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<document id="D775B4606D3D5DCE6565212AFEA5F9F5" ID-CLB-Dataset="8728" ID-DOI="10.11646/zootaxa.4821.3.2" ID-GBIF-Dataset="a9792973-949f-4327-82ad-8a38612624b4" ID-ISSN="1175-5326" ID-Zenodo-Dep="4400933" ID-ZooBank="E831BB59-C95B-4C30-B827-131723233045" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609331543331" checkinUser="plazi" docAuthor="Park, Kyu-Tek, Bae, Yang-Seop, Zhao, Shengnan &amp; Li, Houhun" docDate="2020" docId="03E3878EFFE7FFB0FF7FFB0CFCCAC5E8" docLanguage="en" docName="zootaxa.4821.3.2.pdf" docOrigin="Zootaxa 4821 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dichomeris sclerospinula Park &amp; Bae &amp; Zhao &amp; Li 2020, sp. nov." docType="treatment" docUuid="3382249F-F160-42E0-826D-6B9BF384C03E" docUuidSource="ZooBank" docVersion="9" lastPageNumber="444" masterDocId="FFDAFFF6FFEFFFB9FFE8FF8AFFEAC41D" masterDocTitle="Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam" masterLastPageNumber="461" masterPageNumber="435" pageNumber="443" updateTime="1698849544545" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="B437CC65BBA3949503443CBBE1008806" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="47ED1F5597BE501640214D33B90F6054">
<mods:title id="8CF88060BD3F1B773E41BEAC4B556413">Thirteen new species of genera Dichomeris Hübner, 1818 and Helcystogramma Zeller, 1877 (Lepidoptera, Gelechiidae, Dichomeridinae) with twenty-one newly recorded species from Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="7BCA2D9561F88A902712EAB077B9B46C" type="personal">
<mods:role id="CEB01B5912853D59B7B6482A6112C828">
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<mods:namePart id="4DDA9F479BE477518C50F8E5F03195A4">Park, Kyu-Tek</mods:namePart>
<mods:affiliation id="D690D680B958F828A3EDDFA21BD48F31">BioResource and Environmental Center, Division of Life Sciences, Incheon National University, Incheon, 22012 Korea</mods:affiliation>
</mods:name>
<mods:name id="D6805B2212B65B1684D4AA549783A263" type="personal">
<mods:role id="25783A3E896CBEFAF41458A36C89BA58">
<mods:roleTerm id="573FA877EDF6EFAF093DBC3A8E05A451">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB13D2CE17C175112C66A772A9ED558A">Bae, Yang-Seop</mods:namePart>
<mods:nameIdentifier id="1A876D1D2252ECF7514B438B26C85FFB" type="ORCID">0000-0001-7356-5633</mods:nameIdentifier>
<mods:affiliation id="BFDBF67CCFD8172F7E0C88B7F5CD1C44">BioResource and Environmental Center, Division of Life Sciences, Incheon National University, Incheon, 22012 Korea &amp; https: // orcid. org / 0000 - 0001 - 7356 - 5633</mods:affiliation>
</mods:name>
<mods:name id="0304B2093A800AD31786FCB55E6CE32A" type="personal">
<mods:role id="1C04E0E20F1AE47D00740ECCCD6ECD5D">
<mods:roleTerm id="0044E04C84F8DD3B53B3A24CFF32D47B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="80156D4C2818224C65649AFF91C0DC7D">Zhao, Shengnan</mods:namePart>
<mods:affiliation id="3089753253F7F5FCC1C4891D289D2101">College of Life Sciences, Nankai University, Tianjin 300071, P. R. China</mods:affiliation>
</mods:name>
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<mods:roleTerm id="B2528D16A99AEB0F4FB14F6D03917735">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC36700C03D8F1F620D846D4CCA00C1D">Li, Houhun</mods:namePart>
<mods:affiliation id="128DF47F87C80722503B10B09A7E58EA">College of Life Sciences, Nankai University, Tianjin 300071, P. R. China</mods:affiliation>
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<mods:typeOfResource id="4B1822DF60EB22B16F2D667C2B6997F6">text</mods:typeOfResource>
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<mods:title id="BB87F070681637B4D23361DD617B8126">Zootaxa</mods:title>
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<mods:part id="569098ADD7D9FD64FAEA27220CB01332">
<mods:date id="B8F3F04095015985310AFEBC762FE7BB">2020</mods:date>
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<mods:number id="8C4195F907F4A21EA6109720B8B076C2">2020-08-03</mods:number>
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<mods:number id="64FC39019EB2E1B0A48BE5ABB2E74E16">4821</mods:number>
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<mods:classification id="0BC85DD6CAB51F69EF8C17FB0B8A3A7E">journal article</mods:classification>
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<mods:identifier id="31E4A0C7E3418F5CBB031BF3A7EC654A" type="DOI">10.11646/zootaxa.4821.3.2</mods:identifier>
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<treatment id="03E3878EFFE7FFB0FF7FFB0CFCCAC5E8" ID-DOI="http://doi.org/10.5281/zenodo.4400972" ID-GBIF-Taxon="177013228" ID-Zenodo-Dep="4400972" LSID="urn:lsid:zoobank.org:act:3382249F-F160-42E0-826D-6B9BF384C03E" httpUri="http://treatment.plazi.org/id/03E3878EFFE7FFB0FF7FFB0CFCCAC5E8" lastPageId="9" lastPageNumber="444" pageId="8" pageNumber="443">
<subSubSection id="C3506513FFE7FFB1FF7FFB0CFC5EC0D8" pageId="8" pageNumber="443" type="nomenclature">
<paragraph id="8BF53698FFE7FFB1FF7FFB0CFD2AC0BC" blockId="8.[151,948,1158,1257]" box="[151,704,1158,1185]" pageId="8" pageNumber="443">
<heading id="D0BD81F4FFE7FFB1FF7FFB0CFD2AC0BC" bold="true" box="[151,704,1158,1185]" fontSize="11" level="1" pageId="8" pageNumber="443" reason="1">
<emphasis id="B93EEA8AFFE7FFB1FF7FFB0CFD2AC0BC" bold="true" box="[151,704,1158,1185]" pageId="8" pageNumber="443">
<taxonomicName id="4C4A4D1BFFE7FFB1FF7FFB0CFDBFC0BD" ID-CoL="98Z8J" authority="Park &amp; Li" authorityName="Park &amp; Bae &amp; Zhao &amp; Li" authorityYear="2020" box="[151,597,1158,1185]" class="Insecta" family="Gelechiidae" genus="Dichomeris" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="443" phylum="Arthropoda" rank="species" species="sclerospinula" status="sp. nov.">
<emphasis id="B93EEA8AFFE7FFB1FF7FFB0CFE26C0BC" bold="true" box="[151,460,1158,1185]" italics="true" pageId="8" pageNumber="443">Dichomeris sclerospinula</emphasis>
Park &amp; Li
</taxonomicName>
,
<taxonomicNameLabel id="A20D57F1FFE7FFB1FD8BFB0DFD2AC0BC" box="[611,704,1159,1185]" pageId="8" pageNumber="443" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF53698FFE7FFB1FF7FFB21FC5EC0D8" blockId="8.[151,948,1158,1257]" box="[151,948,1195,1221]" pageId="8" pageNumber="443">
<uri id="FFDB3A9AFFE7FFB1FF7FFB21FC5EC0D8" box="[151,948,1195,1221]" pageId="8" pageNumber="443">
urn:lsid:zoobank.org:act:
<uuid id="FFEC0C4DFFE7FFB1FE40FB21FC5EC0D8" box="[424,948,1195,1221]" pageId="8" pageNumber="443">3382249F-F160-42E0-826D-6B9BF384C03E</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3506513FFE7FFB1FF7FFB45FECDC0F4" box="[151,295,1230,1257]" pageId="8" pageNumber="443" type="description">
<paragraph id="8BF53698FFE7FFB1FF7FFB45FECDC0F4" blockId="8.[151,948,1158,1257]" box="[151,295,1230,1257]" pageId="8" pageNumber="443">
(
<figureCitation id="13712A1DFFE7FFB1FF77FB45FF1CC0F4" box="[159,246,1230,1257]" captionStart="FIGURES 25–30" captionStartId="14.[152,270,1862,1886]" captionTargetBox="[158,1426,190,1823]" captionTargetId="figure-32@14.[150,1438,181,1837]" captionTargetPageId="14" captionText="FIGURES 25–30. Adults. 25. Dichomeris sclerospinula sp. nov., holotype, male; 26. D. sexafurca Li &amp; Zheng, male; 27. D. spinosella sp. nov., holotype, male; 28, D. unilobata sp. nov., holotype, male; 29. D. splendiptera Li &amp; Park, male; 30. D. summata Meyrick, female;" figureDoi="http://doi.org/10.5281/zenodo.4400943" httpUri="https://zenodo.org/record/4400943/files/figure.png" pageId="8" pageNumber="443">Figs 25</figureCitation>
,
<figureCitation id="13712A1DFFE7FFB1FEEAFB45FEF5C0F4" box="[258,287,1231,1257]" captionStart="FIGURES 41–46" captionStartId="21.[152,270,1897,1921]" captionTargetBox="[247,1334,194,1857]" captionTargetId="figure-19@21.[238,1350,181,1873]" captionTargetPageId="21" captionText="FIGURES 41–46. Male genitalia. 41. Dichomeris ochriella sp. nov., holotype, gen. slide no. CIS-6277; 42. D. myriaodontosa sp. nov., holotype, gen. slide no. CIS-6675; 43. D. quadratella sp. nov., paratype, gen. slide no. CIS-6285; 44. D. sclerospinula sp. nov., paratype, gen. slide no. NKLEP059; 45. D. spinosella sp. nov., holotype, gen. slide no. NKLEP003; 46. D. unilobata sp. nov., holotype, gen. slide no. CIS-6320. Scales = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4400951" httpUri="https://zenodo.org/record/4400951/files/figure.png" pageId="8" pageNumber="443">44</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3506513FFE7FFB1FF7FFA9DFC98C164" pageId="8" pageNumber="443" type="materials_examined">
<paragraph id="8BF53698FFE7FFB1FF7FFA9DFC98C164" blockId="8.[151,1437,1302,2013]" pageId="8" pageNumber="443">
<emphasis id="B93EEA8AFFE7FFB1FF7FFA9DFEAFC12D" bold="true" box="[151,325,1302,1329]" pageId="8" pageNumber="443">Type material.</emphasis>
<materialsCitation id="3B223CC5FFE7FFB1FEA4FA9DFDECC148" ID-GBIF-Occurrence="3022744306" collectingDate="2006-04-26" collectorName="Park &amp; Kim &amp; Kang" country="Vietnam" elevation="950" location="Tam Dao Nat. Park" pageId="8" pageNumber="443" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc" typeStatus="holotype">
<typeStatus id="54F1883AFFE7FFB1FEA4FA9DFE5EC12C" box="[332,436,1303,1329]" pageId="8" pageNumber="443" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4CFD11FFE7FFB1FE53FA9DFE38C12C" box="[443,466,1303,1329]" pageId="8" pageNumber="443" type="male">♂</specimenCount>
,
<emphasis id="B93EEA8AFFE7FFB1FE36FA9CFD84C12D" bold="true" box="[478,622,1302,1328]" pageId="8" pageNumber="443">
<collectingCountry id="F35D7608FFE7FFB1FE36FA9CFD80C12D" box="[478,618,1302,1328]" name="Vietnam" pageId="8" pageNumber="443">VIETNAM</collectingCountry>
:
</emphasis>
<collectingRegion id="498EF87AFFE7FFB1FD9DFA9DFCDAC12C" box="[629,816,1303,1329]" country="Vietnam" name="Vinh Phuc" pageId="8" pageNumber="443">Vinh Phuc Prov.</collectingRegion>
:
<location id="8E956043FFE7FFB1FCD1FA9DFBFEC12C" LSID="urn:lsid:plazi:treatment:03E3878EFFE7FFB0FF7FFB0CFCCAC5E8:8E956043FFE7FFB1FCD1FA9DFBFEC12C" box="[825,1044,1303,1329]" country="Vietnam" name="Tam Dao Nat. Park" pageId="8" pageNumber="443" stateProvince="Vinh Phuc">Tam Dao Nat. Park</location>
,
<quantity id="4CB29B7DFFE7FFB1FBF7FA9CFB82C12C" box="[1055,1128,1302,1329]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="8" pageNumber="443" unit="m" value="950.0">
<elevation id="0067D1ABFFE7FFB1FBF7FA9CFB82C12C" box="[1055,1128,1302,1329]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="8" pageNumber="443" unit="m" value="950.0">950 m</elevation>
</quantity>
,
<date id="FFF41058FFE7FFB1FB9BFA9DFB04C12C" box="[1139,1262,1303,1329]" pageId="8" pageNumber="443" value="2006-04-26">
<collectingDate id="EFB0E9B0FFE7FFB1FB9BFA9DFB04C12C" box="[1139,1262,1303,1329]" pageId="8" pageNumber="443" value="2006-04-26">26 iv 2006</collectingDate>
</date>
, leg.
<collectorName id="26BF534EFFE7FFB1FAC1FA9DFAB5C12C" box="[1321,1375,1303,1329]" pageId="8" pageNumber="443">Park</collectorName>
,
<collectorName id="26BF534EFFE7FFB1FA82FA9DFA76C12C" box="[1386,1436,1303,1329]" pageId="8" pageNumber="443">Kim</collectorName>
&amp;
<collectorName id="26BF534EFFE7FFB1FF5CFAB1FF19C148" box="[180,243,1339,1365]" pageId="8" pageNumber="443">Kang</collectorName>
, gen. slide no. nklep023
</materialsCitation>
.
<materialsCitation id="3B223CC5FFE7FFB1FDF8FAB1FC85C164" ID-GBIF-Occurrence="3022744366" collectingDate="2005-05-04" collectingDateMax="2005-05-05" collectingDateMin="2005-05-04" collectionCode="NIBR" collectorName="KT Park &amp; Kim" country="Vietnam" elevation="1000" location="Tam Dao Nat. Park" pageId="8" pageNumber="443" specimenCount="6" specimenCount-male="6" stateProvince="Vinh Phuc" typeStatus="paratype">
<typeStatus id="54F1883AFFE7FFB1FDF8FAB1FD6BC148" box="[528,641,1339,1365]" pageId="8" pageNumber="443" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D4CFD11FFE7FFB1FD64FAB1FD5BC148" box="[652,689,1339,1365]" pageId="8" pageNumber="443" type="male">6♂</specimenCount>
, same locality as holotype,
<quantity id="4CB29B7DFFE7FFB1FC0EFAB1FBD7C148" box="[998,1085,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="443" unit="m" value="1000.0">
<elevation id="0067D1ABFFE7FFB1FC0EFAB1FBD7C148" box="[998,1085,1339,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="443" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="FFF41058FFE7FFB1FBA1FAB0FB0CC148" box="[1097,1254,1338,1365]" pageId="8" pageNumber="443" value="2005-05-04" valueMax="2005-05-05" valueMin="2005-05-04">
<collectingDate id="EFB0E9B0FFE7FFB1FBA1FAB0FB0CC148" box="[1097,1254,1338,1365]" pageId="8" pageNumber="443" value="2005-05-04" valueMax="2005-05-05" valueMin="2005-05-04">04–05 v 2005</collectingDate>
</date>
, leg.
<collectorName id="26BF534EFFE7FFB1FAC9FAB1FA95C148" box="[1313,1407,1339,1365]" pageId="8" pageNumber="443">KT Park</collectorName>
&amp; SR.
<collectorName id="26BF534EFFE7FFB1FF2FFAD5FF16C164" box="[199,252,1375,1401]" pageId="8" pageNumber="443">Kim</collectorName>
, gen. slide nos. CIS-6305, nklep059.
<collectorName id="26BF534EFFE7FFB1FD76FAD5FD28C164" box="[670,706,1375,1401]" pageId="8" pageNumber="443">All</collectorName>
types in
<collectionCode id="ED5BAE5DFFE7FFB1FCC0FAD5FC85C164" box="[808,879,1375,1401]" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="8" pageNumber="443">NIBR</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3506513FFE7FFB0FF2FFA08FEB0C500" lastPageId="9" lastPageNumber="444" pageId="8" pageNumber="443" type="description">
<paragraph id="8BF53698FFE7FFB1FF2FFA08FE8CC318" blockId="8.[151,1437,1302,2013]" pageId="8" pageNumber="443">
<emphasis id="B93EEA8AFFE7FFB1FF2FFA08FEB3C181" bold="true" box="[199,345,1410,1436]" pageId="8" pageNumber="443">Description.</emphasis>
Adult (
<figureCitation id="13712A1DFFE7FFB1FE44FA09FDEAC180" box="[428,512,1410,1437]" captionStart="FIGURES 25–30" captionStartId="14.[152,270,1862,1886]" captionTargetBox="[158,1426,190,1823]" captionTargetId="figure-32@14.[150,1438,181,1837]" captionTargetPageId="14" captionText="FIGURES 25–30. Adults. 25. Dichomeris sclerospinula sp. nov., holotype, male; 26. D. sexafurca Li &amp; Zheng, male; 27. D. spinosella sp. nov., holotype, male; 28, D. unilobata sp. nov., holotype, male; 29. D. splendiptera Li &amp; Park, male; 30. D. summata Meyrick, female;" figureDoi="http://doi.org/10.5281/zenodo.4400943" httpUri="https://zenodo.org/record/4400943/files/figure.png" pageId="8" pageNumber="443">Fig. 25</figureCitation>
). Wingspan 12.5–14.5 mm. Head dark brown; frons gray. Ocellus present. Antenna with scape dark brown on dorsal surface, pale yellow on ventral surface; flagellum alternately dark-brown and yellowish brown on dorsal surface. Second segment of labial palpus dark brown on outer surface, grayish brown on inner surface except yellow near base, with gray-tipped triangular scale tuft dorsally; 3rd segment smoothly clothed, slightly shorter than 2nd segment, dark brown ventrally. Thorax and tegula dark brown; mesothoracic anepisternum without hair pencils in male. Fore- and midleg dark brown on outer surface, femora grayish white, tibiae and tarsi pale yellow on inner surface; hindleg grayish brown on outer surface, yellowish white apically, pale yellow on inner surface, tibia clothed with long, yellowish brown scales. Forewing elongate; ground color dark brown, with ill-defined yellowish–white postmedian line from costal 3/4 to before tornus, strongly convex outward, followed by more darkened area; apex acute; termen oblique; fringe dark brown. Hind wing dark grayish brown; fringe concolorous with ground color.
</paragraph>
<paragraph id="8BF53698FFE7FFB0FF2FF885FEB0C500" blockId="8.[151,1437,1302,2013]" lastBlockId="9.[151,1436,151,501]" lastPageId="9" lastPageNumber="444" pageId="8" pageNumber="443">
Male genitalia (
<figureCitation id="13712A1DFFE7FFB1FE9FF885FE26C334" box="[375,460,1807,1833]" captionStart="FIGURES 41–46" captionStartId="21.[152,270,1897,1921]" captionTargetBox="[247,1334,194,1857]" captionTargetId="figure-19@21.[238,1350,181,1873]" captionTargetPageId="21" captionText="FIGURES 41–46. Male genitalia. 41. Dichomeris ochriella sp. nov., holotype, gen. slide no. CIS-6277; 42. D. myriaodontosa sp. nov., holotype, gen. slide no. CIS-6675; 43. D. quadratella sp. nov., paratype, gen. slide no. CIS-6285; 44. D. sclerospinula sp. nov., paratype, gen. slide no. NKLEP059; 45. D. spinosella sp. nov., holotype, gen. slide no. NKLEP003; 46. D. unilobata sp. nov., holotype, gen. slide no. CIS-6320. Scales = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4400951" httpUri="https://zenodo.org/record/4400951/files/figure.png" pageId="8" pageNumber="443">Fig. 44</figureCitation>
): Uncus elongated, semiovate, densely setose on ventral surface. Gnathos strongly bent at basal 1/4, L-shaped, with acute apex upturned; culcitula irregular-shaped, densely spined. Valva subequal to length of tegumen plus uncus, nearly patrallel sided in basal 2/3, then inflated distally, with round apex; free ventral lobes small, about 1/5 length of valva, with round apex. Vinculum with two pairs of heavily sclerotized lateral lobes, symmetrical in both arms: posterior one, digitate, about 1/3 length of lateral arm, with round apex, densely setose; anterior one slightly shorter, spine-like; valvella setose, about equal to length of dorsal lobe. Saccal region narrow, slightly concave medially on anterior margin. Sica single, bifurcated apically, about equal to length of vinculum, nearly parallel sided. Aedeagus stout, as long as valva, nearly parallel-sided, bluntly rounded at base and apex; two short, sclerotized lobes from zone, at lateral side respectively, symmetrical, about 1/5 length of aedeagus, slightly bifurcate distally.
</paragraph>
</subSubSection>
<subSubSection id="C3506513FFE6FFB0FF2FFEACFDB6C55F" box="[199,604,294,322]" pageId="9" pageNumber="444" type="distribution">
<paragraph id="8BF53698FFE6FFB0FF2FFEACFDB6C55F" blockId="9.[151,1436,151,501]" box="[199,604,294,322]" pageId="9" pageNumber="444">
<emphasis id="B93EEA8AFFE6FFB0FF2FFEACFE8BC55D" bold="true" box="[199,353,294,320]" pageId="9" pageNumber="444">Distribution.</emphasis>
<collectingCountry id="F35D7608FFE6FFB0FE8FFEADFE2CC55C" box="[359,454,295,321]" name="Vietnam" pageId="9" pageNumber="444">Vietnam</collectingCountry>
(
<collectingRegion id="498EF87AFFE6FFB0FE3CFEADFDBAC55F" box="[468,592,295,322]" country="Vietnam" name="Vinh Phuc" pageId="9" pageNumber="444">Vinh Phuc</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3506513FFE6FFB0FF2FFEC0FC81C5B0" pageId="9" pageNumber="444" type="diagnosis">
<paragraph id="8BF53698FFE6FFB0FF2FFEC0FC81C5B0" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="444">
<emphasis id="B93EEA8AFFE6FFB0FF2FFEC0FEA8C579" bold="true" box="[199,322,330,356]" pageId="9" pageNumber="444">Diagnosis.</emphasis>
This species can be distinguished from congeners by the dark-grayish brown forewing. It can be easily distinguished from congers by having the gnathos strongly bent into L-shape, vinculum with two pairs of strong lateral lobes, and a strong sica parallel sided, bifurcated apically.
</paragraph>
</subSubSection>
<subSubSection id="C3506513FFE6FFB0FF2FFE3DFCCAC5E8" pageId="9" pageNumber="444" type="etymology">
<paragraph id="8BF53698FFE6FFB0FF2FFE3DFCCAC5E8" blockId="9.[151,1436,151,501]" pageId="9" pageNumber="444">
<emphasis id="B93EEA8AFFE6FFB0FF2FFE3DFEA4C5CC" bold="true" box="[199,334,439,465]" pageId="9" pageNumber="444">Etymology.</emphasis>
The specific name is derived from the Latin,
<emphasis id="B93EEA8AFFE6FFB0FCA9FE3DFC72C5CC" box="[833,920,439,465]" italics="true" pageId="9" pageNumber="444">-sclero-</emphasis>
(= sclerotized) and
<emphasis id="B93EEA8AFFE6FFB0FB9AFE3DFB2EC5CC" box="[1138,1220,439,465]" italics="true" pageId="9" pageNumber="444">spinula</emphasis>
(= spine), referring to the sclerotized spine on vinculum of the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>
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