treatments-xml/data/03/DE/87/03DE8787FFF7FFB1FF09FC0DFD3E4B42.xml

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<mods:title id="DEF02A3AF8DF82C6EFA55320956BF91F">Systematics and phylogeny of Cristacoxidae (Copepoda, Harpacticoida): a review</mods:title>
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<subSubSection id="C36D651AFFF7FFB3FF09FC0DFD254D89" box="[151,765,994,1020]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BC83691FFF7FFB3FF09FC0DFD254D89" blockId="33.[151,1437,994,2028]" box="[151,765,994,1020]" pageId="33" pageNumber="34">
<heading id="D08081FDFFF7FFB3FF09FC0DFD254D89" bold="true" box="[151,765,994,1020]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<emphasis id="B903EA83FFF7FFB3FF09FC0DFD254D89" bold="true" box="[151,765,994,1020]" pageId="33" pageNumber="34">
Monophyly of 
<taxonomicName id="4C774D12FFF7FFB3FED3FC0DFD254D89" authority="George (2002)" authorityName="George" authorityYear="2002" box="[333,765,994,1020]" class="Maxillopoda" family="Cristacoxidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="family">
Cristacoxidae 
<emphasis id="B903EA83FFF7FFB3FD9FFC0DFD9E4D89" bold="true" box="[513,582,994,1020]" italics="true" pageId="33" pageNumber="34">sensu</emphasis>
<bibRefCitation id="EFE64B60FFF7FFB3FDD2FC0DFD254D89" author="George" box="[588,765,994,1020]" pageId="33" pageNumber="34" refString="George, K. H. (2002) New phylogenetic aspects of the Cristacoxidae Huys (Copepoda, Harpacticoida), including the description of a new genus from the Magellan Region. Vie et Milieu, 52 (1), 31 - 41." type="journal article" year="2002">George (2002)</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C36D651AFFF7FFB3FF58FBE7FA454999" pageId="33" pageNumber="34" type="description">
<paragraph id="8BC83691FFF7FFB3FF58FBE7FC5E4BE2" blockId="33.[151,1437,994,2028]" pageId="33" pageNumber="34">
Although George’s (2002) assessment of the position of 
<taxonomicName id="4C774D12FFF7FFB3FCF6FBE7FBE84A54" box="[872,1072,1032,1057]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FCF6FBE7FBE84A54" box="[872,1072,1032,1057]" italics="true" pageId="33" pageNumber="34">Laophontisochra</emphasis>
</taxonomicName>
was hampered by the absence of males, the evidence supporting his assignment of the genus to the 
<taxonomicName id="4C774D12FFF7FFB3FC49FBC2FBA34A32" box="[983,1147,1069,1095]" class="Maxillopoda" family="Cristacoxidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="family">Cristacoxidae</taxonomicName>
was nevertheless weak. Both 
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<emphasis id="B903EA83FFF7FFB3FF47FBBCFE7B4A19" box="[217,419,1107,1132]" italics="true" pageId="33" pageNumber="34">Laophontisochra</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF7FFB3FE40FBBCFD8C4A19" box="[478,596,1107,1132]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FE40FBBCFD8C4A19" box="[478,596,1107,1132]" italics="true" pageId="33" pageNumber="34">Acuticoxa</emphasis>
</taxonomicName>
have a radically divergent morphology, exhibiting only four (7, 9, 10, 11) of the 
<specimenCount id="9D71FD18FFF7FFB3FE8DFB97FE524AE7" box="[275,394,1144,1170]" pageId="33" pageNumber="34" type="female">12 female</specimenCount>
cristacoxid autapomorphies listed above. At least for some of these the question arises whether the apomorphic state observed is homologous to the typical cristacoxid condition: the inner basal spine/seta of P1 is displaced onto the anterior surface of the basis (character 7), P2–P4 exp-2 has lost the inner seta (character 9; however, the homologue of exp-2 is not expressed as a discrete segment, but its outer spine is), P2–P4 exp-3 has only two outer elements instead of three (character 10; however, these elements are not typically elongate and setiform as in 
<taxonomicName id="4C774D12FFF7FFB3FDD7FADCFCFA4B39" box="[585,802,1331,1356]" class="Maxillopoda" family="Canthocamptidae" genus="Noodtorthopsyllus" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FDD7FADCFCFA4B39" box="[585,802,1331,1356]" italics="true" pageId="33" pageNumber="34">Noodtorthopsyllus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF7FFB3FCC4FADCFC284B39" box="[858,1008,1331,1356]" class="Maxillopoda" family="Cristacoxidae" genus="Cubanocleta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FCC4FADCFC284B39" box="[858,1008,1331,1356]" italics="true" pageId="33" pageNumber="34">Cubanocleta</emphasis>
</taxonomicName>
), and P3–P4 endopods 1-segmented (character 11: the validity of this character is difficult to assess since the endopods are either lost (P 
<quantity id="4C8F9B74FFF7FFB3FAF0FAB7FA444B07" box="[1390,1436,1368,1394]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="33" pageNumber="34" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="4C774D12FFF7FFB3FF09FA92FE874BE3" box="[151,351,1405,1430]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FF09FA92FE874BE3" box="[151,351,1405,1430]" italics="true" pageId="33" pageNumber="34">Laophontisochra</emphasis>
</taxonomicName>
) or represented by a small unisetose segment).
</paragraph>
<paragraph id="8BC83691FFF7FFB3FF58FA4DFA454999" blockId="33.[151,1437,994,2028]" pageId="33" pageNumber="34">
<bibRefCitation id="EFE64B60FFF7FFB3FF58FA4DFEA94BC9" author="George" box="[198,369,1442,1468]" pageId="33" pageNumber="34" refString="George, K. H. (2002) New phylogenetic aspects of the Cristacoxidae Huys (Copepoda, Harpacticoida), including the description of a new genus from the Magellan Region. Vie et Milieu, 52 (1), 31 - 41." type="journal article" year="2002">George (2002)</bibRefCitation>
advocated a basal split of the 
<taxonomicName id="4C774D12FFF7FFB3FD51FA4DFCA84BC9" box="[719,880,1442,1468]" class="Maxillopoda" family="Cristacoxidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="family">Cristacoxidae</taxonomicName>
into two lineages, 
<emphasis id="B903EA83FFF7FFB3FBD2FA4CFBB74BC9" box="[1100,1135,1443,1468]" italics="true" pageId="33" pageNumber="34">i.e.</emphasis>
a clade uniting the “more derived” traditional cristacoxid genera (
<taxonomicName id="4C774D12FFF7FFB3FD3AFA27FCEA4B94" box="[676,818,1480,1505]" class="Maxillopoda" family="Canthocamptidae" genus="Cristacoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FD3AFA27FCEA4B94" box="[676,818,1480,1505]" italics="true" pageId="33" pageNumber="34">Cristacoxa</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C774D12FFF7FFB3FCDAFA27FBEB4B94" box="[836,1075,1480,1505]" class="Maxillopoda" family="Canthocamptidae" genus="Noodtorthopsyllus" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FCDAFA27FBEB4B94" box="[836,1075,1480,1505]" italics="true" pageId="33" pageNumber="34">Noodtorthopsyllus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C774D12FFF7FFB3FBDBFA27FB314B94" box="[1093,1257,1480,1505]" class="Maxillopoda" family="Cristacoxidae" genus="Cubanocleta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FBDBFA27FB314B94" box="[1093,1257,1480,1505]" italics="true" pageId="33" pageNumber="34">Cubanocleta</emphasis>
</taxonomicName>
) and a “more plesiomorphic” clade comprising 
<taxonomicName id="4C774D12FFF7FFB3FDCCFA02FCF74873" box="[594,815,1517,1542]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FDCCFA02FCF74873" box="[594,815,1517,1542]" italics="true" pageId="33" pageNumber="34">Laophontisochra</emphasis>
</taxonomicName>
(and by inference, 
<taxonomicName id="4C774D12FFF7FFB3FBACFA02FB6C4873" box="[1074,1204,1517,1542]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FBACFA02FB6C4873" box="[1074,1204,1517,1542]" italics="true" pageId="33" pageNumber="34">Acuticoxa</emphasis>
</taxonomicName>
). The sistergroup relationship (and therefore the monophyly of the family) was based on three character states, 
<emphasis id="B903EA83FFF7FFB3FA82F9FCFA9A4859" box="[1308,1346,1555,1580]" italics="true" pageId="33" pageNumber="34">i.e.</emphasis>
(1) the presence of cristae on the coxa of P1, (2) large maxillipeds, and (3) “atrophy of antennary exopod and abexopodal seta”. Each of these character states leave room for interpretation. 
<taxonomicName id="4C774D12FFF7FFB3FBE1F9B2FABF4803" box="[1151,1383,1629,1654]" class="Maxillopoda" family="Canthocamptidae" genus="Noodtorthopsyllus" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FBE1F9B2FABF4803" box="[1151,1383,1629,1654]" italics="true" pageId="33" pageNumber="34">Noodtorthopsyllus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF7FFB3FF09F96CFEF748E9" box="[151,303,1667,1692]" class="Maxillopoda" family="Cristacoxidae" genus="Cubanocleta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FF09F96CFEF748E9" box="[151,303,1667,1692]" italics="true" pageId="33" pageNumber="34">Cubanocleta</emphasis>
</taxonomicName>
have paired serrate crests on the coxa and a single one on the praecoxa of leg 1, in addition to serially homologous crests on the coxae of P2 and P3 and occasionally a lobate outgrowth on the coxa of P4. In 
<taxonomicName id="4C774D12FFF7FFB3FF29F922FEF54893" box="[183,301,1741,1766]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FF29F922FEF54893" box="[183,301,1741,1766]" italics="true" pageId="33" pageNumber="34">Acuticoxa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF7FFB3FEF6F922FDE94893" box="[360,561,1741,1766]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FEF6F922FDE94893" box="[360,561,1741,1766]" italics="true" pageId="33" pageNumber="34">Laophontisochra</emphasis>
</taxonomicName>
a single non-serrate, lobate or spinous outgrowth (not a genuine crest!) is present on the coxa of P1 but no such structure is discernible on the praecoxa. Serially homologous spinous projections are also present on the coxae of P2–P4 of 
<taxonomicName id="4C774D12FFF7FFB3FCABF8F7FC684944" box="[821,944,1816,1841]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FCABF8F7FC684944" box="[821,944,1816,1841]" italics="true" pageId="33" pageNumber="34">Acuticoxa</emphasis>
</taxonomicName>
but not in 
<taxonomicName id="4C774D12FFF7FFB3FBA7F8F7FAD34944" box="[1081,1291,1816,1841]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FBA7F8F7FAD34944" box="[1081,1291,1816,1841]" italics="true" pageId="33" pageNumber="34">Laophontisochra</emphasis>
</taxonomicName>
. The single pointed projection on the P1 coxa in 
<taxonomicName id="4C774D12FFF7FFB3FDD1F8D2FD1D4923" box="[591,709,1853,1878]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FDD1F8D2FD1D4923" box="[591,709,1853,1878]" italics="true" pageId="33" pageNumber="34">Acuticoxa</emphasis>
</taxonomicName>
clearly does not resemble the paired serrate cristae exhibited by 
<taxonomicName id="4C774D12FFF7FFB3FF20F88CFE474909" box="[190,415,1891,1916]" class="Maxillopoda" family="Canthocamptidae" genus="Noodtorthopsyllus" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FF20F88CFE474909" box="[190,415,1891,1916]" italics="true" pageId="33" pageNumber="34">Noodtorthopsyllus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF7FFB3FE42F88CFDA04909" box="[476,632,1891,1916]" class="Maxillopoda" family="Cristacoxidae" genus="Cubanocleta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF7FFB3FE42F88CFDA04909" box="[476,632,1891,1916]" italics="true" pageId="33" pageNumber="34">Cubanocleta</emphasis>
</taxonomicName>
and is probably the result of convergent evolution. Secondly, the maxillipeds are fundamentally different between 
<emphasis id="B903EA83FFF7FFB3FD7BF867FBEA49D4" box="[741,1074,1928,1954]" italics="true" pageId="33" pageNumber="34">
<taxonomicName id="4C774D12FFF7FFB3FD7BF867FC6849D4" box="[741,944,1928,1953]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Laophontisochra</taxonomicName>
- 
<taxonomicName id="4C774D12FFF7FFB3FC25F867FBEA49D4" box="[955,1074,1928,1953]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Acuticoxa</taxonomicName>
</emphasis>
(
<emphasis id="B903EA83FFF7FFB3FBDAF867FBA849D4" box="[1092,1136,1928,1953]" italics="true" pageId="33" pageNumber="34">L-A</emphasis>
) and 
<emphasis id="B903EA83FFF7FFB3FB28F867FEE849B3" italics="true" pageId="33" pageNumber="34">
<taxonomicName id="4C774D12FFF7FFB3FB28F867FA4A49D4" box="[1206,1426,1928,1953]" class="Maxillopoda" family="Canthocamptidae" genus="Noodtorthopsyllus" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Noodtorthopsyllus</taxonomicName>
- 
<taxonomicName id="4C774D12FFF7FFB3FF09F842FEE849B3" box="[151,304,1965,1990]" class="Maxillopoda" family="Cristacoxidae" genus="Cubanocleta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Cubanocleta</taxonomicName>
</emphasis>
(
<emphasis id="B903EA83FFF7FFB3FEDCF842FEAC49B3" box="[322,372,1965,1990]" italics="true" pageId="33" pageNumber="34">N-C</emphasis>
). In the former two genera the syncoxa is unarmed and the endopod is represented by a geniculate claw bearing a minute accompanying seta along its outer margin (secondarily lost in 
<emphasis id="B903EA83FFF7FFB3FA1FF83CFA454999" box="[1409,1437,2003,2028]" italics="true" pageId="33" pageNumber="34">A.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36D651AFFF4FFB1FF09FF77FD3E4B42" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="reference_group">
<paragraph id="8BC83691FFF4FFB0FF09FF77FE984A47" blockId="34.[151,1437,152,2007]" pageId="34" pageNumber="35">
<taxonomicName id="4C774D12FFF4FFB0FF09FF77FEFA4EC4" box="[151,290,152,177]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biarticulata">
<emphasis id="B903EA83FFF4FFB0FF09FF77FEFA4EC4" box="[151,290,152,177]" italics="true" pageId="34" pageNumber="35">biarticulata</emphasis>
</taxonomicName>
); in the latter the syncoxa has a pinnate seta and the endopodal claw is not geniculate, bearing a short accompanying seta along the outer margin and, more distally, a distinct long seta along the inner margin. Given these morphological differences, using maxillipedal size as evidence in support of a sistergroup relationship between 
<emphasis id="B903EA83FFF4FFB0FE0AFEE7FE184F54" box="[404,448,264,289]" italics="true" pageId="34" pageNumber="35">L-A</emphasis>
and 
<emphasis id="B903EA83FFF4FFB0FE65FEE7FDF44F54" box="[507,556,264,289]" italics="true" pageId="34" pageNumber="35">N-C</emphasis>
appears a weak argument. Finally, adult cristacoxids consistently lack the antennary exopod (its former position in copepodid I is indicated by a membranous insert in the adult) and the abexopodal seta on the allobasis. Such consistency is not observed in 
<taxonomicName id="4C774D12FFF4FFB0FC5EFEBDFB504F1E" box="[960,1160,338,363]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FC5EFEBDFB504F1E" box="[960,1160,338,363]" italics="true" pageId="34" pageNumber="35">Laophontisochra</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C774D12FFF4FFB0FB30FEBDFAFB4F1E" box="[1198,1315,338,363]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FB30FEBDFAFB4F1E" box="[1198,1315,338,363]" italics="true" pageId="34" pageNumber="35">Acuticoxa</emphasis>
</taxonomicName>
where the “atrophied” condition prevails, with the antennary exopod being absent (
<taxonomicName id="4C774D12FFF4FFB0FC70FE97FB724FE4" box="[1006,1194,376,401]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="ubatubaensis">
<emphasis id="B903EA83FFF4FFB0FC70FE97FB724FE4" box="[1006,1194,376,401]" italics="true" pageId="34" pageNumber="35">A. ubatubaensis</emphasis>
</taxonomicName>
) or represented by a seta (
<taxonomicName id="4C774D12FFF4FFB0FF4BFE72FEB74FC3" box="[213,367,413,438]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="maryamae">
<emphasis id="B903EA83FFF4FFB0FF4BFE72FEB74FC3" box="[213,367,413,438]" italics="true" pageId="34" pageNumber="35">L. maryamae</emphasis>
</taxonomicName>
) or a minute unisetose segment (
<taxonomicName id="4C774D12FFF4FFB0FD6FFE72FC434FC3" box="[753,923,413,438]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="biarticulata">
<emphasis id="B903EA83FFF4FFB0FD6FFE72FC434FC3" box="[753,923,413,438]" italics="true" pageId="34" pageNumber="35">A. biarticulata</emphasis>
</taxonomicName>
), and the abexopodal seta being lost only in 
<taxonomicName id="4C774D12FFF4FFB0FF09FE2DFEE94FAE" box="[151,305,450,475]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="maryamae">
<emphasis id="B903EA83FFF4FFB0FF09FE2DFEE94FAE" box="[151,305,450,475]" italics="true" pageId="34" pageNumber="35">L. maryamae</emphasis>
</taxonomicName>
. 
<bibRefCitation id="EFE64B60FFF4FFB0FEA1FE2DFE324FA9" author="George" box="[319,490,450,476]" pageId="34" pageNumber="35" refString="George, K. H. (2002) New phylogenetic aspects of the Cristacoxidae Huys (Copepoda, Harpacticoida), including the description of a new genus from the Magellan Region. Vie et Milieu, 52 (1), 31 - 41." type="journal article" year="2002">George (2002)</bibRefCitation>
pointed out that including 
<taxonomicName id="4C774D12FFF4FFB0FCB8FE2DFC364FAE" box="[806,1006,450,475]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FCB8FE2DFC364FAE" box="[806,1006,450,475]" italics="true" pageId="34" pageNumber="35">Laophontisochra</emphasis>
</taxonomicName>
in the 
<taxonomicName id="4C774D12FFF4FFB0FBA0FE2DFB074FA9" box="[1086,1247,450,476]" class="Maxillopoda" family="Cristacoxidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Cristacoxidae</taxonomicName>
would draw the family to a much more basal position in the Laophontoidea because of certain primitive characters displayed by 
<taxonomicName id="4C774D12FFF4FFB0FF23FDE2FE854C53" box="[189,349,525,550]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="maryamae">
<emphasis id="B903EA83FFF4FFB0FF23FDE2FE854C53" box="[189,349,525,550]" italics="true" pageId="34" pageNumber="35">L. maryamae</emphasis>
</taxonomicName>
such as the presence of two basal setae on the mandibular palp, a character thus far only reported from some primitive 
<taxonomicName id="4C774D12FFF4FFB0FD91FDDDFD604C39" box="[527,696,562,588]" class="Maxillopoda" family="Laophontidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Laophontidae</taxonomicName>
(subfamily Esolinae; 
<emphasis id="B903EA83FFF4FFB0FC55FDDDFC324C3E" box="[971,1002,562,587]" italics="true" pageId="34" pageNumber="35">cf.</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FC6DFDDDFB0B4C39" author="Huys" box="[1011,1235,562,588]" pageId="34" pageNumber="35" refString="Huys, R. &amp; Lee, W. (2000) Basal resolution of laophontid phylogeny and the paraphyly of Esola Edwards. Bulletin of the Natural History Museum (Zoology), 66, 49 - 107." type="journal article" year="2000">Huys &amp; Lee 2000</bibRefCitation>
) and effectively excluding both 
<taxonomicName id="4C774D12FFF4FFB0FEC5FDB7FE004C04" box="[347,472,600,625]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FEC5FDB7FE004C04" box="[347,472,600,625]" italics="true" pageId="34" pageNumber="35">Acuticoxa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF4FFB0FD89FDB7FD344C04" box="[535,748,600,625]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FD89FDB7FD344C04" box="[535,748,600,625]" italics="true" pageId="34" pageNumber="35">Laophontisochra</emphasis>
</taxonomicName>
from the clade [
<taxonomicName id="4C774D12FFF4FFB0FC25FDB7FB7E4C07" box="[955,1190,600,626]" class="Maxillopoda" family="Adenopleurellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Adenopleurellidae</taxonomicName>
+ 
<taxonomicName id="4C774D12FFF4FFB0FB56FDB7FA5A4C07" box="[1224,1410,600,626]" class="Maxillopoda" family="Orthopsyllidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Orthopsyllidae</taxonomicName>
+ 
<taxonomicName id="4C774D12FFF4FFB0FF09FD92FEB94CE2" box="[151,353,637,663]" class="Maxillopoda" family="Laophontopsidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Laophontopsidae</taxonomicName>
+ 
<taxonomicName id="4C774D12FFF4FFB0FEE1FD92FDFF4CE2" box="[383,551,637,663]" class="Maxillopoda" family="Cristacoxidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Cristacoxidae</taxonomicName>
]. Two of the cristacoxid autapomorphies mentioned above (characters 1 and 13) cannot be verified in either 
<taxonomicName id="4C774D12FFF4FFB0FD8EFD4DFD024CCE" box="[528,730,674,699]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FD8EFD4DFD024CCE" box="[528,730,674,699]" italics="true" pageId="34" pageNumber="35">Laophontisochra</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C774D12FFF4FFB0FC9CFD4DFCA04CCE" box="[770,888,674,699]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FC9CFD4DFCA04CCE" box="[770,888,674,699]" italics="true" pageId="34" pageNumber="35">Acuticoxa</emphasis>
</taxonomicName>
since they are based on male character states. However, the recent discovery of a closely related genus from Paranaguá Bay (
<collectingCountry id="F3607601FFF4FFB0FBD2FD27FB424C97" box="[1100,1178,712,738]" name="Brazil" pageId="34" pageNumber="35">Brazil</collectingCountry>
), represented by both sexes (named hereafter “Genus X”), showed that neither the spermatophore nor the P3 sexual dimorphism are of the cristacoxid 
<typeStatus id="54CC8833FFF4FFB0FEF2FCFDFE464D59" box="[364,414,786,812]" pageId="34" pageNumber="35">type</typeStatus>
(P.H.C. Corgosinho &amp; M. Büntzow, pers. commn), and that the male sixth legs bear 3 setae (except for the basal 
<taxonomicName id="4C774D12FFF4FFB0FE41FCD7FD4B4D27" box="[479,659,824,850]" class="Maxillopoda" family="Normanellidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Normanellidae</taxonomicName>
all other laophontoidean families have 2 setae; 
<emphasis id="B903EA83FFF4FFB0FB7EFCD7FB274D24" box="[1248,1279,824,849]" italics="true" pageId="34" pageNumber="35">cf.</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FA99FCD7FF0C4D02" author="Huys" pageId="34" pageNumber="35" refString="Huys, R. &amp; Lee, W. (1998 / 99) On the relationships of the Normanellidae and the recognition of Cletopsyllidae grad. nov. (Copepoda, Harpacticoida). Zoologischer Anzeiger, 267 - 290." type="journal article" year="1998">Huys &amp; Lee 1998</bibRefCitation>
/99). Neither 
<taxonomicName id="4C774D12FFF4FFB0FEF1FCB2FE3C4D03" box="[367,484,861,886]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FEF1FCB2FE3C4D03" box="[367,484,861,886]" italics="true" pageId="34" pageNumber="35">Acuticoxa</emphasis>
</taxonomicName>
nor 
<taxonomicName id="4C774D12FFF4FFB0FD84FCB2FD3A4D03" box="[538,738,861,886]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FD84FCB2FD3A4D03" box="[538,738,861,886]" italics="true" pageId="34" pageNumber="35">Laophontisochra</emphasis>
</taxonomicName>
have setiform outer elements on the distal exopod segment of P2–P4. The absence of this character would place these genera outside the clade [
<taxonomicName id="4C774D12FFF4FFB0FB32FC6DFA5A4DE9" box="[1196,1410,898,924]" class="Maxillopoda" family="Laophontopsidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Laophontopsidae</taxonomicName>
+ 
<taxonomicName id="4C774D12FFF4FFB0FF09FC47FE9A4DB7" box="[151,322,936,962]" class="Maxillopoda" family="Cristacoxidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Cristacoxidae</taxonomicName>
]. Although the males of both genera are unknown, the male of the closely related “Genus X” shows that the segments distal to the geniculation in the male antennule are free. The presence of a single compound segment distal to the geniculation is a synapomorphy uniting the 
<taxonomicName id="4C774D12FFF4FFB0FB18FC1DFABD4A79" box="[1158,1381,1010,1036]" class="Maxillopoda" family="Laophontopsidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Laophontopsidae</taxonomicName>
and 
<taxonomicName id="4C774D12FFF4FFB0FF09FBF7FEE44A47" box="[151,316,1048,1074]" class="Maxillopoda" family="Cristacoxidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Cristacoxidae</taxonomicName>
.
</paragraph>
<paragraph id="8BC83691FFF4FFB0FF58FBD2FEB84937" blockId="34.[151,1437,152,2007]" pageId="34" pageNumber="35">
The coxal projections on legs 
<quantity id="4C8F9B74FFF4FFB0FDB2FBD2FDAF4A22" box="[556,631,1085,1111]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="34" pageNumber="35" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1–4 in</quantity>
<taxonomicName id="4C774D12FFF4FFB0FDE1FBD2FC9F4A23" box="[639,839,1085,1110]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FDE1FBD2FC9F4A23" box="[639,839,1085,1110]" italics="true" pageId="34" pageNumber="35">Laophontisochra</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF4FFB0FC1DFBD2FC204A23" box="[899,1016,1085,1110]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FC1DFBD2FC204A23" box="[899,1016,1085,1110]" italics="true" pageId="34" pageNumber="35">Acuticoxa</emphasis>
</taxonomicName>
may be indicative of a relationship with some genera currently included in the 
<taxonomicName id="4C774D12FFF4FFB0FD5DFB8DFC4F4A09" box="[707,919,1122,1148]" class="Maxillopoda" family="Huntemanniidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Huntemanniidae</taxonomicName>
. 
<bibRefCitation id="EFE64B60FFF4FFB0FC3AFB8DFBA44A09" author="Por" box="[932,1148,1122,1148]" pageId="34" pageNumber="35" refString="Por, F. D. (1986 a) A re-evaluation of the family Cletodidae Sars, Lang (Copepoda, Harpacticoida). In: Schriever, G., Schminke, H. K. &amp; Shih, C. - t. (Eds), Proceedings of the Second International Conference on Copepoda, Ottawa, Canada, 13 - 17 August, 1984. Syllogeus, 58, 420 - 425." type="journal article" year="1986" yearSuffix="a">Por (1986a: 421)</bibRefCitation>
established the family 
<taxonomicName id="4C774D12FFF4FFB0FF09FB67FE864AD7" box="[151,350,1160,1186]" class="Maxillopoda" family="Huntemanniidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Huntemanniidae</taxonomicName>
for the genera 
<taxonomicName id="4C774D12FFF4FFB0FD86FB67FCEA4AD7" authority="Brady, 1880" authorityName="Brady" authorityYear="1880" box="[536,818,1160,1186]" class="Maxillopoda" family="Huntemanniidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FD86FB67FD4A4AD4" box="[536,658,1160,1185]" italics="true" pageId="34" pageNumber="35">Nannopus</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FD05FB67FCEA4AD7" author="Brady" box="[667,818,1160,1186]" pageId="34" pageNumber="35" refString="Brady, G. S. (1880) A Monograph of the free and semi-parasitic Copepoda of the British Islands, 2. The Ray Society, London, 182 pp." type="book" year="1880">Brady, 1880</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C774D12FFF4FFB0FCA1FB67FB5A4AD7" authority="Poppe, 1884" authorityName="Poppe" authorityYear="1884" box="[831,1154,1160,1186]" class="Maxillopoda" family="Huntemanniidae" genus="Huntemannia" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FCA1FB67FC3B4AD4" box="[831,995,1160,1185]" italics="true" pageId="34" pageNumber="35">Huntemannia</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FC75FB67FB5A4AD7" author="Poppe" box="[1003,1154,1160,1186]" pageId="34" pageNumber="35" refString="Poppe, S. A. (1884) Ein neues Copepoden-Genus aus der Jade. Abhandlungen herausgegeben vom naturwissenschaftlichen Verein zu Bremen, 9, 57 - 58." type="journal article" year="1884">Poppe, 1884</bibRefCitation>
</taxonomicName>
[
<typeStatus id="54CC8833FFF4FFB0FB0CFB67FB164AD7" box="[1170,1230,1160,1186]" pageId="34" pageNumber="35">type</typeStatus>
], 
<taxonomicName id="4C774D12FFF4FFB0FB7DFB67FEFC4AB2" authority="T. Scott, 1894" authorityName="T. Scott" authorityYear="1894" class="Maxillopoda" family="Cletodidae" genus="Pontopolites" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FB7DFB67FAA44AD4" box="[1251,1404,1160,1185]" italics="true" pageId="34" pageNumber="35">Pontopolites</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FA1AFB67FEFC4AB2" author="Scott" pageId="34" pageNumber="35" refString="Scott, T. (1894) Additions to the fauna of the Firth of Forth. Part VI. Reports of the Fishery Board for Scotland, Edinburgh, 12 (3), 231 - 271." type="journal article" year="1894">T. Scott, 1894</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C774D12FFF4FFB0FEACFB42FD124AB2" authority="Smirnov, 1946" authorityName="Smirnov" authorityYear="1946" box="[306,714,1197,1223]" class="Maxillopoda" family="Huntemanniidae" genus="Metahuntemannia" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FEACFB42FDD64AB3" box="[306,526,1197,1222]" italics="true" pageId="34" pageNumber="35">Metahuntemannia</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FD8BFB42FD124AB2" author="Smirnov" box="[533,714,1197,1223]" pageId="34" pageNumber="35" refString="Smirnov, S. S. (1946) Novye vidy Copepoda Harpacticoida iz severnogo ledovitogo okeana. [New species of Copepoda- Harpacticoida from the northern Arctic Ocean]. Trudy Dreifuyuschei Ekspeditsii Glavsevmorputi na Ledokol'nom Parokhode ' G. Sedov', 1937 - 1940 gg, 3, 231 - 263. [In Russian]." type="journal article" year="1946">Smirnov, 1946</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C774D12FFF4FFB0FD46FB42FC084AB2" ID-CoL="8KV9W" authority="Por, 1986" authorityName="Por" authorityYear="1986" box="[728,976,1197,1223]" class="Insecta" family="Carabidae" genus="Beckeria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FD46FB42FC9B4AB3" box="[728,835,1197,1222]" italics="true" pageId="34" pageNumber="35">Beckeria</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FCD2FB42FC084AB2" author="Por" box="[844,976,1197,1223]" pageId="34" pageNumber="35" refString="Por, F. D. (1986 b) New deepsea Harpacticoidea (Copepoda) of cletodid type, collected in the Indian Ocean by R / V &quot; Anton Bruun &quot; in 1964. Crustaceana, 50, 78 - 98." type="journal article" year="1986" yearSuffix="b">Por, 1986b</bibRefCitation>
</taxonomicName>
and possibly 
<taxonomicName id="4C774D12FFF4FFB0FBE4FB42FEFB4A99" authority="Scott &amp; Scott, 1893" authorityName="Scott &amp; Scott" authorityYear="1893" class="Maxillopoda" family="Huntemanniidae" genus="Pseudocletodes" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FBE4FB42FAEE4AB3" box="[1146,1334,1197,1222]" italics="true" pageId="34" pageNumber="35">Pseudocletodes</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FAA0FB42FEFB4A99" author="Scott" pageId="34" pageNumber="35" refString="Scott, T. &amp; Scott, A. (1893) On some new or rare Crustacea from Scotland. Annals and Magazine of natural History, (6) 12, 237 - 246." type="journal article" year="1893">Scott &amp; Scott, 1893</bibRefCitation>
</taxonomicName>
. Unfortunately, it has remained unnoticed that 
<bibRefCitation id="EFE64B60FFF4FFB0FCCCFB3DFBF74A99" author="Brady" box="[850,1071,1234,1260]" pageId="34" pageNumber="35" refString="Brady, G. S. (1880) A Monograph of the free and semi-parasitic Copepoda of the British Islands, 2. The Ray Society, London, 182 pp." type="book" year="1880">Brady (1880: 100)</bibRefCitation>
had already established a new subfamily Nannopinae within the 
<taxonomicName id="4C774D12FFF4FFB0FDAFFB17FD0F4B67" box="[561,727,1272,1298]" class="Maxillopoda" family="Harpacticidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Harpacticidae</taxonomicName>
for 
<taxonomicName id="4C774D12FFF4FFB0FC95FB17FBF84B67" authority="Brady, 1880" authorityName="Brady" authorityYear="1880" box="[779,1056,1272,1298]" class="Maxillopoda" family="Huntemanniidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FC95FB17FC5C4B64" box="[779,900,1272,1297]" italics="true" pageId="34" pageNumber="35">Nannopus</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FC12FB17FBF84B67" author="Brady" box="[908,1056,1272,1298]" pageId="34" pageNumber="35" refString="Brady, G. S. (1880) A Monograph of the free and semi-parasitic Copepoda of the British Islands, 2. The Ray Society, London, 182 pp." type="book" year="1880">Brady, 1880</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54CC8833FFF4FFB0FBAEFB17FBBD4B67" box="[1072,1125,1272,1298]" pageId="34" pageNumber="35">type</typeStatus>
genus) and 
<taxonomicName id="4C774D12FFF4FFB0FB67FB17FEFF4B42" authority="Brady, 1880" authorityName="Brady" authorityYear="1880" class="Maxillopoda" family="Laophontidae" genus="Platychelipus" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FB67FB17FA444B64" box="[1273,1436,1272,1297]" italics="true" pageId="34" pageNumber="35">Platychelipus</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FF09FAF2FEFF4B42" author="Brady" box="[151,295,1309,1335]" pageId="34" pageNumber="35" refString="Brady, G. S. (1880) A Monograph of the free and semi-parasitic Copepoda of the British Islands, 2. The Ray Society, London, 182 pp." type="book" year="1880">Brady, 1880</bibRefCitation>
</taxonomicName>
(now placed in the 
<taxonomicName id="4C774D12FFF4FFB0FD91FAF2FD6E4B42" box="[527,694,1309,1335]" class="Maxillopoda" family="Laophontidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Laophontidae</taxonomicName>
). According to the Principle of Coordination applied to familygroup names (ICZN Art. 36.1) 
<bibRefCitation id="EFE64B60FFF4FFB0FD9CFAADFD474B29" author="Brady" box="[514,671,1346,1372]" pageId="34" pageNumber="35" refString="Brady, G. S. (1880) A Monograph of the free and semi-parasitic Copepoda of the British Islands, 2. The Ray Society, London, 182 pp." type="book" year="1880">Brady (1880)</bibRefCitation>
is deemed also to have simultaneously established the coordinate family name Nannopidae. Since the family-group name 
<taxonomicName id="4C774D12FFF4FFB0FCB1FA87FC294BF7" box="[815,1009,1384,1410]" class="Maxillopoda" family="Huntemanniidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Huntemanniidae</taxonomicName>
was used by 
<bibRefCitation id="EFE64B60FFF4FFB0FB0CFA87FAC64BF7" author="Por" box="[1170,1310,1384,1410]" pageId="34" pageNumber="35" refString="Por, F. D. (1986 a) A re-evaluation of the family Cletodidae Sars, Lang (Copepoda, Harpacticoida). In: Schriever, G., Schminke, H. K. &amp; Shih, C. - t. (Eds), Proceedings of the Second International Conference on Copepoda, Ottawa, Canada, 13 - 17 August, 1984. Syllogeus, 58, 420 - 425." type="journal article" year="1986" yearSuffix="a">Por (1986a)</bibRefCitation>
to include the genus 
<taxonomicName id="4C774D12FFF4FFB0FE8EFA62FE5F4BD3" box="[272,391,1421,1446]" class="Maxillopoda" family="Huntemanniidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FE8EFA62FE5F4BD3" box="[272,391,1421,1446]" italics="true" pageId="34" pageNumber="35">Nannopus</emphasis>
</taxonomicName>
, it must sink as a junior synonym of Nannopinae 
<bibRefCitation id="EFE64B60FFF4FFB0FC47FA62FBB64BD2" author="Brady" box="[985,1134,1421,1447]" pageId="34" pageNumber="35" refString="Brady, G. S. (1880) A Monograph of the free and semi-parasitic Copepoda of the British Islands, 2. The Ray Society, London, 182 pp." type="book" year="1880">Brady, 1880</bibRefCitation>
. Huys (2009) pointed out that the second part of the generic name 
<taxonomicName id="4C774D12FFF4FFB0FDE1FA5CFD2F4BB9" box="[639,759,1459,1484]" class="Maxillopoda" family="Huntemanniidae" genus="Nannopus" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FDE1FA5CFD2F4BB9" box="[639,759,1459,1484]" italics="true" pageId="34" pageNumber="35">Nannopus</emphasis>
</taxonomicName>
is derived from the Greek stem Oυς, meaning foot, and hence the family name must be corrected to 
<taxonomicName id="4C774D12FFF4FFB0FD3DFA37FC8A4B87" box="[675,850,1496,1522]" class="Maxillopoda" family="Nannopodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Nannopodidae</taxonomicName>
. Since Por’s (1986a) proposal, 
<bibRefCitation id="EFE64B60FFF4FFB0FB5DFA37FF3D4862" author="Dahms" pageId="34" pageNumber="35" refString="Dahms, H. - U. &amp; Pottek, M. (1992) Metahuntemannia Smirnov, 1946 and Talpina gen. nov. (Copepoda, Harpacticoida) from the deep-sea of the high Antarctic Weddell Sea with a description of eight new species. Microfauna Marina, 7, 7 - 78." type="journal article" year="1992">Dahms and Pottek (1992)</bibRefCitation>
have relegated 
<taxonomicName id="4C774D12FFF4FFB0FE3EFA12FDD14863" ID-CoL="8KV9W" box="[416,521,1533,1558]" class="Insecta" family="Carabidae" genus="Beckeria" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FE3EFA12FDD14863" box="[416,521,1533,1558]" italics="true" pageId="34" pageNumber="35">Beckeria</emphasis>
</taxonomicName>
to a junior subjective syonym of 
<taxonomicName id="4C774D12FFF4FFB0FC06FA12FA444862" authority="Kihara and Huys (2009)" authorityName="Kihara and Huys" authorityYear="2009" box="[920,1436,1533,1559]" class="Maxillopoda" family="Huntemanniidae" genus="Metahuntemannia" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FC06FA12FBB54863" box="[920,1133,1533,1558]" italics="true" pageId="34" pageNumber="35">Metahuntemannia</emphasis>
, 
<bibRefCitation id="EFE64B60FFF4FFB0FBE3FA12FA444862" author="Kihara" box="[1149,1436,1533,1559]" pageId="34" pageNumber="35" refString="Kihara, T. C. &amp; Huys, R. (2009) Contributions to the taxonomy of the Normanellidae (Copepoda, Harpacticoida): description of a new genus from the Brazilian continental shelf and re-assignment of Pseudocletodes vararensis Scott &amp; Scott, 1893 (ex Nannopodidae). Zootaxa, 2233, 1 - 38." type="journal article" year="2009">Kihara and Huys (2009)</bibRefCitation>
</taxonomicName>
have assigned 
<taxonomicName id="4C774D12FFF4FFB0FECAF9CCFDC34849" box="[340,539,1571,1596]" class="Maxillopoda" family="Huntemanniidae" genus="Pseudocletodes" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FECAF9CCFDC34849" box="[340,539,1571,1596]" italics="true" pageId="34" pageNumber="35">Pseudocletodes</emphasis>
</taxonomicName>
to the 
<taxonomicName id="4C774D12FFF4FFB0FDE6F9CDFCEE4849" box="[632,822,1570,1596]" class="Maxillopoda" family="Normanellidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Normanellidae</taxonomicName>
and two genera have been added to the family, 
<taxonomicName id="4C774D12FFF4FFB0FF09F9A7FE0B4817" authority="Wells, 1985" authorityName="Wells" authorityYear="1985" box="[151,467,1608,1634]" class="Maxillopoda" family="Nannopodidae" genus="Rosacletodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FF09F9A7FEE14814" box="[151,313,1608,1633]" italics="true" pageId="34" pageNumber="35">Rosacletodes</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FEDFF9A7FE0B4817" author="Wells" box="[321,467,1608,1634]" pageId="34" pageNumber="35" refString="Wells, J. B. J. (1985) Keys to aid in the identification of marine harpacticoid copepods. Amendment Bulletin no. 5. Zoological Publications Victoria University of Wellington, 80, 1 - 19." type="journal article" year="1985">Wells, 1985</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF4FFB0FD8FF9A7FCCA4817" authority="Huys, 2009" authorityName="Huys" authorityYear="2009" box="[529,786,1608,1634]" class="Maxillopoda" family="Canthocamptidae" genus="Pottekia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FD8FF9A7FDAF4814" box="[529,631,1608,1633]" italics="true" pageId="34" pageNumber="35">Pottekia</emphasis>
Huys, 2009
</taxonomicName>
. The former was proposed by 
<bibRefCitation id="EFE64B60FFF4FFB0FB14F9A7FAF14817" author="Wells" box="[1162,1321,1608,1634]" pageId="34" pageNumber="35" refString="Wells, J. B. J. (1985) Keys to aid in the identification of marine harpacticoid copepods. Amendment Bulletin no. 5. Zoological Publications Victoria University of Wellington, 80, 1 - 19." type="journal article" year="1985">Wells (1985)</bibRefCitation>
as a new replacement name for the cletodid genus 
<taxonomicName id="4C774D12FFF4FFB0FD3CF982FBC548F2" authority="Pallares, 1982" authorityName="Pallares" authorityYear="1982" box="[674,1053,1645,1671]" class="Maxillopoda" family="Nannopodidae" genus="Echinocletodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FD3CF982FCB948F3" box="[674,865,1645,1670]" italics="true" pageId="34" pageNumber="35">Echinocletodes</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FCF4F982FBC548F2" author="Pallares" box="[874,1053,1645,1671]" pageId="34" pageNumber="35" refString="Pallares, R. E. (1982) Copepodos harpacticoides marinos de Tierra del Fuego (Argentina). IV. Bahia Tethis. Contribuciones cientifica del Centro de Investigacion de Biologia marina, Buenos Aires, 186, 3 - 39." type="journal article" year="1982">Pallares, 1982</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54CC8833FFF4FFB0FBB0F982FBBE48F2" box="[1070,1126,1645,1671]" pageId="34" pageNumber="35">type</typeStatus>
species 
<taxonomicName id="4C774D12FFF4FFB0FB4DF982FE9048D9" authority="Pallares, 1982" authorityName="Pallares" authorityYear="1982" class="Maxillopoda" family="Nannopodidae" genus="Echinocletodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="kuehnemanni">
<emphasis id="B903EA83FFF4FFB0FB4DF982FA4548F3" box="[1235,1437,1645,1670]" italics="true" pageId="34" pageNumber="35">E. kuehnemanni</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FF09F97DFE9048D9" author="Pallares" box="[151,328,1682,1708]" pageId="34" pageNumber="35" refString="Pallares, R. E. (1982) Copepodos harpacticoides marinos de Tierra del Fuego (Argentina). IV. Bahia Tethis. Contribuciones cientifica del Centro de Investigacion de Biologia marina, Buenos Aires, 186, 3 - 39." type="journal article" year="1982">Pallares, 1982</bibRefCitation>
</taxonomicName>
), a junior homonym of 
<taxonomicName id="4C774D12FFF4FFB0FDF7F97CFC6C48D9" authority="Lang, 1936" authorityName="Lang" authorityYear="1936" box="[617,948,1682,1708]" class="Maxillopoda" family="Nannopodidae" genus="Echinocletodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF4FFB0FDF7F97CFCFA48D9" box="[617,802,1683,1708]" italics="true" pageId="34" pageNumber="35">Echinocletodes</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FCB4F97DFC6C48D9" author="Lang" box="[810,948,1682,1708]" pageId="34" pageNumber="35" refString="Lang, K. (1936) Die Familie der Cletodidae Sars, 1909. Zoologische Jahrbucher fur Systematik, 68, 445 - 480." type="journal article" year="1936">Lang, 1936</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54CC8833FFF4FFB0FC58F97DFC2448D9" box="[966,1020,1682,1708]" pageId="34" pageNumber="35">type</typeStatus>
species 
<taxonomicName id="4C774D12FFF4FFB0FBFAF97CFF0D48A7" authority="T. Scott, 1903" authorityName="T. Scott" authorityYear="1903" class="Maxillopoda" family="Cletodidae" genus="Cletodes" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="armata">
<emphasis id="B903EA83FFF4FFB0FBFAF97CFAF548D9" box="[1124,1325,1683,1708]" italics="true" pageId="34" pageNumber="35">Cletodes armata</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FAA8F97DFF0D48A7" author="Scott" pageId="34" pageNumber="35" refString="Scott, T. (1903) Notes on some Copepoda from the Faroe Channel. Journal of the Linnean Society, Zoology, 29, 1 - 11." type="journal article" year="1903">T. Scott, 1903</bibRefCitation>
</taxonomicName>
; 
<emphasis id="B903EA83FFF4FFB0FF7EF957FF2548A4" box="[224,253,1720,1745]" italics="true" pageId="34" pageNumber="35">cf.</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FE9AF957FE0D48A7" author="Huys" box="[260,469,1720,1746]" pageId="34" pageNumber="35" refString="Huys, R., Bodiou, J. - Y. &amp; Bodin, P. (1996 a) A revision of Psamathea (Harpacticoida: Leptastacidae) with description of P. brittanica sp. nov. Vie et Milieu, 46, 7 - 19." type="journal article" year="1996" yearSuffix="a">
Huys 
<emphasis id="B903EA83FFF4FFB0FED6F957FE5A48A4" box="[328,386,1720,1745]" italics="true" pageId="34" pageNumber="35">et al.</emphasis>
1996a
</bibRefCitation>
: 75), and is of particular interest here. 
<bibRefCitation id="EFE64B60FFF4FFB0FC0DF957FBE848A7" author="Bodin" box="[915,1072,1720,1746]" pageId="34" pageNumber="35" refString="Bodin, P. (1997) Catalogue of the new marine harpacticoid copepods (1997 edition). Documents de Travail, Institut royal des Sciences naturelles de Belgique, 89, 1 - 304." type="journal article" year="1997">Bodin (1997)</bibRefCitation>
did not assign the genus to any of the families defined by 
<bibRefCitation id="EFE64B60FFF4FFB0FE49F932FDBF4882" author="Por" box="[471,615,1757,1783]" pageId="34" pageNumber="35" refString="Por, F. D. (1986 a) A re-evaluation of the family Cletodidae Sars, Lang (Copepoda, Harpacticoida). In: Schriever, G., Schminke, H. K. &amp; Shih, C. - t. (Eds), Proceedings of the Second International Conference on Copepoda, Ottawa, Canada, 13 - 17 August, 1984. Syllogeus, 58, 420 - 425." type="journal article" year="1986" yearSuffix="a">Por (1986a)</bibRefCitation>
while 
<bibRefCitation id="EFE64B60FFF4FFB0FD24F932FCB24882" author="George" box="[698,874,1757,1783]" pageId="34" pageNumber="35" refString="George, K. H. (2008) Argestes angolaensis sp. nov. (Copepoda: Harpacticoida: Argestidae) from the Angola Basin (Southeast Atlantic), and the phylogenetic characterization of the taxon Argestes Sars, including the redescription of A. mollis Sars, 1910, and A. reductus (Ito, 1983). Zootaxa, 1866, 223 - 262." type="journal article" year="2008">George (2008)</bibRefCitation>
considered 
<taxonomicName id="4C774D12FFF4FFB0FC63F932FA5E4882" authority="Pallares, 1982" authorityName="Pallares" authorityYear="1982" box="[1021,1414,1757,1783]" class="Maxillopoda" family="Nannopodidae" genus="Echinocletodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="kuehnemanni">
<emphasis id="B903EA83FFF4FFB0FC63F932FB674883" box="[1021,1215,1757,1782]" italics="true" pageId="34" pageNumber="35">A. kuehnemanni</emphasis>
(
<bibRefCitation id="EFE64B60FFF4FFB0FB51F932FAA64882" author="Pallares" box="[1231,1406,1757,1783]" pageId="34" pageNumber="35" refString="Pallares, R. E. (1982) Copepodos harpacticoides marinos de Tierra del Fuego (Argentina). IV. Bahia Tethis. Contribuciones cientifica del Centro de Investigacion de Biologia marina, Buenos Aires, 186, 3 - 39." type="journal article" year="1982">Pallares, 1982</bibRefCitation>
)
</taxonomicName>
a 
<emphasis id="B903EA83FFF4FFB0FF09F8ECFE4F4969" box="[151,407,1795,1820]" italics="true" pageId="34" pageNumber="35">species incertae sedis</emphasis>
in the 
<taxonomicName id="4C774D12FFF4FFB0FE77F8EDFDB34969" box="[489,619,1794,1820]" class="Maxillopoda" family="Argestidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Argestidae</taxonomicName>
. Both 
<bibRefCitation id="EFE64B60FFF4FFB0FD26F8EDFC444969" box="[696,924,1794,1820]" pageId="34" pageNumber="35" refString="Huys, R., Bodiou, J. - Y. &amp; Bodin, P. (1996 a) A revision of Psamathea (Harpacticoida: Leptastacidae) with description of P. brittanica sp. nov. Vie et Milieu, 46, 7 - 19." type="journal article">
Huys 
<emphasis id="B903EA83FFF4FFB0FD60F8ECFCE14969" box="[766,825,1795,1820]" italics="true" pageId="34" pageNumber="35">et al.</emphasis>
(1996a)
</bibRefCitation>
and 
<bibRefCitation id="EFE64B60FFF4FFB0FC49F8EDFBB74969" author="Wells" box="[983,1135,1794,1820]" pageId="34" pageNumber="35" refString="Wells, J. B. J. (2007) An annotated checklist and keys to the species of Copepoda Harpacticoida (Crustacea). Zootaxa, 1568, 1 - 872." type="journal article" year="2007">Wells (2007)</bibRefCitation>
listed it as a genus in the 
<taxonomicName id="4C774D12FFF4FFB0FF09F8C7FE854937" box="[151,349,1832,1858]" class="Maxillopoda" family="Huntemanniidae" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Huntemanniidae</taxonomicName>
.
</paragraph>
<paragraph id="8BC83691FFF4FFB1FF5BF8A2FAC24D92" blockId="34.[151,1437,152,2007]" lastBlockId="35.[151,1437,152,1335]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
Pallares’ (1982) description of 
<taxonomicName id="4C774D12FFF4FFB0FDD6F8A2FC4E4913" box="[584,918,1869,1894]" class="Maxillopoda" family="Nannopodidae" genus="Rosacletodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="kuehnemanni">
<emphasis id="B903EA83FFF4FFB0FDD6F8A2FC4E4913" box="[584,918,1869,1894]" italics="true" pageId="34" pageNumber="35">Rosacletodes kuehnemanni</emphasis>
</taxonomicName>
, based on material from Tierra del Fuego (
<collectingCountry id="F3607601FFF4FFB0FF01F89DFEC549F9" box="[159,285,1906,1932]" name="Argentina" pageId="34" pageNumber="35">Argentina</collectingCountry>
), clearly shows spinulose projections on the coxae of P1–P4 which are virtually identical to the structure found on the coxa of P 
<quantity id="4C8F9B74FFF4FFB0FDA3F877FDB349C7" box="[573,619,1944,1970]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="34" pageNumber="35" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4C774D12FFF4FFB0FDEAF877FCC549C4" box="[628,797,1944,1969]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="maryamae">
<emphasis id="B903EA83FFF4FFB0FDEAF877FCC549C4" box="[628,797,1944,1969]" italics="true" pageId="34" pageNumber="35">L. maryamae</emphasis>
</taxonomicName>
. Such structures were also recently described for 
<taxonomicName id="4C774D12FFF4FFB0FF09F852FDE649A2" authority="Poppe, 1884" authorityName="Poppe" authorityYear="1884" box="[151,574,1981,2007]" class="Maxillopoda" family="Huntemanniidae" genus="Huntemannia" kingdom="Animalia" order="Harpacticoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="jadensis">
<emphasis id="B903EA83FFF4FFB0FF09F852FE7949A3" box="[151,417,1981,2006]" italics="true" pageId="34" pageNumber="35">Huntemannia jadensis</emphasis>
<bibRefCitation id="EFE64B60FFF4FFB0FE37F852FDE649A2" author="Poppe" box="[425,574,1981,2007]" pageId="34" pageNumber="35" refString="Poppe, S. A. (1884) Ein neues Copepoden-Genus aus der Jade. Abhandlungen herausgegeben vom naturwissenschaftlichen Verein zu Bremen, 9, 57 - 58." type="journal article" year="1884">Poppe, 1884</bibRefCitation>
</taxonomicName>
by 
<bibRefCitation id="EFE64B60FFF4FFB0FDF2F852FC3949A2" author="Kornev" box="[620,993,1981,2007]" pageId="34" pageNumber="35" refString="Kornev, P. N. &amp; Chertoprud, E. S. (2008) Harpacticoid Copepods of the White Sea: Morphology, Systematics, Ecology. KMK Scientific Press Ltd, Moscow, 379 pp." type="book" year="2008">Kornev and Chertoprud (2008)</bibRefCitation>
. There is also a gross resemblance in the general morphology of the swimming legs (P2–P4) between 
<taxonomicName id="4C774D12FFF5FFB1FC1BFF77FBF84EC4" box="[901,1056,152,177]" class="Maxillopoda" family="Nannopodidae" genus="Rosacletodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FC1BFF77FBF84EC4" box="[901,1056,152,177]" italics="true" pageId="35" pageNumber="36">Rosacletodes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF5FFB1FBC9FF77FAC74EC4" box="[1111,1311,152,177]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FBC9FF77FAC74EC4" box="[1111,1311,152,177]" italics="true" pageId="35" pageNumber="36">Laophontisochra</emphasis>
</taxonomicName>
, including the strongly reduced endopods (represented by a single seta in 
<taxonomicName id="4C774D12FFF5FFB1FC2AFF52FB814EA3" box="[948,1113,189,214]" class="Maxillopoda" family="Nannopodidae" genus="Rosacletodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FC2AFF52FB814EA3" box="[948,1113,189,214]" italics="true" pageId="35" pageNumber="36">Rosacletodes</emphasis>
</taxonomicName>
) and exopods (at most 2- segmented in the Ƥ), the enlarged outer spine on P2–P4 exp-1 (proximal outer spine if only one segment expressed), and the outer basal seta originating from an articulated setophore (this is shown for at least P4–P 
<quantity id="4C8F9B74FFF5FFB1FA11FEE7FF764F32" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="35" pageNumber="36" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="4C774D12FFF5FFB1FF2AFEC2FEA94F33" box="[180,369,301,326]" class="Maxillopoda" family="Nannopodidae" genus="Rosacletodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kuehnemanni">
<emphasis id="B903EA83FFF5FFB1FF2AFEC2FEA94F33" box="[180,369,301,326]" italics="true" pageId="35" pageNumber="36">R. kuehnemanni</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF5FFB1FE37FEC2FD9C4F33" box="[425,580,301,326]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="maryamae">
<emphasis id="B903EA83FFF5FFB1FE37FEC2FD9C4F33" box="[425,580,301,326]" italics="true" pageId="35" pageNumber="36">L. maryamae</emphasis>
</taxonomicName>
and has as yet to be confirmed for P2–P3). Additional similarity is found in the plesiomorphic condition of the mandibular palp (2-segmented, 2 basal setae, exopod represented by single seta, endopod by 3) and the apomorphic reduction of caudal ramus seta V. 
<taxonomicName id="4C774D12FFF5FFB1FBCBFE97FB284FE4" box="[1109,1264,376,401]" class="Maxillopoda" family="Nannopodidae" genus="Rosacletodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FBCBFE97FB284FE4" box="[1109,1264,376,401]" italics="true" pageId="35" pageNumber="36">Rosacletodes</emphasis>
</taxonomicName>
shows several plesiomorphic character states not found in 
<taxonomicName id="4C774D12FFF5FFB1FD5CFE72FC4F4FC3" box="[706,919,413,438]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FD5CFE72FC4F4FC3" box="[706,919,413,438]" italics="true" pageId="35" pageNumber="36">Laophontisochra</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C774D12FFF5FFB1FC5DFE72FB984FC3" box="[963,1088,413,438]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FC5DFE72FB984FC3" box="[963,1088,413,438]" italics="true" pageId="35" pageNumber="36">Acuticoxa</emphasis>
</taxonomicName>
such as the 5-segmented Ƥ antennule (with aesthetasc on segment IV), the trisetose antennary exopod and the biramous Ƥ P5 with six elements on the endopodal lobe and five on the exopod. Prehensility of the P1 endopod as displayed in the nannopodid genera 
<taxonomicName id="4C774D12FFF5FFB1FE08FDE2FDAA4C53" box="[406,626,525,550]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FE08FDE2FDAA4C53" box="[406,626,525,550]" italics="true" pageId="35" pageNumber="36">Laophontisochra</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF5FFB1FD2DFDE2FCEC4C53" box="[691,820,525,550]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FD2DFDE2FCEC4C53" box="[691,820,525,550]" italics="true" pageId="35" pageNumber="36">Acuticoxa</emphasis>
</taxonomicName>
is a morphological adaptation that has evolved secondarily and convergently in response to changing environments in many other harpacticoid families such as the 
<taxonomicName id="4C774D12FFF5FFB1FF7EFDB7FE7B4C07" box="[224,419,600,626]" class="Maxillopoda" family="Ectinosomatidae" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="family">Ectinosomatidae</taxonomicName>
(
<emphasis id="B903EA83FFF5FFB1FE2AFDB7FE064C04" box="[436,478,600,625]" italics="true" pageId="35" pageNumber="36">e.g.</emphasis>
<bibRefCitation id="EFE64B60FFF5FFB1FE7BFDB7FCDA4C07" author="Hicks" box="[485,770,600,626]" pageId="35" pageNumber="36" refString="Hicks, G. R. F. &amp; Schriever, G. (1983) A new genus and species of Ectinosomatidae (Copepoda, Harpacticoida) based on the original material of Dr. h. c. Walter Klie. Mitteilungen aus dem zoologischen Museum der Universitat Kiel, 2 (1), 1 - 7." type="journal article" year="1983">Hicks &amp; Schriever 1983</bibRefCitation>
), 
<taxonomicName id="4C774D12FFF5FFB1FC88FDB7FC334C07" box="[790,1003,600,626]" class="Maxillopoda" family="Pseudotachidiidae" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="family">Pseudotachidiidae</taxonomicName>
(
<emphasis id="B903EA83FFF5FFB1FC62FDB7FBFE4C04" box="[1020,1062,600,625]" italics="true" pageId="35" pageNumber="36">e.g.</emphasis>
<bibRefCitation id="EFE64B60FFF5FFB1FBB3FDB7FB6E4C07" author="Hicks" box="[1069,1206,600,626]" pageId="35" pageNumber="36" refString="Hicks, G. R. F. (1988) Systematics of the Donsiellidae Lang (Copepoda, Harpacticoida). Journal of natural History, 22, 639 - 684." type="journal article" year="1988">Hicks 1988</bibRefCitation>
), 
<taxonomicName id="4C774D12FFF5FFB1FB54FDB7FA444C07" box="[1226,1436,600,626]" class="Maxillopoda" family="Cylindropsyllidae" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="family">Cylindropsyllidae</taxonomicName>
(
<emphasis id="B903EA83FFF5FFB1FF3FFD92FF174CE3" box="[161,207,637,662]" italics="true" pageId="35" pageNumber="36">e.g.</emphasis>
<bibRefCitation id="EFE64B60FFF5FFB1FF49FD92FE284CE2" author="Huys" box="[215,496,637,663]" pageId="35" pageNumber="36" refString="Huys, R. &amp; Willems, K. A. (1993) A revision of Cylinula and two new species of Cylindropsyllus (Copepoda, Harpacticoida, Cylindropsyllidae). Zoologica Scripta, 22, 347 - 362." type="journal article" year="1993">Huys &amp; Willems 1993</bibRefCitation>
) and 
<taxonomicName id="4C774D12FFF5FFB1FDA9FD92FD3A4CE2" box="[567,738,637,663]" class="Maxillopoda" family="Leptastacidae" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="family">Leptastacidae</taxonomicName>
(
<bibRefCitation id="EFE64B60FFF5FFB1FD6DFD92FC014CE2" author="Huys" box="[755,985,637,663]" pageId="35" pageNumber="36" refString="Huys, R., Gee, J. M., Moore, C. G. &amp; Hamond, R. (1996 b) Marine and brackish water harpacticoid copepods. Part 1. In: Barnes, R. S. K. &amp; J. H. Crothers (Eds), Synopses of the British Fauna (New Series), 51: i - viii, 1 - 352. Field Studies Council, Shrewsbury." type="journal article" year="1996" yearSuffix="b">
Huys 
<emphasis id="B903EA83FFF5FFB1FCA0FD92FCA64CE3" box="[830,894,637,662]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
1996b
</bibRefCitation>
). The non-prehensile P1 endopod in 
<taxonomicName id="4C774D12FFF5FFB1FF09FD4DFEE14CCE" box="[151,313,674,699]" class="Maxillopoda" family="Nannopodidae" genus="Rosacletodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FF09FD4DFEE14CCE" box="[151,313,674,699]" italics="true" pageId="35" pageNumber="36">Rosacletodes</emphasis>
</taxonomicName>
has an identical segmentation and armature pattern as that of the prehensile ramus in both 
<taxonomicName id="4C774D12FFF5FFB1FF09FD27FEBC4C94" box="[151,356,712,737]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FF09FD27FEBC4C94" box="[151,356,712,737]" italics="true" pageId="35" pageNumber="36">Laophontisochra</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF5FFB1FE3EFD27FDC04C94" box="[416,536,712,737]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FE3EFD27FDC04C94" box="[416,536,712,737]" italics="true" pageId="35" pageNumber="36">Acuticoxa</emphasis>
</taxonomicName>
, the only difference being the much shorter proximal endopod segment in 
<taxonomicName id="4C774D12FFF5FFB1FF09FD02FEEA4D73" box="[151,306,749,774]" class="Maxillopoda" family="Nannopodidae" genus="Rosacletodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FF09FD02FEEA4D73" box="[151,306,749,774]" italics="true" pageId="35" pageNumber="36">Rosacletodes</emphasis>
</taxonomicName>
. Prehensility of the P1 endopod is here considered a synapomorphy (in conjunction with the 4- segmented Ƥ antennule, reduced antennary exopod, geniculate endopodal claw on the maxilliped, etc.) supporting the sistergroup relationship between 
<taxonomicName id="4C774D12FFF5FFB1FD56FCD7FC484D24" box="[712,912,824,849]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FD56FCD7FC484D24" box="[712,912,824,849]" italics="true" pageId="35" pageNumber="36">Laophontisochra</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF5FFB1FC57FCD7FBE64D24" box="[969,1086,824,849]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FC57FCD7FBE64D24" box="[969,1086,824,849]" italics="true" pageId="35" pageNumber="36">Acuticoxa</emphasis>
</taxonomicName>
within a larger encompassing clade characterized by coxal projections on the swimming legs. This clade further includes 
<taxonomicName id="4C774D12FFF5FFB1FB6BFCB2FA4D4D03" box="[1269,1429,861,886]" class="Maxillopoda" family="Nannopodidae" genus="Rosacletodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FB6BFCB2FA4D4D03" box="[1269,1429,861,886]" italics="true" pageId="35" pageNumber="36">Rosacletodes</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C774D12FFF5FFB1FF09FC6DFEE04DEE" box="[151,312,898,923]" class="Maxillopoda" family="Huntemanniidae" genus="Huntemannia" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FF09FC6DFEE04DEE" box="[151,312,898,923]" italics="true" pageId="35" pageNumber="36">Huntemannia</emphasis>
</taxonomicName>
and “Genus X”, all of which show varying degrees of coxal modification in leg 1, being most extreme in the latter genus (P.H.C. Corgosinho &amp; M. Büntzow, pers. commn). Based on the arguments presented above, 
<taxonomicName id="4C774D12FFF5FFB1FEFDFC22FDF34D93" box="[355,555,973,998]" class="Maxillopoda" family="Nannopodidae" genus="Laophontisochra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FEFDFC22FDF34D93" box="[355,555,973,998]" italics="true" pageId="35" pageNumber="36">Laophontisochra</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF5FFB1FDFAFC22FD014D93" box="[612,729,973,998]" class="Maxillopoda" family="Nannopodidae" genus="Acuticoxa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FDFAFC22FD014D93" box="[612,729,973,998]" italics="true" pageId="35" pageNumber="36">Acuticoxa</emphasis>
</taxonomicName>
are here formally assigned to the 
<taxonomicName id="4C774D12FFF5FFB1FBF9FC22FACE4D92" box="[1127,1302,973,999]" class="Maxillopoda" family="Nannopodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="family">Nannopodidae</taxonomicName>
.
</paragraph>
<paragraph id="8BC83691FFF5FFB1FF5BFC1DFD3E4B42" blockId="35.[151,1437,152,1335]" pageId="35" pageNumber="36">
The family 
<taxonomicName id="4C774D12FFF5FFB1FED1FC1DFE224A79" box="[335,506,1010,1036]" class="Maxillopoda" family="Nannopodidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="family">Nannopodidae</taxonomicName>
is heterogeneous at present; in particular, both 
<taxonomicName id="4C774D12FFF5FFB1FBB5FC1DFAD84A7E" box="[1067,1280,1010,1035]" class="Maxillopoda" family="Huntemanniidae" genus="Metahuntemannia" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FBB5FC1DFAD84A7E" box="[1067,1280,1010,1035]" italics="true" pageId="35" pageNumber="36">Metahuntemannia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF5FFB1FAA4FC1DFA434A7E" box="[1338,1435,1010,1035]" class="Maxillopoda" family="Canthocamptidae" genus="Pottekia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FAA4FC1DFA434A7E" box="[1338,1435,1010,1035]" italics="true" pageId="35" pageNumber="36">Pottekia</emphasis>
</taxonomicName>
(= 
<taxonomicName id="4C774D12FFF5FFB1FF24FBF7FECC4A44" box="[186,276,1048,1073]" class="Maxillopoda" family="Canthocamptidae" genus="Talpina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FF24FBF7FECC4A44" box="[186,276,1048,1073]" italics="true" pageId="35" pageNumber="36">Talpina</emphasis>
</taxonomicName>
) are radically divergent from the other nannopodid genera. The sexual dimorphism expressed on the P4 endopod (distal inner seta of Ƥ modified into a serrate curved spine in 3; 
<emphasis id="B903EA83FFF5FFB1FBDEFBD2FAE14A23" box="[1088,1337,1085,1110]" italics="true" pageId="35" pageNumber="36">
cf. 
<taxonomicName id="4C774D12FFF5FFB1FBFDFBD2FAE14A23" authority="Dahms &amp; Pottek, 1992" authorityName="Dahms &amp; Pottek" authorityYear="1992" box="[1123,1337,1085,1110]" class="Maxillopoda" family="Canthocamptidae" genus="Pottekia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pectinata">Pottekia pectinata</taxonomicName>
</emphasis>
<bibRefCitation id="EFE64B60FFF5FFB1FAA1FBD2FEB24A09" author="Dahms" pageId="35" pageNumber="36" refString="Dahms, H. - U. &amp; Pottek, M. (1992) Metahuntemannia Smirnov, 1946 and Talpina gen. nov. (Copepoda, Harpacticoida) from the deep-sea of the high Antarctic Weddell Sea with a description of eight new species. Microfauna Marina, 7, 7 - 78." type="journal article" year="1992">(Dahms &amp; Pottek, 1992))</bibRefCitation>
clearly indicates an affinity with genera such as 
<taxonomicName id="4C774D12FFF5FFB1FC51FB8CFA404A09" authority="Huys &amp; Thistle, 1989" authorityName="Huys &amp; Thistle" authorityYear="1989" box="[975,1432,1122,1148]" class="Maxillopoda" family="Canthocamptidae" genus="Bathycamptus" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FC51FB8CFBA54A09" box="[975,1149,1123,1148]" italics="true" pageId="35" pageNumber="36">Bathycamptus</emphasis>
<bibRefCitation id="EFE64B60FFF5FFB1FB1BFB8DFA404A09" author="Huys" box="[1157,1432,1122,1148]" pageId="35" pageNumber="36" refString="Huys, R. &amp; Thistle, D. (1989) Bathycamptus eckmani gen. et spec. nov. (Copepoda, Harpacticoida) with a review of the taxonomic status of certain other deepwater harpacticoids. Hydrobiologia, 185, 101 - 126." type="journal article" year="1989">Huys &amp; Thistle, 1989</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="4C774D12FFF5FFB1FF09FB67FE294AD7" authority="Mielke, 1975" authorityName="Mielke" authorityYear="1975" box="[151,497,1160,1186]" class="Maxillopoda" family="Pseudotachidiidae" genus="Micropsammis" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FF09FB67FE9F4AD4" box="[151,327,1160,1185]" italics="true" pageId="35" pageNumber="36">Micropsammis</emphasis>
<bibRefCitation id="EFE64B60FFF5FFB1FED0FB67FE294AD7" author="Mielke" box="[334,497,1160,1186]" pageId="35" pageNumber="36" refString="Mielke, W. (1975) Systematik der Copepoda eines Sandstrandes der Nordseeinsel Sylt. Mikrofauna des Meeresbodens, 52, 1 - 134." type="journal article" year="1975">Mielke, 1975</bibRefCitation>
</taxonomicName>
; and 
<taxonomicName id="4C774D12FFF5FFB1FDAFFB67FBD74AD7" authority="George &amp; Schminke, 2003" authorityName="George &amp; Schminke" authorityYear="2003" box="[561,1039,1160,1186]" class="Maxillopoda" family="Canthocamptidae" genus="Isthmiocaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FDAFFB67FD1E4AD4" box="[561,710,1160,1185]" italics="true" pageId="35" pageNumber="36">Isthmiocaris</emphasis>
<bibRefCitation id="EFE64B60FFF5FFB1FD50FB67FBD74AD7" author="George" box="[718,1039,1160,1186]" pageId="35" pageNumber="36" refString="George, K. H. &amp; Schminke, H. K. (2003) Isthmiocaris longitelson gen. et sp. nov., a strongly derived harpacticoid (Copepoda) from the Magellan region, and its systematic affinities to certain &quot; canthocamptid &quot; taxa. Journal of crustacean Biology, 23, 119 - 130." type="journal article" year="2003">George &amp; Schminke, 2003</bibRefCitation>
</taxonomicName>
(and almost certainly the closely related 
<taxonomicName id="4C774D12FFF5FFB1FF70FB42FD494AB2" authority="Huys &amp; Thistle, 1989" authorityName="Huys &amp; Thistle" authorityYear="1989" box="[238,657,1197,1223]" class="Maxillopoda" family="Canthocamptidae" genus="Perucamptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FF70FB42FE514AB3" box="[238,393,1197,1222]" italics="true" pageId="35" pageNumber="36">Perucamptus</emphasis>
<bibRefCitation id="EFE64B60FFF5FFB1FE0FFB42FD494AB2" author="Huys" box="[401,657,1197,1223]" pageId="35" pageNumber="36" refString="Huys, R. &amp; Thistle, D. (1989) Bathycamptus eckmani gen. et spec. nov. (Copepoda, Harpacticoida) with a review of the taxonomic status of certain other deepwater harpacticoids. Hydrobiologia, 185, 101 - 126." type="journal article" year="1989">Huys &amp; Thistle, 1989</bibRefCitation>
</taxonomicName>
– male unknown at present!) (
<bibRefCitation id="EFE64B60FFF5FFB1FC67FB42FAEB4AB2" author="George" box="[1017,1331,1197,1223]" pageId="35" pageNumber="36" refString="George, K. H. &amp; Schminke, H. K. (2003) Isthmiocaris longitelson gen. et sp. nov., a strongly derived harpacticoid (Copepoda) from the Magellan region, and its systematic affinities to certain &quot; canthocamptid &quot; taxa. Journal of crustacean Biology, 23, 119 - 130." type="journal article" year="2003">George &amp; Schminke 2003</bibRefCitation>
; 
<bibRefCitation id="EFE64B60FFF5FFB1FAA1FB42FEEB4A99" author="Huys" pageId="35" pageNumber="36" refString="Huys, R. &amp; Thistle, D. (1989) Bathycamptus eckmani gen. et spec. nov. (Copepoda, Harpacticoida) with a review of the taxonomic status of certain other deepwater harpacticoids. Hydrobiologia, 185, 101 - 126." type="journal article" year="1989">Huys &amp; Thistle 1989</bibRefCitation>
; 
<bibRefCitation id="EFE64B60FFF5FFB1FEDFFB3DFE074A99" author="Mielke" box="[321,479,1234,1260]" pageId="35" pageNumber="36" refString="Mielke, W. (1975) Systematik der Copepoda eines Sandstrandes der Nordseeinsel Sylt. Mikrofauna des Meeresbodens, 52, 1 - 134." type="journal article" year="1975">Mielke 1975</bibRefCitation>
), all of which are currently assigned to the subfamily 
<taxonomicName id="4C774D12FFF5FFB1FBEDFB3DFA964A99" box="[1139,1358,1234,1260]" class="Maxillopoda" family="Canthocamptidae" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="subFamily" subFamily="Hemimesochrinae">Hemimesochrinae</taxonomicName>
in the 
<taxonomicName id="4C774D12FFF5FFB1FF09FB17FDC94B67" authority="Wells 2007" authorityName="Wells" authorityYear="2007" box="[151,529,1272,1298]" class="Maxillopoda" family="Canthocamptidae" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="family">
Canthocamptidae (
<bibRefCitation id="EFE64B60FFF5FFB1FEE7FB17FDD04B67" author="Wells" box="[377,520,1272,1298]" pageId="35" pageNumber="36" refString="Wells, J. B. J. (2007) An annotated checklist and keys to the species of Copepoda Harpacticoida (Crustacea). Zootaxa, 1568, 1 - 872." type="journal article" year="2007">Wells 2007</bibRefCitation>
)
</taxonomicName>
. Pending a revision of the latter family, 
<taxonomicName id="4C774D12FFF5FFB1FC68FB17FB084B64" box="[1014,1232,1272,1297]" class="Maxillopoda" family="Huntemanniidae" genus="Metahuntemannia" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FC68FB17FB084B64" box="[1014,1232,1272,1297]" italics="true" pageId="35" pageNumber="36">Metahuntemannia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C774D12FFF5FFB1FA95FB17FAB74B64" box="[1291,1391,1272,1297]" class="Maxillopoda" family="Canthocamptidae" genus="Pottekia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B903EA83FFF5FFB1FA95FB17FAB74B64" box="[1291,1391,1272,1297]" italics="true" pageId="35" pageNumber="36">Pottekia</emphasis>
</taxonomicName>
are here tentatively assigned to the 
<taxonomicName id="4C774D12FFF5FFB1FD97FAF2FD3A4B42" box="[521,738,1309,1335]" class="Maxillopoda" family="Canthocamptidae" kingdom="Animalia" order="Harpacticoida" pageId="35" pageNumber="36" phylum="Arthropoda" rank="subFamily" subFamily="Hemimesochrinae">Hemimesochrinae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>