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<document ID-DOI="10.5281/zenodo.277240" ID-GBIF-Dataset="907ae2b6-4805-4348-96af-640f649d491d" ID-ISSN="1175-5326" ID-Zenodo-Dep="277240" checkinTime="1460277928943" checkinUser="plazi" docAuthor="Eichenberger, Beata &amp; Kranz-Baltensperger, Yvonne" docDate="2011" docId="03B0741CDF02F26DFF3CFDACFB56F8EF" docLanguage="en" docName="zt02823p031.pdf" docOrigin="Zootaxa 2823" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trilacuna clarissa Eichenberger, n. sp." docType="treatment" docVersion="10" lastPageNumber="10" masterDocId="FF890C64DF0BF264FFABFFC0FFBEFFA5" masterDocTitle="New Trilacuna species from Thailand, Malaysia and Sumatra (Araneae, Oonopidae)" masterLastPageNumber="31" masterPageNumber="1" pageNumber="10" updateTime="1678114192562" updateUser="tatiana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New Trilacuna species from Thailand, Malaysia and Sumatra (Araneae, Oonopidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Eichenberger, Beata</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kranz-Baltensperger, Yvonne</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="pubDate">
<mods:number>2011-04-18</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>2823</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>31</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.277240</mods:identifier>
<mods:identifier type="GBIF-Dataset">907ae2b6-4805-4348-96af-640f649d491d</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">277240</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4431123" ID-GBIF-Taxon="119396382" ID-Zenodo-Dep="4431123" LSID="urn:lsid:plazi:treatment:03B0741CDF02F26DFF3CFDACFB56F8EF" httpUri="http://treatment.plazi.org/id/03B0741CDF02F26DFF3CFDACFB56F8EF" lastPageNumber="10" pageId="9" pageNumber="10">
<subSubSection pageId="9" pageNumber="10" type="nomenclature">
<paragraph blockId="9.[151,620,619,678]" box="[151,620,619,646]" pageId="9" pageNumber="10">
<heading bold="true" box="[151,620,619,646]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis bold="true" box="[151,620,619,646]" pageId="9" pageNumber="10">
<taxonomicName ID-CoL="7D65T" authority="Eichenberger" authorityName="Eichenberger" box="[151,547,619,646]" class="Arachnida" family="Oonopidae" genus="Trilacuna" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="clarissa" status="n. sp.">
<emphasis bold="true" box="[151,370,620,646]" italics="true" pageId="9" pageNumber="10">Trilacuna clarissa</emphasis>
Eichenberger
</taxonomicName>
<taxonomicNameLabel box="[554,620,619,645]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="9.[151,620,619,678]" box="[151,274,654,678]" pageId="9" pageNumber="10">
(
<figureCitation box="[159,266,654,678]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStartId-0="10.[151,250,1920,1943]" captionStartId-1="11.[151,249,1889,1912]" captionStartId-2="12.[151,249,1900,1923]" captionTargetBox-0="[152,1436,195,1880]" captionTargetBox-1="[151,1436,185,1872]" captionTargetBox-2="[151,1436,193,1879]" captionTargetId-0="figure@10.[151,1436,195,1896]" captionTargetId-1="figure@11.[151,1436,185,1886]" captionTargetId-2="figure@12.[151,1436,193,1893]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURE 7: Trilacuna clarissa n. sp. Male: A. habitus dorsal view; B. habitus ventral view; C. habitus lateral view; D. sternum ventral view; E. carapace dorsal view; F. sternum ventral view, SEM; G. carapace dorsal view, SEM; H. sternum surface ventral view, SEM. Abbreviations: boc, booklung covers. Scale bars: A – G 0.5 mm; H 100 Μm." captionText-1="FIGURE 8: Trilacuna clarissa n. sp. Male: A. carapace anterior view; B. carapace anterior view, SEM; C. endites ventral view; D. endites ventral view, SEM; E. right endite, ventral view, distal inner margin; F. tibia I prolateral view, SEM; G. tibia I prolateral view, smooth stripe with pore; H. gonopore ventral view. Abbreviations: ehb, elevated hair base; ldi, labium deep incision; coh, comb-like hairs; fuh, funnel-shaped hairs. Scale bars: A 0.5 mm; B 300 Μm; C – D, H 200 Μm; E 20 Μm; F 100 Μm; G 10 Μm." captionText-2="FIGURE 9: Trilacuna clarissa n. sp. Male: A – B. left palp, prolateral view; C. right palp, prolateral view, SEM; D. right palp, dorsal view, SEM; E. left embolus, prolateral view; F. left embolus, prolateral view, SEM; G. right embolus, dorsal view, SEM; H. left embolus, dorsal view, SEM; I. left embolus, dorsal view, distal tip, SEM. Abbreviations: emb, embolus; bsh, brush of thickened hairs; pbb, prolateral broadish branch; vb, ventral branch. Scale bars: A – D 100 Μm; E – H 20 Μm; I 10 Μm." httpUri-0="https://zenodo.org/record/277247/files/figure.png" httpUri-1="https://zenodo.org/record/277248/files/figure.png" httpUri-2="https://zenodo.org/record/277249/files/figure.png" pageId="9" pageNumber="10">
Figs.
<number box="[223,266,654,678]" pageId="9" pageNumber="10" value="8.0" valueMax="9.0" valueMin="7.0">7–9</number>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="materials_examined">
<paragraph blockId="9.[151,1436,726,1867]" pageId="9" pageNumber="10">
<emphasis box="[151,311,726,751]" italics="true" pageId="9" pageNumber="10">Type material</emphasis>
:
<materialsCitation ID-GBIF-Occurrence="3014674309" collectedFrom="primary forest" collectingDate="2006-06-13" collectionCode="MHNG" collectorName="P. Schwendinger" county="Rimbo Panti Nature Reserve" elevation="350" latitude="0.34611112" location="West Sumatra Province: Rimbo Panti Nature Reserve, about 30 km N of Lubuksikaping" longLatPrecision="21" longitude="100.06917" municipality="West Sumatra Province" pageId="9" pageNumber="10" specimenCode="PBI_OON 00012418" specimenCount="1" specimenCount-male="3" stateProvince="Sumatra" typeStatus="holotype">
<typeStatus box="[331,434,726,750]" pageId="9" pageNumber="10">
<emphasis box="[331,434,726,750]" italics="true" pageId="9" pageNumber="10">Holotype</emphasis>
</typeStatus>
male (
<specimenCode box="[520,771,726,750]" pageId="9" pageNumber="10">PBI_OON 00012418</specimenCode>
):
<collectingRegion box="[794,892,726,750]" pageId="9" pageNumber="10">Sumatra</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:03B0741CDF02F26DFF3CFDACFB56F8EF:8EC693D1DF02F26DFC27FD17FDC9FCB7" county="Rimbo Panti Nature Reserve" latitude="0.34611112" longLatPrecision="21" longitude="100.06917" municipality="West Sumatra Province" name="West Sumatra Province: Rimbo Panti Nature Reserve, about 30 km N of Lubuksikaping" pageId="9" pageNumber="10" stateProvince="Sumatra">
West Sumatra Province: Rimbo Panti Nature Reserve, about
<quantity box="[324,395,762,786]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="km" value="30.0">30 km</quantity>
N of Lubuksikaping
</location>
,
<geoCoordinate box="[642,763,762,786]" degrees="0" direction="north" minutes="20" orientation="latitude" pageId="9" pageNumber="10" precision="15" seconds="46" value="0.34611112">
0°
<number box="[666,697,762,786]" pageId="9" pageNumber="10" value="20.0">20</number>
'
<number box="[699,730,762,786]" pageId="9" pageNumber="10" value="46.0">46</number>
&quot;N
</geoCoordinate>
,
<geoCoordinate box="[774,919,762,786]" degrees="100" direction="east" minutes="04" orientation="longitude" pageId="9" pageNumber="10" precision="15" seconds="09" value="100.06917">
<number box="[774,818,762,786]" pageId="9" pageNumber="10" value="100.0">100</number>
°04'09&quot;E
</geoCoordinate>
,
<quantity box="[930,1059,762,786]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="9" pageNumber="10" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">
<elevation box="[930,1059,762,786]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="9" pageNumber="10" unit="m" value="350.0" valueMax="400.0" valueMin="300.0">300–400 m</elevation>
</quantity>
,
<collectedFrom box="[1071,1234,762,786]" pageId="9" pageNumber="10">primary forest</collectedFrom>
,
<collectingDate box="[1245,1432,762,786]" pageId="9" pageNumber="10">
<date box="[1245,1365,762,786]" pageId="9" pageNumber="10" value="2011-06-13">
<number box="[1245,1271,762,786]" pageId="9" pageNumber="10" value="11.0">11</number>
/
<number box="[1280,1307,762,786]" pageId="9" pageNumber="10" value="13.0">13</number>
June
</date>
<number box="[1373,1432,762,786]" pageId="9" pageNumber="10" value="2006.0">2006</number>
</collectingDate>
, leg.
<collectorName box="[198,386,798,822]" pageId="9" pageNumber="10">P. Schwendinger</collectorName>
, deposited in
<collectionCode box="[538,624,798,822]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="10" type="Museum">MHNG</collectionCode>
.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3014674350" collectingDate="2006-06-13" collectionCode="MHNG" collectorName="P. Schwendinger" county="Rimbo Panti Nature Reserve" elevation="350" latitude="0.34611112" location="Sumatra" longLatPrecision="21" longitude="100.06917" municipality="West Sumatra Province" pageId="9" pageNumber="10" specimenCode="PBI_OON 00031971" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount box="[634,708,798,822]" pageId="9" pageNumber="10" type="male">
<number box="[634,648,798,822]" pageId="9" pageNumber="10" value="1.0">1</number>
male
</specimenCount>
<typeStatus box="[715,811,799,823]" pageId="9" pageNumber="10">paratype</typeStatus>
(
<specimenCode box="[825,1068,798,822]" pageId="9" pageNumber="10">PBI_OON 00031971</specimenCode>
), collected with holotype, deposited in
<collectionCode box="[228,314,834,858]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="10" type="Museum">MHNG</collectionCode>
.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3014674325" box="[325,1290,834,859]" collectingDate="2006-06-13" collectionCode="MHNG, NMBE" collectorName="P. Schwendinger" county="Rimbo Panti Nature Reserve" elevation="350" latitude="0.34611112" location="Sumatra" longLatPrecision="21" longitude="100.06917" municipality="West Sumatra Province" pageId="9" pageNumber="10" specimenCount="6" specimenCount-male="3" typeStatus="holotype">
<specimenCount box="[325,400,834,858]" pageId="9" pageNumber="10" type="male">
<number box="[325,339,834,858]" pageId="9" pageNumber="10" value="1.0">1</number>
male
</specimenCount>
<typeStatus box="[407,503,835,859]" pageId="9" pageNumber="10">paratype</typeStatus>
(PBI_OON 00031972), collected with
<typeStatus box="[945,1046,834,858]" pageId="9" pageNumber="10">holotype</typeStatus>
, deposited in
<collectionCode box="[1200,1286,834,858]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="9" pageNumber="10" type="Museum">MHNG</collectionCode>
.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3014674332" collectingDate="2006-06-13" collectionCode="NMBE" collectorName="P. Schwendinger" county="Rimbo Panti Nature Reserve" elevation="350" latitude="0.34611112" location="Sumatra" longLatPrecision="21" longitude="100.06917" municipality="West Sumatra Province" pageId="9" pageNumber="10" specimenCode="PBI_OON 00031973" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount box="[1296,1371,834,858]" pageId="9" pageNumber="10" type="male">
<number box="[1296,1310,834,858]" pageId="9" pageNumber="10" value="1.0">1</number>
male
</specimenCount>
<typeStatus pageId="9" pageNumber="10">paratype</typeStatus>
(
<specimenCode box="[213,457,870,894]" pageId="9" pageNumber="10">PBI_OON 00031973</specimenCode>
), collected with holotype, deposited in
<collectionCode box="[895,980,871,894]" collectionName="Switzerland, Bern, Naturhistorische Museums" country="Switzerland" httpUri="http://grbio.org/cool/5hwi-0wgz" name="Naturhistorisches Museum der Burgergemeinde Bern" pageId="9" pageNumber="10">NMBE</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="etymology">
<paragraph blockId="9.[151,1436,726,1867]" pageId="9" pageNumber="10">
<emphasis box="[199,319,907,931]" italics="true" pageId="9" pageNumber="10">Etymology</emphasis>
: The species epithet refers to the name of Agatha Christie, born as Agatha Mary
<taxonomicName box="[1261,1351,906,930]" class="Arachnida" family="Oonopidae" genus="Trilacuna" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="29" phylum="Arthropoda" rank="species" species="clarissa" status="n. sp.">Clarissa</taxonomicName>
Miller, who was a brilliant and famous English author of novels, short stories and plays.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="diagnosis">
<paragraph blockId="9.[151,1436,726,1867]" pageId="9" pageNumber="10">
<emphasis box="[199,313,980,1003]" italics="true" pageId="9" pageNumber="10">Diagnosis</emphasis>
: This species can be distinguished from all other
<taxonomicName box="[865,974,979,1003]" class="Arachnida" family="Oonopidae" genus="Trilacuna" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis box="[865,974,979,1003]" italics="true" pageId="9" pageNumber="10">Trilacuna</emphasis>
</taxonomicName>
species by the sternum surface with relatively small, ovoid pits (figs.
<number box="[479,496,1015,1038]" pageId="9" pageNumber="10" value="7.0">7</number>
. D, F, H) and abundant, evenly scattered, plumose setae (fig.
<number box="[1184,1201,1015,1038]" pageId="9" pageNumber="10" value="7.0">7</number>
. H).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="description">
<paragraph blockId="9.[151,1436,726,1867]" box="[199,953,1050,1075]" pageId="9" pageNumber="10">
<emphasis box="[199,331,1052,1075]" italics="true" pageId="9" pageNumber="10">Description</emphasis>
: Description based on
<specimenCount box="[589,675,1050,1074]" pageId="9" pageNumber="10" type="male">
<number box="[589,603,1050,1074]" pageId="9" pageNumber="10" value="4.0">4</number>
males
</specimenCount>
(plus
<specimenCount box="[745,831,1050,1074]" pageId="9" pageNumber="10" type="male">
<number box="[745,759,1050,1074]" pageId="9" pageNumber="10" value="2.0">2</number>
males
</specimenCount>
for SEM).
</paragraph>
<paragraph blockId="9.[151,1436,726,1867]" pageId="9" pageNumber="10">
<collectionCode box="[199,283,1087,1111]" pageId="9" pageNumber="10">MALE</collectionCode>
: Body length
<quantity box="[438,527,1086,1110]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="10" unit="mm" value="1.1">1.1 mm</quantity>
, proportion carapace length:abdominal length
<number box="[1050,1065,1086,1110]" pageId="9" pageNumber="10" value="1.0">1</number>
:
<number box="[1071,1086,1086,1110]" pageId="9" pageNumber="10" value="1.0">1</number>
. Uniformly orange-brown species (figs.
<number box="[267,284,1123,1146]" pageId="9" pageNumber="10" value="7.0">7</number>
. A–C).
</paragraph>
<paragraph blockId="9.[151,1436,726,1867]" pageId="9" pageNumber="10">
<emphasis box="[199,309,1159,1182]" italics="true" pageId="9" pageNumber="10">Carapace</emphasis>
broadly oval, strongly elevated, surface smooth, without spikes, lateral margin straight, with blunt denticles, with second, strongly rebordered, straight margin below (figs.
<number box="[968,985,1195,1218]" pageId="9" pageNumber="10" value="7.0">7</number>
. E, G;
<number box="[1063,1079,1194,1218]" pageId="9" pageNumber="10" value="8.0">8</number>
. B).
<emphasis box="[1134,1187,1196,1219]" italics="true" pageId="9" pageNumber="10">Eyes</emphasis>
posterior row straight from above, procurved from front (figs.
<number box="[602,619,1231,1254]" pageId="9" pageNumber="10" value="7.0">7</number>
. E;
<number box="[663,679,1230,1254]" pageId="9" pageNumber="10" value="8.0">8</number>
. B).
<emphasis box="[733,826,1231,1254]" italics="true" pageId="9" pageNumber="10">Sternum</emphasis>
with radial furrows of small pits between coxae, anterior margin with transverse groove.
<emphasis box="[558,689,1267,1291]" italics="true" pageId="9" pageNumber="10">Mouthparts</emphasis>
: Chelicerae promargin with row of flattened hairs (fsh) in combination with feather-like hairs (flh), (see fig.
<number box="[651,668,1302,1326]" pageId="9" pageNumber="10" value="2.0">2</number>
. E); labium triangular, not fused to sternum, anterior margin deeply incised (ldi), (figs.
<number box="[368,384,1338,1362]" pageId="9" pageNumber="10" value="8.0">8</number>
. C–D); endites slender, distally only slightly branched, with distal patch of fork-like serrated setae (coh), outer part with some broadish, funnel shaped, distally serrated setae (fuh), (fig.
<number box="[1178,1194,1374,1398]" pageId="9" pageNumber="10" value="8.0">8</number>
. E).
</paragraph>
<paragraph blockId="9.[151,1436,726,1867]" pageId="9" pageNumber="10">
<emphasis box="[199,305,1411,1435]" italics="true" pageId="9" pageNumber="10">Abdomen</emphasis>
: Broadly-ovoid, dorsal scutum not fused to epigastric scutum, postepigastric scutum short, almost circular, with short, posteriorly directed lateral apodemes, posterior spiracles not connected by groove or dark belt (fig.
<number box="[205,221,1482,1506]" pageId="9" pageNumber="10" value="8.0">8</number>
. H), booklung covers roundish.
</paragraph>
<paragraph blockId="9.[151,1436,726,1867]" pageId="9" pageNumber="10">
<emphasis box="[199,252,1520,1543]" italics="true" pageId="9" pageNumber="10">Legs</emphasis>
: Ventral apex of tibiae III and IV with one or few specialized hairs with fine, plumose, curved hair tip (see fig.
<number box="[196,227,1554,1578]" pageId="9" pageNumber="10" value="16.0">16</number>
. H); metatarsi III, IV and tarsus IV with harpoon-like hairs (see fig.
<number box="[986,1003,1554,1578]" pageId="9" pageNumber="10" value="2.0">2</number>
. G); tibiae I and II ventral surface with smooth stripes containing pores (figs.
<number box="[578,594,1590,1614]" pageId="9" pageNumber="10" value="8.0">8</number>
. F–G), less distinct stripes also observed on femora I–IV and metatarsi III– IV, pores not observed but absence unproved. Femora without spines.
</paragraph>
<paragraph blockId="9.[151,1436,726,1867]" pageId="9" pageNumber="10">
<emphasis box="[199,362,1662,1686]" italics="true" pageId="9" pageNumber="10">Male genitalia</emphasis>
:
<emphasis box="[376,490,1662,1686]" italics="true" pageId="9" pageNumber="10">Male palp</emphasis>
(figs.
<number box="[561,578,1662,1686]" pageId="9" pageNumber="10" value="9.0">9</number>
. A–I) embolus (emb) light, short and straight; retrolateral-dorsally framed by a brush of thickened hairs (bsh) (figs.
<number box="[574,591,1698,1722]" pageId="9" pageNumber="10" value="9.0">9</number>
. E–I); conductor-complex about as long as embolus, divided in a prolateral, broadish branch (pbb) and in a ventral branch (vb), both surrounded by numerous setae (figs.
<number box="[1217,1234,1734,1758]" pageId="9" pageNumber="10" value="9.0">9</number>
. E–I). Bulb about twice as long as cymbium, stout, tapering apically (figs.
<number box="[784,801,1770,1794]" pageId="9" pageNumber="10" value="9.0">9</number>
. A–C). Femur elongated, slender (figs.
<number box="[1244,1261,1770,1794]" pageId="9" pageNumber="10" value="9.0">9</number>
. A–B).
</paragraph>
<paragraph blockId="9.[151,1436,726,1867]" box="[199,438,1806,1831]" pageId="9" pageNumber="10">FEMALE: unknown.</paragraph>
</subSubSection>
<subSubSection box="[199,1256,1842,1867]" pageId="9" pageNumber="10" type="distribution">
<paragraph blockId="9.[151,1436,726,1867]" box="[199,1256,1842,1867]" pageId="9" pageNumber="10">
<emphasis box="[199,336,1843,1867]" italics="true" pageId="9" pageNumber="10">Distribution</emphasis>
: Known only from the
<typeStatus box="[600,648,1843,1867]" pageId="9" pageNumber="10">type</typeStatus>
locality, the Rimbo Panti Nature Reserve on Sumatra.
</paragraph>
</subSubSection>
</treatment>
</document>
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