245 lines
32 KiB
XML
245 lines
32 KiB
XML
<document id="91EBFA10F1ABDB0C90839BE99CF3D05D" ID-CLB-Dataset="31287" ID-DOI="10.11646/zootaxa.4358.2.1" ID-GBIF-Dataset="d226f633-d454-4e32-b12f-6d00c30bddfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="1067996" ID-ZooBank="8861BCC0-022F-4803-98E8-D28B90F666E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511949109352" checkinUser="plazi" docAuthor="Dow, Rory A., Stokvis, Frank & Ngiam, Robin W. J." docDate="2017" docId="03C3A90C9877FFADFF6A8DE4FF7BE8C2" docLanguage="en" docName="zootaxa.4358.2.1.pdf.imf" docOrigin="Zootaxa 4358 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptogomphus Dow & Stokvis & Ngiam 2017" docType="treatment" docVersion="8" lastPageNumber="238" masterDocId="FFFAD1749852FF8AFFFD8823FF9DEF7A" masterDocTitle="Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)" masterLastPageNumber="257" masterPageNumber="201" pageNumber="238" updateTime="1698505057438" updateUser="ExternalLinkService">
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<mods:title id="41A2F7C808284CE90EB35091D626F8B5">Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)</mods:title>
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<mods:namePart id="F7397CB19701BCAE9C84493D478759C7">Dow, Rory A.</mods:namePart>
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<mods:namePart id="26427EBE69FBBF94A483E76ED5999240">Stokvis, Frank</mods:namePart>
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<mods:namePart id="FB7F98C2A19D11530E57B06F0A1173D6">Ngiam, Robin W. J.</mods:namePart>
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<mods:title id="55953404B4A52F1CD20487AE7F8A4EA8">Zootaxa</mods:title>
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<mods:part id="41C87D955C040457A1096C66C7F9605D">
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<mods:date id="26768E57FBE5EF0693C9BBB5A170F4B4">2017</mods:date>
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<treatment id="03C3A90C9877FFADFF6A8DE4FF7BE8C2" ID-DOI="http://doi.org/10.5281/zenodo.5631260" ID-GBIF-Taxon="137580770" ID-Zenodo-Dep="5631260" LSID="urn:lsid:plazi:treatment:03C3A90C9877FFADFF6A8DE4FF7BE8C2" httpUri="http://treatment.plazi.org/id/03C3A90C9877FFADFF6A8DE4FF7BE8C2" lastPageId="39" lastPageNumber="238" pageId="37" pageNumber="238">
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<subSubSection id="C3704B919877FFAFFF6A8DE4FDA2E979" pageId="37" pageNumber="238" type="nomenclature">
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<paragraph id="8BD5181A9877FFAFFF6A8DE4FD46EA9B" blockId="37.[151,731,1479,1539]" box="[151,731,1479,1505]" pageId="37" pageNumber="238">
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<heading id="D09DAF769877FFAFFF6A8DE4FD46EA9B" bold="true" box="[151,731,1479,1505]" fontSize="11" level="1" pageId="37" pageNumber="238" reason="1">
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<emphasis id="B91EC4089877FFAFFF6A8DE4FD46EA9B" bold="true" box="[151,731,1479,1505]" pageId="37" pageNumber="238">
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<taxonomicName id="4C6A63999877FFAFFF6A8DE4FED7EA9B" ID-CoL="5D3K" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[151,330,1479,1505]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="238" phylum="Arthropoda" rank="genus">
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<emphasis id="B91EC4089877FFAFFF6A8DE4FED7EA9B" bold="true" box="[151,330,1479,1505]" italics="true" pageId="37" pageNumber="238">Leptogomphus</emphasis>
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</taxonomicName>
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sp. cf
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<taxonomicName id="4C6A63999877FFAFFE678DE4FD46EA9B" authority="Laidlaw, 1912" authorityName="Laidlaw" authorityYear="1912" box="[410,731,1479,1505]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="williamsoni">
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<emphasis id="B91EC4089877FFAFFE678DE4FDBBEA9B" bold="true" box="[410,550,1479,1505]" italics="true" pageId="37" pageNumber="238">williamsoni</emphasis>
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<bibRefCitation id="EFFB65EB9877FFAFFDD08DE4FD46EA9B" author="Laidlaw" box="[557,731,1479,1505]" pageId="37" pageNumber="238" refString="Laidlaw, F. F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. (Appendix VIII to Moulton, J. C., 1912.) Journal of the Straits Branch of the Royal Asiatic Society, 63, 92 - 99, 1 pl." type="journal article" year="1912">Laidlaw, 1912</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BD5181A9877FFAFFF6A8DC9FDA2E979" blockId="37.[151,731,1479,1539]" box="[151,575,1514,1539]" pageId="37" pageNumber="238">
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(
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<figureCitation id="1351049F9877FFAFFF628DC9FF78E979" box="[159,229,1514,1539]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="37" pageNumber="238">Figs 1</figureCitation>
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,
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<figureCitation id="1351049F9877FFAFFF0E8DC8FE9EE979" box="[243,259,1515,1539]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="37" pageNumber="238">3</figureCitation>
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,
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<figureCitation id="1351049F9877FFAFFEF28DC8FEB0E979" box="[271,301,1515,1539]" captionStart="FIGURES 24–27" captionStartId="20.[151,264,1441,1464]" captionTargetBox="[163,1439,268,1402]" captionTargetId="figure@20.[163,1439,268,1402]" captionTargetPageId="20" captionText="FIGURES 24–27. Female head dorsal: (24) L pendleburyi SAR16_GOM8; (25) L. schieli paratype SAR15_GOM6; (26) L. williamsoni SAR13_14_GOM51; (27) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068012/files/figure.png" pageId="37" pageNumber="238">27</figureCitation>
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,
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<figureCitation id="1351049F9877FFAFFEC48DC9FECAE979" box="[313,343,1514,1539]" captionStart="FIGURES 32–35" captionStartId="23.[151,264,1357,1380]" captionTargetBox="[151,1436,265,1336]" captionTargetId="figure@23.[151,1436,265,1336]" captionTargetPageId="23" captionText="FIGURES 32–35. Female head dorsal detail; h—occipital horn; p—pit; s—occipital spur; t—tubercle: (32) L pendleburyi SAR16_GOM8; (33) L. schieli paratype SAR15_GOM6; (34) L. williamsoni SAR13_14_GOM51; (35) L. sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068016/files/figure.png" pageId="37" pageNumber="238">35</figureCitation>
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,
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<figureCitation id="1351049F9877FFAFFE9E8DC9FE1FE979" box="[355,386,1514,1539]" captionStart="FIGURES 42–45" captionStartId="27.[151,264,1498,1521]" captionTargetBox="[163,1439,186,1462]" captionTargetId="figure@27.[163,1439,186,1462]" captionTargetPageId="27" captionText="FIGURES 42–45. Female head frontal: (42) L pendleburyi SAR16_GOM8; (43) L. schieli paratype SAR15_GOM6; (44) L. williamsoni SAR13_14_GOM51; (45) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068020/files/figure.png" pageId="37" pageNumber="238">45</figureCitation>
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,
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<figureCitation id="1351049F9877FFAFFE708DC9FE36E979" box="[397,427,1514,1539]" captionStart="FIGURES 46–51" captionStartId="29.[151,264,1672,1695]" captionTargetBox="[156,1432,375,1651]" captionTargetId="figure@29.[156,1432,375,1651]" captionTargetPageId="29" captionText="FIGURES 46–51. Female head frontal detail; h—occipital horn; ot—occipital tube; t—tubercle; th—tubercle horn: (46) L. coomansi SAR11_12_GOM48; (47) L pendleburyi SAR16_GOM8; (48) L. schieli paratype SAR15_GOM6; (49) L. species SAR11_12_GOM33; (50) L. williamsoni SAR13_14_GOM51; (51) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068022/files/figure.png" pageId="37" pageNumber="238">51</figureCitation>
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,
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<figureCitation id="1351049F9877FFAFFE4A8DC8FE48E978" box="[439,469,1515,1538]" captionStart="FIGURES 74–77" captionStartId="37.[151,264,1356,1379]" captionTargetBox="[151,1435,194,1335]" captionTargetId="figure@37.[151,1436,193,1335]" captionTargetPageId="37" captionText="FIGURES 74–77. Female synthorax dorsal: (74) L pendleburyi SAR16_GOM8; (75) L. schieli paratype SAR15_GOM6; (76) L. williamsoni SAR13_14_GOM51; (77) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068034/files/figure.png" pageId="37" pageNumber="238">77</figureCitation>
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,
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<figureCitation id="1351049F9877FFAFFE1C8DC8FE62E979" box="[481,511,1515,1539]" captionStart="FIGURES 84–87" captionStartId="39.[151,264,1427,1450]" captionTargetBox="[151,1435,194,1406]" captionTargetId="figure@39.[151,1436,193,1406]" captionTargetPageId="39" captionText="FIGURES 84–87. Female synthorax lateral: (84) L pendleburyi SAR16_GOM8; (85) L. schieli paratype SAR15_GOM6; (86) L. williamsoni SAR13_14_GOM51, right hand side, flipped for consistency;(87) L. sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068038/files/figure.png" pageId="37" pageNumber="238">87</figureCitation>
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,
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<figureCitation id="1351049F9877FFAFFDF68DC8FDABE979" box="[523,566,1515,1539]" captionStart="FIGURE 129" captionStartId="51.[151,250,1608,1630]" captionTargetBox="[151,1436,193,1587]" captionTargetId="figure@51.[151,1436,193,1587]" captionTargetPageId="51" captionText="FIGURE 129. Distribution of L. williamsoni (black square) and L.sp.cf williamsoni (red triangle)." httpUri="https://zenodo.org/record/1068060/files/figure.png" pageId="37" pageNumber="238">129</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3704B919877FFAFFF6A8E0EFB02E9FA" pageId="37" pageNumber="238" type="reference_group">
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<paragraph id="8BD5181A9877FFAFFF6A8E0EFD28E918" blockId="37.[151,1437,1581,1664]" pageId="37" pageNumber="238">
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<treatmentCitationGroup id="AB7A3F349877FFAFFF6A8E0EFD28E918" pageId="37" pageNumber="238">
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<materialsCitation id="3B0212479877FFAFFF6A8E0EFA9AE939" ID-GBIF-Occurrence="1702634993" box="[151,1287,1581,1603]" country="Malaysia" location="Gunung Penrissen" municipality="Annah Rais" pageId="37" pageNumber="238" specimenCount="1" specimenCount-female="1">
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<taxonomicName id="4C6A63999877FFAFFF6A8E0EFDA1E939" ID-CoL="3TQ4T" authority="Laidlaw 1912" authorityName="Laidlaw" authorityYear="1912" box="[151,572,1581,1603]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="williamsoni">
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<emphasis id="B91EC4089877FFAFFF6A8E0EFE34E939" box="[151,425,1581,1603]" italics="true" pageId="37" pageNumber="238">Leptogomphus williamsoni</emphasis>
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<bibRefCitation id="EFFB65EB9877FFAFFE528E0EFDA1E939" author="Laidlaw" box="[431,572,1581,1603]" pageId="37" pageNumber="238" refString="Laidlaw, F. F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. (Appendix VIII to Moulton, J. C., 1912.) Journal of the Straits Branch of the Royal Asiatic Society, 63, 92 - 99, 1 pl." type="journal article" year="1912">Laidlaw 1912</bibRefCitation>
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</taxonomicName>
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||
;—Dow 2012: 7, 10 (
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<specimenCount id="9D6CD3939877FFAFFCE58E0EFCB7E939" box="[792,810,1581,1603]" pageId="37" pageNumber="238" type="female">♀</specimenCount>
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,
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||
<location id="8EB54EC19877FFAFFCCB8E0EFC6DE939" LSID="urn:lsid:plazi:treatment:03C3A90C9877FFADFF6A8DE4FF7BE8C2:8EB54EC19877FFAFFCCB8E0EFC6DE939" box="[822,1008,1581,1603]" country="Malaysia" municipality="Annah Rais" name="Gunung Penrissen" pageId="37" pageNumber="238">Gunung Penrissen</location>
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||
and
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<collectingMunicipality id="6BB182609877FFAFFBDF8E0EFB06E939" box="[1058,1179,1581,1603]" pageId="37" pageNumber="238">Annah Rais</collectingMunicipality>
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,
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||
<collectingCountry id="F37D588A9877FFAFFB5B8E0EFA9DE939" box="[1190,1280,1581,1603]" name="Malaysia" pageId="37" pageNumber="238">Sarawak</collectingCountry>
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)
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||
</materialsCitation>
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||
;—
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||
<materialsCitation id="3B0212479877FFAFFADB8E0EFD2CE918" ID-GBIF-Occurrence="1702634992" collectorName="Kubah NP" country="Malaysia" location="Sarawak" pageId="37" pageNumber="238" specimenCount="1">
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?
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||
<treatmentCitation id="0ACB3E0B9877FFAFFAD28E0EFEB2E918" author="Dow" page="19" pageId="37" pageNumber="238" year="2013">
|
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<bibRefCitation id="EFFB65EB9877FFAFFAD28E0EFE97E918" author="Dow" pageId="37" pageNumber="238" refString="Dow, R. A., Reels, G. T. & Butler, S. G. (2013 b) Previously unpublished Odonata records from Sarawak, Borneo. Part II. Kubah National Park. Faunistic Studies in South-East Asian and Pacific Island Odonata, 6, 1 - 21." type="journal article" year="2013" yearSuffix="b">
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||
Dow
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||
<emphasis id="B91EC4089877FFAFFA948E0DFA00E939" box="[1385,1437,1581,1603]" italics="true" pageId="37" pageNumber="238">et al.</emphasis>
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2013b
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</bibRefCitation>
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||
: 19
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||
</treatmentCitation>
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,
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<figureCitation id="1351049F9877FFAFFEC48E6FFE1FE918" box="[313,386,1612,1634]" captionStart="FIGURES18–23" captionStartId="18.[151,264,1895,1918]" captionTargetBox="[183,1420,198,1847]" captionTargetId="figure@18.[183,1420,198,1847]" captionTargetPageId="18" captionText="FIGURES18–23.Female headdorsal:(18) L.coomansi SAR11_12_GOM48;(19)L.mariaeholotype;(20) L.sp.cf coomansi PomS260; (21) L. pasia holotype; (22) L.sii paratype SAR13_14_GOM44; (23) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068010/files/figure.png" pageId="37" pageNumber="238">Fig. 20</figureCitation>
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(larvae,
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<collectorName id="269F7DCC9877FFAFFE278E6FFDD9E918" box="[474,580,1612,1634]" pageId="37" pageNumber="238">Kubah NP</collectorName>
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||
,
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||
<collectingCountry id="F37D588A9877FFAFFDB38E6FFD37E918" box="[590,682,1612,1634]" name="Malaysia" pageId="37" pageNumber="238">Sarawak</collectingCountry>
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)
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||
</materialsCitation>
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.
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||
</treatmentCitationGroup>
|
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</paragraph>
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<paragraph id="8BD5181A9877FFAFFF6A8E49FB02E9FA" blockId="37.[151,1437,1581,1664]" box="[151,1183,1642,1664]" pageId="37" pageNumber="238">
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<taxonomicName id="4C6A63999877FFAFFF6A8E49FEB1E9FA" ID-CoL="5D3K" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[151,300,1642,1664]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="238" phylum="Arthropoda" rank="genus">
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<emphasis id="B91EC4089877FFAFFF6A8E49FEB1E9FA" box="[151,300,1642,1664]" italics="true" pageId="37" pageNumber="238">Leptogomphus</emphasis>
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</taxonomicName>
|
||
species;—Dow
|
||
<emphasis id="B91EC4089877FFAFFE2E8E48FD9BE9FA" box="[467,518,1642,1664]" italics="true" pageId="37" pageNumber="238">et al.</emphasis>
|
||
2016: 11 (females and larvae from Gunung Penrissen discussed).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B919877FFAFFF6A8E8FFED4E803" pageId="37" pageNumber="238" type="materials_examined">
|
||
<paragraph id="8BD5181A9877FFAFFF6A8E8FFED4E803" blockId="37.[151,1437,1708,2022]" pageId="37" pageNumber="238">
|
||
<emphasis id="B91EC4089877FFAFFF6A8E8FFDC3E9BF" bold="true" box="[151,606,1708,1733]" pageId="37" pageNumber="238">
|
||
Material studied.
|
||
<collectingRegion id="49AED6F89877FFAFFE938E8FFE47E9BF" box="[366,474,1708,1733]" country="Malaysia" name="Sarawak" pageId="37" pageNumber="238">
|
||
<collectingCountry id="F37D588A9877FFAFFE938E8FFE47E9BF" box="[366,474,1708,1733]" name="Malaysia" pageId="37" pageNumber="238">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37D588A9877FFAFFE1A8E8FFDC5E9BF" box="[487,600,1708,1733]" name="Malaysia" pageId="37" pageNumber="238">Malaysia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D6CD3939877FFAFFD9B8E8EFD0FE9BF" box="[614,658,1709,1733]" pageId="37" pageNumber="238" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED7B80DF9877FFAFFD5F8E8FFD46E9BF" box="[674,731,1708,1733]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="37" pageNumber="238">SAR</collectionCode>
|
||
11_12_GOM59, used in illustrations), boulder stream, Borneo Highlands Resort, Gunung Penrissen, Kuching Division,
|
||
<geoCoordinate id="EE5E7EDD9877FFAFFCDB8EF2FC05E990" box="[806,920,1745,1770]" direction="north" orientation="latitude" pageId="37" pageNumber="238" precision="1" value="1.13641">1.13641N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9877FFAFFC588EF2FBB2E990" box="[933,1071,1745,1770]" direction="east" orientation="longitude" pageId="37" pageNumber="238" precision="1" value="110.21555">110.21555E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA9877FFAFFBC08EF2FB59E993" box="[1085,1220,1745,1770]" pageId="37" pageNumber="238" value="2012-07-24">24 vii 2012</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD3939877FFAFFEAC8ED6FEE6E877" box="[337,379,1781,1805]" pageId="37" pageNumber="238" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED7B80DF9877FFAFFE748ED6FE7CE874" box="[393,481,1781,1806]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="37" pageNumber="238">RMNH</collectionCode>
|
||
.5008436; teneral), same location,
|
||
<date id="FFD43EDA9877FFAFFC9F8ED6FC41E877" box="[866,988,1781,1806]" pageId="37" pageNumber="238" value="2014-06-29">29 vi 2014</date>
|
||
, leg. R.A. Dow, in
|
||
<collectionCode id="ED7B80DF9877FFAFFB4C8ED5FA94E877" box="[1201,1289,1782,1805]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="37" pageNumber="238">RMNH</collectionCode>
|
||
;
|
||
<specimenCount id="9D6CD3939877FFAFFAEE8ED6FAA0E877" box="[1299,1341,1781,1805]" pageId="37" pageNumber="238" type="female">1 ♀</specimenCount>
|
||
, stream,
|
||
<geoCoordinate id="EE5E7EDD9877FFAFFF6A8F3AFE94E848" box="[151,265,1817,1842]" direction="north" orientation="latitude" pageId="37" pageNumber="238" precision="1" value="1.12157">1.12157N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9877FFAFFEE88F3AFE03E84B" box="[277,414,1817,1841]" direction="east" orientation="longitude" pageId="37" pageNumber="238" precision="1" value="110.22701">110.22701E</geoCoordinate>
|
||
, same area,
|
||
<date id="FFD43EDA9877FFAFFDD48F3AFD30E84B" box="[553,685,1817,1842]" pageId="37" pageNumber="238" value="2012-07-26">26 vii 2012</date>
|
||
, leg, R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD3939877FFAFFBBF8F3AFBF0E84B" box="[1090,1133,1817,1841]" pageId="37" pageNumber="238" type="female">1 ♀</specimenCount>
|
||
, small high gradient forest stream on Mount Matang, Matang Range, Kuching Division,
|
||
<geoCoordinate id="EE5E7EDD9877FFAFFCBA8F1EFC24E82C" box="[839,953,1853,1878]" direction="north" orientation="latitude" pageId="37" pageNumber="238" precision="1" value="1.59632">1.59632N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD9877FFAFFC398F1EFBD3E82F" box="[964,1102,1853,1877]" direction="east" orientation="longitude" pageId="37" pageNumber="238" precision="1" value="110.20949">110.20949E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA9877FFAFFBA48F1EFB53E82F" box="[1113,1230,1853,1878]" pageId="37" pageNumber="238" value="2017-07-02">2 vii 2017</date>
|
||
, leg. R.A. Dow, in collection Dow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B919877FFADFF3A8FA6FF7BE8C2" lastPageId="39" lastPageNumber="240" pageId="37" pageNumber="238" type="discussion">
|
||
<paragraph id="8BD5181A9877FFADFF3A8FA6FF7BE8C2" blockId="37.[151,1437,1708,2022]" lastBlockId="39.[151,1437,1520,1976]" lastPageId="39" lastPageNumber="240" pageId="37" pageNumber="238">
|
||
<emphasis id="B91EC4089877FFAFFF3A8FA6FEA6E8E4" bold="true" box="[199,315,1925,1950]" pageId="37" pageNumber="238">Remarks.</emphasis>
|
||
Females of
|
||
<taxonomicName id="4C6A63999877FFAFFE258FA6FDE0E8E4" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[472,637,1925,1950]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="238" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EC4089877FFAFFE258FA6FDE0E8E4" box="[472,637,1925,1950]" italics="true" pageId="37" pageNumber="238">Leptogomphus</emphasis>
|
||
</taxonomicName>
|
||
collected on Gunung Penrissen and in the Matang Range but not associated with
|
||
<taxonomicName id="4C6A63999877FFAFFEB48F89FE2DE8B8" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[329,432,1961,1986]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="schieli">
|
||
<emphasis id="B91EC4089877FFAFFEB48F89FE2DE8B8" box="[329,432,1961,1986]" italics="true" pageId="37" pageNumber="238">L. schieli</emphasis>
|
||
</taxonomicName>
|
||
resemble
|
||
<taxonomicName id="4C6A63999877FFAFFDDE8F89FD5EE8B8" authorityName="Laidlaw" authorityYear="1912" box="[547,707,1961,1986]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="238" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089877FFAFFDDE8F89FD5EE8B8" box="[547,707,1961,1986]" italics="true" pageId="37" pageNumber="238">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
, but are atypical in some respects, and may represent yet another species. The chief differences are in the head (
|
||
<figureCitation id="1351049F9877FFAFFD4F8FEEFC94E89F" box="[690,777,1997,2022]" captionStart="FIGURES 24–27" captionStartId="20.[151,264,1441,1464]" captionTargetBox="[163,1439,268,1402]" captionTargetId="figure@20.[163,1439,268,1402]" captionTargetPageId="20" captionText="FIGURES 24–27. Female head dorsal: (24) L pendleburyi SAR16_GOM8; (25) L. schieli paratype SAR15_GOM6; (26) L. williamsoni SAR13_14_GOM51; (27) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068012/files/figure.png" pageId="37" pageNumber="238">Figs 27</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9877FFAFFCE88FEEFCAEE89C" box="[789,819,1997,2022]" captionStart="FIGURES 32–35" captionStartId="23.[151,264,1357,1380]" captionTargetBox="[151,1436,265,1336]" captionTargetId="figure@23.[151,1436,265,1336]" captionTargetPageId="23" captionText="FIGURES 32–35. Female head dorsal detail; h—occipital horn; p—pit; s—occipital spur; t—tubercle: (32) L pendleburyi SAR16_GOM8; (33) L. schieli paratype SAR15_GOM6; (34) L. williamsoni SAR13_14_GOM51; (35) L. sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068016/files/figure.png" pageId="37" pageNumber="238">35</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9877FFAFFCC28FEEFCC3E89C" box="[831,862,1997,2022]" captionStart="FIGURES 42–45" captionStartId="27.[151,264,1498,1521]" captionTargetBox="[163,1439,186,1462]" captionTargetId="figure@27.[163,1439,186,1462]" captionTargetPageId="27" captionText="FIGURES 42–45. Female head frontal: (42) L pendleburyi SAR16_GOM8; (43) L. schieli paratype SAR15_GOM6; (44) L. williamsoni SAR13_14_GOM51; (45) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068020/files/figure.png" pageId="37" pageNumber="238">45</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9877FFAFFC948FEEFC1AE89C" box="[873,903,1997,2022]" captionStart="FIGURES 46–51" captionStartId="29.[151,264,1672,1695]" captionTargetBox="[156,1432,375,1651]" captionTargetId="figure@29.[156,1432,375,1651]" captionTargetPageId="29" captionText="FIGURES 46–51. Female head frontal detail; h—occipital horn; ot—occipital tube; t—tubercle; th—tubercle horn: (46) L. coomansi SAR11_12_GOM48; (47) L pendleburyi SAR16_GOM8; (48) L. schieli paratype SAR15_GOM6; (49) L. species SAR11_12_GOM33; (50) L. williamsoni SAR13_14_GOM51; (51) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068022/files/figure.png" pageId="37" pageNumber="238">51</figureCitation>
|
||
), where the occipital horns are placed close together, and there is an approximately crescentic depression surrounding the tubercles behind the lateral ocelli except at the front. It was only possible to include one specimen of this form in the molecular analysis, and only COI was successfully amplified; the sample is distinct from the other
|
||
<taxonomicName id="4C6A63999875FFADFC538E1AFBCDE92B" authorityName="Laidlaw" authorityYear="1912" box="[942,1104,1592,1617]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089875FFADFC538E1AFBCDE92B" box="[942,1104,1592,1617]" italics="true" pageId="39" pageNumber="240">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
samples (
|
||
<figureCitation id="1351049F9875FFADFB3C8E1BFA94E92A" box="[1217,1289,1592,1617]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="39" pageNumber="240">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9875FFADFAE58E1BFABAE92A" box="[1304,1319,1592,1616]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="39" pageNumber="240">3</figureCitation>
|
||
), but with only one sample and one marker we do not regard this as providing definitive support for the notion that this is a separate species. This is especially so since
|
||
<taxonomicName id="4C6A63999875FFADFD718EA2FCB2E9E3" authorityName="Laidlaw" authorityYear="1912" box="[652,815,1664,1689]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089875FFADFD718EA2FCB2E9E3" box="[652,815,1664,1689]" italics="true" pageId="39" pageNumber="240">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
is rather variable in the COI marker (see the Remarks under that species). In the absence of the male and more molecular data, we have decided to leave the issue of the status of this form open, nor do we provide a complete description; we do however provide a full set of illustrations. If it should prove to be a separate species, the
|
||
<taxonomicName id="4C6A63999875FFADFCDA8ECFFC51E87F" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[807,972,1772,1797]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="39" pageNumber="240" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91EC4089875FFADFCDA8ECFFC51E87F" box="[807,972,1772,1797]" italics="true" pageId="39" pageNumber="240">Leptogomphus</emphasis>
|
||
</taxonomicName>
|
||
larva from Kubah National Park reported in
|
||
<bibRefCitation id="EFFB65EB9875FFADFF498F32FE16E853" box="[180,395,1808,1833]" pageId="39" pageNumber="240" refString="Dow, R. A., Reels, G. T. & Butler, S. G. (2013 b) Previously unpublished Odonata records from Sarawak, Borneo. Part II. Kubah National Park. Faunistic Studies in South-East Asian and Pacific Island Odonata, 6, 1 - 21." type="journal article">
|
||
Dow
|
||
<emphasis id="B91EC4089875FFADFF0F8F32FEB6E853" box="[242,299,1808,1833]" italics="true" pageId="39" pageNumber="240">et al.</emphasis>
|
||
(2013b)
|
||
</bibRefCitation>
|
||
as
|
||
<taxonomicName id="4C6A63999875FFADFE4C8F32FDCFE853" authorityName="Laidlaw" authorityYear="1912" box="[433,594,1808,1833]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="39" pageNumber="240" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089875FFADFE4C8F32FDCFE853" box="[433,594,1808,1833]" italics="true" pageId="39" pageNumber="240">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
probably belongs here, since Kubah is in the Matang Range, where one of the females was collected. A female from Annah Rais at the foot of Gunung Penrissen, previously in the possession of the first author, probably belongs here as well; the specimen was not available to us to check for this study. The measurements of the three non-teneral specimens available are: abdomen without appendages
|
||
<quantity id="4C92B5FF9875FFADFB3F8F5FFAA8E8EF" box="[1218,1333,1916,1941]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="39" pageNumber="240" unit="mm" value="34.5" valueMax="35.0" valueMin="34.0">34–35mm</quantity>
|
||
, Hw
|
||
<quantity id="4C92B5FF9875FFADFA8F8F5FFF7FE8C2" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.0" metricValueMin="2.9" pageId="39" pageNumber="240" unit="mm" value="29.5" valueMax="30.0" valueMin="29.0">29– 30mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |