233 lines
29 KiB
XML
233 lines
29 KiB
XML
<document id="7FF3284B6C887F3C41BE563A70BF8CBD" ID-CLB-Dataset="7413" ID-DOI="10.11646/zootaxa.4951.1.2" ID-GBIF-Dataset="d7a40f10-4145-499c-97be-3f9bc5088d2f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4655625" ID-ZooBank="557825A0-714A-426A-917F-1C9AB7372C30" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617265349256" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Makarkin, Vladimir N." docDate="2021" docId="03DE878EFFF5C44BFF67FF30CF0F83A5" docLanguage="en" docName="zootaxa.4951.1.2.pdf" docOrigin="Zootaxa 4951 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaraphidia antiquissima Archibald & Makarkin 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="46" masterDocId="FFE7FFF6FFF6C44EFFF0FFA7CC6F8000" masterDocTitle="Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation" masterLastPageNumber="79" masterPageNumber="41" pageNumber="44" updateTime="1698935824968" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="A1113B11B929B26E82CF6985AB3D433F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="102D185941D2A975E259A947AA64E9CB">
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<mods:title id="35F0F55A7C7EDCBA014D95CD53CFC548">Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation</mods:title>
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</mods:titleInfo>
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<mods:name id="789370336A732394FA7F6ABE83218CF7" type="personal">
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<mods:roleTerm id="4740C1050425A628A5D7280E049A438B">Author</mods:roleTerm>
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<mods:namePart id="39B5F9DA91CAA1A0D318ED653D053BC9">Archibald, S. Bruce</mods:namePart>
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<mods:nameIdentifier id="4CB6F6FF352ABEF643502E35CF5CD750" type="ORCID">0000-0002-4397-2497</mods:nameIdentifier>
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<mods:affiliation id="BFC203B473D15AD5A267FBB0C0F53795">Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada. https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:name id="2FB497742E538CC58E074FCF847DF62A" type="personal">
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<mods:roleTerm id="E3460722FC5739E407B0918C1FD980BE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A41FCBF5DB108F2C7C304A93B1636345">Makarkin, Vladimir N.</mods:namePart>
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<mods:affiliation id="F3FC2994A5886E79F9611E0E8FD70918">Federal Scientific Centre of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivo-</mods:affiliation>
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<mods:typeOfResource id="EC194094CAB7247C10D6F3F51C18AC0C">text</mods:typeOfResource>
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<mods:title id="F8A083777313000CD84E72524A00A26E">Zootaxa</mods:title>
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<mods:part id="B2EA029F0B72E045D2F8CF9B6DA48867">
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<mods:date id="655D8AB13FC4E80CA2761B12A608D6A9">2021</mods:date>
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<mods:number id="6DC62313D9FB287D8D513C002915AC98">2021-04-01</mods:number>
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<mods:number id="0ED4D73D02BB8725C32CCFC45C869FAF">4951</mods:number>
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<mods:detail id="C3EF7BF64AAA770A9208D225390DE0AD" type="issue">
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<mods:number id="6DBFF507419D408E2A314140B0363D0D">1</mods:number>
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<mods:start id="D3AD78E2C4C77CAD0E67EF212D51EE3D">41</mods:start>
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<mods:classification id="288CDA5E754183578A24FEDDCEC03BFE">journal article</mods:classification>
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<mods:identifier id="0BC92D7DBBDA8D638FBC125FD1A7B651" type="CLB-Dataset">7413</mods:identifier>
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<mods:identifier id="22284AD9FC92296D31F5200C0C299213" type="DOI">10.11646/zootaxa.4951.1.2</mods:identifier>
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<treatment id="03DE878EFFF5C44BFF67FF30CF0F83A5" ID-DOI="http://doi.org/10.5281/zenodo.4655633" ID-GBIF-Taxon="180999345" ID-Zenodo-Dep="4655633" LSID="urn:lsid:plazi:treatment:03DE878EFFF5C44BFF67FF30CF0F83A5" httpUri="http://treatment.plazi.org/id/03DE878EFFF5C44BFF67FF30CF0F83A5" lastPageId="5" lastPageNumber="46" pageId="3" pageNumber="44">
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<subSubSection id="C36D6513FFF5C44DFF67FF30CE2680B1" box="[151,585,151,178]" pageId="3" pageNumber="44" type="nomenclature">
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<paragraph id="8BC83698FFF5C44DFF67FF30CE2680B1" blockId="3.[151,585,151,214]" box="[151,585,151,178]" pageId="3" pageNumber="44">
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<heading id="D08081F4FFF5C44DFF67FF30CE2680B1" bold="true" box="[151,585,151,178]" fontSize="11" level="1" pageId="3" pageNumber="44" reason="1">
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<emphasis id="B903EA8AFFF5C44DFF67FF30CE2680B1" bold="true" box="[151,585,151,178]" pageId="3" pageNumber="44">
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<taxonomicName id="4C774D1BFFF5C44DFF67FF30CD8A80B1" authority="Archibald & Makarkin, 2021" authorityName="Archibald & Makarkin" authorityYear="2021" box="[151,485,151,178]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="antiquissima" status="sp. nov.">
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<emphasis id="B903EA8AFFF5C44DFF67FF30CD8A80B1" bold="true" box="[151,485,151,178]" italics="true" pageId="3" pageNumber="44">Megaraphidia antiquissima</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23057F1FFF5C44DFE1CFF30CE2680B1" box="[492,585,151,177]" pageId="3" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36D6513FFF5C44DFF67FF1CCD3780D6" box="[151,344,187,214]" pageId="3" pageNumber="44" type="description">
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||
<paragraph id="8BC83698FFF5C44DFF67FF1CCD3780D6" blockId="3.[151,585,151,214]" box="[151,344,187,214]" pageId="3" pageNumber="44">
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||
<figureCitation id="134C2A1DFFF5C44DFF67FF1CCCB380D5" box="[151,220,187,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1381,1405]" captionTargetBox="[152,1436,777,1353]" captionTargetId="figure-221@3.[152,1436,777,1357]" captionTargetPageId="3" captionText="FIGURE 1. Megaraphidia antiquissima sp. nov., holotype RBCM P1555, specimen as preserved. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655627" httpUri="https://zenodo.org/record/4655627/files/figure.png" pageId="3" pageNumber="44">Figs 1</figureCitation>
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,
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<figureCitation id="134C2A1DFFF5C44DFF1AFF1CCC9580D5" box="[234,250,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[206,1380,192,1913]" captionTargetId="figure-26@4.[195,1393,181,1919]" captionTargetPageId="4" captionText="FIGURE 2. Wing venation of Megaraphidia antiquissima sp. nov., holotype RBCM P1555. A, right forewing, B, left forewing; C, right forewing; D, left hind wing. Scale bar = 3 mm (all to scale)." figureDoi="http://doi.org/10.5281/zenodo.4655631" httpUri="https://zenodo.org/record/4655631/files/figure.png" pageId="3" pageNumber="44">2</figureCitation>
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,
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<figureCitation id="134C2A1DFFF5C44DFEF6FF1CCD3780D6" box="[262,344,187,214]" captionStart="FIGURE 10" captionStartId="14.[151,250,1402,1426]" captionTargetBox="[151,1436,597,1379]" captionTargetId="figure-227@14.[151,1436,597,1379]" captionTargetPageId="14" captionText="FIGURE 10. Pterostigmata of Megaraphidia. A, M. antiquissima sp. nov., right wings of the holotype RBCM P1555; B, M. antiquissima sp. nov., left wings of same specimen; C, M. ootsa sp. nov., right hind wing of the holotype RBCM P1556; D, M. ootsa sp. nov., left wings of same specimen; E, M. klondika sp. nov., holotype, an overlapping forewing and hind wing SR03-01-01; F, M. klondika sp. nov., specimen SR-16-004-001-1. RAf, RA of forewing; Rah, RA of hind wing. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4655665" httpUri="https://zenodo.org/record/4655665/files/figure.png" pageId="3" pageNumber="44">10A, B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6513FFF5C44DFF67FEA4C9F68166" pageId="3" pageNumber="44" type="materials_examined">
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<paragraph id="8BC83698FFF5C44DFF67FEA4CFF38142" blockId="3.[151,1437,259,753]" pageId="3" pageNumber="44">
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<emphasis id="B903EA8AFFF5C44DFF67FEA4CD29811D" bold="true" box="[151,326,259,285]" pageId="3" pageNumber="44">Type material.</emphasis>
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<materialsCitation id="3B1F3CC5FFF5C44DFEBEFEA4CFF38142" ID-GBIF-Occurrence="3071122312" collectionCode="RBCM" collectorName="John Leahy" pageId="3" pageNumber="44" specimenCode="P1555" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54CC883AFFF5C44DFEBEFEA4CDD9811D" box="[334,438,259,285]" pageId="3" pageNumber="44" type="holotype">Holotype</typeStatus>
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<collectionCode id="ED66AE5DFFF5C44DFE4EFEA4CE60811D" box="[446,527,259,285]" country="Canada" httpUri="http://grbio.org/cool/gayt-20cg" name="Royal British Columbia Museum" pageId="3" pageNumber="44" type="Museum">RBCM</collectionCode>
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<specimenCode id="DBD19EE3FFF5C44DFDE7FEA4CE0E811E" box="[535,609,259,286]" collectionCode="RBCM" country="Canada" httpUri="http://grbio.org/cool/gayt-20cg" name="Royal British Columbia Museum" pageId="3" pageNumber="44" type="Museum">P1555</specimenCode>
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, collected by
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<collectorName id="2682534EFFF5C44DFD0EFEA4CFED811D" box="[766,898,259,285]" pageId="3" pageNumber="44">John Leahy</collectorName>
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, unknown date, deposited in the
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<collectionCode id="ED66AE5DFFF5C44DFB07FEA4C927811D" box="[1271,1352,259,285]" country="Canada" httpUri="http://grbio.org/cool/gayt-20cg" name="Royal British Columbia Museum" pageId="3" pageNumber="44" type="Museum">RBCM</collectionCode>
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collections. A nearly complete female specimen preserved in dorsal aspect.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BC83698FFF5C44DFF37FEECC9F68166" blockId="3.[151,1437,259,753]" box="[199,1433,331,358]" pageId="3" pageNumber="44">
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<materialsCitation id="3B1F3CC5FFF5C44DFF37FEECC9F98166" ID-GBIF-Occurrence="3071122308" box="[199,1430,331,358]" country="Canada" county="Eocene" location="Hoodoo Face" municipality="McAbee" pageId="3" pageNumber="44" specimenCount="1" stateProvince="British Columbia" typeStatus="holotype">
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<emphasis id="B903EA8AFFF5C44DFF37FEECCD918165" bold="true" box="[199,510,331,357]" pageId="3" pageNumber="44">
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<typeStatus id="54CC883AFFF5C44DFF37FEECCD6D8165" box="[199,258,331,357]" pageId="3" pageNumber="44">Type</typeStatus>
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locality and horizon.
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</emphasis>
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||
<location id="8EA86043FFF5C44DFDF5FEECCEF48166" LSID="urn:lsid:plazi:treatment:03DE878EFFF5C44BFF67FF30CF0F83A5:8EA86043FFF5C44DFDF5FEECCEF48166" box="[517,667,331,358]" country="Canada" county="Eocene" municipality="McAbee" name="Hoodoo Face" pageId="3" pageNumber="44" stateProvince="British Columbia">Hoodoo Face</location>
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beds,
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<collectingMunicipality id="6BACACE2FFF5C44DFD13FEECCF298165" box="[739,838,331,357]" pageId="3" pageNumber="44">McAbee</collectingMunicipality>
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,
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<collectingRegion id="49B3F87AFFF5C44DFCA2FEECC8798165" box="[850,1046,331,357]" country="Canada" name="British Columbia" pageId="3" pageNumber="44">British Columbia</collectingRegion>
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,
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<collectingCountry id="F3607608FFF5C44DFBD1FEECC8168165" box="[1057,1145,331,357]" name="Canada" pageId="3" pageNumber="44">Canada</collectingCountry>
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; early
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<collectingCounty id="62A94E14FFF5C44DFB33FEEBC97A8166" box="[1219,1301,332,358]" pageId="3" pageNumber="44">Eocene</collectingCounty>
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(Ypresian)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6513FFF5C44DFF37FEC8CC8F81AE" pageId="3" pageNumber="44" type="etymology">
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<paragraph id="8BC83698FFF5C44DFF37FEC8CC8F81AE" blockId="3.[151,1437,259,753]" pageId="3" pageNumber="44">
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<emphasis id="B903EA8AFFF5C44DFF37FEC8CD218189" bold="true" box="[199,334,367,393]" pageId="3" pageNumber="44">Etymology.</emphasis>
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From the Latin
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<emphasis id="B903EA8AFFF5C44DFDFFFED7CEC68189" box="[527,681,368,393]" italics="true" pageId="3" pageNumber="44">antiquissimus</emphasis>
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[
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<emphasis id="B903EA8AFFF5C44DFD4CFED7CF608189" box="[700,783,368,393]" italics="true" pageId="3" pageNumber="44">-a, -um</emphasis>
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], oldest, most ancient, as this is the oldest record of the genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36D6513FFF5C44DFF37FE10CFF18261" pageId="3" pageNumber="44" type="diagnosis">
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<paragraph id="8BC83698FFF5C44DFF37FE10CFF18261" blockId="3.[151,1437,259,753]" pageId="3" pageNumber="44">
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<emphasis id="B903EA8AFFF5C44DFF37FE10CD2D81D1" bold="true" box="[199,322,439,465]" pageId="3" pageNumber="44">Diagnosis.</emphasis>
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May be distinguished from other species of
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<taxonomicName id="4C774D1BFFF5C44DFCA4FE10CF9B81D1" authorityName="Cockerell" authorityYear="1907" box="[852,1012,439,465]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
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<emphasis id="B903EA8AFFF5C44DFCA4FE10CF9B81D1" box="[852,1012,439,465]" italics="true" pageId="3" pageNumber="44">Megaraphidia</emphasis>
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</taxonomicName>
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by head broad, not cuneate [clearly cuneate,
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<emphasis id="B903EA8AFFF5C44DFF0DFE7BCD7781F5" box="[253,280,476,501]" italics="true" pageId="3" pageNumber="44">i.e</emphasis>
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, narrowed posteriorly in
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<taxonomicName id="4C774D1BFFF5C44DFDCCFE7BCECF81F5" authorityName="Archibald & Makarkin" authorityYear="2021" box="[572,672,476,501]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="ootsa">
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<emphasis id="B903EA8AFFF5C44DFDCCFE7BCECF81F5" box="[572,672,476,501]" italics="true" pageId="3" pageNumber="44">M. ootsa</emphasis>
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</taxonomicName>
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]; ovipositor relatively short (0.28 body length) [
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<quantity id="4C8F9B7DFFF5C44DFB3CFE7CC97081F5" box="[1228,1311,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="8.636" pageId="3" pageNumber="44" unit="in" value="0.34">0.34 in</quantity>
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<taxonomicName id="4C774D1BFFF5C44DFAD8FE7BC9E381F5" authorityName="Archibald & Makarkin" authorityYear="2021" box="[1320,1420,476,501]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="ootsa">
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<emphasis id="B903EA8AFFF5C44DFAD8FE7BC9E381F5" box="[1320,1420,476,501]" italics="true" pageId="3" pageNumber="44">M. ootsa</emphasis>
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</taxonomicName>
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], relatively narrow costal space in forewing (2.3 times wider than subcostal space) [broader in
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<taxonomicName id="4C774D1BFFF5C44DFB53FDA7C95B8219" baseAuthorityName="Cockerell" baseAuthorityYear="1909" box="[1187,1332,511,537]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="exhumata">
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<emphasis id="B903EA8AFFF5C44DFB53FDA7C95B8219" box="[1187,1332,511,537]" italics="true" pageId="3" pageNumber="44">M. exhumata</emphasis>
|
||
</taxonomicName>
|
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(2.6),
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<taxonomicName id="4C774D1BFFF5C44DFA8EFDA7CC98823D" authorityName="Archibald & Makarkin" authorityYear="2021" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="klondika">
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<emphasis id="B903EA8AFFF5C44DFA8EFDA7CC98823D" italics="true" pageId="3" pageNumber="44">M. klondika</emphasis>
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</taxonomicName>
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||
(2.9),
|
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<taxonomicName id="4C774D1BFFF5C44DFEB2FD83CDA7823D" authorityName="Archibald & Makarkin" authorityYear="2021" box="[322,456,547,573]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="klondika">
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<emphasis id="B903EA8AFFF5C44DFEB2FD83CDA7823D" box="[322,456,547,573]" italics="true" pageId="3" pageNumber="44">M. klondika</emphasis>
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</taxonomicName>
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(2.9),
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<taxonomicName id="4C774D1BFFF5C44DFDE3FD83CEE1823D" authorityName="Cockerell" authorityYear="1907" box="[531,654,547,573]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="elegans">
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<emphasis id="B903EA8AFFF5C44DFDE3FD83CEE1823D" box="[531,654,547,573]" italics="true" pageId="3" pageNumber="44">M. elegans</emphasis>
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</taxonomicName>
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(3.0); narrower in
|
||
<taxonomicName id="4C774D1BFFF5C44DFC92FD83CF88823D" authorityName="Archibald & Makarkin" authorityYear="2021" box="[866,999,547,573]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="hopkinsi">
|
||
<emphasis id="B903EA8AFFF5C44DFC92FD83CF88823D" box="[866,999,547,573]" italics="true" pageId="3" pageNumber="44">M. hopkinsi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C774D1BFFF5C44DFC05FD83C838823D" authorityName="Archibald & Makarkin" authorityYear="2021" box="[1013,1111,548,573]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="ootsa">
|
||
<emphasis id="B903EA8AFFF5C44DFC05FD83C838823D" box="[1013,1111,548,573]" italics="true" pageId="3" pageNumber="44">M. ootsa</emphasis>
|
||
</taxonomicName>
|
||
(2.0)]; basal
|
||
<emphasis id="B903EA8AFFF5C44DFB1CFD84C97F823D" box="[1260,1296,547,573]" italics="true" pageId="3" pageNumber="44">doi</emphasis>
|
||
shorter than distal in hind wing [both approximately equal in length
|
||
<taxonomicName id="4C774D1BFFF5C44DFCF8FDEFCFE28261" authorityName="Archibald & Makarkin" authorityYear="2021" box="[776,909,583,609]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="klondika">
|
||
<emphasis id="B903EA8AFFF5C44DFCF8FDEFCFE28261" box="[776,909,583,609]" italics="true" pageId="3" pageNumber="44">M. klondika</emphasis>
|
||
</taxonomicName>
|
||
].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36D6513FFF5C44BFF37FDCCCF0F83A5" lastPageId="5" lastPageNumber="46" pageId="3" pageNumber="44" type="description">
|
||
<paragraph id="8BC83698FFF5C44DFF37FDCCCDA182F1" blockId="3.[151,1437,259,753]" pageId="3" pageNumber="44">
|
||
<emphasis id="B903EA8AFFF5C44DFF37FDCCCD368285" bold="true" box="[199,345,619,645]" pageId="3" pageNumber="44">Description.</emphasis>
|
||
Female. Body length (without ovipositor) approximately
|
||
<quantity id="4C8F9B7DFFF5C44DFC0CFDCCC83D8286" box="[1020,1106,619,646]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="44" unit="mm" value="15.0">15 mm</quantity>
|
||
. Head broad, apparently not cuneate (incompletely preserved). Prothorax probably relatively long (poorly preserved), approximately 3.0–
|
||
<quantity id="4C8F9B7DFFF5C44DFA8AFD28CCAC82CE" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="44" unit="mm" value="3.5">3.5 mm</quantity>
|
||
long (0.20–0.23 body length). Ovipositor relatively short, approximately
|
||
<quantity id="4C8F9B7DFFF5C44DFC05FD14C82582CE" box="[1013,1098,691,718]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="44" unit="mm" value="4.2">4.2 mm</quantity>
|
||
long. Details of body, legs not discernible by preservation.
|
||
</paragraph>
|
||
<caption id="DF086610FFF5C44DFF67FAC2C952857D" ID-DOI="http://doi.org/10.5281/zenodo.4655627" ID-Zenodo-Dep="4655627" box="[151,1341,1381,1405]" httpUri="https://zenodo.org/record/4655627/files/figure.png" pageId="3" pageNumber="44" startId="3.[151,250,1381,1405]" targetBox="[152,1436,777,1353]" targetPageId="3">
|
||
<paragraph id="8BC83698FFF5C44DFF67FAC2C952857D" blockId="3.[151,1341,1381,1405]" box="[151,1341,1381,1405]" pageId="3" pageNumber="44">
|
||
<emphasis id="B903EA8AFFF5C44DFF67FAC2CD62857D" bold="true" box="[151,269,1381,1405]" pageId="3" pageNumber="44">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="4C774D1BFFF5C44DFEE9FAC2CE5C857D" authority="Archibald & Makarkin, 2021" authorityName="Archibald & Makarkin" authorityYear="2021" box="[281,563,1381,1405]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="antiquissima" status="sp. nov.">
|
||
<emphasis id="B903EA8AFFF5C44DFEE9FAC2CE5C857D" box="[281,563,1381,1405]" italics="true" pageId="3" pageNumber="44">Megaraphidia antiquissima</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B903EA8AFFF5C44DFDC8FAC2CEE6857D" bold="true" box="[568,649,1381,1405]" pageId="3" pageNumber="44">
|
||
<taxonomicNameLabel id="A23057F1FFF5C44DFDC8FAC2CEE6857D" box="[568,649,1381,1405]" pageId="3" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype RBCM P1555, specimen as preserved. Scale bar = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC83698FFF5C44BFF37FA0CC95D811D" blockId="3.[151,1437,1451,2018]" lastBlockId="5.[151,1437,151,933]" lastPageId="5" lastPageNumber="46" pageId="3" pageNumber="44">
|
||
Forewing approximately
|
||
<quantity id="4C8F9B7DFFF5C44DFE16FA0BCE5885C6" box="[486,567,1452,1478]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="44" unit="mm" value="12.0">12 mm</quantity>
|
||
long, approximately
|
||
<quantity id="4C8F9B7DFFF5C44DFCDCFA0CCFEB85C6" box="[812,900,1451,1478]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="44" unit="mm" value="3.5">3.5 mm</quantity>
|
||
wide (length/width: 3.43). Costal space broad, widest at proximal 1/5 of length; seven simple subcostal veinlets (right forewing: preserved, probably nine to ten total), all relatively closely spaced. ScP relatively long, terminating on costal margin opposite crossvein 2ra-rp; length from termination of ScP to pterostigma approximately equal to length of pterostigma. Subcostal space moderately broad, with basal crossvein located in middle between origin of RP, divergence of M, CuA; distal crossvein 2scp-r (forming basal margin of pterostigma) straight, slightly inclined toward base (
|
||
<emphasis id="B903EA8AFFF5C44DFC0BF9C7C8798679" box="[1019,1046,1632,1657]" italics="true" pageId="3" pageNumber="44">i.e</emphasis>
|
||
., joins C slightly proximad joining RA). Pterostigma relatively short, rather strongly pigmented. Three RA branches: one incorporated in pterostigma medially well discernible; second forming distal margin of pterostigma; third distad pterostigma; none forked. RA entering margin about halfway between pterostigma, apex. RA space with two crossveins forming two radial cells: 2ra-rp long, located well proximad pterostigma, opposite termination of ScP; 3ra-rp long, joins distal part of pterostigma along RA. RP originates in proximal part of wing (approximately 0.39 wing length), with four (right wing) or six (left wing) pectinate branches: RP1 deeply forked; its anterior branch simple, posterior branch forked once (both wings). RP3 deeply forked (right wing), and RP4 shallowly forked (right wing); other branches simple. One long intraradial crossvein rp1-rp2 between stem of RP, RP1. Three crossveins between RP, MA: 1r-m, 2r-m connecting stem of RP, MA; 3r-m connecting RP1, anterior branch of MA. M fused with CuA for short distance; forked well proximad origin of RP. MA deeply dichotomously forked twice. MP zigzagged, with three simple branches. Two intramedian crossveins form two
|
||
<emphasis id="B903EA8AFFF3C44BFDB9FF30CE0280B1" box="[585,621,151,177]" italics="true" pageId="5" pageNumber="46">doi</emphasis>
|
||
, basal cell somewhat shorter than apical. Anterior trace of CuA strongly zigzagged, simple, fused with MP for short distance forming part of posterior margin of basal
|
||
<emphasis id="B903EA8AFFF3C44BFB36FF1CC88580D5" box="[1222,1258,187,213]" italics="true" pageId="5" pageNumber="46">doi</emphasis>
|
||
; posterior trace of CuA simple. CuP probably simple (distal portion not visible). Crossvein icu between CuA, CuP long. Crossvein cu-aa between CuP, AA1 closer to origin of CuA than to icu. Only basal AA1 discernible, AA2, AA3 not.
|
||
</paragraph>
|
||
<caption id="DF086610FFF2C44AFF67F830CF2487D3" ID-DOI="http://doi.org/10.5281/zenodo.4655631" ID-Zenodo-Dep="4655631" httpUri="https://zenodo.org/record/4655631/files/figure.png" pageId="4" pageNumber="45" startId="4.[151,250,1943,1967]" targetBox="[206,1380,192,1913]" targetPageId="4">
|
||
<paragraph id="8BC83698FFF2C44AFF67F830CF2487D3" blockId="4.[151,1437,1943,2003]" pageId="4" pageNumber="45">
|
||
<emphasis id="B903EA8AFFF2C44AFF67F830CD7D87AF" bold="true" box="[151,274,1943,1967]" pageId="4" pageNumber="45">FIGURE 2.</emphasis>
|
||
Wing venation of
|
||
<taxonomicName id="4C774D1BFFF2C44AFE3BF830CE8C87AF" authority="Archibald & Makarkin, 2021" authorityName="Archibald & Makarkin" authorityYear="2021" box="[459,739,1943,1967]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="antiquissima" status="sp. nov.">
|
||
<emphasis id="B903EA8AFFF2C44AFE3BF830CE8C87AF" box="[459,739,1943,1967]" italics="true" pageId="4" pageNumber="45">Megaraphidia antiquissima</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B903EA8AFFF2C44AFD18F830CF5787AF" bold="true" box="[744,824,1943,1967]" pageId="4" pageNumber="45">
|
||
<taxonomicNameLabel id="A23057F1FFF2C44AFD18F830CF5787AF" box="[744,824,1943,1967]" pageId="4" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype RBCM P1555. A, right forewing, B, left forewing; C, right forewing; D, left hind wing. Scale bar = 3 mm (all to scale).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC83698FFF3C44BFF37FE80CF0F83A5" blockId="5.[151,1437,151,933]" pageId="5" pageNumber="46">
|
||
Hind wing approximately
|
||
<quantity id="4C8F9B7DFFF3C44BFE01FE80CE508141" box="[497,575,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="46" unit="mm" value="11.0">11 mm</quantity>
|
||
long, approximately
|
||
<quantity id="4C8F9B7DFFF3C44BFCC1FE81CFF98141" box="[817,918,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" pageId="5" pageNumber="46" unit="mm" value="3.05">3.05 mm</quantity>
|
||
wide (length/width: 3.61). Costal space moderately broad; four to five simple subcostal veinlets preserved, rather closely spaced. ScP relatively long; length from termination of ScP to pterostigma approximately equal to its length. In subcostal space, one crossvein detected (2scp-r, forming basal margin of pterostigma), straight, distinctly inclined toward base. Pterostigma elongate, slightly longer than in forewing. RA with three branches: first, within pterostigma, easily visible; second, forming distal margin of pterostigma, straight, oblique to margin; third, simple. RA terminating on margin about mid-way between pterostigma, apex. Three crossveins between RA, RP: 1ra-rp slightly distad 2r-m; 2ra-rp approximately opposite termination of ScP; 3ra-rp located in distal half of pterostigma along RA. RP originating at approximately 0.36 wing length. Anterior trace of RP simple apically, with four (left wing) to five (right wing) branches, in both wings one proximad 3ra-rp, three to four at beginning or distad 3ra-rp. RP1 deeply forked, its anterior branch simple, posterior forked once; RP2, RP
|
||
<quantity id="4C8F9B7DFFF3C44BFE65FD28CDAF82A9" box="[405,448,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="46" unit="in" value="3.0">3 in</quantity>
|
||
left wing forked; other branches simple. One intraradial crossvein (rp1-rp2), between stem of RP, RP1. Three crossveins between R, M: 1r-m long, running subparallel to R; 2r-m connects stem of RP, MA; 3r-m connecting RP1, anterior branch of MA. M forked approximately at level of origin of RP. MA deeply dichotomously forked twice. MP pectinately branched, with probably three branches (right wing: poorly preserved), four probably simple branches (left wing). Two intramedian crossveins form two
|
||
<emphasis id="B903EA8AFFF3C44BFBC9FCB8C8328339" box="[1081,1117,799,825]" italics="true" pageId="5" pageNumber="46">doi</emphasis>
|
||
; basal
|
||
<emphasis id="B903EA8AFFF3C44BFB40FCB8C8BB8339" box="[1200,1236,799,825]" italics="true" pageId="5" pageNumber="46">doi</emphasis>
|
||
markedly shorter than distal; one indistinct crossvein distad 2im (this may be artefact). One crossvein 2m-cu between M, Cu detected, connecting MP, CuA. Anterior trace of CuA simple distally, with one simple (but incompletely preserved) branch. CuP or CuP+AA1 partially preserved; anal veins not preserved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |