treatments-xml/data/03/DB/55/03DB55320206FFF7EDF87E10FF6B59AC.xml

<document id="256B66688CBA525F233618B3FE4659AB" ID-DOI="10.11646/zootaxa.4088.3.6" ID-GBIF-Dataset="715978f0-8f48-4e85-8865-c3ba9ddae313" ID-ISSN="1175-5326" ID-Zenodo-Dep="266698" ID-ZooBank="B87BB845-FB3D-4536-8465-9DAB4B62AC53" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1457603093683" checkinUser="plazi" docAuthor="Pauwels, Olivier S. G., Sumontha, Montri &amp; Bauer, Aaron M." docDate="2016" docId="03DB55320206FFF7EDF87E10FF6B59AC" docLanguage="en" docName="15580-5108-1-PB.pdf.imf" docOrigin="Zootaxa 4088 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyrtodactylus phetchaburiensis Pauwels, Sumontha &amp; Bauer, 2016, sp. nov." docType="treatment" docVersion="10" lastPageNumber="415" masterDocId="FFE22D4A0207FFF1ED6F7A47FF985C31" masterDocTitle="A new Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Phetchaburi Province, Thailand" masterLastPageNumber="419" masterPageNumber="409" pageNumber="410" updateTime="1698177604698" updateUser="plazi">
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<mods:title id="EA81F35A6FAFC9E84B0F0CD5ED895852">A new Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Phetchaburi Province, Thailand</mods:title>
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<mods:namePart id="F50FBC47B4642AD5A75E4FCF6E1E1684">Pauwels, Olivier S. G.</mods:namePart>
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<mods:namePart id="00C269C049849755609F296A004C0E4F">Sumontha, Montri</mods:namePart>
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<treatment id="03DB55320206FFF7EDF87E10FF6B59AC" ID-DOI="http://doi.org/10.5281/zenodo.6085358" ID-GBIF-Taxon="117198867" ID-Zenodo-Dep="6085358" LSID="urn:lsid:plazi:treatment:03DB55320206FFF7EDF87E10FF6B59AC" httpUri="http://treatment.plazi.org/id/03DB55320206FFF7EDF87E10FF6B59AC" lastPageId="6" lastPageNumber="415" pageId="1" pageNumber="410">
<subSubSection id="C368B7AF0206FFF0EDF87E10FE9858A2" pageId="1" pageNumber="410" type="nomenclature">
<paragraph id="8BCDE4240206FFF0EDF87E10FDE05843" blockId="1.[151,632,1111,1171]" box="[151,632,1111,1138]" pageId="1" pageNumber="410">
<heading id="D08553480206FFF0EDF87E10FDE05843" bold="true" box="[151,632,1111,1138]" fontSize="11" level="1" pageId="1" pageNumber="410" reason="1">
<emphasis id="B90638360206FFF0EDF87E10FDE05843" bold="true" box="[151,632,1111,1138]" pageId="1" pageNumber="410">
<taxonomicName id="4C729FA70206FFF0EDF87E10FD8A5840" box="[151,530,1111,1137]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="410" phylum="Chordata" rank="species" species="phetchaburiensis" status="sp. nov.">
<emphasis id="B90638360206FFF0EDF87E10FD8A5840" bold="true" box="[151,530,1111,1137]" italics="true" pageId="1" pageNumber="410">Cyrtodactylus phetchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235854D0206FFF0EF757E1FFDE05843" box="[538,632,1112,1138]" pageId="1" pageNumber="410" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCDE4240206FFF0EDF87E3DFE9858A2" blockId="1.[151,632,1111,1171]" box="[151,256,1146,1171]" pageId="1" pageNumber="410">
<figureCitation id="1349F8A10206FFF0EDF87E3DFE9858A2" box="[151,256,1146,1171]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1001,1022]" captionStartId-1="2.[151,250,1543,1564]" captionStartId-2="4.[151,250,1035,1056]" captionTargetBox-0="[151,1436,193,979]" captionTargetBox-1="[150,1436,1072,1521]" captionTargetBox-2="[151,1432,155,1005]" captionTargetId-0="figure@2.[151,1436,193,979]" captionTargetId-1="figure@2.[150,1436,1072,1521]" captionTargetId-2="figure@4.[151,1436,155,1013]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="4" captionText-0="FIGURE 1. Adult male holotype of Cyrtodactylus phetchaburiensis sp. nov. (IRSNB 2682). Scale bar = 10 mm." captionText-1="FIGURE 2. Adult female paratype of Cyrtodactylus phetchaburiensis sp. nov. (IRSNB 2683). Scale bar = 10 mm." captionText-2="FIGURE 3. Specimens tentatively referable to Cyrtodactylus phetchaburiensis sp. nov. but showing variation in dorsal colour pattern (see Discussion)." httpUri-0="https://zenodo.org/record/266699/files/figure.png" httpUri-1="https://zenodo.org/record/266700/files/figure.png" httpUri-2="https://zenodo.org/record/266701/files/figure.png" pageId="1" pageNumber="410">Figs. 1–3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C368B7AF0206FFF0EDF87E84FBB65A56" pageId="1" pageNumber="410" type="materials_examined">
<paragraph id="8BCDE4240206FFF0EDF87E84FD085912" blockId="1.[151,1437,1218,2035]" pageId="1" pageNumber="410">
<emphasis id="B90638360206FFF0EDF87E84FE9358EB" bold="true" box="[151,267,1219,1242]" pageId="1" pageNumber="410">Holotype.</emphasis>
IRSNB 2682 (formerly IRSNB 16682), adult male (
<figureCitation id="1349F8A10206FFF0EE0F7E85FC3958EA" box="[864,929,1218,1243]" captionStart="FIGURE 1" captionStartId="2.[151,250,1001,1022]" captionTargetBox="[151,1436,193,979]" captionTargetId="figure@2.[151,1436,193,979]" captionTargetPageId="2" captionText="FIGURE 1. Adult male holotype of Cyrtodactylus phetchaburiensis sp. nov. (IRSNB 2682). Scale bar = 10 mm." httpUri="https://zenodo.org/record/266699/files/figure.png" pageId="1" pageNumber="410">Fig. 1</figureCitation>
); Thailand, Phetchaburi Province, Tha Yang District, Klatluang Subdistrict, Khao (= Mountain) Tomo, Tham (= Cave) Khao Tomo (12º47.98’N, 99º44.36’E), collected by local villagers, September 2003.
</paragraph>
<paragraph id="8BCDE4240206FFF0EDA87F68FDEA595A" blockId="1.[151,1437,1218,2035]" pageId="1" pageNumber="410">
<emphasis id="B90638360206FFF0EDA87F68FEA35977" bold="true" box="[199,315,1327,1350]" pageId="1" pageNumber="410">Paratype.</emphasis>
IRSNB 2683 (formerly IRSNB 16650), adult female (
<figureCitation id="1349F8A10206FFF0EECC7F69FC705976" box="[931,1000,1326,1351]" captionStart="FIGURE 2" captionStartId="2.[151,250,1543,1564]" captionTargetBox="[150,1436,1072,1521]" captionTargetId="figure@2.[150,1436,1072,1521]" captionTargetPageId="2" captionText="FIGURE 2. Adult female paratype of Cyrtodactylus phetchaburiensis sp. nov. (IRSNB 2683). Scale bar = 10 mm." httpUri="https://zenodo.org/record/266700/files/figure.png" pageId="1" pageNumber="410">Fig. 2</figureCitation>
); Thailand, Phetchaburi Province, Tha Yang District, collector unknown, no date.
</paragraph>
<paragraph id="8BCDE4240206FFF0EDA87F31FBB65A56" blockId="1.[151,1437,1218,2035]" pageId="1" pageNumber="410">
<emphasis id="B90638360206FFF0EDA87F31FE5659BE" bold="true" box="[199,462,1398,1423]" pageId="1" pageNumber="410">Additional specimens.</emphasis>
Other Phetchaburi material initially referred to 
<taxonomicName id="4C729FA70206FFF0EE877F30FBFD59BF" box="[1000,1125,1398,1423]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="410" phylum="Chordata" rank="species" species="oldhami">
<emphasis id="B90638360206FFF0EE877F30FBFD59BF" box="[1000,1125,1398,1423]" italics="true" pageId="1" pageNumber="410">C. oldhami</emphasis>
</taxonomicName>
is here tentatively assigned to 
<taxonomicName id="4C729FA70206FFF0EDD77FDDFD875983" box="[184,543,1434,1458]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="410" phylum="Chordata" rank="species" species="phetchaburiensis" status="sp. nov.">
<emphasis id="B90638360206FFF0EDD77FDDFD875983" box="[184,543,1434,1458]" italics="true" pageId="1" pageNumber="410">Cyrtodactylus phetchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235854D0206FFF0EF457FDCFD1E5983" box="[554,646,1435,1458]" pageId="1" pageNumber="410" rank="species">
<emphasis id="B90638360206FFF0EF457FDCFD195983" bold="true" box="[554,641,1435,1458]" pageId="1" pageNumber="410">sp. nov</emphasis>
.
</taxonomicNameLabel>
, but with some reservation (see 
<emphasis id="B90638360206FFF0E9647FDDFB115982" bold="true" box="[1035,1161,1434,1459]" pageId="1" pageNumber="410">Discussion</emphasis>
): THNHM 1247–1248, Thailand, Phetchaburi Province, Tha Yang District, Kaeng Krachan National Park, along Pala-U Waterfall; THNHM 1256–1257, 1283–1284, Thailand, Phetchaburi Province, Tha Yang District, Kaeng Krachan National Park, Khao Phanoen Thung; THNHM 20245–20249 (formerly TNRC 52-3868–3872) (
<figureCitation id="1349F8A10206FFF0E9AB7C41FA885A2E" box="[1220,1296,1542,1567]" captionStart="FIGURE 3" captionStartId="4.[151,250,1035,1056]" captionTargetBox="[151,1432,155,1005]" captionTargetId="figure@4.[151,1436,155,1013]" captionTargetPageId="4" captionText="FIGURE 3. Specimens tentatively referable to Cyrtodactylus phetchaburiensis sp. nov. but showing variation in dorsal colour pattern (see Discussion)." httpUri="https://zenodo.org/record/266701/files/figure.png" pageId="1" pageNumber="410">Fig. 3</figureCitation>
), Thailand, Phetchaburi Province, Tha Yang District, Kaeng Krachan National Park, near km 23 on the road from Thung Peak (12°47’N, 99°23’E, 900 m elevation), collected by J. Nabhitabhata, 22 June 1990.
</paragraph>
</subSubSection>
<subSubSection id="C368B7AF0206FFF0EDA87C35FB6D5BC2" pageId="1" pageNumber="410" type="diagnosis">
<paragraph id="8BCDE4240206FFF0EDA87C35FB6D5BC2" blockId="1.[151,1437,1218,2035]" pageId="1" pageNumber="410">
<emphasis id="B90638360206FFF0EDA87C35FEA25ABA" bold="true" box="[199,314,1650,1675]" pageId="1" pageNumber="410">Diagnosis</emphasis>
. A medium-sized 
<taxonomicName id="4C729FA70206FFF0EF687C35FD3C5ABB" box="[519,676,1650,1674]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="410" phylum="Chordata" rank="genus">
<emphasis id="B90638360206FFF0EF687C35FD3C5ABB" box="[519,676,1650,1674]" italics="true" pageId="1" pageNumber="410">Cyrtodactylus</emphasis>
</taxonomicName>
(SVL to at least 63.2 mm); body slender; limbs and digits relatively long, slender; original tail longer than SVL; enlarged pair of first postmental scales in contact with one another behind mental, a smaller pair of enlarged chin shields (second postmentals) lateral to these; small, mostly keeled tubercles in 20 regular longitudinal rows on dorsum; 33 scales across mid-venter between lowest rows of flank tubercles; ventrolateral folds weakly developed and atuberculate; enlarged row of femoral scales present; five precloacal pores in male, femoral pores and precloacal groove absent; 5–6 broad basal lamellae and 11 narrow distal lamellae beneath digit IV of pes; most of post-pygal portion of tail atuberculate, a single median row of transversely enlarged subcaudal scales present. Colour pattern of large, dark markings with diffuse edges on a fawn background extending from shoulders to sacrum. A pair of dark scapular patches and a broad occipital band outlined by thin white borders. Flanks pale brown with scattered lighter blotches. Tail distinctly banded with thicker brown annuli, becoming darker posteriorly, alternating with narrower white annuli or bands.
</paragraph>
</subSubSection>
<caption id="DF0DB4AC0205FFF3EDF879AEFA8A5FCE" box="[151,1298,1001,1023]" httpUri="https://zenodo.org/record/266699/files/figure.png" pageId="2" pageNumber="411" targetBox="[151,1436,193,979]" targetPageId="2">
<paragraph id="8BCDE4240205FFF3EDF879AEFA8A5FCE" blockId="2.[151,1298,1001,1023]" box="[151,1298,1001,1023]" pageId="2" pageNumber="411">
<emphasis id="B90638360205FFF3EDF879AEFE8B5FCF" bold="true" box="[151,275,1001,1022]" pageId="2" pageNumber="411">FIGURE 1.</emphasis>
Adult male holotype of 
<taxonomicName id="4C729FA70205FFF3EF6479AEFCD45FCE" box="[523,844,1001,1023]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="411" phylum="Chordata" rank="species" species="phetchaburiensis" status="sp. nov.">
<emphasis id="B90638360205FFF3EF6479AEFCD45FCE" box="[523,844,1001,1023]" italics="true" pageId="2" pageNumber="411">Cyrtodactylus phetchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235854D0205FFF3EE3D79AEFC3B5FCF" box="[850,931,1001,1022]" pageId="2" pageNumber="411" rank="species">
<emphasis id="B90638360205FFF3EE3D79AEFC3B5FCF" bold="true" box="[850,931,1001,1022]" pageId="2" pageNumber="411">sp. nov.</emphasis>
</taxonomicNameLabel>
(IRSNB 2682). Scale bar = 10 mm.
</paragraph>
</caption>
<caption id="DF0DB4AC0205FFF3EDF87C40FABC5A2C" box="[151,1316,1543,1565]" httpUri="https://zenodo.org/record/266700/files/figure.png" pageId="2" pageNumber="411" targetBox="[150,1436,1072,1521]" targetPageId="2">
<paragraph id="8BCDE4240205FFF3EDF87C40FABC5A2C" blockId="2.[151,1316,1543,1565]" box="[151,1316,1543,1565]" pageId="2" pageNumber="411">
<emphasis id="B90638360205FFF3EDF87C40FE8A5A2D" bold="true" box="[151,274,1543,1564]" pageId="2" pageNumber="411">FIGURE 2.</emphasis>
Adult female paratype of 
<taxonomicName id="4C729FA70205FFF3EF727C40FCC65A2C" box="[541,862,1543,1565]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="411" phylum="Chordata" rank="species" species="phetchaburiensis" status="sp. nov.">
<emphasis id="B90638360205FFF3EF727C40FCC65A2C" box="[541,862,1543,1565]" italics="true" pageId="2" pageNumber="411">Cyrtodactylus phetchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235854D0205FFF3EE0B7C4FFC2D5A2C" box="[868,949,1544,1565]" pageId="2" pageNumber="411" rank="species">
<emphasis id="B90638360205FFF3EE0B7C4FFC2D5A2C" bold="true" box="[868,949,1544,1565]" pageId="2" pageNumber="411">sp. nov.</emphasis>
</taxonomicNameLabel>
(IRSNB 2683). Scale bar = 10 mm.
</paragraph>
</caption>
<subSubSection id="C368B7AF0205FFF5EDA87C03FB135B1D" lastPageId="4" lastPageNumber="413" pageId="2" pageNumber="411" type="description">
<paragraph id="8BCDE4240205FFF2EDA87C03FD3E5DFE" blockId="2.[151,1437,1604,2025]" lastBlockId="3.[151,1437,151,1869]" lastPageId="3" lastPageNumber="412" pageId="2" pageNumber="411">
<emphasis id="B90638360205FFF3EDA87C03FE7E5A6D" bold="true" box="[199,486,1604,1629]" pageId="2" pageNumber="411">Description of holotype.</emphasis>
Adult male. SVL 57.5 mm; TailL 70.8 mm (original). Head long (HeadL/SVL ratio 0.31), moderately narrow (HeadW/HeadL = 0.60), not markedly depressed (HeadH/HeadL = 0.40), clearly distinct from neck. Loreal region weakly inflated, canthus rostralis not prominent. Snout moderate in length (SnEye/HeadL = 0.38), somewhat acuminate; longer than eye diameter (OrbD/SnEye = 0.65); scales on snout small, oval to oblong, weakly conical with apex directed posteriorly, somewhat heterogeneous, distinctly larger than those on crown, interorbital and occipital regions. Eye moderately large (OrbD/HeadL = 0.25); pupil vertical with crenelated margins; supraciliaries short, minute spines on posterior 60% of scales; anterior supraciliaries largest and bearing a central keel that becomes less pronounced posteriorly. Ear opening obliquely elliptical, demarcated posteroventrally by a small fold of skin, large (EarL/HeadL = 0.11); eye to ear distance slightly greater than diameter of eyes (EyeEar/OrbD = 1.10). Rostral much wider (3.0 mm) than deep (1.9 mm), rostral crease about one half height of rostral. Two enlarged, rhomboidal supranasals separated by a single, small, pentagonal internasal. Rostral in contact with first supralabials, nostrils, internasal and supranasals. Nostrils oval, slightly posterolaterally-directed, each surrounded by supranasal, rostral, first supralabial, two distinct postnasals, and posteroventrally by a crescentic nasal, occluded posteromedially by a narial valve. Two to three rows of small scales separate orbit from supralabials. Mental triangular with acute posteriorly-oriented apex separating anterior 40% of postmentals, wider (2.4 mm) than deep (1.8 mm). A pair of enlarged postmentals in contact behind mental, each bordered anteromedially by mental, anterolaterally by first infralabial, posterolaterally by an enlarged lateral chinshield (= second postmental) approximately 40% size of first postmental, and posteriorly by three slightly enlarged granules. Supralabials to midorbital position 8 (both sides); enlarged supralabials to angle of jaws 12 (right) to 13 (left). Infralabials 11 (both sides). Interorbital scale rows across narrowest point of frontal bone 15; 47 scales between left and right supraciliary rows.
</paragraph>
<paragraph id="8BCDE4240204FFF2EDA87B9BFF795F6C" blockId="3.[151,1437,151,1869]" pageId="3" pageNumber="412">Body slender, elongate (TrunkL/SVL = 0.41) with faintly demarcated ventrolateral folds lacking serrations, tubercles, or spines. Dorsal scales exclusive of tubercles largely homogeneous, conical with slightly posteriorlyoriented apices; regularly distributed tubercles (5–8 times size of adjacent scales), most circular, some partly flattened posteriorly, extending from crown and dorsal postorbital region to tail base, on trunk smallest on flanks, largest in paravertebral position and over sacrum, smaller tubercles on postorbital region, crown, occiput, and nape; tubercles conical and erect anteriorly, becoming slightly recumbent and strongly keeled posterior to shoulders, in 20 rows at midbody, typically separated from one another by 1–3 dorsal granules. Ventral scales much larger than dorsals, smooth, oval subimbricate to imbricate, largest on posterior abdomen and especially in precloacal region where they are approximately 3–4 times the size of those at midbody and assume a subtriangular shape. Midbody scale rows across belly to ventrolateral folds ~33. Gular region with homogeneous, smooth, juxtaposed granular scales.</paragraph>
<paragraph id="8BCDE4240204FFF2EDA87920FDBF5964" blockId="3.[151,1437,151,1869]" pageId="3" pageNumber="412">Three (left) and two (right) precloacal pores separated by a single poreless scale. No femoral pores. No precloacal groove. Anteroventral scales of thigh much larger than posteroventral and posterior, with a distinct single row of enlarged femoral scales. Two slightly enlarged, smooth postcloacal spurs, anterior larger than posterior. Hemipeneal bulges evident. Scales on palm and sole smooth, rounded to oval, flattened to slightly domed. Scalation on dorsal surfaces of hind limbs and distal fore limb similar to body dorsum with enlarged, conical, partly keeled tubercles interspersed among smaller scales, tubercles separated from one another by 1–2 small scales; proximal portion of forearm lacking tubercles. Fore and hindlimbs relatively long, slender (ForeaL/ SVL = 0.15; CrusL/SVL = 0.19). Digits relatively short, slender, inflected at interphalangeal joints, especially antepenultimate joint of longer digits, all bearing robust, slightly recurved claws. Basal subdigital lamellae broad, squarish to rectangular with rounded corners, distalmost generally largest, without scansorial surfaces (3-4-4-4-4) both manus, 3-3-3-6-4 left pes, 3-4-4-5-4 right pes); narrow lamellae distal to digital inflection and not including ventral claw sheath: 7-8-10-10-8 (left manus), 7-8-8-9-8 (right manus), 8-9-11-11-11 (left pes), 7-9-11-11-11 (right pes); basalmost usually fragmented; no interdigital webbing. Relative length of digits: IV&gt; III&gt; II ~ V&gt; I (manus); IV ~ V ~ III&gt; II&gt; I (pes).</paragraph>
<paragraph id="8BCDE4240204FFF2EDA87F18FAD95949" blockId="3.[151,1437,151,1869]" box="[199,1345,1375,1400]" pageId="3" pageNumber="412">Original tail, long, subcylindrical, gently tapering to pointed tip; longer than SVL (TailL/SVL = 1.23).</paragraph>
<paragraph id="8BCDE4240204FFF2EDA87FC3FD065A45" blockId="3.[151,1437,151,1869]" pageId="3" pageNumber="412">Scales of tail dorsum regular, flat, juxtaposed to weakly imbricate; squarish basally, becoming more rectangular posteriorly; those on basal segments weakly keeled and with slightly pointed free margins, arranged in visible but indistinct segments of 7 scale rows basally. Basalmost postpygal segments with a transverse row 6 enlarged, flattened, keeled tubercles, each 3–4 times size of adjacent scales, decreasing to 4 tubercles in next segment and to 2 in the fifth and sixth postpygal segments, becoming smaller and more weakly keeled posteriorly and lost in the seventh and more posterior segments. Subcaudal scales much enlarged, medial row forming a single series of enlarged plates, two per tail segment.</paragraph>
<paragraph id="8BCDE4240204FFF2EDA87CC6FB765AD1" blockId="3.[151,1437,151,1869]" pageId="3" pageNumber="412">Measurements (in mm; holotype followed by paratype): SVL 57.5/63.2, ForeaL 8.8/9.2, CrusL 11.1/11.9, TailL 70.8/72.6, TailW 5.4/5.4, TrunkL 23.8/27.8, HeadL 17.8/17.6, HeadW 10.7/11.3, HeadH 7.1/7.5, OrbD 4.4/ 4.2, EyeEar 4.8/4.9, SnEye 6.7/6.9, NarEye 4.6/5.2, Interorb 2.4/2.3, EarL 1.9/1.7, Internar 2.1/2.1.</paragraph>
<paragraph id="8BCDE4240204FFF2EDA87CABFE2F5B7C" blockId="3.[151,1437,151,1869]" pageId="3" pageNumber="412">Coloration in ethanol: Dorsum pale fawn with a series of large, darker, diffusely-edged markings from shoulders to sacrum. A pair of dark scapular patches outlined by a thin white border. Flanks pale brown with scattered lighter blotches.</paragraph>
<caption id="DF0DB4AC0203FFF5EDF87E4CFE095871" httpUri="https://zenodo.org/record/266701/files/figure.png" pageId="4" pageNumber="413" targetBox="[151,1432,155,1005]" targetPageId="4">
<paragraph id="8BCDE4240203FFF5EDF87E4CFE095871" blockId="4.[151,1436,1035,1088]" pageId="4" pageNumber="413">
<emphasis id="B90638360203FFF5EDF87E4CFE8A5811" bold="true" box="[151,274,1035,1056]" pageId="4" pageNumber="413">FIGURE 3.</emphasis>
Specimens tentatively referable to 
<taxonomicName id="4C729FA70203FFF5EF167E4CFC235810" box="[633,955,1035,1057]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="413" phylum="Chordata" rank="species" species="phetchaburiensis" status="sp. nov.">
<emphasis id="B90638360203FFF5EF167E4CFC235810" box="[633,955,1035,1057]" italics="true" pageId="4" pageNumber="413">Cyrtodactylus phetchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235854D0203FFF5EEAE7E4BFB8A5810" box="[961,1042,1036,1057]" pageId="4" pageNumber="413" rank="species">
<emphasis id="B90638360203FFF5EEAE7E4BFB8A5810" bold="true" box="[961,1042,1036,1057]" pageId="4" pageNumber="413">sp. nov.</emphasis>
</taxonomicNameLabel>
but showing variation in dorsal colour pattern (see Discussion).
</paragraph>
</caption>
<paragraph id="8BCDE4240203FFF5EDA87E20FD6458DD" blockId="4.[151,1437,1127,2016]" pageId="4" pageNumber="413">A well demarcated medium brown band with diffuse white margins extending between postorbital regions of left and right sides across the occiput, band posteriorly bifid across mid-occiput. Crown and snout mid-brown with some lighter and darker irregular patches. Area below occipital band pale cream, posterior supralabials whitish, becoming cream with diffuse darker centres anteriorly.</paragraph>
<paragraph id="8BCDE4240203FFF5EDA87EB0FC9F5991" blockId="4.[151,1437,1127,2016]" pageId="4" pageNumber="413">Limbs pale brown, without distinct markings, feet darker than limbs. Venter beige with all scales bearing tiny brown to black flecks, enlarged precloacal scales paler than remainder of venter. Tail distinctly banded with thicker brown annuli, becoming darker posteriorly, alternating with narrower white annuli; 13 white bands in total, basalmost merely dorsal spots, becoming more entire posteriorly, but not extending onto enlarged median subcaudals, which are brown with a darker free margin.</paragraph>
<paragraph id="8BCDE4240203FFF5EDA87FECFB0C5AD5" blockId="4.[151,1437,1127,2016]" pageId="4" pageNumber="413">
<emphasis id="B90638360203FFF5EDA87FECFEA359F2" bold="true" box="[199,315,1451,1476]" pageId="4" pageNumber="413">Variation.</emphasis>
The female paratype agrees in nearly all respects of pholidosis with the holotype, but lacks any trace of precloacal pores and has smaller, less prominent postcloacal spurs. The paratype is, however, more boldly patterned. The white stripes on the neck and shoulders fully enclose a dark brown patch that is itself bisected by a white band perpendicular to the long axis of the body. The dark brown blotches on the trunk are discrete and have irregular margins and two parallel rows along the dorsum, with some sequential blotches partly confluent with one another. A more diffuse series of dark blotches is present on each flank. The pale background colour forms an irregular vertebral stripe separating the two paravertebral rows of dark blotches. The crown is distinctly lighter than the occipital band. All labial scales are whitish, without dark centers. The original tail bears 11 white bands, all with dark brown margins; those at midportion of tail enclosing dark brown spots or dashes.
</paragraph>
<paragraph id="8BCDE4240203FFF5EDA87CA8FB135B1D" blockId="4.[151,1437,1127,2016]" pageId="4" pageNumber="413">
Additional material tentatively referred to 
<taxonomicName id="4C729FA70203FFF5EFD27CA8FBBD5B36" box="[701,1061,1775,1799]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="413" phylum="Chordata" rank="species" species="phetchaburiensis" status="sp. nov.">
<emphasis id="B90638360203FFF5EFD27CA8FBBD5B36" box="[701,1061,1775,1799]" italics="true" pageId="4" pageNumber="413">Cyrtodactylus phetchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235854D0203FFF5E95D7CB7FB095B36" box="[1074,1169,1776,1799]" pageId="4" pageNumber="413" rank="species">
<emphasis id="B90638360203FFF5E95D7CB7FB125B36" bold="true" box="[1074,1162,1776,1799]" pageId="4" pageNumber="413">sp. nov</emphasis>
.
</taxonomicNameLabel>
is similar in scalation features to the holotype, but shows significant variation in dorsal pattern (see 
<emphasis id="B90638360203FFF5EE927D54FBE35B1D" bold="true" box="[1021,1147,1811,1836]" pageId="4" pageNumber="413">Discussion</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C368B7AF0203FFF5EDA87D7FFE685B45" pageId="4" pageNumber="413" type="etymology">
<paragraph id="8BCDE4240203FFF5EDA87D7FFE685B45" blockId="4.[151,1437,1127,2016]" pageId="4" pageNumber="413">
<emphasis id="B90638360203FFF5EDA87D7FFED65B7E" bold="true" box="[199,334,1848,1871]" pageId="4" pageNumber="413">Etymology.</emphasis>
The specific epithet refers to Phetchaburi Province, south-western Thailand, to which the new species appears to be endemic.
</paragraph>
</subSubSection>
<subSubSection id="C368B7AF0203FFF7EDA87D38FF6B59AC" lastPageId="6" lastPageNumber="415" pageId="4" pageNumber="413" type="distribution">
<paragraph id="8BCDE4240203FFF4EDA87D38FC3A5D9C" blockId="4.[151,1437,1127,2016]" lastBlockId="5.[151,1437,151,429]" lastPageId="5" lastPageNumber="414" pageId="4" pageNumber="413">
<emphasis id="B90638360203FFF5EDA87D38FDD65BA9" bold="true" box="[199,590,1919,1944]" pageId="4" pageNumber="413">Distribution and natural history.</emphasis>
The types are from the Tha Yang District of Phetchaburi Province (
<figureCitation id="1349F8A10203FFF5E8267D38FA165BA9" box="[1353,1422,1919,1944]" captionStart="FIGURE 4" captionStartId="5.[151,250,1126,1147]" captionTargetBox="[151,1436,481,1103]" captionTargetId="figure@5.[151,1437,481,1104]" captionTargetPageId="5" captionText="FIGURE 4. Map of the Phetchaburi Province (right) showing the type locality of Cyrtodactylus phetchaburiensis sp. nov. (white circle with black dot). Other specimens referred to the new species have been collected at localities throughout Kaeng Krachan National Park, which occupies most of the western half of the province and abuts the border with Myanmar. Inset map at left shows the relative position of Phetchaburi Province, marked in black." httpUri="https://zenodo.org/record/266702/files/figure.png" pageId="4" pageNumber="413">Fig. 4</figureCitation>
). The holotype was found by day at about two meters above the ground within a cave-like crevice formed by large karst boulders in a secondary forest area. Ecological information from other referred specimens, from Kaeng Krachan and Tha Yang districts, suggests that the species occurs in moist evergreen forest and climbs on vegetation-covered walls and tree trunks (Nabhitabhata in Ulber 1993; Pauwels and Chan-ard 2006). Ulber (1993: 195) quoted Jarujin Nabhitabhata’s field notes: ‘‘All specimens collected in moist evergreen forest on the hill, found at night crawling on vertical steep wayside walls of the forest tracks which are covered with dense vegetation and roots in cracks on the walls. Their reddish brown colour blends well with the colour of the lateritic soil. A young was dug out of the rock pile on the forest track’’. It is highly probable that the species will be found on adjacent hills and mountains covered with primary and mature secondary forests in the northern part of Prachuap Khiri Khan Province and in the southern part of Ratchaburi Province.
</paragraph>
<caption id="DF0DB4AC0202FFF4EDF87E21FC0058E6" httpUri="https://zenodo.org/record/266702/files/figure.png" pageId="5" pageNumber="414" targetBox="[151,1436,481,1103]" targetPageId="5">
<paragraph id="8BCDE4240202FFF4EDF87E21FC0058E6" blockId="5.[151,1436,1125,1239]" pageId="5" pageNumber="414">
<emphasis id="B90638360202FFF4EDF87E21FE8D584A" bold="true" box="[151,277,1126,1147]" pageId="5" pageNumber="414">FIGURE 4.</emphasis>
Map of the Phetchaburi Province (right) showing the type locality of 
<taxonomicName id="4C729FA70202FFF4EE957E22FAA6584A" box="[1018,1342,1125,1147]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="species" species="phetchaburiensis" status="sp. nov.">
<emphasis id="B90638360202FFF4EE957E22FAA6584A" box="[1018,1342,1125,1147]" italics="true" pageId="5" pageNumber="414">Cyrtodactylus phetchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235854D0202FFF4E8277E21FA03584A" box="[1352,1435,1126,1147]" pageId="5" pageNumber="414" rank="species">
<emphasis id="B90638360202FFF4E8277E21FA03584A" bold="true" box="[1352,1435,1126,1147]" pageId="5" pageNumber="414">sp. nov.</emphasis>
</taxonomicNameLabel>
(white circle with black dot). Other specimens referred to the new species have been collected at localities throughout Kaeng Krachan National Park, which occupies most of the western half of the province and abuts the border with Myanmar. Inset map at left shows the relative position of Phetchaburi Province, marked in black.
</paragraph>
</caption>
<paragraph id="8BCDE4240202FFF4EDA87EB9FF405A6A" blockId="5.[151,1436,1278,2023]" pageId="5" pageNumber="414">
<emphasis id="B90638360202FFF4EDA87EB9FDCA5926" bold="true" box="[199,594,1278,1303]" pageId="5" pageNumber="414">Comparisons with other species.</emphasis>
Wood 
<emphasis id="B90638360202FFF4EFC77EB8FD435927" box="[680,731,1278,1302]" italics="true" pageId="5" pageNumber="414">et al</emphasis>
. (2012) established that there was a geographically coherent pattern of phylogenetic relationships among 
<taxonomicName id="4C729FA70202FFF4EFF77F65FCAD590B" box="[664,821,1314,1338]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="genus">
<emphasis id="B90638360202FFF4EFF77F65FCAD590B" box="[664,821,1314,1338]" italics="true" pageId="5" pageNumber="414">Cyrtodactylus</emphasis>
</taxonomicName>
. The Thai material they sampled did not constitute a monophyletic group, but fell out in several clusters in a more inclusive clade that included most members of the genus exclusive of those occurring west of the Salween River. Some groups within this large clade are especially well-circumscribed geographically and are also generally morphologically distinctive from the new species and other Thai 
<taxonomicName id="4C729FA70202FFF4EC7B7FF5FE2959FB" box="[276,433,1458,1482]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="genus">
<emphasis id="B90638360202FFF4EC7B7FF5FE2959FB" box="[276,433,1458,1482]" italics="true" pageId="5" pageNumber="414">Cyrtodactylus</emphasis>
</taxonomicName>
. These include the members of the Philippine/Bornean clade, the Australo-Papuan clade and the Lesser Sunda clade (see Wood 
<emphasis id="B90638360202FFF4EF287F90FDE059DF" box="[583,632,1494,1518]" italics="true" pageId="5" pageNumber="414">et al</emphasis>
. 2012), as well as the distinctive peninsular Indian/Sri Lankan members of the subgenus 
<taxonomicName id="4C729FA70202FFF4EC3F7FBDFCA75A22" authority="Gray (Agarwal &amp; Karanth 2015)" authorityName="Gray (Agarwal &amp; Karanth" authorityYear="2015" box="[336,831,1530,1555]" class="Reptilia" family="Gekkonidae" genus="Geckoella" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="genus">
<emphasis id="B90638360202FFF4EC3F7FBDFE595A23" box="[336,449,1530,1554]" italics="true" pageId="5" pageNumber="414">Geckoella</emphasis>
Gray (Agarwal &amp; Karanth 2015)
</taxonomicName>
. However, because Thai, Indochinese, and peninsular Malaysian taxa are less strictly geographically constrained, it is appropriate to compare the new species with all of these.
</paragraph>
<paragraph id="8BCDE4240202FFF4EDA87C2FFD2E5BD6" blockId="5.[151,1436,1278,2023]" pageId="5" pageNumber="414">
Ngo &amp; Grismer (2012) and Pauwels &amp; Sumontha (2014) provided a summary of 
<taxonomicName id="4C729FA70202FFF4E9147C21FA805A4F" box="[1147,1304,1638,1662]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="genus">
<emphasis id="B90638360202FFF4E9147C21FA805A4F" box="[1147,1304,1638,1662]" italics="true" pageId="5" pageNumber="414">Cyrtodactylus</emphasis>
</taxonomicName>
from these regions that have precloacal pores but lack femoral pores, the condition shown by 
<taxonomicName id="4C729FA70202FFF4E93B7CCCFAAE5A93" box="[1108,1334,1674,1699]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="species" species="phetchaburiensis" status="sp. nov.">
<emphasis id="B90638360202FFF4E93B7CCCFAAE5A93" box="[1108,1334,1674,1699]" italics="true" pageId="5" pageNumber="414">C. phetchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235854D0202FFF4E82F7CCCFA045A93" box="[1344,1436,1675,1698]" pageId="5" pageNumber="414" rank="species">
<emphasis id="B90638360202FFF4E82F7CCCFA045A93" bold="true" box="[1344,1436,1675,1698]" pageId="5" pageNumber="414">sp. nov.</emphasis>
</taxonomicNameLabel>
Among these taxa, the new species may be distinguished by its possession of enlarged femoral scales from 
<taxonomicName id="4C729FA70202FFF4E8ED7CE8FE1E5ADA" authority="Grismer, C." authorityName="Grismer, C." class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="species" species="aurensis">
<emphasis id="B90638360202FFF4E8ED7CE8FF6D5ADB" italics="true" pageId="5" pageNumber="414">C. aurensis</emphasis>
Grismer, 
<emphasis id="B90638360202FFF4EC037C94FE1E5ADA" box="[364,390,1747,1771]" italics="true" pageId="5" pageNumber="414">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360202FFF4ECFF7C95FE7E5ADB" box="[400,486,1746,1770]" italics="true" pageId="5" pageNumber="414">bobrovi</emphasis>
Nguyen, Le, Pham, Ngo, Hoang, Pham &amp; Ziegler, 
<taxonomicName id="4C729FA70202FFF4E9297C94FF665B3E" authority="David, Teynie &amp; Ohler, C." authorityName="David, Teynie &amp; Ohler, C." class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="species" species="buchardi">
<emphasis id="B90638360202FFF4E9297C94FB575ADB" box="[1094,1231,1746,1771]" italics="true" pageId="5" pageNumber="414">C. buchardi</emphasis>
David, Teynié &amp; Ohler, 
<emphasis id="B90638360202FFF4ED8B7CB0FF665B3E" box="[228,254,1783,1807]" italics="true" pageId="5" pageNumber="414">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360202FFF4EC6B7CB1FE205B3F" box="[260,440,1782,1806]" italics="true" pageId="5" pageNumber="414">chauquangensis</emphasis>
Hoang, Orlov, Ananjeva, Johns, Hoang &amp; Dau, 
<taxonomicName id="4C729FA70202FFF4EEB47CB0FEB25B02" authority="Heidrich, Rosler, Vu, Bohme &amp; Ziegler, C." authorityName="Heidrich, Rosler, Vu, Bohme &amp; Ziegler, C." class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="species" species="cryptus">
<emphasis id="B90638360202FFF4EEB47CB0FBD75B3F" box="[987,1103,1783,1807]" italics="true" pageId="5" pageNumber="414">C. cryptus</emphasis>
Heidrich, Rösler, Vu, Böhme &amp; Ziegler, 
<emphasis id="B90638360202FFF4EC7F7D5CFEB25B02" box="[272,298,1819,1843]" italics="true" pageId="5" pageNumber="414">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360202FFF4EC5E7D5DFEF55B03" box="[305,365,1818,1842]" italics="true" pageId="5" pageNumber="414">durio</emphasis>
Grismer, Shahrul, Quah, Muin, Chan, Grismer &amp; Norhayati, 
<taxonomicName id="4C729FA70202FFF4E9757D5CFB405B02" authority="Dring, C." authorityName="Dring, C." box="[1050,1240,1818,1843]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="species" species="elok">
<emphasis id="B90638360202FFF4E9757D5CFBF15B03" box="[1050,1129,1818,1843]" italics="true" pageId="5" pageNumber="414">C. elok</emphasis>
Dring, 
<emphasis id="B90638360202FFF4E9D17D5CFB405B02" box="[1214,1240,1819,1843]" italics="true" pageId="5" pageNumber="414">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360202FFF4E9B17D5DFAC65B03" box="[1246,1374,1818,1842]" italics="true" pageId="5" pageNumber="414">hontreensis</emphasis>
Ngo, Grismer &amp; Grismer, 
<taxonomicName id="4C729FA70202FFF4EC117D78FC0B5B66" authority="Nguyen, Orlov &amp; Darevsky, C." authorityName="Nguyen, Orlov &amp; Darevsky, C." box="[382,915,1854,1879]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="species" species="nigriocularis">
<emphasis id="B90638360202FFF4EC117D78FDA85B67" box="[382,560,1854,1879]" italics="true" pageId="5" pageNumber="414">C. nigriocularis</emphasis>
Nguyen, Orlov &amp; Darevsky, 
<emphasis id="B90638360202FFF4EE177D78FC0B5B66" box="[888,915,1855,1879]" italics="true" pageId="5" pageNumber="414">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360202FFF4EEF67D78FC5D5B67" box="[921,965,1855,1878]" italics="true" pageId="5" pageNumber="414">otai</emphasis>
Nguyen, Le, Pham, Ngo, Hoang, Pham &amp; Ziegler, 
<taxonomicName id="4C729FA70202FFF4ED9A7D24FB9D5B4A" authority="Schneider, Nguyen, Schmitz, Kingsada, Auer &amp; Ziegler, C." authorityName="Schneider, Nguyen, Schmitz, Kingsada, Auer &amp; Ziegler, C." box="[245,1029,1890,1915]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="species" species="pageli">
<emphasis id="B90638360202FFF4ED9A7D24FEC55B4B" box="[245,349,1890,1915]" italics="true" pageId="5" pageNumber="414">C. pageli</emphasis>
Schneider, Nguyen, Schmitz, Kingsada, Auer &amp; Ziegler, 
<emphasis id="B90638360202FFF4EE847D24FB9D5B4A" box="[1003,1029,1891,1915]" italics="true" pageId="5" pageNumber="414">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360202FFF4E9627D24FBE75B4B" box="[1037,1151,1891,1914]" italics="true" pageId="5" pageNumber="414">pantiensis</emphasis>
Grismer, Chan, Grismer, Wood &amp; Belabut, 
<taxonomicName id="4C729FA70202FFF4EC077DC0FBD45BAE" authority="Rosler, Vu, Nguyen, Ngo &amp; Ziegler, C." authorityName="Rosler, Vu, Nguyen, Ngo &amp; Ziegler, C." box="[360,1100,1926,1951]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="species" species="pseudoquadrivirgatus">
<emphasis id="B90638360202FFF4EC077DC0FD185BAF" box="[360,640,1926,1951]" italics="true" pageId="5" pageNumber="414">C. pseudoquadrivirgatus</emphasis>
Rösler, Vu, Nguyen, Ngo &amp; Ziegler, 
<emphasis id="B90638360202FFF4E95D7DC0FBD45BAE" box="[1074,1100,1927,1951]" italics="true" pageId="5" pageNumber="414">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360202FFF4E93A7DC1FB2E5BAF" box="[1109,1206,1926,1950]" italics="true" pageId="5" pageNumber="414">spelaeus</emphasis>
Nazarov, Poyarkov, Orlov, Nguyen, Milto, Martynov, Konstantinov &amp; Chulisov, 
<taxonomicName id="4C729FA70202FFF4EE277DECFB565BF2" authority="Rosler &amp; Glaw, C." authorityName="Rosler &amp; Glaw, C." box="[840,1230,1962,1987]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="B90638360202FFF4EE277DECFC695BF3" box="[840,1009,1963,1987]" italics="true" pageId="5" pageNumber="414">C. stresemanni</emphasis>
Rösler &amp; Glaw, 
<emphasis id="B90638360202FFF4E9DB7DECFB565BF2" box="[1204,1230,1963,1987]" italics="true" pageId="5" pageNumber="414">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360202FFF4E9B97DEDFAD35BF3" box="[1238,1355,1962,1986]" italics="true" pageId="5" pageNumber="414">sumonthai</emphasis>
Bauer, Pauwels &amp; Chanhome, and 
<taxonomicName id="4C729FA70202FFF4ECA17D88FD2A5BD6" authority="Smith" authorityName="Smith" box="[462,690,1998,2023]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="414" phylum="Chordata" rank="species" species="sworderi">
<emphasis id="B90638360202FFF4ECA17D88FDCA5BD7" box="[462,594,1998,2023]" italics="true" pageId="5" pageNumber="414">C. sworderi</emphasis>
(Smith)
</taxonomicName>
.
</paragraph>
<paragraph id="8BCDE4240201FFF7EDA87ADFFE125DFE" blockId="6.[151,1437,151,1437]" pageId="6" pageNumber="415">
By its possession of enlarged subcaudal scales, 
<taxonomicName id="4C729FA70201FFF7EF8F7AD0FBDC5C9E" box="[736,1092,151,175]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="phetchaburiensis" status="sp. nov.">
<emphasis id="B90638360201FFF7EF8F7AD0FBDC5C9E" box="[736,1092,151,175]" italics="true" pageId="6" pageNumber="415">Cyrtodactylus phetchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235854D0201FFF7E9237ADFFB3F5C9E" box="[1100,1191,152,175]" pageId="6" pageNumber="415" rank="species">
<emphasis id="B90638360201FFF7E9237ADFFB3F5C9E" bold="true" box="[1100,1191,152,175]" pageId="6" pageNumber="415">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguished from 
<taxonomicName id="4C729FA70201FFF7EDF87AFBFBC95CE5" authority="Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang &amp; Ziegler, C." authorityName="Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang &amp; Ziegler, C." box="[151,1105,188,213]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="bidoupimontis">
<emphasis id="B90638360201FFF7EDF87AFBFEC35CE5" box="[151,347,188,212]" italics="true" pageId="6" pageNumber="415">C. bidoupimontis</emphasis>
Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang &amp; Ziegler, 
<emphasis id="B90638360201FFF7E9587AFBFBC95CE5" box="[1079,1105,188,212]" italics="true" pageId="6" pageNumber="415">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360201FFF7E9347AFBFB275CE5" box="[1115,1215,188,212]" italics="true" pageId="6" pageNumber="415">buchardi</emphasis>
, 
<taxonomicName id="4C729FA70201FFF7E9BF7AFBFC1A5CC9" authority="Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang &amp; Ziegler, C." authorityName="Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang &amp; Ziegler, C." class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="bugiamapensis">
<emphasis id="B90638360201FFF7E9BF7AFBFA045CE5" box="[1232,1436,188,212]" italics="true" pageId="6" pageNumber="415">C. bugiamapensis</emphasis>
Nazarov, Poyarkov, Orlov, Phung, Nguyen, Hoang &amp; Ziegler, 
<emphasis id="B90638360201FFF7EE077AA7FC1A5CC9" box="[872,898,224,248]" italics="true" pageId="6" pageNumber="415">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360201FFF7EEE37AA7FB885CC6" box="[908,1040,224,247]" italics="true" pageId="6" pageNumber="415">cattienensis</emphasis>
Geissler, Nazarov, Orlov, Böhme, Phung, Nguyen &amp; Ziegler, 
<taxonomicName id="4C729FA70201FFF7ECB67B43FD2A5D2D" box="[473,690,260,284]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="chauquangensis">
<emphasis id="B90638360201FFF7ECB67B43FD2A5D2D" box="[473,690,260,284]" italics="true" pageId="6" pageNumber="415">C. chauquangensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C729FA70201FFF7EFAA7B43FCA45D2A" box="[709,828,260,284]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="cryptus">
<emphasis id="B90638360201FFF7EFAA7B43FCA45D2A" box="[709,828,260,284]" italics="true" pageId="6" pageNumber="415">C. cryptus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C729FA70201FFF7EE207B43FE895D71" authority="Schneider, Phung, Le, Nguyen &amp; Ziegler, C." authorityName="Schneider, Phung, Le, Nguyen &amp; Ziegler, C." class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="cucdongensis">
<emphasis id="B90638360201FFF7EE207B43FB935D2D" box="[847,1035,260,284]" italics="true" pageId="6" pageNumber="415">C. cucdongensis</emphasis>
Schneider, Phung, Le, Nguyen &amp; Ziegler, 
<emphasis id="B90638360201FFF7ED997B6FFE895D71" box="[246,273,296,320]" italics="true" pageId="6" pageNumber="415">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360201FFF7EC767B60FECD5D0E" box="[281,341,295,319]" italics="true" pageId="6" pageNumber="415">durio</emphasis>
, 
<taxonomicName id="4C729FA70201FFF7EC0A7B6FFDF85D71" authority="Smith" authorityName="Smith" box="[357,608,295,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="irregularis">
<emphasis id="B90638360201FFF7EC0A7B6FFE675D0E" box="[357,511,295,320]" italics="true" pageId="6" pageNumber="415">C. irregularis</emphasis>
(Smith)
</taxonomicName>
, 
<taxonomicName id="4C729FA70201FFF7EF027B6FFCDD5D71" authority="Ngo, C." authorityName="Ngo, C." box="[621,837,296,320]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="martini">
<emphasis id="B90638360201FFF7EF027B6FFD7A5D0E" box="[621,738,296,320]" italics="true" pageId="6" pageNumber="415">C. martini</emphasis>
Ngo, 
<emphasis id="B90638360201FFF7EE447B6FFCDD5D71" box="[811,837,296,320]" italics="true" pageId="6" pageNumber="415">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360201FFF7EE227B60FC5C5D0E" box="[845,964,295,319]" italics="true" pageId="6" pageNumber="415">metropolis</emphasis>
Grismer, Wood, Onn, Anuar &amp; Muin, 
<taxonomicName id="4C729FA70201FFF7E8ED7B6FFE915D52" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="pantiensis">
<emphasis id="B90638360201FFF7E8ED7B6FFE915D52" italics="true" pageId="6" pageNumber="415">C. pantiensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C729FA70201FFF7EC737B0BFC985D55" authority="Ulber &amp; Grossmann, C." authorityName="Ulber &amp; Grossmann, C." box="[284,768,332,357]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="papilionoides">
<emphasis id="B90638360201FFF7EC737B0BFE415D55" box="[284,473,332,356]" italics="true" pageId="6" pageNumber="415">C. papilionoides</emphasis>
Ulber &amp; Grossmann, 
<emphasis id="B90638360201FFF7EF897B0BFC985D55" box="[742,768,332,356]" italics="true" pageId="6" pageNumber="415">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360201FFF7EE637B0BFCEB5D55" box="[780,883,332,356]" italics="true" pageId="6" pageNumber="415">payacola</emphasis>
Johnson, Quah, Anuar, Muin, Wood, Grismer, Greer, Chan, Norhayati, Bauer &amp; Grismer, 
<taxonomicName id="4C729FA70201FFF7EFEF7B37FC0F5DB6" box="[640,919,367,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="pseudoquadrivirgatus">
<emphasis id="B90638360201FFF7EFEF7B37FC0F5DB6" box="[640,919,367,392]" italics="true" pageId="6" pageNumber="415">C. pseudoquadrivirgatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C729FA70201FFF7EECA7B37FB7D5DB9" authority="Taylor, C." authorityName="Taylor, C." box="[933,1253,367,392]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="quadrivirgatus">
<emphasis id="B90638360201FFF7EECA7B37FBF45DB6" box="[933,1132,367,392]" italics="true" pageId="6" pageNumber="415">C. quadrivirgatus</emphasis>
Taylor, 
<emphasis id="B90638360201FFF7E9A47B37FB7D5DB9" box="[1227,1253,368,392]" italics="true" pageId="6" pageNumber="415">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360201FFF7E9827B37FAEB5DB6" box="[1261,1395,368,391]" italics="true" pageId="6" pageNumber="415">stresemanni</emphasis>
, 
<taxonomicName id="4C729FA70201FFF7E8ED7B37FF625D9D" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="sworderi">
<emphasis id="B90638360201FFF7E8ED7B37FF625D9D" italics="true" pageId="6" pageNumber="415">C. sworderi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C729FA70201FFF7EC667BD3FBE45D9C" authority="Schneider, Nguyen, Duc Le, Nophaseud, Bonkowski &amp; Ziegler" authorityName="Schneider, Nguyen, Duc Le, Nophaseud, Bonkowski &amp; Ziegler" box="[265,1148,404,429]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="vilaphongi">
<emphasis id="B90638360201FFF7EC667BD3FE3B5D9D" box="[265,419,404,428]" italics="true" pageId="6" pageNumber="415">C. vilaphongi</emphasis>
Schneider, Nguyen, Duc Le, Nophaseud, Bonkowski &amp; Ziegler
</taxonomicName>
, and 
<taxonomicName id="4C729FA70201FFF7E9D67BD3FEA55DFE" authority="Nazarov, Orlov, Nguyen" authorityName="Nazarov, Orlov, Nguyen" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="ziegleri">
<emphasis id="B90638360201FFF7E9D67BD3FAA85D9D" box="[1209,1328,404,428]" italics="true" pageId="6" pageNumber="415">C. ziegleri</emphasis>
Nazarov, Orlov, Nguyen
</taxonomicName>
&amp; Ho.
</paragraph>
<paragraph id="8BCDE4240201FFF7EDA87B9AFB3C5E51" blockId="6.[151,1437,151,1437]" pageId="6" pageNumber="415">
By its possession of 20 longitudinal rows of dorsal tubercles at midbody, 
<taxonomicName id="4C729FA70201FFF7E9627B9BFAEA5DC5" box="[1037,1394,476,500]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="phetchaburiensis" status="sp. nov.">
<emphasis id="B90638360201FFF7E9627B9BFAEA5DC5" box="[1037,1394,476,500]" italics="true" pageId="6" pageNumber="415">Cyrtodactylus phetchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235854D0201FFF7E8157B9AFF505E26" pageId="6" pageNumber="415" rank="species">
<emphasis id="B90638360201FFF7E8157B9AFF505E26" bold="true" pageId="6" pageNumber="415">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguishable from 
<taxonomicName id="4C729FA70201FFF7EC8D7847FDF15E26" box="[482,617,511,536]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="buchardi">
<emphasis id="B90638360201FFF7EC8D7847FDF15E26" box="[482,617,511,536]" italics="true" pageId="6" pageNumber="415">C. buchardi</emphasis>
</taxonomicName>
(25), 
<taxonomicName id="4C729FA70201FFF7EFDC7847FC0C5E26" authority="Ngo" authorityName="Ngo" box="[691,916,512,536]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="eisenmanae">
<emphasis id="B90638360201FFF7EFDC7847FCC25E26" box="[691,858,512,536]" italics="true" pageId="6" pageNumber="415">C. eisenmanae</emphasis>
Ngo
</taxonomicName>
(14), 
<taxonomicName id="4C729FA70201FFF7EEB17847FBA85E26" box="[990,1072,511,536]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="elok">
<emphasis id="B90638360201FFF7EEB17847FBA85E26" box="[990,1072,511,536]" italics="true" pageId="6" pageNumber="415">C. elok</emphasis>
</taxonomicName>
(5–10), 
<taxonomicName id="4C729FA70201FFF7E9FA7847FAA15E26" box="[1173,1337,511,536]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="hontreensis">
<emphasis id="B90638360201FFF7E9FA7847FAA15E26" box="[1173,1337,511,536]" italics="true" pageId="6" pageNumber="415">C. hontreensis</emphasis>
</taxonomicName>
(14), 
<taxonomicName id="4C729FA70201FFF7E8ED7847FEB15E0D" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="nigriocularis">
<emphasis id="B90638360201FFF7E8ED7847FEB15E0D" italics="true" pageId="6" pageNumber="415">C. nigriocularis</emphasis>
</taxonomicName>
(0), 
<taxonomicName id="4C729FA70201FFF7EC0F7863FE5F5E0D" box="[352,455,548,572]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="pageli">
<emphasis id="B90638360201FFF7EC0F7863FE5F5E0D" box="[352,455,548,572]" italics="true" pageId="6" pageNumber="415">C. pageli</emphasis>
</taxonomicName>
(9–14), 
<taxonomicName id="4C729FA70201FFF7EF467863FD7A5E0D" box="[553,738,548,572]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="papilionoides">
<emphasis id="B90638360201FFF7EF467863FD7A5E0D" box="[553,738,548,572]" italics="true" pageId="6" pageNumber="415">C. papilionoides</emphasis>
</taxonomicName>
(12–14), 
<taxonomicName id="4C729FA70201FFF7EE3D7863FC4C5E0D" box="[850,980,548,572]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="spelaeus">
<emphasis id="B90638360201FFF7EE3D7863FC4C5E0D" box="[850,980,548,572]" italics="true" pageId="6" pageNumber="415">C. spelaeus</emphasis>
</taxonomicName>
(10), 
<taxonomicName id="4C729FA70201FFF7E9767863FB585E0A" box="[1049,1216,548,572]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="stresemanni">
<emphasis id="B90638360201FFF7E9767863FB585E0A" box="[1049,1216,548,572]" italics="true" pageId="6" pageNumber="415">C. stresemanni</emphasis>
</taxonomicName>
(13), 
<taxonomicName id="4C729FA70201FFF7E8697863FA045E0D" box="[1286,1436,548,572]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="sumonthai">
<emphasis id="B90638360201FFF7E8697863FA045E0D" box="[1286,1436,548,572]" italics="true" pageId="6" pageNumber="415">C. sumonthai</emphasis>
</taxonomicName>
(12), and 
<taxonomicName id="4C729FA70201FFF7EC6C780FFBFF5E51" authority="Sumontha, Pauwels, Suwannakarn, Nutatheera &amp; Sodob" authorityName="Sumontha, Pauwels, Suwannakarn, Nutatheera &amp; Sodob" box="[259,1127,583,608]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="wangkulangkulae">
<emphasis id="B90638360201FFF7EC6C780FFE725E6E" box="[259,490,583,608]" italics="true" pageId="6" pageNumber="415">C. wangkulangkulae</emphasis>
Sumontha, Pauwels, Suwannakarn, Nutatheera &amp; Sodob
</taxonomicName>
(10).
</paragraph>
<paragraph id="8BCDE4240201FFF7EDA8782BFC845FF9" blockId="6.[151,1437,151,1437]" pageId="6" pageNumber="415">
Among remaining species 
<taxonomicName id="4C729FA70201FFF7EC9F782BFD485EB5" box="[496,720,620,644]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="phetchaburiensis" status="sp. nov.">
<emphasis id="B90638360201FFF7EC9F782BFD485EB5" box="[496,720,620,644]" italics="true" pageId="6" pageNumber="415">C. phetchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235854D0201FFF7EFB9782AFCA85EB5" box="[726,816,621,644]" pageId="6" pageNumber="415" rank="species">
<emphasis id="B90638360201FFF7EFB9782AFCA85EB5" bold="true" box="[726,816,621,644]" pageId="6" pageNumber="415">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguished by a lower number of lamellae beneath digit IV of the pes (16–17) from 
<taxonomicName id="4C729FA70201FFF7EF6C78D7FE895EFC" authority="Kunya, Panmongkol, Pauwels, Sumontha, Meewasana, Bunkhwamdi &amp; Dangsri" authorityName="Kunya, Panmongkol, Pauwels, Sumontha, Meewasana, Bunkhwamdi &amp; Dangsri" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="doisuthep">
<emphasis id="B90638360201FFF7EF6C78D7FD085E96" box="[515,656,655,680]" italics="true" pageId="6" pageNumber="415">C. doisuthep</emphasis>
Kunya, Panmongkol, Pauwels, Sumontha, Meewasana, Bunkhwamdi &amp; Dangsri
</taxonomicName>
(19), 
<taxonomicName id="4C729FA70201FFF7EC3178F3FD705EFC" authority="Chan &amp; Norhayati" authorityName="Chan &amp; Norhayati" box="[350,744,692,717]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="leegrismeri">
<emphasis id="B90638360201FFF7EC3178F3FD9B5EFD" box="[350,515,692,716]" italics="true" pageId="6" pageNumber="415">C. leegrismeri</emphasis>
Chan &amp; Norhayati
</taxonomicName>
(18–20), 
<taxonomicName id="4C729FA70201FFF7EE0F78F3FC8E5EC1" authority="Nguyen, Yang, Le, Nguyen, Orlov, Hoang, Nguyen, Jin, Rao, Hoang Che, Murphy &amp; Zhang" authorityName="Nguyen, Yang, Le, Nguyen, Orlov, Hoang, Nguyen, Jin, Rao, Hoang Che, Murphy &amp; Zhang" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="puhuensis">
<emphasis id="B90638360201FFF7EE0F78F3FC6D5EFD" box="[864,1013,692,716]" italics="true" pageId="6" pageNumber="415">C. puhuensis</emphasis>
Nguyen, Yang, Le, Nguyen, Orlov, Hoang, Nguyen, Jin, Rao, Hoang Che, Murphy &amp; Zhang
</taxonomicName>
(23), 
<taxonomicName id="4C729FA70201FFF7EE34789FFB7F5EC1" authority="Pauwels &amp; Sumontha" authorityName="Pauwels &amp; Sumontha" box="[859,1255,727,752]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="samroiyot">
<emphasis id="B90638360201FFF7EE34789FFC735EDE" box="[859,1003,728,752]" italics="true" pageId="6" pageNumber="415">C. samroiyot</emphasis>
Pauwels &amp; Sumontha
</taxonomicName>
(20), 
<taxonomicName id="4C729FA70201FFF7E844789FFDF35F24" authority="Pauwels, Sumontha, Latinne &amp; Grismer" authorityName="Pauwels, Sumontha, Latinne &amp; Grismer" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="sanook">
<emphasis id="B90638360201FFF7E844789FFA035EDE" box="[1323,1435,727,752]" italics="true" pageId="6" pageNumber="415">C. sanook</emphasis>
Pauwels, Sumontha, Latinne &amp; Grismer
</taxonomicName>
(20), 
<taxonomicName id="4C729FA70201FFF7EFD778BBFC805F22" box="[696,792,764,788]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="surin">
<emphasis id="B90638360201FFF7EFD778BBFC805F22" box="[696,792,764,788]" italics="true" pageId="6" pageNumber="415">C. surin</emphasis>
</taxonomicName>
Chan-ard &amp; Makchai (18), 
<taxonomicName id="4C729FA70201FFF7E90678BBFEE75F09" authority="David, Nguyen, Schneider &amp; Ziegler" authorityName="David, Nguyen, Schneider &amp; Ziegler" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="teyniei">
<emphasis id="B90638360201FFF7E90678BBFB405F22" box="[1129,1240,764,788]" italics="true" pageId="6" pageNumber="415">C. teyniei</emphasis>
David, Nguyen, Schneider &amp; Ziegler
</taxonomicName>
(19–20), 
<taxonomicName id="4C729FA70201FFF7EC9E7967FBE75F09" authority="Nguyen, Kingsada, Rosler, Auer &amp; Ziegler" authorityName="Nguyen, Kingsada, Rosler, Auer &amp; Ziegler" box="[497,1151,799,824]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="wayakonei">
<emphasis id="B90638360201FFF7EC9E7967FD135F06" box="[497,651,799,824]" italics="true" pageId="6" pageNumber="415">C. wayakonei</emphasis>
Nguyen, Kingsada, Rösler, Auer &amp; Ziegler
</taxonomicName>
(19–20); and by a lower number of precloacal pores (5) than 
<taxonomicName id="4C729FA70201FFF7EF5B7903FC875F6C" authority="Smith" authorityName="Smith" box="[564,799,836,861]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B90638360201FFF7EF5B7903FD595F6D" box="[564,705,836,860]" italics="true" pageId="6" pageNumber="415">C. angularis</emphasis>
(Smith)
</taxonomicName>
(6), 
<taxonomicName id="4C729FA70201FFF7EE387903FC7D5F6D" box="[855,997,836,860]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="doisuthep">
<emphasis id="B90638360201FFF7EE387903FC7D5F6D" box="[855,997,836,860]" italics="true" pageId="6" pageNumber="415">C. doisuthep</emphasis>
</taxonomicName>
(6), 
<taxonomicName id="4C729FA70201FFF7E9727903FAB95F6C" authority="Smith" authorityName="Smith" box="[1053,1313,836,861]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="intermedius">
<emphasis id="B90638360201FFF7E9727903FB5A5F6D" box="[1053,1218,836,860]" italics="true" pageId="6" pageNumber="415">C. intermedius</emphasis>
(Smith)
</taxonomicName>
(8–10), 
<taxonomicName id="4C729FA70201FFF7E8ED7903FDF35FB1" authority="Ngo, Grismer &amp; Grismer" authorityName="Ngo, Grismer &amp; Grismer" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="phuquocensis">
<emphasis id="B90638360201FFF7E8ED7903FEB75F4E" italics="true" pageId="6" pageNumber="415">C. phuquocensis</emphasis>
Ngo, Grismer &amp; Grismer
</taxonomicName>
(7–9), 
<taxonomicName id="4C729FA70201FFF7EFA5792FFCF85F4E" box="[714,864,872,896]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="samroiyot">
<emphasis id="B90638360201FFF7EFA5792FFCF85F4E" box="[714,864,872,896]" italics="true" pageId="6" pageNumber="415">C. samroiyot</emphasis>
</taxonomicName>
(7), 
<taxonomicName id="4C729FA70201FFF7EECC792FFB8D5F4E" box="[931,1045,872,896]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="teyniei">
<emphasis id="B90638360201FFF7EECC792FFB8D5F4E" box="[931,1045,872,896]" italics="true" pageId="6" pageNumber="415">C. teyniei</emphasis>
</taxonomicName>
(14 in female holotype) and 
<taxonomicName id="4C729FA70201FFF7E8ED792FFE975F95" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="wayakonei">
<emphasis id="B90638360201FFF7E8ED792FFE975F95" italics="true" pageId="6" pageNumber="415">C. wayakonei</emphasis>
</taxonomicName>
(6–8), and a greater number of precloacal pores than 
<taxonomicName id="4C729FA70201FFF7EE0B79CBFA9A5F94" authority="Darevsky &amp; Szczerbak" authorityName="Darevsky &amp; Szczerbak" box="[868,1282,908,933]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="paradoxus">
<emphasis id="B90638360201FFF7EE0B79CBFC625F95" box="[868,1018,908,932]" italics="true" pageId="6" pageNumber="415">C. paradoxus</emphasis>
Darevsky &amp; Szczerbak
</taxonomicName>
(0–4), and 
<taxonomicName id="4C729FA70201FFF7E8ED79CBFD4A5FF6" authority="Phung, Van Schingen, Ziegler &amp; Nguyen" authorityName="Phung, Van Schingen, Ziegler &amp; Nguyen" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="thuongae">
<emphasis id="B90638360201FFF7E8ED79CBFF675FF6" italics="true" pageId="6" pageNumber="415">C. thuongae</emphasis>
Phung, Van Schingen, Ziegler &amp; Nguyen
</taxonomicName>
(0–1).
</paragraph>
<paragraph id="8BCDE4240201FFF7EDA87992FF6B59AC" blockId="6.[151,1437,151,1437]" pageId="6" pageNumber="415">
In addition, the colour pattern of 
<taxonomicName id="4C729FA70201FFF7EF517993FC875FDD" box="[574,799,980,1004]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="phetchaburiensis" status="sp. nov.">
<emphasis id="B90638360201FFF7EF517993FC875FDD" box="[574,799,980,1004]" italics="true" pageId="6" pageNumber="415">C. phetchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235854D0201FFF7EE487992FC195FDD" box="[807,897,981,1004]" pageId="6" pageNumber="415" rank="species">
<emphasis id="B90638360201FFF7EE487992FC195FDD" bold="true" box="[807,897,981,1004]" pageId="6" pageNumber="415">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinctly different from all Thai, Malay, and Indochinese 
<taxonomicName id="4C729FA70201FFF7EC4A79B0FE5A583E" box="[293,450,1015,1039]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="genus">
<emphasis id="B90638360201FFF7EC4A79B0FE5A583E" box="[293,450,1015,1039]" italics="true" pageId="6" pageNumber="415">Cyrtodactylus</emphasis>
</taxonomicName>
that share the lack of femoral pores, presence of six or fewer precloacal pores, enlarged femoral scales present, broad subcaudal scutes, and 19–24 dorsal rows of tubercles (or condition unknown). Specifically, it differs from 
<taxonomicName id="4C729FA70201FFF7EC837E07FC995869" authority="Smith" authorityName="Smith" box="[492,769,1087,1112]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="condorensis">
<emphasis id="B90638360201FFF7EC837E07FD025866" box="[492,666,1087,1112]" italics="true" pageId="6" pageNumber="415">C. condorensis</emphasis>
(Smith)
</taxonomicName>
, 
<taxonomicName id="4C729FA70201FFF7EE7B7E07FC325866" box="[788,938,1087,1112]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="doisuthep">
<emphasis id="B90638360201FFF7EE7B7E07FC325866" box="[788,938,1087,1112]" italics="true" pageId="6" pageNumber="415">C. doisuthep</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C729FA70201FFF7EEAF7E07FBFF5866" box="[960,1127,1087,1112]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="leegrismeri">
<emphasis id="B90638360201FFF7EEAF7E07FBFF5866" box="[960,1127,1087,1112]" italics="true" pageId="6" pageNumber="415">C. leegrismeri</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C729FA70201FFF7E9127E07FA8E5866" box="[1149,1302,1087,1112]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="puhuensis">
<emphasis id="B90638360201FFF7E9127E07FA8E5866" box="[1149,1302,1087,1112]" italics="true" pageId="6" pageNumber="415">C. puhuensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C729FA70201FFF7E8437E07FD42584D" authority="Panitvong, Sumontha, Tunprasert &amp; Pauwels, C." authorityName="Panitvong, Sumontha, Tunprasert &amp; Pauwels, C." class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="saiyok">
<emphasis id="B90638360201FFF7E8437E07FA035866" box="[1324,1435,1087,1112]" italics="true" pageId="6" pageNumber="415">C. saiyok</emphasis>
Panitvong, Sumontha, Tunprasert &amp; Pauwels, 
<emphasis id="B90638360201FFF7EFAF7E23FD42584D" box="[704,730,1124,1148]" italics="true" pageId="6" pageNumber="415">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360201FFF7EF887E23FCCE584A" box="[743,854,1124,1147]" italics="true" pageId="6" pageNumber="415">samroiyot</emphasis>
, 
<taxonomicName id="4C729FA70201FFF7EE047E23FC7A584D" box="[875,994,1124,1148]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="sanook">
<emphasis id="B90638360201FFF7EE047E23FC7A584D" box="[875,994,1124,1148]" italics="true" pageId="6" pageNumber="415">C. sanook</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C729FA70201FFF7EE987E23FA04584D" authority="Ngo &amp; Grismer, C." authorityName="Ngo &amp; Grismer, C." box="[1015,1436,1124,1149]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="thochuensis">
<emphasis id="B90638360201FFF7EE987E23FB3B584D" box="[1015,1187,1124,1148]" italics="true" pageId="6" pageNumber="415">C. thochuensis</emphasis>
Ngo &amp; Grismer, 
<emphasis id="B90638360201FFF7E8ED7E23FA04584D" box="[1410,1436,1124,1148]" italics="true" pageId="6" pageNumber="415">C.</emphasis>
</taxonomicName>
<emphasis id="B90638360201FFF7EDF87EC0FF6758AE" box="[151,255,1159,1183]" italics="true" pageId="6" pageNumber="415">thuongae</emphasis>
and 
<taxonomicName id="4C729FA70201FFF7EC567ECFFD155891" authority="Ngo &amp; Chan" authorityName="Ngo &amp; Chan" box="[313,653,1159,1184]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="yangbayensis">
<emphasis id="B90638360201FFF7EC567ECFFE6A58AE" box="[313,498,1159,1184]" italics="true" pageId="6" pageNumber="415">C. yangbayensis</emphasis>
Ngo &amp; Chan
</taxonomicName>
in lacking body bands or well-defined transverse markings, from 
<taxonomicName id="4C729FA70201FFF7E8ED7ECFFE9B58F5" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="angularis">
<emphasis id="B90638360201FFF7E8ED7ECFFE9B58F5" italics="true" pageId="6" pageNumber="415">C. angularis</emphasis>
</taxonomicName>
in lacking W-shaped dorsal markings, from 
<taxonomicName id="4C729FA70201FFF7EF9C7EEBFC1458F5" box="[755,908,1196,1220]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="wayakonei">
<emphasis id="B90638360201FFF7EF9C7EEBFC1458F5" box="[755,908,1196,1220]" italics="true" pageId="6" pageNumber="415">C. wayakonei</emphasis>
</taxonomicName>
in lacking a reticulated pattern on the head and in possessing a well-defined nuchal loop, from 
<taxonomicName id="4C729FA70201FFF7EFDE7E97FCD158D6" box="[689,841,1231,1256]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="paradoxus">
<emphasis id="B90638360201FFF7EFDE7E97FCD158D6" box="[689,841,1231,1256]" italics="true" pageId="6" pageNumber="415">C. paradoxus</emphasis>
</taxonomicName>
in lacking a thin, pale vertebral stripe and in having an entire (
<emphasis id="B90638360201FFF7EC667EB3FEC9593A" box="[265,337,1268,1291]" italics="true" pageId="6" pageNumber="415">versus</emphasis>
bisected) nuchal loop, from 
<taxonomicName id="4C729FA70201FFF7EFFA7EB3FCBD593A" box="[661,805,1268,1292]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="peguensis">
<emphasis id="B90638360201FFF7EFFA7EB3FCBD593A" box="[661,805,1268,1292]" italics="true" pageId="6" pageNumber="415">C. peguensis</emphasis>
</taxonomicName>
in lacking a reticulated pattern on the head and discrete dark dorsal spots separated by narrow pale margins or reticulations, and from 
<taxonomicName id="4C729FA70201FFF7EE987F5FFBEB591E" box="[1015,1139,1303,1328]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="oldhami">
<emphasis id="B90638360201FFF7EE987F5FFBEB591E" box="[1015,1139,1303,1328]" italics="true" pageId="6" pageNumber="415">C. oldhami</emphasis>
</taxonomicName>
in having a diffuse pattern of dark blotches on a light background rather than four longitudinal series of white spots. In addition, 
<taxonomicName id="4C729FA70201FFF7E8ED7F7BFECE5946" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="415" phylum="Chordata" rank="species" species="phetchaburiensis" status="sp. nov.">
<emphasis id="B90638360201FFF7E8ED7F7BFECE5946" italics="true" pageId="6" pageNumber="415">C. phetchaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235854D0201FFF7EC327F27FE2F5946" box="[349,439,1376,1399]" pageId="6" pageNumber="415" rank="species">
<emphasis id="B90638360201FFF7EC327F27FE2F5946" bold="true" box="[349,439,1376,1399]" pageId="6" pageNumber="415">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from all of these species is having discrete, dark scapular patches with thin white borders.
</paragraph>
</subSubSection>
</treatment>
</document>