313 lines
82 KiB
XML
313 lines
82 KiB
XML
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<mods:title id="26110D8CE82BF9DFC959CFD96A045D12">Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species</mods:title>
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<mods:name id="24E7113D3AEF5F596D7C95809C3D6F24" type="personal">
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<mods:namePart id="ACAA80E14B86D59641E8BC2DEEAE599B">Toda, Masanori J.</mods:namePart>
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<mods:affiliation id="AC148E7EF640C1BFEACE3D93C2CE0B49">Hokkaido University Museum, Hokkaido University, Sapporo 060 - 0810, Japan</mods:affiliation>
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<mods:namePart id="E0B26DE69C1A6B532E949BF3855473B7">Bänziger, Hans</mods:namePart>
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<mods:affiliation id="AB78C82FF817790FA32DB43E2986EF10">Department of Entomology and Plant Pathology, Faculty of Agriculture, Chiang Mai University, Chiang Mai 50200, Thailand hans. banziger @ cmu. ac. th; https: // orcid. org / 0000 - 0001 - 8913 - 3034</mods:affiliation>
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<mods:namePart id="EC66011B75FE9CD81EB4ACB6A03BD06C">Sati, Pradeep C.</mods:namePart>
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<mods:affiliation id="9F2AF2C6E3ABCEC51AD9C2DF715ECD86">Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & satipradeep 28 @ gmail. com; https: // orcid. org / 0000 - 0001 - 6809 - 8575</mods:affiliation>
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<mods:namePart id="110F7CA268FEBBB60A6B6983C5ABFFF7">Fartyal, Rajendra S.</mods:namePart>
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<mods:affiliation id="163197F563296C22C2ED547CB774BCD2">Systematic & Cytogenetic Laboratory, Department of Zoology and Biotechnology, HNB Garhwal University, Srinagar-Garhwal, 246174, Uttarakhand, India. & rs. fartyal @ hnbgu. ac. in; https: // orcid. org / 0000 - 0002 - 8844 - 6263</mods:affiliation>
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<mods:namePart id="06D80AA3B76B94E66B8F3C0EAE036268">Suwito, Awit</mods:namePart>
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<mods:affiliation id="89C33FA443FE61B1939AADB22BE2D71A">Zoology Division (Museum Zoologicum Bogoriense), Research Center for Biology-LIPI, Jl. Raya Jakarta-Bogor Km. 46, Cibinong 16911, West Java, Indonesia. awitswt @ yahoo. com</mods:affiliation>
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<mods:namePart id="AFD355CBC62F849A4FBA8F8CEC2E0A0D">Katoh, Toru</mods:namePart>
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<mods:affiliation id="737CE53E2D68676CB0DD7C4B8697A36F">Department of Biological Sciences, Faculty of Science, Hokkaido University, Sapporo 060 - 0810, Japan t _ katoh @ sci. hokudai. ac. jp; https: // orcid. org / 0000 - 0003 - 3376 - 8545</mods:affiliation>
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<mods:title id="22011B2849942F4B6E206ACEE19A3E51">Zootaxa</mods:title>
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<treatment id="03C86C16EB2E5753FF6EDBBEFE55FD01" ID-DOI="http://doi.org/10.5281/zenodo.5221266" ID-GBIF-Taxon="164612796" ID-Zenodo-Dep="5221266" LSID="urn:lsid:plazi:treatment:03C86C16EB2E5753FF6EDBBEFE55FD01" httpUri="http://treatment.plazi.org/id/03C86C16EB2E5753FF6EDBBEFE55FD01" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
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Subgenus
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<taxonomicName id="4C61A683EB2E5752FEEEDBBEFDEDF97A" ID-CoL="89RX5" authority="Chen, 2011" authorityName="Chen" authorityYear="2011" box="[279,518,1718,1745]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subGenus" subGenus="Ashima">
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<emphasis id="B9150112EB2E5752FEEEDBBEFE9FF97B" bold="true" box="[279,372,1718,1745]" italics="true" pageId="8" pageNumber="9">Ashima</emphasis>
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Chen, 2011
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<taxonomicName id="4C61A683EB2E5752FF6EDBF3FEF4F8B9" ID-CoL="89RX5" authority="Chen" authorityName="Chen" authorityYear="2011" box="[151,287,1786,1811]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subGenus" subGenus="Ashima">
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<emphasis id="B9150112EB2E5752FF6EDBF3FF08F8B9" box="[151,227,1787,1811]" italics="true" pageId="8" pageNumber="9">Ashima</emphasis>
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Chen
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</taxonomicName>
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in
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<treatmentCitation id="0AC0FB11EB2E5752FEC6DBF3FDF8F8B9" author="Cao, H. L. & Wang, X. L. & Gao, J. J. & Prigent, S. R. & Watabe, H. & Zhang, Y. P. & Chen, H. W." box="[319,531,1786,1811]" page="682" pageId="8" pageNumber="9" year="2011">
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<bibRefCitation id="EFF0A0F1EB2E5752FEC6DBF3FDF8F8B9" author="Cao" box="[319,531,1786,1811]" etAl="et al." firstAuthor="Cao" pageId="8" pageNumber="9" pagination="677 - 685" refId="ref33633" refString="Cao, H. L., Wang, X. L., Gao, J. J., Prigent, S. R., Watabe, H., Zhang, Y. P. & Chen, H. W. (2011) Phylogeny of the African and Asian Phortica (Drosophilidae) deduced from nuclear and mitochondrial DNA sequences. Molecular Phylogenetics and Evolution, 61, 677 - 685. https: // doi. org / 10.1016 / j. ympev. 2011.08.002" type="journal article" year="2011">
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Cao
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<emphasis id="B9150112EB2E5752FE97DBF3FE70F8B9" box="[366,411,1787,1811]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
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., 2011: 682
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</bibRefCitation>
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</treatmentCitation>
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(as subgenus of
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<taxonomicName id="4C61A683EB2E5752FD44DBF3FCFDF8B9" ID-CoL="9G2PY" authorityName="Schiner" authorityYear="1862" box="[701,790,1787,1811]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="B9150112EB2E5752FD44DBF3FCFDF8B9" box="[701,790,1787,1811]" italics="true" pageId="8" pageNumber="9">Phortica</emphasis>
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</taxonomicName>
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);
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||
<treatmentCitation id="0AC0FB11EB2E5752FCD5DBF3FBF8F8B9" author="Gong, L. & Zhu, L. & Chen, H. W." box="[812,1043,1787,1811]" page="438" pageId="8" pageNumber="9" year="2018">
|
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<bibRefCitation id="EFF0A0F1EB2E5752FCD5DBF3FBF8F8B9" author="Gong" box="[812,1043,1787,1811]" etAl="et al." firstAuthor="Gong" pageId="8" pageNumber="9" pagination="434 - 446" refId="ref34842" refString="Gong, L., Zhu, L. & Chen, H. W. (2018) Six new species of the subgenera Alloparadisa and Ashima (Diptera: Drosophilidae: Phortica) from Yunnan, China, with DNA barcoding information. Annales de la Societe entomologique de France, New Series, 54, 434 - 446. https: // doi. org / 10.1080 / 00379271.2018.1491327" type="journal article" year="2018">
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Gong
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||
<emphasis id="B9150112EB2E5752FC93DBF3FC7CF8B9" box="[874,919,1787,1811]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
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., 2018: 438
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</bibRefCitation>
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</treatmentCitation>
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.
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<typeStatus id="54DA63A2EB2E5752FBE5DBF3FBA5F8B9" box="[1052,1102,1787,1811]" pageId="8" pageNumber="9">Type</typeStatus>
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species:
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<taxonomicName id="4C61A683EB2E5752FB52DBF3FE79F89B" ID-CoL="8DBXC" authority="Senior-White, 1921" authorityName="Senior-White" authorityYear="1921" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="xyleboriphaga">
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<emphasis id="B9150112EB2E5752FB52DBF3FA77F8B9" box="[1195,1436,1787,1811]" italics="true" pageId="8" pageNumber="9">Phortica xyleboriphaga</emphasis>
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<bibRefCitation id="EFF0A0F1EB2E5752FF3EDA11FE79F89B" author="Senior-White" box="[199,402,1817,1842]" firstAuthor="Senior-White" pageId="8" pageNumber="9" pagination="381 - 396" refId="ref36831" refString="Senior-White, R. A. (1921) New Ceylon Diptera. Spolia Zeylanica, 11, 381 - 396. https: // doi. org / 10.5962 / bhl. part. 12654" type="journal article" year="1921">Senior-White, 1921</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<subSubSection id="C37B8E8BEB2E5753FF6EDA57FB08FDB1" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
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<paragraph id="8BDEDD00EB2E5752FF6EDA57FB88F84C" blockId="8.[151,1436,1887,2022]" pageId="8" pageNumber="9">
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<emphasis id="B9150112EB2E5752FF6EDA57FEF9F8D0" bold="true" box="[151,274,1887,1914]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
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Projections at anterolateral corners of the cibarium longer than 1/2 width of cibarial, anterior margin (ch.10-1;
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<figureCitation id="135AC185EB2E5752FEFDDA8CFEB7F834" box="[260,348,1924,1950]" captionStart="FIGURE 2" captionStartId="12.[151,250,1856,1881]" captionTargetBox="[151,1435,181,1830]" captionTargetId="figure@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURE 2. Phortica (Ashima) efragmentata Toda, sp. nov. (♂ holotype).A, Antenna; B, palpus; C, cibarium (dorsal view); D, proboscis (lateral view); E, fore tarsus; F, abdomen (dorsal view); G, legs; H, abdominal sternites. Abbreviations: a s, anterior sensilla; m s, medial sensilla; p s, posterior sensilla; s c, sensilla campaniformia; s-I–IV, sternites I–IV; t-I–IV, tergites I–IV. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm in A–D; 0.5 mm in E–H." figureDoi="http://doi.org/10.5281/zenodo.3884616" httpUri="https://zenodo.org/record/3884616/files/figure.png" pageId="8" pageNumber="9">Fig. 2C</figureCitation>
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); mid-dorsally constricted epandrium (ch.29-1;
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<figureCitation id="135AC185EB2E5752FC8FDA8CFC00F834" box="[886,1003,1924,1950]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="8" pageNumber="9">Figs 3B,D</figureCitation>
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||
,
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||
<figureCitation id="135AC185EB2E5752FC0EDA8CFBF1F834" box="[1015,1050,1924,1950]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="8" pageNumber="9">4B</figureCitation>
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||
,
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||
<figureCitation id="135AC185EB2E5752FBDFDA8BFBA0F837" box="[1062,1099,1923,1949]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen & Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="8" pageNumber="9">6D</figureCitation>
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,
|
||
<figureCitation id="135AC185EB2E5752FBAEDA8CFB93F834" box="[1111,1144,1924,1950]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="8" pageNumber="9">7E</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2E5752FB7DDA8CFB42F834" box="[1156,1193,1924,1950]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="8" pageNumber="9">8G</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2E5752FB4CDA8CFB3EF834" box="[1205,1237,1924,1950]" captionStart="FIGURE 9" captionStartId="32.[151,250,1936,1961]" captionTargetBox="[166,1420,633,1909]" captionTargetId="figure@32.[166,1420,633,1910]" captionTargetPageId="32" captionText="FIGURE 9. Phortica (Ashima) takehiroi Toda, sp. nov. (♂ paratype). A, Head (caudal view); B, antenna; C, palpus and prob- ocsis; D, prementum; E, abdominal sternites; F, periphallic organs (caudolateral view); G–I, phallic organs (lateral, ventrolateral and ventral views, respectively). Scale bars: 0.1 mm in B–D, F–I; 0.5 mm in A, E." figureDoi="http://doi.org/10.5281/zenodo.3884632" httpUri="https://zenodo.org/record/3884632/files/figure.png" pageId="8" pageNumber="9">9F</figureCitation>
|
||
); less sclerotized, broad lobe-like surstylus (ch.34-1;
|
||
<figureCitation id="135AC185EB2E5752FDDCDAA0FD94F868" box="[549,639,1960,1986]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="8" pageNumber="9">Figs 3E</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2E5752FD72DAA0FD45F868" box="[651,686,1960,1986]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="8" pageNumber="9">4B</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2E5752FD42DAAFFD35F868" box="[699,734,1959,1986]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati & Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="8" pageNumber="9">5C</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2E5752FD12DAAFFCFBF86B" box="[747,784,1959,1985]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen & Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="8" pageNumber="9">6D</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2E5752FCE4DAA0FCD5F868" box="[797,830,1960,1986]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="8" pageNumber="9">7E</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2E5752FCB2DAA0FC87F868" box="[843,876,1960,1986]" captionStart="FIGURE 9" captionStartId="32.[151,250,1936,1961]" captionTargetBox="[166,1420,633,1909]" captionTargetId="figure@32.[166,1420,633,1910]" captionTargetPageId="32" captionText="FIGURE 9. Phortica (Ashima) takehiroi Toda, sp. nov. (♂ paratype). A, Head (caudal view); B, antenna; C, palpus and prob- ocsis; D, prementum; E, abdominal sternites; F, periphallic organs (caudolateral view); G–I, phallic organs (lateral, ventrolateral and ventral views, respectively). Scale bars: 0.1 mm in B–D, F–I; 0.5 mm in A, E." figureDoi="http://doi.org/10.5281/zenodo.3884632" httpUri="https://zenodo.org/record/3884632/files/figure.png" pageId="8" pageNumber="9">9F</figureCitation>
|
||
); hypandrium with a pair of anterolateral, inward extensions articulated to base of pregonite (ch.42-1;
|
||
<figureCitation id="135AC185EB2E5752FD1ADAC4FCB9F84C" box="[739,850,1996,2022]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="8" pageNumber="9">Figs 3C,F</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2E5752FCA4DAC4FC69F84C" box="[861,898,1996,2022]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="8" pageNumber="9">4D</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2E5752FC74DAC3FC44F84C" box="[909,943,1995,2022]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati & Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="8" pageNumber="9">5E</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2E5752FC42DAC3FC34F84F" box="[955,991,1995,2021]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen & Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="8" pageNumber="9">6G</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2E5752FC12DAC4FBE2F84C" box="[1003,1033,1996,2022]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="8" pageNumber="9">7F</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2E5752FBEDDAC4FBC4F84C" box="[1044,1071,1996,2022]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="8" pageNumber="9">8J</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2E5752FBC2DAC4FBBDF84C" box="[1083,1110,1996,2022]" captionStart="FIGURE 9" captionStartId="32.[151,250,1936,1961]" captionTargetBox="[166,1420,633,1909]" captionTargetId="figure@32.[166,1420,633,1910]" captionTargetPageId="32" captionText="FIGURE 9. Phortica (Ashima) takehiroi Toda, sp. nov. (♂ paratype). A, Head (caudal view); B, antenna; C, palpus and prob- ocsis; D, prementum; E, abdominal sternites; F, periphallic organs (caudolateral view); G–I, phallic organs (lateral, ventrolateral and ventral views, respectively). Scale bars: 0.1 mm in B–D, F–I; 0.5 mm in A, E." figureDoi="http://doi.org/10.5281/zenodo.3884632" httpUri="https://zenodo.org/record/3884632/files/figure.png" pageId="8" pageNumber="9">9I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDEDD00EB2F5753FF3EDD90FB08FDB1" blockId="9.[151,1437,152,683]" pageId="9" pageNumber="10">
|
||
<emphasis id="B9150112EB2F5753FF3EDD90FE42FF19" bold="true" box="[199,425,152,179]" pageId="9" pageNumber="10">Shared characters.</emphasis>
|
||
Characters described as commonly seen across this subgenus (the
|
||
<taxonomicName id="4C61A683EB2F5753FB58DD90FA8EFF18" authorityName="Duda (Duda" authorityYear="1923" box="[1185,1381,152,178]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="foliiseta">
|
||
<emphasis id="B9150112EB2F5753FB58DD90FA8EFF18" box="[1185,1381,152,178]" italics="true" pageId="9" pageNumber="10">Phortica foliiseta</emphasis>
|
||
</taxonomicName>
|
||
species complex at that time) by
|
||
<bibRefCitation id="EFF0A0F1EB2F5753FE17DDB4FD25FF7D" author="Chen" box="[494,718,188,215]" etAl="et al." firstAuthor="Chen" pageId="9" pageNumber="10" pagination="419 - 429" refId="ref34109" refString="Chen, H. W., Toda, M. J. & Gao, J. J. (2005 b) The Phortica (s. str.) foliiseta species-complex (Diptera, Drosophilidae) from China and its adjacent countries. Acta Zootaxonomica Sinica, 30, 419 - 429." type="journal article" year="2005" yearSuffix="b">
|
||
Chen
|
||
<emphasis id="B9150112EB2F5753FDCBDDB5FD8EFF7C" box="[562,613,188,214]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
|
||
. (2005b)
|
||
</bibRefCitation>
|
||
are not repeated here. Occipital setae on lower portion shorter than postocular setae (ch.2-0;
|
||
<figureCitation id="135AC185EB2F5753FE15DDE8FDA2FF51" box="[492,585,224,251]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="9" pageNumber="10">Figs 7A</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FDACDDE9FD90FF51" box="[597,635,225,251]" captionStart="FIGURE 9" captionStartId="32.[151,250,1936,1961]" captionTargetBox="[166,1420,633,1909]" captionTargetId="figure@32.[166,1420,633,1910]" captionTargetPageId="32" captionText="FIGURE 9. Phortica (Ashima) takehiroi Toda, sp. nov. (♂ paratype). A, Head (caudal view); B, antenna; C, palpus and prob- ocsis; D, prementum; E, abdominal sternites; F, periphallic organs (caudolateral view); G–I, phallic organs (lateral, ventrolateral and ventral views, respectively). Scale bars: 0.1 mm in B–D, F–I; 0.5 mm in A, E." figureDoi="http://doi.org/10.5281/zenodo.3884632" httpUri="https://zenodo.org/record/3884632/files/figure.png" pageId="9" pageNumber="10">9A</figureCitation>
|
||
). Pedicel with only 1 thick seta (ch.4-0;
|
||
<figureCitation id="135AC185EB2F5753FBBEDDE8FB48FF51" box="[1095,1187,224,251]" captionStart="FIGURE 2" captionStartId="12.[151,250,1856,1881]" captionTargetBox="[151,1435,181,1830]" captionTargetId="figure@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURE 2. Phortica (Ashima) efragmentata Toda, sp. nov. (♂ holotype).A, Antenna; B, palpus; C, cibarium (dorsal view); D, proboscis (lateral view); E, fore tarsus; F, abdomen (dorsal view); G, legs; H, abdominal sternites. Abbreviations: a s, anterior sensilla; m s, medial sensilla; p s, posterior sensilla; s c, sensilla campaniformia; s-I–IV, sternites I–IV; t-I–IV, tergites I–IV. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm in A–D; 0.5 mm in E–H." figureDoi="http://doi.org/10.5281/zenodo.3884616" httpUri="https://zenodo.org/record/3884616/files/figure.png" pageId="9" pageNumber="10">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FB49DDE9FB3EFF51" box="[1200,1237,225,251]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="9" pageNumber="10">4A</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FB18DDE8FAEDFF51" box="[1249,1286,224,251]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati & Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="9" pageNumber="10">5A</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FAEBDDE8FADCFF50" box="[1298,1335,224,250]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen & Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="9" pageNumber="10">6A</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FABADDE9FA8DFF51" box="[1347,1382,225,251]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="9" pageNumber="10">7C</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FA8ADDE9FA7CFF51" box="[1395,1431,225,251]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="9" pageNumber="10">8A</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FF6EDC0DFF57FEB5" box="[151,188,261,287]" captionStart="FIGURE 9" captionStartId="32.[151,250,1936,1961]" captionTargetBox="[166,1420,633,1909]" captionTargetId="figure@32.[166,1420,633,1910]" captionTargetPageId="32" captionText="FIGURE 9. Phortica (Ashima) takehiroi Toda, sp. nov. (♂ paratype). A, Head (caudal view); B, antenna; C, palpus and prob- ocsis; D, prementum; E, abdominal sternites; F, periphallic organs (caudolateral view); G–I, phallic organs (lateral, ventrolateral and ventral views, respectively). Scale bars: 0.1 mm in B–D, F–I; 0.5 mm in A, E." figureDoi="http://doi.org/10.5281/zenodo.3884632" httpUri="https://zenodo.org/record/3884632/files/figure.png" pageId="9" pageNumber="10">9B</figureCitation>
|
||
). Clypeus medially pale (ch.8-1;
|
||
<figureCitation id="135AC185EB2F5753FDC4DC0CFD7FFEB5" box="[573,660,260,287]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="9" pageNumber="10">Fig. 8F</figureCitation>
|
||
). Lateral membrane of proboscis scale-like patterned (ch.13-1;
|
||
<figureCitation id="135AC185EB2F5753FA89DC0CFF56FEE9" captionStart="FIGURE 2" captionStartId="12.[151,250,1856,1881]" captionTargetBox="[151,1435,181,1830]" captionTargetId="figure@12.[151,1435,181,1830]" captionTargetPageId="12" captionText="FIGURE 2. Phortica (Ashima) efragmentata Toda, sp. nov. (♂ holotype).A, Antenna; B, palpus; C, cibarium (dorsal view); D, proboscis (lateral view); E, fore tarsus; F, abdomen (dorsal view); G, legs; H, abdominal sternites. Abbreviations: a s, anterior sensilla; m s, medial sensilla; p s, posterior sensilla; s c, sensilla campaniformia; s-I–IV, sternites I–IV; t-I–IV, tergites I–IV. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm in A–D; 0.5 mm in E–H." figureDoi="http://doi.org/10.5281/zenodo.3884616" httpUri="https://zenodo.org/record/3884616/files/figure.png" pageId="9" pageNumber="10">Fig. 2D</figureCitation>
|
||
). Prominent postpronotal seta 1. Epandrium not extended at anteroventral corner (ch.30-1;
|
||
<figureCitation id="135AC185EB2F5753FB4BDC20FAE6FEE9" box="[1202,1293,296,323]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="9" pageNumber="10">Figs 3D</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FAE1DC21FAD0FEE9" box="[1304,1339,297,323]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="9" pageNumber="10">4B</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FABFDC20FA80FEE8" box="[1350,1387,296,322]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen & Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="9" pageNumber="10">6D</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FA8FDC21FA7CFEE9" box="[1398,1431,297,323]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="9" pageNumber="10">7E</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FF6EDC45FF50FECD" box="[151,187,333,359]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="9" pageNumber="10">8G</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FF31DC45FF02FECD" box="[200,233,333,359]" captionStart="FIGURE 9" captionStartId="32.[151,250,1936,1961]" captionTargetBox="[166,1420,633,1909]" captionTargetId="figure@32.[166,1420,633,1910]" captionTargetPageId="32" captionText="FIGURE 9. Phortica (Ashima) takehiroi Toda, sp. nov. (♂ paratype). A, Head (caudal view); B, antenna; C, palpus and prob- ocsis; D, prementum; E, abdominal sternites; F, periphallic organs (caudolateral view); G–I, phallic organs (lateral, ventrolateral and ventral views, respectively). Scale bars: 0.1 mm in B–D, F–I; 0.5 mm in A, E." figureDoi="http://doi.org/10.5281/zenodo.3884632" httpUri="https://zenodo.org/record/3884632/files/figure.png" pageId="9" pageNumber="10">9F</figureCitation>
|
||
). Surstylus completely or largely covered with epandrium (ch.36-1;
|
||
<figureCitation id="135AC185EB2F5753FC11DC44FBA8FECD" box="[1000,1091,332,359]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="9" pageNumber="10">Figs 4B</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FBB6DC45FB9BFECD" box="[1103,1136,333,359]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="9" pageNumber="10">7E</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FB85DC45FB4AFECD" box="[1148,1185,333,359]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="9" pageNumber="10">8G</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FB54DC45FB26FECD" box="[1197,1229,333,359]" captionStart="FIGURE 9" captionStartId="32.[151,250,1936,1961]" captionTargetBox="[166,1420,633,1909]" captionTargetId="figure@32.[166,1420,633,1910]" captionTargetPageId="32" captionText="FIGURE 9. Phortica (Ashima) takehiroi Toda, sp. nov. (♂ paratype). A, Head (caudal view); B, antenna; C, palpus and prob- ocsis; D, prementum; E, abdominal sternites; F, periphallic organs (caudolateral view); G–I, phallic organs (lateral, ventrolateral and ventral views, respectively). Scale bars: 0.1 mm in B–D, F–I; 0.5 mm in A, E." figureDoi="http://doi.org/10.5281/zenodo.3884632" httpUri="https://zenodo.org/record/3884632/files/figure.png" pageId="9" pageNumber="10">9F</figureCitation>
|
||
), with pubescence (ch.38-0;
|
||
<figureCitation id="135AC185EB2F5753FEFADC78FEB1FE21" box="[259,346,368,395]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="9" pageNumber="10">Figs 3E</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FE9CDC79FE6CFE21" box="[357,391,369,395]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="9" pageNumber="10">4B</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FE6BDC78FE5EFE21" box="[402,437,368,395]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati & Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="9" pageNumber="10">5C</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FE46DC78FE0FFE20" box="[447,484,368,394]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen & Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="9" pageNumber="10">6D</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FE17DC79FDFAFE21" box="[494,529,369,395]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="9" pageNumber="10">7E</figureCitation>
|
||
). Cercus longer than twice of width (ch.39-0;
|
||
<figureCitation id="135AC185EB2F5753FBF2DC78FB94FE21" box="[1035,1151,368,395]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="9" pageNumber="10">Figs 3B,D</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FB73DC79FB47FE21" box="[1162,1196,369,395]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="9" pageNumber="10">4B</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FB4EDC78FB31FE21" box="[1207,1242,368,395]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati & Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="9" pageNumber="10">5C</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FB1DDC78FAE2FE20" box="[1252,1289,368,394]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen & Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="9" pageNumber="10">6D</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FAEADC79FADEFE21" box="[1299,1333,369,395]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="9" pageNumber="10">7E</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FAC6DC79FA8FFE21" box="[1343,1380,369,395]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="9" pageNumber="10">8G</figureCitation>
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||
,
|
||
<figureCitation id="135AC185EB2F5753FA97DC79FA64FE21" box="[1390,1423,369,395]" captionStart="FIGURE 9" captionStartId="32.[151,250,1936,1961]" captionTargetBox="[166,1420,633,1909]" captionTargetId="figure@32.[166,1420,633,1910]" captionTargetPageId="32" captionText="FIGURE 9. Phortica (Ashima) takehiroi Toda, sp. nov. (♂ paratype). A, Head (caudal view); B, antenna; C, palpus and prob- ocsis; D, prementum; E, abdominal sternites; F, periphallic organs (caudolateral view); G–I, phallic organs (lateral, ventrolateral and ventral views, respectively). Scale bars: 0.1 mm in B–D, F–I; 0.5 mm in A, E." figureDoi="http://doi.org/10.5281/zenodo.3884632" httpUri="https://zenodo.org/record/3884632/files/figure.png" pageId="9" pageNumber="10">9F</figureCitation>
|
||
). Pregonite articulated to tip of aedeagal guide (ch.54-1;
|
||
<figureCitation id="135AC185EB2F5753FD06DC9CFCB1FE05" box="[767,858,404,431]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="9" pageNumber="10">Figs 3G</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FC9FDC9DFC6BFE05" box="[870,896,405,431]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="9" pageNumber="10">4I</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FC72DC9CFC5BFE05" box="[907,944,404,431]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati & Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="9" pageNumber="10">5G</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FC42DC9CFC37FE04" box="[955,988,404,430]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen & Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="9" pageNumber="10">6E</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FC11DC9DFBE7FE05" box="[1000,1036,405,431]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="9" pageNumber="10">7G</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FBE1DC9DFBD7FE05" box="[1048,1084,405,431]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="9" pageNumber="10">8D</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FBB1DC9DFB87FE05" box="[1096,1132,405,431]" captionStart="FIGURE 9" captionStartId="32.[151,250,1936,1961]" captionTargetBox="[166,1420,633,1909]" captionTargetId="figure@32.[166,1420,633,1910]" captionTargetPageId="32" captionText="FIGURE 9. Phortica (Ashima) takehiroi Toda, sp. nov. (♂ paratype). A, Head (caudal view); B, antenna; C, palpus and prob- ocsis; D, prementum; E, abdominal sternites; F, periphallic organs (caudolateral view); G–I, phallic organs (lateral, ventrolateral and ventral views, respectively). Scale bars: 0.1 mm in B–D, F–I; 0.5 mm in A, E." figureDoi="http://doi.org/10.5281/zenodo.3884632" httpUri="https://zenodo.org/record/3884632/files/figure.png" pageId="9" pageNumber="10">9C</figureCitation>
|
||
), without sensilla on apical portion (ch.51-0;
|
||
<figureCitation id="135AC185EB2F5753FEA6DCB0FE33FE79" box="[351,472,440,467]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="9" pageNumber="10">Figs 3G,H</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FE1CDCB1FDE3FE79" box="[485,520,441,467]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="9" pageNumber="10">4C</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FDECDCB0FDD1FE79" box="[533,570,440,467]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati & Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="9" pageNumber="10">5D</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FDBEDCB0FD83FE78" box="[583,616,440,466]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen & Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="9" pageNumber="10">6E</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FD8FDCB1FD71FE79" box="[630,666,441,467]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="9" pageNumber="10">7G</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FD51DCB1FD2AFE79" box="[680,705,441,467]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="9" pageNumber="10">8I</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FD36DCB1FD1FFE79" box="[719,756,441,467]" captionStart="FIGURE 9" captionStartId="32.[151,250,1936,1961]" captionTargetBox="[166,1420,633,1909]" captionTargetId="figure@32.[166,1420,633,1910]" captionTargetPageId="32" captionText="FIGURE 9. Phortica (Ashima) takehiroi Toda, sp. nov. (♂ paratype). A, Head (caudal view); B, antenna; C, palpus and prob- ocsis; D, prementum; E, abdominal sternites; F, periphallic organs (caudolateral view); G–I, phallic organs (lateral, ventrolateral and ventral views, respectively). Scale bars: 0.1 mm in B–D, F–I; 0.5 mm in A, E." figureDoi="http://doi.org/10.5281/zenodo.3884632" httpUri="https://zenodo.org/record/3884632/files/figure.png" pageId="9" pageNumber="10">9G</figureCitation>
|
||
). Phallapodeme vertically flat plate, with posterior, rod-like process apically articulated to aedeagal median rod (ch.60,61-1;
|
||
<figureCitation id="135AC185EB2F5753FC88DCD4FC02FE5D" box="[881,1001,476,503]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="9" pageNumber="10">Figs 3G,H</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FC0FDCD5FBF2FE5D" box="[1014,1049,477,503]" captionStart="FIGURE 4" captionStartId="18.[151,250,1804,1829]" captionTargetBox="[151,1436,525,1774]" captionTargetId="figure@18.[151,1436,525,1779]" captionTargetPageId="18" captionText="FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884622" httpUri="https://zenodo.org/record/3884622/files/figure.png" pageId="9" pageNumber="10">4C</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FBDCDCD4FBA1FE5D" box="[1061,1098,476,503]" captionStart="FIGURE 5" captionStartId="21.[151,250,1902,1927]" captionTargetBox="[151,1435,561,1876]" captionTargetId="figure@21.[151,1435,561,1877]" captionTargetPageId="21" captionText="FIGURE 5. Phortica (Ashima) watabei Sati & Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B." figureDoi="http://doi.org/10.5281/zenodo.3884624" httpUri="https://zenodo.org/record/3884624/files/figure.png" pageId="9" pageNumber="10">5D</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FBAFDCD4FB9CFE5C" box="[1110,1143,476,502]" captionStart="FIGURE 6" captionStartId="24.[151,250,1907,1932]" captionTargetBox="[151,1435,417,1882]" captionTargetId="figure@24.[151,1435,417,1882]" captionTargetPageId="24" captionText="FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen & Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C." figureDoi="http://doi.org/10.5281/zenodo.3884626" httpUri="https://zenodo.org/record/3884626/files/figure.png" pageId="9" pageNumber="10">6E</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FB7DDCD5FB43FE5D" box="[1156,1192,477,503]" captionStart="FIGURE 7" captionStartId="25.[151,250,1911,1936]" captionTargetBox="[215,1371,181,1885]" captionTargetId="figure@25.[215,1372,181,1885]" captionTargetPageId="25" captionText="FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D." figureDoi="http://doi.org/10.5281/zenodo.3884628" httpUri="https://zenodo.org/record/3884628/files/figure.png" pageId="9" pageNumber="10">7G</figureCitation>
|
||
,
|
||
<figureCitation id="135AC185EB2F5753FB4CDCD5FB24FE5D" box="[1205,1231,477,503]" captionStart="FIGURE 8" captionStartId="30.[151,250,1887,1912]" captionTargetBox="[163,1424,346,1861]" captionTargetId="figure@30.[163,1424,345,1861]" captionTargetPageId="30" captionText="FIGURE 8. Phortica (Ashima) kerinciensis Toda, sp. nov. (♂ paratype). A, Antenna; B, posterior portion of scutum (dorsal view); C, medial and posterior sensilla on cibarium (dorsal view); D, fore tarsus; E, abdominal sternites; F, habitus (ventral view); G, periphallic organs (caudolateral view); H, aedeagus and postgonites; I–K, phallic organs (lateral, ventral and ventrolateral views, respectively; B-type). Scale bars: 0.1 mm in A, C, D, G–K; 0.5 mm in B, E, F." figureDoi="http://doi.org/10.5281/zenodo.3884630" httpUri="https://zenodo.org/record/3884630/files/figure.png" pageId="9" pageNumber="10">8I</figureCitation>
|
||
). Aedeagal sheath with more or less sclerotized postgonite(s) (ch.64-1;
|
||
<figureCitation id="135AC185EB2F5753FD1FDF08FCC2FDB1" box="[742,809,512,539]" captionStart="FIGURE 3" captionStartId="13.[151,250,1744,1769]" captionTargetBox="[180,1408,181,1717]" captionTargetId="figure@13.[180,1408,181,1718]" captionTargetPageId="13" captionText="FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3884620" httpUri="https://zenodo.org/record/3884620/files/figure.png" pageId="9" pageNumber="10">Figs 3</figureCitation>
|
||
F–H, 4D, 5D,E, 6F,H, 7F, 8I–K, 9G–I).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37B8E8BEB2F5753FF3EDF2DFE55FD01" pageId="9" pageNumber="10" type="key">
|
||
<paragraph id="8BDEDD00EB2F5753FF3EDF2DFE55FD01" blockId="9.[151,1437,152,683]" pageId="9" pageNumber="10">
|
||
<emphasis id="B9150112EB2F5753FF3EDF2DFF15FD95" bold="true" box="[199,254,549,575]" pageId="9" pageNumber="10">Key.</emphasis>
|
||
The digital, multiple-entry key “
|
||
<taxonomicName id="4C61A683EB2F5753FD88DF2CFD20FD94" ID-CoL="89RX5" authorityName="Chen" authorityYear="2011" box="[625,715,548,574]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="42" phylum="Arthropoda" rank="subGenus" subGenus="Ashima">Ashima</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C61A683EB2F5753FD22DF2CFCDEFD94" authorityName="Linnaeus" authorityYear="1758" box="[731,821,548,574]" kingdom="Plantae" order="Diptera" pageId="9" pageNumber="10" rank="order">Diptera</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C61A683EB2F5753FCB9DF2CFC08FD94" ID-CoL="9GR" authorityName="Rondani" authorityYear="1856" box="[832,995,548,574]" class="Insecta" family="Drosophilidae" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="family">Drosophilidae</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C61A683EB2F5753FC17DF2CFBBBFD95" ID-CoL="9G2PY" authorityName="Schiner" authorityYear="1862" box="[1006,1104,548,575]" class="Insecta" family="Drosophilidae" genus="Phortica" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Phortica</taxonomicName>
|
||
)” to all species of this subgenus has been constructed largely based on the character matrix of Appendix 1, and its database package including software instructions is available from the following website: https://drive.google.com/open?id=10ht68FhP1DJDg h2T33kZn7I2ro8YGGX0.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |