Eichelkraut" authorityYear="1993" box="[198,338,1976,2002]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="B938E511E922FFA8FF30F4F3FEE658B2" box="[198,338,1976,2002]" italics="true" pageId="30" pageNumber="481">L. angulatus</emphasis>
</taxonomicName>
by the pubescent area of the propleuron extended posteriorly, which does not occur in the last two species.
<taxonomicName id="4C4C4280E922FFA8FF01F497FDB65896" authorityName=": La Rivers" authorityYear="1971" box="[247,514,2013,2038]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B938E511E922FFA8FF01F497FDB65896" box="[247,514,2013,2038]" italics="true" pageId="30" pageNumber="481">Limnocoris panamensis</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="4C4C4280E922FFA8FCC0F497FC765896" authorityName="Nieser, Gonzalez &amp; Eichelkraut" authorityYear="1993" box="[822,962,2012,2038]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="B938E511E922FFA8FCC0F497FC765896" box="[822,962,2012,2038]" italics="true" pageId="30" pageNumber="481">L. angulatus</emphasis>
</taxonomicName>
by having the posterolateral corner of the pronotum rounded in brachypterous specimens, male abdominal tergum VIII with the right medial lobe twisted in the distal half, and the female subgenital plate with the posterior margin broadly rounded; whereas in
<taxonomicName id="4C4C4280E923FFA9FAD0F3F4FF0C5F9B" authorityName="Nieser, Gonzalez &amp; Eichelkraut" authorityYear="1993" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="B938E511E923FFA9FAD0F3F4FF0C5F9B" italics="true" pageId="31" pageNumber="482">L. angulatus</emphasis>
</taxonomicName>
the posterolateral corner of pronotum is acute, male abdominal tergum VIII with the right medial lobe weakly twisted, and the female subgenital plate tapering posteriorly and the posterior margin truncate medially.
</paragraph>
</subSubSection>
<subSubSection id="C3566A88E923FFA9FF31F262FCD45ECF" pageId="31" pageNumber="482" type="distribution">
<paragraph id="8BF33903E923FFA9FF31F262FEFE5EEB" blockId="31.[151,1437,153,1152]" pageId="31" pageNumber="482">
<emphasis id="B938E511E923FFA9FF31F262FED55E23" bold="true" box="[199,353,296,323]" pageId="31" pageNumber="482">Distribution.</emphasis>
This species is distributed from northern
<collectingCountry id="F35B7993E923FFA9FCDDF263FC105E23" box="[811,932,297,323]" name="Costa Rica" pageId="31" pageNumber="482">Costa Rica</collectingCountry>
(
<collectingRegion id="4988F7E1E923FFA9FC47F263FB825E23" box="[945,1078,297,323]" country="Costa Rica" name="Guanacaste" pageId="31" pageNumber="482">Guanacaste</collectingRegion>
) to the central region of
<collectingCountry id="F35B7993E923FFA9FAB2F263FA295E23" box="[1348,1437,297,323]" name="Panama" pageId="31" pageNumber="482">Panamá</collectingCountry>
(
<collectingRegion id="4988F7E1E923FFA9FF69F207FEBD5E07" box="[159,265,333,359]" country="Panama" name="Veraguas" pageId="31" pageNumber="482">Veraguas</collectingRegion>
) (
<figureCitation id="13772586E923FFA9FED6F207FE3C5E07" box="[288,392,333,359]" captionStart="FIGURE 20" captionStartId="42.[151,250,2002,2027]" captionTargetBox="[188,1396,171,1974]" captionTargetId="figure@42.[182,1405,164,1981]" captionTargetPageId="42" captionText="FIGURE 20. (AB) Distribution records for species of Limnocoris in North America." httpUri="https://zenodo.org/record/3274091/files/figure.png" pageId="31" pageNumber="482">Fig. 20A</figureCitation>
). Label data suggest that
<taxonomicName id="4C4C4280E923FFA9FD55F204FCF05E07" authorityName=": La Rivers" authorityYear="1971" box="[675,836,334,359]" class="Insecta" family="Naucoridae" genus="Limnocoris" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="panamensis">
<emphasis id="B938E511E923FFA9FD55F204FCF05E07" box="[675,836,334,359]" italics="true" pageId="31" pageNumber="482">L. panamensis</emphasis>
</taxonomicName>
occurs in regions with elevations above 1,000 meters above sea level.
</paragraph>
<paragraph id="8BF33903E923FFA9FF31F2DEFCD45ECF" blockId="31.[151,1437,153,1152]" box="[199,864,404,431]" pageId="31" pageNumber="482">
<emphasis id="B938E511E923FFA9FF31F2DEFE115ECF" bold="true" box="[199,421,404,431]" pageId="31" pageNumber="482">Published records.</emphasis>
<collectingCountry id="F35B7993E923FFA9FE5AF2DFFD9F5ECF" box="[428,555,405,431]" name="Costa Rica" pageId="31" pageNumber="482">Costa Rica</collectingCountry>
: Chiriquí (
<bibRefCitation id="EFDD44F2E923FFA9FD57F2DFFCE75ECF" author="La Rivers, I." box="[673,851,405,431]" pageId="31" pageNumber="482" pagination="1 - 10" refId="ref32890" refString="La Rivers, I. (1970) New species of Limnocoris from Latin America (Hemiptea, Naucoridae). Biological Society of Nevada Occasional Papers, 23, 1 - 10." type="journal article" year="1970">La Rivers 1970</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3566A88E923FFA9FF31F2F3FAE75BE0" pageId="31" pageNumber="482" type="materials_examined">
<paragraph id="8BF33903E923FFA9FF31F2F3FEFE5D7B" blockId="31.[151,1437,153,1152]" pageId="31" pageNumber="482">
<emphasis id="B938E511E923FFA9FF31F2F3FE595EB3" bold="true" box="[199,493,440,467]" pageId="31" pageNumber="482">Type material examined</emphasis>
. All specimens brachypterous.
<typeStatus id="54F787A1E923FFA9FCAFF2F3FC595EB3" box="[857,1005,441,467]" pageId="31" pageNumber="482" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="9D4AF28AE923FFA9FC00F2F3FBBF5EB3" box="[1014,1035,441,467]" pageId="31" pageNumber="482" type="female"></specimenCount>
,
<emphasis id="B938E511E923FFA9FBEAF2F2FB295EB3" bold="true" box="[1052,1181,440,467]" pageId="31" pageNumber="482">
<collectingCountry id="F35B7993E923FFA9FBEAF2F2FB2C5EB3" box="[1052,1176,440,467]" name="Panama" pageId="31" pageNumber="482">PANAMA</collectingCountry>
,
</emphasis>
[
<collectingRegion id="4988F7E1E923FFA9FB46F2F2FAA35EB3" box="[1200,1303,440,467]" country="Panama" name="Chiriqui" pageId="31" pageNumber="482">
<emphasis id="B938E511E923FFA9FB46F2F2FAA35EB3" bold="true" box="[1200,1303,440,467]" pageId="31" pageNumber="482">Chiriquí</emphasis>
</collectingRegion>
], Poterillos [=Potrerillos],
<date id="FFF21FC3E923FFA9FEC8F296FE0A5E97" box="[318,446,476,503]" pageId="31" pageNumber="482" value="1935-03-27">27.III.1935</date>
, J. MacSwain Collector, Type No. 13422 (
<collectionCode id="ED5DA1C6E923FFA9FC50F297FC565E97" box="[934,994,477,503]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="31" pageNumber="482" type="Museum">CAS</collectionCode>
).
<typeStatus id="54F787A1E923FFA9FC00F297FB215E97" box="[1014,1173,477,503]" pageId="31" pageNumber="482" type="paratype">PARATYPES</typeStatus>
: same data as
<typeStatus id="54F787A1E923FFA9FACCF297FA285E97" box="[1338,1436,477,503]" pageId="31" pageNumber="482" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D4AF28AE923FFA9FF57F14BFF715D7B" box="[161,197,513,539]" pageId="31" pageNumber="482" type="male">1♂</specimenCount>
,
<specimenCount id="9D4AF28AE923FFA9FF24F14BFF405D7B" box="[210,244,513,539]" pageId="31" pageNumber="482" type="female">1♀</specimenCount>
,
<collectionCode id="ED5DA1C6E923FFA9FEF5F14BFE8A5D7B" box="[259,318,513,539]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="31" pageNumber="482" type="Museum">CAS</collectionCode>
).
</paragraph>
<paragraph id="8BF33903E923FFA9FF31F16EFAE75BE0" blockId="31.[151,1437,153,1152]" pageId="31" pageNumber="482">
<materialsCitation id="3B24335EE923FFA9FF31F16EFC4A5D03" ID-GBIF-Occurrence="2273889337" collectingDate="2002-11-25" collectionCode="UMC" collectorName="W. D. Shepard" country="Costa Rica" county="Rio Cataratas" location="Fortuna" municipality="San Ramon" pageId="31" pageNumber="482" specimenCount="1" specimenCount-male="1" stateProvince="Alajuela">
<emphasis id="B938E511E923FFA9FF31F16EFD865D5F" bold="true" box="[199,562,548,575]" pageId="31" pageNumber="482">Additional material examined</emphasis>
.
<emphasis id="B938E511E923FFA9FDB4F16EFCD15D5F" bold="true" box="[578,869,548,575]" pageId="31" pageNumber="482">
<collectingCountry id="F35B7993E923FFA9FDB4F16EFD455D5F" box="[578,753,548,575]" name="Costa Rica" pageId="31" pageNumber="482">COSTA RICA</collectingCountry>
,
<collectingRegion id="4988F7E1E923FFA9FCF6F16EFCD15D5F" box="[768,869,548,575]" country="Costa Rica" name="Alajuela" pageId="31" pageNumber="482">Alajuela</collectingRegion>
</emphasis>
,
<collectingCounty id="6292418FE923FFA9FC83F16FFBA65D5F" box="[885,1042,549,575]" pageId="31" pageNumber="482">Rio Cataratas</collectingCounty>
, on road from
<collectingMunicipality id="6B97A379E923FFA9FB36F16FFAF05D5F" box="[1216,1348,549,575]" pageId="31" pageNumber="482">San Ramon</collectingMunicipality>
to
<location id="8E936FD8E923FFA9FA9BF16FFF7E5D03" LSID="urn:lsid:plazi:treatment:03E58815E920FFA9FF61F7E7FAE75BE0:8E936FD8E923FFA9FA9BF16FFF7E5D03" country="Costa Rica" county="Rio Cataratas" municipality="San Ramon" name="Fortuna" pageId="31" pageNumber="482" stateProvince="Alajuela">Fortuna</location>
,
<collectingDate id="EFB6E62BE923FFA9FF20F102FEC75D03" box="[214,371,584,611]" pageId="31" pageNumber="482" value="2002-11-25">
<date id="FFF21FC3E923FFA9FF20F102FEC75D03" box="[214,371,584,611]" pageId="31" pageNumber="482" value="2002-11-25">25 Nov. 2002</date>
</collectingDate>
,
<collectorName id="26B95CD5E923FFA9FE88F103FDAB5D03" box="[382,543,585,611]" pageId="31" pageNumber="482">W.D. Shepard</collectorName>
, WDS-A-1498 (
<specimenCount id="9D4AF28AE923FFA9FD2CF103FD485D03" box="[730,764,585,611]" pageId="31" pageNumber="482" type="male">1♂</specimenCount>
brachypterous,
<collectionCode id="ED5DA1C6E923FFA9FC44F103FC435D03" box="[946,1015,585,611]" pageId="31" pageNumber="482">UMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B24335EE923FFA9FBFCF102FE285DCB" ID-GBIF-Occurrence="2273889312" collectingDate="23. March" collectionCode="USNM" collectorName="W. E. Steiner &amp; J. M. Hill &amp; S. E. Frye" country="Costa Rica" location="Rio Pejibaye" municipality="Pejibaye" pageId="31" pageNumber="482" specimenCount="1" specimenCount-female="1" stateProvince="Cartago">
<collectingRegion id="4988F7E1E923FFA9FBFCF102FBD85D03" box="[1034,1132,584,611]" country="Costa Rica" name="Cartago" pageId="31" pageNumber="482">
<emphasis id="B938E511E923FFA9FBFCF102FBD85D03" bold="true" box="[1034,1132,584,611]" pageId="31" pageNumber="482">Cartago</emphasis>
</collectingRegion>
,
<collectingMunicipality id="6B97A379E923FFA9FB8DF103FB6A5D03" box="[1147,1246,585,611]" pageId="31" pageNumber="482">Pejibaye</collectingMunicipality>
,
<collectingDate id="EFB6E62BE923FFA9FB1CF103FAEF5D03" box="[1258,1371,585,611]" pageId="31" pageNumber="482">
<date id="FFF21FC3E923FFA9FB1CF103FAEF5D03" box="[1258,1371,585,611]" pageId="31" pageNumber="482">23.March</date>
</collectingDate>
.1987, among rocks and leaf packs in sunlit rapids of
<location id="8E936FD8E923FFA9FD40F127FCFE5DE7" LSID="urn:lsid:plazi:treatment:03E58815E920FFA9FF61F7E7FAE75BE0:8E936FD8E923FFA9FD40F127FCFE5DE7" box="[694,842,621,647]" country="Costa Rica" municipality="Pejibaye" name="Rio Pejibaye" pageId="31" pageNumber="482" stateProvince="Cartago">Rio Pejibaye</location>
,
<collectorName id="26B95CD5E923FFA9FCAEF127FC5D5DE7" box="[856,1001,621,647]" pageId="31" pageNumber="482">W.E. Steiner</collectorName>
,
<collectorName id="26B95CD5E923FFA9FC00F127FBE95DE7" box="[1014,1117,621,647]" pageId="31" pageNumber="482">J.M. Hill</collectorName>
&amp;
<collectorName id="26B95CD5E923FFA9FB73F127FB445DE7" box="[1157,1264,621,647]" pageId="31" pageNumber="482">S.E. Frye</collectorName>
collectors (
<specimenCount id="9D4AF28AE923FFA9FA8AF127FA295DE7" box="[1404,1437,621,647]" pageId="31" pageNumber="482" type="female">1♀</specimenCount>
macropterous,
<collectionCode id="ED5DA1C6E923FFA9FEC8F1DBFE205DCB" box="[318,404,657,683]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="482" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B24335EE923FFA9FE5EF1DBFE4B5DAF" ID-GBIF-Occurrence="2273889435" collectingDate="2003-06-22" collectionCode="EMEC" collectorName="Tapanti NP &amp; William D. Shepard" country="Costa Rica" elevation="2" location="Km" pageId="31" pageNumber="482" specimenCount="1" specimenCount-male="1" stateProvince="Cartago">
<collectorName id="26B95CD5E923FFA9FE5EF1DBFD935DCB" box="[424,551,657,683]" pageId="31" pageNumber="482">Tapanti NP</collectorName>
, 5.1
<location id="8E936FD8E923FFA9FDA8F1DBFD3C5DCB" LSID="urn:lsid:plazi:treatment:03E58815E920FFA9FF61F7E7FAE75BE0:8E936FD8E923FFA9FDA8F1DBFD3C5DCB" box="[606,648,657,683]" country="Costa Rica" name="Km" pageId="31" pageNumber="482" stateProvince="Cartago">Km</location>
abv gate,
<collectingDate id="EFB6E62BE923FFA9FD08F1DBFC345DCB" box="[766,896,657,683]" pageId="31" pageNumber="482" value="2003-06-22">
<date id="FFF21FC3E923FFA9FD08F1DBFC345DCB" box="[766,896,657,683]" pageId="31" pageNumber="482" value="2003-06-22">22.VI.2003</date>
</collectingDate>
,
<elevation id="0061DE30E923FFA9FC7AF1DBFC535DCB" box="[908,999,657,683]" metricMagnitude="0" metricUnit="m" metricValue="1.517904" pageId="31" pageNumber="482" unit="ft" value="4.98">
<quantity id="4CB494E6E923FFA9FC7AF1DBFC535DCB" box="[908,999,657,683]" metricMagnitude="0" metricUnit="m" metricValue="1.517904" pageId="31" pageNumber="482" unit="ft" value="4.98">4.980 ft</quantity>
</elevation>
, unnamed stream,
<collectorName id="26B95CD5E923FFA9FB4DF1DBFA285DCB" box="[1211,1436,657,683]" pageId="31" pageNumber="482">William D. Shepard</collectorName>
leg (
<specimenCount id="9D4AF28AE923FFA9FF3DF1FFFF5A5DAF" box="[203,238,693,719]" pageId="31" pageNumber="482" type="male">1♂</specimenCount>
brachypterous,
<collectionCode id="ED5DA1C6E923FFA9FE52F1FFFE435DAF" box="[420,503,693,719]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="31" pageNumber="482" type="University or college">EMEC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B24335EE923FFA9FDFCF1FEFE715D93" ID-GBIF-Occurrence="2273889313" collectingDate="1969-12-26" collectionCode="USNM" collectorName="Polhemus" country="Costa Rica" location="El Santo de Angel" pageId="31" pageNumber="482" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Guanacaste">
<collectingRegion id="4988F7E1E923FFA9FDFCF1FEFD225DAF" box="[522,662,692,719]" country="Costa Rica" name="Guanacaste" pageId="31" pageNumber="482">
<emphasis id="B938E511E923FFA9FDFCF1FEFD225DAF" bold="true" box="[522,662,692,719]" pageId="31" pageNumber="482">Guanacaste</emphasis>
</collectingRegion>
,
<location id="8E936FD8E923FFA9FD50F1FFFCC35DAF" LSID="urn:lsid:plazi:treatment:03E58815E920FFA9FF61F7E7FAE75BE0:8E936FD8E923FFA9FD50F1FFFCC35DAF" box="[678,887,693,719]" country="Costa Rica" name="El Santo de Angel" pageId="31" pageNumber="482" stateProvince="Guanacaste">El Santo de Angel</location>
, CL-1271,
<collectingDate id="EFB6E62BE923FFA9FC0EF1FFFB375DAF" box="[1016,1155,693,719]" pageId="31" pageNumber="482" value="1969-12-26">
<date id="FFF21FC3E923FFA9FC0EF1FFFB375DAF" box="[1016,1155,693,719]" pageId="31" pageNumber="482" value="1969-12-26">26.XII.1969</date>
</collectingDate>
, J.T.
<collectorName id="26B95CD5E923FFA9FB36F1FFFA9A5DAF" box="[1216,1326,693,719]" pageId="31" pageNumber="482">Polhemus</collectorName>
(
<specimenCount id="9D4AF28AE923FFA9FAB6F1FFFAD15DAF" box="[1344,1381,693,719]" pageId="31" pageNumber="482" type="male">3♂</specimenCount>
,
<specimenCount id="9D4AF28AE923FFA9FA84F1FFFA225DAF" box="[1394,1430,693,719]" pageId="31" pageNumber="482" type="female">3♀</specimenCount>
, all brachypterous,
<collectionCode id="ED5DA1C6E923FFA9FE91F193FE095D93" box="[359,445,729,755]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="482" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B24335EE923FFA9FE39F193FE095C77" ID-GBIF-Occurrence="2273889358" collectingDate="2001-06-14" collectionCode="UMC" collectorName="Rincon de la Vieja NP &amp; W. D. Shepard" country="Costa Rica" location="Quebrada Catarata" pageId="31" pageNumber="482" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste">
<collectorName id="26B95CD5E923FFA9FE39F193FD7D5D93" box="[463,713,729,755]" pageId="31" pageNumber="482">Rincon de la Vieja NP</collectorName>
,
<location id="8E936FD8E923FFA9FD22F193FC135D93" LSID="urn:lsid:plazi:treatment:03E58815E920FFA9FF61F7E7FAE75BE0:8E936FD8E923FFA9FD22F193FC135D93" box="[724,935,729,755]" country="Costa Rica" name="Quebrada Catarata" pageId="31" pageNumber="482" stateProvince="Guanacaste">Quebrada Catarata</location>
,
<collectingDate id="EFB6E62BE923FFA9FC44F193FBFF5D93" box="[946,1099,729,755]" pageId="31" pageNumber="482" value="2001-06-14">
<date id="FFF21FC3E923FFA9FC44F193FBFF5D93" box="[946,1099,729,755]" pageId="31" pageNumber="482" value="2001-06-14">14 June 2001</date>
</collectingDate>
,
<collectorName id="26B95CD5E923FFA9FBA3F193FB415D93" box="[1109,1269,729,755]" pageId="31" pageNumber="482">W.D. Shepard</collectorName>
, WDS-A-1387 (
<specimenCount id="9D4AF28AE923FFA9FF57F1B7FF705C77" box="[161,196,765,791]" pageId="31" pageNumber="482" type="male">1♂</specimenCount>
macropterous,
<collectionCode id="ED5DA1C6E923FFA9FE86F1B7FE015C77" box="[368,437,765,791]" pageId="31" pageNumber="482">UMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B24335EE923FFA9FE31F1B6FDFD5C5B" ID-GBIF-Occurrence="2273889319" collectingDate="1995-05-11" collectionCode="BYUC" collectorName="Baumann &amp; Houseman" country="Costa Rica" location="Arroyo La Paz Chiquita" pageId="31" pageNumber="482" specimenCount="8" specimenCount-female="8" stateProvince="Heredia">
<collectingRegion id="4988F7E1E923FFA9FE31F1B6FD935C77" box="[455,551,764,791]" country="Costa Rica" name="Heredia" pageId="31" pageNumber="482">
<emphasis id="B938E511E923FFA9FE31F1B6FD935C77" bold="true" box="[455,551,764,791]" pageId="31" pageNumber="482">Heredia</emphasis>
</collectingRegion>
,
<location id="8E936FD8E923FFA9FDC5F1B7FCF45C77" LSID="urn:lsid:plazi:treatment:03E58815E920FFA9FF61F7E7FAE75BE0:8E936FD8E923FFA9FDC5F1B7FCF45C77" box="[563,832,765,791]" country="Costa Rica" name="Arroyo La Paz Chiquita" pageId="31" pageNumber="482" stateProvince="Heredia">Arroyo La Paz Chiquita</location>
,
<collectingDate id="EFB6E62BE923FFA9FCBDF1B6FC0B5C77" box="[843,959,764,791]" pageId="31" pageNumber="482" value="1995-05-11">
<date id="FFF21FC3E923FFA9FCBDF1B6FC0B5C77" box="[843,959,764,791]" pageId="31" pageNumber="482" value="1995-05-11">11.V.1995</date>
</collectingDate>
, W.
<collectorName id="26B95CD5E923FFA9FC19F1B7FBEE5C77" box="[1007,1114,765,791]" pageId="31" pageNumber="482">Baumann</collectorName>
&amp; R.M.
<collectorName id="26B95CD5E923FFA9FB48F1B7FA825C77" box="[1214,1334,765,791]" pageId="31" pageNumber="482">Houseman</collectorName>
(
<specimenCount id="9D4AF28AE923FFA9FAB0F1B7FADC5C77" box="[1350,1384,765,791]" pageId="31" pageNumber="482" type="female">7♀</specimenCount>
bra- chypterous,
<specimenCount id="9D4AF28AE923FFA9FEE9F06BFEF65C5B" box="[287,322,801,827]" pageId="31" pageNumber="482" type="female">1♀</specimenCount>
macropterous,
<collectionCode id="ED5DA1C6E923FFA9FE18F06BFDF55C5B" box="[494,577,801,827]" pageId="31" pageNumber="482">BYUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B24335EE923FFA9FDA4F06BFC055C3F" ID-GBIF-Occurrence="2273889424" collectingDate="06. June- 14. June" collectionCode="USNM" collectorName="Erwin &amp; Hevel" country="Costa Rica" latitude="1.4" location="Central Expedition" longLatPrecision="78" longitude="-1.7" municipality="Monteverde area" pageId="31" pageNumber="482" specimenCount="1" specimenCount-female="1" stateProvince="Puntarenas">
<collectingRegion id="4988F7E1E923FFA9FDA4F06BFD6E5C5B" box="[594,730,801,827]" country="Costa Rica" name="Puntarenas" pageId="31" pageNumber="482">
<emphasis id="B938E511E923FFA9FDA4F06BFD6E5C5B" bold="true" box="[594,730,801,827]" pageId="31" pageNumber="482">Puntarenas</emphasis>
</collectingRegion>
,
<collectingMunicipality id="6B97A379E923FFA9FD1EF06BFC135C5B" box="[744,935,801,827]" pageId="31" pageNumber="482">Monteverde area</collectingMunicipality>
,
<geoCoordinate id="EE785FC4E923FFA9FC47F06BFC445C5B" box="[945,1008,801,827]" degrees="1.400" direction="north" orientation="latitude" pageId="31" pageNumber="482" precision="55" value="1.4">1.400</geoCoordinate>
<geoCoordinate id="EE785FC4E923FFA9FC06F06BFB8A5C5B" box="[1008,1086,801,827]" degrees="1.700" direction="west" orientation="longitude" pageId="31" pageNumber="482" precision="55" value="-1.7">1.700</geoCoordinate>
meters,
<collectingDate id="EFB6E62BE923FFA9FB6AF06BFAEF5C5B" box="[1180,1371,801,827]" pageId="31" pageNumber="482">
<date id="FFF21FC3E923FFA9FB6AF06BFAEF5C5B" box="[1180,1371,801,827]" pageId="31" pageNumber="482">06.June14.June</date>
</collectingDate>
.1973,
<collectorName id="26B95CD5E923FFA9FF61F00FFF6F5C3F" box="[151,219,837,863]" pageId="31" pageNumber="482">Erwin</collectorName>
&amp;
<collectorName id="26B95CD5E923FFA9FEF6F00FFEF25C3F" box="[256,326,837,863]" pageId="31" pageNumber="482">Hevel</collectorName>
,
<location id="8E936FD8E923FFA9FEA7F00FFD9C5C3F" LSID="urn:lsid:plazi:treatment:03E58815E920FFA9FF61F7E7FAE75BE0:8E936FD8E923FFA9FEA7F00FFD9C5C3F" box="[337,552,837,863]" country="Costa Rica" latitude="1.4" longLatPrecision="78" longitude="-1.7" municipality="Monteverde area" name="Central Expedition" pageId="31" pageNumber="482" stateProvince="Puntarenas">Central Expedition</location>
, 1973 (
<specimenCount id="9D4AF28AE923FFA9FD88F00FFD2B5C3F" box="[638,671,837,863]" pageId="31" pageNumber="482" type="female">1♀</specimenCount>
brachypterous,
<collectionCode id="ED5DA1C6E923FFA9FCA5F00FFC1D5C3F" box="[851,937,837,863]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="482" type="Museum">USNM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B24335EE923FFA9FC4DF00FFAE35CE3" ID-GBIF-Occurrence="2273889322" collectingDate="2003-06-20" collectionCode="EMEC" collectorName="Rio Bella Vista" country="Costa Rica" elevation="1" location="Alturas Biol. Station" pageId="31" pageNumber="482" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Puntarenas">
<location id="8E936FD8E923FFA9FC4DF00FFB115C3F" LSID="urn:lsid:plazi:treatment:03E58815E920FFA9FF61F7E7FAE75BE0:8E936FD8E923FFA9FC4DF00FFB115C3F" box="[955,1189,837,863]" country="Costa Rica" name="Alturas Biol. Station" pageId="31" pageNumber="482" stateProvince="Puntarenas">Alturas Biol. Station</location>
,
<collectingDate id="EFB6E62BE923FFA9FB46F00FFA875C3F" box="[1200,1331,837,863]" pageId="31" pageNumber="482" value="2003-06-20">
<date id="FFF21FC3E923FFA9FB46F00FFA875C3F" box="[1200,1331,837,863]" pageId="31" pageNumber="482" value="2003-06-20">20.VI.2003</date>
</collectingDate>
,
<elevation id="0061DE30E923FFA9FAC8F00FFA2C5C3F" box="[1342,1432,837,863]" metricMagnitude="0" metricUnit="m" metricValue="1.328928" pageId="31" pageNumber="482" unit="ft" value="4.36">
<quantity id="4CB494E6E923FFA9FAC8F00FFA2C5C3F" box="[1342,1432,837,863]" metricMagnitude="0" metricUnit="m" metricValue="1.328928" pageId="31" pageNumber="482" unit="ft" value="4.36">4.360 ft</quantity>
</elevation>
,
<collectorName id="26B95CD5E923FFA9FF61F023FEFC5CE3" box="[151,328,873,899]" hasSafeAffix="true" pageId="31" pageNumber="482">Rio Bella Vista</collectorName>
, William D. Shepard leg (
<specimenCount id="9D4AF28AE923FFA9FD83F023FD2C5CE3" box="[629,664,873,899]" pageId="31" pageNumber="482" type="male">1♂</specimenCount>
brachypterous,
<specimenCount id="9D4AF28AE923FFA9FCB9F023FCC55CE3" box="[847,881,873,899]" pageId="31" pageNumber="482" type="female">2♀</specimenCount>
brachypterous,
<specimenCount id="9D4AF28AE923FFA9FBDFF023FBFF5CE3" box="[1065,1099,873,899]" pageId="31" pageNumber="482" type="female">3♀</specimenCount>
macropterous,
<collectionCode id="ED5DA1C6E923FFA9FB0BF023FAE45CE3" box="[1277,1360,873,899]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="31" pageNumber="482" type="University or college">EMEC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B24335EE923FFA9FA92F023FE9C5CAB" ID-GBIF-Occurrence="2273889316" collectingDate="2000-01-24" collectionCode="EMEC" collectorName="C. B. Barr" country="Costa Rica" latitude="10.309167" location="Estac. Biol. Monteverde" longLatPrecision="13" longitude="-84.802666" municipality="Moquina" pageId="31" pageNumber="482" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Puntarenas">
Que- brada
<collectingMunicipality id="6B97A379E923FFA9FF2BF0C7FEF15CC7" box="[221,325,909,935]" pageId="31" pageNumber="482">Móquina</collectingMunicipality>
,
<location id="8E936FD8E923FFA9FEA7F0C7FDD25CC7" LSID="urn:lsid:plazi:treatment:03E58815E920FFA9FF61F7E7FAE75BE0:8E936FD8E923FFA9FEA7F0C7FDD25CC7" box="[337,614,909,935]" country="Costa Rica" latitude="10.309167" longLatPrecision="13" longitude="-84.802666" municipality="Moquina" name="Estac. Biol. Monteverde" pageId="31" pageNumber="482" stateProvince="Puntarenas">Estac. Biol. Monteverde</location>
,
<geoCoordinate id="EE785FC4E923FFA9FD84F0C6FD405CC7" box="[626,756,908,935]" degrees="10" direction="north" minutes="18.55" orientation="latitude" pageId="31" pageNumber="482" precision="9" value="10.309167">10°18.55N</geoCoordinate>
/
<geoCoordinate id="EE785FC4E923FFA9FCF3F0C7FC245CC7" box="[773,912,909,935]" degrees="84" direction="west" minutes="48.16" orientation="longitude" pageId="31" pageNumber="482" precision="9" value="-84.802666">84°48.16W</geoCoordinate>
,
<collectingDate id="EFB6E62BE923FFA9FC6AF0C7FBBE5CC7" box="[924,1034,909,935]" pageId="31" pageNumber="482" value="2000-01-24">
<date id="FFF21FC3E923FFA9FC6AF0C7FBBE5CC7" box="[924,1034,909,935]" pageId="31" pageNumber="482" value="2000-01-24">24.I.2000</date>
</collectingDate>
,
<collectorName id="26B95CD5E923FFA9FBE0F0C7FB375CC7" box="[1046,1155,909,935]" pageId="31" pageNumber="482">C.B. Barr</collectorName>
(
<specimenCount id="9D4AF28AE923FFA9FB63F0C7FB0D5CC7" box="[1173,1209,909,935]" pageId="31" pageNumber="482" type="male">1♂</specimenCount>
,
<specimenCount id="9D4AF28AE923FFA9FB30F0C7FB5E5CC7" box="[1222,1258,909,935]" pageId="31" pageNumber="482" type="female">3♀</specimenCount>
, all brachypter- ous,
<collectionCode id="ED5DA1C6E923FFA9FF3BF0FBFE945CAB" box="[205,288,945,971]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="31" pageNumber="482" type="University or college">EMEC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B24335EE923FFA9FEC2F0FBFAB85C8F" ID-GBIF-Occurrence="2273889486" collectingDate="2003-06-20" collectionCode="EMEC" collectorName="C. B. Barr" country="Costa Rica" elevation="1" latitude="12.4575" location="Las Alturas Biological Station" longLatPrecision="13" longitude="-75.592834" municipality="Rio Bella Vista" pageId="31" pageNumber="482" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Puntarenas">
<collectingMunicipality id="6B97A379E923FFA9FEC2F0FBFE505CAB" box="[308,484,945,971]" pageId="31" pageNumber="482">Rio Bella Vista</collectingMunicipality>
,
<location id="8E936FD8E923FFA9FE07F0FBFCF25CAB" LSID="urn:lsid:plazi:treatment:03E58815E920FFA9FF61F7E7FAE75BE0:8E936FD8E923FFA9FE07F0FBFCF25CAB" box="[497,838,945,971]" country="Costa Rica" latitude="12.4575" longLatPrecision="13" longitude="-75.592834" municipality="Rio Bella Vista" name="Las Alturas Biological Station" pageId="31" pageNumber="482" stateProvince="Puntarenas">Las Alturas Biological Station</location>
<locationDeviation id="9A412021E923FFA9FCB9F0FBFC6B5CAB" box="[847,991,945,971]" location="Alturas" pageId="31" pageNumber="482">N of Alturas</locationDeviation>
,
<collectingDate id="EFB6E62BE923FFA9FC1DF0FBFBDA5CAB" box="[1003,1134,945,971]" pageId="31" pageNumber="482" value="2003-06-20">
<date id="FFF21FC3E923FFA9FC1DF0FBFBDA5CAB" box="[1003,1134,945,971]" pageId="31" pageNumber="482" value="2003-06-20">20.VI.2003</date>
</collectingDate>
,
<geoCoordinate id="EE785FC4E923FFA9FB8CF0FAFB485CAB" box="[1146,1276,944,971]" degrees="12" direction="north" minutes="27.45" orientation="latitude" pageId="31" pageNumber="482" precision="9" value="12.4575">12°27.45N</geoCoordinate>
/
<geoCoordinate id="EE785FC4E923FFA9FAF8F0FAFA2C5CAB" box="[1294,1432,944,971]" degrees="75" direction="west" minutes="35.57" orientation="longitude" pageId="31" pageNumber="482" precision="9" value="-75.592834">75°35.57W</geoCoordinate>
,
<elevation id="0061DE30E923FFA9FF61F09FFF475C8F" box="[151,243,981,1007]" metricMagnitude="0" metricUnit="m" metricValue="1.328928" pageId="31" pageNumber="482" unit="ft" value="4.36">
<quantity id="4CB494E6E923FFA9FF61F09FFF475C8F" box="[151,243,981,1007]" metricMagnitude="0" metricUnit="m" metricValue="1.328928" pageId="31" pageNumber="482" unit="ft" value="4.36">4.360 ft</quantity>
</elevation>
elevation,
<collectorName id="26B95CD5E923FFA9FE81F09FFE5C5C8F" box="[375,488,981,1007]" pageId="31" pageNumber="482">C.B. Barr</collectorName>
coll (
<specimenCount id="9D4AF28AE923FFA9FDC7F09FFDE35C8F" box="[561,599,981,1007]" pageId="31" pageNumber="482" type="male">2♂</specimenCount>
,
<specimenCount id="9D4AF28AE923FFA9FD91F09FFD3F5C8F" box="[615,651,981,1007]" pageId="31" pageNumber="482" type="female">2♀</specimenCount>
, all brachypterous,
<specimenCount id="9D4AF28AE923FFA9FC83F09FFC2D5C8F" box="[885,921,981,1007]" pageId="31" pageNumber="482" type="male">1♂</specimenCount>
,
<specimenCount id="9D4AF28AE923FFA9FC5CF09FFC7A5C8F" box="[938,974,981,1007]" pageId="31" pageNumber="482" type="female">2♀</specimenCount>
, all macropterous,
<collectionCode id="ED5DA1C6E923FFA9FB47F09FFAB05C8F" box="[1201,1284,981,1007]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="31" pageNumber="482" type="University or college">EMEC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B24335EE923FFA9FAEDF09EFBC95B74" ID-GBIF-Occurrence="2273889315" collectingDate="2003-06-20" collectionCode="USNM" collectorName="C. B. Barr" country="Panama" elevation="1" latitude="12.4575" location="Las Alturas Biological Station" longLatPrecision="13" longitude="-75.592834" municipality="Rio Bella Vista" pageId="31" pageNumber="482" specimenCount="17" specimenCount-female="11" specimenCount-male="6" stateProvince="Chiriqui">
<emphasis id="B938E511E923FFA9FAEDF09EFF4A5B73" bold="true" pageId="31" pageNumber="482">
<collectingCountry id="F35B7993E923FFA9FAEDF09EFA235C8F" box="[1307,1431,980,1007]" name="Panama" pageId="31" pageNumber="482">PANAMA</collectingCountry>
,
<collectingRegion id="4988F7E1E923FFA9FF61F0B2FF4A5B73" box="[151,254,1016,1043]" country="Panama" name="Chiriqui" pageId="31" pageNumber="482">Chiriquí</collectingRegion>
</emphasis>
, same data as holotype (
<specimenCount id="9D4AF28AE923FFA9FDEFF0B3FD8A5B73" box="[537,574,1017,1043]" pageId="31" pageNumber="482" type="male">3♂</specimenCount>
,
<specimenCount id="9D4AF28AE923FFA9FDBDF0B3FDC45B73" box="[587,624,1017,1043]" pageId="31" pageNumber="482" type="female">6♀</specimenCount>
,
<collectionCode id="ED5DA1C6E923FFA9FD88F0B3FD675B73" box="[638,723,1017,1043]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="482" type="Museum">USNM</collectionCode>
;
<specimenCount id="9D4AF28AE923FFA9FD29F0B3FCB05B73" box="[735,772,1017,1043]" pageId="31" pageNumber="482" type="male">3♂</specimenCount>
,
<specimenCount id="9D4AF28AE923FFA9FCE7F0B2FC815B73" box="[785,821,1016,1043]" pageId="31" pageNumber="482" type="female">5♀</specimenCount>
, EMEC—all brachypterous)
</materialsCitation>
;
<materialsCitation id="3B24335EE923FFA9FB7FF0B3FA8A5B58" ID-GBIF-Occurrence="2273889396" collectingDate="1993-01" collectionCode="USNM, UMC" collectorName="J. T. Polhemus" country="Panama" location="Jurutungo" municipality="Rio Candela" pageId="31" pageNumber="482" specimenCount="10" specimenCount-female="5" specimenCount-male="5" stateProvince="Chiriqui">
<collectingMunicipality id="6B97A379E923FFA9FB7FF0B3FAAD5B73" box="[1161,1305,1017,1043]" pageId="31" pageNumber="482">Rio Candela</collectingMunicipality>
,
<location id="8E936FD8E923FFA9FAD3F0B3FA2C5B73" LSID="urn:lsid:plazi:treatment:03E58815E920FFA9FF61F7E7FAE75BE0:8E936FD8E923FFA9FAD3F0B3FA2C5B73" box="[1317,1432,1017,1043]" country="Panama" municipality="Rio Candela" name="Jurutungo" pageId="31" pageNumber="482" stateProvince="Chiriqui">Jurutungo</location>
,
<collectingDate id="EFB6E62BE923FFA9FF61F757FE845B57" box="[151,304,1053,1079]" pageId="31" pageNumber="482" value="1993-01">
<date id="FFF21FC3E923FFA9FF61F757FE845B57" box="[151,304,1053,1079]" pageId="31" pageNumber="482" value="1993-01">January 1993</date>
</collectingDate>
, CL2806, coll.
<collectorName id="26B95CD5E923FFA9FE2DF757FDC35B57" box="[475,631,1053,1079]" pageId="31" pageNumber="482">J.T. Polhemus</collectorName>
(
<specimenCount id="9D4AF28AE923FFA9FD71F757FD195B57" box="[647,685,1053,1079]" pageId="31" pageNumber="482" type="male">4♂</specimenCount>
,
<specimenCount id="9D4AF28AE923FFA9FD4EF757FD695B57" box="[696,733,1053,1079]" pageId="31" pageNumber="482" type="female">4♀</specimenCount>
,
<collectionCode id="ED5DA1C6E923FFA9FD1EF757FC895B57" box="[744,829,1053,1079]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="482" type="Museum">USNM</collectionCode>
;
<specimenCount id="9D4AF28AE923FFA9FCBEF757FCD85B57" box="[840,876,1053,1079]" pageId="31" pageNumber="482" type="male">1♂</specimenCount>
,
<collectionCode id="ED5DA1C6E923FFA9FC8EF757FC0F5B57" box="[888,955,1053,1079]" pageId="31" pageNumber="482">UMC</collectionCode>
;
<specimenCount id="9D4AF28AE923FFA9FC30F757FC5D5B57" box="[966,1001,1053,1079]" pageId="31" pageNumber="482" type="female">1♀</specimenCount>
, MZUSP—all brachypterous)
</materialsCitation>
;
<materialsCitation id="3B24335EE923FFA9FABEF757FB785B3B" ID-GBIF-Occurrence="2273889329" collectingDate="01. March" collectionCode="USNM" collectorName="Fred J. Foster" country="Panama" location="Volcan" pageId="31" pageNumber="482" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Chiriqui">
<location id="8E936FD8E923FFA9FABEF757FA2C5B57" LSID="urn:lsid:plazi:treatment:03E58815E920FFA9FF61F7E7FAE75BE0:8E936FD8E923FFA9FABEF757FA2C5B57" box="[1352,1432,1053,1079]" country="Panama" name="Volcan" pageId="31" pageNumber="482" stateProvince="Chiriqui">Volcán</location>
,
<collectingDate id="EFB6E62BE923FFA9FF61F70BFEB35B3B" box="[151,263,1089,1115]" pageId="31" pageNumber="482">
<date id="FFF21FC3E923FFA9FF61F70BFEB35B3B" box="[151,263,1089,1115]" pageId="31" pageNumber="482">01.March</date>
</collectingDate>
.1924,
<collectorName id="26B95CD5E923FFA9FEB9F70BFE535B3B" box="[335,487,1089,1115]" pageId="31" pageNumber="482">Fred J. Foster</collectorName>
(
<specimenCount id="9D4AF28AE923FFA9FE01F70BFDAD5B3B" box="[503,537,1089,1115]" pageId="31" pageNumber="482" type="male">1♂</specimenCount>
brachypterous,
<specimenCount id="9D4AF28AE923FFA9FD3DF70BFD5A5B3B" box="[715,750,1089,1115]" pageId="31" pageNumber="482" type="female">2♀</specimenCount>
brachypterous,
<specimenCount id="9D4AF28AE923FFA9FC56F70BFC775B3B" box="[928,963,1089,1115]" pageId="31" pageNumber="482" type="female">1♀</specimenCount>
macropterous,
<collectionCode id="ED5DA1C6E923FFA9FB98F70BFB705B3B" box="[1134,1220,1089,1115]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="482" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B24335EE923FFA9FB20F70BFAFB5BE0" ID-GBIF-Occurrence="2273889450" collectingDate="1993-01-15" collectionCode="USNM" collectorName="J. T. Polhemus" country="Panama" location="Santa Fe" pageId="31" pageNumber="482" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Veraguas">
<collectingRegion id="4988F7E1E923FFA9FB20F70BFAF15B3B" box="[1238,1349,1089,1115]" country="Panama" name="Veraguas" pageId="31" pageNumber="482">
<emphasis id="B938E511E923FFA9FB20F70BFAF15B3B" bold="true" box="[1238,1349,1089,1115]" pageId="31" pageNumber="482">Veraguas</emphasis>
</collectingRegion>
, stream near
<location id="8E936FD8E923FFA9FF38F72FFE875B1F" LSID="urn:lsid:plazi:treatment:03E58815E920FFA9FF61F7E7FAE75BE0:8E936FD8E923FFA9FF38F72FFE875B1F" box="[206,307,1125,1151]" country="Panama" name="Santa Fe" pageId="31" pageNumber="482" stateProvince="Veraguas">Santa Fe</location>
,
<collectingDate id="EFB6E62BE923FFA9FEC8F72EFE1F5B1F" box="[318,427,1124,1151]" pageId="31" pageNumber="482" value="1993-01-15">
<date id="FFF21FC3E923FFA9FEC8F72EFE1F5B1F" box="[318,427,1124,1151]" pageId="31" pageNumber="482" value="1993-01-15">15.I.1993</date>
</collectingDate>
, CL-2832,
<collectorName id="26B95CD5E923FFA9FDDCF72FFD735B1F" box="[554,711,1125,1151]" pageId="31" pageNumber="482">J.T. Polhemus</collectorName>
col (
<specimenCount id="9D4AF28AE923FFA9FCF7F72FFC925B1F" box="[769,806,1125,1151]" pageId="31" pageNumber="482" type="male">2♂</specimenCount>
,
<specimenCount id="9D4AF28AE923FFA9FCC4F72FFCE25B1F" box="[818,854,1125,1151]" pageId="31" pageNumber="482" type="female">2♀</specimenCount>
,
<collectionCode id="ED5DA1C6E923FFA9FC95F72FFC0C5B1F" box="[867,952,1125,1151]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="482" type="Museum">USNM</collectionCode>
;
<specimenCount id="9D4AF28AE923FFA9FC35F72FFC535B1F" box="[963,999,1125,1151]" pageId="31" pageNumber="482" type="male">1♂</specimenCount>
,
<specimenCount id="9D4AF28AE923FFA9FC02F72FFBA35B1F" box="[1012,1047,1125,1151]" pageId="31" pageNumber="482" type="female">1♀</specimenCount>
, UMC—all brachypterous)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>