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<document id="B2C43AE3624D6DDAC5268B6CE6953AB5" ID-DOI="10.24199/j.mmv.2023.82.06" ID-ISSN="1447-2554" ID-Zenodo-Dep="12214602" ID-ZooBank="urn:lsid:zoobank.org:pub:B48A3062-C822-4DD1-ADCC-A577C9D2CD32" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718999013290" checkinUser="felipe" docAuthor="Mah, Christopher L." docDate="2023" docId="03E33A5AFFE0136AFCF8F974FD01FDE9" docLanguage="en" docName="MemMusVictoria.82.119-131.pdf" docOrigin="Memoirs of Museum Victoria 82" docSource="http://dx.doi.org/10.24199/j.mmv.2023.82.06" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Poraniomorpha tartarus Mah, 2023, n. sp." docType="treatment" docUuid="F714BE6E-130F-4454-8B31-2278F3724E07" docUuidSource="ZooBank" docVersion="2" lastPageNumber="129" masterDocId="FFDA4222FFE61360FFD1FFB3FFC8FFB9" masterDocTitle="New occurrence of Poraniidae (Valvatacea, Asteroidea) in Australia with a new genus and species from deep-sea settings" masterLastPageNumber="131" masterPageNumber="119" pageNumber="125" updateTime="1719016232216" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0A3584B0180815D80AD49F0586B09065">New occurrence of Poraniidae (Valvatacea, Asteroidea) in Australia with a new genus and species from deep-sea settings</mods:title>
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<mods:namePart id="81F161A40A0E03EF67FCD07188F4A434">Mah, Christopher L.</mods:namePart>
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<mods:date id="33A9A51374F41F864681B01C809DFD15">2023</mods:date>
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<mods:number id="26350C0BC4CDAB9120BCE1194AF3DD1B">2023-08-08</mods:number>
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<mods:identifier id="F703EDF00A9BB2DE81D82F3CBC75B26B" type="DOI">10.24199/j.mmv.2023.82.06</mods:identifier>
<mods:identifier id="47C36CF513C3F3364B12DDE01F372895" type="ISSN">1447-2554</mods:identifier>
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<treatment id="03E33A5AFFE0136AFCF8F974FD01FDE9" LSID="urn:lsid:zoobank.org:act:F714BE6E-130F-4454-8B31-2278F3724E07" httpUri="http://treatment.plazi.org/id/03E33A5AFFE0136AFCF8F974FD01FDE9" lastPageId="10" lastPageNumber="129" pageId="6" pageNumber="125">
<subSubSection id="C350D8C7FFE01366FCF8F974FBA4F965" box="[809,1132,1735,1757]" pageId="6" pageNumber="125" type="nomenclature">
<paragraph id="8BF58B4CFFE01366FCF8F974FBA4F965" blockId="6.[809,1132,1735,1757]" box="[809,1132,1735,1757]" pageId="6" pageNumber="125">
<heading id="D0BD3C20FFE01366FCF8F974FBA4F965" bold="true" box="[809,1132,1735,1757]" fontSize="9" level="2" pageId="6" pageNumber="125" reason="2">
<emphasis id="B93E575EFFE01366FCF8F974FBA4F965" bold="true" box="[809,1132,1735,1757]" pageId="6" pageNumber="125">
<taxonomicName id="4C4AF0CFFFE01366FCF8F974FBE5F964" ID-CoL="BSBYJ" authorityName="Mah" authorityYear="2023" box="[809,1069,1735,1757]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="6" pageNumber="125" phylum="Echinodermata" rank="species" species="tartarus" status="sp. nov.">
<emphasis id="B93E575EFFE01366FCF8F974FBE5F964" bold="true" box="[809,1069,1735,1757]" italics="true" pageId="6" pageNumber="125">Poraniomorpha tartarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DEA25FFE01366FBE2F974FBA4F965" box="[1075,1132,1735,1756]" pageId="6" pageNumber="125" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C350D8C7FFE01368FCF8F940FD23FC35" lastPageId="8" lastPageNumber="127" pageId="6" pageNumber="125" type="description">
<paragraph id="8BF58B4CFFE01366FCF8F940FC60F8B0" blockId="6.[809,936,1779,1801]" box="[809,936,1779,1801]" pageId="6" pageNumber="125">
<figureCitation id="137197C9FFE01366FCF8F940FC6CF8B0" box="[809,932,1779,1801]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="6" pageNumber="125">Figure 3a–e</figureCitation>
.
</paragraph>
<paragraph id="8BF58B4CFFE01366FCF8F8ACFB7FF8D6" blockId="6.[809,1440,1823,1903]" pageId="6" pageNumber="125">
<emphasis id="B93E575EFFE01366FCF8F8ACFC52F88D" box="[809,922,1823,1844]" italics="true" pageId="6" pageNumber="125">Etymology.</emphasis>
The species epithet is named for Tartarus, the mythical Greek underworld, alluding to this species’ occurrence at great depth. Noun held in apposition.
</paragraph>
<caption id="DF35DBC4FFE11367FF42F8F2FA8FF8D7" ID-DOI="http://doi.org/10.5281/zenodo.12214606" ID-Zenodo-Dep="12214606" httpUri="https://zenodo.org/record/12214606/files/figure.png" pageId="7" pageNumber="126" startId="7.[147,207,1857,1876]" targetBox="[207,1389,227,1839]" targetPageId="7" targetType="figure">
<paragraph id="8BF58B4CFFE11367FF42F8F2FA8FF8D7" blockId="7.[147,1449,1857,1902]" pageId="7" pageNumber="126">
Figure 2.
<taxonomicName id="4C4AF0CFFFE11367FF3BF8F2FDDCF8ED" authority="NMV F" authorityName="NMV F" box="[234,532,1857,1876]" class="Asteroidea" family="Poraniidae" genus="Marginaster" kingdom="Animalia" order="Valvatida" pageId="7" pageNumber="126" phylum="Echinodermata" rank="species" species="paucispinus">
<emphasis id="B93E575EFFE11367FF3BF8F2FE0FF8EA" box="[234,455,1857,1875]" italics="true" pageId="7" pageNumber="126">Marginaster paucispinus</emphasis>
NMV F
</taxonomicName>
240386: a, abactinal view; b, actinal view; USNM holotype 032641: c, abactinal view; d, actinal view. MNHN-2013-1518: e, abactinal view; f, actinal view. Scale bars a, b=1.0 mm, c, d, e, f=1.5 mm. Photos by Melanie Mackenzie NMV.
</paragraph>
</caption>
<paragraph id="8BF58B4CFFEE1368FF5DFF55FE72FD77" blockId="8.[139,772,229,719]" pageId="8" pageNumber="127">
<emphasis id="B93E575EFFEE1368FF5DFF55FF32FF42" box="[140,250,230,251]" italics="true" pageId="8" pageNumber="127">Diagnosis.</emphasis>
Body strongly stellate (
<figureCitation id="137197C9FFEE1368FDC0FF56FDAAFF43" box="[529,610,229,251]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="8" pageNumber="127">Fig. 3a</figureCitation>
). Actinolateral fringe rounded with rounded edge (
<figureCitation id="137197C9FFEE1368FDC6FEB1FD4EFEA1" box="[535,646,258,280]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="8" pageNumber="127">Fig. 3c, d</figureCitation>
). Abactinal surface hard, very resistant to the touch. Surface covered by minute, pointed granules embedded in dermal integument (
<figureCitation id="137197C9FFEE1368FF42FEE9FF30FEC9" box="[147,248,346,368]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="8" pageNumber="127">Fig. 3b, d</figureCitation>
). Drying of specimen suggests flattened marginals are polygonal plates, approximately 48 per interradius. Actinal region narrow, surface is flat, with approximately 50 shallow segments corresponding with adambulacral and marginal plates but also bisecting the actinal intermediate region extending to the oral plate. Actinal surface also covered by small granules, covered by dermis, 5–30 per actinal segment, each with a hyaline tip invested in the dermal integument. Dark brown colour adjacent to the adambulacral spination on the disk and along the arms (
<figureCitation id="137197C9FFEE1368FE66FDD2FDF9FDCF" box="[439,561,609,631]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="8" pageNumber="127">Fig. 3c, d, e</figureCitation>
). One or two furrow spines, with 1–2 enlarged subambulacral spines, each approximately twice the thickness of and more elongate than the furrow spines (
<figureCitation id="137197C9FFEE1368FE9CFD0AFE66FD77" box="[333,430,697,719]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="8" pageNumber="127">Fig. 3d, e</figureCitation>
).
</paragraph>
<paragraph id="8BF58B4CFFEE1368FF5DFD56FD23FC35" blockId="8.[137,773,741,1668]" pageId="8" pageNumber="127">
<emphasis id="B93E575EFFEE1368FF5DFD56FF31FD43" box="[140,249,741,762]" italics="true" pageId="8" pageNumber="127">Comments.</emphasis>
Although
<taxonomicName id="4C4AF0CFFFEE1368FEBCFD55FE15FD42" authorityName="Mah" authorityYear="2023" box="[365,477,742,763]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="tartarus" status="n. sp.">
<emphasis id="B93E575EFFEE1368FEBCFD55FE15FD42" box="[365,477,742,763]" italics="true" pageId="8" pageNumber="127">P. tartarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DEA25FFEE1368FE3BFD56FDECFD43" box="[490,548,741,762]" pageId="8" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
possesses a rounded actinolateral edge, unlike other
<taxonomicName id="4C4AF0CFFFEE1368FE16FCB1FD7BFCA1" box="[455,691,770,792]" pageId="8" pageNumber="127">
<emphasis id="B93E575EFFEE1368FE16FCB1FDADFCAE" box="[455,613,770,791]" italics="true" pageId="8" pageNumber="127">Poraniomorpha</emphasis>
species
</taxonomicName>
, several other characters – including irregular imbricate plates, the pointed granules invested in the dermal tissue, and the distinctively enlarged subambulacral and furrow spination on the adambulacral plates (
<figureCitation id="137197C9FFEE1368FF1FFCC4FEE1FC35" box="[206,297,887,909]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="8" pageNumber="127">Fig. 3d, e</figureCitation>
) – support placement within
<taxonomicName id="4C4AF0CFFFEE1368FD96FCC4FD2DFC35" authorityName="Danielssen and Koren" authorityYear="1881" box="[583,741,887,908]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="genus">
<emphasis id="B93E575EFFEE1368FD96FCC4FD2DFC35" box="[583,741,887,908]" italics="true" pageId="8" pageNumber="127">Poraniomorpha</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C350D8C7FFEE1368FF63FC27FE4DF9B6" pageId="8" pageNumber="127" type="reference_group">
<paragraph id="8BF58B4CFFEE1368FF63FC27FD30FAC4" blockId="8.[137,773,741,1668]" pageId="8" pageNumber="127">
<taxonomicName id="4C4AF0CFFFEE1368FF63FC27FE7DFC13" ID-CoL="BSBYJ" authorityName="Mah" authorityYear="2023" box="[178,437,916,938]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="tartarus" status="sp. nov.">
<emphasis id="B93E575EFFEE1368FF63FC27FE7DFC13" box="[178,437,916,938]" italics="true" pageId="8" pageNumber="127">Poraniomorpha tartarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DEA25FFEE1368FE11FC26FE32FC13" box="[448,506,917,938]" pageId="8" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
invites comparison with Atlantic species such as
<taxonomicName id="4C4AF0CFFFEE1368FE5EFC01FD7FFC7E" ID-CoL="4LVVN" authority="(Verrill, 1895)" baseAuthorityName="Verrill" baseAuthorityYear="1895" box="[399,695,945,967]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="abyssicola">
<emphasis id="B93E575EFFEE1368FE5EFC01FDD1FC7F" box="[399,537,945,967]" italics="true" pageId="8" pageNumber="127">P. abyssicola</emphasis>
(
<bibRefCitation id="EFDBF6BDFFEE1368FDF8FC02FD78FC7E" author="Verrill, A. E." box="[553,688,945,967]" pageId="8" pageNumber="127" pagination="127 - 141" refId="ref13031" refString="Verrill, A. E. 1895. Distribution of the echinoderms of north - eastern America. American Journal of Science 3 (49): 127 - 141." type="journal article" year="1895">Verrill, 1895</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C4AF0CFFFEE1368FD3EFC01FE5AFC5D" ID-CoL="6VVVV" authority="(Stuxberg, 1878)" baseAuthorityName="Stuxberg" baseAuthorityYear="1878" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="tumida">
<emphasis id="B93E575EFFEE1368FD3EFC01FF1CFC5A" italics="true" pageId="8" pageNumber="127">P. tumida</emphasis>
(
<bibRefCitation id="EFDBF6BDFFEE1368FF38FC7DFE43FC5D" author="Stuxberg, A." box="[233,395,974,996]" pageId="8" pageNumber="127" pagination="27 - 40" refId="ref12935" refString="Stuxberg, A. 1878. Ecninodermer fran Novaja Semljas haf samlade under Nordenskioldska expeditionerna 1875 och 1876. Ofversigt af Kon gl. Vetenskaps-Akademiens Forhandlingar 3: 27 - 40." type="journal article" year="1878">Stuxberg, 1878</bibRefCitation>
)
</taxonomicName>
, which it resembles very closely. They share similar abactinal plate morphology, displaying closely articulated irregular-shaped, imbricate, mound-like plates bearing a cover of spinose granules covering the surface such that plate boundaries are obscured. The disk in these species is strongly arched and arms are similarly elongate and tapering. Furrow spines are relatively few in both species (
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<taxonomicName id="4C4AF0CFFFEE1368FEADFB2FFE39FB08" ID-CoL="BSBYJ" authorityName="Mah" authorityYear="2023" box="[380,497,1180,1201]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="tartarus" status="n. sp.">
<emphasis id="B93E575EFFEE1368FEADFB2FFE39FB08" box="[380,497,1180,1201]" italics="true" pageId="8" pageNumber="127">P. tartarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DEA25FFEE1368FE2EFB28FDF4FB09" box="[511,572,1179,1200]" pageId="8" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
versus
<quantity id="4CB226A9FFEE1368FD49FB28FD28FB08" box="[664,736,1179,1201]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="7.62" metricValueMin="5.08" pageId="8" pageNumber="127" unit="in" value="2.5" valueMax="3.0" valueMin="2.0">2–3 in</quantity>
<taxonomicName id="4C4AF0CFFFEE1368FD3EFB2FFF31FB74" ID-CoL="4LVVN" baseAuthorityName="Verrill" baseAuthorityYear="1895" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="abyssicola">
<emphasis id="B93E575EFFEE1368FD3EFB2FFF31FB74" italics="true" pageId="8" pageNumber="127">P. abyssicola</emphasis>
</taxonomicName>
but
<quantity id="4CB226A9FFEE1368FEF9FB0BFEA2FB77" box="[296,362,1208,1230]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="8" pageNumber="127" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">3–5 in</quantity>
<taxonomicName id="4C4AF0CFFFEE1368FEA3FB0AFE1CFB74" ID-CoL="6VVVV" baseAuthorityName="Stuxberg" baseAuthorityYear="1878" box="[370,468,1208,1230]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="tumida">
<emphasis id="B93E575EFFEE1368FEA3FB0AFE1CFB74" box="[370,468,1208,1230]" italics="true" pageId="8" pageNumber="127">P. tumida</emphasis>
</taxonomicName>
).
<taxonomicName id="4C4AF0CFFFEE1368FE38FB0BFD21FB77" ID-CoL="BSBYJ" authorityName="Mah" authorityYear="2023" box="[489,745,1208,1230]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="tartarus" status="sp. nov.">
<emphasis id="B93E575EFFEE1368FE38FB0BFD21FB77" box="[489,745,1208,1230]" italics="true" pageId="8" pageNumber="127">Poraniomorpha tartarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DEA25FFEE1368FD20FB0AFF60FB52" pageId="8" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
is distinguished by the enlarged subambulacral spine, the absence of most papulae from the abactinal surface, and the difference in actinal plate texture, which possesses distinct transverse segments and lacks the numerous pointed spinelets seen in
<taxonomicName id="4C4AF0CFFFEE1368FF0AFAF8FEAAFAE6" ID-CoL="4LVVN" baseAuthorityName="Verrill" baseAuthorityYear="1895" box="[219,354,1354,1376]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="abyssicola">
<emphasis id="B93E575EFFEE1368FF0AFAF8FEAAFAE6" box="[219,354,1354,1376]" italics="true" pageId="8" pageNumber="127">P. abyssicola</emphasis>
</taxonomicName>
. Neither
<taxonomicName id="4C4AF0CFFFEE1368FE12FAF8FD82FAE6" ID-CoL="4LVVN" baseAuthorityName="Verrill" baseAuthorityYear="1895" box="[451,586,1354,1376]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="abyssicola">
<emphasis id="B93E575EFFEE1368FE12FAF8FD82FAE6" box="[451,586,1354,1376]" italics="true" pageId="8" pageNumber="127">P. abyssicola</emphasis>
</taxonomicName>
nor
<taxonomicName id="4C4AF0CFFFEE1368FDA8FAF8FD14FAE6" ID-CoL="6VVVV" baseAuthorityName="Stuxberg" baseAuthorityYear="1878" box="[633,732,1354,1376]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="tumida">
<emphasis id="B93E575EFFEE1368FDA8FAF8FD14FAE6" box="[633,732,1354,1376]" italics="true" pageId="8" pageNumber="127">P. tumida</emphasis>
</taxonomicName>
are known to display the distinct colouration of
<taxonomicName id="4C4AF0CFFFEE1368FD9FFADBFD75FAC4" ID-CoL="BSBYJ" authorityName="Mah" authorityYear="2023" box="[590,701,1384,1405]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="tartarus" status="n. sp.">
<emphasis id="B93E575EFFEE1368FD9FFADBFD75FAC4" box="[590,701,1384,1405]" italics="true" pageId="8" pageNumber="127">P. tartarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DEA25FFEE1368FD12FADBFD30FAC4" box="[707,760,1384,1405]" pageId="8" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BF58B4CFFEE1368FF63FA36FE4DF9B6" blockId="8.[137,773,741,1668]" pageId="8" pageNumber="127">
<taxonomicName id="4C4AF0CFFFEE1368FF63FA36FE76FA23" ID-CoL="BSBYJ" authorityName="Mah" authorityYear="2023" box="[178,446,1413,1434]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="tartarus" status="sp. nov.">
<emphasis id="B93E575EFFEE1368FF63FA36FE76FA23" box="[178,446,1413,1434]" italics="true" pageId="8" pageNumber="127">Poraniomorpha tartarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DEA25FFEE1368FE03FA36FDDDFA23" box="[466,533,1413,1434]" pageId="8" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
is the first known occurrence of this genus in the Southern Hemisphere and is also the deepest known species of the genus (
<quantity id="4CB226A9FFEE1368FDB9FA0CFD30FA6D" box="[616,760,1471,1493]" metricMagnitude="3" metricUnit="m" metricValue="3.8515" metricValueMax="3.853" metricValueMin="3.85" pageId="8" pageNumber="127" unit="m" value="3851.5" valueMax="3853.0" valueMin="3850.0">3850–3853 m</quantity>
).
<taxonomicName id="4C4AF0CFFFEE1368FF5FFA6FFE6DFA48" ID-CoL="4LVVN" baseAuthorityName="Verrill" baseAuthorityYear="1895" box="[142,421,1500,1521]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="abyssicola">
<emphasis id="B93E575EFFEE1368FF5FFA6FFE6DFA48" box="[142,421,1500,1521]" italics="true" pageId="8" pageNumber="127">Poraniomorpha abyssicola</emphasis>
</taxonomicName>
occurs at comparable depth in the Atlantic (
<quantity id="4CB226A9FFEE1368FF3AFA4AFEB2F9B6" box="[235,378,1529,1551]" metricMagnitude="3" metricUnit="m" metricValue="3.3579999999999997" metricValueMax="3.7399999999999998" metricValueMin="2.976" pageId="8" pageNumber="127" unit="m" value="3358.0" valueMax="3740.0" valueMin="2976.0">2976–3740 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C350D8C7FFEE136AFF63F9A4FD01FDE9" lastPageId="10" lastPageNumber="129" pageId="8" pageNumber="127" type="description">
<paragraph id="8BF58B4CFFEE1368FF63F9A4FCC8F93D" blockId="8.[137,773,741,1668]" pageId="8" pageNumber="127">
Although very little is known about the biology of
<taxonomicName id="4C4AF0CFFFEE1368FF5DF987FEACF9F0" box="[140,356,1588,1609]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="undetermined">
<emphasis id="B93E575EFFEE1368FF5DF987FEE3F9F0" box="[140,299,1588,1609]" italics="true" pageId="8" pageNumber="127">Poraniomorpha</emphasis>
spp.
</taxonomicName>
, the NOAA vessel
<taxonomicName id="4C4AF0CFFFEE1368FD92F987FCCAF9F0" authority="Explorer" authorityName="Explorer" box="[579,770,1588,1609]" class="Insecta" family="Pentatomidae" genus="Okeanos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="B93E575EFFEE1368FD92F987FCCAF9F0" box="[579,770,1588,1609]" italics="true" pageId="8" pageNumber="127">Okeanos Explorer</emphasis>
</taxonomicName>
observed
<taxonomicName id="4C4AF0CFFFEE1368FF3FF9E1FEBBF9DF" baseAuthorityName="Verrill" baseAuthorityYear="1895" box="[238,371,1617,1639]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="abyssicola">
<emphasis id="B93E575EFFEE1368FF3FF9E1FEBBF9DF" box="[238,371,1617,1639]" italics="true" pageId="8" pageNumber="127">P. abyssicola</emphasis>
</taxonomicName>
feeding on a sponge at
<quantity id="4CB226A9FFEE1368FDBEF9E2FD76F9DE" box="[623,702,1617,1639]" metricMagnitude="3" metricUnit="m" metricValue="3.403" pageId="8" pageNumber="127" unit="m" value="3403.0">3403 m</quantity>
(
<bibRefCitation id="EFDBF6BDFFEE1368FD1CF9E2FF08F93A" author="Mah, C. L." pageId="8" pageNumber="127" pagination="201 - 260" refId="ref11461" refString="Mah, C. L. 2020. New species, occurrence records and observations of predation by deep-sea Asteroidea (Echinodermata) from the North Atlantic by NOAA Ship Okeanos Explorer. Zootaxa 4766 (2): 201 - 260. https: // doi. org / 10.11646 / zootaxa. 4766.2.1" type="journal article" year="2020">Mah, 2020</bibRefCitation>
).
<taxonomicName id="4C4AF0CFFFEE1368FF01F9DDFE0EF93D" authorityName="Mah" authorityYear="2023" box="[208,454,1646,1668]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="8" pageNumber="127" phylum="Echinodermata" rank="species" species="tartarus" status="sp. nov.">
<emphasis id="B93E575EFFEE1368FF01F9DDFE0EF93D" box="[208,454,1646,1668]" italics="true" pageId="8" pageNumber="127">Poraniomorpha tartarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DEA25FFEE1368FE1DF9DCFDC8F93D" box="[460,512,1647,1668]" pageId="8" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
may feed on similar prey.
</paragraph>
<paragraph id="8BF58B4CFFEE1368FF5DF928FE84F974" blockId="8.[139,770,1691,1742]" pageId="8" pageNumber="127">
<emphasis id="B93E575EFFEE1368FF5DF928FECFF909" box="[140,263,1691,1712]" italics="true" pageId="8" pageNumber="127">Occurrence.</emphasis>
<materialsCitation id="3B228111FFEE1368FEDDF928FE80F974" county="Known" elevation="3852" location="Sea" municipality="East Gippsland" pageId="8" pageNumber="127" specimenCount="1" stateProvince="Victoria, Tasman">
<collectingCounty id="6294F3C0FFEE1368FEDDF928FE9EF909" box="[268,342,1691,1712]" pageId="8" pageNumber="127">Known</collectingCounty>
only off
<collectingMunicipality id="6B911136FFEE1368FE60F928FD84F908" box="[433,588,1691,1713]" pageId="8" pageNumber="127">East Gippsland</collectingMunicipality>
,
<collectingRegion id="498E45AEFFEE1368FD87F928FCCAF909" box="[598,770,1691,1713]" isEnumeration="true" pageId="8" pageNumber="127">Victoria, Tasman</collectingRegion>
<location id="8E95DD97FFEE1368FF5AF90BFF79F974" LSID="urn:lsid:plazi:treatment:03E33A5AFFE0136AFCF8F974FD01FDE9:8E95DD97FFEE1368FF5AF90BFF79F974" box="[139,177,1720,1741]" county="Known" municipality="East Gippsland" name="Sea" pageId="8" pageNumber="127" stateProvince="Victoria, Tasman">Sea</location>
,
<quantity id="4CB226A9FFEE1368FF6DF90BFE80F974" box="[188,328,1720,1742]" metricMagnitude="3" metricUnit="m" metricValue="3.8515" metricValueMax="3.853" metricValueMin="3.85" pageId="8" pageNumber="127" unit="m" value="3851.5" valueMax="3853.0" valueMin="3850.0">
<elevation id="00676C7FFFEE1368FF6DF90BFE80F974" box="[188,328,1720,1742]" metricMagnitude="3" metricUnit="m" metricValue="3.8515" metricValueMax="3.853" metricValueMin="3.85" pageId="8" pageNumber="127" unit="m" value="3851.5" valueMax="3853.0" valueMin="3850.0">3850–3853 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF58B4CFFEE1368FF5DF956FEADF8D6" blockId="8.[140,771,1764,1903]" pageId="8" pageNumber="127">
<emphasis id="B93E575EFFEE1368FF5DF956FEC9F943" box="[140,257,1765,1786]" italics="true" pageId="8" pageNumber="127">Description</emphasis>
. Body strongly stellate (R/r=3.7–3.9) (
<figureCitation id="137197C9FFEE1368FD46F957FD33F940" box="[663,763,1764,1786]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="8" pageNumber="127">Fig. 3a, c</figureCitation>
) with elongate arms, round in cross-section. Disk and arms confluent, disk strongly convex, rms thick, tips strongly upturned. Interradial arcs acute. Actinolateral fringe rounded with no distinct edge.
</paragraph>
<paragraph id="8BF58B4CFFEE1368FC9EFF56FC34FCC0" blockId="8.[809,1442,229,1903]" pageId="8" pageNumber="127">
Disk plates not evident, thick integument covers all of body surface (
<figureCitation id="137197C9FFEE1368FC6DFEB1FBD5FEA1" box="[956,1053,258,280]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="8" pageNumber="127">Fig. 3a, b</figureCitation>
). Body texture hard to touch. General surface topology rough, presenting a wrinkled appearance, covered by minute, pointed granules invested in dermal integument (
<figureCitation id="137197C9FFEE1368FC60FEEAFBC8FED6" box="[945,1024,345,367]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="8" pageNumber="127">Fig. 3b</figureCitation>
). Individual plates irregular in shape, surface texture mound-like. Granular cover is complete, evenly distributed, approximately 5–6 along a 1.0 mm line. Papulae mostly absent from abactinal surface with some occurring interradially (5–8 observed between arms in each interradius), with few present proximally on the lateral sides of each arm. Anus at center of disk, flanked by 8–10 angular granules around edge. Dermal granules around anus slightly larger than those elsewhere. Madreporite convex, outline circular, large, approximately 1.5 (smaller specimen) to 3.0 (larger specimen) cm in diameter (
<figureCitation id="137197C9FFEE1368FB56FDC8FB2FFD28" box="[1159,1255,635,657]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="8" pageNumber="127">Fig. 3a, b</figureCitation>
). On the
<typeStatus id="54F135EEFFEE1368FA90FDC8FA54FD28" box="[1345,1436,635,657]" pageId="8" pageNumber="127" type="holotype">holotype</typeStatus>
, madreporite adjacent to contact with superomarginal plates. Basal portion of madreporite covered by small dermal granules. No pedicellariae observed. Interradii each with a discrete fold each bearing 10–15 slender grooves that synchronise with those on marginal to actinal plates. Shallow transverse fasciolar channels extend from abactinal surface along lateral surface aligned with those on actinal surface and adambulacral plates.
</paragraph>
<paragraph id="8BF58B4CFFEE1368FC9EFC33FB46FB9F" blockId="8.[809,1442,229,1903]" pageId="8" pageNumber="127">Marginal plates completely obscured by dermal integument. Drying shows marginal plates, flattened, polygonal, approximately 48 per interradius (armtip to armtip), boundaries are obscured, exact count uncertain. Single row of approximately 5–8 papulae, in mostly single pores, along lateral surface of arm.</paragraph>
<paragraph id="8BF58B4CFFEE1368FC9EFB9DFB88FAF1" blockId="8.[809,1442,229,1903]" pageId="8" pageNumber="127">
Actinal surface flat with approximately 50 shallow transverse grooves tracking from marginal to adambulacral plates forming segments. These grooves bisecting the actinal intermediate region tracking from the oral plate (
<figureCitation id="137197C9FFEE1368FAE0FB37FA5EFB23" box="[1329,1430,1156,1178]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="8" pageNumber="127">Fig. 3d, e</figureCitation>
). Actinal surface covered by small granules, 5–30 per actinal segment, covered by dermis, each with a hyaline tip invested in the dermal integument. A dark colour pattern, especially evident on NMV F241811 (
<figureCitation id="137197C9FFEE1368FB9EFB4BFB52FAB7" box="[1103,1178,1272,1294]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="8" pageNumber="127">Fig. 3c</figureCitation>
), present adjacent to the adambulacral spine series and around the mouth extending interradially on to the disk.
</paragraph>
<paragraph id="8BF58B4CFFEE1368FC9EFAE3FB9DF942" blockId="8.[809,1442,229,1903]" pageId="8" pageNumber="127">
Furrow spines (1–2) large and prominent, covered by dermal integument, conical tip, blunt spines widely spaced interlacing with furrow spines on opposing side (
<figureCitation id="137197C9FFEE1368FACDFA3AFA5DFA27" box="[1308,1429,1417,1439]" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="8" pageNumber="127">Fig. 3b, d, e</figureCitation>
). Subambulacral spine mostly single, but two are present on approximately 40% of adambulacrals, especially on NMV F241811, approximately twice the thickness of each furrow spine arranged transversely relative to the furrow spine. Remaining adambulacral plates with single, short subambulacral spine, variably blunt and smooth, a minority of spines with notched or roughened tip. At least one of the larger subambulacral spines comparable in size with one of the furrow spines. Those secondary subambulacrals spines smaller, less than half the height and thickness of the furrow spine. Dermal integument covers the adambulacral plate; no other accessories are present.
</paragraph>
<paragraph id="8BF58B4CFFEE1368FC9EF8B1FC43F8D6" blockId="8.[809,1442,229,1903]" pageId="8" pageNumber="127">Oral plates with four furrow spines and one spine from each oral plate projecting into the mouth (two total). Oral plate surface, with a total of 4–6 suboral spines (two or three per half).</paragraph>
<caption id="DF35DBC4FFEF1369FF42F895FEBDF8D7" ID-DOI="http://doi.org/10.5281/zenodo.12214608" ID-Zenodo-Dep="12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="9" pageNumber="128" startId="9.[147,207,1830,1849]" targetBox="[147,1447,227,1799]" targetPageId="9" targetType="figure">
<paragraph id="8BF58B4CFFEF1369FF42F895FEBDF8D7" blockId="9.[147,1450,1830,1902]" pageId="9" pageNumber="128">
Figure 3.
<taxonomicName id="4C4AF0CFFFEF1369FF3EF895FE02F880" authorityName="Mah" authorityYear="2023" box="[239,458,1830,1849]" class="Asteroidea" family="Poraniidae" genus="Poraniomorpha" kingdom="Animalia" order="Valvatida" pageId="9" pageNumber="128" phylum="Echinodermata" rank="species" species="tartarus" status="sp. nov.">
<emphasis id="B93E575EFFEF1369FF3EF895FE02F880" box="[239,458,1830,1849]" italics="true" pageId="9" pageNumber="128">Poraniomorpha tartarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DEA25FFEF1369FE01F894FDCBF883" box="[464,515,1831,1850]" pageId="9" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
, holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm.
</paragraph>
</caption>
<paragraph id="8BF58B4CFFEC136AFF63FF56FE96FECC" blockId="10.[140,772,229,373]" pageId="10" pageNumber="129">
Colour in life white to dark brown on disk, arms mottled, dark brown to white, interradii dark brown. Underside is white with dark brown in each interradius around mouth (
<figureCitation id="137197C9FFEC136AFD0BFE91FF3AFEEF" captionStart="Figure 3" captionStartId="9.[147,207,1830,1849]" captionTargetBox="[147,1447,227,1799]" captionTargetId="figure-7@9.[147,1447,227,1799]" captionTargetPageId="9" captionText="Figure 3. Poraniomorpha tartarus n. sp., holotype. NMV F 241811: a, abactinal; b, abactinal surface showing madreporite, surface texture; c, actinal view; d, actinal intermediate region and inferomarginals; e, actinal view furrow spines, oral region. Scale bars a=9.0 mm, b=5.0 mm, c=9.0 mm, d, e=2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12214608" httpUri="https://zenodo.org/record/12214608/files/figure.png" pageId="10" pageNumber="129">Fig. 3c, d, e</figureCitation>
). Dark colouration present in patches along adambulacral series.
</paragraph>
<paragraph id="8BF58B4CFFEC136AFF5DFE3EFDC9FE44" blockId="10.[139,771,397,593]" pageId="10" pageNumber="129">
<emphasis id="B93E575EFFEC136AFF5DFE3EFE83FE1B" box="[140,331,397,418]" italics="true" pageId="10" pageNumber="129">Material examined</emphasis>
.
<materialsCitation id="3B228111FFEC136AFE87FE3EFDC9FE44" collectingDate="2017-05-24" collectionCode="NMV, RV, R" collectorName="Coll. O'Hara" country="Australia" elevation="3852" latitude="-38.479" location="East Gippsland" longLatPrecision="70" longitude="150.185" municipality="Tasman Sea" pageId="10" pageNumber="129" specimenCode="F 241811" specimenCount="1" stateProvince="Victoria" typeStatus="holotype">
<typeStatus id="54F135EEFFEC136AFE87FE3EFE70FE1A" box="[342,440,397,419]" pageId="10" pageNumber="129" type="holotype">Holotype</typeStatus>
.
<collectionCode id="ED5B1389FFEC136AFE6EFE3EFE32FE1B" box="[447,506,397,418]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="10" pageNumber="129" type="Museum">NMV</collectionCode>
<specimenCode id="DBEC2337FFEC136AFDD1FE3EFD96FE1B" box="[512,606,397,418]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="10" pageNumber="129" type="Museum">F 241811</specimenCode>
<location id="8E95DD97FFEC136AFDB2FE3EFD37FE1A" LSID="urn:lsid:plazi:treatment:03E33A5AFFE0136AFCF8F974FD01FDE9:8E95DD97FFEC136AFDB2FE3EFD37FE1A" box="[611,767,397,419]" country="Australia" latitude="-38.479" longLatPrecision="70" longitude="150.185" municipality="Tasman Sea" name="East Gippsland" pageId="10" pageNumber="129" stateProvince="Victoria">East Gippsland</location>
,
<collectingRegion id="498E45AEFFEC136AFF5AFE18FF17FE78" box="[139,223,427,449]" country="Australia" name="Victoria" pageId="10" pageNumber="129">Victoria</collectingRegion>
,
<collectingMunicipality id="6B911136FFEC136AFF3EFE1FFEB9FE79" box="[239,369,427,449]" pageId="10" pageNumber="129">Tasman Sea</collectingMunicipality>
,
<collectingCountry id="F35DCBDCFFEC136AFE51FE18FE28FE78" box="[384,480,427,449]" name="Australia" pageId="10" pageNumber="129">Australia</collectingCountry>
, -
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,
<geoCoordinate id="EE7EED8BFFEC136AFD52FE18FD35FE78" box="[643,765,427,449]" degrees="150.185" direction="east" orientation="longitude" pageId="10" pageNumber="129" precision="55" value="150.185">150.185° E</geoCoordinate>
,
<quantity id="4CB226A9FFEC136AFF5DFE7AFED4FE66" box="[140,284,457,479]" metricMagnitude="3" metricUnit="m" metricValue="3.8515" metricValueMax="3.853" metricValueMin="3.85" pageId="10" pageNumber="129" unit="m" value="3851.5" valueMax="3853.0" valueMin="3850.0">
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</quantity>
,
<collectorName id="26BFEE9AFFEC136AFEF9FE79FE66FE66" box="[296,430,458,480]" pageId="10" pageNumber="129">Coll. O’Hara</collectorName>
et al. aboard
<collectionCode id="ED5B1389FFEC136AFDEFFE79FD97FE66" box="[574,607,458,479]" country="Italy" name="Collection of Leptospira Strains" pageId="10" pageNumber="129">RV</collectionCode>
<taxonomicName id="4C4AF0CFFFEC136AFDB9FE79FD2BFE66" box="[616,739,458,479]" class="Actinopterygii" family="Nemichthyidae" genus="Investigator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="10" pageNumber="129" phylum="Chordata" rank="genus">
<emphasis id="B93E575EFFEC136AFDB9FE79FD2BFE66" box="[616,739,458,479]" italics="true" pageId="10" pageNumber="129">Investigator</emphasis>
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<date id="FFF4AD8CFFEC136AFD38FE79FF3EFE44" pageId="10" pageNumber="129" value="2017-05-24">
<collectingDate id="EFB05464FFEC136AFD38FE79FF3EFE44" pageId="10" pageNumber="129" value="2017-05-24">24 May 2017</collectingDate>
</date>
, 1 wet spec.
<collectionCode id="ED5B1389FFEC136AFEA9FE5BFE42FE47" box="[376,394,488,510]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="129">R</collectionCode>
=5.4, r=1.3.
</materialsCitation>
</paragraph>
<paragraph id="8BF58B4CFFEC136AFF63FDB5FD01FDE9" blockId="10.[139,771,397,593]" pageId="10" pageNumber="129">
<materialsCitation id="3B228111FFEC136AFF63FDB5FF04FDE9" collectionCode="NMV" collectorName="Coll. O'Hara" country="Australia" elevation="3852" latitude="-38.479" location="East Gippsland" longLatPrecision="70" longitude="150.185" municipality="Tasman Sea" pageId="10" pageNumber="129" specimenCode="F 241807" specimenCount="1" stateProvince="Victoria" typeStatus="paratype">
<typeStatus id="54F135EEFFEC136AFF63FDB5FECFFDA0" box="[178,263,518,537]" pageId="10" pageNumber="129" type="paratype">Paratype</typeStatus>
.
<collectionCode id="ED5B1389FFEC136AFEDFFDB5FE8CFDA0" box="[270,324,518,537]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="10" pageNumber="129" type="Museum">NMV</collectionCode>
<specimenCode id="DBEC2337FFEC136AFE98FDB5FE55FDA0" box="[329,413,518,537]" collectionCode="NMV" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="10" pageNumber="129" type="Museum">F 241807</specimenCode>
<location id="8E95DD97FFEC136AFE72FDB5FDF8FDA0" LSID="urn:lsid:plazi:treatment:03E33A5AFFE0136AFCF8F974FD01FDE9:8E95DD97FFEC136AFE72FDB5FDF8FDA0" box="[419,560,518,537]" country="Australia" latitude="-38.479" longLatPrecision="70" longitude="150.185" municipality="Tasman Sea" name="East Gippsland" pageId="10" pageNumber="129" stateProvince="Victoria">East Gippsland</location>
,
<collectingRegion id="498E45AEFFEC136AFDE8FDB5FD4EFDA0" box="[569,646,518,537]" country="Australia" name="Victoria" pageId="10" pageNumber="129">Victoria</collectingRegion>
,
<collectingMunicipality id="6B911136FFEC136AFD5EFDB5FD37FDA0" box="[655,767,518,537]" pageId="10" pageNumber="129">Tasman Sea</collectingMunicipality>
,
<collectingCountry id="F35DCBDCFFEC136AFF5DFD92FF2BFD8D" box="[140,227,545,564]" name="Australia" pageId="10" pageNumber="129">Australia</collectingCountry>
, -
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,
<geoCoordinate id="EE7EED8BFFEC136AFE8BFD92FE09FD8C" box="[346,449,545,565]" degrees="150.185" direction="east" orientation="longitude" pageId="10" pageNumber="129" precision="55" value="150.185">150.185° E</geoCoordinate>
,
<quantity id="4CB226A9FFEC136AFE1AFD92FD83FD8C" box="[459,587,545,565]" metricMagnitude="3" metricUnit="m" metricValue="3.8515" metricValueMax="3.853" metricValueMin="3.85" pageId="10" pageNumber="129" unit="m" value="3851.5" valueMax="3853.0" valueMin="3850.0">
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</quantity>
,
<collectorName id="26BFEE9AFFEC136AFD87FD92FD06FD8C" box="[598,718,545,565]" pageId="10" pageNumber="129">Coll. O’Hara</collectorName>
et al. aboard
</materialsCitation>
<materialsCitation id="3B228111FFEC136AFF03FD8DFD84FDE8" box="[210,588,573,593]" collectingDate="2017-05-24" collectionCode="RV" country="Australia" pageId="10" pageNumber="129" specimenCount="1" stateProvince="Victoria" typeStatus="paratype">
<collectionCode id="ED5B1389FFEC136AFF03FD8DFF38FDE8" box="[210,240,574,593]" country="Italy" name="Collection of Leptospira Strains" pageId="10" pageNumber="129">RV</collectionCode>
<taxonomicName id="4C4AF0CFFFEC136AFF25FD8DFEABFDE9" box="[244,355,574,592]" class="Actinopterygii" family="Nemichthyidae" genus="Investigator" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anguilliformes" pageId="10" pageNumber="129" phylum="Chordata" rank="genus">
<emphasis id="B93E575EFFEC136AFF25FD8DFEABFDE9" box="[244,355,574,592]" italics="true" pageId="10" pageNumber="129">Investigator</emphasis>
</taxonomicName>
<date id="FFF4AD8CFFEC136AFEB8FD8EFE29FDE9" box="[361,481,573,593]" pageId="10" pageNumber="129" value="2017-05-24">
<collectingDate id="EFB05464FFEC136AFEB8FD8EFE29FDE9" box="[361,481,573,593]" pageId="10" pageNumber="129" value="2017-05-24">24 May 2017</collectingDate>
</date>
, 1 wet spec
</materialsCitation>
. R=5.7, r=1.3.
</paragraph>
</subSubSection>
</treatment>
</document>
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