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<document id="E8183ABE6E37F02A84502BF32F5A99D8" ID-DOI="10.11646/zootaxa.3655.1.1" ID-GBIF-Dataset="a6990774-66c7-4586-b14e-354b42d0857b" ID-ISSN="1175-5326" ID-Zenodo-Dep="223358" ID-ZooBank="FC302AA5-49CC-41B0-9A66-23C11AB4EBAE" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460612493272" checkinUser="plazi" docAuthor="Gardini, Giulio" docDate="2013" docId="03DE87C1FFD1F95E6B99FAFA1CC3A147" docLanguage="en" docName="zt03655p151.pdf" docOrigin="Zootaxa 3655 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Chthonius Gardini, 2013, n. sp." docType="treatment" docVersion="8" lastPageNumber="61" masterDocId="FFE7FFB9FFEBF9636B0EFF8C1851A078" masterDocTitle="A revision of the species of the pseudoscorpion subgenus Chthonius (Ephippiochthonius) (Arachnida, Pseudoscorpiones, Chthoniidae) from Italy and neighbouring areas" masterLastPageNumber="151" masterPageNumber="1" pageNumber="59" updateTime="1698329375031" updateUser="plazi">
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<mods:title id="B39F27844DF2C64E54845C7DF7AF999F">A revision of the species of the pseudoscorpion subgenus Chthonius (Ephippiochthonius) (Arachnida, Pseudoscorpiones, Chthoniidae) from Italy and neighbouring areas</mods:title>
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<mods:name id="1123A15E6DA7FF99A6C9F80BA5024A9B" type="personal">
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<mods:namePart id="1B2CAF7007E78973C58DCE851C4BE912">Gardini, Giulio</mods:namePart>
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<mods:date id="9960B4805A5BB12CD1703C27705FE874">2013</mods:date>
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<treatment id="03DE87C1FFD1F95E6B99FAFA1CC3A147" ID-DOI="http://doi.org/10.5281/zenodo.6155959" ID-GBIF-Taxon="119568573" ID-Zenodo-Dep="6155959" LSID="urn:lsid:plazi:treatment:03DE87C1FFD1F95E6B99FAFA1CC3A147" httpUri="http://treatment.plazi.org/id/03DE87C1FFD1F95E6B99FAFA1CC3A147" lastPageId="61" lastPageNumber="61" pageId="58" pageNumber="59">
<subSubSection id="C36D655CFFD1F9596B99FAFA192BA5C9" pageId="58" pageNumber="59" type="nomenclature">
<paragraph id="8BC836D7FFD1F9596B99FAFA1A4BA5E8" blockId="58.[151,538,1398,1457]" box="[151,538,1398,1424]" pageId="58" pageNumber="59">
<heading id="D08081BBFFD1F9596B99FAFA1A4BA5E8" bold="true" box="[151,538,1398,1424]" fontSize="11" level="1" pageId="58" pageNumber="59" reason="1">
<emphasis id="B903EAC5FFD1F9596B99FAFA1A4BA5E8" bold="true" box="[151,538,1398,1424]" pageId="58" pageNumber="59">
<emphasis id="B903EAC5FFD1F9596B99FAFA1981A5E8" bold="true" box="[151,464,1398,1424]" italics="true" pageId="58" pageNumber="59">
<taxonomicName id="4C774D54FFD1F9596B99FAFA1947A5E8" ID-CoL="62LNY" box="[151,278,1398,1424]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.)
<taxonomicName id="4C774D54FFD1F9596A5DFAFA1981A5E8" box="[339,464,1398,1424]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="intemelius" status="sp. nov.">intemelius</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A23057BEFFD1F9596AD6FAFA1A4BA5E8" box="[472,538,1398,1424]" pageId="58" pageNumber="59" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836D7FFD1F9596B99FA15192BA5C9" blockId="58.[151,538,1398,1457]" box="[151,378,1433,1457]" pageId="58" pageNumber="59">
(
<figureCitation id="134C2A52FFD1F9596B91FA15196AA5C9" box="[159,315,1433,1457]" captionStart-0="FIGURES 112 118" captionStart-1="FIGURES 119 120" captionStartId-0="59.[151,264,1663,1686]" captionStartId-1="61.[151,264,1912,1935]" captionTargetBox-0="[211,1380,183,1640]" captionTargetBox-1="[261,1324,401,1877]" captionTargetId-0="figure@59.[182,1405,155,1659]" captionTargetId-1="figure@61.[251,1336,375,1890]" captionTargetPageId-0="59" captionTargetPageId-1="61" captionText-0="FIGURES 112 118. Chthonius (E.) intemelius n. sp. 112, male holotype, anterior margin of carapace; 113, female paratype, id.; 114, female paratype, id.; 115, male holotype, anterolateral portion of carapace; 116, id., right chelicera; 117, male paratype, fingers of right chelicera; 118, female paratype, right chelicera." captionText-1="FIGURES 119 120. Chthonius (E.) intemelius n. sp. 119, male holotype, right chela, lateral view; 120, female paratype, id. (Scale bar 0.1 mm)." httpUri-0="https://zenodo.org/record/223384/files/figure.png" httpUri-1="https://zenodo.org/record/223385/files/figure.png" pageId="58" pageNumber="59">Figs 112120</figureCitation>
,
<figureCitation id="134C2A52FFD1F9596A48FA151920A5C9" box="[326,369,1433,1457]" captionStart="FIGURES 290 293" captionStartId="146.[151,264,1815,1838]" captionTargetBox="[151,1435,193,1793]" captionTargetId="figure@146.[151,1436,193,1794]" captionTargetPageId="146" captionText="FIGURES 290 293. Distribution maps for the Italian species of the Chthonius fuscimanus group. 290, Chthonius (E.) boldorii Beier, 1934; 291, Chthonius (E.) genuensis Gardini, 1990, C. (E.) gestroi Simon, 1896, C. (E.) monguzzii n. sp., C. (E.) pieltaini Beier, 1930, C. (E.) troglophilus Beier, 1930 and C. (E.) zoiai Gardini, 1990; 292, Chthonius (E.) etruscus n. sp., C. (E.) latellai n. sp., C. (E.) magrinii n. sp. and C. (E.) sulphureus n. sp.; 293, Chthonius (E.) aeneae n. sp., C. (E.) gallii n. sp., C. (E.) intemelius n. sp. and C. (E.) ligur n. sp." httpUri="https://zenodo.org/record/223424/files/figure.png" pageId="58" pageNumber="59">293</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36D655CFFD1F9596B99FA6E1CC2A581" box="[151,1171,1504,1530]" pageId="58" pageNumber="59" type="materials_examined">
<paragraph id="8BC836D7FFD1F9596B99FA6E1CC2A581" blockId="58.[151,1436,1504,2033]" box="[151,1171,1504,1530]" pageId="58" pageNumber="59">
<emphasis id="B903EAC5FFD1F9596B99FA6E1968A582" bold="true" box="[151,313,1505,1530]" pageId="58" pageNumber="59">
<typeStatus id="54CC8875FFD1F9596B99FA6E1883A582" box="[151,210,1506,1530]" pageId="58" pageNumber="59">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F3607647FFD1F9596A4EFA6D1925A581" box="[320,372,1505,1529]" name="Italy" pageId="58" pageNumber="59">Italy</collectingCountry>
, Liguria, Imperia Prov., Apricale, Mt Cianela (43°53ʹ45ʺN 7°40ʹ36ʺE).
</paragraph>
</subSubSection>
<subSubSection id="C36D655CFFD1F9596BC9F9891A52A665" box="[199,515,1541,1566]" pageId="58" pageNumber="59" type="distribution">
<paragraph id="8BC836D7FFD1F9596BC9F9891A52A665" blockId="58.[151,1436,1504,2033]" box="[199,515,1541,1566]" pageId="58" pageNumber="59">
<emphasis id="B903EAC5FFD1F9596BC9F9891931A666" bold="true" box="[199,352,1541,1566]" pageId="58" pageNumber="59">Distribution.</emphasis>
<collectingCountry id="F3607647FFD1F9596A69F98919CEA665" box="[359,415,1541,1565]" name="Italy" pageId="58" pageNumber="59">Italy</collectingCountry>
: Liguria.
</paragraph>
</subSubSection>
<subSubSection id="C36D655CFFD1F9596BC9F9A51BA6A719" pageId="58" pageNumber="59" type="diagnosis">
<paragraph id="8BC836D7FFD1F9596BC9F9A51BA6A719" blockId="58.[151,1436,1504,2033]" pageId="58" pageNumber="59">
<emphasis id="B903EAC5FFD1F9596BC9F9A51901A63A" bold="true" box="[199,336,1577,1602]" pageId="58" pageNumber="59">Diagnosis (</emphasis>
Ƥ
<emphasis id="B903EAC5FFD1F9596A75F9A519DBA63A" bold="true" box="[379,394,1577,1602]" pageId="58" pageNumber="59">).</emphasis>
A microphthalmic epigean
<taxonomicName id="4C774D54FFD1F95969D3F9A51C12A639" authority="(Ephippiochthonius)" baseAuthorityName="Ephippiochthonius" box="[733,1091,1577,1601]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="58" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="B903EAC5FFD1F95969D3F9A51C12A639" box="[733,1091,1577,1601]" italics="true" pageId="58" pageNumber="59">Chthonius (Ephippiochthonius)</emphasis>
</taxonomicName>
from Ligurian pre-Alps that differs from other species of the
<taxonomicName id="4C774D54FFD1F9596927F9C21A9FA61D" box="[553,718,1613,1637]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="fuscimanus">
<emphasis id="B903EAC5FFD1F9596927F9C21A9FA61D" box="[553,718,1613,1637]" italics="true" pageId="58" pageNumber="59">C. fuscimanus</emphasis>
</taxonomicName>
group in the following combination of characters: movable cheliceral finger with isolated subapical tooth (
<emphasis id="B903EAC5FFD1F95969ACF9FD1AE6A6F1" box="[674,695,1649,1673]" italics="true" pageId="58" pageNumber="59">di</emphasis>
); spinneret absent or vestigial in males; pedipalpal hand (in lateral view) without a marked rounded hump distad of the trichobothria
<emphasis id="B903EAC5FFD1F9596871F9191BEEA6D5" box="[895,959,1685,1709]" italics="true" pageId="58" pageNumber="59">ib-isb</emphasis>
, weakly depressed at level of
<emphasis id="B903EAC5FFD1F9596E1DF9191D02A6D5" box="[1299,1363,1685,1709]" italics="true" pageId="58" pageNumber="59">ib-isb</emphasis>
; fixed and movable chelal fingers respectively with
<date id="FFC91017FFD1F95969ACF9351B1DA6A9" box="[674,844,1721,1745]" pageId="58" pageNumber="59" value="2011-06-12" valueMax="2011-06-05" valueMin="2011-06-12">1112 and 56</date>
triangular teeth (proximal rounded teeth excepted); pedipalpal fixed finger at level of
<emphasis id="B903EAC5FFD1F9596916F9521A01A68D" box="[536,592,1758,1781]" italics="true" pageId="58" pageNumber="59">est-it</emphasis>
with 5 teeth occupying
<quantity id="4C8F9B32FFD1F959686CF9511BE9A68D" box="[866,952,1757,1781]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="58" pageNumber="59" unit="mm" value="0.1">0.1 mm</quantity>
(distance between successive apices
<quantity id="4C8F9B32FFD1F9596E53F9511896A761" metricMagnitude="-5" metricUnit="m" metricValue="2.1" pageId="58" pageNumber="59" unit="mm" value="0.021">0.021 mm</quantity>
); proximal half of movable chelal finger with thick and undulated marginal lamina; length of chela 0.510.53 (3), 0.560.55 (Ƥ) mm, length of movable finger 0.300.31 (3), 0.3150.325 (Ƥ) mm; chela 5.25.4 (3), 4.64.75 (Ƥ) times as long as deep; ratio of pedipalpal femur/carapace 1.151.2 (3Ƥ).
</paragraph>
</subSubSection>
<subSubSection id="C36D655CFFD1F9596BC9F8E119DDA789" pageId="58" pageNumber="59" type="materials_examined">
<paragraph id="8BC836D7FFD1F9596BC9F8E119DDA789" blockId="58.[151,1436,1504,2033]" pageId="58" pageNumber="59">
<emphasis id="B903EAC5FFD1F9596BC9F8E1192AA7FE" bold="true" box="[199,379,1901,1926]" pageId="58" pageNumber="59">
<typeStatus id="54CC8875FFD1F9596BC9F8E11953A7FD" box="[199,258,1901,1925]" pageId="58" pageNumber="59">Type</typeStatus>
material.
</emphasis>
ITALY—
<emphasis id="B903EAC5FFD1F9596AFDF8E21B46A7FD" box="[499,791,1902,1925]" italics="true" pageId="58" pageNumber="59">Liguria: Imperia Prov.—</emphasis>
1 3 (
<typeStatus id="54CC8875FFD1F9596853F8E11B94A7FD" box="[861,965,1901,1925]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
),
<date id="FFC91017FFD1F95968D0F8E11C17A7FD" box="[990,1094,1901,1925]" pageId="58" pageNumber="59" value="1918-03-22">22 3 18</date>
Ƥ 8T (
<typeStatus id="54CC8875FFD1F9596FA4F8E11D4AA7FD" box="[1194,1307,1901,1925]" pageId="58" pageNumber="59" type="paratype">paratypes</typeStatus>
), Apricale, halfway between Baiardo and Apricale, Monte Cianela (43°53ʹ45ʺN 7°40ʹ36ʺE), ±
<quantity id="4C8F9B32FFD1F9596F36F81D1C2FA7D1" box="[1080,1150,1937,1961]" metricMagnitude="2" metricUnit="m" metricValue="5.9" pageId="58" pageNumber="59" unit="m" value="590.0">590 m</quantity>
a.s.l.,
<date id="FFC91017FFD1F9596FC8F81D1D6BA7D1" box="[1222,1338,1937,1961]" pageId="58" pageNumber="59" value="1981-11-06">6.XI.1981</date>
, S. Zoia, G. Gardini &amp; A. Boato leg., sieved in
<taxonomicName id="4C774D54FFD1F959694AF83A1A80A7B5" box="[580,721,1973,1997]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="58" pageNumber="59" phylum="Tracheophyta" rank="species" species="ilex">
<emphasis id="B903EAC5FFD1F959694AF83A1A80A7B5" box="[580,721,1973,1997]" italics="true" pageId="58" pageNumber="59">Quercus ilex</emphasis>
</taxonomicName>
wood on flysch (
<typeStatus id="54CC8875FFD1F9596898F8391BABA7B5" box="[918,1018,1973,1997]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
deposited in MHNG,
<typeStatus id="54CC8875FFD1F9596FFDF83A1D0FA7B6" box="[1267,1374,1974,1998]" pageId="58" pageNumber="59" type="paratype">paratypes</typeStatus>
in G. Gardini coll., Genoa).
</paragraph>
</subSubSection>
<caption id="DF08665FFFD0F9586B99F9F31B2AA6AB" httpUri="https://zenodo.org/record/223384/files/figure.png" pageId="59" pageNumber="60" targetBox="[211,1380,183,1640]" targetPageId="59">
<paragraph id="8BC836D7FFD0F9586B99F9F31B2AA6AB" blockId="59.[151,1437,1663,1747]" pageId="59" pageNumber="60">
<emphasis id="B903EAC5FFD0F9586B99F9F31935A6ED" bold="true" box="[151,356,1663,1686]" pageId="59" pageNumber="60">FIGURES 112118</emphasis>
.
<emphasis id="B903EAC5FFD0F9586A7DF90C1A2BA6EE" box="[371,634,1664,1686]" italics="true" pageId="59" pageNumber="60">
<taxonomicName id="4C774D54FFD0F9586A7DF90C198AA6EE" box="[371,475,1664,1686]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="genus">Chthonius</taxonomicName>
(E.)
<taxonomicName id="4C774D54FFD0F958691CF90C1A2BA6EE" box="[530,634,1664,1686]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="intemelius" status="sp. nov.">intemelius</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A23057BEFFD0F958698CF90D1AEBA6EE" box="[642,698,1665,1686]" pageId="59" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
112, male holotype, anterior margin of carapace; 113, female paratype, id.; 114, female paratype, id.; 115, male holotype, anterolateral portion of carapace; 116, id., right chelicera; 117, male paratype, fingers of right chelicera; 118, female paratype, right chelicera.
</paragraph>
</caption>
<subSubSection id="C36D655CFFD0F9586BC9F9711C5BA741" pageId="59" pageNumber="60" type="etymology">
<paragraph id="8BC836D7FFD0F9586BC9F9711C5BA741" blockId="59.[151,1436,1789,2030]" pageId="59" pageNumber="60">
<emphasis id="B903EAC5FFD0F9586BC9F97119CCA76E" bold="true" box="[199,413,1789,1814]" italics="true" pageId="59" pageNumber="60">Derivatio nominis.</emphasis>
The Intemelii were the inhabitants of Albium Intemelium (Strabo,
<emphasis id="B903EAC5FFD0F9586F85F9711D56A76D" box="[1163,1287,1789,1813]" italics="true" pageId="59" pageNumber="60">Geography</emphasis>
,
<date id="FFC91017FFD0F9586E1BF9721D18A76D" box="[1301,1353,1789,1813]" pageId="59" pageNumber="60">IV 6</date>
, 1), the current Ventimiglia, in which the villages of Baiardo and Apricale are situated.
</paragraph>
</subSubSection>
<subSubSection id="C36D655CFFD0F95F6BC9F8C91C4DA6A7" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60" type="description">
<paragraph id="8BC836D7FFD0F95F6BC9F8C919FCA3BF" blockId="59.[151,1436,1789,2030]" lastBlockId="60.[151,1437,151,2013]" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">
<emphasis id="B903EAC5FFD0F9586BC9F8C91982A726" bold="true" box="[199,467,1861,1886]" pageId="59" pageNumber="60">Description of adults (</emphasis>
Ƥ
<emphasis id="B903EAC5FFD0F9586AF0F8C91A5FA726" bold="true" box="[510,526,1861,1886]" pageId="59" pageNumber="60">).</emphasis>
Integument weakly pigmented; carapace, tergites, chelicerae and pedipalps pale yellow-brown; marked hispid granulation on lateral surface of carapace and on cheliceral palm. Carapace 1.01.1 times longer than broad, subquadrate, posteriorly slightly constricted; anterior margin (figs 112114) strongly dentate between median macrosetae, sometimes with small epistome; ocular area as in fig. 115, anterior eyes with tapetum and weak convex lens (diameter
<quantity id="4C8F9B32FFD0F9586965F8591B7FA795" box="[619,814,2005,2029]" metricMagnitude="-5" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="59" pageNumber="60" unit="mm" value="0.0265" valueMax="0.028" valueMin="0.025">0.0250.028 mm</quantity>
), posterior eyes absent, reduced to a pale cuticular area with hispid granulation; distance from anterior eyes to anterior margin of carapace
<quantity id="4C8F9B32FFD7F95F6F67FF1B1D42A0D7" box="[1129,1299,151,175]" metricMagnitude="-5" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="60" pageNumber="61" unit="mm" value="0.025" valueMax="0.03" valueMin="0.02">0.020.03 mm</quantity>
; chaetotaxy mm
<quantity id="4C8F9B32FFD7F95F6BC7FF30195BA0AC" box="[201,266,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="60" pageNumber="61" unit="mm" value="4.0">4mm</quantity>
:6:4:2:2 (18), macrosetae thick; length of anteromedian macrosetae 0.0700075 mm. Chaetotaxy tergites
<date id="FFC91017FFD7F95F6B99FF6C188FA08F" box="[151,222,223,247]" pageId="60" pageNumber="61">IX 4</date>
:4:4:4:6:6:6:6:6:4. Chaetotaxy sternites
<date id="FFC91017FFD7F95F6993FF6C1AABA08F" box="[669,762,223,247]" pageId="60" pageNumber="61" value="2010-02" valueMax="2010-10-31" valueMin="2010-02-10">IIX 10</date>
:(3)8(3):(2)7(2):8:6:6:6:6:7; sternite III rarely with (3)10(3) setae; genital opening of males flanked by 5 or 6 (rarely 8) setae on each side. Chelicerae (figs 116118) 2.02.1 times as long as broad, palm with 6 setae and 2 microsetae laterally; fixed finger with 67 teeth proximally reduced in size and 510 proximal microtubercles; movable finger with a large isolated subapical tooth (
<emphasis id="B903EAC5FFD7F95F6E16FEC01D7CA11C" box="[1304,1325,332,356]" italics="true" pageId="60" pageNumber="61">di</emphasis>
), 5 teeth proximally reduced in size and 47 proximal microtubercles;
<emphasis id="B903EAC5FFD7F95F685FFEFC1B36A1F0" box="[849,871,368,392]" italics="true" pageId="60" pageNumber="61">gl</emphasis>
ratio 0.560.62; spinneret mostly absent in males (rarely weakly raised), prominent and apically rounded in females; rallum with 11 blades; serrulae interior and exterior respectively with 12 and 14 blades. Coxal setae: pedipalp 5 (including 2 on manducatory process),
<date id="FFC91017FFD7F95F6E5FFE341D3EA1B7" box="[1361,1391,439,463]" pageId="60" pageNumber="61">I 3</date>
+ 3 marginal microsetae,
<date id="FFC91017FFD7F95F6A85FE5119E6A18C" box="[395,439,476,500]" pageId="60" pageNumber="61">II 4</date>
,
<date id="FFC91017FFD7F95F6ACAFE5119A8A18C" box="[452,505,476,500]" pageId="60" pageNumber="61">III 5</date>
,
<date id="FFC91017FFD7F95F6908FE511A6DA18C" box="[518,572,476,500]" pageId="60" pageNumber="61">IV 6</date>
; coxa II with 68 (rarely 5) coxal spines, coxa III with 45 (rarely 3) coxal spines; intercoxal tubercle bisetose. Pedipalp: femur 5.15.3 (3Ƥ) times as long as broad, with laterobasal hispid granulation; chela (figs 119120) 5.25.4 (3), 4.64.75 (Ƥ) times as long as deep; hand of chela 2.152.2 (3), 2.0 (Ƥ) times as long as deep, depressed at level of
<emphasis id="B903EAC5FFD7F95F69A7FDC41AB8A218" box="[681,745,584,608]" italics="true" pageId="60" pageNumber="61">ib-isb</emphasis>
, in males with dorsal straight profile between the base and
<emphasis id="B903EAC5FFD7F95F6E73FDC418E9A2FC" italics="true" pageId="60" pageNumber="61">ibisb</emphasis>
; ventrodistal surface of chelal hand with weak hispid granulation; fixed chelal finger with 1112 (rarely 13) triangular teeth, weakly reclined, with dental canals: the first 23 distal teeth reduced in size, the proximal one tooth with rounded tip; base of fixed finger with 69 microtubercles; tip of fixed finger with a modified accessory tooth (
<emphasis id="B903EAC5FFD7F95F6BECFD5418A9A288" box="[226,248,728,752]" italics="true" pageId="60" pageNumber="61">td</emphasis>
) on antiaxial face; fixed chelal finger at level of
<emphasis id="B903EAC5FFD7F95F6829FD551B0EA288" box="[807,863,729,752]" italics="true" pageId="60" pageNumber="61">est-it</emphasis>
with 5 teeth occupying
<quantity id="4C8F9B32FFD7F95F6F7DFD5B1C9BA297" box="[1139,1226,727,751]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="60" pageNumber="61" unit="mm" value="0.1">0.1 mm</quantity>
(distance between successive apices
<quantity id="4C8F9B32FFD7F95F6A6BFD70198DA36C" box="[357,476,764,788]" metricMagnitude="-5" metricUnit="m" metricValue="2.1" pageId="60" pageNumber="61" unit="mm" value="0.021">0.021 mm</quantity>
); distal half of movable chelal finger with 6 (rarely 5) upright, triangular teeth, with dental canals; proximal half of movable chelal finger with 13 vestigial teeth, marginal lamina thick, undulated; coupled sensilla
<emphasis id="B903EAC5FFD7F95F6A5CFCC9193CA324" box="[338,365,837,860]" italics="true" pageId="60" pageNumber="61">pc</emphasis>
mostly at level of
<emphasis id="B903EAC5FFD7F95F6930FCC81A06A324" box="[574,599,836,860]" italics="true" pageId="60" pageNumber="61">sb</emphasis>
; trichobothria as in figs 119120,
<emphasis id="B903EAC5FFD7F95F68D7FCC81C17A324" box="[985,1094,836,860]" italics="true" pageId="60" pageNumber="61">eb-esb-ist</emphasis>
placed in a straight line; basal apodeme of movable finger strongly sclerotized, subrectangular, apically triangular; ratio of movable finger/hand of chela 1.41.5 (3), 1.21.35 (Ƥ); ratio of pedipalpal femur/movable finger 1.2 (3Ƥ); ratio of pedipalpal femur/ carapace 1.151.2 (3Ƥ).
</paragraph>
<paragraph id="8BC836D7FFD7F95F6BC9FC591BF6A404" blockId="60.[151,1437,151,2013]" pageId="60" pageNumber="61">Measurements (in mm). Body length 1.01.05 (3), 1.051.1 (Ƥ). Carapace 0.310.32 × 0.280.30 (0.240.26 anteriorly) (3), 0.3150.33 × 0.320.33 (0.280.29 anteriorly) (Ƥ). Chelicerae 0.230.245 × 0.110.115 (3), 0.26 0.27 × 0.1250.13 (Ƥ), movable finger length 0.1150.12 (3), 0.1250.13 (Ƥ). Pedipalp: femur 0.360.37 × 0.07 (3), 0.370.39 × 0.070.08 (Ƥ); chela 0.510.53 × 0.0950.10 (3), 0.560.57 × 0.12 (Ƥ); hand length 0.2050.22 (3), 0.24 (Ƥ); movable finger length 0.300.31 (3), 0.3150.325 (Ƥ).</paragraph>
<paragraph id="8BC836D7FFD7F95F6BC9FB0B1BE7A60C" blockId="60.[151,1437,151,2013]" pageId="60" pageNumber="61">
<emphasis id="B903EAC5FFD7F95F6BC9FB0B1A42A4D8" bold="true" box="[199,531,1159,1184]" pageId="60" pageNumber="61">Description of tritonymph.</emphasis>
Integument with weak pigmentation and weak cuticular hispid granulation. Carapace 1.1 times longer than broad, anterior margin dentate, without epistome; anterior eyes with tapetum and weak convex lens, posterior eyes absent; chaetotaxy m
<quantity id="4C8F9B32FFD7F95F6804FB431B65A49F" box="[778,820,1231,1255]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="60" pageNumber="61" unit="m" value="4.0">4m</quantity>
:6:4:2:2 (18), length of anteromedian macrosetae
<quantity id="4C8F9B32FFD7F95F6E53FB4318AEA574" metricMagnitude="-5" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="60" pageNumber="61" unit="mm" value="0.055" valueMax="0.06" valueMin="0.05">0.05 0.06 mm</quantity>
. Chaetotaxy of tergites as in adults. Chaetotaxy of sternites
<date id="FFC91017FFD7F95F68A8FB791BADA574" box="[934,1020,1268,1292]" pageId="60" pageNumber="61">IIIX 5</date>
:(2)6(2):(1)6(1):7:6:6:6:6. Chelicerae 2.0 times as long as broad, palm with 5 setae and 1 lateral microseta; fixed finger with 56 teeth, movable finger with an isolated subapical tooth (
<emphasis id="B903EAC5FFD7F95F6904FAB01A71A52C" box="[522,544,1340,1364]" italics="true" pageId="60" pageNumber="61">di</emphasis>
) and 5 teeth; spinneret as in females; rallum with 9 blades; serrula interior with 10, exterior with 12 blades;
<emphasis id="B903EAC5FFD7F95F6AD5FAEC19A0A500" box="[475,497,1376,1400]" italics="true" pageId="60" pageNumber="61">gl</emphasis>
ratio 0.61. Coxal setae: pedipalp 5 (including 2 on manducatory process),
<date id="FFC91017FFD7F95F6E42FAED1D3DA50F" box="[1356,1388,1375,1400]" pageId="60" pageNumber="61">I 3</date>
+ 2 marginal microsetae,
<date id="FFC91017FFD7F95F6A83FA0919E8A5E4" box="[397,441,1412,1436]" pageId="60" pageNumber="61">II 4</date>
,
<date id="FFC91017FFD7F95F6AC6FA0919AFA5E4" box="[456,510,1412,1436]" pageId="60" pageNumber="61">III 5</date>
,
<date id="FFC91017FFD7F95F6902FA091A15A5E4" box="[524,580,1412,1436]" pageId="60" pageNumber="61">IV 5</date>
; coxa II with 58 coxal spines, coxa III with 35 coxal spines; intercoxal tubercle. Pedipalp: femur 5.15.2 times as long as broad; chela 5.05.2 times as long as deep; hand of chela 2.1 2.25 times as long as deep; fixed finger with 11 triangular teeth, the first two proximal apically rounded; base of fixed finger with microtubercles; tip of fixed finger with a modified accessory tooth (
<emphasis id="B903EAC5FFD7F95F6F64FA7C1CD1A670" box="[1130,1152,1520,1544]" italics="true" pageId="60" pageNumber="61">td</emphasis>
) on antiaxial face; distal half of movable finger with 56 triangular teeth and 23 vestigial teeth, marginal lamina weakly undulated; coupled sensilla
<emphasis id="B903EAC5FFD7F95F6A5BF9B51921A628" box="[341,368,1593,1616]" italics="true" pageId="60" pageNumber="61">pc</emphasis>
mostly nearer to
<emphasis id="B903EAC5FFD7F95F6934F9B41A19A628" box="[570,584,1592,1616]" italics="true" pageId="60" pageNumber="61">b</emphasis>
than
<emphasis id="B903EAC5FFD7F95F6984F9B41AF2A628" box="[650,675,1592,1616]" italics="true" pageId="60" pageNumber="61">sb</emphasis>
; ratio of movable finger/hand of chela 1.31.4; ratio of pedipalpal femur/movable finger 1.151.2; ratio of pedipalpal femur/carapace 1.0.
</paragraph>
<paragraph id="8BC836D7FFD7F95F6BC9F90C1C4DA6A7" blockId="60.[151,1437,151,2013]" pageId="60" pageNumber="61">Measurements (in mm). Body length 0.80. Carapace 0.260.29 × 0.230.25 (0.200.22 anteriorly). Chelicerae 0.180.20 × 0.090.10, movable finger length 0.090.10. Pedipalp: femur 0.260.28 × 0.050.055; chela 0.39 0.425 × 0.0750.085; hand length 0.170.18; movable finger length 0.220.245.</paragraph>
</subSubSection>
<subSubSection id="C36D655CFFD7F95E6BC9F9601CC3A147" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="discussion">
<paragraph id="8BC836D7FFD7F95E6BC9F9601CC3A147" blockId="60.[151,1437,151,2013]" lastBlockId="61.[151,1436,151,320]" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
<emphasis id="B903EAC5FFD7F95F6BC9F960196AA77C" bold="true" box="[199,315,1772,1796]" pageId="60" pageNumber="61">Remarks.</emphasis>
Among the species of the
<taxonomicName id="4C774D54FFD7F95F6965F9601B35A77C" box="[619,868,1772,1796]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="fuscimanus">
<emphasis id="B903EAC5FFD7F95F6965F9601B35A77C" box="[619,868,1772,1796]" italics="true" pageId="60" pageNumber="61">Chthonius fuscimanus</emphasis>
</taxonomicName>
group,
<taxonomicName id="4C774D54FFD7F95F68B2F9611C00A77C" box="[956,1105,1772,1796]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="intemelius" status="sp. nov.">
<emphasis id="B903EAC5FFD7F95F68B2F9611C00A77C" box="[956,1105,1772,1796]" italics="true" pageId="60" pageNumber="61">C. intemelius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057BEFFD7F95F6F57F9611CC7A77C" box="[1113,1174,1773,1796]" pageId="60" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
is closely related to
<taxonomicName id="4C774D54FFD7F95F6E8CF961188BA750" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B903EAC5FFD7F95F6E8CF961188BA750" italics="true" pageId="60" pageNumber="61">C. nanus</emphasis>
</taxonomicName>
, a common and widespread epigean species also present in the environs of Baiardo. It is difficult to verbally describe the differences between the two species, which are nevertheless recognizable at first sight:
<taxonomicName id="4C774D54FFD7F95F6E09F8B91DCDA734" box="[1287,1436,1844,1868]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="intemelius">
<emphasis id="B903EAC5FFD7F95F6E09F8B91DCDA734" box="[1287,1436,1844,1868]" italics="true" pageId="60" pageNumber="61">C. intemelius</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C774D54FFD7F95F6A28F8D419DAA708" box="[294,395,1880,1904]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B903EAC5FFD7F95F6A28F8D419DAA708" box="[294,395,1880,1904]" italics="true" pageId="60" pageNumber="61">C. nanus</emphasis>
</taxonomicName>
in the shape of pedipalpal hand (less elongate and more depressed at level of
<emphasis id="B903EAC5FFD7F95F6E08F8D41D17A708" box="[1286,1350,1880,1904]" italics="true" pageId="60" pageNumber="61">ib-isb</emphasis>
and, in lateral view, with a more square profile in
<taxonomicName id="4C774D54FFD7F95F6960F8F11B50A7EC" box="[622,769,1916,1940]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="intemelius">
<emphasis id="B903EAC5FFD7F95F6960F8F11B50A7EC" box="[622,769,1916,1940]" italics="true" pageId="60" pageNumber="61">C. intemelius</emphasis>
</taxonomicName>
), in the reduction of the eyes (anterior ones with weak lens, posterior absent in
<taxonomicName id="4C774D54FFD7F95F6A65F82D19AFA7C0" box="[363,510,1952,1976]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="intemelius">
<emphasis id="B903EAC5FFD7F95F6A65F82D19AFA7C0" box="[363,510,1952,1976]" italics="true" pageId="60" pageNumber="61">C. intemelius</emphasis>
</taxonomicName>
), in the shape of spinneret (absent or rarely vestigial in males of
<taxonomicName id="4C774D54FFD7F95F6FDEF82D1D35A7C0" box="[1232,1380,1952,1976]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="intemelius">
<emphasis id="B903EAC5FFD7F95F6FDEF82D1D35A7C0" box="[1232,1380,1952,1976]" italics="true" pageId="60" pageNumber="61">C. intemelius</emphasis>
</taxonomicName>
) and in a smaller average size [length of chela
<quantity id="4C8F9B32FFD7F95F6979F8481B4DA7A4" box="[631,796,1988,2012]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" metricValueMax="5.3" metricValueMin="5.1" pageId="60" pageNumber="61" unit="mm" value="0.52" valueMax="0.53" valueMin="0.51">0.510.53 mm</quantity>
(3)
<quantity id="4C8F9B32FFD7F95F685BF8481BABA7A4" box="[853,1018,1988,2012]" metricMagnitude="-4" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="5.699999999999999" metricValueMin="5.6000000000000005" pageId="60" pageNumber="61" unit="mm" value="0.565" valueMax="0.57" valueMin="0.56">0.560.57 mm</quantity>
(Ƥ) in
<taxonomicName id="4C774D54FFD7F95F6F5CF8491CB6A7A4" box="[1106,1255,1988,2012]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="intemelius">
<emphasis id="B903EAC5FFD7F95F6F5CF8491CB6A7A4" box="[1106,1255,1988,2012]" italics="true" pageId="60" pageNumber="61">C. intemelius</emphasis>
</taxonomicName>
,
<quantity id="4C8F9B32FFD7F95F6FF9F8481DCDA7A4" box="[1271,1436,1988,2012]" metricMagnitude="-4" metricUnit="m" metricValue="5.85" metricValueMax="6.5" metricValueMin="5.2" pageId="60" pageNumber="61" unit="mm" value="0.585" valueMax="0.65" valueMin="0.52">0.520.65 mm</quantity>
(3),
<quantity id="4C8F9B32FFD6F95E6BC2FF1B193EA0D7" box="[204,367,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="6.45" metricValueMax="7.3" metricValueMin="5.6" pageId="61" pageNumber="62" unit="mm" value="0.645" valueMax="0.73" valueMin="0.56">0.560.73 mm</quantity>
(Ƥ) in
<taxonomicName id="4C774D54FFD6F95E6ACFFF141A74A0C8" box="[449,549,152,176]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="B903EAC5FFD6F95E6ACFFF141A74A0C8" box="[449,549,152,176]" italics="true" pageId="61" pageNumber="62">C. nanus</emphasis>
</taxonomicName>
]. It differs from
<taxonomicName id="4C774D54FFD6F95E69EFFF141BD1A0C8" box="[737,896,152,176]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="tyrrhenicus">
<emphasis id="B903EAC5FFD6F95E69EFFF141BD1A0C8" box="[737,896,152,176]" italics="true" pageId="61" pageNumber="62">C. tyrrhenicus</emphasis>
</taxonomicName>
(which seems to be absent from the Province of Imperia) in the shape of the pedipalpal hand (with a less square profile in
<taxonomicName id="4C774D54FFD6F95E68D4FF311C3FA0AC" box="[986,1134,188,212]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="intemelius">
<emphasis id="B903EAC5FFD6F95E68D4FF311C3FA0AC" box="[986,1134,188,212]" italics="true" pageId="61" pageNumber="62">C. intemelius</emphasis>
</taxonomicName>
), the reduction of the eyes and the shape of the spinneret in males.
<taxonomicName id="4C774D54FFD6F95E695BFF6C1AB9A080" box="[597,744,224,248]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="intemelius">
<emphasis id="B903EAC5FFD6F95E695BFF6C1AB9A080" box="[597,744,224,248]" italics="true" pageId="61" pageNumber="62">C. intemelius</emphasis>
</taxonomicName>
differs from the epigean
<taxonomicName id="4C774D54FFD6F95E6F0CFF6C1C06A080" box="[1026,1111,224,248]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ligur">
<emphasis id="B903EAC5FFD6F95E6F0CFF6C1C06A080" box="[1026,1111,224,248]" italics="true" pageId="61" pageNumber="62">C. ligur</emphasis>
</taxonomicName>
, from Baiardo, chiefly in the presence of an isolated subapical tooth (
<emphasis id="B903EAC5FFD6F95E6953FE881A22A164" box="[605,627,260,284]" italics="true" pageId="61" pageNumber="62">di</emphasis>
) on the movable cheliceral finger (absent in
<taxonomicName id="4C774D54FFD6F95E6F7EFE891C96A164" box="[1136,1223,260,284]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="ligur">
<emphasis id="B903EAC5FFD6F95E6F7EFE891C96A164" box="[1136,1223,260,284]" italics="true" pageId="61" pageNumber="62">C. ligur</emphasis>
</taxonomicName>
). For comparisons of
<taxonomicName id="4C774D54FFD6F95E6BBBFEA41918A138" box="[181,329,296,320]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="intemelius">
<emphasis id="B903EAC5FFD6F95E6BBBFEA41918A138" box="[181,329,296,320]" italics="true" pageId="61" pageNumber="62">C. intemelius</emphasis>
</taxonomicName>
with
<taxonomicName id="4C774D54FFD6F95E6A87FEA419A9A138" box="[393,504,296,320]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="aeneae">
<emphasis id="B903EAC5FFD6F95E6A87FEA419A9A138" box="[393,504,296,320]" italics="true" pageId="61" pageNumber="62">C. aeneae</emphasis>
</taxonomicName>
from eastern Liguria, see Remarks under the latter species.
</paragraph>
</subSubSection>
</treatment>
</document>