445 lines
55 KiB
XML
445 lines
55 KiB
XML
<document id="687C7F4D7121DBFFC56480685838BB93" ID-CLB-Dataset="20368" ID-DOI="10.15553/c2020v752a1" ID-GBIF-Dataset="1b2f3915-6b17-4a42-9895-d8a11f37fd81" ID-ISSN="2235-3658" ID-Zenodo-Dep="5724844" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637765588461" checkinUser="carolina" docAuthor="Taylor, Charlotte M., Gereau, Roy E. & Schmidt, Heidi H." docDate="2020" docId="03DFC37DFFDEFFD6FFEEF8F683FD9052" docLanguage="en" docName="Candollea.75.2.159-182.pdf" docOrigin="Candollea 75 (2)" docStyle="DocumentStyle:1E5CC43F38D0DBC531513809D1267F7B.3:Candollea.2015-.journal_article" docStyleId="1E5CC43F38D0DBC531513809D1267F7B" docStyleName="Candollea.2015-.journal_article" docStyleVersion="3" docTitle="Psychotria eumachioides C. M. Taylor 2020, sp. nov." docType="treatment" docVersion="10" lastPageNumber="171" masterDocId="FFE6BB05FFD5FFDAFF9FFFDA8044963C" masterDocTitle="Some distinctive new species of Psychotria (Rubiaceae, Psychotrieae) from Madagascar" masterLastPageNumber="182" masterPageNumber="159" pageNumber="170" updateTime="1699470178107" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="9311CE3C9FEBF15DF3E846DFDF0A5CF1">Some distinctive new species of Psychotria (Rubiaceae, Psychotrieae) from Madagascar</mods:title>
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<mods:namePart id="2761EF53E422595EF7BDBE99353690CE">Taylor, Charlotte M.</mods:namePart>
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<mods:namePart id="58001D519239885179C662D9A4407A43">Gereau, Roy E.</mods:namePart>
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<mods:namePart id="D62C0F0A92DC9ECB207D99DF3CDBD76A">Schmidt, Heidi H.</mods:namePart>
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<mods:title id="8EC802394B1198F2A70DEFB1F7CD0F2A">Candollea</mods:title>
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<mods:part id="4A3D00215ADA937322CDF580118133C7">
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<mods:date id="8CE8B70F6D903F811C3E620AD51988AF">2020</mods:date>
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<treatment id="03DFC37DFFDEFFD6FFEEF8F683FD9052" ID-DOI="http://doi.org/10.5281/zenodo.6335854" ID-GBIF-Taxon="193878538" ID-Zenodo-Dep="6335854" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052" httpUri="http://treatment.plazi.org/id/03DFC37DFFDEFFD6FFEEF8F683FD9052" lastPageId="12" lastPageNumber="171" pageId="11" pageNumber="170">
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<subSubSection id="C36C21E0FFDEFFD1FFEEF8F682A19178" box="[113,741,1835,1861]" pageId="11" pageNumber="170" type="nomenclature">
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<paragraph id="8BC9726BFFDEFFD1FFEEF8F682A19178" blockId="11.[113,741,1835,1861]" box="[113,741,1835,1861]" pageId="11" pageNumber="170">
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<heading id="D081C507FFDEFFD1FFEEF8F682A19178" box="[113,741,1835,1861]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="170" reason="2">
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<taxonomicName id="4C7609E8FFDEFFD1FFEEF8F681B79179" ID-CoL="9V5GC" authority="C.M. Tayor" authorityName="C. M. Taylor" authorityYear="2020" box="[113,499,1836,1861]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="170" phylum="Tracheophyta" rank="species" species="eumachioides" status="sp. nov.">
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<emphasis id="B902AE79FFDEFFD1FFEEF8F6812B9178" bold="true" box="[113,367,1836,1860]" italics="true" pageId="11" pageNumber="170">Psychotria eumachioides</emphasis>
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C.M. Tayor
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</taxonomicName>
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,
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<emphasis id="B902AE79FFDEFFD1FE62F8F682159178" bold="true" box="[509,593,1836,1860]" pageId="11" pageNumber="170">
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<taxonomicNameLabel id="A2311302FFDEFFD1FE62F8F682159178" box="[509,593,1836,1860]" pageId="11" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
(
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<figureCitation id="134D6EEEFFDEFFD1FDFFF8F6829D9178" box="[608,729,1835,1860]" captionStart="Fig" captionStartId="4.[136,167,1737,1756]" captionTargetBox="[208,1410,296,1675]" captionTargetId="figure-18@4.[246,1346,300,1675]" captionTargetPageId="4" captionText="Fig. 1. – Psychotria ambatovensis C.M. Taylor: A. Flowering branch; B. Stem apex with bases of two petioles and stipules in process of dehiscence; C. Hypanthium and calyx limb of flower; D. Corolla at anthesis; E. Fruit; F. Cross-section of pyrene and seed. Psychotria eumachioides C.M. Taylor: G. Fruiting branch; H. Stem apex with stipule and basal part of two petioles; I. Portion of stem below leaves with two nodes, stipules (upper node), and stipule scar (lower node); J. Portion of inflorescence with three flowers from which corollas have fallen and one flower at anthesis; K. Cross-section of pyrene and seed. Psychotria notopleuroides C.M. Taylor: L. Flowering stem and part of basal prostrate stem; M. Portion of inflorescence with two flower buds and one flower at anthesis; N. Cross-section of pyrene and seed. [A–D: Andriamiarinoro 164; E–F: Antilahimena 6408; G, K: Rabehevitra 1059; H–J: De Block 1172; L: Schatz et al. 3666; M: Labat & Andrianjafy 3377; N: Zjhra & Hutcheon 378]" figureDoi="http://doi.org/10.5281/zenodo.5724846" httpUri="https://zenodo.org/record/5724846/files/figure.png" pageId="11" pageNumber="170">Fig. 1G–K</figureCitation>
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).
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C21E0FFDEFFD1FF08F88D85269773" pageId="11" pageNumber="170" type="materials_examined">
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<paragraph id="8BC9726BFFDEFFD1FF08F88D85269773" blockId="11.[151,762,1879,1935]" lastBlockId="11.[839,1450,279,335]" pageId="11" pageNumber="170">
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<materialsCitation id="3B1E7836FFDEFFD1FF08F88D849A9773" ID-GBIF-Occurrence="3697348399" collectingDate="2004-05-18" collectionCode="V" collectorName="Rabehevitra et al" country="Madagascar" latitude="-13.712777" location="Antsiranana" longLatPrecision="21" longitude="50.090275" pageId="11" pageNumber="170" specimenCode="MO-5899412" specimenCount="1" stateProvince="Reg. S AVA" typeStatus="holotype">
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<emphasis id="B902AE79FFDEFFD1FF08F88D817F91B3" bold="true" pageId="11" pageNumber="170">
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||
<typeStatus id="54CDCCC9FFDEFFD1FF08F88D81679153" box="[151,291,1879,1903]" pageId="11" pageNumber="170" type="holotype">Holotypus</typeStatus>
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:
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<collectingCountry id="F36132FBFFDEFFD1FEA1F88D81B19153" box="[318,501,1879,1903]" name="Madagascar" pageId="11" pageNumber="170">MADAGASCAR</collectingCountry>
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.
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<collectingRegion id="49B2BC89FFDEFFD1FD96F88D82DA9153" box="[521,670,1879,1903]" pageId="11" pageNumber="170">Reg. S AVA</collectingRegion>
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[Prov.
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<location id="8EA924B0FFDEFFD1FF08F8AD816B91B3" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFDEFFD1FF08F8AD816B91B3" box="[151,303,1911,1935]" country="Madagascar" latitude="-13.712777" longLatPrecision="21" longitude="50.090275" name="Antsiranana" pageId="11" pageNumber="170" stateProvince="Reg. S AVA">Antsiranana</location>
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]:
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</emphasis>
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<location id="8EA924B0FFDEFFD1FEDBF8AD81EF91B3" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFDEFFD1FEDBF8AD81EF91B3" box="[324,427,1911,1935]" country="Madagascar" latitude="-13.712777" longLatPrecision="21" longitude="50.090275" name="Vohemar" pageId="11" pageNumber="170" stateProvince="Reg. S AVA">Vohémar</location>
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,
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||
<location id="8EA924B0FFDEFFD1FE27F8AD827491B3" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFDEFFD1FE27F8AD827491B3" box="[440,560,1911,1935]" country="Madagascar" latitude="-13.712777" longLatPrecision="21" longitude="50.090275" name="Tsarabaria" pageId="11" pageNumber="170" stateProvince="Reg. S AVA">Tsarabaria</location>
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||
,
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||
<location id="8EA924B0FFDEFFD1FDA1F8AD82F391B3" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFDEFFD1FDA1F8AD82F391B3" box="[574,695,1911,1935]" country="Madagascar" latitude="-13.712777" longLatPrecision="21" longitude="50.090275" name="Manakana" pageId="11" pageNumber="170" stateProvince="Reg. S AVA">Manakana</location>
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||
,
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||
<location id="8EA924B0FFDEFFD1FD5AF8AD83B99713" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFDEFFD1FD5AF8AD83B99713" box="[709,1021,279,1935]" country="Madagascar" latitude="-13.712777" longLatPrecision="21" longitude="50.090275" name="foret d'Ambondrobe" pageId="11" pageNumber="170" stateProvince="Reg. S AVA">forêt d’Ambondrobe</location>
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,
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||
<geoCoordinate id="EE4214ACFFDEFFD1FB91FECD84C89713" box="[1038,1164,279,303]" degrees="13" direction="south" minutes="42" orientation="latitude" pageId="11" pageNumber="170" precision="15" seconds="46" value="-13.712777">13°42'46"S</geoCoordinate>
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<geoCoordinate id="EE4214ACFFDEFFD1FB07FECD85589713" box="[1176,1308,279,303]" degrees="50" direction="east" minutes="05" orientation="longitude" pageId="11" pageNumber="170" precision="15" seconds="25" value="50.090275">50°05'25"E</geoCoordinate>
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,
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<date id="FFC854ABFFDEFFD1FAB2FECD85E19713" box="[1325,1445,279,303]" pageId="11" pageNumber="170" value="2004-05-18">
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<collectingDate id="EF8CAD43FFDEFFD1FAB2FECD85E19713" box="[1325,1445,279,303]" pageId="11" pageNumber="170" value="2004-05-18">18.V.2004</collectingDate>
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</date>
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||
,
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||
<emphasis id="B902AE79FFDEFFD1FCD8FEED84759773" box="[839,1073,311,335]" italics="true" pageId="11" pageNumber="170">
|
||
<collectorName id="268317BDFFDEFFD1FCD8FEED83B39773" box="[839,1015,311,335]" pageId="11" pageNumber="170">Rabehevitra et al</collectorName>
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||
. 1059
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</emphasis>
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||
(
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<specimenCode id="DBD0DA10FFDEFFD1FBDFFEED84939773" box="[1088,1239,311,335]" pageId="11" pageNumber="170">MO-5899412</specimenCode>
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!;
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</materialsCitation>
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<materialsCitation id="3B1E7836FFDEFFD1FB7AFEED85269773" ID-GBIF-Occurrence="3697348340" box="[1253,1378,311,335]" collectionCode="TAN" latitude="-13.712777" longLatPrecision="21" longitude="50.090275" pageId="11" pageNumber="170" specimenCount="1" typeStatus="isotype">
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<typeStatus id="54CDCCC9FFDEFFD1FB7AFEED85549773" box="[1253,1296,311,335]" pageId="11" pageNumber="170" type="isotype">isotype</typeStatus>
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:
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<collectionCode id="ED67EAAEFFDEFFD1FA86FEED85129773" box="[1305,1366,311,335]" pageId="11" pageNumber="170">TAN</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C21E0FFDEFFD6FCD8FEB8818597B3" lastPageId="12" lastPageNumber="171" pageId="11" pageNumber="170" type="description">
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<paragraph id="8BC9726BFFDEFFD1FCD8FEB885E997E6" blockId="11.[839,1453,354,474]" pageId="11" pageNumber="170">
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<emphasis id="B902AE79FFDEFFD1FCD8FEB885E997E6" italics="true" pageId="11" pageNumber="170">
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<taxonomicName id="4C7609E8FFDEFFD1FCD8FEB885499746" authority="C. M. Taylor" authorityName="C. M. Taylor" authorityYear="2020" box="[839,1293,354,378]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="170" phylum="Tracheophyta" rank="species" species="eumachioides">Psychotria eumachioides C.M. Taylor</taxonomicName>
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differs f rom
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<taxonomicName id="4C7609E8FFDEFFD1FCD8FE58852997A5" authority="(Bremek.) Razafim. & B. Bremer" authorityName="Razafim. & B. Bremer" baseAuthorityName="Bremek." box="[839,1389,386,410]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="170" phylum="Tracheophyta" rank="species" species="rubropedicellata">P. rubropedicellata (Bremek.) Razafim. & B. Bremer</taxonomicName>
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by its stems that become yellowed and indurated with age, narrower oblanceolate leaves that lack domatia, and shorter inflorescences.
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</emphasis>
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</paragraph>
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<paragraph id="8BC9726BFFDEFFD6FCD8FDD8818597B3" blockId="11.[801,1458,514,1946]" lastBlockId="12.[136,787,278,399]" lastPageId="12" lastPageNumber="171" pageId="11" pageNumber="170">
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<emphasis id="B902AE79FFDEFFD1FCD8FDD883CF9426" box="[839,907,514,538]" italics="true" pageId="11" pageNumber="170">Shrubs</emphasis>
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, at least sometimes apparently deciduous, collected in flower and fruit variously at
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<quantity id="4C8EDF8EFFDEFFD1FBE4FDFB84979405" box="[1147,1235,545,570]" metricMagnitude="0" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="11" pageNumber="170" unit="m" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 m</quantity>
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tall, branched with lateral stems usually comprising 5 to 10 nodes separated by reduced internodes; stems glabrous, weakly flattened becoming subterete, with bark often becoming yellowed or brown and thickened with age.
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<emphasis id="B902AE79FFDEFFD1FC6EFD7884719485" box="[1009,1077,674,697]" italics="true" pageId="11" pageNumber="170">Leaves</emphasis>
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opposite;
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<emphasis id="B902AE79FFDEFFD1FB00FD7884999486" box="[1183,1245,674,698]" italics="true" pageId="11" pageNumber="170">petiole</emphasis>
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<quantity id="4C8EDF8EFFDEFFD1FB7BFD78850D9485" box="[1252,1353,673,698]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.8" metricValueMin="0.4" pageId="11" pageNumber="170" unit="mm" value="16.0" valueMax="28.0" valueMin="4.0">4–28 mm</quantity>
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, glabrous or moderately to densely puberulous or pilosulous with trichomes
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<quantity id="4C8EDF8EFFDEFFD1FCE5FD3883BA94C5" box="[890,1022,737,762]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="11" pageNumber="170" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.1–0.3 mm</quantity>
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;
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<emphasis id="B902AE79FFDEFFD1FB96FD38847A94C6" box="[1033,1086,738,762]" italics="true" pageId="11" pageNumber="170">blade</emphasis>
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oblanceolate, 2.8–9 ×
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<quantity id="4C8EDF8EFFDEFFD1FAB0FD3B85EA94C5" box="[1327,1454,737,762]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="3.2" metricValueMin="0.5" pageId="11" pageNumber="170" unit="cm" value="1.85" valueMax="3.2" valueMin="0.5">0.5–3.2 cm</quantity>
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, at base tapered and acute, at apex broadly obtuse to rounded, drying papyraceous, on both surfaces glabrous or moderately puberulous to pilosulous with trichomes
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<quantity id="4C8EDF8EFFDEFFD1FB52FC98850A9565" box="[1229,1358,833,858]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="11" pageNumber="170" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.1–0.3 mm</quantity>
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; secondary veins 7 to 12 pairs, weakly looping to interconnect or reticulating near margins, without domatia, without intersecondary veins, adaxially venation plane or costa thinly impressed, secondary veins thickened, and remaining venation not evident, abaxially costa prominent, secondary veins plane to thickened, and remaining venation not evident.
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<emphasis id="B902AE79FFDEFFD1FB2EFBD885459226" box="[1201,1281,1026,1050]" italics="true" pageId="11" pageNumber="170">Stipules</emphasis>
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interpetiolar or shortly fused around stem, deciduous leaving thickened scar, ovate to triangular,
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<quantity id="4C8EDF8EFFDEFFD1FC70FB98840D9265" box="[1007,1097,1089,1114]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="11" pageNumber="170" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
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, abaxially glabrous or moderately pilosulous with trichomes
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<quantity id="4C8EDF8EFFDEFFD1FBD0FBB8849F9245" box="[1103,1243,1121,1146]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="11" pageNumber="170" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.1 – 0.3 mm</quantity>
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, adaxially sparsely pilosulous with trichomes
|
||
<quantity id="4C8EDF8EFFDEFFD1FBDBFB58848892A5" box="[1092,1228,1153,1178]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="11" pageNumber="170" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.1–0.3 mm</quantity>
|
||
, acute to acuminate, at tip sometimes glandular or bifid with lobes
|
||
<quantity id="4C8EDF8EFFDEFFD1FA80FB7885EB9285" box="[1311,1455,1185,1210]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="11" pageNumber="170" unit="mm" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 mm</quantity>
|
||
.
|
||
<emphasis id="B902AE79FFDEFFD1FCBEFB1883F592E6" box="[801,945,1218,1242]" italics="true" pageId="11" pageNumber="170">Inflorescences</emphasis>
|
||
terminal, cymose, pedunculate, sparsely to moderately pilosulous with trichomes
|
||
<quantity id="4C8EDF8EFFDEFFD1FB24FB38857B92C5" box="[1211,1343,1249,1274]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="11" pageNumber="170" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.1–0.3 mm</quantity>
|
||
; peduncle
|
||
<quantity id="4C8EDF8EFFDEFFD1FCBEFADB83C99325" box="[801,909,1281,1306]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="11" pageNumber="170" unit="cm" value="0.75" valueMax="1.0" valueMin="0.5">0.5– 1 cm</quantity>
|
||
; flower-bearing portion rounded-corymbiform, 1–1.5 ×
|
||
<quantity id="4C8EDF8EFFDEFFD1FCEAFAF883819305" box="[885,965,1313,1338]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="11" pageNumber="170" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
|
||
, branched to 1 or 2 orders, c. 10- to 25-flowered; bracts reduced or usually triangular,
|
||
<quantity id="4C8EDF8EFFDEFFD1FB7FFA9885269365" box="[1248,1378,1345,1370]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="11" pageNumber="170" unit="mm" value="0.3" valueMax="0.5" valueMin="0.1">0.1–0.5 mm</quantity>
|
||
, acute; pedicels
|
||
<quantity id="4C8EDF8EFFDEFFD1FC1CFAB883BF9345" box="[899,1019,1377,1402]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="11" pageNumber="170" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1 – 1.5 mm</quantity>
|
||
.
|
||
<emphasis id="B902AE79FFDEFFD1FB9FFAB884119346" box="[1024,1109,1378,1402]" italics="true" pageId="11" pageNumber="170">Flowers</emphasis>
|
||
all pedicellate in umbelliform groups of 5 to 7, 5-merous; hypanthium cylindrical to ellipsoid, c.
|
||
<quantity id="4C8EDF8EFFDEFFD1FCA6FA7883C89385" box="[825,908,1442,1466]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="170" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, moderately pilosulous with trichomes
|
||
<quantity id="4C8EDF8EFFDEFFD1FAB9FA7885E39385" box="[1318,1447,1441,1466]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="11" pageNumber="170" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.1–0.3 mm</quantity>
|
||
;
|
||
<emphasis id="B902AE79FFDEFFD1FCBEFA18831093E6" box="[801,852,1474,1498]" italics="true" pageId="11" pageNumber="170">calyx</emphasis>
|
||
limb c.
|
||
<quantity id="4C8EDF8EFFDEFFD1FC32FA18844793E5" box="[941,1027,1474,1498]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="170" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, externally moderately to sparsely pilosulous with trichomes
|
||
<quantity id="4C8EDF8EFFDEFFD1FBBBFA3884F493C5" box="[1060,1200,1505,1530]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="11" pageNumber="170" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.1 – 0.3 mm</quantity>
|
||
, lobed for 1/3– 1/2 of its length, lobes ovate, acute to shortly acuminate;
|
||
<emphasis id="B902AE79FFDEFFD1FAAFF9D885379026" box="[1328,1395,1538,1562]" italics="true" pageId="11" pageNumber="170">corolla</emphasis>
|
||
funnelform, yellow, externally sparsely pilosulous with trichomes
|
||
<quantity id="4C8EDF8EFFDEFFD1FCBEF99883E89065" box="[801,940,1601,1626]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="11" pageNumber="170" unit="mm" value="0.2" valueMax="0.3" valueMin="0.1">0.1– 0.3 mm</quantity>
|
||
, tube c.
|
||
<quantity id="4C8EDF8EFFDEFFD1FB90F99884159065" box="[1039,1105,1602,1626]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="170" unit="mm" value="3.0">3 mm</quantity>
|
||
, c.
|
||
<quantity id="4C8EDF8EFFDEFFD1FBE5F998848A9065" box="[1146,1230,1602,1626]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="170" unit="mm" value="1.2">1.2 mm</quantity>
|
||
diam. near middle, internally glabrous except densely hirtellous in upper part of tube, lobes narrowly triangular, c.
|
||
<quantity id="4C8EDF8EFFDEFFD1FB34F95884AF90A5" box="[1195,1259,1666,1690]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="170" unit="mm" value="2.0">2 mm</quantity>
|
||
, adaxially galeate, abaxially with rounded thickening near tip;
|
||
<emphasis id="B902AE79FFDEFFD1FB6CF97885059085" box="[1267,1345,1698,1721]" italics="true" pageId="11" pageNumber="170">stamens</emphasis>
|
||
with filaments c.
|
||
<quantity id="4C8EDF8EFFDEFFD1FCE0F91B838990E5" box="[895,973,1729,1753]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="170" unit="mm" value="1.5">1.5 mm</quantity>
|
||
, inserted in upper part of corolla tube, anthers c.
|
||
<quantity id="4C8EDF8EFFDEFFD1FCA6F93883CD90C5" box="[825,905,1762,1786]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="170" unit="mm" value="1.2">1.2 mm</quantity>
|
||
, exserted;
|
||
<emphasis id="B902AE79FFDEFFD1FC69F938846590C6" box="[1014,1057,1762,1786]" italics="true" pageId="11" pageNumber="170">style</emphasis>
|
||
c.
|
||
<quantity id="4C8EDF8EFFDEFFD1FBDFF938843B90C5" box="[1088,1151,1762,1786]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="170" unit="mm" value="2.0">2 mm</quantity>
|
||
,
|
||
<emphasis id="B902AE79FFDEFFD1FB16F938849090C5" box="[1161,1236,1762,1785]" italics="true" pageId="11" pageNumber="170">stigmas</emphasis>
|
||
c.
|
||
<quantity id="4C8EDF8EFFDEFFD1FB6DF938850190C5" box="[1266,1349,1762,1786]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="170" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, included and positioned above pubescent internal ring of corolla tube.
|
||
<emphasis id="B902AE79FFDEFFD1FCBEF8F883F89106" box="[801,956,1826,1850]" italics="true" pageId="11" pageNumber="170">Infructescences</emphasis>
|
||
similar to inflorescences except sometimes displaced to pseudoaxillary by subsequent stem growth from an axillary bud, and with axes elongating and deciduous as fruits fail to develop or fall off; pedicels elongating, to
|
||
<quantity id="4C8EDF8EFFDEFFD1FAF8F85B85E391A5" box="[1383,1447,1921,1945]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="170" unit="mm" value="6.0">6 mm</quantity>
|
||
, becoming fleshy.
|
||
<emphasis id="B902AE79FFD9FFD6FEA3FEC2813E9713" box="[316,378,280,303]" italics="true" pageId="12" pageNumber="171">Fruits</emphasis>
|
||
didymous, c. 5 ×
|
||
<quantity id="4C8EDF8EFFD9FFD6FDAEFECD822B9712" box="[561,623,279,303]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="171" unit="mm" value="8.0">8 mm</quantity>
|
||
, glabrous, juicy, orange to red;
|
||
<emphasis id="B902AE79FFD9FFD6FEBBFEE281299773" box="[292,365,312,335]" italics="true" pageId="12" pageNumber="171">pyrenes</emphasis>
|
||
subglobose, abaxially smooth, adaxially plane;
|
||
<emphasis id="B902AE79FFD9FFD6FF51FE8D81739753" box="[206,311,343,367]" italics="true" pageId="12" pageNumber="171">endosperm</emphasis>
|
||
abaxially entire (i.e., not ruminate), adaxially with a few deep ruminations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C21E0FFD9FFD6FF31FE6D82A09492" pageId="12" pageNumber="171" type="etymology">
|
||
<paragraph id="8BC9726BFFD9FFD6FF31FE6D82A09492" blockId="12.[136,787,439,687]" pageId="12" pageNumber="171">
|
||
<emphasis id="B902AE79FFD9FFD6FF31FE6D815997F3" box="[174,285,439,463]" italics="true" pageId="12" pageNumber="171">Etymology</emphasis>
|
||
. – The species epithet refers to the similarity of this new species to some species of
|
||
<taxonomicName id="4C7609E8FFD9FFD6FDBFFE0D82C397D3" box="[544,647,471,495]" class="Magnoliopsida" family="Rubiaceae" genus="Eumachia" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B902AE79FFD9FFD6FDBFFE0D82C397D3" box="[544,647,471,495]" italics="true" pageId="12" pageNumber="171">Eumachia</emphasis>
|
||
</taxonomicName>
|
||
, as discussed below.
|
||
<taxonomicName id="4C7609E8FFD9FFD6FF50FE2D81709433" box="[207,308,503,527]" class="Magnoliopsida" family="Rubiaceae" genus="Eumachia" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B902AE79FFD9FFD6FF50FE2D81709433" box="[207,308,503,527]" italics="true" pageId="12" pageNumber="171">Eumachia</emphasis>
|
||
</taxonomicName>
|
||
has been commonly confused with
|
||
<taxonomicName id="4C7609E8FFD9FFD6FD33FE2D83569433" box="[684,786,503,527]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B902AE79FFD9FFD6FD33FE2D83569433" box="[684,786,503,527]" italics="true" pageId="12" pageNumber="171">Psychotria</emphasis>
|
||
</taxonomicName>
|
||
but these are not closely related (
|
||
<bibRefCitation id="EFE70F9AFFD9FFD6FE46FDCD82DA9413" author="TAYLOR, C. M. & S. G. RAZAFIMANDIMBISON & L. BARRABE & J. G. JARDIM & M. R. V. BARBOSA" box="[473,670,535,559]" journalOrPublisher="Candollea" pageId="12" pageNumber="171" pagination="289 - 318" part="72" refId="ref23444" refString="TAYLOR, C. M., S. G. RAZAFIMANDIMBISON, L. BARRABE, J. G. JARDIM & M. R. V. BARBOSA (2017). Eumachia expanded, a pantropical genus distinct from Psychotria (Rubiaceae, Palicoureeae). Candollea 72: 289 - 318." title="Eumachia expanded, a pantropical genus distinct from Psychotria (Rubiaceae, Palicoureeae)" type="journal article" year="2017">TAYLOR et al., 2017</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C7609E8FFD9FFD6FD31FDCD83569413" box="[686,786,535,559]" class="Magnoliopsida" family="Rubiaceae" genus="Eumachia" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B902AE79FFD9FFD6FD31FDCD83569413" box="[686,786,535,559]" italics="true" pageId="12" pageNumber="171">Eumachia</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C7609E8FFD9FFD6FE93FDED81379473" box="[268,371,567,591]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B902AE79FFD9FFD6FE93FDED81379473" box="[268,371,567,591]" italics="true" pageId="12" pageNumber="171">Psychotria</emphasis>
|
||
</taxonomicName>
|
||
in its stipules that are generally persistent and have one or two gland-tipped lobes, a lack of persistent colleters at the stipule insertion, and molecular sequences that indicate its relationships are in another tribe,
|
||
<taxonomicName id="4C7609E8FFD9FFD6FDFBFD4D82999493" authorityName="Robbr. & Manen" authorityYear="2006" box="[612,733,663,687]" class="Magnoliopsida" family="Rubiaceae" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="tribe" tribe="Palicoureeae">
|
||
<emphasis id="B902AE79FFD9FFD6FDFBFD4D82999493" box="[612,733,663,687]" italics="true" pageId="12" pageNumber="171">Palicoureeae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C21E0FFD9FFD6FF31FD0D835095B3" pageId="12" pageNumber="171" type="multiple">
|
||
<paragraph id="8BC9726BFFD9FFD6FF31FD0D835095B3" blockId="12.[136,788,727,911]" pageId="12" pageNumber="171">
|
||
<emphasis id="B902AE79FFD9FFD6FF31FD0D824894D3" box="[174,524,727,751]" italics="true" pageId="12" pageNumber="171">Habitat, distribution and phenology</emphasis>
|
||
. –
|
||
<taxonomicName id="4C7609E8FFD9FFD6FDB5FD0D835694D3" authorityName="C. M. Taylor" authorityYear="2020" box="[554,786,727,751]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="species" species="eumachioides">
|
||
<emphasis id="B902AE79FFD9FFD6FDB5FD0D835694D3" box="[554,786,727,751]" italics="true" pageId="12" pageNumber="171">Psychotria eumachioides</emphasis>
|
||
</taxonomicName>
|
||
has been collected in gallery forest in subhumid or dry bioclimate zones (
|
||
<bibRefCitation id="EFE70F9AFFD9FFD6FE8FFCCD81EC9513" author="CORNET" box="[272,424,791,815]" firstAuthor="CORNET" pageId="12" pageNumber="171" refId="ref23327" refString="CORNET, A. (1974). Essai de cartographie bioclimatique a Madagascar. Notic. Explic. ORSTOM 55." type="journal volume" year="1974">CORNET, 1974</bibRefCitation>
|
||
), from near sea level to c.
|
||
<quantity id="4C8EDF8EFFD9FFD6FD21FCCD82B19512" box="[702,757,791,815]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="171" unit="m" value="40.0">
|
||
<elevation id="005B9558FFD9FFD6FD21FCCD82B19512" box="[702,757,791,815]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="171" unit="m" value="40.0">40 m</elevation>
|
||
</quantity>
|
||
in northwestern
|
||
<collectingCountry id="F36132FBFFD9FFD6FE81FCED81DB9573" box="[286,415,823,847]" name="Madagascar" pageId="12" pageNumber="171">Madagascar</collectingCountry>
|
||
(
|
||
<collectingRegion id="49B2BC89FFD9FFD6FE32FCED82709573" box="[429,564,823,847]" country="Madagascar" name="Antsiranana" pageId="12" pageNumber="171">Antsiranana</collectingRegion>
|
||
), variously in littoral forest and on lakeshores and river banks.
|
||
<collectingCounty id="62A80AE7FFD9FFD6FDA5FC8D82089552" box="[570,588,855,878]" pageId="12" pageNumber="171">It</collectingCounty>
|
||
has been collected with flowers in
|
||
<collectingMunicipality id="6BADE811FFD9FFD6FEB1FCAD813A95B3" box="[302,382,887,911]" pageId="12" pageNumber="171">January</collectingMunicipality>
|
||
, and with fruits in
|
||
<location id="8EA924B0FFD9FFD6FDD6FCAD82ED95B3" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFD9FFD6FDD6FCAD82ED95B3" box="[585,681,887,911]" country="Madagascar" county="It" municipality="January" name="February" pageId="12" pageNumber="171" stateProvince="Antsiranana">February</location>
|
||
and
|
||
<location id="8EA924B0FFD9FFD6FD42FCAD834B95B3" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFD9FFD6FD42FCAD834B95B3" box="[733,783,887,911]" country="Madagascar" county="It" municipality="January" name="May" pageId="12" pageNumber="171" stateProvince="Antsiranana">May</location>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C21E0FFD9FFD6FF31FC6D83519173" pageId="12" pageNumber="171" type="conservation">
|
||
<paragraph id="8BC9726BFFD9FFD6FF31FC6D83519173" blockId="12.[136,789,951,1871]" pageId="12" pageNumber="171">
|
||
<emphasis id="B902AE79FFD9FFD6FF31FC6D812B95F3" box="[174,367,951,975]" italics="true" pageId="12" pageNumber="171">Conservation status</emphasis>
|
||
. –
|
||
<taxonomicName id="4C7609E8FFD9FFD6FE12FC6D823295F3" authorityName="C. M. Taylor" authorityYear="2020" box="[397,630,951,975]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="species" species="eumachioides">
|
||
<emphasis id="B902AE79FFD9FFD6FE12FC6D823295F3" box="[397,630,951,975]" italics="true" pageId="12" pageNumber="171">Psychotria eumachioides</emphasis>
|
||
</taxonomicName>
|
||
is known from
|
||
<specimenCount id="9D70B9E2FFD9FFD6FF17FC0D815F95D3" box="[136,283,983,1007]" pageId="12" pageNumber="171" type="generic">four specimen</specimenCount>
|
||
collections representing four unique occurrences in gallery forest in subhumid or dry bioclimatic zones at
|
||
<quantity id="4C8EDF8EFFD9FFD6FF17FBCD80989212" box="[136,220,1046,1071]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="4.0" metricValueMin="0.0" pageId="12" pageNumber="171" unit="m" value="20.0" valueMax="40.0" valueMin="0.0">0–40 m</quantity>
|
||
elevation. The EOO of the species is
|
||
<quantity id="4C8EDF8EFFD9FFD6FDF5FBCD82FC9213" box="[618,696,1047,1071]" metricMagnitude="5" metricUnit="m" metricValue="8.959999999999999" pageId="12" pageNumber="171" unit="km" value="896.0">896 km</quantity>
|
||
², within the limits for “Endangered” under IUCN Red List Criterion B1; and the AOO is
|
||
<quantity id="4C8EDF8EFFD9FFD6FEC0FB8D81DA9253" box="[351,414,1111,1135]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="171" unit="km" value="12.0">12 km</quantity>
|
||
², also within the limits for “Endangered” under Criterion B2 (
|
||
<bibRefCitation id="EFE70F9AFFD9FFD6FE38FBAD826A92B3" author="IUCN" box="[423,558,1143,1167]" firstAuthor="IUCN" pageId="12" pageNumber="171" refId="ref23474" refString="IUCN (2012). IUCN Red List Categories and Criteria. Version 3.1. Ed. 2. Gland & Cambridge." type="book" year="2012">IUCN, 2012</bibRefCitation>
|
||
). Two occurrences are within the generally well-protected Loky-Manambato PA, though there are reports of some artisanal gold mining within the protected area (C. Birkinshaw, pers. comm.). A third occurrence is near the northern boundary of, but distinctly outside, Analamerana PA, and the remaining occurrence is not in a formally recognized protected area, although the label indicates “forêt d’Ambondrobe”. Forested areas without formal protection in the vicinity of Loky-Manambato PA are subject to degradation by small-scale slash and burn agriculture and resource exploitation including logging, hunting and mining (
|
||
<bibRefCitation id="EFE70F9AFFD9FFD6FF0EFA0D81C993D3" author="GOODMAN" box="[145,397,1495,1519]" etAl="et al." firstAuthor="GOODMAN" pageId="12" pageNumber="171" refId="ref23413" refString="GOODMAN, S. M., M. J. RAHERILALAO & S. WOHLHAUSER (ed.) (2018). Les aires protegees terrestres de Madagascar: leur histoire, description et biote / The terrestrial protected areas of Madagascar: their history, description, and biota. 3 vol. Association Vahatra, Antananarivo." type="book" year="2018">GOODMAN et al., 2018</bibRefCitation>
|
||
). The northernmost collection site outside Analamerana PA is stated to be in dry, semi-deciduous, very degraded forest, and it constitutes one “location” (sensu
|
||
<bibRefCitation id="EFE70F9AFFD9FFD6FF17F9ED81569073" author="IUCN" box="[136,274,1591,1615]" firstAuthor="IUCN" pageId="12" pageNumber="171" refId="ref23474" refString="IUCN (2012). IUCN Red List Categories and Criteria. Version 3.1. Ed. 2. Gland & Cambridge." type="book" year="2012">IUCN, 2012</bibRefCitation>
|
||
) with respect to the known threats. This is the northernmost location and its disappearance would significantly reduce both EOO and AOO. The two occurrences within Loky-Manambato constitute a second location, and the single occurrence at Ambondrobe Forest is a third location. With only three locations, documented degradation at the northernmost location, and general habitat degradation in unprotected sites, the Red List status of
|
||
<taxonomicName id="4C7609E8FFD9FFD6FDC1F8C282BE9113" authorityName="C. M. Taylor" authorityYear="2020" box="[606,762,1815,1839]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="species" species="eumachioides">
|
||
<emphasis id="B902AE79FFD9FFD6FDC1F8C282BE9113" box="[606,762,1815,1839]" italics="true" pageId="12" pageNumber="171">P. eumachioides</emphasis>
|
||
</taxonomicName>
|
||
is assessed as “Endangered” [EN B1ab(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v)].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C21E0FFD9FFD6FCC1FEC285F79393" pageId="12" pageNumber="171" type="discussion">
|
||
<paragraph id="8BC9726BFFD9FFD6FCC1FEC2858192D3" blockId="12.[824,1477,279,1455]" pageId="12" pageNumber="171">
|
||
<emphasis id="B902AE79FFD9FFD6FCC1FEC284849713" box="[862,1216,279,303]" italics="true" pageId="12" pageNumber="171">
|
||
Notes. –
|
||
<taxonomicName id="4C7609E8FFD9FFD6FC20FECD84849713" authorityName="C. M. Taylor" authorityYear="2020" box="[959,1216,279,303]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="species" species="eumachioides">Psychotria eumachioides</taxonomicName>
|
||
</emphasis>
|
||
is characterized by its stems with shortened internodes and an epidermis that often becomes yellowed and indurated with age; medium-sized oblanceolate leaves that are obtuse to rounded at the apex and lack domatia; triangular to ovate, acute, medium-sized stipules that are deciduous and leave a thickened scar; corymbiform several-flowered inflorescences that are overtopped by the leaves; flowers that are pedicellate in umbelliform cymes; developed calyx limbs; somewhat small yellow corollas; orange to red fruits borne on thickened pedicels; smooth subglobose pyrenes; and endosperm with several adaxial ruminations. The plants dry with a generally green color. The dried trichomes are mostly white, though some trichomes on old stipules turned brown. The stems do have developd internodes but these are quite short, so the growth pattern can almost be considered a brachyblast and its arrangement is accentuated by thickened petiole and stipule scars. This growth pattern may be related to the habitat of this species, which lives in relatively dry areas and is apparently deciduous.
|
||
<taxonomicName id="4C7609E8FFD9FFD6FBEBFC8D852C9553" authorityName="C. M. Taylor" authorityYear="2020" box="[1140,1384,855,879]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="species" species="eumachioides">
|
||
<emphasis id="B902AE79FFD9FFD6FBEBFC8D852C9553" box="[1140,1384,855,879]" italics="true" pageId="12" pageNumber="171">Psychotria eumachioides</emphasis>
|
||
</taxonomicName>
|
||
appears to produce flowers and new leaves together, then the fruit and stem development proceed concurrently. The few flowers seen resemble the short-styled form of distylous
|
||
<taxonomicName id="4C7609E8FFD9FFD6FAC8FC6D83C695D3" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="B902AE79FFD9FFD6FAC8FC6D858695F3" box="[1367,1474,951,975]" italics="true" pageId="12" pageNumber="171">Psychotria</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, but whether this species is distylous is not yet known. A fruiting specimen is chosen as the
|
||
<typeStatus id="54CDCCC9FFD9FFD6FB47FC2D85439233" box="[1240,1287,1015,1039]" pageId="12" pageNumber="171">type</typeStatus>
|
||
because most of characters that diagnose this new species are in the fruits and seeds.
|
||
<taxonomicName id="4C7609E8FFD9FFD6FCE6FBED84219273" authorityName="C. M. Taylor" authorityYear="2020" box="[889,1125,1079,1103]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="species" species="eumachioides">
|
||
<emphasis id="B902AE79FFD9FFD6FCE6FBED84219273" box="[889,1125,1079,1103]" italics="true" pageId="12" pageNumber="171">Psychotria eumachioides</emphasis>
|
||
</taxonomicName>
|
||
agrees with
|
||
<bibRefCitation id="EFE70F9AFFD9FFD6FB79FBED85F29273" author="BREMEKAMP" box="[1254,1462,1079,1103]" firstAuthor="BREMEKAMP" pageId="12" pageNumber="171" pagination="5" refId="ref23275" refString="BREMEKAMP, C. E. B. (1963). Sur quelques genres de Psychotriees (Rubiacees) et sur leurs representants malgaches et comoriens. Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Tweede Sect. 54 (5)." type="book chapter" year="1963">BREMEKAMP (1963)</bibRefCitation>
|
||
’s
|
||
<taxonomicName id="4C7609E8FFD9FFD6FCA7FB8283DA9253" authorityName="Aublet" authorityYear="1775" box="[824,926,1112,1135]" class="Magnoliopsida" family="Rubiaceae" genus="Mapouria" higherTaxonomySource="GBIF,IPNI" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B902AE79FFD9FFD6FCA7FB8283DA9253" box="[824,926,1112,1135]" italics="true" pageId="12" pageNumber="171">Mapouria</emphasis>
|
||
</taxonomicName>
|
||
, and keys there to either Group VI or Group V but does not fully agree with either. It agrees with Group IV in its lack of domatia, but differs in its fruits that are not blue or white at maturity; it agrees with Group V in its yellow to red fruits, but differs in its lack of domatia and entire stipules.
|
||
</paragraph>
|
||
<paragraph id="8BC9726BFFD9FFD6FCC1FB2D85F79393" blockId="12.[824,1477,279,1455]" pageId="12" pageNumber="171">
|
||
<taxonomicName id="4C7609E8FFD9FFD6FCC1FB2D84019333" authorityName="C. M. Taylor" authorityYear="2020" box="[862,1093,1271,1295]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="species" species="eumachioides">
|
||
<emphasis id="B902AE79FFD9FFD6FCC1FB2D84019333" box="[862,1093,1271,1295]" italics="true" pageId="12" pageNumber="171">Psychotria eumachioides</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C7609E8FFD9FFD6FB57FB22853C9333" box="[1224,1400,1271,1295]" class="Magnoliopsida" family="Rubiaceae" genus="Psychotria" kingdom="Plantae" order="Gentianales" pageId="12" pageNumber="171" phylum="Tracheophyta" rank="species" species="rubropedicellata">
|
||
<emphasis id="B902AE79FFD9FFD6FB57FB22853C9333" box="[1224,1400,1271,1295]" italics="true" pageId="12" pageNumber="171">P. rubropedicellata</emphasis>
|
||
</taxonomicName>
|
||
, which has stems with regularly developed, elongated internodes, larger leaf blades 6.5–16 ×
|
||
<quantity id="4C8EDF8EFFD9FFD6FB95FAED84219372" box="[1034,1125,1334,1359]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="10.0" metricValueMin="2.0" pageId="12" pageNumber="171" unit="cm" value="6.0" valueMax="10.0" valueMin="2.0">2–10 cm</quantity>
|
||
with pilosulous domatia and tips that are sharply obtuse to acute, stipules with two linear lobes, often larger inflorescences
|
||
<quantity id="4C8EDF8EFFD9FFD6FBFEFAAD84F793B2" box="[1121,1203,1398,1423]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="12" pageNumber="171" unit="cm" value="4.5" valueMax="7.0" valueMin="2.0">2–7 cm</quantity>
|
||
long, and usually longer fruiting pedicels,
|
||
<quantity id="4C8EDF8EFFD9FFD6FC71FA4D84149392" box="[1006,1104,1430,1455]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="12" pageNumber="171" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">5–12 mm</quantity>
|
||
long, that become fleshy and red.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C21E0FFD9FFD6FCC1FA0C83FD9052" pageId="12" pageNumber="171" type="materials_examined">
|
||
<paragraph id="8BC9726BFFD9FFD6FCC1FA0C83FD9052" blockId="12.[824,1475,1493,1646]" pageId="12" pageNumber="171">
|
||
<materialsCitation id="3B1E7836FFD9FFD6FCC1FA0C84C69022" ID-GBIF-Occurrence="3697348318" collectingDate="2003-02-02" collectionCode="BR, MO, P, TAN" collectorName="De Block" country="Madagascar" county="Analamera" elevation="41" latitude="-12.673612" location="Antsiranana" longLatPrecision="21" longitude="49.544445" municipality="Irodo" pageId="12" pageNumber="171" specimenCount="1" stateProvince="Reg. Sava" typeStatus="paratype">
|
||
<emphasis id="B902AE79FFD9FFD6FCC1FA0C83E193D4" box="[862,933,1494,1512]" italics="true" pageId="12" pageNumber="171">
|
||
<typeStatus id="54CDCCC9FFD9FFD6FCC1FA0C83E193D4" box="[862,933,1494,1512]" pageId="12" pageNumber="171" type="paratype">Paratypi</typeStatus>
|
||
</emphasis>
|
||
. –
|
||
<emphasis id="B902AE79FFD9FFD6FC20FA0F847F93D4" bold="true" box="[959,1083,1493,1512]" pageId="12" pageNumber="171">
|
||
<collectingCountry id="F36132FBFFD9FFD6FC20FA0F847F93D4" box="[959,1083,1493,1512]" name="Madagascar" pageId="12" pageNumber="171">MADAGASCAR</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B902AE79FFD9FFD6FBDAFA0F851E93D4" bold="true" box="[1093,1370,1493,1512]" pageId="12" pageNumber="171">
|
||
<collectingRegion id="49B2BC89FFD9FFD6FBDAFA0F84D993D4" box="[1093,1181,1493,1512]" pageId="12" pageNumber="171">Reg. Sava</collectingRegion>
|
||
[Prov.
|
||
<location id="8EA924B0FFD9FFD6FB43FA0F851493D4" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFD9FFD6FB43FA0F851493D4" box="[1244,1360,1493,1512]" country="Madagascar" county="Analamera" latitude="-12.673612" longLatPrecision="21" longitude="49.544445" municipality="Irodo" name="Antsiranana" pageId="12" pageNumber="171" stateProvince="Reg. Sava">Antsiranana</location>
|
||
]:
|
||
</emphasis>
|
||
<location id="8EA924B0FFD9FFD6FAFFFA0F85FB93D4" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFD9FFD6FAFFFA0F85FB93D4" box="[1376,1471,1493,1512]" country="Madagascar" county="Analamera" latitude="-12.673612" longLatPrecision="21" longitude="49.544445" municipality="Irodo" name="Analamera" pageId="12" pageNumber="171" stateProvince="Reg. Sava">Analamera</location>
|
||
,
|
||
<location id="8EA924B0FFD9FFD6FCA7FA2A838E903F" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFD9FFD6FCA7FA2A838E903F" box="[824,970,1520,1539]" country="Madagascar" county="Analamera" latitude="-12.673612" longLatPrecision="21" longitude="49.544445" municipality="Irodo" name="bank of Irodo riv." pageId="12" pageNumber="171" stateProvince="Reg. Sava">bank of Irodo riv.</location>
|
||
,
|
||
<location id="8EA924B0FFD9FFD6FC4EFA2A8431903F" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFD9FFD6FC4EFA2A8431903F" box="[977,1141,1520,1539]" country="Madagascar" county="Analamera" latitude="-12.673612" longLatPrecision="21" longitude="49.544445" municipality="Irodo" name="side of Irodo village" pageId="12" pageNumber="171" stateProvince="Reg. Sava">side of Irodo village</location>
|
||
,
|
||
<geoCoordinate id="EE4214ACFFD9FFD6FBE3FA2A8497903F" box="[1148,1235,1520,1539]" degrees="12" direction="south" minutes="40" orientation="latitude" pageId="12" pageNumber="171" precision="15" seconds="25" value="-12.673612">12°40'25"S</geoCoordinate>
|
||
<geoCoordinate id="EE4214ACFFD9FFD6FB47FA2A8572903F" box="[1240,1334,1520,1539]" degrees="49" direction="east" minutes="32" orientation="longitude" pageId="12" pageNumber="171" precision="15" seconds="40" value="49.544445">49°32'40"E</geoCoordinate>
|
||
,
|
||
<quantity id="4C8EDF8EFFD9FFD6FAA1FA2A8523903F" box="[1342,1383,1520,1539]" metricMagnitude="1" metricUnit="m" metricValue="4.1" pageId="12" pageNumber="171" unit="m" value="41.0">
|
||
<elevation id="005B9558FFD9FFD6FAA1FA2A8523903F" box="[1342,1383,1520,1539]" metricMagnitude="1" metricUnit="m" metricValue="4.1" pageId="12" pageNumber="171" unit="m" value="41.0">41 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingDate id="EF8CAD43FFD9FFD6FAF0FA2A85FB903F" box="[1391,1471,1520,1539]" pageId="12" pageNumber="171">
|
||
<date id="FFC854ABFFD9FFD6FAF0FA2A85C9903F" box="[1391,1421,1520,1539]" pageId="12" pageNumber="171" value="2002-01-10">10.I</date>
|
||
.2002
|
||
</collectingDate>
|
||
,
|
||
<emphasis id="B902AE79FFD9FFD6FCA7F9D183919022" box="[824,981,1546,1566]" italics="true" pageId="12" pageNumber="171">
|
||
<collectorName id="268317BDFFD9FFD6FCA7F9D1833B9021" box="[824,895,1546,1565]" pageId="12" pageNumber="171">De Block</collectorName>
|
||
et al. 1172
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="ED67EAAEFFD9FFD6FC40F9D183BA9022" box="[991,1022,1547,1566]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="12" pageNumber="171">BR</collectionCode>
|
||
,
|
||
<collectionCode id="ED67EAAEFFD9FFD6FB98F9D184699022" box="[1031,1069,1547,1566]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="12" pageNumber="171" type="Herbarium">MO</collectionCode>
|
||
,
|
||
<collectionCode id="ED67EAAEFFD9FFD6FBA9F9D184059022" box="[1078,1089,1547,1566]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="12" pageNumber="171" type="Herbarium">P</collectionCode>
|
||
,
|
||
<collectionCode id="ED67EAAEFFD9FFD6FBD7F9D1843C9022" box="[1096,1144,1547,1566]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="12" pageNumber="171" type="Herbarium">TAN</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1E7836FFD9FFD6FB19F9D1832B906F" ID-GBIF-Occurrence="3697348384" collectingDate="2003-02-02" collectionCode="MO, TEF" collectorName="Rabevohitra et al" country="Madagascar" county="Analamera" elevation="41" latitude="-12.673612" location="Nosy Be" longLatPrecision="21" longitude="49.544445" municipality="Irodo" pageId="12" pageNumber="171" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
|
||
<location id="8EA924B0FFD9FFD6FB19F9D1848B9022" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFD9FFD6FB19F9D1848B9022" box="[1158,1231,1547,1566]" country="Madagascar" county="Analamera" latitude="-12.673612" longLatPrecision="21" longitude="49.544445" municipality="Irodo" name="Nosy Be" pageId="12" pageNumber="171" stateProvince="Antsiranana">Nosy Be</location>
|
||
,
|
||
<location id="8EA924B0FFD9FFD6FB48F9D185D69021" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFD9FFD6FB48F9D185D69021" box="[1239,1426,1546,1566]" country="Madagascar" county="Analamera" latitude="-12.673612" longLatPrecision="21" longitude="49.544445" municipality="Irodo" name="au bord du lac Sahaka" pageId="12" pageNumber="171" stateProvince="Antsiranana">au bord du lac Sahaka</location>
|
||
,
|
||
<location id="8EA924B0FFD9FFD6FA05F9D083D69004" LSID="urn:lsid:plazi:treatment:03DFC37DFFDEFFD6FFEEF8F683FD9052:8EA924B0FFD9FFD6FA05F9D083D69004" country="Madagascar" county="Analamera" latitude="-12.673612" longLatPrecision="21" longitude="49.544445" municipality="Irodo" name="foret d'Analabe" pageId="12" pageNumber="171" stateProvince="Antsiranana">forêt d’Analabe</location>
|
||
,
|
||
<geoCoordinate id="EE4214ACFFD9FFD6FC04F9FF83B29004" box="[923,1014,1573,1592]" degrees="13" direction="south" minutes="04" orientation="latitude" pageId="12" pageNumber="171" precision="15" seconds="14" value="-13.070556">13°04'14"S</geoCoordinate>
|
||
<geoCoordinate id="EE4214ACFFD9FFD6FC63F9FF84199004" box="[1020,1117,1573,1592]" degrees="49" direction="east" minutes="53" orientation="longitude" pageId="12" pageNumber="171" precision="15" seconds="40" value="49.894447">49°53'40"E</geoCoordinate>
|
||
,
|
||
<date id="FFC854ABFFD9FFD6FBF8F9FF84FE9004" box="[1127,1210,1573,1592]" pageId="12" pageNumber="171" value="2003-02-02">
|
||
<collectingDate id="EF8CAD43FFD9FFD6FBF8F9FF84FE9004" box="[1127,1210,1573,1592]" pageId="12" pageNumber="171" value="2003-02-02">2.II.2003</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="B902AE79FFD9FFD6FB5BF9FF85CE9004" box="[1220,1418,1573,1592]" italics="true" pageId="12" pageNumber="171">
|
||
<collectorName id="268317BDFFD9FFD6FB5BF9FF85139004" box="[1220,1367,1573,1592]" pageId="12" pageNumber="171">Rabevohitra et al</collectorName>
|
||
. 4463
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="ED67EAAEFFD9FFD6FA08F9FF85FB9004" box="[1431,1471,1573,1592]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="12" pageNumber="171" type="Herbarium">MO</collectionCode>
|
||
,
|
||
<collectionCode id="ED67EAAEFFD9FFD6FCA7F99A8320906F" box="[824,868,1600,1619]" collectionName="TEF" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12774" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="12" pageNumber="171" type="Herbarium">TEF</collectionCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1E7836FFD9FFD6FCEBF99A83FD9052" ID-GBIF-Occurrence="3697348350" collectingDate="2003-05-13" collectionCode="MO, P, TEF" collectorName="Razakamalala et al" country="Madagascar" county="Analamera" elevation="41" latitude="-12.673612" location="Irodo village" longLatPrecision="21" longitude="49.544445" municipality="Irodo" pageId="12" pageNumber="171" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
|
||
<emphasis id="B902AE79FFD9FFD6FCEBF99A8386906F" box="[884,962,1600,1619]" italics="true" pageId="12" pageNumber="171">ibid. loco,</emphasis>
|
||
<geoCoordinate id="EE4214ACFFD9FFD6FC59F99A8464906F" box="[966,1056,1600,1619]" degrees="13" direction="south" minutes="04" orientation="latitude" pageId="12" pageNumber="171" precision="15" seconds="43" value="-13.078611">13°04'43"S</geoCoordinate>
|
||
<geoCoordinate id="EE4214ACFFD9FFD6FBB9F99A84C2906F" box="[1062,1158,1600,1619]" degrees="49" direction="east" minutes="54" orientation="longitude" pageId="12" pageNumber="171" precision="15" seconds="04" value="49.90111">49°54'04"E</geoCoordinate>
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,
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||
<date id="FFC854ABFFD9FFD6FB0FF99A84A1906F" box="[1168,1253,1600,1619]" pageId="12" pageNumber="171" value="2003-05-13">
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||
<collectingDate id="EF8CAD43FFD9FFD6FB0FF99A84A1906F" box="[1168,1253,1600,1619]" pageId="12" pageNumber="171" value="2003-05-13">13.V.2003</collectingDate>
|
||
</date>
|
||
,
|
||
<emphasis id="B902AE79FFD9FFD6FB71F99A8585906F" box="[1262,1473,1600,1619]" italics="true" pageId="12" pageNumber="171">
|
||
<collectorName id="268317BDFFD9FFD6FB71F99A85D5906F" box="[1262,1425,1600,1619]" pageId="12" pageNumber="171">Razakamalala et al</collectorName>
|
||
. 1272
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="ED67EAAEFFD9FFD6FCA1F98183219052" box="[830,869,1627,1646]" pageId="12" pageNumber="171">MO</collectionCode>
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||
,
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||
<collectionCode id="ED67EAAEFFD9FFD6FCF1F981833D9052" box="[878,889,1627,1646]" pageId="12" pageNumber="171">P</collectionCode>
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||
,
|
||
<collectionCode id="ED67EAAEFFD9FFD6FC1EF98183EA9052" box="[897,942,1627,1646]" collectionName="TEF" pageId="12" pageNumber="171">TEF</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |