887 lines
124 KiB
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887 lines
124 KiB
XML
<document id="B3FA6AC25C7FA7D64780719D0FCE5BDE" ID-CLB-Dataset="31287" ID-DOI="10.11646/zootaxa.4358.2.1" ID-GBIF-Dataset="d226f633-d454-4e32-b12f-6d00c30bddfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="1067996" ID-ZooBank="8861BCC0-022F-4803-98E8-D28B90F666E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511949109352" checkinUser="plazi" docAuthor="Dow, Rory A., Stokvis, Frank & Ngiam, Robin W. J." docDate="2017" docId="03C3A90C984FFFA9FF6A8F04FAF5ED1A" docLanguage="en" docName="zootaxa.4358.2.1.pdf.imf" docOrigin="Zootaxa 4358 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptogomphus williamsoni Laidlaw 1912" docType="treatment" docVersion="9" lastPageNumber="236" masterDocId="FFFAD1749852FF8AFFFD8823FF9DEF7A" masterDocTitle="Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)" masterLastPageNumber="257" masterPageNumber="201" pageNumber="230" updateTime="1698505057438" updateUser="ExternalLinkService">
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<mods:title id="8B453E21C3C5E35610E235413EC5B5F9">Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)</mods:title>
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<mods:namePart id="21D908C2C4876B86BF07032C5E9B209E">Dow, Rory A.</mods:namePart>
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<mods:namePart id="5DF7A492A30584A9252C8160208F55BE">Stokvis, Frank</mods:namePart>
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<mods:namePart id="661E7F11D47D8818D9D61CFEE9A0A5C0">Ngiam, Robin W. J.</mods:namePart>
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<mods:date id="6D9D4EE3E525FBA55C5B2919C60DA706">2017</mods:date>
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<treatment id="03C3A90C984FFFA9FF6A8F04FAF5ED1A" ID-DOI="http://doi.org/10.5281/zenodo.5631258" ID-GBIF-Taxon="137580776" ID-Zenodo-Dep="5631258" LSID="urn:lsid:plazi:treatment:03C3A90C984FFFA9FF6A8F04FAF5ED1A" httpUri="http://treatment.plazi.org/id/03C3A90C984FFFA9FF6A8F04FAF5ED1A" lastPageId="35" lastPageNumber="236" pageId="29" pageNumber="230">
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<subSubSection id="C3704B91984FFF97FF6A8F04FB08E819" pageId="29" pageNumber="230" type="nomenclature">
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<paragraph id="8BD5181A984FFF97FF6A8F04FD0EE83B" blockId="29.[151,1173,1831,1891]" box="[151,659,1831,1857]" pageId="29" pageNumber="230">
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<heading id="D09DAF76984FFF97FF6A8F04FD0EE83B" bold="true" box="[151,659,1831,1857]" fontSize="11" level="1" pageId="29" pageNumber="230" reason="1">
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<taxonomicName id="4C6A6399984FFF97FF6A8F04FD0EE83B" ID-CoL="3TQ4T" authority="Laidlaw, 1912" authorityName="Laidlaw" authorityYear="1912" box="[151,659,1831,1857]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="williamsoni">
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<emphasis id="B91EC408984FFF97FF6A8F04FD0EE83B" bold="true" box="[151,659,1831,1857]" pageId="29" pageNumber="230">
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<emphasis id="B91EC408984FFF97FF6A8F04FE40E83B" bold="true" box="[151,477,1831,1857]" italics="true" pageId="29" pageNumber="230">Leptogomphus williamsoni</emphasis>
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<bibRefCitation id="EFFB65EB984FFF97FE198F04FD0EE83B" author="Laidlaw" box="[484,659,1831,1857]" pageId="29" pageNumber="230" refString="Laidlaw, F. F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. (Appendix VIII to Moulton, J. C., 1912.) Journal of the Straits Branch of the Royal Asiatic Society, 63, 92 - 99, 1 pl." type="journal article" year="1912">Laidlaw, 1912</bibRefCitation>
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<paragraph id="8BD5181A984FFF97FF6A8F69FB08E819" blockId="29.[151,1173,1831,1891]" box="[151,1173,1866,1891]" pageId="29" pageNumber="230">
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(
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<figureCitation id="1351049F984FFF97FF628F69FF78E819" box="[159,229,1866,1891]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="29" pageNumber="230">Figs 1</figureCitation>
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<figureCitation id="1351049F984FFF97FF0E8F68FE9EE819" box="[243,259,1867,1891]" captionStart="FIGURE 2" captionStartId="5.[151,250,1294,1316]" captionTargetBox="[243,1384,200,1237]" captionTargetId="figure@5.[151,1436,193,1273]" captionTargetPageId="5" captionText="FIGURE 2. ITS gene tree for 20 specimens of Leptogomphus and three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068000/files/figure.png" pageId="29" pageNumber="230">2</figureCitation>
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<figureCitation id="1351049F984FFF97FEF28F68FE82E819" box="[271,287,1867,1891]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="29" pageNumber="230">3</figureCitation>
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<figureCitation id="1351049F984FFF97FED68F68FEA6E819" box="[299,315,1867,1891]" captionStart="FIGURES 4–9" captionStartId="9.[151,264,1713,1736]" captionTargetBox="[151,1436,193,1692]" captionTargetId="figure@9.[151,1436,193,1692]" captionTargetPageId="9" captionText="FIGURES 4–9. Male head dorsal: (4) L. coomansi SAR09_10_GOM4; (5) L. pendleburyi SAR13_14_GOM13; (6) L. pasia SAB12_GOM1; (7) L. sii holotype; (8) L.schieli holotype; (9) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068004/files/figure.png" pageId="29" pageNumber="230">9</figureCitation>
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<figureCitation id="1351049F984FFF97FEBA8F68FEF8E819" box="[327,357,1867,1891]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1933,1956]" captionTargetBox="[151,1436,841,1912]" captionTargetId="figure@16.[151,1436,841,1912]" captionTargetPageId="16" captionText="FIGURES 14–17. Male head frontal: (14) L. pasia SAB12_GOM1; (15) L. sii holotype; (16) L. schieli holotype; (17) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="29" pageNumber="230">17</figureCitation>
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<figureCitation id="1351049F984FFF97FE8C8F69FE12E819" box="[369,399,1866,1891]" captionStart="FIGURES 24–27" captionStartId="20.[151,264,1441,1464]" captionTargetBox="[163,1439,268,1402]" captionTargetId="figure@20.[163,1439,268,1402]" captionTargetPageId="20" captionText="FIGURES 24–27. Female head dorsal: (24) L pendleburyi SAR16_GOM8; (25) L. schieli paratype SAR15_GOM6; (26) L. williamsoni SAR13_14_GOM51; (27) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068012/files/figure.png" pageId="29" pageNumber="230">26</figureCitation>
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<figureCitation id="1351049F984FFF97FE668F68FE27E819" box="[411,442,1867,1891]" captionStart="FIGURES 32–35" captionStartId="23.[151,264,1357,1380]" captionTargetBox="[151,1436,265,1336]" captionTargetId="figure@23.[151,1436,265,1336]" captionTargetPageId="23" captionText="FIGURES 32–35. Female head dorsal detail; h—occipital horn; p—pit; s—occipital spur; t—tubercle: (32) L pendleburyi SAR16_GOM8; (33) L. schieli paratype SAR15_GOM6; (34) L. williamsoni SAR13_14_GOM51; (35) L. sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068016/files/figure.png" pageId="29" pageNumber="230">34</figureCitation>
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<figureCitation id="1351049F984FFF97FE388F68FE7EE819" box="[453,483,1867,1891]" captionStart="FIGURES 42–45" captionStartId="27.[151,264,1498,1521]" captionTargetBox="[163,1439,186,1462]" captionTargetId="figure@27.[163,1439,186,1462]" captionTargetPageId="27" captionText="FIGURES 42–45. Female head frontal: (42) L pendleburyi SAR16_GOM8; (43) L. schieli paratype SAR15_GOM6; (44) L. williamsoni SAR13_14_GOM51; (45) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068020/files/figure.png" pageId="29" pageNumber="230">44</figureCitation>
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<figureCitation id="1351049F984FFF97FE128F69FD90E819" box="[495,525,1866,1891]" captionStart="FIGURES 46–51" captionStartId="29.[151,264,1672,1695]" captionTargetBox="[156,1432,375,1651]" captionTargetId="figure@29.[156,1432,375,1651]" captionTargetPageId="29" captionText="FIGURES 46–51. Female head frontal detail; h—occipital horn; ot—occipital tube; t—tubercle; th—tubercle horn: (46) L. coomansi SAR11_12_GOM48; (47) L pendleburyi SAR16_GOM8; (48) L. schieli paratype SAR15_GOM6; (49) L. species SAR11_12_GOM33; (50) L. williamsoni SAR13_14_GOM51; (51) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068022/files/figure.png" pageId="29" pageNumber="230">50</figureCitation>
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<figureCitation id="1351049F984FFF97FDE48F69FDAAE819" box="[537,567,1866,1891]" captionStart="FIGURES 56–59" captionStartId="33.[151,264,1277,1300]" captionTargetBox="[156,1432,193,1256]" captionTargetId="figure@33.[156,1432,193,1256]" captionTargetPageId="33" captionText="FIGURES 56–59. Male synthorax dorsal: (56) L. pasia SAB12_GOM1; (57) L. sii holotype; (58) L. schieli holotype; (59) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068026/files/figure.png" pageId="29" pageNumber="230">59</figureCitation>
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<figureCitation id="1351049F984FFF97FDBE8F69FDFCE819" box="[579,609,1866,1891]" captionStart="FIGURES 60–63" captionStartId="34.[151,264,1601,1624]" captionTargetBox="[156,1432,517,1580]" captionTargetId="figure@34.[156,1432,517,1580]" captionTargetPageId="34" captionText="FIGURES 60–63. Male synthorax lateral: (60) L. coomansi SAR09_10_GOM4; (61) L. sp. cf coomansi SAB12_GOM4; (62) L. schieli holotype; (63) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068028/files/figure.png" pageId="29" pageNumber="230">63</figureCitation>
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<figureCitation id="1351049F984FFF97FD938F69FD11E819" box="[622,652,1866,1891]" captionStart="FIGURES 74–77" captionStartId="37.[151,264,1356,1379]" captionTargetBox="[151,1435,194,1335]" captionTargetId="figure@37.[151,1436,193,1335]" captionTargetPageId="37" captionText="FIGURES 74–77. Female synthorax dorsal: (74) L pendleburyi SAR16_GOM8; (75) L. schieli paratype SAR15_GOM6; (76) L. williamsoni SAR13_14_GOM51; (77) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068034/files/figure.png" pageId="29" pageNumber="230">76</figureCitation>
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<figureCitation id="1351049F984FFF97FD6A8F69FD28E819" box="[663,693,1866,1891]" captionStart="FIGURES 84–87" captionStartId="39.[151,264,1427,1450]" captionTargetBox="[151,1435,194,1406]" captionTargetId="figure@39.[151,1436,193,1406]" captionTargetPageId="39" captionText="FIGURES 84–87. Female synthorax lateral: (84) L pendleburyi SAR16_GOM8; (85) L. schieli paratype SAR15_GOM6; (86) L. williamsoni SAR13_14_GOM51, right hand side, flipped for consistency;(87) L. sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068038/files/figure.png" pageId="29" pageNumber="230">86</figureCitation>
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<figureCitation id="1351049F984FFF97FD3C8F68FD42E819" box="[705,735,1867,1891]" captionStart="FIGURES 88–93" captionStartId="40.[151,264,1280,1303]" captionTargetBox="[156,1432,195,1258]" captionTargetId="figure@40.[156,1432,195,1258]" captionTargetPageId="40" captionText="FIGURES 88–93. Male accessory genitalia, lateral view, all scale bars 1mm: (88) L. coomansi SAR09_10_GOM4; (89) L. pasia SAB12_GOM1; (90) L. pendleburyi SAR13_14_GOM13; (91) L. schieli holotype; (92) L. sii holotype; (93) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068040/files/figure.png" pageId="29" pageNumber="230">93</figureCitation>
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<figureCitation id="1351049F984FFF97FD168F68FC97E819" box="[747,778,1867,1891]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="29" pageNumber="230">99</figureCitation>
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<figureCitation id="1351049F984FFF97FCE88F69FCDFE819" box="[789,834,1866,1891]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="29" pageNumber="230">105</figureCitation>
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<figureCitation id="1351049F984FFF97FCB08F68FCEAE819" box="[845,887,1867,1891]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="29" pageNumber="230">111</figureCitation>
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<figureCitation id="1351049F984FFF97FC7E8F68FC33E819" box="[899,942,1867,1891]" captionStart="FIGURES 112–117" captionStartId="44.[151,264,1808,1831]" captionTargetBox="[156,1432,193,1787]" captionTargetId="figure@44.[156,1432,193,1787]" captionTargetPageId="44" captionText="FIGURES 112–117. Male anal appendages in ventral view (all scale bars 0.5mm): (112) L. coomansi SAR09_10_GOM4; (113) L.pasia SAB12_GOM1;(114) L.pendleburyi SAR13_14_GOM13; (115) L. schieli holotype; (116) L.sii holotype; (117) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068048/files/figure.png" pageId="29" pageNumber="230">117</figureCitation>
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<figureCitation id="1351049F984FFF97FC478F69FC7BE819" box="[954,998,1866,1891]" captionStart="FIGURES 120–125" captionStartId="46.[151,264,1891,1914]" captionTargetBox="[151,1435,372,1869]" captionTargetId="figure@46.[151,1436,371,1870]" captionTargetPageId="46" captionText="FIGURES 120–125. Female vulvar scale (all scale bars 0.5mm): (120) L. coomansi SAR11_12_GOM48; (121) L. pasia holotype; (122) L. pendleburyi SAR13_14_GOM15; (123) L. schieli paratype SAR15_GOM6; (124) L. sii paratype SAR13_14_GOM44; (125) L.williamsoni SAR13_14_GOM51." httpUri="https://zenodo.org/record/1068052/files/figure.png" pageId="29" pageNumber="230">125</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984FFF97FC0F8F68FB83E819" box="[1010,1054,1867,1891]" captionStart="FIGURE 129" captionStartId="51.[151,250,1608,1630]" captionTargetBox="[151,1436,193,1587]" captionTargetId="figure@51.[151,1436,193,1587]" captionTargetPageId="51" captionText="FIGURE 129. Distribution of L. williamsoni (black square) and L.sp.cf williamsoni (red triangle)." httpUri="https://zenodo.org/record/1068060/files/figure.png" pageId="29" pageNumber="230">129</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984FFF97FBD78F68FBCBE819" box="[1066,1110,1867,1891]" captionStart="FIGURES 130–132" captionStartId="52.[151,264,1763,1786]" captionTargetBox="[453,1133,193,1742]" captionTargetId="figure@52.[453,1133,193,1742]" captionTargetPageId="52" captionText="FIGURES 130–132. Habitus of some Bornean Leptogomphus species:(130) L. pasia holotype (left hand side, flipped to right for consistency with other images); (131) L.sii holotype male; (132) L. williamsoni male SAR13_14_GOM56." httpUri="https://zenodo.org/record/1068062/files/figure.png" pageId="29" pageNumber="230">132</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984FFF97FB9F8F68FB10E819" box="[1122,1165,1867,1891]" captionStart="FIGURES 133–137" captionStartId="53.[151,264,1171,1194]" captionTargetBox="[151,1436,193,1150]" captionTargetId="figure@53.[151,1436,193,1150]" captionTargetPageId="53" captionText="FIGURES 133–137. Photographs in life of some Bornean Leptogomphus species: (133) L. coomansi male (left hand side, flippedto right for consistency with other images), Sungai Sbong, Kapit Division, Sarawak, photograph by G.T.Reels; (134) L. coomansi female, Ranchan Recreational Park, Serian, Serian Division, Sarawak; (135) L. pendleburyi female, Bukit Tanggan, Ulu Mujok, Lanjak Entimau WildlifeSanctuary, Sarikei Division, Sarawak, photograph by Robin Ngiam; (136) L. pendleburyi male, Sungai Kahei area, Ulu Balui, Kapit Division, Sarawak, photograph by Robin Ngiam; (137) L. williamsoni immature male, semi-teneral, Sungai Kahei area, Ulu Balui, Kapit Division,Sarawak, photograph by Robin Ngiam." httpUri="https://zenodo.org/record/1068064/files/figure.png" pageId="29" pageNumber="230">137</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B91984FFF94FF6A8FB1FC6EEF91" lastPageId="30" lastPageNumber="231" pageId="29" pageNumber="230" type="reference_group">
|
||
<paragraph id="8BD5181A984FFF94FF6A8FB1FC6EEF91" blockId="29.[151,1436,1938,2021]" lastBlockId="30.[199,1436,152,235]" lastPageId="30" lastPageNumber="231" pageId="29" pageNumber="230">
|
||
<treatmentCitationGroup id="AB7A3F34984FFF94FF6A8FB1FC6EEF91" lastPageId="30" lastPageNumber="231" pageId="29" pageNumber="230">
|
||
<taxonomicName id="4C6A6399984FFF97FF6A8FB1FDF6E8D2" ID-CoL="3TQ4T" authority="Laidlaw 1912: 94" authorityName="Laidlaw" authorityPageNumber="94" authorityYear="1912" box="[151,619,1938,1960]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="29" pageNumber="230" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC408984FFF97FF6A8FB1FE37E8D2" box="[151,426,1938,1960]" italics="true" pageId="29" pageNumber="230">Leptogomphus williamsoni</emphasis>
|
||
<treatmentCitation id="0ACB3E0B984FFF97FE4E8FB1FDF6E8D2" author="Laidlaw" box="[435,619,1938,1960]" page="94" pageId="29" pageNumber="230" year="1912">
|
||
<bibRefCitation id="EFFB65EB984FFF97FE4E8FB1FDDBE8D2" author="Laidlaw" box="[435,582,1938,1960]" pageId="29" pageNumber="230" refString="Laidlaw, F. F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. (Appendix VIII to Moulton, J. C., 1912.) Journal of the Straits Branch of the Royal Asiatic Society, 63, 92 - 99, 1 pl." type="journal article" year="1912">Laidlaw 1912</bibRefCitation>
|
||
: 94
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
–95,
|
||
<figureCitation id="1351049F984FFF97FD628FB1FD7CE8D2" box="[671,737,1938,1960]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="29" pageNumber="230">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984FFF97FD128FB1FD66E8D2" box="[751,763,1938,1960]" captionStart="FIGURE 2" captionStartId="5.[151,250,1294,1316]" captionTargetBox="[243,1384,200,1237]" captionTargetId="figure@5.[151,1436,193,1273]" captionTargetPageId="5" captionText="FIGURE 2. ITS gene tree for 20 specimens of Leptogomphus and three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068000/files/figure.png" pageId="29" pageNumber="230">2</figureCitation>
|
||
(original description male, Madihit, Sarawak);—
|
||
<treatmentCitation id="0ACB3E0B984FFF97FB048FB1FF73E8BD" author="Lieftinck" page="247" pageId="29" pageNumber="230" year="1948">
|
||
<bibRefCitation id="EFFB65EB984FFF97FB048FB1FA05E8D2" author="Lieftinck" box="[1273,1432,1938,1960]" pageId="29" pageNumber="230" refString="Lieftinck, M. A. (1948) Descriptions and records of South - east Asiatic Odonata. Treubia, 19, 221 - 278." type="journal article" year="1948">Lieftinck 1948</bibRefCitation>
|
||
: 247
|
||
</treatmentCitation>
|
||
–249, Plate 8,
|
||
<figureCitation id="1351049F984FFF97FE828F92FE5AE8BD" box="[383,455,1969,1991]" captionStart="FIGURES10–13" captionStartId="14.[151,264,1864,1887]" captionTargetBox="[151,1436,772,1843]" captionTargetId="figure@14.[151,1436,772,1843]" captionTargetPageId="14" captionText="FIGURES10–13. Male head frontal: (10) L.coomansi SAR09_10_GOM4; (11) L.coomansi holotype; (12) L.sp. cf coomansi SAB12_GOM4; (13) L.pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068006/files/figure.png" pageId="29" pageNumber="230">Fig. 10</figureCitation>
|
||
(description of ♀, Kutai);—
|
||
<treatmentCitation id="0ACB3E0B984FFF97FD188F92FC39E8BD" author="Lieftinck" box="[741,932,1969,1991]" page="83" pageId="29" pageNumber="230" year="1954">
|
||
<bibRefCitation id="EFFB65EB984FFF97FD188F92FC1EE8BD" author="Lieftinck" box="[741,899,1969,1991]" pageId="29" pageNumber="230" refString="Lieftinck, M. A. (1954) Handlist of Malaysian Odonata. A catalogue of the dragonflies of the Malay Peninsula, Sumatra, Java and Borneo, including the adjacent small islands. Treubia, 22 (Supplement), i - xiii + 1 - 202." type="journal article" year="1954">Lieftinck 1954</bibRefCitation>
|
||
: 83
|
||
</treatmentCitation>
|
||
;—
|
||
<treatmentCitation id="0ACB3E0B984FFF97FC3F8F92FB0DE8BD" author="Kimmins" box="[962,1168,1969,1991]" page="299" pageId="29" pageNumber="230" year="1969">
|
||
<bibRefCitation id="EFFB65EB984FFF97FC3F8F92FBFDE8BD" author="Kimmins" box="[962,1120,1969,1991]" pageId="29" pageNumber="230" refString="Kimmins, D. E. (1969) A list of the type - specimens of Odonata in the British Museum (Natural History) part II. Bulletin of the British Museum (Natural History), Entomology, 23 (7), 289 - 314. https: // doi. org / 10.5962 / bhl. part. 15135" type="journal article" year="1969">Kimmins 1969</bibRefCitation>
|
||
: 299
|
||
</treatmentCitation>
|
||
(note on type);—
|
||
<treatmentCitation id="0ACB3E0B984FFF97FABF8F92FE1BE89F" author="Tsuda" page="38" pageId="29" pageNumber="230" year="1989">
|
||
<bibRefCitation id="EFFB65EB984FFF97FABF8F92FEF9E89F" author="Tsuda" pageId="29" pageNumber="230" refString="Tsuda, S. and Kitagawa, K. (1989) Odonata of Southeast Asia collected by the late Mr. M. Iwasaki. Part 3. Odonata of North Borneo (Sabah, Malaysia). Gracile, 40, 37 - 39." type="journal article" year="1989">Tsuda & Kitagawa 1989</bibRefCitation>
|
||
: 38
|
||
</treatmentCitation>
|
||
(♂, Poring);—van
|
||
<treatmentCitation id="0ACB3E0B984FFF97FDB68FECFD53E89F" author="Tol" box="[587,718,1999,2021]" page="98" pageId="29" pageNumber="230" year="1990">
|
||
<bibRefCitation id="EFFB65EB984FFF97FDB68FECFD34E89F" author="Tol" box="[587,681,1999,2021]" pageId="29" pageNumber="230" refString="van Tol, J. (1990) Key to the Malesian species of Leptogomphus Selys, with the description of a new species from Sabah (Odonata, Gomphidae). Tijdschrift voor Entomologie, 133, 97 - 105." type="journal article" year="1990">Tol 1990</bibRefCitation>
|
||
: 98
|
||
</treatmentCitation>
|
||
, 99, 101, 102,
|
||
<figureCitation id="1351049F984FFF97FC9F8FECFC32E89F" box="[866,943,1999,2021]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1933,1956]" captionTargetBox="[151,1436,841,1912]" captionTargetId="figure@16.[151,1436,841,1912]" captionTargetPageId="16" captionText="FIGURES 14–17. Male head frontal: (14) L. pasia SAB12_GOM1; (15) L. sii holotype; (16) L. schieli holotype; (17) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="29" pageNumber="230">Figs 17</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984FFF97FC448FECFC66E89F" box="[953,1019,1999,2021]" captionStart="FIGURES 24–27" captionStartId="20.[151,264,1441,1464]" captionTargetBox="[163,1439,268,1402]" captionTargetId="figure@20.[163,1439,268,1402]" captionTargetPageId="20" captionText="FIGURES 24–27. Female head dorsal: (24) L pendleburyi SAR16_GOM8; (25) L. schieli paratype SAR15_GOM6; (26) L. williamsoni SAR13_14_GOM51; (27) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068012/files/figure.png" pageId="29" pageNumber="230">26, 27</figureCitation>
|
||
,
|
||
<tableCitation id="C6E82DA1984FFF97FBF88FECFBD2E89F" box="[1029,1103,1999,2021]" captionStart="TABLE 1" captionStartId="2.[151,239,358,381]" captionText="TABLE 1. Primer combinations used for amplification of ITS and COI." pageId="29" pageNumber="230">Table 1</tableCitation>
|
||
,
|
||
<tableCitation id="C6E82DA1984FFF97FBA68FECFBF5E89F" box="[1115,1128,1999,2021]" captionStart="TABLE 2" captionStartId="3.[151,239,256,279]" captionTargetBox="[159,1375,443,1403]" captionText="TABLE 2. Leptogomphus from Borneo used in the molecular analyses; collectors and more details of location given in material lists under each species. Collection codes and BOLD Process IDs are given along with basic location details. The collection codes can be used to locate the sequences on the BOLD website, and also appear as BOLD Sample IDs there." pageId="29" pageNumber="230">2</tableCitation>
|
||
(key);—van
|
||
<treatmentCitation id="0ACB3E0B984FFF97FB138FECFAE1E89F" author="Tol" box="[1262,1404,1999,2021]" page="236" pageId="29" pageNumber="230" year="1992">
|
||
<bibRefCitation id="EFFB65EB984FFF97FB138FECFAD1E89F" author="Tol" box="[1262,1356,1999,2021]" pageId="29" pageNumber="230" refString="van Tol, J. (1992) An annotated index to names of Odonata used in publications by M. A. Lieftinck. Zoologische Verhandelingen, 279, 1 - 263." type="journal article" year="1992">Tol 1992</bibRefCitation>
|
||
: 236
|
||
</treatmentCitation>
|
||
;—
|
||
<treatmentCitation id="0ACB3E0B984CFF94FF3A88BBFED0EFD4" author="Orr" box="[199,333,152,174]" page="40" pageId="30" pageNumber="231" year="2003">
|
||
<bibRefCitation id="EFFB65EB984CFF94FF3A88BBFEB5EFD4" author="Orr" box="[199,296,152,174]" pageId="30" pageNumber="231" refString="Orr, A. G. (2003) A guide to the dragonflies of Borneo: their identification and biology. Natural History Publications (Borneo), Kota Kinabalu, x + 195 pp." type="book" year="2003">Orr 2003</bibRefCitation>
|
||
: 40
|
||
</treatmentCitation>
|
||
;—
|
||
<treatmentCitation id="0ACB3E0B984CFF94FE9688BBFDF5EFD4" author="Dow" box="[363,616,152,174]" page="14" pageId="30" pageNumber="231" year="2012">
|
||
<bibRefCitation id="EFFB65EB984CFF94FE9688BBFDD9EFD4" author="Dow" box="[363,580,152,174]" pageId="30" pageNumber="231" refString="Dow, R. A. & Ngiam, R. W. J. (2012) Odonata collected in the Hose Mountains, Kapit Division, Sarawak, Malaysia in April 2011. International Dragonfly Fund Report, 44, 1 - 18." type="journal article" year="2012">Dow & Ngiam 2012</bibRefCitation>
|
||
: 14
|
||
</treatmentCitation>
|
||
(both sexes, Hose Mountains, Sarawak);—
|
||
<treatmentCitation id="0ACB3E0B984CFF94FBDB88BBFAAFEFD4" author="Ngiam" box="[1062,1330,152,174]" page="307" pageId="30" pageNumber="231" year="2013">
|
||
<bibRefCitation id="EFFB65EB984CFF94FBDB88BBFB63EFD4" author="Ngiam" box="[1062,1278,152,174]" pageId="30" pageNumber="231" refString="Ngiam, R. W. J. & Dow, R. A. (2013) The larva of Leptogomphus risi Laidlaw from Singapore with a comparison to Leptogomphus williamsoni Laidlaw from Sarawak and congeners (Odonata: Anisoptera: Gomphidae). Nature in Singapore, 6, 307 - 312." type="journal article" year="2013">Ngiam & Dow 2013</bibRefCitation>
|
||
: 307
|
||
</treatmentCitation>
|
||
, 309–311,
|
||
<figureCitation id="1351049F984CFF94FF3A8895FE89EFB6" box="[199,276,182,204]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="30" pageNumber="231">Figs 1b</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984CFF94FEDC8895FEA0EFB6" box="[289,317,182,204]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="30" pageNumber="231">3b</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984CFF94FEB48895FECBEFB6" box="[329,342,182,204]" captionStart="FIGURES 4–9" captionStartId="9.[151,264,1713,1736]" captionTargetBox="[151,1436,193,1692]" captionTargetId="figure@9.[151,1436,193,1692]" captionTargetPageId="9" captionText="FIGURES 4–9. Male head dorsal: (4) L. coomansi SAR09_10_GOM4; (5) L. pendleburyi SAR13_14_GOM13; (6) L. pasia SAB12_GOM1; (7) L. sii holotype; (8) L.schieli holotype; (9) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068004/files/figure.png" pageId="30" pageNumber="231">4</figureCitation>
|
||
(♀, larva, Hose Mountains, Sarawak);—
|
||
<treatmentCitation id="0ACB3E0B984CFF94FD018895FC67EFB6" author="Dow" box="[764,1018,182,204]" page="35" pageId="30" pageNumber="231" year="2014">
|
||
<bibRefCitation id="EFFB65EB984CFF94FD018895FC48EFB6" author="Dow" box="[764,981,182,204]" pageId="30" pageNumber="231" refString="Dow, R. A. & Ngiam, R. W. J. (2014) Odonata from logged and unlogged forest in the Ulu Balui and Ulu Baleh, Kapit Division, Sarawak, in June and September 2013. International Dragonfly Fund Report, 73, 1 - 48." type="journal article" year="2014">Dow & Ngiam 2014</bibRefCitation>
|
||
: 35
|
||
</treatmentCitation>
|
||
,
|
||
<figureCitation id="1351049F984CFF94FBFB8895FBD2EFB6" box="[1030,1103,182,204]" captionStart="FIGURES28–31" captionStartId="22.[151,264,1357,1380]" captionTargetBox="[151,1435,266,1336]" captionTargetId="figure@22.[151,1436,265,1336]" captionTargetPageId="22" captionText="FIGURES28–31. Female head dorsaldetail; h—occipital horn; ot—occipital tube; p—pit; s—occipital spur; t—tubercle;thtubercle horn: (28) L. coomansi SAR11_12_GOM48; (29) L. mariae holotype; (30) L. sp. cf coomansi PomS260; (31) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068014/files/figure.png" pageId="30" pageNumber="231">Fig. 28</figureCitation>
|
||
(both sexes, Ulu Baleh and Ulu Balui, Sarawak);—
|
||
<treatmentCitation id="0ACB3E0B984CFF94FE7488F6FDFDEF91" author="Dow" box="[393,608,213,235]" page="27" pageId="30" pageNumber="231" year="2015">
|
||
<bibRefCitation id="EFFB65EB984CFF94FE7488F6FDA0EF91" author="Dow" box="[393,573,213,235]" pageId="30" pageNumber="231" refString="Dow, R. A., Reels, G. T. & Ngiam, R. W. J. (2015 a) Previously unpublished Odonata records from Sarawak, Borneo, Part III. Sri Aman, Sibu and Kapit Divisions. Faunistic Studies in South-East Asian and Pacific Island Odonata, 9, 1 - 34." type="journal article" year="2015" yearSuffix="a">
|
||
Dow
|
||
<emphasis id="B91EC408984CFF94FE3F88F5FE73EF91" box="[450,494,213,235]" italics="true" pageId="30" pageNumber="231">et al</emphasis>
|
||
. 2015a
|
||
</bibRefCitation>
|
||
: 27
|
||
</treatmentCitation>
|
||
(locations in Kapit Division, Sarawak).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B91984CFF94FF6A8934FA96E862" pageId="30" pageNumber="231" type="materials_examined">
|
||
<paragraph id="8BD5181A984CFF94FF6A8934FF6EEE0D" blockId="30.[151,1437,279,2033]" pageId="30" pageNumber="231">
|
||
<emphasis id="B91EC408984CFF94FF6A8934FD02EE4A" bold="true" box="[151,671,279,304]" pageId="30" pageNumber="231">
|
||
Material studied.
|
||
<typeStatus id="54D1A6B8984CFF94FE898934FE32EE4A" box="[372,431,279,304]" pageId="30" pageNumber="231">Type</typeStatus>
|
||
material.
|
||
<typeStatus id="54D1A6B8984CFF94FDCF8934FD02EE4A" box="[562,671,279,304]" pageId="30" pageNumber="231" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<specimenCount id="9D6CD393984CFF94FD57893BFD44EE55" box="[682,729,280,304]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED7B80DF984CFF94FD168934FCC5EE4A" box="[747,856,279,304]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="30" pageNumber="231">NHMUK</collectionCode>
|
||
010595516), Madihit,
|
||
<collectingRegion id="49AED6F8984CFF94FBA18934FB22EE4A" box="[1116,1215,279,304]" country="Malaysia" name="Sarawak" pageId="30" pageNumber="231">
|
||
<collectingCountry id="F37D588A984CFF94FBA18934FB22EE4A" box="[1116,1215,279,304]" name="Malaysia" pageId="30" pageNumber="231">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
, Borneo,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FAC9893BFA05EE4A" box="[1332,1432,280,304]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="5" value="3.8707">3.8707N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FF6A891FFE8DEE2F" box="[151,272,316,341]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="5" value="115.3866">115.3866E</geoCoordinate>
|
||
(Batu Lawi, Madihit on route taken by Moulton to Batu Lawi), ca
|
||
<quantity id="4C92B5FF984CFF94FC08891FFBA5EE2F" box="[1013,1080,316,341]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="30" pageNumber="231" unit="m" value="600.0">600m</quantity>
|
||
,
|
||
<date id="FFD43EDA984CFF94FBBE891FFB30EE2E" box="[1091,1197,316,341]" pageId="30" pageNumber="231" value="1911-06-04">4 vi 1911</date>
|
||
, leg. J.C. Moulton, in
|
||
<collectionCode id="ED7B80DF984CFF94FF6A8943FF72EE0D" box="[151,239,352,375]" pageId="30" pageNumber="231">BMNH</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A984CFF94FF3A89A7FA96E862" blockId="30.[151,1437,279,2033]" pageId="30" pageNumber="231">
|
||
<emphasis id="B91EC408984CFF94FF3A89A7FD1EEEE7" bold="true" box="[199,643,388,413]" pageId="30" pageNumber="231">
|
||
Other Material.
|
||
<collectingRegion id="49AED6F8984CFF94FE6F89A7FE60EEE7" box="[402,509,388,413]" country="Malaysia" name="Sarawak" pageId="30" pageNumber="231">
|
||
<collectingCountry id="F37D588A984CFF94FE6F89A7FE60EEE7" box="[402,509,388,413]" name="Malaysia" pageId="30" pageNumber="231">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37D588A984CFF94FDF189A7FDE0EEE7" box="[524,637,388,413]" name="Malaysia" pageId="30" pageNumber="231">Malaysia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D6CD393984CFF94FD7189A7FD24EEE6" box="[652,697,388,412]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
, small tributary to Sungai Engkari, Nanga Segerak,
|
||
<collectionCode id="ED7B80DF984CFF94FAE689A7FAF7EEE6" box="[1307,1386,388,412]" pageId="30" pageNumber="231">LEWS</collectionCode>
|
||
, ulu Engkari, Sri Aman Division,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FE208984FDD2EEBA" box="[477,591,423,448]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="1.41376">1.41376N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FDA7898BFD7CEEBA" box="[602,737,424,448]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="112.00449">112.00449E</geoCoordinate>
|
||
(Ulu Engkari),
|
||
<date id="FFD43EDA984CFF94FC6C8984FB8FEEBA" box="[913,1042,423,448]" pageId="30" pageNumber="231" value="2016-07-15">15 vii 2016</date>
|
||
, leg. G.T. Reels, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FF6A89EFFF58EE9E" box="[151,197,460,484]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
(teneral), tributary to Sungai Segerak, same area,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FCEA89EFFC15EE9F" box="[791,904,460,485]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="1.41186">1.41186N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FC6A89EFFBBDEE9F" box="[919,1056,460,485]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="112.00518">112.00518E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA984CFF94FBCD89EFFB27EE9F" box="[1072,1210,460,485]" pageId="30" pageNumber="231" value="2016-07-17">17 vii 2016</date>
|
||
, leg. G.T. Reels, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FEAF89D3FEE0ED7D" box="[338,381,496,520]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED7B80DF984CFF94FE7189CCFE5BED72" box="[396,454,495,520]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="30" pageNumber="231">SAR</collectionCode>
|
||
16_GOM1, used in
|
||
<figureCitation id="1351049F984CFF94FD5989CCFD6AED72" box="[676,759,495,520]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="30" pageNumber="231">Fig. 99</figureCitation>
|
||
), seepage by small stream next to field station, Nanga Bloh, ulu Katibas,
|
||
<collectionCode id="ED7B80DF984CFF94FED48A37FEE5ED56" box="[297,376,532,556]" pageId="30" pageNumber="231">LEWS</collectionCode>
|
||
, Kapit Division,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FDC18A37FD33ED57" box="[572,686,532,557]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="1.64617">1.64617N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FD468A37FCD8ED57" box="[699,837,532,557]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="112.27592">112.27592E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA984CFF94FCAF8A37FC4FED57" box="[850,978,532,557]" pageId="30" pageNumber="231" value="2016-03-12">12 iii 2016</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FA8D8A37FA01ED56" box="[1392,1436,532,556]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(teneral), Sungai Nyungan, same area,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FDAC8A14FD5EED2A" box="[593,707,567,592]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="1.65723">1.65723N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FD2D8A1BFCD1ED2A" box="[720,844,568,592]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="5" value="112.2432">112.2432E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA984CFF94FCA48A1BFC45ED2A" box="[857,984,567,592]" pageId="30" pageNumber="231" value="2016-03-14">14 iii 2016</date>
|
||
, leg. E. Jangoh, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FA8C8A1BFA01ED35" box="[1393,1436,568,592]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(teneral), Sungai Menyarin, same area,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FDAC8A7FFD5EED0F" box="[593,707,604,629]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="1.65518">1.65518N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FD338A7FFCC5ED0E" box="[718,856,604,628]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="112.21832">112.21832E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA984CFF94FC998A7FFC7DED0F" box="[868,992,604,629]" pageId="30" pageNumber="231" value="2016-03-15">15 iii 2016</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FA8F8A7FFA01ED0E" box="[1394,1436,604,628]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED7B80DF984CFF94FF628A5CFF64EDE2" box="[159,249,639,664]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="231">RMNH</collectionCode>
|
||
.INS.501379), tributary to Sungai Ulu Yong, Kapit town area, Kapit Division,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FB868AA3FB59EDE2" box="[1147,1220,640,664]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="555" value="1.94">1.94N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FB2C8AA3FAA1EDE2" box="[1233,1340,640,664]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="55" value="112.828">112.828E</geoCoordinate>
|
||
(Sungai Ulu Yong),
|
||
<date id="FFD43EDA984CFF94FEE28A87FE0AEDC6" box="[287,407,676,700]" pageId="30" pageNumber="231" value="2009-10-21">21 x 2009</date>
|
||
, leg. R.A. Dow, in
|
||
<collectionCode id="ED7B80DF984CFF94FD878A87FD4AEDC6" box="[634,727,676,700]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="231">RMNH</collectionCode>
|
||
;
|
||
<specimenCount id="9D6CD393984CFF94FD268A87FC9FEDC6" box="[731,770,676,700]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
, small stream in disturbed forest, Nanga Gaat area, Kapit Division,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FEF48AEBFEE6ED9A" box="[265,379,712,736]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="1.74723">1.74723N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FE778AEBFD9EED9A" box="[394,515,712,736]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="5" value="113.3487">113.3487E</geoCoordinate>
|
||
(Nanga Gaat area),
|
||
<date id="FFD43EDA984CFF94FD0E8AE4FCFAED9A" box="[755,871,711,736]" pageId="30" pageNumber="231" value="2015-09-05">5 ix 2015</date>
|
||
, leg. M. Budi, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FB038AEBFAB6EDA5" box="[1278,1323,712,736]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
(teneral), stream in virgin forest, Sungai Kahei area, Ulu Balui,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FCF98ACFFCC6EC7F" box="[772,859,748,773]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="55" value="2.046">2.046N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FC958ACFFC48EC7F" box="[872,981,748,773]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="55" value="114.572">114.572E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA984CFF94FC1F8ACFFBFCEC7E" box="[994,1121,748,773]" pageId="30" pageNumber="231" value="2013-06-15">15 vi 2013</date>
|
||
, leg. M. Budi, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FF238B33FE97EC5D" box="[222,266,784,808]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
(teneral), same area,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FDFE8B33FDC4EC52" box="[515,601,784,808]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="55" value="2.038">2.038N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FD9B8B2CFD49EC52" box="[614,724,783,808]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="55" value="114.602">114.602E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA984CFF94FD1C8B33FCFDEC52" box="[737,864,783,808]" pageId="30" pageNumber="231" value="2013-09-10">10 ix 2013</date>
|
||
, leg. L. Southwell, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FADD8B33FAD1EC5D" box="[1312,1356,784,808]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(semiteneral), stream
|
||
<quantity id="4C92B5FF984CFF94FEAD8B17FE59EC36" box="[336,452,820,844]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="30" pageNumber="231" unit="m" value="850.0" valueMax="900.0" valueMin="800.0">800-900m</quantity>
|
||
in disturbed forest, same area,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FCD28B17FC18EC36" box="[815,901,820,844]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="55" value="2.012">2.012N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FC6F8B17FB9DEC37" box="[914,1024,820,845]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="55" value="114.672">114.672E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA984CFF94FBF08B17FB10EC36" box="[1037,1165,820,845]" pageId="30" pageNumber="231" value="2013-06-16">16 vi 2013</date>
|
||
, leg. L. Southwell;
|
||
<specimenCount id="9D6CD393984CFF94FA8D8B17FA01EC36" box="[1392,1436,820,844]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(teneral), streams
|
||
<quantity id="4C92B5FF984CFF94FE908B74FE7EEC0A" box="[365,483,855,880]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="8.4" metricValueMin="6.6" pageId="30" pageNumber="231" unit="m" value="750.0" valueMax="840.0" valueMin="660.0">660-840m</quantity>
|
||
, Gunung Kajang area, Hose Mountains, Kapit Division,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FB6F8B74FA99EC0A" box="[1170,1284,855,880]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="2.29768">2.29768N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FAE88B74FA01EC0A" box="[1301,1436,855,880]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="113.69733">113.69733E</geoCoordinate>
|
||
(Gunung Kajang area),
|
||
<date id="FFD43EDA984CFF94FE508B5FFDBCECEE" box="[429,545,892,917]" pageId="30" pageNumber="231" value="2011-04-09">9 iv 2011</date>
|
||
, leg. R.A. Dow;
|
||
<specimenCount id="9D6CD393984CFF94FD0C8B5FFCBDECEE" box="[753,800,892,916]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED7B80DF984CFF94FCC98B5FFC13ECEF" box="[820,910,892,917]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="231">RMNH</collectionCode>
|
||
.INS.503842), stream at ca
|
||
<quantity id="4C92B5FF984CFF94FB328B5FFA8FECEE" box="[1231,1298,892,916]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="30" pageNumber="231" unit="m" value="740.0">740m</quantity>
|
||
, same area,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FF6A8BBCFE94ECC2" box="[151,265,927,952]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="2.29768">2.29768N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FEEB8BBCFE00ECC2" box="[278,413,927,952]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="113.69733">113.69733E</geoCoordinate>
|
||
(Gunung Kajang area),
|
||
<date id="FFD43EDA984CFF94FD498B83FCACECC2" box="[692,817,927,952]" pageId="30" pageNumber="231" value="2011-04-13">13 iv 2011</date>
|
||
, leg. M. Kibi, in
|
||
<collectionCode id="ED7B80DF984CFF94FC048B83FBCDECCD" box="[1017,1104,928,951]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="231">RMNH</collectionCode>
|
||
;
|
||
<specimenCount id="9D6CD393984CFF94FBA08B83FB14ECCD" box="[1117,1161,928,952]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(teneral), Sungai Sagan Besai, foot of Hose mountains, Kapit Division,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FD4E8BE7FCBBECA7" box="[691,806,964,989]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="2.06478">2.06478N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FCCC8BE7FC25ECA7" box="[817,952,964,989]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="113.67565">113.67565E</geoCoordinate>
|
||
(general coordinates for area),
|
||
<date id="FFD43EDA984CFF94FAE18BE7FA0AECA6" box="[1308,1431,964,989]" pageId="30" pageNumber="231" value="2011-09-25">25 ix 2011</date>
|
||
, leg. M. Budi, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FDF38BCBFDA5EC85" box="[526,568,1000,1024]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED7B80DF984CFF94FDBA8BC4FD3CEB7A" box="[583,673,999,1024]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="231">RMNH</collectionCode>
|
||
.INS.504046; teneral), streams ca
|
||
<quantity id="4C92B5FF984CFF94FBE08BC4FB0EEB7A" box="[1053,1171,999,1024]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="9.4" metricValueMin="8.6" pageId="30" pageNumber="231" unit="m" value="900.0" valueMax="940.0" valueMin="860.0">860-940m</quantity>
|
||
, Stone Park, Batu Laga Plateau,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FEF88C2FFEEBEB5E" box="[261,374,1036,1060]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="2.38279">2.38279N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FE6D8C2FFD8AEB5F" box="[400,535,1036,1061]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="114.06441">114.06441E</geoCoordinate>
|
||
(Stone Park area),
|
||
<date id="FFD43EDA984CFF94FCD98C2FFC24EB5E" box="[804,953,1036,1061]" pageId="30" pageNumber="231" value="2011-09-29">29 ix 2011</date>
|
||
, leg. L. Southwell, in
|
||
<collectionCode id="ED7B80DF984CFF94FB098C2EFAD1EB5E" box="[1268,1356,1037,1060]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="231">RMNH</collectionCode>
|
||
;
|
||
<specimenCount id="9D6CD393984CFF94FA998C2FFA01EB5E" box="[1380,1436,1036,1060]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED7B80DF984CFF94FF628C0CFF45EB32" box="[159,216,1071,1096]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="30" pageNumber="231">SAR</collectionCode>
|
||
11_12_GOM56, used for description and illustrations), Bukit Naong, Tatau district, Bintulu Division,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FF6A8C77FE94EB17" box="[151,265,1108,1133]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="2.68512">2.68512N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FEE98C77FE03EB17" box="[276,414,1108,1133]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="112.86433">112.86433E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA984CFF94FE548C77FDBBEB16" box="[425,550,1108,1133]" pageId="30" pageNumber="231" value="2012-03-22">22 iii 2012</date>
|
||
, leg. N. Megom, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FC3C8C77FC76EB16" box="[961,1003,1108,1132]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED7B80DF984CFF94FC078C77FBC9EB17" box="[1018,1108,1108,1133]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="231">RMNH</collectionCode>
|
||
.INS.506248; teneral), stream in block A1M, Tatau district,
|
||
<collectingRegion id="49AED6F8984CFF94FE1F8C54FDDFEBEA" box="[482,578,1143,1168]" country="Malaysia" name="Sarawak" pageId="30" pageNumber="231">
|
||
<collectingCountry id="F37D588A984CFF94FE1F8C54FDDFEBEA" box="[482,578,1143,1168]" name="Malaysia" pageId="30" pageNumber="231">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
Planted Forest Project, Bintulu Division,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FBE58C54FBE1EBEA" box="[1048,1148,1143,1168]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="5" value="2.6959">2.6959N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FB7A8C54FA8CEBEA" box="[1159,1297,1143,1168]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="112.88629">112.88629E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA984CFF94FAE18C5BFA05EBEA" box="[1308,1432,1143,1168]" pageId="30" pageNumber="231" value="2012-03-24">24 iii 2012</date>
|
||
, leg. R.A. Dow, in
|
||
<collectionCode id="ED7B80DF984CFF94FE9B8CBEFE23EBCE" box="[358,446,1181,1204]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="231">RMNH</collectionCode>
|
||
;
|
||
<specimenCount id="9D6CD393984CFF94FE378CBFFD97EBCE" box="[458,522,1180,1204]" pageId="30" pageNumber="231" type="male">2 ♂♂</specimenCount>
|
||
(one teneral), same location,
|
||
<date id="FFD43EDA984CFF94FCA68CBFFC44EBCE" box="[859,985,1180,1205]" pageId="30" pageNumber="231" value="2012-03-25">25 iii 2012</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FA8C8CBFFA01EBCE" box="[1393,1436,1180,1204]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED7B80DF984CFF94FF638C9CFF65EBA2" box="[158,248,1215,1240]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="231">RMNH</collectionCode>
|
||
.INS.506250, teneral), same data, in
|
||
<collectionCode id="ED7B80DF984CFF94FD638CE2FD6BEBA2" box="[670,758,1217,1240]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="231">RMNH</collectionCode>
|
||
;
|
||
<specimenCount id="9D6CD393984CFF94FCF98CE3FCACEBA2" box="[772,817,1216,1240]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
, small high gradient stream, Tubau,
|
||
<collectingRegion id="49AED6F8984CFF94FB208C9CFAA0EBA2" box="[1245,1341,1215,1240]" country="Malaysia" name="Sarawak" pageId="30" pageNumber="231">
|
||
<collectingCountry id="F37D588A984CFF94FB208C9CFAA0EBA2" box="[1245,1341,1215,1240]" name="Malaysia" pageId="30" pageNumber="231">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
Planted Forest Project, Bintulu Division,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FDF78CC7FDE0EB87" box="[522,637,1252,1277]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="3.14335">3.14335N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FD758CC7FD79EB87" box="[648,740,1252,1277]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="555" value="113.65">113.65E</geoCoordinate>
|
||
(Tubau area),
|
||
<date id="FFD43EDA984CFF94FC758CC7FB8EEB86" box="[904,1043,1252,1277]" pageId="30" pageNumber="231" value="2009-08-17">17 viii 2009</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FF6A8D2BFF5EEA5A" box="[151,195,1288,1312]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
, small stream, same area,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FE118D24FDC2EA5A" box="[492,607,1287,1312]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="3.14335">3.14335N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FD978D24FD5AEA5A" box="[618,711,1287,1312]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="555" value="113.65">113.65E</geoCoordinate>
|
||
(Tubau area),
|
||
<date id="FFD43EDA984CFF94FC908D2BFC64EA5A" box="[877,1017,1287,1312]" pageId="30" pageNumber="231" value="2009-08-31">31 viii 2009</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FA738D2BFF31EA3E" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
, stream in Lio Mato water catchment area, Lio Mato, Upper Baram, Miri Division,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FBA48D0FFB3CEA3F" box="[1113,1185,1324,1349]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="555" value="3.16">3.16N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FB518D0FFA87EA3F" box="[1196,1306,1324,1349]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="55" value="115.234">115.234E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA984CFF94FAD88D0FFA05EA3E" box="[1317,1432,1324,1348]" pageId="30" pageNumber="231" value="2009-10-13">13 x 2009</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FDE08D73FDD5EA12" box="[541,584,1360,1384]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(semi-teneral), by logging road from Sungai Sii to Sungai Alah, Sungai Sii, upper Baram, Miri Division,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FE238D57FDDFEAF7" box="[478,578,1396,1421]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="5" value="3.0151">3.0151N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FDB08D57FD5BEAF6" box="[589,710,1396,1420]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="5" value="114.9097">114.9097E</geoCoordinate>
|
||
(Sungai Alah),
|
||
<date id="FFD43EDA984CFF94FC848D57FC61EAF6" box="[889,1020,1396,1421]" pageId="30" pageNumber="231" value="2014-07-17">17 vii 2014</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FA738D57FF31EACA" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
(
|
||
<collectionCode id="ED7B80DF984CFF94FF3E8DB4FF60EACA" box="[195,253,1431,1456]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="30" pageNumber="231">SAR</collectionCode>
|
||
13_14_GOM51; used in illustrations), tributary to Sungai Sii downstream from camp, same area,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FF6A8D9FFE94EAAE" box="[151,265,1468,1492]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="2.99147">2.99147N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FEE58D9FFE02EAAF" box="[280,415,1468,1493]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="114.90256">114.90256E</geoCoordinate>
|
||
(Sungai Sii),
|
||
<date id="FFD43EDA984CFF94FDBB8D9FFD4DEAAE" box="[582,720,1468,1493]" pageId="30" pageNumber="231" value="2014-07-17">17 vii 2014</date>
|
||
, leg. R.W.J. Ngiam, to be deposited in
|
||
<collectionCode id="ED7B80DF984CFF94FB5F8D9FFB41EAAE" box="[1186,1244,1468,1492]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="30" pageNumber="231">ZRC</collectionCode>
|
||
;
|
||
<specimenCount id="9D6CD393984CFF94FB168D9FFA84EAAE" box="[1259,1305,1468,1492]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
, stream on Gunung Kalulong, Upper Baram, Miri Division,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FD428DC3FCAFEA82" box="[703,818,1504,1528]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="3.19038">3.19038N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FCC08DFCFC25EA82" box="[829,952,1503,1528]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="5" value="114.6968">114.6968E</geoCoordinate>
|
||
,
|
||
<quantity id="4C92B5FF984CFF94FC388DC3FBA7EA82" box="[965,1082,1504,1528]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="30" pageNumber="231" unit="m" value="750.0" valueMax="800.0" valueMin="700.0">700-800m</quantity>
|
||
,
|
||
<date id="FFD43EDA984CFF94FBBB8DC3FB57EA82" box="[1094,1226,1503,1528]" pageId="30" pageNumber="231" value="2010-07-17">17 vii 2010</date>
|
||
, leg. R.A. Dow, in collection Dow;
|
||
<specimenCount id="9D6CD393984CFF94FEAC8E27FEE7E966" box="[337,378,1540,1564]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
(semi-teneral), Deer Cave Stream upstream of waterfall, outside Garden of Eden,
|
||
<collectionCode id="ED7B80DF984CFF94FAF28E27FAFDE966" box="[1295,1376,1540,1564]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13141" name="Gros Morne National Park" pageId="30" pageNumber="231">GMNP</collectionCode>
|
||
, Miri Division,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FEF88E04FEEAE93A" box="[261,375,1575,1600]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="4.02785">4.02785N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FE7E8E0BFD91E93A" box="[387,524,1576,1600]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="114.83978">114.83978E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA984CFF94FDE58E0BFDE0E93A" box="[536,637,1576,1600]" pageId="30" pageNumber="231" value="2014-05-08">8 v 2014</date>
|
||
, leg. P.O.M. Steinhoff, in coll. Steinhoff;
|
||
<specimenCount id="9D6CD393984CFF94FBB38E0BFBE5E93A" box="[1102,1144,1576,1600]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
(teneral) same location,
|
||
<date id="FFD43EDA984CFF94FA738E0BFF7BE91E" pageId="30" pageNumber="231" value="2014-05-09">9 v 2014</date>
|
||
, leg. P.O.M. Steinhoff, in coll. Steinhoff;
|
||
<specimenCount id="9D6CD393984CFF94FD4E8E6FFD41E91E" box="[691,732,1612,1636]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
, parallel tributary of Deer Cave Stream, upstream of Garden of Eden Waterfall, same NP,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FE388E4CFDAAE9F2" box="[453,567,1647,1672]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="4.02656">4.02656N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FDB88E53FD53E9F2" box="[581,718,1648,1672]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="114.84297">114.84297E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA984CFF94FD218E53FCD8E9F2" box="[732,837,1648,1672]" pageId="30" pageNumber="231" value="2014-05-08">8 v 2014</date>
|
||
, leg. P.O.M. Steinhoff, in coll. Steinhoff;
|
||
<specimenCount id="9D6CD393984CFF94FADB8E53FACFE9F2" box="[1318,1362,1648,1672]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
, same location,
|
||
<date id="FFD43EDA984CFF94FEF98EB7FEE2E9D6" box="[260,383,1684,1708]" pageId="30" pageNumber="231" value="2014-05-22">22 v 2014</date>
|
||
, leg. P.O.M. Steinhoff, in coll. Steinhoff;
|
||
<specimenCount id="9D6CD393984CFF94FC928EB7FC00E9D6" box="[879,925,1684,1708]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
, stream systems at “camp 2”, lower slopes Gunung Mulu, same NP,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FE478E94FDB1E9AA" box="[442,556,1719,1744]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="1" value="4.04752">4.04752N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FDC48E94FD5EE9AA" box="[569,707,1719,1744]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="1" value="114.86622">114.86622E</geoCoordinate>
|
||
,
|
||
<date id="FFD43EDA984CFF94FD2D8E9BFCD5E9AA" box="[720,840,1720,1744]" pageId="30" pageNumber="231" value="2014-05-31">31 v 2014</date>
|
||
, leg. P.O.M. Steinhoff, in coll. Steinhoff;
|
||
<specimenCount id="9D6CD393984CFF94FADA8E9BFAC8E9AA" box="[1319,1365,1720,1744]" pageId="30" pageNumber="231" type="male">1 ♂</specimenCount>
|
||
, same location,
|
||
<date id="FFD43EDA984CFF94FEFD8EFFFEE3E98E" box="[256,382,1756,1781]" pageId="30" pageNumber="231" value="2014-06-19">19 vi 2014</date>
|
||
, leg. P.O.M. Steinhoff, in coll. Steinhoff;
|
||
<specimenCount id="9D6CD393984CFF94FCA58EFFFC1EE98E" box="[856,899,1756,1780]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
, second order tributary to Sungai Lansat, same NP,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FF388EDCFE81E862" box="[197,284,1791,1816]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="55" value="4.006">4.006N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FEDA8F23FE19E862" box="[295,388,1792,1816]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="555" value="114.82">114.82E</geoCoordinate>
|
||
(Long (mouth of) Lansat),
|
||
<date id="FFD43EDA984CFF94FD428F23FCDCE862" box="[703,833,1791,1816]" pageId="30" pageNumber="231" value="2014-07-23">23 vii 2014</date>
|
||
, leg. P.O.M. Steinhoff, in coll. Steinhoff.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B91984CFFA8FF3A8F07FD99E85B" lastPageId="34" lastPageNumber="235" pageId="30" pageNumber="231" type="description">
|
||
<paragraph id="8BD5181A984CFF94FF3A8F07FB15E8FE" blockId="30.[151,1437,279,2033]" pageId="30" pageNumber="231">
|
||
<materialsCitation id="3B021247984CFF94FF3A8F07FE41E81A" ID-GBIF-Occurrence="1702634970" collectingDate="1937-06" collectionCode="RMNH" collectorName="M. E. Walsh" country="Indonesia" latitude="0.977" location="Dagaunan" longLatPrecision="55" longitude="117.9803" municipality="Sangkulirang" pageId="30" pageNumber="231" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Timur">
|
||
<emphasis id="B91EC408984CFF94FF3A8F07FDAAE847" bold="true" box="[199,567,1828,1853]" pageId="30" pageNumber="231">
|
||
<collectingRegion id="49AED6F8984CFF94FF3A8F07FE36E847" box="[199,427,1828,1853]" country="Indonesia" name="Kalimantan Timur" pageId="30" pageNumber="231">Kalimantan Timur</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37D588A984CFF94FE448F07FDAFE847" box="[441,562,1828,1853]" name="Indonesia" pageId="30" pageNumber="231">Indonesia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D6CD393984CFF94FDBD8F07FDF1E846" box="[576,620,1828,1852]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
,
|
||
<location id="8EB54EC1984CFF94FD818F06FD6DE847" LSID="urn:lsid:plazi:treatment:03C3A90C984FFFA9FF6A8F04FAF5ED1A:8EB54EC1984CFF94FD818F06FD6DE847" box="[636,752,1829,1853]" country="Indonesia" latitude="0.977" longLatPrecision="55" longitude="117.9803" municipality="Sangkulirang" name="Dagaunan" pageId="30" pageNumber="231" stateProvince="Kalimantan Timur">Dagaunan</location>
|
||
,
|
||
<collectingMunicipality id="6BB18260984CFF94FD008F07FC08E847" box="[765,917,1828,1853]" pageId="30" pageNumber="231">Sangkulirang</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FC5F8F07FC65E846" box="[930,1016,1828,1852]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="55" value="0.977">0.977N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FBF88F07FBE3E846" box="[1029,1150,1828,1852]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="5" value="117.9803">117.9803E</geoCoordinate>
|
||
(Sangkulirang),
|
||
<collectingDate id="EF90C732984CFF94FAC28F07FA05E846" box="[1343,1432,1828,1853]" pageId="30" pageNumber="231" value="1937-06">
|
||
<date id="FFD43EDA984CFF94FAC28F07FA05E846" box="[1343,1432,1828,1853]" pageId="30" pageNumber="231" value="1937-06">vi 1937</date>
|
||
</collectingDate>
|
||
, leg.
|
||
<collectorName id="269F7DCC984CFF94FF378F6AFEC8E81A" box="[202,341,1863,1888]" pageId="30" pageNumber="231">M.E. Walsh</collectorName>
|
||
, in
|
||
<collectionCode id="ED7B80DF984CFF94FE798F6AFE41E81A" box="[388,476,1865,1888]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="231">RMNH</collectionCode>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B021247984CFF94FE178F6BFB19E8FE" ID-GBIF-Occurrence="1702634975" collectingDate="1956-09-30" collectionCode="RMNH" collectorName="A. M. R. Wegner" country="Indonesia" county="Tabang" elevation="125" latitude="0.5667" location="Tabang" longLatPrecision="7" longitude="116.0333" municipality="Bengen River" pageId="30" pageNumber="231" specimenCount="1" specimenCount-female="1" stateProvince="East Kalimantan">
|
||
<specimenCount id="9D6CD393984CFF94FE178F6BFD8AE81A" box="[490,535,1864,1888]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
(semi-teneral),
|
||
<location id="8EB54EC1984CFF94FD2D8F64FCB8E81A" LSID="urn:lsid:plazi:treatment:03C3A90C984FFFA9FF6A8F04FAF5ED1A:8EB54EC1984CFF94FD2D8F64FCB8E81A" box="[720,805,1863,1888]" country="Indonesia" county="Tabang" latitude="0.5667" longLatPrecision="7" longitude="116.0333" municipality="Bengen River" name="Tabang" pageId="30" pageNumber="231" stateProvince="East Kalimantan">Tabang</location>
|
||
,
|
||
<collectingMunicipality id="6BB18260984CFF94FCCE8F6AFC4EE81A" box="[819,979,1863,1889]" pageId="30" pageNumber="231">Bengen River</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="49AED6F8984CFF94FC1F8F6AFB3FE81A" box="[994,1186,1863,1888]" country="Indonesia" name="Kalimantan Timur" pageId="30" pageNumber="231">East Kalimantan</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FB4C8F64FA88E81A" box="[1201,1301,1863,1888]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="5" value="0.5667">0.5667N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FADE8F64FA01E81A" box="[1315,1436,1863,1888]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="5" value="116.0333">116.0333E</geoCoordinate>
|
||
(approximate coordinates for
|
||
<collectingCounty id="62B46096984CFF94FE1E8F4FFDA4E8FF" box="[483,569,1900,1925]" pageId="30" pageNumber="231">Tabang</collectingCounty>
|
||
),
|
||
<elevation id="0047FF29984CFF94FDB18F4FFD13E8FF" box="[588,654,1900,1925]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="30" pageNumber="231" unit="m" value="125.0">
|
||
<quantity id="4C92B5FF984CFF94FDB18F4FFD13E8FF" box="[588,654,1900,1925]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="30" pageNumber="231" unit="m" value="125.0">125m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EF90C732984CFF94FD678F4FFC88E8FF" box="[666,789,1900,1925]" pageId="30" pageNumber="231" value="1956-09-30">
|
||
<date id="FFD43EDA984CFF94FD678F4FFC88E8FF" box="[666,789,1900,1925]" pageId="30" pageNumber="231" value="1956-09-30">30 ix 1956</date>
|
||
</collectingDate>
|
||
, leg.
|
||
<collectorName id="269F7DCC984CFF94FCAD8F4FFB98E8FF" box="[848,1029,1900,1925]" pageId="30" pageNumber="231">A.M.R. Wegner</collectorName>
|
||
, in
|
||
<collectionCode id="ED7B80DF984CFF94FBD18F4EFB19E8FE" box="[1068,1156,1901,1924]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="231">RMNH</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A984CFF94FF3A8FACFA01E8B6" blockId="30.[151,1437,279,2033]" pageId="30" pageNumber="231">
|
||
<materialsCitation id="3B021247984CFF94FF3A8FACFA01E8B6" ID-GBIF-Occurrence="1702634969" collectingDate="1996-11-11" collectorName="M. Bedjanic & Dow." country="Indonesia" county="Banjarbaru" elevation="250" latitude="3.5059" location="Sungai Kuinam" longLatPrecision="7" longitude="115.0923" municipality="Aranio District" pageId="30" pageNumber="231" specimenCount="1" specimenCount-female="1" stateProvince="Kalimantan Selatan">
|
||
<emphasis id="B91EC408984CFF94FF3A8FACFDDCE8D2" bold="true" box="[199,577,1935,1960]" pageId="30" pageNumber="231">
|
||
<collectingRegion id="49AED6F8984CFF94FF3A8FACFE2BE8D2" box="[199,438,1935,1960]" country="Indonesia" name="Kalimantan Selatan" pageId="30" pageNumber="231">Kalimantan Selatan</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37D588A984CFF94FE3F8FACFDA6E8D2" box="[450,571,1935,1960]" name="Indonesia" pageId="30" pageNumber="231">Indonesia</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="9D6CD393984CFF94FDBA8FB3FDEDE8D2" box="[583,624,1936,1960]" pageId="30" pageNumber="231" type="female">1 ♀</specimenCount>
|
||
,
|
||
<location id="8EB54EC1984CFF94FD838FACFCB2E8D2" LSID="urn:lsid:plazi:treatment:03C3A90C984FFFA9FF6A8F04FAF5ED1A:8EB54EC1984CFF94FD838FACFCB2E8D2" box="[638,815,1935,1960]" country="Indonesia" county="Banjarbaru" latitude="3.5059" longLatPrecision="7" longitude="115.0923" municipality="Aranio District" name="Sungai Kuinam" pageId="30" pageNumber="231" stateProvince="Kalimantan Selatan">Sungai Kuinam</location>
|
||
,
|
||
<location id="8EB54EC1984CFF94FCC78FB3FC78E8D2" LSID="urn:lsid:plazi:treatment:03C3A90C984FFFA9FF6A8F04FAF5ED1A:8EB54EC1984CFF94FCC78FB3FC78E8D2" box="[826,997,1935,1960]" country="Indonesia" county="Banjarbaru" latitude="3.5059" longLatPrecision="7" longitude="115.0923" municipality="Aranio District" name="Gunung Pisang" pageId="30" pageNumber="231" stateProvince="Kalimantan Selatan">Gunung Pisang</location>
|
||
primary forest,
|
||
<locationDeviation id="9A670138984CFF94FB658FACFF7BE8B7" pageId="30" pageNumber="231">
|
||
<quantity id="4C92B5FF984CFF94FB658FACFB4DE8D2" box="[1176,1232,1935,1960]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="231" unit="km" value="5.0">5 km</quantity>
|
||
SSE of Belangian village
|
||
</locationDeviation>
|
||
,
|
||
<collectingMunicipality id="6BB18260984CFF94FF0C8F97FE00E8B7" box="[241,413,1972,1997]" pageId="30" pageNumber="231">Aranio District</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62B46096984CFF94FE558F97FDB5E8B7" box="[424,552,1972,1997]" pageId="30" pageNumber="231">Banjarbaru</collectingCounty>
|
||
, -
|
||
<geoCoordinate id="EE5E7EDD984CFF94FDC78F97FD02E8B7" box="[570,671,1972,1997]" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="5" value="3.5059">3.5059N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5E7EDD984CFF94FD568F97FCBBE8B7" box="[683,806,1972,1997]" direction="east" orientation="longitude" pageId="30" pageNumber="231" precision="5" value="115.0923">115.0923E</geoCoordinate>
|
||
,
|
||
<elevation id="0047FF29984CFF94FCCF8F97FCE9E8B7" box="[818,884,1972,1997]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="30" pageNumber="231" unit="m" value="250.0">
|
||
<quantity id="4C92B5FF984CFF94FCCF8F97FCE9E8B7" box="[818,884,1972,1997]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="30" pageNumber="231" unit="m" value="250.0">250m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EF90C732984CFF94FC7D8F97FC64E8B7" box="[896,1017,1972,1997]" pageId="30" pageNumber="231" value="1996-11-11">
|
||
<date id="FFD43EDA984CFF94FC7D8F97FC64E8B7" box="[896,1017,1972,1997]" pageId="30" pageNumber="231" value="1996-11-11">11 xi 1996</date>
|
||
</collectingDate>
|
||
, leg.
|
||
<collectorName id="269F7DCC984CFF94FBC88F96FB5FE8B7" box="[1077,1218,1972,1997]" pageId="30" pageNumber="231">M. Bedjanič</collectorName>
|
||
, in collection
|
||
<collectorName id="269F7DCC984CFF94FAA28F96FA01E8B6" box="[1375,1436,1973,1996]" pageId="30" pageNumber="231">Dow.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD5181A984CFF95FF3A8FFBFD39EE1F" blockId="30.[151,1437,279,2033]" lastBlockId="31.[151,1437,151,357]" lastPageId="31" lastPageNumber="232" pageId="30" pageNumber="231">
|
||
<emphasis id="B91EC408984CFF94FF3A8FFBFCA7E88B" bold="true" box="[199,826,2008,2033]" pageId="30" pageNumber="231">Description of male (based on SAR11_12_GOM56).</emphasis>
|
||
<emphasis id="B91EC408984CFF94FCB98FFBFC1DE88B" box="[836,896,2008,2033]" italics="true" pageId="30" pageNumber="231">Head</emphasis>
|
||
(
|
||
<figureCitation id="1351049F984CFF94FC6F8FFBFC40E88A" box="[914,989,2008,2033]" captionStart="FIGURES 4–9" captionStartId="9.[151,264,1713,1736]" captionTargetBox="[151,1436,193,1692]" captionTargetId="figure@9.[151,1436,193,1692]" captionTargetPageId="9" captionText="FIGURES 4–9. Male head dorsal: (4) L. coomansi SAR09_10_GOM4; (5) L. pendleburyi SAR13_14_GOM13; (6) L. pasia SAB12_GOM1; (7) L. sii holotype; (8) L.schieli holotype; (9) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068004/files/figure.png" pageId="30" pageNumber="231">Figs 9</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984CFF94FC118FFBFB96E88A" box="[1004,1035,2008,2032]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1933,1956]" captionTargetBox="[151,1436,841,1912]" captionTargetId="figure@16.[151,1436,841,1912]" captionTargetPageId="16" captionText="FIGURES 14–17. Male head frontal: (14) L. pasia SAB12_GOM1; (15) L. sii holotype; (16) L. schieli holotype; (17) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="30" pageNumber="231">17</figureCitation>
|
||
). Median lobe labium dark brown, pale basally except laterally; lateral lobes pale, hooks dark brown and black. Labrum black with pale transverse basal mark. Mandible bases pale, genae pale adjacent to mandible bases, otherwise black. Clypeus mostly black with rather indistinct brown markings at anterior corners of postclypeus, very faint mark centre anteclypeus. Ante- and postfrons not very sharply divided, postfrons almost entirely pale bluish (
|
||
<figureCitation id="1351049F984DFF95FBC88927FB13EE66" box="[1077,1166,260,285]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1933,1956]" captionTargetBox="[151,1436,841,1912]" captionTargetId="figure@16.[151,1436,841,1912]" captionTargetPageId="16" captionText="FIGURES 14–17. Male head frontal: (14) L. pasia SAB12_GOM1; (15) L. sii holotype; (16) L. schieli holotype; (17) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="31" pageNumber="232">Fig. 17</figureCitation>
|
||
). Pair of prominent ca transverse tubercles behind lateral ocelli; ocelli orange. Vertex and occiput black with large pale mark on vertex behind and extending onto rear parts tubercles.
|
||
</paragraph>
|
||
<caption id="DF154892984DFF95FF6A8DFEFCD0E968" httpUri="https://zenodo.org/record/1068024/files/figure.png" pageId="31" pageNumber="232" targetBox="[151,1436,409,1479]" targetPageId="31">
|
||
<paragraph id="8BD5181A984DFF95FF6A8DFEFCD0E968" blockId="31.[151,1436,1501,1555]" pageId="31" pageNumber="232">
|
||
<emphasis id="B91EC408984DFF95FF6A8DFEFECAEA89" bold="true" box="[151,343,1501,1524]" pageId="31" pageNumber="232">FIGURES 52–55.</emphasis>
|
||
Male synthorax dorsal: (52)
|
||
<taxonomicName id="4C6A6399984DFF95FD708DFCFC91EA8E" authorityName="Laidlaw" authorityYear="1936" box="[653,780,1503,1524]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408984DFF95FD708DFCFC91EA8E" box="[653,780,1503,1524]" italics="true" pageId="31" pageNumber="232">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
SAR09_10_GOM4; (53)
|
||
<taxonomicName id="4C6A6399984DFF95FBE38DFCFB00EA8E" authorityName="Laidlaw" authorityYear="1936" box="[1054,1181,1503,1524]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408984DFF95FBE38DFCFB00EA8E" box="[1054,1181,1503,1524]" italics="true" pageId="31" pageNumber="232">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
holotype; (54)
|
||
<emphasis id="B91EC408984DFF95FAB98DFCFAC4EA8E" box="[1348,1369,1503,1524]" italics="true" pageId="31" pageNumber="232">L.</emphasis>
|
||
sp. cf
|
||
<taxonomicName id="4C6A6399984DFF95FF6A8DDDFF65E969" authorityName="Laidlaw" authorityYear="1936" box="[151,248,1534,1555]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC408984DFF95FF6A8DDDFF65E969" box="[151,248,1534,1555]" italics="true" pageId="31" pageNumber="232">coomansi</emphasis>
|
||
</taxonomicName>
|
||
SAB12_GOM4; (55)
|
||
<taxonomicName id="4C6A6399984DFF95FE268DDDFDF2E968" authorityName="Steinhoff" authorityYear="2015" box="[475,623,1532,1555]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="31" pageNumber="232" phylum="Arthropoda" rank="species" species="pendleburyi">
|
||
<emphasis id="B91EC408984DFF95FE268DDDFDF2E968" box="[475,623,1532,1555]" italics="true" pageId="31" pageNumber="232">L. pendleburyi</emphasis>
|
||
</taxonomicName>
|
||
SAR13_14_GOM13.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD5181A984DFF95FF3A8E19FA00E8A4" blockId="31.[151,1437,1594,2014]" pageId="31" pageNumber="232">
|
||
<emphasis id="B91EC408984DFF95FF3A8E19FE85E929" box="[199,280,1594,1619]" italics="true" pageId="31" pageNumber="232">Thorax</emphasis>
|
||
. Prothorax mostly dark brown, pale transverse stripe anteriorly on anterior pronotal lobe. Pair of lateral posterior pale marks, central posterior mark (divided centrally), pair small pale marks, widely separated dorsalanteriorly on middle pronotal lobe. Propleuron mottled dark brown and pale. Tiny central pale mark on posterior pronotal lobe. Synthorax dark brown with pale yellow-green markings as follows (
|
||
<figureCitation id="1351049F984DFF95FB5B8E86FA95E9C4" box="[1190,1288,1701,1726]" captionStart="FIGURES 56–59" captionStartId="33.[151,264,1277,1300]" captionTargetBox="[156,1432,193,1256]" captionTargetId="figure@33.[156,1432,193,1256]" captionTargetPageId="33" captionText="FIGURES 56–59. Male synthorax dorsal: (56) L. pasia SAB12_GOM1; (57) L. sii holotype; (58) L. schieli holotype; (59) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068026/files/figure.png" pageId="31" pageNumber="232">Figs 59</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F984DFF95FAE38E86FAA3E9C4" box="[1310,1342,1701,1726]" captionStart="FIGURES 60–63" captionStartId="34.[151,264,1601,1624]" captionTargetBox="[156,1432,517,1580]" captionTargetId="figure@34.[156,1432,517,1580]" captionTargetPageId="34" captionText="FIGURES 60–63. Male synthorax lateral: (60) L. coomansi SAR09_10_GOM4; (61) L. sp. cf coomansi SAB12_GOM4; (62) L. schieli holotype; (63) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068028/files/figure.png" pageId="31" pageNumber="232">63</figureCitation>
|
||
): broad mesothoracic collar, just divided at middorsal carina, separated from short narrow dorsal thoracic stripes that extend just past apex of antealar crest. Broad antehumeral stripes. Most of rest of synthorax laterally pale, except a broad brown stripe around mesopleural suture, small brown area on metepisternum above spiracle, ca diagonal brown mark centrally on metepisternum and another brown mark adjacent to antealar carina, Metepimeron brown below metakatepisternum, this extended along lower margin to antealar carina. Mesokatepisternum dark brown above, pale below contiguous with antehumeral stripes, metakatepisternum mostly pale. Metapoststernum pale with brown lateral marks in anterior ca two-thirds. Legs robust and relatively short. Coxae mottled pale, grey and brown, trochanters black and pale. Rest of legs black except inner surface of anterior femora with pale stripe.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9872FFAAFF6A88B4FC9FEE67" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="233">
|
||
Wings: sectors of arculus separated at origin with 6 (right) or 5 (left) cross veins up to and at first bifurcation of superior sector in Fw, 4 (right) or 3 (left) in Hw. Discoidal field with 2 rows of cells from origin, transitioning to just three rows well before level of nodus in both wings. 14 Ax in Fw,
|
||
<quantity id="4C92B5FF9872FFAAFC4988C3FC42EF82" box="[948,991,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="32" pageNumber="233" unit="in" value="9.0">9 in</quantity>
|
||
Hw, 11 (left) or 12 (right) Px in Fw,
|
||
<quantity id="4C92B5FF9872FFAAFA7C88C3FF30EE67" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="32" pageNumber="233" unit="in" value="11.0">11 in</quantity>
|
||
Hw. Pt pale brown, covering ca 3-4 underlying cells.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9872FFAAFF3A8904FBC4EB6A" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="233">
|
||
<emphasis id="B91EC4089872FFAAFF3A8904FEACEE3A" box="[199,305,295,320]" italics="true" pageId="32" pageNumber="233">Abdomen</emphasis>
|
||
. Slender after base of S3, expanding moderately from base of S7, maximum width and height reached apical part of S8, then almost constant. Very dark brown to black with pale markings as follows: S1 mostly pale laterally, obscure pale marks dorsally. S2 central lateral pale mark running from base, including auricle, narrowly divided from posterior yellow mark that extends anteriorly along lower margin tergite, narrow yellowish middorsal stripe not quite reaching apex of segment, wider apically. S3 with basal lateral pale marking, S4–7 with small lateral markings at base. S3–7 with narrow, irregular middorsal line, on S7 slight expanded apically. S8 pale brown behind posterior carina. S10 distinctive yellow-green subcircular mark centrally on dorsum. Cerci (
|
||
<figureCitation id="1351049F9872FFAAFACE89DCFA0AED62" box="[1331,1431,511,536]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="32" pageNumber="233">Figs 105</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9872FFAAFF6A8A07FF5DED46" box="[151,192,548,572]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="32" pageNumber="233">111</figureCitation>
|
||
) black and dark brown, broad at base in lateral view (
|
||
<figureCitation id="1351049F9872FFAAFCC68A07FC00ED46" box="[827,925,548,573]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="32" pageNumber="233">Fig. 111</figureCitation>
|
||
), tapering to sharp upturned tip, small basal dorsal mound visible, upper margin sloping steeply down from this to after mid-length, then gently curved to apex, along lower margin sloping down from base to ca half length, then abruptly up and in to apex, very prominent corner at upturn, first half of upturned section of lower margin bearing small teeth. In dorsal view (
|
||
<figureCitation id="1351049F9872FFAAFB0E8AACFACEEDD2" box="[1267,1363,655,680]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="32" pageNumber="233">Fig. 105</figureCitation>
|
||
) outer margin running straight from base or slightly outward, then inward to tips at ca half length; prominent corner on lower margin clearly visible. Black, brown at base laterally, this more extensive dorsally. Epiproct (
|
||
<figureCitation id="1351049F9872FFAAFB058AF4FAC0ED8A" box="[1272,1373,727,752]" captionStart="FIGURES 100–105" captionStartId="42.[151,264,1215,1238]" captionTargetBox="[163,1392,207,1164]" captionTargetId="figure@42.[163,1439,186,1178]" captionTargetPageId="42" captionText="FIGURES 100–105. Male anal appendages dorsal view (all scale bars 0.5mm): (100) L. coomansi SAR09_10_GOM4; (101) L. pasia SAB12_GOM1; (102) L. pendleburyi SAR13_14_GOM13; (103) L. schieli holotype; (104) L. sii holotype; (105) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068044/files/figure.png" pageId="32" pageNumber="233">Figs 105</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9872FFAAFA978AFBFA09ED8A" box="[1386,1428,728,752]" captionStart="FIGURES 112–117" captionStartId="44.[151,264,1808,1831]" captionTargetBox="[156,1432,193,1787]" captionTargetId="figure@44.[156,1432,193,1787]" captionTargetPageId="44" captionText="FIGURES 112–117. Male anal appendages in ventral view (all scale bars 0.5mm): (112) L. coomansi SAR09_10_GOM4; (113) L.pasia SAB12_GOM1;(114) L.pendleburyi SAR13_14_GOM13; (115) L. schieli holotype; (116) L.sii holotype; (117) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068048/files/figure.png" pageId="32" pageNumber="233">117</figureCitation>
|
||
) black and dark brown, pale central basal area ventrally, just shorter than cerci, deeply divided in an almost semicircular shape, outer margins gently divergent to near apices, apices rounded. In lateral view (
|
||
<figureCitation id="1351049F9872FFAAFACC8B3CFA09EC42" box="[1329,1428,799,824]" captionStart="FIGURES 106–111" captionStartId="43.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure@43.[151,1436,193,1799]" captionTargetPageId="43" captionText="FIGURES 106–111. Male anal appendages and abdominal S10 in lateral view (all scale bars 0.5mm): (106) L. coomansi SAR09_10_GOM4; (107) L. pasia SAB12_GOM1; (108) L. pendleburyi SAR13_14_GOM13; (109) L. schieli holotype; (110) L. sii holotype; (111) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068046/files/figure.png" pageId="32" pageNumber="233">Fig. 111</figureCitation>
|
||
) directed down from base then up before mid-length, apices just under tips of cerci, turned almost to rear; basal part of upper margin obscured by cerci. Accessory genitalia as shown in
|
||
<figureCitation id="1351049F9872FFAAFC468B44FBAAECFA" box="[955,1079,871,896]" captionStart="FIGURES 88–93" captionStartId="40.[151,264,1280,1303]" captionTargetBox="[156,1432,195,1258]" captionTargetId="figure@40.[156,1432,195,1258]" captionTargetPageId="40" captionText="FIGURES 88–93. Male accessory genitalia, lateral view, all scale bars 1mm: (88) L. coomansi SAR09_10_GOM4; (89) L. pasia SAB12_GOM1; (90) L. pendleburyi SAR13_14_GOM13; (91) L. schieli holotype; (92) L. sii holotype; (93) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068040/files/figure.png" pageId="32" pageNumber="233">Figures 93</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9872FFAAFBBA8B4BFBFEECFA" box="[1095,1123,872,896]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="32" pageNumber="233">99</figureCitation>
|
||
(
|
||
<figureCitation id="1351049F9872FFAAFB8B8B44FB57ECFA" box="[1142,1226,871,896]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="32" pageNumber="233">Fig. 99</figureCitation>
|
||
shows a different specimen), with anterior hamule moderate sized, tapering toward sharp apex, slightly hooked to rear at apex. Posterior hamule large, ca twice length of anterior hamule, directed postero-ventrally, anterior margin converging on posterior margin for much of length, then whole curving gently forward so directed down, tapering to sharp tip, directed slightly outward. Penis vesicle prominent, ca half height of posterior hamule.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9872FFAAFF3A8C3EFB08EB4F" blockId="32.[151,1437,151,2013]" box="[199,1173,1052,1077]" pageId="32" pageNumber="233">Measurements (mm). Hw 25.5, abdomen excluding anal appendages 29.5, cerci ca 1.5.</paragraph>
|
||
<paragraph id="8BD5181A9872FFAAFF3A8C62FD37EB92" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="233">
|
||
<emphasis id="B91EC4089872FFAAFF3A8C62FE98EB22" box="[199,261,1089,1112]" italics="true" pageId="32" pageNumber="233">Penis</emphasis>
|
||
. Examined in SAR16_GOM1 (
|
||
<figureCitation id="1351049F9872FFAAFD948C1CFD20EB22" box="[617,701,1087,1112]" captionStart="FIGURES 94–99" captionStartId="41.[151,264,1932,1955]" captionTargetBox="[156,1432,529,1911]" captionTargetId="figure@41.[156,1432,529,1911]" captionTargetPageId="41" captionText="FIGURES 94–99. Penis in lateral view (all scale bars 0.5mm): (94) L. coomansi SAR07_8_GOM18; (95) L. pasia SAB12_GOM1; (96) L. pendleburyi SAB12_GOM3; (97)L. schieli holotype; (98)L. sii paratype SAR13_14_GOM49; (99) L. williamsoni SAR16_GOM1." httpUri="https://zenodo.org/record/1068042/files/figure.png" pageId="32" pageNumber="233">Fig. 99</figureCitation>
|
||
), very similar to that of
|
||
<taxonomicName id="4C6A63999872FFAAFC328C62FBABEB22" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[975,1078,1087,1112]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="schieli">
|
||
<emphasis id="B91EC4089872FFAAFC328C62FBABEB22" box="[975,1078,1087,1112]" italics="true" pageId="32" pageNumber="233">L. schieli</emphasis>
|
||
</taxonomicName>
|
||
. Penultimate segment extended to rear of join with terminal segment, bifurcated in this part, apices slender, curved down, shorter than in
|
||
<taxonomicName id="4C6A63999872FFAAFAD38C46FA08EB07" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[1326,1429,1124,1149]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="233" phylum="Arthropoda" rank="species" species="schieli">
|
||
<emphasis id="B91EC4089872FFAAFAD38C46FA08EB07" box="[1326,1429,1124,1149]" italics="true" pageId="32" pageNumber="233">L. schieli</emphasis>
|
||
</taxonomicName>
|
||
. Terminal segment extended well beyond penultimate segment, approximately dumbbell shaped in lateral view, then shallowly contracted before strongly expanding dorsally, apically with short rearward directed, curled, cornua; short, rearward directed flap just below cornua.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9872FFAAFF3A8CD7FE64E8EF" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="233">
|
||
<emphasis id="B91EC4089872FFAAFF3A8CD7FDBFEA77" bold="true" box="[199,546,1268,1293]" pageId="32" pageNumber="233">Variation in examined males.</emphasis>
|
||
There are frequently well-defined yellow markings on the clypeus, particularly in teneral and semi-teneral individuals; this is the case in the
|
||
<typeStatus id="54D1A6B89872FFAAFCB38D34FC2DEA4A" box="[846,944,1303,1328]" pageId="32" pageNumber="233" type="holotype">holotype</typeStatus>
|
||
where there are lower lateral marks and an upper central mark on the post clypeus. There is no yellow mark on the posterior pronotal lobe in any other individual, and the smaller, dorsal-anterior marks on the middle pronotal lobe are frequently absent, but sometimes joined to the larger lateral ones. The dorsal marking on the head is frequently smaller, and sometimes divided into two. The dorsal thoracic stripes are sometimes shorter than in
|
||
<figureCitation id="1351049F9872FFAAFC888D84FC75EABA" box="[885,1000,1447,1472]" captionStart="FIGURES 56–59" captionStartId="33.[151,264,1277,1300]" captionTargetBox="[156,1432,193,1256]" captionTargetId="figure@33.[156,1432,193,1256]" captionTargetPageId="33" captionText="FIGURES 56–59. Male synthorax dorsal: (56) L. pasia SAB12_GOM1; (57) L. sii holotype; (58) L. schieli holotype; (59) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068026/files/figure.png" pageId="32" pageNumber="233">Figure 59</figureCitation>
|
||
. There are frequently more extensive brown markings on the metepisternum. Most mature individuals have a dark central area on the metapoststernum near the abdomen. The coxae are entirely yellowish in the
|
||
<typeStatus id="54D1A6B89872FFAAFCDA8DCCFC10E972" box="[807,909,1519,1544]" pageId="32" pageNumber="233" type="holotype">holotype</typeStatus>
|
||
. There are often pale stripes on the upper inner surface of the anterior femur, and less often pale marks in various positions on the middle and posterior femur in mature individuals; in teneral individuals the legs are extensively pale. In some mature individuals Pt is darker than in the male described. The yellow middorsal mark on S7 is often not at all expanded apically and in the
|
||
<typeStatus id="54D1A6B89872FFAAFAC78E7FFA01E90F" box="[1338,1436,1628,1653]" pageId="32" pageNumber="233" type="holotype">holotype</typeStatus>
|
||
and one semi-teneral from GMNP there is a narrow yellow middorsal mark on S8 as well. Small, lateral apical markings are occasionally present, especially in teneral and semi-teneral individuals, on any of S7, S8 or S10 (and are presumably possible on S9). In the
|
||
<typeStatus id="54D1A6B89872FFAAFDAF8EE4FD29E99A" box="[594,692,1735,1760]" pageId="32" pageNumber="233" type="holotype">holotype</typeStatus>
|
||
there is a fairly large lateral marking on S8, approximately in the form of an elongate triangle based at the apex of the segment, but there is no basal mark on S7. The dorsal pale mark on S10 is sometimes smaller, in one individual it is divided and in the non-teneral male from Nanga Bloh it is completely absent. In the
|
||
<typeStatus id="54D1A6B89872FFAAFE458F17FD87E837" box="[440,538,1844,1869]" pageId="32" pageNumber="233" type="holotype">holotype</typeStatus>
|
||
the cerci and epiproct have substantial paler brown areas. The shape of the area between the arms of the epiproct varies from that in the male described here to a broad “v” shape (this is the condition seen in the
|
||
<typeStatus id="54D1A6B89872FFAAFE7B8F5FFE70E8EF" box="[390,493,1916,1941]" pageId="32" pageNumber="233" type="holotype">holotype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9872FFAAFF3A8F82FD9EE8A6" blockId="32.[151,1437,151,2013]" pageId="32" pageNumber="233">
|
||
Measurements (mm). Hw 24–26, abdomen excluding anal appendages 29–31, 13–15 Ax in Fw,
|
||
<quantity id="4C92B5FF9872FFAAFAED8F83FAFBE8C3" box="[1296,1382,1952,1977]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" pageId="32" pageNumber="233" unit="in" value="10.0" valueMax="11.0" valueMin="9.0">9–11 in</quantity>
|
||
Hw, 10–13 Px in Fw, 8–11 Px in Hw.
|
||
</paragraph>
|
||
<caption id="DF1548929873FFABFF6A8CDEFE76EA48" httpUri="https://zenodo.org/record/1068026/files/figure.png" pageId="33" pageNumber="234" targetBox="[156,1432,193,1256]" targetPageId="33">
|
||
<paragraph id="8BD5181A9873FFABFF6A8CDEFE76EA48" blockId="33.[151,1437,1277,1330]" pageId="33" pageNumber="234">
|
||
<emphasis id="B91EC4089873FFABFF6A8CDEFEC8EA69" bold="true" box="[151,341,1277,1300]" pageId="33" pageNumber="234">FIGURES 56–59.</emphasis>
|
||
Male synthorax dorsal: (56)
|
||
<taxonomicName id="4C6A63999873FFABFD828CDCFD4FEA6E" authorityName="van Tol" authorityYear="1990" box="[639,722,1279,1300]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="pasia">
|
||
<emphasis id="B91EC4089873FFABFD828CDCFD4FEA6E" box="[639,722,1279,1300]" italics="true" pageId="33" pageNumber="234">L. pasia</emphasis>
|
||
</taxonomicName>
|
||
SAB12_GOM1; (57)
|
||
<taxonomicName id="4C6A63999873FFABFC4A8CDCFC76EA6E" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[951,1003,1279,1300]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="sii">
|
||
<emphasis id="B91EC4089873FFABFC4A8CDCFC76EA6E" box="[951,1003,1279,1300]" italics="true" pageId="33" pageNumber="234">L. sii</emphasis>
|
||
</taxonomicName>
|
||
holotype; (58)
|
||
<taxonomicName id="4C6A63999873FFABFB778CDCFB75EA69" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[1162,1256,1277,1300]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="schieli">
|
||
<emphasis id="B91EC4089873FFABFB778CDCFB75EA69" box="[1162,1256,1277,1300]" italics="true" pageId="33" pageNumber="234">L. schieli</emphasis>
|
||
</taxonomicName>
|
||
holotype; (59)
|
||
<taxonomicName id="4C6A63999873FFABFA7A8CDCFE90EA48" authorityName="Laidlaw" authorityYear="1912" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="234" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089873FFABFA7A8CDCFE90EA48" italics="true" pageId="33" pageNumber="234">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
SAR11_12_GOM56.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD5181A9873FFABFF3A8D7AFCE4E930" blockId="33.[151,1437,1369,2006]" pageId="33" pageNumber="234">
|
||
<emphasis id="B91EC4089873FFABFF3A8D7AFDCAEA08" bold="true" box="[199,599,1369,1394]" pageId="33" pageNumber="234">Descriptive notes on the female.</emphasis>
|
||
<bibRefCitation id="EFFB65EB9873FFABFD9E8D7AFCBDEA08" author="Lieftinck" box="[611,800,1369,1394]" pageId="33" pageNumber="234" refString="Lieftinck, M. A. (1948) Descriptions and records of South - east Asiatic Odonata. Treubia, 19, 221 - 278." type="journal article" year="1948">Lieftinck (1948)</bibRefCitation>
|
||
gave a detailed description of a female from eastern
|
||
<collectingRegion id="49AED6F89873FFABFF6A8D5EFE87EAEC" box="[151,282,1405,1430]" country="Indonesia" name="Kalimantan Utara" pageId="33" pageNumber="234">Kalimantan</collectingRegion>
|
||
(the female from Dagaunan listed above), including illustrations of the dorsum of the head, the markings of the thorax and the vulvar scale. The other females that we have examined agree well with Lieftinck’s description in most respects; females from south-west
|
||
<collectingRegion id="49AED6F89873FFABFCFF8DE6FCFFEAA4" box="[770,866,1477,1502]" country="Malaysia" name="Sarawak" pageId="33" pageNumber="234">
|
||
<collectingCountry id="F37D588A9873FFABFCFF8DE6FCFFEAA4" box="[770,866,1477,1502]" name="Malaysia" pageId="33" pageNumber="234">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
are discussed separately below. There is variation of course, with individuals differing in some details of markings (similar to the variation seen in males), wing venation and in measurements. There are two main areas of difference from the female described by Lieftinck: the markings of the abdomen and the structures on the occipital plate.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9873FFABFF3A8E76FBBCE8F4" blockId="33.[151,1437,1369,2006]" pageId="33" pageNumber="234">
|
||
<bibRefCitation id="EFFB65EB9873FFABFF3A8E76FE5CE914" author="Lieftinck" box="[199,449,1621,1646]" pageId="33" pageNumber="234" refString="Lieftinck, M. A. (1948) Descriptions and records of South - east Asiatic Odonata. Treubia, 19, 221 - 278." type="journal article" year="1948">Lieftinck (1948: 249)</bibRefCitation>
|
||
described the markings of the abdomen of his female as follows: “Segm. 1 largely yellowish laterally, 2 without visible light side-markings (apparently discoloured); 3–7 each with a fairly large, oval, basal lateral yellow spot. 4 and 5 moreover with a longitudinal streak of the same colour placed on each side of segment a little distal to the middle, and 2–7 with a continuous middorsal line, widest on 2. Terminal segments black.” In all specimens examined by us, S1 is yellow dorsally as well as laterally, S2 has a lateral yellow stripe running the length of the segment and including the small, rounded auricle. S3 has a stripe running from base for most of the segment, narrowly divided before its midpoint. S4–7 have the basal lateral spot; the “longitudinal streak” is present on S4–6 and usually on S7, but diminishes in size on successive segments, so that it is typically a small spot on S6 and S7. The dorsal markings agree with Lieftinck’s description.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9873FFA8FF3A8FB9FD8FEEAA" blockId="33.[151,1437,1369,2006]" lastBlockId="34.[151,1437,151,464]" lastPageId="34" lastPageNumber="235" pageId="33" pageNumber="234">
|
||
In the female described and illustrated in
|
||
<bibRefCitation id="EFFB65EB9873FFABFD648FBAFCC9E8C8" author="Lieftinck" box="[665,852,1945,1970]" pageId="33" pageNumber="234" refString="Lieftinck, M. A. (1948) Descriptions and records of South - east Asiatic Odonata. Treubia, 19, 221 - 278." type="journal article" year="1948">Lieftinck (1948)</bibRefCitation>
|
||
, and that shown in
|
||
<figureCitation id="1351049F9873FFABFBD28FBAFB35E8C8" box="[1071,1192,1945,1970]" captionStart="FIGURES 24–27" captionStartId="20.[151,264,1441,1464]" captionTargetBox="[163,1439,268,1402]" captionTargetId="figure@20.[163,1439,268,1402]" captionTargetPageId="20" captionText="FIGURES 24–27. Female head dorsal: (24) L pendleburyi SAR16_GOM8; (25) L. schieli paratype SAR15_GOM6; (26) L. williamsoni SAR13_14_GOM51; (27) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068012/files/figure.png" pageId="33" pageNumber="234">Figures 26</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9873FFABFB498FB9FB4FE8C8" box="[1204,1234,1946,1970]" captionStart="FIGURES 32–35" captionStartId="23.[151,264,1357,1380]" captionTargetBox="[151,1436,265,1336]" captionTargetId="figure@23.[151,1436,265,1336]" captionTargetPageId="23" captionText="FIGURES 32–35. Female head dorsal detail; h—occipital horn; p—pit; s—occipital spur; t—tubercle: (32) L pendleburyi SAR16_GOM8; (33) L. schieli paratype SAR15_GOM6; (34) L. williamsoni SAR13_14_GOM51; (35) L. sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068016/files/figure.png" pageId="33" pageNumber="234">34</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9873FFABFB238FB9FB61E8C8" box="[1246,1276,1946,1970]" captionStart="FIGURES 42–45" captionStartId="27.[151,264,1498,1521]" captionTargetBox="[163,1439,186,1462]" captionTargetId="figure@27.[163,1439,186,1462]" captionTargetPageId="27" captionText="FIGURES 42–45. Female head frontal: (42) L pendleburyi SAR16_GOM8; (43) L. schieli paratype SAR15_GOM6; (44) L. williamsoni SAR13_14_GOM51; (45) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068020/files/figure.png" pageId="33" pageNumber="234">44</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9873FFABFAF48FBAFAB8E8C8" box="[1289,1317,1945,1970]" captionStart="FIGURES 46–51" captionStartId="29.[151,264,1672,1695]" captionTargetBox="[156,1432,375,1651]" captionTargetId="figure@29.[156,1432,375,1651]" captionTargetPageId="29" captionText="FIGURES 46–51. Female head frontal detail; h—occipital horn; ot—occipital tube; t—tubercle; th—tubercle horn: (46) L. coomansi SAR11_12_GOM48; (47) L pendleburyi SAR16_GOM8; (48) L. schieli paratype SAR15_GOM6; (49) L. species SAR11_12_GOM33; (50) L. williamsoni SAR13_14_GOM51; (51) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068022/files/figure.png" pageId="33" pageNumber="234">50</figureCitation>
|
||
here, both occipital horns and occipital spurs are present. The occipital spurs are short, robust and widely separated; they are present in all females we have examined, although they show some variation in length. The occipital horns, long and sharp in those figured and in Lieftinck’s illustration, are entirely absent in five of the
|
||
<specimenCount id="9D6CD3939870FFA8FB48889FFAAFEFAF" box="[1205,1330,188,213]" pageId="34" pageNumber="235" type="female">17 females</specimenCount>
|
||
we have examined, and only present on one side in another four. Close examination gives no indication that the horns have been broken in these cases; they are evidently undeveloped. In other individuals one or both horns are clearly broken, but there is also some variation in length. In the female from Tubau the basal half of the horns is swollen and brown. Between the occipital horns there are a pair of gentle swellings on the occipital ridge, visible in Lieftinck’s (1948) illustration, but not mentioned in his description. These are typically present but variably developed, and most conspicuous in females that lack the horns. There is a shallow pit or depression outside of the tubercles behind the lateral ocelli.
|
||
</paragraph>
|
||
<caption id="DF1548929870FFA8FF6A8E62FD62E90C" httpUri="https://zenodo.org/record/1068028/files/figure.png" pageId="34" pageNumber="235" targetBox="[156,1432,517,1580]" targetPageId="34">
|
||
<paragraph id="8BD5181A9870FFA8FF6A8E62FD62E90C" blockId="34.[151,1435,1601,1655]" pageId="34" pageNumber="235">
|
||
<emphasis id="B91EC4089870FFA8FF6A8E62FEC9E92D" bold="true" box="[151,340,1601,1624]" pageId="34" pageNumber="235">FIGURES 60–63.</emphasis>
|
||
Male synthorax lateral: (60)
|
||
<taxonomicName id="4C6A63999870FFA8FD838E60FD67E922" authorityName="Laidlaw" authorityYear="1936" box="[638,762,1603,1624]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC4089870FFA8FD838E60FD67E922" box="[638,762,1603,1624]" italics="true" pageId="34" pageNumber="235">L. coomansi</emphasis>
|
||
</taxonomicName>
|
||
SAR09_10_GOM4; (61)
|
||
<emphasis id="B91EC4089870FFA8FBF98E60FB84E922" box="[1028,1049,1603,1624]" italics="true" pageId="34" pageNumber="235">L.</emphasis>
|
||
sp. cf
|
||
<taxonomicName id="4C6A63999870FFA8FBA08E60FB23E922" authorityName="Laidlaw" authorityYear="1936" box="[1117,1214,1603,1624]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="coomansi">
|
||
<emphasis id="B91EC4089870FFA8FBA08E60FB23E922" box="[1117,1214,1603,1624]" italics="true" pageId="34" pageNumber="235">coomansi</emphasis>
|
||
</taxonomicName>
|
||
SAB12_GOM4; (62)
|
||
<taxonomicName id="4C6A63999870FFA8FF6A8E41FF69E90C" authorityName="Dow & Stokvis & Ngiam" authorityYear="2017" box="[151,244,1632,1655]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="schieli">
|
||
<emphasis id="B91EC4089870FFA8FF6A8E41FF69E90C" box="[151,244,1632,1655]" italics="true" pageId="34" pageNumber="235">L. schieli</emphasis>
|
||
</taxonomicName>
|
||
holotype; (63)
|
||
<taxonomicName id="4C6A63999870FFA8FE6D8E41FDBCE90C" authorityName="Laidlaw" authorityYear="1912" box="[400,545,1632,1655]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="235" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089870FFA8FE6D8E41FDBCE90C" box="[400,545,1632,1655]" italics="true" pageId="34" pageNumber="235">L. williamsoni</emphasis>
|
||
</taxonomicName>
|
||
SAR11_12_GOM56.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD5181A9870FFA8FF3A8EBCFAF0E9A0" blockId="34.[151,1436,1693,2006]" pageId="34" pageNumber="235">
|
||
In addition, there is some variation in the dorsal thoracic markings (
|
||
<figureCitation id="1351049F9870FFA8FC1C8EBEFBA6E9CC" box="[993,1083,1693,1718]" captionStart="FIGURES 74–77" captionStartId="37.[151,264,1356,1379]" captionTargetBox="[151,1435,194,1335]" captionTargetId="figure@37.[151,1436,193,1335]" captionTargetPageId="37" captionText="FIGURES 74–77. Female synthorax dorsal: (74) L pendleburyi SAR16_GOM8; (75) L. schieli paratype SAR15_GOM6; (76) L. williamsoni SAR13_14_GOM51; (77) L.sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068034/files/figure.png" pageId="34" pageNumber="235">Figs 76</figureCitation>
|
||
,
|
||
<figureCitation id="1351049F9870FFA8FBB78EBEFBF4E9CC" box="[1098,1129,1693,1718]" captionStart="FIGURES 84–87" captionStartId="39.[151,264,1427,1450]" captionTargetBox="[151,1435,194,1406]" captionTargetId="figure@39.[151,1436,193,1406]" captionTargetPageId="39" captionText="FIGURES 84–87. Female synthorax lateral: (84) L pendleburyi SAR16_GOM8; (85) L. schieli paratype SAR15_GOM6; (86) L. williamsoni SAR13_14_GOM51, right hand side, flipped for consistency;(87) L. sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068038/files/figure.png" pageId="34" pageNumber="235">86</figureCitation>
|
||
), for instance in the semiteneral female from Tabang the dorsal thoracic stripes are almost completely joined to the mesothoracic collar.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9870FFA8FF3A8EC4FD99E85B" blockId="34.[151,1436,1693,2006]" pageId="34" pageNumber="235">
|
||
Measurements (mm). Hw 25–28.5, abdomen excluding anal appendages 27–33, 13–17 Ax in Fw,
|
||
<quantity id="4C92B5FF9870FFA8FAEF8EC5FAF5E984" box="[1298,1384,1765,1790]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="34" pageNumber="235" unit="in" value="10.5" valueMax="12.0" valueMin="9.0">9–12 in</quantity>
|
||
Hw, 10–12 Px in Fw, 9–12 Px in Hw.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B919870FFA9FF3A8F0EFEBBEEAA" lastPageId="35" lastPageNumber="236" pageId="34" pageNumber="235" type="discussion">
|
||
<paragraph id="8BD5181A9870FFA8FF3A8F0EFEF0E8F4" blockId="34.[151,1436,1693,2006]" pageId="34" pageNumber="235">
|
||
<emphasis id="B91EC4089870FFA8FF3A8F0EFEA6E83C" bold="true" box="[199,315,1837,1862]" pageId="34" pageNumber="235">Remarks.</emphasis>
|
||
The fact that either two, one or no occipital horns are present on the occipital plate of females demonstrates that in this species the occipital horns are variable and non-diagnostic. The occipital spurs, however, are always present.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9870FFA8FF3A8FBAFC85E8AC" blockId="34.[151,1436,1693,2006]" pageId="34" pageNumber="235">
|
||
<bibRefCitation id="EFFB65EB9870FFA8FF3A8FBAFEEAE8C8" author="Laidlaw" box="[199,375,1945,1970]" pageId="34" pageNumber="235" refString="Laidlaw, F. F. (1912) List of the Odonata taken on an expedition to Mt. Batu Lawi together with descriptions of supposed new species. (Appendix VIII to Moulton, J. C., 1912.) Journal of the Straits Branch of the Royal Asiatic Society, 63, 92 - 99, 1 pl." type="journal article" year="1912">Laidlaw (1912)</bibRefCitation>
|
||
states that the abdomen plus anal appendages of the
|
||
<typeStatus id="54D1A6B89870FFA8FC1C8FBAFBDEE8C8" box="[993,1091,1945,1970]" pageId="34" pageNumber="235" type="holotype">holotype</typeStatus>
|
||
measure
|
||
<quantity id="4C92B5FF9870FFA8FB498FB9FA95E8CB" box="[1204,1288,1946,1970]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="34" pageNumber="235" unit="mm" value="34.0">34 mm</quantity>
|
||
, but the first author measured it as
|
||
<quantity id="4C92B5FF9870FFA8FE708F9DFE46E8AC" box="[397,475,1982,2006]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="34" pageNumber="235" unit="mm" value="31.0">31 mm</quantity>
|
||
excluding anal appendages.
|
||
</paragraph>
|
||
<paragraph id="8BD5181A9871FFA9FF3A88B4FEBBEEAA" blockId="35.[151,1436,151,608]" pageId="35" pageNumber="236">
|
||
<taxonomicName id="4C6A63999871FFA9FF3A88B4FE6BEFCA" authorityName="Laidlaw" authorityYear="1912" box="[199,502,151,176]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089871FFA9FF3A88B4FE6BEFCA" box="[199,502,151,176]" italics="true" pageId="35" pageNumber="236">Leptogomphus williamsoni</emphasis>
|
||
</taxonomicName>
|
||
exhibits considerable variation in its COI marker, with the five sequences available only agreeing in 90.6% of sites. With ITS, sequences from the same five specimens agree at 90.8% of sites, but this relatively low agreement is almost entirely due to the fact that most of the section of 28S included is excluded in one sample due to poor quality, and a smaller part of 16S in the same sample. When the comparison only includes the parts of 16S and 28S present in all sequences analysed, the sequences are identical at more than 99% of sites; this agreement is evident in
|
||
<figureCitation id="1351049F9871FFA9FE2F896FFDADEE1E" box="[466,560,332,357]" captionStart="FIGURE 2" captionStartId="5.[151,250,1294,1316]" captionTargetBox="[243,1384,200,1237]" captionTargetId="figure@5.[151,1436,193,1273]" captionTargetPageId="5" captionText="FIGURE 2. ITS gene tree for 20 specimens of Leptogomphus and three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068000/files/figure.png" pageId="35" pageNumber="236">Figure 2</figureCitation>
|
||
where the sequences appear almost identical. Given the variation seen in COI in this species, we do not consider that the difference seen in this marker with
|
||
<emphasis id="B91EC4089871FFA9FBFC8952FB85EEF2" box="[1025,1048,369,392]" italics="true" pageId="35" pageNumber="236">L.</emphasis>
|
||
sp. cf
|
||
<taxonomicName id="4C6A63999871FFA9FB9F894CFB78EEF2" authorityName="Laidlaw" authorityYear="1912" box="[1122,1253,367,392]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089871FFA9FB9F894CFB78EEF2" box="[1122,1253,367,392]" italics="true" pageId="35" pageNumber="236">williamsoni</emphasis>
|
||
</taxonomicName>
|
||
(see below), for which no ITS sequence was obtained, to provide conclusive evidence that the two are separate species, although this is likely.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3704B919871FFA9FF3A89FFFAF5ED1A" pageId="35" pageNumber="236" type="description">
|
||
<paragraph id="8BD5181A9871FFA9FF3A89FFFAF5ED1A" blockId="35.[151,1436,151,608]" pageId="35" pageNumber="236">
|
||
<taxonomicName id="4C6A63999871FFA9FF3A89FFFE6BEE8F" authorityName="Laidlaw" authorityYear="1912" box="[199,502,476,501]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="236" phylum="Arthropoda" rank="species" species="williamsoni">
|
||
<emphasis id="B91EC4089871FFA9FF3A89FFFE6BEE8F" box="[199,502,476,501]" italics="true" pageId="35" pageNumber="236">Leptogomphus williamsoni</emphasis>
|
||
</taxonomicName>
|
||
is known from
|
||
<collectingRegion id="49AED6F89871FFA9FD5B89FFFC94EE8F" box="[678,777,476,501]" country="Malaysia" name="Sarawak" pageId="35" pageNumber="236">
|
||
<collectingCountry id="F37D588A9871FFA9FD5B89FFFC94EE8F" box="[678,777,476,501]" name="Malaysia" pageId="35" pageNumber="236">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
,
|
||
<collectingRegion id="49AED6F89871FFA9FCE889FFFCC4EE8F" box="[789,857,476,501]" country="Malaysia" name="Sabah" pageId="35" pageNumber="236">Sabah</collectingRegion>
|
||
and east and south-east
|
||
<collectingRegion id="49AED6F89871FFA9FB9789FFFB70EE8F" box="[1130,1261,476,501]" country="Indonesia" name="Kalimantan Utara" pageId="35" pageNumber="236">Kalimantan</collectingRegion>
|
||
(
|
||
<figureCitation id="1351049F9871FFA9FB0189FFFAC3EE8E" box="[1276,1374,476,501]" captionStart="FIGURE 129" captionStartId="51.[151,250,1608,1630]" captionTargetBox="[151,1436,193,1587]" captionTargetId="figure@51.[151,1436,193,1587]" captionTargetPageId="51" captionText="FIGURE 129. Distribution of L. williamsoni (black square) and L.sp.cf williamsoni (red triangle)." httpUri="https://zenodo.org/record/1068060/files/figure.png" pageId="35" pageNumber="236">Fig. 129</figureCitation>
|
||
). It is likely to occur in parts of
|
||
<collectingCountry id="F37D588A9871FFA9FE4B89DCFD9FED62" box="[438,514,511,536]" name="Brunei" pageId="35" pageNumber="236">Brunei</collectingCountry>
|
||
and to be more widely distributed in
|
||
<collectingRegion id="49AED6F89871FFA9FC5E89DCFBBBED62" box="[931,1062,511,536]" country="Indonesia" name="Kalimantan Utara" pageId="35" pageNumber="236">Kalimantan</collectingRegion>
|
||
than we currently know. Records from
|
||
<collectingRegion id="49AED6F89871FFA9FF2A8A07FEAAED47" box="[215,311,548,573]" country="Malaysia" name="Sarawak" pageId="35" pageNumber="236">
|
||
<collectingCountry id="F37D588A9871FFA9FF2A8A07FEAAED47" box="[215,311,548,573]" name="Malaysia" pageId="35" pageNumber="236">Sarawak</collectingCountry>
|
||
</collectingRegion>
|
||
west of the Lupar River may belong to a separate species and are discussed as
|
||
<emphasis id="B91EC4089871FFA9FB1D8A06FB6AED46" box="[1248,1271,549,572]" italics="true" pageId="35" pageNumber="236">L.</emphasis>
|
||
species sp. cf
|
||
<emphasis id="B91EC4089871FFA9FF6A8A64FE8FED1A" box="[151,274,583,608]" italics="true" pageId="35" pageNumber="236">wiliamsoni</emphasis>
|
||
below. This species occurs at streams in mixed dipterocarp forest in hilly and mountainous terrain.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |