243 lines
38 KiB
XML
243 lines
38 KiB
XML
<document id="281578929468F1DEB7F2C4C27F897149" ID-CLB-Dataset="31287" ID-DOI="10.11646/zootaxa.4358.2.1" ID-GBIF-Dataset="d226f633-d454-4e32-b12f-6d00c30bddfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="1067996" ID-ZooBank="8861BCC0-022F-4803-98E8-D28B90F666E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511949109352" checkinUser="plazi" docAuthor="Dow, Rory A., Stokvis, Frank & Ngiam, Robin W. J." docDate="2017" docId="03C3A90C987AFFA0FF6A8DB4FA0EE9A5" docLanguage="en" docName="zootaxa.4358.2.1.pdf.imf" docOrigin="Zootaxa 4358 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptogomphus Dow & Stokvis & Ngiam 2017" docType="treatment" docVersion="8" lastPageNumber="243" masterDocId="FFFAD1749852FF8AFFFD8823FF9DEF7A" masterDocTitle="Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)" masterLastPageNumber="257" masterPageNumber="201" pageNumber="241" updateTime="1698505057438" updateUser="ExternalLinkService">
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<mods:title id="F83C2A2C54CF21F681FF6AAB9C972486">Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)</mods:title>
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<mods:namePart id="697B7F592511C74C4C4641A270624771">Dow, Rory A.</mods:namePart>
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<mods:namePart id="917C191373F74E4C7A814B0FD8FE72C2">Stokvis, Frank</mods:namePart>
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<mods:namePart id="9566866778DFD3EC14FC256799184913">Ngiam, Robin W. J.</mods:namePart>
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<mods:title id="5772153466E754B46E5E58C529D43CF1">Zootaxa</mods:title>
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<mods:part id="4D8D6E0DD29D282C2617974555F34222">
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<mods:date id="D56D13521CE639A9A200EB17E7DE2CB1">2017</mods:date>
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<mods:number id="80E046C8A1E8D733A23933AC056DB06E">2017-11-29</mods:number>
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<mods:number id="DBCD89661EA8BEB316E371A8A74143AE">4358</mods:number>
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<treatment id="03C3A90C987AFFA0FF6A8DB4FA0EE9A5" ID-DOI="http://doi.org/10.5281/zenodo.5631262" ID-GBIF-Taxon="137580769" ID-Zenodo-Dep="5631262" LSID="urn:lsid:plazi:treatment:03C3A90C987AFFA0FF6A8DB4FA0EE9A5" httpUri="http://treatment.plazi.org/id/03C3A90C987AFFA0FF6A8DB4FA0EE9A5" lastPageId="42" lastPageNumber="243" pageId="40" pageNumber="241">
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<subSubSection id="C3704B91987AFFA2FF6A8DB4FDC7EAA8" pageId="40" pageNumber="241" type="nomenclature">
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<paragraph id="8BD5181A987AFFA2FF6A8DB4FE3AEACB" blockId="40.[151,602,1431,1490]" box="[151,423,1431,1457]" pageId="40" pageNumber="241">
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<heading id="D09DAF76987AFFA2FF6A8DB4FE3AEACB" bold="true" box="[151,423,1431,1457]" fontSize="11" level="1" pageId="40" pageNumber="241" reason="1">
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<emphasis id="B91EC408987AFFA2FF6A8DB4FE3AEACB" bold="true" box="[151,423,1431,1457]" pageId="40" pageNumber="241">
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<taxonomicName id="4C6A6399987AFFA2FF6A8DB4FED7EACB" ID-CoL="5D3K" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[151,330,1431,1457]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="40" pageNumber="241" phylum="Arthropoda" rank="genus">
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<emphasis id="B91EC408987AFFA2FF6A8DB4FED7EACB" bold="true" box="[151,330,1431,1457]" italics="true" pageId="40" pageNumber="241">Leptogomphus</emphasis>
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</taxonomicName>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD5181A987AFFA2FF6A8D9AFDC7EAA8" blockId="40.[151,602,1431,1490]" box="[151,602,1465,1490]" pageId="40" pageNumber="241">
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(
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<figureCitation id="1351049F987AFFA2FF628D9AFF78EAA8" box="[159,229,1465,1490]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="40" pageNumber="241">Figs 1</figureCitation>
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,
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<figureCitation id="1351049F987AFFA2FF0E8D99FE9EEAA8" box="[243,259,1466,1490]" captionStart="FIGURE 2" captionStartId="5.[151,250,1294,1316]" captionTargetBox="[243,1384,200,1237]" captionTargetId="figure@5.[151,1436,193,1273]" captionTargetPageId="5" captionText="FIGURE 2. ITS gene tree for 20 specimens of Leptogomphus and three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068000/files/figure.png" pageId="40" pageNumber="241">2</figureCitation>
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,
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<figureCitation id="1351049F987AFFA2FEF28D99FE82EAA8" box="[271,287,1466,1490]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="40" pageNumber="241">3</figureCitation>
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,
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<figureCitation id="1351049F987AFFA2FED68D99FED4EAA8" box="[299,329,1466,1490]" captionStart="FIGURES18–23" captionStartId="18.[151,264,1895,1918]" captionTargetBox="[183,1420,198,1847]" captionTargetId="figure@18.[183,1420,198,1847]" captionTargetPageId="18" captionText="FIGURES18–23.Female headdorsal:(18) L.coomansi SAR11_12_GOM48;(19)L.mariaeholotype;(20) L.sp.cf coomansi PomS260; (21) L. pasia holotype; (22) L.sii paratype SAR13_14_GOM44; (23) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068010/files/figure.png" pageId="40" pageNumber="241">23</figureCitation>
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,
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<figureCitation id="1351049F987AFFA2FEA88D99FEEEEAA8" box="[341,371,1466,1490]" captionStart="FIGURES28–31" captionStartId="22.[151,264,1357,1380]" captionTargetBox="[151,1435,266,1336]" captionTargetId="figure@22.[151,1436,265,1336]" captionTargetPageId="22" captionText="FIGURES28–31. Female head dorsaldetail; h—occipital horn; ot—occipital tube; p—pit; s—occipital spur; t—tubercle;thtubercle horn: (28) L. coomansi SAR11_12_GOM48; (29) L. mariae holotype; (30) L. sp. cf coomansi PomS260; (31) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068014/files/figure.png" pageId="40" pageNumber="241">31</figureCitation>
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,
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<figureCitation id="1351049F987AFFA2FE828D99FE00EAA8" box="[383,413,1466,1490]" captionStart="FIGURES 36–41" captionStartId="25.[151,264,1973,1996]" captionTargetBox="[204,1387,183,1958]" captionTargetId="figure@25.[204,1387,183,1958]" captionTargetPageId="25" captionText="FIGURES 36–41. Female head frontal: (36) L. coomansi SAR11_12_GOM48; (37) L. mariae holotype; (38) L. sp. cf coomansi PomS260; (39) L.pasia holotype; (40) L.sii paratype SAR13_14_GOM44; (41) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068018/files/figure.png" pageId="40" pageNumber="241">41</figureCitation>
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,
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<figureCitation id="1351049F987AFFA2FE548D99FE5AEAA8" box="[425,455,1466,1490]" captionStart="FIGURES 46–51" captionStartId="29.[151,264,1672,1695]" captionTargetBox="[156,1432,375,1651]" captionTargetId="figure@29.[156,1432,375,1651]" captionTargetPageId="29" captionText="FIGURES 46–51. Female head frontal detail; h—occipital horn; ot—occipital tube; t—tubercle; th—tubercle horn: (46) L. coomansi SAR11_12_GOM48; (47) L pendleburyi SAR16_GOM8; (48) L. schieli paratype SAR15_GOM6; (49) L. species SAR11_12_GOM33; (50) L. williamsoni SAR13_14_GOM51; (51) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068022/files/figure.png" pageId="40" pageNumber="241">49</figureCitation>
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,
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<figureCitation id="1351049F987AFFA2FE2E8D99FE6FEAA8" box="[467,498,1466,1490]" captionStart="FIGURES 68–73" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[156,1432,193,1894]" captionTargetId="figure@36.[156,1432,193,1894]" captionTargetPageId="36" captionText="FIGURES 68–73. Female synthorax dorsal: (68) L. coomansi SAR11_12_GOM48; (69) L. mariae holotype; (70) L. sp. cf coomansi PomS260; (71) L.pasia holotype; (72) L.sii paratype SAR13_14_GOM44; (73) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068032/files/figure.png" pageId="40" pageNumber="241">73</figureCitation>
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,
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<figureCitation id="1351049F987AFFA2FE038D99FD81EAA8" box="[510,540,1466,1490]" captionStart="FIGURES 78–83" captionStartId="38.[151,264,1952,1975]" captionTargetBox="[169,1406,182,1934]" captionTargetId="figure@38.[169,1406,182,1934]" captionTargetPageId="38" captionText="FIGURES 78–83. Female synthorax lateral: (78) L. coomansi SAR11_12_GOM48; (79) L. mariae holotype, right hand side, flipped for consistency; (80) L. sp. cf coomansi PomS260; (81) L. pasia holotype; (82) L. sii paratype SAR13_14_GOM44; (83) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068036/files/figure.png" pageId="40" pageNumber="241">83</figureCitation>
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,
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<figureCitation id="1351049F987AFFA2FDDA8D99FDCFEAA8" box="[551,594,1466,1490]" captionStart="FIGURE128" captionStartId="50.[151,250,1610,1632]" captionTargetBox="[151,1436,193,1589]" captionTargetId="figure@50.[151,1436,193,1589]" captionTargetPageId="50" captionText="FIGURE128. Distributionof L. pendleburyi (yellowcircle), L. schieli (black square) and L. species from Pulong Tau National Park, Sarawak (white square)." httpUri="https://zenodo.org/record/1068058/files/figure.png" pageId="40" pageNumber="241">128</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3704B91987AFFA2FF6A8DDFFA0AE94C" pageId="40" pageNumber="241" type="materials_examined">
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<paragraph id="8BD5181A987AFFA2FF6A8DDFFA0AE94C" blockId="40.[151,1437,1532,2024]" pageId="40" pageNumber="241">
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<emphasis id="B91EC408987AFFA2FF6A8DDFFEF5E96F" bold="true" box="[151,360,1532,1557]" pageId="40" pageNumber="241">Material studied.</emphasis>
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<specimenCount id="9D6CD393987AFFA2FE8F8DDEFE3DE96E" box="[370,416,1533,1557]" pageId="40" pageNumber="241" type="female">1 ♀</specimenCount>
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(
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<collectionCode id="ED7B80DF987AFFA2FE4F8DDEFE71E96F" box="[434,492,1533,1557]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="40" pageNumber="241">SAR</collectionCode>
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11_12_GOM33:
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<collectionCode id="ED7B80DF987AFFA2FD518DDFFC9BE96F" box="[684,774,1532,1557]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="40" pageNumber="241">RMNH</collectionCode>
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.INS.506307), tributary of Pa’Lungan, ca
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<quantity id="4C92B5FF987AFFA2FB1A8DDEFAA3E96F" box="[1255,1342,1533,1557]" metricMagnitude="0" metricUnit="m" metricValue="1.3" pageId="40" pageNumber="241" unit="m" value="1.3">1,300m</quantity>
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, Pulong Tau National Park, Miri Division,
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<collectingCountry id="F37D588A987AFFA2FDEA8E3EFDE7E94C" box="[535,634,1565,1590]" name="Malaysia" pageId="40" pageNumber="241">
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<collectingRegion id="49AED6F8987AFFA2FDEA8E3EFDE7E94C" box="[535,634,1565,1590]" country="Malaysia" name="Sarawak" pageId="40" pageNumber="241">Sarawak</collectingRegion>
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</collectingCountry>
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,
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<collectingCountry id="F37D588A987AFFA2FD7B8E3EFD72E94C" box="[646,751,1565,1590]" name="Malaysia" pageId="40" pageNumber="241">Malaysia</collectingCountry>
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,
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<geoCoordinate id="EE5E7EDD987AFFA2FD078E3DFCF1E94C" box="[762,876,1566,1590]" direction="north" orientation="latitude" pageId="40" pageNumber="241" precision="1" value="3.84912">3.84912N</geoCoordinate>
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,
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<geoCoordinate id="EE5E7EDD987AFFA2FC858E3EFB9CE94C" box="[888,1025,1565,1590]" direction="east" orientation="longitude" pageId="40" pageNumber="241" precision="1" value="115.51678">115.51678E</geoCoordinate>
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,
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<date id="FFD43EDA987AFFA2FBF08E3DFBE7E94C" box="[1037,1146,1565,1590]" pageId="40" pageNumber="241" value="2012-04-09">9 iv 2012</date>
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, leg. Fernado, in
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<collectionCode id="ED7B80DF987AFFA2FAC18E3CFA09E94C" box="[1340,1428,1567,1590]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="40" pageNumber="241">RMNH</collectionCode>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3704B91987AFFA0FF3A8E1CFC6FE97D" lastPageId="42" lastPageNumber="243" pageId="40" pageNumber="241" type="description">
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<paragraph id="8BD5181A987AFFA2FF3A8E1CFA09E85A" blockId="40.[151,1437,1532,2024]" pageId="40" pageNumber="241">
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<emphasis id="B91EC408987AFFA2FF3A8E1CFEC4E922" bold="true" box="[199,345,1599,1624]" pageId="40" pageNumber="241">Description.</emphasis>
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<emphasis id="B91EC408987AFFA2FEA28E1CFE01E922" box="[351,412,1599,1624]" italics="true" pageId="40" pageNumber="241">Head</emphasis>
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(
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<figureCitation id="1351049F987AFFA2FE568E1CFD9FE92D" box="[427,514,1599,1624]" captionStart="FIGURES18–23" captionStartId="18.[151,264,1895,1918]" captionTargetBox="[183,1420,198,1847]" captionTargetId="figure@18.[183,1420,198,1847]" captionTargetPageId="18" captionText="FIGURES18–23.Female headdorsal:(18) L.coomansi SAR11_12_GOM48;(19)L.mariaeholotype;(20) L.sp.cf coomansi PomS260; (21) L. pasia holotype; (22) L.sii paratype SAR13_14_GOM44; (23) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068010/files/figure.png" pageId="40" pageNumber="241">Figs 23</figureCitation>
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,
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<figureCitation id="1351049F987AFFA2FDF38E1CFDB1E92D" box="[526,556,1599,1623]" captionStart="FIGURES28–31" captionStartId="22.[151,264,1357,1380]" captionTargetBox="[151,1435,266,1336]" captionTargetId="figure@22.[151,1436,265,1336]" captionTargetPageId="22" captionText="FIGURES28–31. Female head dorsaldetail; h—occipital horn; ot—occipital tube; p—pit; s—occipital spur; t—tubercle;thtubercle horn: (28) L. coomansi SAR11_12_GOM48; (29) L. mariae holotype; (30) L. sp. cf coomansi PomS260; (31) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068014/files/figure.png" pageId="40" pageNumber="241">31</figureCitation>
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,
|
||
<figureCitation id="1351049F987AFFA2FDC48E1CFDCAE92D" box="[569,599,1599,1623]" captionStart="FIGURES 36–41" captionStartId="25.[151,264,1973,1996]" captionTargetBox="[204,1387,183,1958]" captionTargetId="figure@25.[204,1387,183,1958]" captionTargetPageId="25" captionText="FIGURES 36–41. Female head frontal: (36) L. coomansi SAR11_12_GOM48; (37) L. mariae holotype; (38) L. sp. cf coomansi PomS260; (39) L.pasia holotype; (40) L.sii paratype SAR13_14_GOM44; (41) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068018/files/figure.png" pageId="40" pageNumber="241">41</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F987AFFA2FD9E8E1CFD1EE92D" box="[611,643,1599,1623]" captionStart="FIGURES 46–51" captionStartId="29.[151,264,1672,1695]" captionTargetBox="[156,1432,375,1651]" captionTargetId="figure@29.[156,1432,375,1651]" captionTargetPageId="29" captionText="FIGURES 46–51. Female head frontal detail; h—occipital horn; ot—occipital tube; t—tubercle; th—tubercle horn: (46) L. coomansi SAR11_12_GOM48; (47) L pendleburyi SAR16_GOM8; (48) L. schieli paratype SAR15_GOM6; (49) L. species SAR11_12_GOM33; (50) L. williamsoni SAR13_14_GOM51; (51) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068022/files/figure.png" pageId="40" pageNumber="241">49</figureCitation>
|
||
). Median lobe labium brownish, lateral lobes and hooks pale. Labrum, mandible bases, brown. Clypeus brown, postclypeus with pair of widely spaced, small, poorly defined yellow marks. Frons brown, postfrons with pale transverse stripe, just extending onto upper part antefrons (
|
||
<figureCitation id="1351049F987AFFA2FB108EA2FAA2E9E0" box="[1261,1343,1665,1690]" captionStart="FIGURES 36–41" captionStartId="25.[151,264,1973,1996]" captionTargetBox="[204,1387,183,1958]" captionTargetId="figure@25.[204,1387,183,1958]" captionTargetPageId="25" captionText="FIGURES 36–41. Female head frontal: (36) L. coomansi SAR11_12_GOM48; (37) L. mariae holotype; (38) L. sp. cf coomansi PomS260; (39) L.pasia holotype; (40) L.sii paratype SAR13_14_GOM44; (41) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068018/files/figure.png" pageId="40" pageNumber="241">Fig. 41</figureCitation>
|
||
). Vertex and occiput brown. Pair tubercles behind lateral ocelli, outer parts of these produced into a pair of moderately long, robust, rounded horns (referred to as
|
||
<emphasis id="B91EC408987AFFA2FDB98EE7FD7AE9A7" box="[580,743,1732,1757]" italics="true" pageId="40" pageNumber="241">tubercle horns</emphasis>
|
||
in
|
||
<figureCitation id="1351049F987AFFA2FCED8EE7FCFEE9A7" box="[784,867,1732,1757]" captionStart="FIGURES28–31" captionStartId="22.[151,264,1357,1380]" captionTargetBox="[151,1435,266,1336]" captionTargetId="figure@22.[151,1436,265,1336]" captionTargetPageId="22" captionText="FIGURES28–31. Female head dorsaldetail; h—occipital horn; ot—occipital tube; p—pit; s—occipital spur; t—tubercle;thtubercle horn: (28) L. coomansi SAR11_12_GOM48; (29) L. mariae holotype; (30) L. sp. cf coomansi PomS260; (31) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068014/files/figure.png" pageId="40" pageNumber="241">Fig. 31</figureCitation>
|
||
), rest of vertex smooth; ocelli yellowish. Tubular ridge running along hind margin of each compound eye onto occipital plate where it terminates as very robust, rounded projection (referred to as
|
||
<emphasis id="B91EC408987AFFA2FDEB8F24FD27E85A" box="[534,698,1799,1824]" italics="true" pageId="40" pageNumber="241">occipital tubes</emphasis>
|
||
in
|
||
<figureCitation id="1351049F987AFFA2FD238F24FCAEE865" box="[734,819,1799,1824]" captionStart="FIGURES28–31" captionStartId="22.[151,264,1357,1380]" captionTargetBox="[151,1435,266,1336]" captionTargetId="figure@22.[151,1436,265,1336]" captionTargetPageId="22" captionText="FIGURES28–31. Female head dorsaldetail; h—occipital horn; ot—occipital tube; p—pit; s—occipital spur; t—tubercle;thtubercle horn: (28) L. coomansi SAR11_12_GOM48; (29) L. mariae holotype; (30) L. sp. cf coomansi PomS260; (31) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068014/files/figure.png" pageId="40" pageNumber="241">Figs 31</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F987AFFA2FCC28F24FCC3E865" box="[831,862,1799,1823]" captionStart="FIGURES 46–51" captionStartId="29.[151,264,1672,1695]" captionTargetBox="[156,1432,375,1651]" captionTargetId="figure@29.[156,1432,375,1651]" captionTargetPageId="29" captionText="FIGURES 46–51. Female head frontal detail; h—occipital horn; ot—occipital tube; t—tubercle; th—tubercle horn: (46) L. coomansi SAR11_12_GOM48; (47) L pendleburyi SAR16_GOM8; (48) L. schieli paratype SAR15_GOM6; (49) L. species SAR11_12_GOM33; (50) L. williamsoni SAR13_14_GOM51; (51) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068022/files/figure.png" pageId="40" pageNumber="241">49</figureCitation>
|
||
), occipital plate markedly raised in between these.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A987AFFA3FF3A8F0BFB4FEE84" blockId="40.[151,1437,1532,2024]" lastBlockId="41.[151,1437,151,510]" lastPageId="41" lastPageNumber="242" pageId="40" pageNumber="241">
|
||
<emphasis id="B91EC408987AFFA2FF3A8F0BFE85E83B" box="[199,280,1832,1857]" italics="true" pageId="40" pageNumber="241">Thorax</emphasis>
|
||
. Prothorax mostly brown, anterior pronotal lobe with anterior carina pale, middle pronotal lobe with two large indistinct lateral marks and posterior dorsal mark narrowly divided centrally. Pair of small mounds dorsally on middle pronotal lobe, bearing tiny tubercles. Synthorax brown with pale yellow-green markings as follows (
|
||
<figureCitation id="1351049F987AFFA2FF038FAFFEC7E8DF" box="[254,346,1932,1957]" captionStart="FIGURES 68–73" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[156,1432,193,1894]" captionTargetId="figure@36.[156,1432,193,1894]" captionTargetPageId="36" captionText="FIGURES 68–73. Female synthorax dorsal: (68) L. coomansi SAR11_12_GOM48; (69) L. mariae holotype; (70) L. sp. cf coomansi PomS260; (71) L.pasia holotype; (72) L.sii paratype SAR13_14_GOM44; (73) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068032/files/figure.png" pageId="40" pageNumber="241">Figs 73</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F987AFFA2FE978FAEFE17E8DF" box="[362,394,1933,1957]" captionStart="FIGURES 78–83" captionStartId="38.[151,264,1952,1975]" captionTargetBox="[169,1406,182,1934]" captionTargetId="figure@38.[169,1406,182,1934]" captionTargetPageId="38" captionText="FIGURES 78–83. Female synthorax lateral: (78) L. coomansi SAR11_12_GOM48; (79) L. mariae holotype, right hand side, flipped for consistency; (80) L. sp. cf coomansi PomS260; (81) L. pasia holotype; (82) L. sii paratype SAR13_14_GOM44; (83) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068036/files/figure.png" pageId="40" pageNumber="241">83</figureCitation>
|
||
): short mesothoracic collar, broadly divided at middorsal carina, joined to narrow dorsal thoracic stripes, slightly tapering toward wing bases, extending beyond level of apex of antealar crest, terminating just short of antealar carina, inner margins parallel. No antehumeral markings but tiny yellow spot at antealar carina in this position. Narrow pale marking along mesopleural suture, probably due to postmortem discoloration. Stripe running most of length of mesepimeron. Metepisternum with indistinct pale area below spiracle, small yellow mark near antealar carina. Broad stripe occupying much of metepimeron. Mesokatepisternum, metakatepisternum obscurely brown and pale. Venter pale. Legs (only middle leg still attached on left, left anterior leg detached at trochanter but still present in bag, left posterior leg used for DNA extraction) robust and relatively short. Pale brown, likely much darker when mature, tarsal claws pale basally, tips brown, teeth black. Tarsus of left anterior leg with only three basal spines, claws short without tooth, tarsus of right posterior leg with few spines, claws short, lacking teeth; tibia of same leg abnormally short. Wings: sectors of arculus separated at origin with 5 cross veins before first bifurcation of superior sector in Fw, 3 (right) or 4 (left) in Hw. Discoidal field with 2 rows of cells from origin, transitioning to 3 well before nodus in all wings. 15 (left) or 16 (right) Ax in Fw, 11 (left) or 13 (right) in Hw, 13 Px in Fw, 14 (left) or 13 (right) in Hw. Pt white, covering ca 4 underlying cells.
|
||
</paragraph>
|
||
<caption id="DF154892987BFFA3FF6A8FAFFE27E89A" httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="41" pageNumber="242" targetBox="[156,1432,529,1911]" targetPageId="41">
|
||
<paragraph id="8BD5181A987BFFA3FF6A8FAFFE27E89A" blockId="41.[151,1437,1932,2016]" pageId="41" pageNumber="242">
|
||
<emphasis id="B91EC408987BFFA3FF6A8FAFFEC6E8DB" bold="true" box="[151,347,1932,1955]" pageId="41" pageNumber="242">FIGURES 94–99.</emphasis>
|
||
Penis in lateral view (all scale bars 0.5mm): (94)
|
||
<taxonomicName id="4C6A6399987BFFA3FC5D8FAEFBBEE8D8" authorityName="Laidlaw" authorityYear="1936" box="[928,1059,1933,1954]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408987BFFA3FC5D8FAEFBBEE8D8" box="[928,1059,1933,1954]" italics="true" pageId="41" pageNumber="242">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
SAR07_8_GOM18; (95)
|
||
<taxonomicName id="4C6A6399987BFFA3FABE8FAEFA01E8D8" authorityName="van Tol" authorityYear="1990" box="[1347,1436,1933,1954]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="pasia">
|
||
<emphasis id="B91EC408987BFFA3FABE8FAEFA01E8D8" box="[1347,1436,1933,1954]" italics="true" pageId="41" pageNumber="242">L. pasia</emphasis>
|
||
</taxonomicName>
|
||
SAB12_GOM1; (96)
|
||
<taxonomicName id="4C6A6399987BFFA3FE8E8F8FFD9AE8BB" authorityName="Steinhoff" authorityYear="2015" box="[371,519,1963,1985]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="pendleburyi">
|
||
<emphasis id="B91EC408987BFFA3FE8E8F8FFD9AE8BB" box="[371,519,1963,1985]" italics="true" pageId="41" pageNumber="242">L. pendleburyi</emphasis>
|
||
</taxonomicName>
|
||
SAB12_GOM3; (97)
|
||
<taxonomicName id="4C6A6399987BFFA3FD148F8FFCDBE8BB" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[745,838,1963,1985]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="schieli">
|
||
<emphasis id="B91EC408987BFFA3FD148F8FFCDBE8BB" box="[745,838,1963,1985]" italics="true" pageId="41" pageNumber="242">L. schieli</emphasis>
|
||
</taxonomicName>
|
||
holotype; (98)
|
||
<taxonomicName id="4C6A6399987BFFA3FC1F8F8FFB88E8BB" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[994,1045,1964,1985]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="sii">
|
||
<emphasis id="B91EC408987BFFA3FC1F8F8FFB88E8BB" box="[994,1045,1964,1985]" italics="true" pageId="41" pageNumber="242">L. sii</emphasis>
|
||
</taxonomicName>
|
||
paratype SAR13_14_GOM49; (99)
|
||
<taxonomicName id="4C6A6399987BFFA3FA7A8F8FFE90E89A" authorityName="Laidlaw" authorityYear="1912" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="41" pageNumber="242" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC408987BFFA3FA7A8F8FFE90E89A" italics="true" pageId="41" pageNumber="242">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
SAR16_GOM1.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1548929878FFA0FF6A8C9CFE76EA68" httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="42" pageNumber="243" targetBox="[163,1392,207,1164]" targetPageId="42">
|
||
<paragraph id="8BD5181A9878FFA0FF6A8C9CFE76EA68" blockId="42.[151,1437,1215,1298]" pageId="42" pageNumber="243">
|
||
<emphasis id="B91EC4089878FFA0FF6A8C9CFEF3EBAE" bold="true" box="[151,366,1215,1238]" pageId="42" pageNumber="243">FIGURES 100–105.</emphasis>
|
||
Male anal appendages dorsal view (all scale bars 0.5mm): (100)
|
||
<taxonomicName id="4C6A63999878FFA0FBF38CE2FB17EBAC" authorityName="Laidlaw" authorityYear="1936" box="[1038,1162,1217,1238]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC4089878FFA0FBF38CE2FB17EBAC" box="[1038,1162,1217,1238]" italics="true" pageId="42" pageNumber="243">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
SAR09_10_GOM4; (101)
|
||
<taxonomicName id="4C6A63999878FFA0FF6A8CFCFF77EB8E" authorityName="van Tol" authorityYear="1990" box="[151,234,1247,1268]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="pasia">
|
||
<emphasis id="B91EC4089878FFA0FF6A8CFCFF77EB8E" box="[151,234,1247,1268]" italics="true" pageId="42" pageNumber="243">L. pasia</emphasis>
|
||
</taxonomicName>
|
||
SAB12_GOM1; (102)
|
||
<taxonomicName id="4C6A63999878FFA0FE238CFCFDEEEB8E" authorityName="Steinhoff" authorityYear="2015" box="[478,627,1246,1268]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="pendleburyi">
|
||
<emphasis id="B91EC4089878FFA0FE238CFCFDEEEB8E" box="[478,627,1246,1268]" italics="true" pageId="42" pageNumber="243">L. pendleburyi</emphasis>
|
||
</taxonomicName>
|
||
SAR13_14_GOM13; (103)
|
||
<taxonomicName id="4C6A63999878FFA0FC678CFCFC65EB8E" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[922,1016,1246,1268]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="schieli">
|
||
<emphasis id="B91EC4089878FFA0FC678CFCFC65EB8E" box="[922,1016,1246,1268]" italics="true" pageId="42" pageNumber="243">L. schieli</emphasis>
|
||
</taxonomicName>
|
||
holotype; (104)
|
||
<taxonomicName id="4C6A63999878FFA0FB588CFCFB47EB8E" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[1189,1242,1247,1268]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="sii">
|
||
<emphasis id="B91EC4089878FFA0FB588CFCFB47EB8E" box="[1189,1242,1247,1268]" italics="true" pageId="42" pageNumber="243">L. sii</emphasis>
|
||
</taxonomicName>
|
||
holotype; (105)
|
||
<taxonomicName id="4C6A63999878FFA0FA7A8CFCFE90EA68" authorityName="Laidlaw" authorityYear="1912" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089878FFA0FA7A8CFCFE90EA68" italics="true" pageId="42" pageNumber="243">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
SAR11_12_GOM56.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD5181A9878FFA0FF3A8D19FE2BEA99" blockId="42.[151,1437,1338,1759]" pageId="42" pageNumber="243">
|
||
<emphasis id="B91EC4089878FFA0FF3A8D19FEACEA29" box="[199,305,1338,1363]" italics="true" pageId="42" pageNumber="243">Abdomen</emphasis>
|
||
. Compressed due to teneral condition. Dark brown with pale markings as follows: S1 with lateral mark and dorsal mark. S2 faint pale lateral stripe running from small auricle to apex. Lateral basal marks on S3–6, becoming smaller on successive segments. S2–7 with narrow middorsal line, widest on S2. S8–10 entirely dark. Cerci simple conical, tapering to sharp tip, longer than S10. Vulvar scale distorted but at least slightly longer than S9, not fit for illustration.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9878FFA0FF3A8DCCFC6FE97D" blockId="42.[151,1437,1338,1759]" box="[199,1010,1518,1543]" pageId="42" pageNumber="243">Measurements (mm). Hw 39, abdomen excluding anal appendages 39.5.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B919878FFA0FF3A8E31FA0EE9A5" pageId="42" pageNumber="243" type="discussion">
|
||
<paragraph id="8BD5181A9878FFA0FF3A8E31FB20E9ED" blockId="42.[151,1437,1338,1759]" pageId="42" pageNumber="243">
|
||
<emphasis id="B91EC4089878FFA0FF3A8E31FEA6E951" bold="true" box="[199,315,1554,1579]" pageId="42" pageNumber="243">Remarks.</emphasis>
|
||
This species is remarkable for its head structures and for the apparent reduction of pale markings on the metepisternum. It is clearly distinct from all known species morphologically, and in the two molecular markers examined (see
|
||
<figureCitation id="1351049F9878FFA0FEBD8E79FE1BE909" box="[320,390,1626,1651]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="42" pageNumber="243">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9878FFA0FE688E78FE38E909" box="[405,421,1627,1651]" captionStart="FIGURE 2" captionStartId="5.[151,250,1294,1316]" captionTargetBox="[243,1384,200,1237]" captionTargetId="figure@5.[151,1436,193,1273]" captionTargetPageId="5" captionText="FIGURE 2. ITS gene tree for 20 specimens of Leptogomphus and three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068000/files/figure.png" pageId="42" pageNumber="243">2</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9878FFA0FE4C8E78FE5DE909" box="[433,448,1627,1651]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="42" pageNumber="243">3</figureCitation>
|
||
), however we will wait for a male specimen to become available before naming it. The peculiarities noted for the left anterior and right posterior legs appear to be merely deformities.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9878FFA0FF3A8E80FA0EE9A5" blockId="42.[151,1437,1338,1759]" pageId="42" pageNumber="243">
|
||
Our molecular analyses suggest that this species is most closely related to
|
||
<taxonomicName id="4C6A63999878FFA0FBCA8E80FB42E9C1" authorityName="Steinhoff" authorityYear="2015" box="[1079,1247,1698,1723]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="42" pageNumber="243" phylum="Arthropoda" rank="species" species="pendleburyi">
|
||
<emphasis id="B91EC4089878FFA0FBCA8E80FB42E9C1" box="[1079,1247,1698,1723]" italics="true" pageId="42" pageNumber="243">L. pendleburyi</emphasis>
|
||
</taxonomicName>
|
||
. At present this species is known from a single location in Pulong Tau National Park in the Tama Abu Range,
|
||
<collectingRegion id="49AED6F89878FFA0FB488EE5FA88E9A5" box="[1205,1301,1734,1759]" country="Malaysia" name="Sarawak" pageId="42" pageNumber="243">
|
||
<collectingCountry id="F37D588A9878FFA0FB488EE5FA88E9A5" box="[1205,1301,1734,1759]" name="Malaysia" pageId="42" pageNumber="243">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
(
|
||
<figureCitation id="1351049F9878FFA0FAD98EE5FA1AE9A5" box="[1316,1415,1734,1759]" captionStart="FIGURE128" captionStartId="50.[151,250,1610,1632]" captionTargetBox="[151,1436,193,1589]" captionTargetId="figure@50.[151,1436,193,1589]" captionTargetPageId="50" captionText="FIGURE128. Distributionof L. pendleburyi (yellowcircle), L. schieli (black square) and L. species from Pulong Tau National Park, Sarawak (white square)." httpUri="https://zenodo.org/record/1068058/files/figure.png" pageId="42" pageNumber="243">Fig. 128</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |