450 lines
64 KiB
XML
450 lines
64 KiB
XML
<document id="979395DFCFB099E7DC25FA7B516BF302" ID-CLB-Dataset="7413" ID-DOI="10.11646/zootaxa.4951.1.2" ID-GBIF-Dataset="d7a40f10-4145-499c-97be-3f9bc5088d2f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4655625" ID-ZooBank="557825A0-714A-426A-917F-1C9AB7372C30" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617265349256" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Makarkin, Vladimir N." docDate="2021" docId="03DE878EFFF9C45EFF67FD29C9FB8435" docLanguage="en" docName="zootaxa.4951.1.2.pdf" docOrigin="Zootaxa 4951 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Archiinocellia Handlirsch 1910" docType="treatment" docVersion="7" lastPageNumber="57" masterDocId="FFE7FFF6FFF6C44EFFF0FFA7CC6F8000" masterDocTitle="Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation" masterLastPageNumber="79" masterPageNumber="41" pageNumber="56" updateTime="1698935824968" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="2559DFAB659045D11049ECAF1511D0AC" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="BD945A1A799660B1B6284E020BE68D6E">
|
||
<mods:title id="759DF5C762C39F369C7F3EEB69BFA664">Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="BAE7B9CCADA37C5A2A6496B79C0C60B8" type="personal">
|
||
<mods:role id="E4FA76B1C708568338B7BEFD00D4C212">
|
||
<mods:roleTerm id="A3CB6FC9A8F7897EBEF7FD3BCFAEDABA">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8B3E0CEB00A5FC3196D01AA9FF213377">Archibald, S. Bruce</mods:namePart>
|
||
<mods:nameIdentifier id="8BD624DF8593E10FDD1E7CB3063ABD4D" type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
|
||
<mods:affiliation id="5FDE76EF632E789AE247EAB7D7397591">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada. https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="D39D7863A7857D992F126FE355A50D06" type="personal">
|
||
<mods:role id="5653B0F0A2F582BADE710261D73A44F6">
|
||
<mods:roleTerm id="71127F5905E0F1C86FD9DB2BF4606172">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="97B745B852CF8CB84122B4250A370435">Makarkin, Vladimir N.</mods:namePart>
|
||
<mods:affiliation id="4793AB4BEC91492C120A746D5567FFD5">Federal Scientific Centre of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivo-</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="8E89D7ABA1351F2F4ED12CF67A2C3738">text</mods:typeOfResource>
|
||
<mods:relatedItem id="E4AD49C01401EC4B628BA9D4A3656976" type="host">
|
||
<mods:titleInfo id="B9B032F1D1ADDC3EFD2746DC1F8BDEAC">
|
||
<mods:title id="369A0D502B90E0EC0C39287B87E64660">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="7BAC79C22F8DA358210A73E80B5BBC6F">
|
||
<mods:date id="1165516933DFFFB1218CA4036BA7D5B9">2021</mods:date>
|
||
<mods:detail id="5AFA3BD377188EDD6D16A29E98EEC9FB" type="pubDate">
|
||
<mods:number id="97D009AB45F5F7EC112BDFFD9CEB4C8A">2021-04-01</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="68B389EFF1E0C474458D3CB4B42569DC" type="volume">
|
||
<mods:number id="DCE88DF4A42F3ABB2A773C4A08F5CC1C">4951</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="C25556ACC38C5128AEBAA13E81501D2A" type="issue">
|
||
<mods:number id="534CB405320D266D97084552758AD8CF">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="63CAB54723E78A4FDAC8BB3BF39AC600" unit="page">
|
||
<mods:start id="DF28E7D2AFCB0B20EE1D0920678A7A76">41</mods:start>
|
||
<mods:end id="BE9AC7A48349AC2A1C1095821F12BFFE">79</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="9FE9337656D6E1B0C702326D23A16F3B">journal article</mods:classification>
|
||
<mods:identifier id="CB586C73D64FAF5340D40D1F533D57CC" type="CLB-Dataset">7413</mods:identifier>
|
||
<mods:identifier id="C69D67C5B40DCDE1EF692A7E76051A03" type="DOI">10.11646/zootaxa.4951.1.2</mods:identifier>
|
||
<mods:identifier id="8A43FE7A85ED8DEF9D7503034B436AA3" type="GBIF-Dataset">d7a40f10-4145-499c-97be-3f9bc5088d2f</mods:identifier>
|
||
<mods:identifier id="0E0ED68ADCDCC4CC2AB8C2F62E4E1558" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="2810B5FBC0CE71994A1BAE2800B436D6" type="Zenodo-Dep">4655625</mods:identifier>
|
||
<mods:identifier id="A98F4F625C28F09B9F4B28817C6293D7" type="ZooBank">557825A0-714A-426A-917F-1C9AB7372C30</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03DE878EFFF9C45EFF67FD29C9FB8435" ID-DOI="http://doi.org/10.5281/zenodo.4681527" ID-GBIF-Taxon="180999341" ID-Zenodo-Dep="4681527" LSID="urn:lsid:plazi:treatment:03DE878EFFF9C45EFF67FD29C9FB8435" httpUri="http://treatment.plazi.org/id/03DE878EFFF9C45EFF67FD29C9FB8435" lastPageId="16" lastPageNumber="57" pageId="15" pageNumber="56">
|
||
<subSubSection id="C36D6513FFF9C441FF67FD29CE8982A9" box="[151,742,654,681]" pageId="15" pageNumber="56" type="nomenclature">
|
||
<paragraph id="8BC83698FFF9C441FF67FD29CE8982A9" blockId="15.[151,742,654,681]" box="[151,742,654,681]" pageId="15" pageNumber="56">
|
||
<heading id="D08081F4FFF9C441FF67FD29CE8982A9" bold="true" box="[151,742,654,681]" fontSize="11" level="1" pageId="15" pageNumber="56" reason="1">
|
||
<emphasis id="B903EA8AFFF9C441FF67FD29CE8982A9" bold="true" box="[151,742,654,681]" pageId="15" pageNumber="56">
|
||
Genus
|
||
<taxonomicName id="4C774D1BFFF9C441FF1EFD29CE1682A8" ID-CoL="62D3X" authority="Handlirsch, 1910" authorityName="Handlirsch" authorityYear="1910" box="[238,633,654,681]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFF9C441FF1EFD29CDF482A9" bold="true" box="[238,411,654,681]" italics="true" pageId="15" pageNumber="56">Archiinocellia</emphasis>
|
||
<bibRefCitation id="EFE64B69FFF9C441FE52FD29CE1682A8" author="Handlirsch, A." box="[418,633,654,680]" pageId="15" pageNumber="56" pagination="93 - 129" refId="ref23134" refString="Handlirsch, A. (1910) Contributions to Canadian Palaeontology. Vol. II. Part III. Canadian fossil insects. V. Insects from the Tertiary lake deposits of the Southern Interior of British Columbia collected by Mr. Lawrence Lambe, in 1906. Memoirs of the Geological Survey of Canada, 12, 93 - 129. https: // doi. org / 10.4095 / 100486" type="journal article" year="1910">Handlirsch, 1910</bibRefCitation>
|
||
</taxonomicName>
|
||
, sit. nov.
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D6513FFF9C441FF67FD71CFB183C9" pageId="15" pageNumber="56" type="diagnosis">
|
||
<paragraph id="8BC83698FFF9C441FF67FD71CFB183C9" blockId="15.[151,1437,726,2013]" pageId="15" pageNumber="56">
|
||
<emphasis id="B903EA8AFFF9C441FF67FD71CD7D82F0" bold="true" box="[151,274,726,752]" pageId="15" pageNumber="56">Diagnosis.</emphasis>
|
||
Relatively small raphidiids (forewing up to
|
||
<quantity id="4C8F9B7DFFF9C441FCF8FD70CF2582F1" box="[776,842,727,753]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="56" unit="mm" value="8.0">8 mm</quantity>
|
||
long), distinguished from other genera of the family by the following combination of character states. Fore- and hind wings: (1) ScP short (length from termination to pterostigma greater than that of pterostigma) [relatively long ScP (length from termination to pterostigma approximately equal to or less than that of short pterostigma) in
|
||
<taxonomicName id="4C774D1BFFF9C441FCD2FCE4CFAD835D" authorityName="Cockerell" authorityYear="1907" box="[802,962,835,861]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFF9C441FCD2FCE4CFAD835D" box="[802,962,835,861]" italics="true" pageId="15" pageNumber="56">Megaraphidia</emphasis>
|
||
</taxonomicName>
|
||
]; (2) pterostigma elongate (4.5–5.1 times longer than width measured along to costal margin) [short in
|
||
<taxonomicName id="4C774D1BFFF9C441FC97FCC0C8688381" authorityName="Cockerell" authorityYear="1907" box="[871,1031,871,897]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFF9C441FC97FCC0C8688381" box="[871,1031,871,897]" italics="true" pageId="15" pageNumber="56">Megaraphidia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C774D1BFFF9C441FBE8FCC0C8D98381" box="[1048,1206,871,897]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFF9C441FBE8FCC0C8D98381" box="[1048,1206,871,897]" italics="true" pageId="15" pageNumber="56">Archiraphidia</emphasis>
|
||
</taxonomicName>
|
||
]; (3) 3ra-rp located within the pterostigma; (4) RP1 twice forked, proximal fork proximad level of 3ra-rp. Hind wing: (5) two
|
||
<emphasis id="B903EA8AFFF9C441FACFFC2CC90C83A5" box="[1343,1379,907,933]" italics="true" pageId="15" pageNumber="56">doi</emphasis>
|
||
[one in
|
||
<taxonomicName id="4C774D1BFFF9C441FF43FC08CD3E83C9" box="[179,337,943,969]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFF9C441FF43FC08CD3E83C9" box="[179,337,943,969]" italics="true" pageId="15" pageNumber="56">Archiraphidia</emphasis>
|
||
</taxonomicName>
|
||
]; (6) CuA with one branch [three in
|
||
<emphasis id="B903EA8AFFF9C441FD1CFC08CFA183C9" box="[748,974,943,969]" italics="true" pageId="15" pageNumber="56">Florissontoraphidia</emphasis>
|
||
].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D6513FFF9C441FF37FC74CFC38459" pageId="15" pageNumber="56" type="type_taxon">
|
||
<paragraph id="8BC83698FFF9C441FF37FC74CF5C83ED" blockId="15.[151,1437,726,2013]" box="[199,819,978,1005]" pageId="15" pageNumber="56">
|
||
<emphasis id="B903EA8AFFF9C441FF37FC74CD0D83EC" bold="true" box="[199,354,978,1005]" pageId="15" pageNumber="56">
|
||
<typeStatus id="54CC883AFFF9C441FF37FC74CD6D83ED" box="[199,258,979,1005]" pageId="15" pageNumber="56">Type</typeStatus>
|
||
species.
|
||
</emphasis>
|
||
<taxonomicName id="4C774D1BFFF9C441FE99FC74CEE583ED" ID-CoL="G93B" authorityName="Handlirsch" authorityYear="1910" box="[361,650,979,1005]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="oligoneura">
|
||
<emphasis id="B903EA8AFFF9C441FE99FC74CEE583ED" box="[361,650,979,1005]" italics="true" pageId="15" pageNumber="56">Archiinocellia oligoneura</emphasis>
|
||
</taxonomicName>
|
||
, by monotypy.
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFF9C441FF37FC51C85B8411" blockId="15.[151,1437,726,2013]" box="[199,1076,1014,1041]" pageId="15" pageNumber="56">
|
||
<emphasis id="B903EA8AFFF9C441FF37FC51CDFC8410" bold="true" box="[199,403,1014,1040]" pageId="15" pageNumber="56">Species included.</emphasis>
|
||
<taxonomicName id="4C774D1BFFF9C441FE69FC50CE5B8411" ID-CoL="G93B" authorityName="Handlirsch" authorityYear="1910" box="[409,564,1015,1041]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="oligoneura">
|
||
<emphasis id="B903EA8AFFF9C441FE69FC50CE5B8411" box="[409,564,1015,1041]" italics="true" pageId="15" pageNumber="56">A. oligoneura</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C774D1BFFF9C441FDB1FC50CFC88411" authority="(Engel, 2011)" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[577,935,1015,1041]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="protomaculata" status="comb. nov.">
|
||
<emphasis id="B903EA8AFFF9C441FDB1FC50CF6A8411" box="[577,773,1015,1041]" italics="true" pageId="15" pageNumber="56">A. protomaculata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE64B69FFF9C441FCE3FC50CFF08411" author="Engel, M. S." box="[787,927,1015,1041]" pageId="15" pageNumber="56" pagination="77 - 87" refId="ref22829" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. https: // doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel, 2011</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A23057F1FFF9C441FC42FC50C85B8411" box="[946,1076,1015,1041]" pageId="15" pageNumber="56" rank="species">
|
||
<emphasis id="B903EA8AFFF9C441FC42FC50C8418411" bold="true" box="[946,1070,1015,1041]" pageId="15" pageNumber="56">comb. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFF9C441FF37FBBDCFC38459" blockId="15.[151,1437,726,2013]" pageId="15" pageNumber="56">
|
||
<emphasis id="B903EA8AFFF9C441FF37FBBDCD358434" bold="true" box="[199,346,1050,1076]" pageId="15" pageNumber="56">Occurrence.</emphasis>
|
||
Ypresian of the Okanagan Highlands at Horsefly,
|
||
<collectingRegion id="49B3F87AFFF9C441FC57FBBCC8018435" box="[935,1134,1051,1077]" country="Canada" name="British Columbia" pageId="15" pageNumber="56">British Columbia</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3607608FFF9C441FB8CFBBCC8BC8435" box="[1148,1235,1051,1077]" name="Canada" pageId="15" pageNumber="56">Canada</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3607608FFF9C441FB12FBBCC9558435" box="[1250,1338,1051,1077]" name="Canada" pageId="15" pageNumber="56">Canada</collectingCountry>
|
||
; the late Ypresian Green River Formation,
|
||
<collectingRegion id="49B3F87AFFF9C441FDE4FB98CE108459" box="[532,639,1087,1113]" country="United States of America" name="Colorado" pageId="15" pageNumber="56">Colorado</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3607608FFF9C441FD7AFB98CFC78459" box="[650,936,1087,1113]" name="United States of America" pageId="15" pageNumber="56">United States of America</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D6513FFF9C45EFF37FBC4C9FB8435" lastPageId="16" lastPageNumber="57" pageId="15" pageNumber="56" type="discussion">
|
||
<paragraph id="8BC83698FFF9C441FF37FBC4CE40850D" blockId="15.[151,1437,726,2013]" pageId="15" pageNumber="56">
|
||
<emphasis id="B903EA8AFFF9C441FF37FBC4CD54847D" bold="true" box="[199,315,1123,1149]" pageId="15" pageNumber="56">Remarks.</emphasis>
|
||
The genus was originally considered to be of unknown family (
|
||
<bibRefCitation id="EFE64B69FFF9C441FBFEFBC4C8BA847D" author="Handlirsch, A." box="[1038,1237,1123,1149]" pageId="15" pageNumber="56" pagination="93 - 129" refId="ref23134" refString="Handlirsch, A. (1910) Contributions to Canadian Palaeontology. Vol. II. Part III. Canadian fossil insects. V. Insects from the Tertiary lake deposits of the Southern Interior of British Columbia collected by Mr. Lawrence Lambe, in 1906. Memoirs of the Geological Survey of Canada, 12, 93 - 129. https: // doi. org / 10.4095 / 100486" type="journal article" year="1910">Handlirsch, 1910</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE64B69FFF9C441FB11FBC4C9F8847D" author="Martynov, A. V." box="[1249,1431,1122,1149]" pageId="15" pageNumber="56" pagination="233 - 246" refId="ref23966" refString="Martynov, A. V. (1925) To the knowledge of fossil insects from Jurassic beds in Turkestan. 1. Raphidioptera. Bulletin of the Academy of Sciences USSR, Series 6, 19, 233 - 246." type="journal article" year="1925">Martynov, 1925</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE64B69FFF9C441FF67FB20CD2084A2" author="Carpenter, F. M." box="[151,335,1159,1186]" pageId="15" pageNumber="56" pagination="89 - 157" refId="ref21922" refString="Carpenter, F. M. (1936) Revision of the Nearctic Raphidiodea (recent and fossil). Proceedings of the American Academy of Arts and Science, 71, 89 - 157. https: // doi. org / 10.2307 / 20023217" type="journal article" year="1936">Carpenter, 1936</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFE64B69FFF9C441FE91FB20CE2784A1" author="Aspock, H. & Aspock, U. & Rausch, H." box="[353,584,1159,1185]" pageId="15" pageNumber="56" refId="ref20769" refString="Aspock, H., Aspock, U. & Rausch, H. (1991) Die Raphidiopteren der Erde. Eine monographische Darstellung der Systematik, Taxonomie, Biologie, Okologie und Chorologie der rezenten Raphidiopteren der Erde, mit einer zusammenfassenden Ubersicht der fossilen Raphidiopteren (Insecta: Neuropteroidea). 2 Vols. Goecke & Evers, Krefeld, 730 and 550 pp." type="book" year="1991">
|
||
Aspöck
|
||
<emphasis id="B903EA8AFFF9C441FE4EFB20CD8084A1" box="[446,495,1159,1185]" italics="true" pageId="15" pageNumber="56">et al</emphasis>
|
||
. (1991)
|
||
</bibRefCitation>
|
||
assigned it to the
|
||
<taxonomicName id="4C774D1BFFF9C441FCE5FB20CFF384A1" authorityName="Navas" authorityYear="1913" box="[789,924,1159,1185]" class="Insecta" family="Inocelliidae" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="family">Inocelliidae</taxonomicName>
|
||
, but more recent authors have treated it again as
|
||
<taxonomicName id="4C774D1BFFF9C441FF46FB0CCD3A84C5" authorityName="Navas" authorityYear="1916" box="[182,341,1195,1221]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="order">Raphidioptera</taxonomicName>
|
||
<emphasis id="B903EA8AFFF9C441FEACFB0CCD9884C5" box="[348,503,1195,1221]" italics="true" pageId="15" pageNumber="56">incertae sedis</emphasis>
|
||
(
|
||
<emphasis id="B903EA8AFFF9C441FDF7FB0CCE4684C4" box="[519,553,1195,1220]" italics="true" pageId="15" pageNumber="56">e.g</emphasis>
|
||
.,
|
||
<bibRefCitation id="EFE64B69FFF9C441FDCEFB0CCEB284C5" author="Oswald, J. D." box="[574,733,1195,1221]" pageId="15" pageNumber="56" pagination="154 - 163" refId="ref24369" refString="Oswald, J. D. (1990) Raphidioptera. In: Grimaldi, D. A. (Ed.), Insects from the Santana Formation, Lower Cretaceous, of Brazil. Bulletin of the American Museum of Natural History, 195, 154 - 163." type="journal article" year="1990">Oswald, 1990</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE64B69FFF9C441FD19FB0CCFCF84C5" author="Carpenter, F. M." box="[745,928,1195,1221]" pageId="15" pageNumber="56" pagination="1 - 655" refId="ref22044" refString="Carpenter, F. M. (1992) Superclass Hexapoda. In: Kaesler, R. L. (Ed.), Treatise on Invertebrate Paleontology. Part R. Arthropoda 4. Vols. 3 & 4. The Geological Society of America, Boulder, Colorado & University of Kansas, Lawrence, Kansas, pp. 1 - 655." type="book chapter" year="1992">Carpenter, 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE64B69FFF9C441FC5CFB0CC87284C5" author="Nel, A." box="[940,1053,1195,1221]" pageId="15" pageNumber="56" pagination="99 - 108" refId="ref24327" refString="Nel, A. (1993) Nouveaux Raphidiopteres fossiles du Cenozoique de France et d'Espagne (Raphidioptera, Raphidiidae, Inocelliidae). Ecole Pratique des Hautes Etudes, Biologie et Evolution des Insectes, 6, 99 - 108." type="journal article" year="1993">Nel, 1993</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE64B69FFF9C441FBD9FB0CC8DB84C5" author="Engel, M. S." box="[1065,1204,1195,1221]" pageId="15" pageNumber="56" pagination="1 - 22" refId="ref22701" refString="Engel, M. S. (2002) The smallest snakefly (Raphidioptera: Mesoraphidiidae): A new species in Cretaceous amber from Myanmar, with a catalog of fossil snakeflies. American Museum Novitates, 3363, 1 - 22." type="journal article" year="2002">Engel, 2002</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE64B69FFF9C441FB4FFB0CC89384C5" author="Engel, M. S." box="[1215,1276,1195,1221]" pageId="15" pageNumber="56" pagination="211 - 214" refId="ref22782" refString="Engel, M. S. (2009) A Miocene snakefly from Stewart Valley, Nevada (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 112, 211 - 214. https: // doi. org / 10.1660 / 062.112.0409" type="journal article" year="2009">2009</bibRefCitation>
|
||
). Here, we assign it to the
|
||
<taxonomicName id="4C774D1BFFF9C441FEDDFB68CDDB84E9" authorityName="Latreille" authorityYear="1810" box="[301,436,1231,1257]" class="Insecta" family="Raphidiidae" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="family">Raphidiidae</taxonomicName>
|
||
by venation characteristic of the family, including the placement of crossvein 2ra-scp at the proximal end of the pterostigma.
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFF9C441FF37FAB0C8818675" blockId="15.[151,1437,726,2013]" pageId="15" pageNumber="56">
|
||
<taxonomicName id="4C774D1BFFF9C441FF37FAB0CE7E8531" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[199,529,1303,1329]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFF9C441FF37FAB0CE7E8531" box="[199,529,1303,1329]" italics="true" pageId="15" pageNumber="56">Archiinocellia protomaculata</emphasis>
|
||
</taxonomicName>
|
||
from the Green River Formation was originally assigned to
|
||
<taxonomicName id="4C774D1BFFF9C441FB44FAB0C8908531" authorityName="Navas" authorityYear="1914" box="[1204,1279,1303,1329]" class="Insecta" family="Raphidiidae" genus="Agulla" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFF9C441FB44FAB0C8908531" box="[1204,1279,1303,1329]" italics="true" pageId="15" pageNumber="56">Agulla</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFE64B69FFF9C441FAFDFAB0C9F78531" author="Engel, M. S." box="[1293,1432,1303,1329]" pageId="15" pageNumber="56" pagination="77 - 87" refId="ref22829" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. https: // doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel, 2011</bibRefCitation>
|
||
:
|
||
<figureCitation id="134C2A1DFFF9C441FF67FA9CCC938555" box="[151,252,1339,1365]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStartId-0="3.[151,250,1381,1405]" captionStartId-1="4.[151,250,1943,1967]" captionStartId-2="5.[151,250,1986,2010]" captionStartId-3="6.[151,250,1943,1967]" captionStartId-4="9.[151,250,1943,1967]" captionStartId-5="10.[151,250,1930,1954]" captionStartId-6="11.[151,250,1581,1605]" captionStartId-7="12.[151,250,1620,1644]" captionStartId-8="13.[151,250,1943,1967]" captionTargetBox-0="[152,1436,777,1353]" captionTargetBox-1="[206,1380,192,1913]" captionTargetBox-2="[151,1436,1353,1956]" captionTargetBox-3="[182,1407,190,1909]" captionTargetBox-4="[187,1401,181,1900]" captionTargetBox-5="[164,1426,181,1893]" captionTargetBox-6="[151,1430,1137,1553]" captionTargetBox-7="[151,1436,664,1596]" captionTargetBox-8="[275,1316,200,1905]" captionTargetId-0="figure-221@3.[152,1436,777,1357]" captionTargetId-1="figure-26@4.[195,1393,181,1919]" captionTargetId-2="figure-459@5.[151,1436,1353,1963]" captionTargetId-3="figure-26@6.[165,1423,181,1919]" captionTargetId-4="figure-19@9.[187,1401,181,1919]" captionTargetId-5="figure-26@10.[151,1436,181,1906]" captionTargetId-6="figure-445@11.[151,1436,1137,1558]" captionTargetId-7="figure-176@12.[151,1436,664,1596]" captionTargetId-8="figure-19@13.[259,1329,181,1919]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="13" captionText-0="FIGURE 1. Megaraphidia antiquissima sp. nov., holotype RBCM P1555, specimen as preserved. Scale bar = 5 mm." captionText-1="FIGURE 2. Wing venation of Megaraphidia antiquissima sp. nov., holotype RBCM P1555. A, right forewing, B, left forewing; C, right forewing; D, left hind wing. Scale bar = 3 mm (all to scale)." captionText-2="FIGURE 3. Megaraphidia ootsa sp. nov., holotype RBCM P1556, specimen as preserved. Scale bars = 5 mm." captionText-3="FIGURE 4. Wing venation of Megaraphidia ootsa sp. nov., holotype RBCM P1556. A, right forewing, B, left forewing; C, right hind wing; D, left hind wing. Scale bar = 3 mm (all to scale)." captionText-4="FIGURE 5. Megaraphidia klondika sp. nov., holotype. A, counterpart UWBM 97073; B, part SR03-01-01; C, forewing venation; D, hind wing venation. Scale bar = 3 mm (all to scale)." captionText-5="FIGURE 6. Megaraphidia klondika sp. nov., specimen SRUI99-71-63. A, part; B, counterpart; C, drawing of the hind wing venation. Scale bar = 3 mm (all to scale)." captionText-6="FIGURE 7. Megaraphidia klondika sp. nov., specimen SR16-004-001-01.01. A, specimen as preserved; B, drawing. Scale bar = 3 mm (both to scale)." captionText-7="FIGURE 8. Megaraphidia hopkinsi sp. nov., holotype UWBM77544. A, specimen as preserved; B, head. gu, gula. Scale bars = 5 mm (A), 1 mm (B)." captionText-8="FIGURE 9. Wing venation of Megaraphidia hopkinsi sp. nov., holotype UWBM77544. A, right forewing; B, left hind wing; C, right forewing; D, left hind wing. Scale bar = 3 mm (all to scale)." figureDoi-0="http://doi.org/10.5281/zenodo.4655627" figureDoi-1="http://doi.org/10.5281/zenodo.4655631" figureDoi-2="http://doi.org/10.5281/zenodo.4655635" figureDoi-3="http://doi.org/10.5281/zenodo.4655637" figureDoi-4="http://doi.org/10.5281/zenodo.4655641" figureDoi-5="http://doi.org/10.5281/zenodo.4655647" figureDoi-6="http://doi.org/10.5281/zenodo.4655655" figureDoi-7="http://doi.org/10.5281/zenodo.4655657" figureDoi-8="http://doi.org/10.5281/zenodo.4655661" httpUri-0="https://zenodo.org/record/4655627/files/figure.png" httpUri-1="https://zenodo.org/record/4655631/files/figure.png" httpUri-2="https://zenodo.org/record/4655635/files/figure.png" httpUri-3="https://zenodo.org/record/4655637/files/figure.png" httpUri-4="https://zenodo.org/record/4655641/files/figure.png" httpUri-5="https://zenodo.org/record/4655647/files/figure.png" httpUri-6="https://zenodo.org/record/4655655/files/figure.png" httpUri-7="https://zenodo.org/record/4655657/files/figure.png" httpUri-8="https://zenodo.org/record/4655661/files/figure.png" pageId="15" pageNumber="56">Figs 1–9</figureCitation>
|
||
). We believe that this is incorrect by its wing venation, which possesses at least three important character states not present in that genus. First, the basal 1r-m in the hind wing of this species is long and subparallel to R (clearly visible in
|
||
<figureCitation id="134C2A1DFFF9C441FE93FA24CDA8859D" box="[355,455,1411,1437]" captionStart="FIGURE 17" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[167,1424,191,1902]" captionTargetId="figure-26@24.[151,1436,181,1919]" captionTargetPageId="24" captionText="FIGURE 17. Venation of Archiinocellia protomaculata (Engel, 2011). A, right forewing of UCM 61231a (female); B, right hind wing of the same specimen; C, right forewing of UCM 62523; D, right hind wing of the same specimen. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655688" httpUri="https://zenodo.org/record/4655688/files/figure.png" pageId="15" pageNumber="56">Fig. 17D</figureCitation>
|
||
and the
|
||
<typeStatus id="54CC883AFFF9C441FDD8FA24CEE7859D" box="[552,648,1411,1437]" pageId="15" pageNumber="56" type="paratype">paratype</typeStatus>
|
||
USNM 31752), but it is crossvein-like in all known species of
|
||
<taxonomicName id="4C774D1BFFF9C441FAA2FA24C9F2859D" authorityName="Navas" authorityYear="1914" box="[1362,1437,1411,1437]" class="Insecta" family="Raphidiidae" genus="Agulla" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFF9C441FAA2FA24C9F2859D" box="[1362,1437,1411,1437]" italics="true" pageId="15" pageNumber="56">Agulla</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<emphasis id="B903EA8AFFF9C441FF3DFA00CC8185C0" box="[205,238,1447,1472]" italics="true" pageId="15" pageNumber="56">e.g</emphasis>
|
||
.,
|
||
<bibRefCitation id="EFE64B69FFF9C441FEF5FA00CDD285C1" author="Carpenter, F. M." box="[261,445,1447,1473]" pageId="15" pageNumber="56" pagination="89 - 157" refId="ref21922" refString="Carpenter, F. M. (1936) Revision of the Nearctic Raphidiodea (recent and fossil). Proceedings of the American Academy of Arts and Science, 71, 89 - 157. https: // doi. org / 10.2307 / 20023217" type="journal article" year="1936">Carpenter, 1936</bibRefCitation>
|
||
:
|
||
<figureCitation id="134C2A1DFFF9C441FE3AFA00CE6485C1" box="[458,523,1447,1473]" captionStart="FIGURE 1" captionStartId="3.[151,250,1381,1405]" captionTargetBox="[152,1436,777,1353]" captionTargetId="figure-221@3.[152,1436,777,1357]" captionTargetPageId="3" captionText="FIGURE 1. Megaraphidia antiquissima sp. nov., holotype RBCM P1555, specimen as preserved. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655627" httpUri="https://zenodo.org/record/4655627/files/figure.png" pageId="15" pageNumber="56">Fig. 1</figureCitation>
|
||
). Secondly, the crossvein 3ra-rp in both wings is located within the pterostigma in this species, whereas it is normally located distad the pterostigma in all known species of
|
||
<taxonomicName id="4C774D1BFFF9C441FB5DFA6CC89785E5" authorityName="Navas" authorityYear="1914" box="[1197,1272,1483,1509]" class="Insecta" family="Raphidiidae" genus="Agulla" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFF9C441FB5DFA6CC89785E5" box="[1197,1272,1483,1509]" italics="true" pageId="15" pageNumber="56">Agulla</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B903EA8AFFF9C441FAF8FA6CC94585E4" box="[1288,1322,1483,1508]" italics="true" pageId="15" pageNumber="56">e.g</emphasis>
|
||
., see
|
||
<bibRefCitation id="EFE64B69FFF9C441FA9BFA6CCD498609" author="Carpenter, F. M." pageId="15" pageNumber="56" pagination="89 - 157" refId="ref21922" refString="Carpenter, F. M. (1936) Revision of the Nearctic Raphidiodea (recent and fossil). Proceedings of the American Academy of Arts and Science, 71, 89 - 157. https: // doi. org / 10.2307 / 20023217" type="journal article" year="1936">Carpenter, 1936</bibRefCitation>
|
||
:
|
||
<figureCitation id="134C2A1DFFF9C441FEC1FA48CD1D8609" box="[305,370,1519,1545]" captionStart="FIGURE 1" captionStartId="3.[151,250,1381,1405]" captionTargetBox="[152,1436,777,1353]" captionTargetId="figure-221@3.[152,1436,777,1357]" captionTargetPageId="3" captionText="FIGURE 1. Megaraphidia antiquissima sp. nov., holotype RBCM P1555, specimen as preserved. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655627" httpUri="https://zenodo.org/record/4655627/files/figure.png" pageId="15" pageNumber="56">Fig. 1</figureCitation>
|
||
); very rarely, adventitiously at the distal end of pterostigma (
|
||
<emphasis id="B903EA8AFFF9C441FBD5FA48C8298608" box="[1061,1094,1519,1544]" italics="true" pageId="15" pageNumber="56">e.g</emphasis>
|
||
., Wognum, 1959: Figs 20, 35;
|
||
<bibRefCitation id="EFE64B69FFF9C441FF67F9B4CD58862D" author="Aspock, U." box="[151,311,1555,1581]" pageId="15" pageNumber="56" pagination="1 - 4" refId="ref20929" refString="Aspock, U. (1987) Agulla faulkneri n. sp. - eine neue Raphidiiden-Spezies aus Kalifornien (Neuropteroidea: Raphidioptera: Raphidiidae). Z eitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 39, 1 - 4." type="journal article" year="1987">Aspöck, 1987</bibRefCitation>
|
||
:
|
||
<figureCitation id="134C2A1DFFF9C441FEB3F9B4CDEB862D" box="[323,388,1555,1581]" captionStart="FIGURE 1" captionStartId="3.[151,250,1381,1405]" captionTargetBox="[152,1436,777,1353]" captionTargetId="figure-221@3.[152,1436,777,1357]" captionTargetPageId="3" captionText="FIGURE 1. Megaraphidia antiquissima sp. nov., holotype RBCM P1555, specimen as preserved. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655627" httpUri="https://zenodo.org/record/4655627/files/figure.png" pageId="15" pageNumber="56">Fig. 1</figureCitation>
|
||
); and only abnormally within the pterostigma in one of the wings (
|
||
<emphasis id="B903EA8AFFF9C441FB8DF9B4C8F1862C" box="[1149,1182,1555,1580]" italics="true" pageId="15" pageNumber="56">e.g</emphasis>
|
||
., Wognum, 1959: Fig. 21). Thirdly, ScP is very short in this species, whereas it is long in all species of
|
||
<taxonomicName id="4C774D1BFFF9C441FBEBF990C8098651" authorityName="Navas" authorityYear="1914" box="[1051,1126,1591,1617]" class="Insecta" family="Raphidiidae" genus="Agulla" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFF9C441FBEBF990C8098651" box="[1051,1126,1591,1617]" italics="true" pageId="15" pageNumber="56">Agulla</emphasis>
|
||
</taxonomicName>
|
||
(from its termination to the pterostigma is nearly equal to the length of pterostigma or slightly shorter in the species of
|
||
<taxonomicName id="4C774D1BFFF9C441FB63F9FCC8B18675" authorityName="Navas" authorityYear="1914" box="[1171,1246,1627,1653]" class="Insecta" family="Raphidiidae" genus="Agulla" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFF9C441FB63F9FCC8B18675" box="[1171,1246,1627,1653]" italics="true" pageId="15" pageNumber="56">Agulla</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFF9C441FF37F9D8CFBA86BD" blockId="15.[151,1437,726,2013]" pageId="15" pageNumber="56">
|
||
We assigned
|
||
<taxonomicName id="4C774D1BFFF9C441FEA9F9D8CECD8699" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[345,674,1663,1689]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFF9C441FEA9F9D8CECD8699" box="[345,674,1663,1689]" italics="true" pageId="15" pageNumber="56">Archiinocellia protomaculata</emphasis>
|
||
</taxonomicName>
|
||
to this genus as its venation is very similar to the preserved venation of
|
||
<taxonomicName id="4C774D1BFFF9C441FF46F904CD3F86BD" authorityName="Handlirsch" authorityYear="1910" box="[182,336,1699,1725]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="oligoneura">
|
||
<emphasis id="B903EA8AFFF9C441FF46F904CD3F86BD" box="[182,336,1699,1725]" italics="true" pageId="15" pageNumber="56">A. oligoneura</emphasis>
|
||
</taxonomicName>
|
||
, agreeing with all character states of the genus diagnosis.
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFF9C45EFF37F960CEAE8219" blockId="15.[151,1437,726,2013]" lastBlockId="16.[151,1437,150,1077]" lastPageId="16" lastPageNumber="57" pageId="15" pageNumber="56">
|
||
Of extant genera, the venation of
|
||
<taxonomicName id="4C774D1BFFF9C441FDB8F960CFFB86E1" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[584,916,1735,1761]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFF9C441FDB8F960CFFB86E1" box="[584,916,1735,1761]" italics="true" pageId="15" pageNumber="56">Archiinocellia protomaculata</emphasis>
|
||
</taxonomicName>
|
||
is most similar to that of
|
||
<taxonomicName id="4C774D1BFFF9C441FB4EF960C9F786E1" authority="Navas, 1916" authorityName="Navas" authorityYear="1916" box="[1214,1432,1735,1761]" class="Insecta" family="Raphidiidae" genus="Alena" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFF9C441FB4EF960C89086E1" box="[1214,1279,1735,1761]" italics="true" pageId="15" pageNumber="56">Alena</emphasis>
|
||
<bibRefCitation id="EFE64B69FFF9C441FAF8F960C9F786E1" author="Navas, L." box="[1288,1432,1735,1761]" pageId="15" pageNumber="56" pagination="507 - 513" refId="ref24287" refString="Navas, L. (1916) Notas sobre el orden de los Rafidiopteros (Ins.). Memorias de la Real Academia de Ciencias y Artes de Barcelona, Series 3, 12, 507 - 513." type="journal article" year="1916">Navás, 1916</bibRefCitation>
|
||
</taxonomicName>
|
||
, the southern-most genus of
|
||
<taxonomicName id="4C774D1BFFF9C441FE26F94CCE328705" authorityName="Latreille" authorityYear="1810" box="[470,605,1771,1797]" class="Insecta" family="Raphidiidae" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="family">Raphidiidae</taxonomicName>
|
||
in the Western Hemisphere, occurring in south-western
|
||
<collectingCountry id="F3607608FFF9C441FB14F94CCC988729" name="United States of America" pageId="15" pageNumber="56">United States of America</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3607608FFF9C441FEDEF8A8CDEB8729" box="[302,388,1807,1833]" name="Mexico" pageId="15" pageNumber="56">Mexico</collectingCountry>
|
||
(south to
|
||
<collectingRegion id="49B3F87AFFF9C441FE05F8A8CE3E872A" box="[501,593,1807,1834]" country="Mexico" name="Chiapas" pageId="15" pageNumber="56">Chiapas</collectingRegion>
|
||
) with ten species (
|
||
<bibRefCitation id="EFE64B69FFF9C441FCD2F8A8CFAD8729" author="Aspock, U." box="[802,962,1806,1833]" pageId="15" pageNumber="56" pagination="537 - 546" refId="ref20837" refString="Aspock, U. (1975) The present state of knowledge on the Raphidioptera of America (Insecta: Neuropteroidea). Polskie Pismo Entomologiczne, 45, 537 - 546." type="journal article" year="1975">Aspöck, 1975</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE64B69FFF9C441FC3DF8A8C8898729" author="Aspock, U. & Aspock, H." box="[973,1254,1807,1833]" pageId="15" pageNumber="56" pagination="53 - 58" refId="ref21256" refString="Aspock, U. & Aspock, H. (2013) Alena Navas, 1916 - the dethroned genus and Alena (Aztekoraphidia) michoacana sp. n. from Mexico (Neuropterida: Raphidioptera: Raphidiidae). Deutsche Entomologische Zeitschrift, 60, 53 - 58." type="journal article" year="2013">Aspöck & Aspöck, 2013</bibRefCitation>
|
||
). A combination of the following features is characteristic of only five species of this genus among extant
|
||
<taxonomicName id="4C774D1BFFF9C441FB87F894C96C874D" authorityName="Latreille" authorityYear="1810" box="[1143,1283,1843,1869]" class="Insecta" family="Raphidiidae" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="family">Raphidiidae</taxonomicName>
|
||
: (A) the basal crossvein 1r-m in the hind wing is long, subparallel to R; (B) very short ScP (from its termination to pterostigma is much longer than the length of pterostigma), and (C) the proximal position of the crossvein 3ra-rp (
|
||
<emphasis id="B903EA8AFFF9C441FAEAF8DCC9598794" box="[1306,1334,1915,1940]" italics="true" pageId="15" pageNumber="56">i.e</emphasis>
|
||
., located within the pterostigma) in both wings. The venation of
|
||
<taxonomicName id="4C774D1BFFF9C441FCE0F838CFBA87B9" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[784,981,1951,1977]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFF9C441FCE0F838CFBA87B9" box="[784,981,1951,1977]" italics="true" pageId="15" pageNumber="56">A. protomaculata</emphasis>
|
||
</taxonomicName>
|
||
shares all of these character states. It is most similar to that of some species of the Mexican genus
|
||
<taxonomicName id="4C774D1BFFF9C441FCEBF864CF3387DD" authorityName="Navas" authorityYear="1916" box="[795,860,1987,2013]" class="Insecta" family="Raphidiidae" genus="Alena" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFF9C441FCEBF864CF3387DD" box="[795,860,1987,2013]" italics="true" pageId="15" pageNumber="56">Alena</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B903EA8AFFF9C441FC9BF864CFFB87DC" box="[875,916,1987,2012]" italics="true" pageId="15" pageNumber="56">e.g.</emphasis>
|
||
,
|
||
<taxonomicName id="4C774D1BFFF9C441FC51F864C97587DD" authority="Carpenter, 1958" authorityName="Carpenter" authorityYear="1958" box="[929,1306,1986,2013]" class="Insecta" family="Raphidiidae" genus="Alena" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="americana">
|
||
<emphasis id="B903EA8AFFF9C441FC51F864C80F87DC" box="[929,1120,1987,2013]" italics="true" pageId="15" pageNumber="56">Alena americana</emphasis>
|
||
<bibRefCitation id="EFE64B69FFF9C441FB95F864C97587DD" author="Carpenter, F. M." box="[1125,1306,1986,2013]" pageId="15" pageNumber="56" pagination="52 - 58" refId="ref22007" refString="Carpenter, F. M. (1958) Mexican snake-flies (Neuroptera: Raphidiodea). Psyche, 65, 52 - 58. https: // doi. org / 10.1155 / 1958 / 14859" type="journal article" year="1958">Carpenter, 1958</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C774D1BFFF9C45EFAD4F864CD2780B2" authority="(Navas, 1914 b)" baseAuthorityName="Navas" baseAuthorityYear="1914" class="Insecta" family="Raphidiidae" genus="Alena" kingdom="Animalia" lastPageId="16" lastPageNumber="57" order="Raphidioptera" pageId="15" pageNumber="56" phylum="Arthropoda" rank="species" species="caudata">
|
||
<emphasis id="B903EA8AFFF9C441FAD4F864C9F387DD" box="[1316,1436,1987,2013]" italics="true" pageId="15" pageNumber="56">A. caudata</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE64B69FFE6C45EFF6FFF30CD2F80B2" author="Navas, L." box="[159,320,151,178]" pageId="16" pageNumber="57" pagination="25 - 26" refId="ref24253" refString="Navas, L. (1914 b) Nevropteres nouveaux de l'Amerique du Nord. IIeme serie. Entomologische Zeitschrift (Frankfurt am Main), 28,18 - 20 + 25 - 26." type="book chapter" year="1914">Navás, 1914b</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; see
|
||
<bibRefCitation id="EFE64B69FFE6C45EFE8FFF30CE5880B1" author="Carpenter, F. M." box="[383,567,150,177]" pageId="16" pageNumber="57" pagination="52 - 58" refId="ref22007" refString="Carpenter, F. M. (1958) Mexican snake-flies (Neuroptera: Raphidiodea). Psyche, 65, 52 - 58. https: // doi. org / 10.1155 / 1958 / 14859" type="journal article" year="1958">Carpenter, 1958</bibRefCitation>
|
||
:
|
||
<figureCitation id="134C2A1DFFE6C45EFDB3FF30CEE980B1" box="[579,646,151,177]" captionStart="FIGURE 1" captionStartId="3.[151,250,1381,1405]" captionTargetBox="[152,1436,777,1353]" captionTargetId="figure-221@3.[152,1436,777,1357]" captionTargetPageId="3" captionText="FIGURE 1. Megaraphidia antiquissima sp. nov., holotype RBCM P1555, specimen as preserved. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655627" httpUri="https://zenodo.org/record/4655627/files/figure.png" pageId="16" pageNumber="57">Fig. 1</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFE64B69FFE6C45EFD63FF30CFC380B1" author="Aspock, U. & Aspock, H." box="[659,940,151,177]" pageId="16" pageNumber="57" pagination="705 - 725" refId="ref21026" refString="Aspock, U. & Aspock, H. (1970) Untersuchungen uber die Raphidiopteren Mexikos (Insecta, Neuropteroidea). Polskie Pismo Entomologiczne, 40, 705 - 725." type="journal article" year="1970">Aspöck & Aspöck, 1970</bibRefCitation>
|
||
:
|
||
<figureCitation id="134C2A1DFFE6C45EFC48FF30CF9380B1" box="[952,1020,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1986,2010]" captionTargetBox="[151,1436,1353,1956]" captionTargetId="figure-459@5.[151,1436,1353,1963]" captionTargetPageId="5" captionText="FIGURE 3. Megaraphidia ootsa sp. nov., holotype RBCM P1556, specimen as preserved. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655635" httpUri="https://zenodo.org/record/4655635/files/figure.png" pageId="16" pageNumber="57">Fig. 3</figureCitation>
|
||
). The only venational feature clearly distinguishing
|
||
<taxonomicName id="4C774D1BFFE6C45EFECEFF1CCE6D80D5" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[318,514,187,213]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE6C45EFECEFF1CCD2080D5" box="[318,335,187,213]" italics="true" pageId="16" pageNumber="57">A</emphasis>
|
||
.
|
||
<emphasis id="B903EA8AFFE6C45EFEAEFF1CCE6D80D5" box="[350,514,187,213]" italics="true" pageId="16" pageNumber="57">protomaculata</emphasis>
|
||
</taxonomicName>
|
||
from extant species of
|
||
<taxonomicName id="4C774D1BFFE6C45EFCFCFF1CCF2280D5" authorityName="Navas" authorityYear="1916" box="[780,845,187,213]" class="Insecta" family="Raphidiidae" genus="Alena" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFE6C45EFCFCFF1CCF2280D5" box="[780,845,187,213]" italics="true" pageId="16" pageNumber="57">Alena</emphasis>
|
||
</taxonomicName>
|
||
is the branching of RP1. It is forked twice, with the proximal fork located very deeply, proximad the level of 3ra-rp, while RP1 is forked once in most species of
|
||
<taxonomicName id="4C774D1BFFE6C45EFAA4FF78C9FA80F9" authorityName="Navas" authorityYear="1916" box="[1364,1429,223,249]" class="Insecta" family="Raphidiidae" genus="Alena" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFE6C45EFAA4FF78C9FA80F9" box="[1364,1429,223,249]" italics="true" pageId="16" pageNumber="57">Alena</emphasis>
|
||
</taxonomicName>
|
||
, and in the few other species of
|
||
<taxonomicName id="4C774D1BFFE6C45EFDF6FEA4CE28811D" authorityName="Navas" authorityYear="1916" box="[518,583,259,285]" class="Insecta" family="Raphidiidae" genus="Alena" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFE6C45EFDF6FEA4CE28811D" box="[518,583,259,285]" italics="true" pageId="16" pageNumber="57">Alena</emphasis>
|
||
</taxonomicName>
|
||
it is forked twice (
|
||
<emphasis id="B903EA8AFFE6C45EFCD9FEA4CF2B811C" box="[809,836,259,284]" italics="true" pageId="16" pageNumber="57">i.e</emphasis>
|
||
.,
|
||
<taxonomicName id="4C774D1BFFE6C45EFCABFEA4CF9A811C" authorityName="Carpenter" authorityYear="1958" box="[859,1013,259,285]" class="Insecta" family="Raphidiidae" genus="Alena" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="americana">
|
||
<emphasis id="B903EA8AFFE6C45EFCABFEA4CF9A811C" box="[859,1013,259,285]" italics="true" pageId="16" pageNumber="57">A. americana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C774D1BFFE6C45EFBF5FEA4CCBD8141" authority="Aspock & Aspock, 1978" authorityName="Aspock & Aspock" authorityYear="1978" class="Insecta" family="Raphidiidae" genus="Alena" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="tenochtitlana">
|
||
<emphasis id="B903EA8AFFE6C45EFBF5FEA4C8D4811D" box="[1029,1211,259,285]" italics="true" pageId="16" pageNumber="57">A. tenochtitlana</emphasis>
|
||
<bibRefCitation id="EFE64B69FFE6C45EFB33FEA4CCBD8141" author="Aspock, U. & Aspock, H." pageId="16" pageNumber="57" pagination="21 - 24" refId="ref21059" refString="Aspock, U. & Aspock, H. (1978) Raphidia (Aliaberaphidia) tenochtitlana n. sp. - eine neue Raphidiiden-Spezies aus Baja California (Mexiko) (Neuropteroidea, Raphidioptera). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 30, 21 - 24." type="journal article" year="1978">Aspöck & Aspöck, 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
), the proximal fork is relatively shallow (distad the level of 3ra-rp). However, the shape of the male terminal segments of
|
||
<taxonomicName id="4C774D1BFFE6C45EFED3FEECCE028165" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[291,621,331,357]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE6C45EFED3FEECCE028165" box="[291,621,331,357]" italics="true" pageId="16" pageNumber="57">Archiinocellia protomaculata</emphasis>
|
||
</taxonomicName>
|
||
fundamentally differs from those of all species of
|
||
<taxonomicName id="4C774D1BFFE6C45EFB6EFEECC8B08165" authorityName="Navas" authorityYear="1916" box="[1182,1247,331,357]" class="Insecta" family="Raphidiidae" genus="Alena" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFE6C45EFB6EFEECC8B08165" box="[1182,1247,331,357]" italics="true" pageId="16" pageNumber="57">Alena</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C774D1BFFE6C45EFAFAFEECCCBE8189" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE6C45EFAFAFEECCCBE8189" italics="true" pageId="16" pageNumber="57">A. protomaculata</emphasis>
|
||
</taxonomicName>
|
||
, the ninth tergite extends laterally to the ventral margin, and is probably fused with the ninth sternite to form a ring, a plesiomorphic condition at the family level, characteristic of all
|
||
<taxonomicName id="4C774D1BFFE6C45EFC3BFE34C83D81AD" authorityName="Latreille" authorityYear="1810" box="[971,1106,403,429]" class="Insecta" family="Raphidiidae" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="family">Raphidiidae</taxonomicName>
|
||
except
|
||
<taxonomicName id="4C774D1BFFE6C45EFB5AFE34C88481AD" authorityName="Navas" authorityYear="1916" box="[1194,1259,403,429]" class="Insecta" family="Raphidiidae" genus="Alena" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFE6C45EFB5AFE34C88481AD" box="[1194,1259,403,429]" italics="true" pageId="16" pageNumber="57">Alena</emphasis>
|
||
</taxonomicName>
|
||
, in which these are widely separated (Haring
|
||
<emphasis id="B903EA8AFFE6C45EFE15FE10CE7881D1" box="[485,535,439,465]" italics="true" pageId="16" pageNumber="57">et al</emphasis>
|
||
., 2010). The apparent absence of sclerotized gonocoxites
|
||
<quantity id="4C8F9B7DFFE6C45EFB56FE10C8BE81D1" box="[1190,1233,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="16" pageNumber="57" unit="in" value="9.0">9 in</quantity>
|
||
<taxonomicName id="4C774D1BFFE6C45EFB29FE10C9F381D1" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[1241,1436,439,465]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE6C45EFB29FE10C9F381D1" box="[1241,1436,439,465]" italics="true" pageId="16" pageNumber="57">A. protomaculata</emphasis>
|
||
</taxonomicName>
|
||
is also noteworthy and probably plesiomorphic at the family level. But we do not know if these terminal characters are also shared by other two species of the genus.
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFE6C45EFF37FD84CE1E8286" blockId="16.[151,1437,150,1077]" pageId="16" pageNumber="57">
|
||
In all extant genera, these are the largest sclerites of the terminalia and are strongly sclerotized (
|
||
<bibRefCitation id="EFE64B69FFE6C45EFAD5FD84CD558261" author="Aspock, U. & Aspock, H." pageId="16" pageNumber="57" pagination="97 - 127" refId="ref21205" refString="Aspock, U. & Aspock, H. (2008) Phylogenetic relevance of the genital sclerites of Neuropterida (Insecta: Holometabola). Systematic Entomology, 33, 97 - 127. https: // doi. org / 10.1111 / j. 1365 - 3113.2007.00396. x" type="journal article" year="2008">Aspöck & Aspöck, 2008</bibRefCitation>
|
||
). Gonocoxites 9 are also well developed in the Baltic amber
|
||
<taxonomicName id="4C774D1BFFE6C45EFC0BFDE0CC828285" authority="Aspock & Aspock" authorityName="Aspock & Aspock" class="Insecta" family="Raphidiidae" genus="Succinoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="exhibens">
|
||
<emphasis id="B903EA8AFFE6C45EFC0BFDE0C94F8261" box="[1019,1312,583,609]" italics="true" pageId="16" pageNumber="57">Succinoraphidia exhibens</emphasis>
|
||
Aspöck & Aspöck
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFE64B69FFE6C45EFF0BFDCCCE7B8285" author="Aspock, U. & Aspock, H." box="[251,532,619,645]" pageId="16" pageNumber="57" pagination="11 - 19" refId="ref21145" refString="Aspock, U. & Aspock, H. (2004) Two significant new snakeflies from Baltic amber, with discussion on autapomorphies of the order and its included taxa (Raphidioptera). Systematic Entomology, 29, 11 - 19. https: // doi. org / 10.1111 / j. 1365 - 3113.2004.00245. x" type="journal article" year="2004">Aspöck & Aspöck, 2004</bibRefCitation>
|
||
:
|
||
<figureCitation id="134C2A1DFFE6C45EFDEFFDCCCE0C8286" box="[543,611,619,646]" captionStart="FIGURE 6" captionStartId="10.[151,250,1930,1954]" captionTargetBox="[164,1426,181,1893]" captionTargetId="figure-26@10.[151,1436,181,1906]" captionTargetPageId="10" captionText="FIGURE 6. Megaraphidia klondika sp. nov., specimen SRUI99-71-63. A, part; B, counterpart; C, drawing of the hind wing venation. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655647" httpUri="https://zenodo.org/record/4655647/files/figure.png" pageId="16" pageNumber="57">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFE6C45EFF37FD28CE2783ED" blockId="16.[151,1437,150,1077]" pageId="16" pageNumber="57">
|
||
<bibRefCitation id="EFE64B69FFE6C45EFF37FD28CE4B82A9" author="Makarkin, V. N. & Archibald, S. B." box="[199,548,655,681]" pageId="16" pageNumber="57" pagination="401 - 444" refId="ref23770" refString="Makarkin, V. N. & Archibald, S. B. (2014) A revision of the late Eocene snakeflies (Raphidioptera) of the Florissant Formation, Colorado, with special reference to the wing venation of the Raphidiomorpha. Zootaxa, 3784 (4), 401 - 444. https: // doi. org / 10.11646 / zootaxa. 3784.4.4" type="journal article" year="2014">Makarkin & Archibald (2014)</bibRefCitation>
|
||
referred
|
||
<emphasis id="B903EA8AFFE6C45EFD62FD28CFA582A9" box="[658,970,655,681]" italics="true" pageId="16" pageNumber="57">
|
||
<taxonomicName id="4C774D1BFFE6C45EFD62FD28CF5B82A9" box="[658,820,655,681]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">Archiraphidia</taxonomicName>
|
||
? somnolenta
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE64B69FFE6C45EFC2CFD28C8EB82A9" author="Scudder, S. H." box="[988,1156,655,681]" pageId="16" pageNumber="57" pagination="1 - 734" refId="ref25198" refString="Scudder, S. H. (1890) The Tertiary Insects of North America. Report of the United States Geological Survey of the Territories, 13, 1 - 734. https: // doi. org / 10.5962 / bhl. title. 44698" type="journal article" year="1890">Scudder, 1890</bibRefCitation>
|
||
) from Florissant to that genus only provisionally. It is similar to
|
||
<taxonomicName id="4C774D1BFFE6C45EFD97FD14CF6982CD" authorityName="Handlirsch" authorityYear="1910" box="[615,774,691,717]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFE6C45EFD97FD14CF6982CD" box="[615,774,691,717]" italics="true" pageId="16" pageNumber="57">Archiinocellia</emphasis>
|
||
</taxonomicName>
|
||
in some ways, sharing with
|
||
<taxonomicName id="4C774D1BFFE6C45EFBBFFD14C91D82CD" authorityName="Handlirsch" authorityYear="1910" box="[1103,1394,691,717]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="oligoneura">
|
||
<emphasis id="B903EA8AFFE6C45EFBBFFD14C91D82CD" box="[1103,1394,691,717]" italics="true" pageId="16" pageNumber="57">Archiinocellia oligoneura</emphasis>
|
||
</taxonomicName>
|
||
the following character states: (1)
|
||
<emphasis id="B903EA8AFFE6C45EFE01FD70CF4A82F1" box="[497,805,727,753]" italics="true" pageId="16" pageNumber="57">
|
||
<taxonomicName id="4C774D1BFFE6C45EFE01FD70CEFC82F1" box="[497,659,727,753]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">Archiraphidia</taxonomicName>
|
||
? somnolenta
|
||
</emphasis>
|
||
has a long pterostigma, 4.5 to 5 times longer than wide (
|
||
<taxonomicName id="4C774D1BFFE6C45EFF51FD5CCD548315" authorityName="Handlirsch" authorityYear="1910" box="[161,315,763,789]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="oligoneura">
|
||
<emphasis id="B903EA8AFFE6C45EFF51FD5CCD548315" box="[161,315,763,789]" italics="true" pageId="16" pageNumber="57">A. oligoneura</emphasis>
|
||
</taxonomicName>
|
||
, 4.6–4.7), while those of
|
||
<taxonomicName id="4C774D1BFFE6C45EFDABFD5CCE968315" box="[603,761,763,789]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFE6C45EFDABFD5CCE968315" box="[603,761,763,789]" italics="true" pageId="16" pageNumber="57">Archiraphidia</emphasis>
|
||
</taxonomicName>
|
||
species are much shorter, 2.3 to 3.2 times longer than wide, and (2) both have two
|
||
<emphasis id="B903EA8AFFE6C45EFE66FCB8CDD58339" box="[406,442,799,825]" italics="true" pageId="16" pageNumber="57">doi</emphasis>
|
||
in the hind wing (one in species of
|
||
<taxonomicName id="4C774D1BFFE6C45EFCBDFCB8CF848339" box="[845,1003,799,825]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFE6C45EFCBDFCB8CF848339" box="[845,1003,799,825]" italics="true" pageId="16" pageNumber="57">Archiraphidia</emphasis>
|
||
</taxonomicName>
|
||
). However, the ScP of
|
||
<emphasis id="B903EA8AFFE6C45EFB01FCB8CD77835D" italics="true" pageId="16" pageNumber="57">
|
||
<taxonomicName id="4C774D1BFFE6C45EFB01FCB8C9FC8339" box="[1265,1427,799,825]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">Archiraphidia</taxonomicName>
|
||
? somnolenta
|
||
</emphasis>
|
||
is slightly longer than in
|
||
<taxonomicName id="4C774D1BFFE6C45EFDB8FCE4CF03835D" authorityName="Handlirsch" authorityYear="1910" box="[584,876,835,861]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="oligoneura">
|
||
<emphasis id="B903EA8AFFE6C45EFDB8FCE4CF03835D" box="[584,876,835,861]" italics="true" pageId="16" pageNumber="57">Archiinocellia oligoneura</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C774D1BFFE6C45EFC58FCE4C801835D" authorityName="Archibald & Makarkin" authorityYear="2021" baseAuthorityName="Engel" baseAuthorityYear="2011" box="[936,1134,835,861]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="species" species="protomaculata">
|
||
<emphasis id="B903EA8AFFE6C45EFC58FCE4C801835D" box="[936,1134,835,861]" italics="true" pageId="16" pageNumber="57">A. protomaculata</emphasis>
|
||
</taxonomicName>
|
||
, somewhat differing from character state (1) of the
|
||
<taxonomicName id="4C774D1BFFE6C45EFE57FCC0CE298381" authorityName="Handlirsch" authorityYear="1910" box="[423,582,871,897]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFE6C45EFE57FCC0CE298381" box="[423,582,871,897]" italics="true" pageId="16" pageNumber="57">Archiinocellia</emphasis>
|
||
</taxonomicName>
|
||
diagnosis, and the proximal-most branching of RP1 is clearly distad the level of 3ra-rp, contradicting diagnostic character state (4). The taxonomic placement of
|
||
<emphasis id="B903EA8AFFE6C45EFBC7FC2CC90583A5" box="[1079,1386,907,933]" italics="true" pageId="16" pageNumber="57">
|
||
<taxonomicName id="4C774D1BFFE6C45EFBC7FC2CC8B683A5" box="[1079,1241,907,933]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">Archiraphidia</taxonomicName>
|
||
? somnolenta
|
||
</emphasis>
|
||
will be clarified by the discovery of more complete specimens; the species does not belong to
|
||
<taxonomicName id="4C774D1BFFE6C45EFB79FC08C94783C9" authorityName="Handlirsch" authorityYear="1910" box="[1161,1320,943,969]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFE6C45EFB79FC08C94783C9" box="[1161,1320,943,969]" italics="true" pageId="16" pageNumber="57">Archiinocellia</emphasis>
|
||
</taxonomicName>
|
||
and probably does not belong to
|
||
<taxonomicName id="4C774D1BFFE6C45EFE53FC74CE2E83ED" box="[419,577,979,1005]" class="Insecta" family="Raphidiidae" genus="Archiraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFE6C45EFE53FC74CE2E83ED" box="[419,577,979,1005]" italics="true" pageId="16" pageNumber="57">Archiraphidia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC83698FFE6C45EFF37FC50C9FB8435" blockId="16.[151,1437,150,1077]" pageId="16" pageNumber="57">
|
||
Of fossil genera, the venation of
|
||
<taxonomicName id="4C774D1BFFE6C45EFDDFFC50CEA68411" authorityName="Handlirsch" authorityYear="1910" box="[559,713,1015,1041]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFE6C45EFDDFFC50CEA68411" box="[559,713,1015,1041]" italics="true" pageId="16" pageNumber="57">Archiinocellia</emphasis>
|
||
</taxonomicName>
|
||
is most similar to that of
|
||
<emphasis id="B903EA8AFFE6C45EFC14FC50C8EB8411" box="[996,1156,1015,1041]" italics="true" pageId="16" pageNumber="57">Archiiraphidia</emphasis>
|
||
and
|
||
<emphasis id="B903EA8AFFE6C45EFB4BFC50C9F98411" box="[1211,1430,1015,1041]" italics="true" pageId="16" pageNumber="57">Florissontoraphidia</emphasis>
|
||
, which share character states A–C. However,
|
||
<taxonomicName id="4C774D1BFFE6C45EFD8DFBBCCF788435" authorityName="Handlirsch" authorityYear="1910" box="[637,791,1051,1077]" class="Insecta" family="Raphidiidae" genus="Archiinocellia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="16" pageNumber="57" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B903EA8AFFE6C45EFD8DFBBCCF788435" box="[637,791,1051,1077]" italics="true" pageId="16" pageNumber="57">Archiinocellia</emphasis>
|
||
</taxonomicName>
|
||
clearly differs from these genera as stated in its diagnosis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |