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<document id="F87A7CB625CED4716DC9312827EB139D" ID="10.11646/zootaxa.4151.1.1" ID-DOI="10.11646/zootaxa.4151.1.1" ID-GBIF-Dataset="8cdcdd3c-dfec-48a4-803b-c0c55290e00b" ID-ISSN="1175-5326" ID-Zenodo-Dep="262273" ID-ZooBank="606E8BD8-5CA2-4643-A0C9-3882302A7019" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1471415195561" checkinUser="plazi" docAuthor="Richard, Miguel, Graber, Werner &amp; Kropf, Christian" docDate="2016" docId="03AF8D64FFE1FFE8FF4EF957FB88EE83" docLanguage="en" docName="zootaxa.4151.1.1.pdf" docOrigin="Zootaxa 4151 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ischnothyreus ligulatus Richard, sp. nov." docType="treatment" docVersion="7" lastPageNumber="26" masterDocId="FF96F51CFFF1FFF1FFD9FF91FFAFEC6C" masterDocTitle="The goblin spider genus Ischnothyreus (Araneae, Oonopidae) in Java and Sumatra" masterLastPageNumber="99" masterPageNumber="1" pageNumber="17" updateTime="1698381915265" updateUser="plazi">
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<mods:title id="F283A330C34FCB23ABA3FE1086E76B33">The goblin spider genus Ischnothyreus (Araneae, Oonopidae) in Java and Sumatra</mods:title>
</mods:titleInfo>
<mods:name id="AE6F64EA1FEAA8233B971C1975A3B100" type="personal">
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<mods:namePart id="B0C8F73B7369DCDDBBFFE760FBF93200">Richard, Miguel</mods:namePart>
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<mods:namePart id="E25459B9C58F823BD354474F345A8C9C">Graber, Werner</mods:namePart>
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<mods:namePart id="6AB456D8B6D591B47FF852A77DA656D1">Kropf, Christian</mods:namePart>
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<mods:title id="CFC03F19414832F74BE33D074489BEB7">Zootaxa</mods:title>
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<mods:date id="E60D12AA7A3CB5256D725E4503B982F5">2016</mods:date>
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<mods:number id="7B95DAC32F75ECAFE44F717656EF2C9D">4151</mods:number>
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<mods:classification id="2B21953E352A405B3FACD82E38CBF313">journal article</mods:classification>
<mods:identifier id="A99773D2B5FB081C0C687C8155B67D38" type="DOI">10.11646/zootaxa.4151.1.1</mods:identifier>
<mods:identifier id="7440BFF8C40CB08C20542400142BFCF0" type="GBIF-Dataset">8cdcdd3c-dfec-48a4-803b-c0c55290e00b</mods:identifier>
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<treatment id="03AF8D64FFE1FFE8FF4EF957FB88EE83" ID-DOI="http://doi.org/10.5281/zenodo.6079881" ID-GBIF-Taxon="124329598" ID-Zenodo-Dep="6079881" LSID="urn:lsid:plazi:treatment:03AF8D64FFE1FFE8FF4EF957FB88EE83" httpUri="http://treatment.plazi.org/id/03AF8D64FFE1FFE8FF4EF957FB88EE83" lastPageId="25" lastPageNumber="26" pageId="16" pageNumber="17">
<subSubSection id="C31C6FF9FFE1FFE1FF4EF957FE8FEB6E" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BB93C72FFE1FFE1FF4EF957FD20EA8C" blockId="16.[151,655,1734,1794]" box="[151,655,1734,1760]" pageId="16" pageNumber="17">
<heading id="D0F18B1EFFE1FFE1FF4EF957FD20EA8C" bold="true" box="[151,655,1734,1760]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B972E060FFE1FFE1FF4EF957FD20EA8C" bold="true" box="[151,655,1734,1760]" pageId="16" pageNumber="17">
<taxonomicName id="4C0647F1FFE1FFE1FF4EF957FD8BEA8C" ID-CoL="3Q665" authority="Richard" authorityName="Richard" box="[151,548,1734,1760]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ligulatus" status="sp. nov.">
<emphasis id="B972E060FFE1FFE1FF4EF957FE1AEA8C" bold="true" box="[151,437,1734,1760]" italics="true" pageId="16" pageNumber="17">Ischnothyreus ligulatus</emphasis>
Richard
</taxonomicName>
,
<taxonomicNameLabel id="A2415D1BFFE1FFE1FDE8F957FD20EA8C" box="[561,655,1734,1760]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB93C72FFE1FFE1FF4EF978FE8FEB6E" blockId="16.[151,655,1734,1794]" box="[151,288,1769,1794]" pageId="16" pageNumber="17">
(
<figureCitation id="133D20F7FFE1FFE1FF46F978FEB8EB6E" box="[159,279,1769,1794]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStart-4="FIGURE 13" captionStart-5="FIGURE 14" captionStart-6="FIGURE 15" captionStart-7="FIGURE 16" captionStartId-0="17.[151,250,1913,1935]" captionStartId-1="18.[151,250,1882,1904]" captionStartId-2="19.[151,250,1550,1572]" captionStartId-3="20.[151,250,1640,1662]" captionStartId-4="21.[151,250,1716,1738]" captionStartId-5="22.[151,250,1878,1900]" captionStartId-6="23.[151,250,1377,1399]" captionStartId-7="24.[151,250,1677,1699]" captionTargetBox-0="[151,1436,373,1884]" captionTargetBox-1="[180,1395,197,1856]" captionTargetBox-2="[307,1275,198,1523]" captionTargetBox-3="[151,1436,183,1618]" captionTargetBox-4="[151,1436,193,1698]" captionTargetBox-5="[151,1436,193,1856]" captionTargetBox-6="[182,1406,189,1354]" captionTargetBox-7="[174,1416,189,1655]" captionTargetId-0="figure@17.[151,1436,373,1892]" captionTargetId-1="figure@18.[180,1406,193,1861]" captionTargetId-2="figure@19.[297,1290,193,1529]" captionTargetId-3="figure@20.[151,1436,183,1621]" captionTargetId-4="figure@21.[151,1436,193,1702]" captionTargetId-5="figure@22.[151,1436,193,1857]" captionTargetId-6="figure@23.[182,1406,189,1355]" captionTargetId-7="figure@24.[174,1418,189,1659]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionTargetPageId-4="21" captionTargetPageId-5="22" captionTargetPageId-6="23" captionTargetPageId-7="24" captionText-0="FIGURE 9. Ischnothyreus ligulatus sp. nov. Male: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, carapace, frontal view; E, sternum, ventral view; F, opisthosoma, dorsal view; G, opisthosoma, ventral view. Scale bar: A = 1 mm." captionText-1="FIGURE 10. Ischnothyreus ligulatus sp. nov. Male: A, left palp, prolateral view; B, H, left palp, retrolate ¬ ral view; C, D, left embolus, prolateral view; E, F, left embolus, prolateral view; G, left palp, prolateral view (mirrored image). rl retrolateral lobe; ssa, spoon-shaped appendage. Scale bars: A, G = 0.2 mm." captionText-2="FIGURE 11. Ischnothyreus ligulatus sp. nov. Male: A, C, left chelicera, anterior view; B, right chelicera, anterior view; D, left chelicera, posterior view. p, process. Scale bars: A, C = 0.1 mm." captionText-3="FIGURE 12. Ischnothyreus ligulatus sp. nov. Male, SEM images: A, carapace, dorsal view; B, mouthparts, ventral view; C, palp, retrolateral view; D, palp, prolateral view; E, embolus, retrolateral view; F, embolus, prolateral view. eo, embolic opening; rl, retrolateral lobe; ssa, spoon-shaped appendage. Scale bars: A = 0.5 mm, B = 0.2 mm, C, D = 0.1 mm, E = 20 µm, F = 10 µm." captionText-4="FIGURE 13. Ischnothyreus ligulatus sp. nov. Male: A, left endite, ventral view; B, chelicerae, ventral view; C, base of left cheliceral fang, anteroventral view; D, base of left cheliceral fang, ventral view; E, sperm pore, ventral view; F, claws of leg III; G, tarsal organ of leg III; H, surface of carapace cuticle, median part of pars cephalica. p, process. Scale bars: A, B, D = 50 µm; C, E, H = 20 µm; F = 10 µm; G = 2 µm." captionText-5="FIGURE 14. Ischnothyreus ligulatus sp. nov. Female: A, habitus, dorsal view; B, habitus, ventral view; C carapace, frontal view; D, habitus, lateral view; E, opisthosoma, dorsal view; F, carapace, dorsal view; G, opisthosoma, lateral view; H, opisthosoma, ventral view. Scale bar: A = 1 mm." captionText-6="FIGURE 15. Ischnothyreus ligulatus sp. nov.: A, B, C, E, female genital area, ventral view; D, dorsal view. a, apodeme; ls, lateral sclerite; nsp, nail-shaped projection; o, opening of apodeme; s, stigma; wt, winding tube. Scale bars: A, E = 0.2 mm; C = 0.1 mm." captionText-7="FIGURE 16. Ischnothyreus ligulatus sp. nov. Female: A, eyes, dorsal view; B, right endite with serrula, ventral view; C, sternum, ventral view; D, mouthparts, ventral view; E, postepigastric scutum, ventral view; F, row of pores, ventral view; G, nail-shaped process, ventral view. nsp, nail-shaped projection; o, opening of apodeme; po, pores. Scale bars: A, C = 0.2 mm; B = 50 µm; D, E = 0.1 mm; F, G = 20 µm." httpUri-0="https://zenodo.org/record/262282/files/figure.png" httpUri-1="https://zenodo.org/record/262283/files/figure.png" httpUri-2="https://zenodo.org/record/262284/files/figure.png" httpUri-3="https://zenodo.org/record/262285/files/figure.png" httpUri-4="https://zenodo.org/record/262286/files/figure.png" httpUri-5="https://zenodo.org/record/262287/files/figure.png" httpUri-6="https://zenodo.org/record/262288/files/figure.png" httpUri-7="https://zenodo.org/record/262289/files/figure.png" pageId="16" pageNumber="17">Figs. 916</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31C6FF9FFE1FFE0FF4EF8A0FAC3ED2C" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BB93C72FFE1FFE1FF4EF8A0FB63EBFE" blockId="16.[151,1437,1841,2010]" pageId="16" pageNumber="17">
<materialsCitation id="3B6E362FFFE1FFE1FF4EF8A0FA36EB01" ID-GBIF-Occurrence="1291639668" collectingDate="1986-11-14" collectingDateMax="1986-11-25" collectingDateMin="1986-11-14" collectionCode="NNM" collectorName="Leg. Suh. Djojosudharmo &amp; Christa Deeleman &amp; Leiden, NL" latitude="-6.746667" location="Ujung Kulon National Park" longLatPrecision="21" longitude="105.33361" pageId="16" pageNumber="17" specimenCount="2" stateProvince="JAVA" typeStatus="holotype">
<emphasis id="B972E060FFE1FFE1FF4EF8A0FEFFEB26" bold="true" box="[151,336,1841,1866]" pageId="16" pageNumber="17">
<typeStatus id="54BD82D0FFE1FFE1FF4EF8A0FF7DEB26" box="[151,210,1841,1866]" pageId="16" pageNumber="17">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54BD82D0FFE1FFE1FEBBF8A0FE60EB26" box="[354,463,1841,1866]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
: male (PBI_OON00052392),
<collectingRegion id="49C2F290FFE1FFE1FC99F8A3FC2AEB26" box="[832,901,1842,1866]" country="Indonesia" name="Jawa Timur" pageId="16" pageNumber="17">JAVA</collectingRegion>
:
<location id="8ED96AA9FFE1FFE1FC42F8A0FB5DEB26" LSID="urn:lsid:plazi:treatment:03AF8D64FFE1FFE8FF4EF957FB88EE83:8ED96AA9FFE1FFE1FC42F8A0FB5DEB26" box="[923,1266,1841,1866]" latitude="-6.746667" longLatPrecision="21" longitude="105.33361" name="Ujung Kulon National Park" pageId="16" pageNumber="17" stateProvince="JAVA">Ujung Kulon National Park</location>
(
<geoCoordinate id="EE325AB5FFE1FFE1FAD4F8A0FA39EB26" box="[1293,1430,1841,1866]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="15" value="-6.746667">6° 44 48″ S</geoCoordinate>
,
<geoCoordinate id="EE325AB5FFE1FFE1FF4EF8C4FE80EB02" box="[151,303,1877,1902]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="15" value="105.33361">105° 20 1″ E</geoCoordinate>
)
<date id="FFB81AB2FFE1FFE1FEE7F8C4FDE6EB02" box="[318,585,1877,1902]" pageId="16" pageNumber="17" value="1986-11-14" valueMax="1986-11-25" valueMin="1986-11-14">
<collectingDate id="EFFCE35AFFE1FFE1FEE7F8C4FDE6EB02" box="[318,585,1877,1902]" pageId="16" pageNumber="17" value="1986-11-14" valueMax="1986-11-25" valueMin="1986-11-14">November 1425, 1986</collectingDate>
</date>
,
<collectorName id="26F359A4FFE1FFE1FD8AF8C7FCDCEB02" box="[595,883,1877,1902]" pageId="16" pageNumber="17">Leg. Suh. Djojosudharmo</collectorName>
<emphasis id="B972E060FFE1FFE1FCA3F8C7FC1DEB02" box="[890,946,1877,1902]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
, coll.
<collectorName id="26F359A4FFE1FFE1FC2FF8C4FB6DEB02" box="[1014,1218,1877,1902]" pageId="16" pageNumber="17">Christa Deeleman</collectorName>
,
<collectionCode id="ED17A4B7FFE1FFE1FB14F8C7FAA1EB01" box="[1229,1294,1878,1901]" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="16" pageNumber="17">NNM</collectionCode>
<collectorName id="26F359A4FFE1FFE1FACCF8C4FA36EB01" box="[1301,1433,1877,1902]" pageId="16" pageNumber="17">Leiden, NL</collectorName>
</materialsCitation>
. Paratypes: 6 males and 8 females (PBI_OON00031890), collected with holotype, NNM Leiden.
</paragraph>
<paragraph id="8BB93C72FFE1FFE0FF1EF80CFAC3ED2C" blockId="16.[151,1437,1841,2010]" lastBlockId="17.[151,1436,151,320]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="B972E060FFE1FFE1FF1EF80CFD92EBDA" bold="true" box="[199,573,1949,1974]" pageId="16" pageNumber="17">Additional material examined.</emphasis>
5 males and 6 females, collected with holotype, NNM Leiden.
<materialsCitation id="3B6E362FFFE1FFE0FAEFF80FFE6EECDC" ID-GBIF-Occurrence="1291639657" collectingDate="1989-11-05" collectionCode="MHNG" collectorName="Agosti &amp; Lobl &amp; Burckhardt" elevation="2600" lastPageId="17" lastPageNumber="18" location="Mt Gede" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1" stateProvince="JAVA">
<specimenCount id="9D00F7FBFFE1FFE1FAEFF80FFA33EBDA" box="[1334,1436,1949,1974]" pageId="16" pageNumber="17" type="female">1 female</specimenCount>
(PBI_OON00012053),
<collectingRegion id="49C2F290FFE1FFE1FE7AF853FE47EBB6" box="[419,488,1986,2010]" country="Indonesia" name="Jawa Timur" pageId="16" pageNumber="17">JAVA</collectingRegion>
: W
<collectingRegion id="49C2F290FFE1FFE1FDC7F853FDFCEBB5" box="[542,595,1986,2009]" country="Indonesia" name="Jawa Timur" pageId="16" pageNumber="17">Java</collectingRegion>
,
<location id="8ED96AA9FFE1FFE1FDBDF853FD60EBB6" LSID="urn:lsid:plazi:treatment:03AF8D64FFE1FFE8FF4EF957FB88EE83:8ED96AA9FFE1FFE1FDBDF853FD60EBB6" box="[612,719,1985,2010]" name="Mt Gede" pageId="16" pageNumber="17" stateProvince="JAVA">Mt Gede</location>
,
<locationDeviation id="9A0B2550FFE1FFE1FD39F850FC0EEBB6" box="[736,929,1985,2010]" pageId="16" pageNumber="17">
<quantity id="4CFE9197FFE1FFE1FD39F850FC8FEBB6" box="[736,800,1985,2010]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="17" unit="km" value="50.0">50km</quantity>
SE Bogor
</locationDeviation>
,
<quantity id="4CFE9197FFE1FFE1FC68F850FBADEBB6" box="[945,1026,1985,2010]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="16" pageNumber="17" unit="m" value="2600.0">
<elevation id="002BDB41FFE1FFE1FC68F850FBADEBB6" box="[945,1026,1985,2010]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="16" pageNumber="17" unit="m" value="2600.0">2600m</elevation>
</quantity>
,
<date id="FFB81AB2FFE1FFE1FBCBF850FB5DEBB6" box="[1042,1266,1985,2010]" pageId="16" pageNumber="17" value="1989-11-05">
<collectingDate id="EFFCE35AFFE1FFE1FBCBF850FB5DEBB6" box="[1042,1266,1985,2010]" pageId="16" pageNumber="17" value="1989-11-05">November 5, 1989</collectingDate>
</date>
,
<collectorName id="26F359A4FFE1FFE1FADBF850FAE0EBB6" box="[1282,1359,1985,2010]" pageId="16" pageNumber="17">Agosti</collectorName>
,
<collectorName id="26F359A4FFE1FFE1FAB9F850FA37EBB6" box="[1376,1432,1985,2010]" pageId="16" pageNumber="17">Löbl</collectorName>
,
<collectorName id="26F359A4FFE0FFE0FF4EFF06FEBAECDC" box="[151,277,151,176]" pageId="17" pageNumber="18">Burckhardt</collectorName>
# 5a,
<collectionCode id="ED17A4B7FFE0FFE0FEB2FF09FE6EECDC" box="[363,449,152,176]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="17" pageNumber="18">MHNG</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B6E362FFFE0FFE0FE09FF09FB92ECC3" ID-GBIF-Occurrence="1291639669" box="[464,1085,151,176]" location="JAVA" pageId="17" pageNumber="18" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="JAVA">
<specimenCount id="9D00F7FBFFE0FFE0FE09FF09FD80ECDC" box="[464,559,151,176]" pageId="17" pageNumber="18" type="male">2 males</specimenCount>
,
<specimenCount id="9D00F7FBFFE0FFE0FD99FF06FD1DECDC" box="[576,690,151,176]" pageId="17" pageNumber="18" type="female">6 females</specimenCount>
(PBI_OON00031897),
<collectingRegion id="49C2F290FFE0FFE0FC15FF09FBBEECDC" box="[972,1041,152,176]" country="Indonesia" name="Jawa Timur" pageId="17" pageNumber="18">JAVA</collectingRegion>
: W
</materialsCitation>
.
<materialsCitation id="3B6E362FFFE0FFE0FB95FF09FBDCECB9" ID-GBIF-Occurrence="1291639663" collectingDate="1986-12" collectionCode="NNM" collectorName="Suh. Djojosudharmo &amp; Coll. Deeleman &amp; Leiden" elevation="1550" location="Cibodas" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Java">
<collectingRegion id="49C2F290FFE0FFE0FB95FF09FB2FECC3" box="[1100,1152,152,175]" country="Indonesia" name="Jawa Timur" pageId="17" pageNumber="18">Java</collectingRegion>
,
<location id="8ED96AA9FFE0FFE0FB4BFF06FB5EECDC" LSID="urn:lsid:plazi:treatment:03AF8D64FFE1FFE8FF4EF957FB88EE83:8ED96AA9FFE0FFE0FB4BFF06FB5EECDC" box="[1170,1265,151,176]" name="Cibodas" pageId="17" pageNumber="18" stateProvince="Java">Cibodas</location>
,
<quantity id="4CFE9197FFE0FFE0FAD8FF06FA37ECDC" box="[1281,1432,151,176]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="17" pageNumber="18" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">
<elevation id="002BDB41FFE0FFE0FAD8FF06FA37ECDC" box="[1281,1432,151,176]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="17" pageNumber="18" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">15001600m</elevation>
</quantity>
,
<date id="FFB81AB2FFE0FFE0FF4EFF2DFECFECB9" box="[151,352,188,213]" pageId="17" pageNumber="18" value="1986-12">
<collectingDate id="EFFCE35AFFE0FFE0FF4EFF2DFECFECB9" box="[151,352,188,213]" pageId="17" pageNumber="18" value="1986-12">December 1986</collectingDate>
</date>
, leg.
<collectorName id="26F359A4FFE0FFE0FE1BFF2DFD6EECB9" box="[450,705,188,213]" pageId="17" pageNumber="18">Suh. Djojosudharmo</collectorName>
,
<collectorName id="26F359A4FFE0FFE0FD06FF2DFC09ECB9" box="[735,934,188,213]" pageId="17" pageNumber="18">Coll. Deeleman</collectorName>
,
<collectionCode id="ED17A4B7FFE0FFE0FC1CFF2CFBA9ECB8" box="[965,1030,189,212]" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">NNM</collectionCode>
<collectorName id="26F359A4FFE0FFE0FBF8FF2DFBDCECB9" box="[1057,1139,188,213]" pageId="17" pageNumber="18">Leiden</collectorName>
</materialsCitation>
.
<materialsCitation id="3B6E362FFFE0FFE0FB48FF2DFF55ED71" ID-GBIF-Occurrence="1291639666" collectingDate="1986-11-14" collectingDateMax="1986-11-25" collectingDateMin="1986-11-14" collectorName="Suh" latitude="-6.746667" location="Ujung Kulon National Park" longLatPrecision="21" longitude="105.33361" pageId="17" pageNumber="18" specimenCount="8" specimenCount-female="2" specimenCount-male="6" stateProvince="JAVA">
<specimenCount id="9D00F7FBFFE0FFE0FB48FF2DFB52ECB9" box="[1169,1277,188,213]" pageId="17" pageNumber="18" type="male">6 males</specimenCount>
,
<specimenCount id="9D00F7FBFFE0FFE0FAC5FF2DFA33ECB9" box="[1308,1436,188,213]" pageId="17" pageNumber="18" type="female">2 females</specimenCount>
(PBI_OON00031886),
<collectingRegion id="49C2F290FFE0FFE0FE79FF71FE4AEC94" box="[416,485,224,248]" country="Indonesia" name="Jawa Timur" pageId="17" pageNumber="18">JAVA</collectingRegion>
:
<location id="8ED96AA9FFE0FFE0FE28FF4EFC82EC94" LSID="urn:lsid:plazi:treatment:03AF8D64FFE1FFE8FF4EF957FB88EE83:8ED96AA9FFE0FFE0FE28FF4EFC82EC94" box="[497,813,223,248]" latitude="-6.746667" longLatPrecision="21" longitude="105.33361" name="Ujung Kulon National Park" pageId="17" pageNumber="18" stateProvince="JAVA">Ujung Kulon National Park</location>
(
<geoCoordinate id="EE325AB5FFE0FFE0FCE6FF4EFC67EC94" box="[831,968,223,248]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="15" value="-6.746667">6° 44 48″ S</geoCoordinate>
,
<geoCoordinate id="EE325AB5FFE0FFE0FC0EFF4EFBDEEC94" box="[983,1137,223,248]" direction="east" orientation="longitude" pageId="17" pageNumber="18" precision="15" value="105.33361">105° 20 1″ E</geoCoordinate>
),
<date id="FFB81AB2FFE0FFE0FB51FF4EFA37EC94" box="[1160,1432,223,248]" pageId="17" pageNumber="18" value="1986-11-14" valueMax="1986-11-25" valueMin="1986-11-14">
<collectingDate id="EFFCE35AFFE0FFE0FB51FF4EFA37EC94" box="[1160,1432,223,248]" pageId="17" pageNumber="18" value="1986-11-14" valueMax="1986-11-25" valueMin="1986-11-14">November 1425, 1986</collectingDate>
</date>
, leg.
<collectorName id="26F359A4FFE0FFE0FF12FE95FF55ED71" box="[203,250,260,285]" pageId="17" pageNumber="18">Suh</collectorName>
</materialsCitation>
.
<materialsCitation id="3B6E362FFFE0FFE0FED1FE95FC84ED71" ID-GBIF-Occurrence="1291639652" box="[264,811,260,285]" collectionCode="NNM" location="Djojosudharmo" pageId="17" pageNumber="18" specimenCount="1" stateProvince="JAVA">
<location id="8ED96AA9FFE0FFE0FED1FE95FE16ED71" LSID="urn:lsid:plazi:treatment:03AF8D64FFE1FFE8FF4EF957FB88EE83:8ED96AA9FFE0FFE0FED1FE95FE16ED71" box="[264,441,260,285]" name="Djojosudharmo" pageId="17" pageNumber="18" stateProvince="JAVA">Djojosudharmo</location>
, Coll. Deeleman,
<collectionCode id="ED17A4B7FFE0FFE0FD54FE94FD61ED70" box="[653,718,261,284]" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">NNM</collectionCode>
Leiden
</materialsCitation>
.
<materialsCitation id="3B6E362FFFE0FFE0FCE0FE95FD57ED2C" ID-GBIF-Occurrence="1291639662" collectingDate="1986-11-21" collectorName="Suh" location="Ujung Kulon National Park" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="JAVA">
<specimenCount id="9D00F7FBFFE0FFE0FCE0FE95FC32ED71" box="[825,925,260,285]" pageId="17" pageNumber="18" type="female">1 female</specimenCount>
(PBI_OON00031899),
<collectingRegion id="49C2F290FFE0FFE0FB6AFE95FB57ED70" box="[1203,1272,260,284]" country="Indonesia" name="Jawa Timur" pageId="17" pageNumber="18">JAVA</collectingRegion>
:
<location id="8ED96AA9FFE0FFE0FADCFE95FE9BED2C" LSID="urn:lsid:plazi:treatment:03AF8D64FFE1FFE8FF4EF957FB88EE83:8ED96AA9FFE0FFE0FADCFE95FE9BED2C" name="Ujung Kulon National Park" pageId="17" pageNumber="18" stateProvince="JAVA">Ujung Kulon National Park</location>
, leaflitter,
<date id="FFB81AB2FFE0FFE0FE72FEB6FD22ED2C" box="[427,653,295,320]" pageId="17" pageNumber="18" value="1986-11-21">
<collectingDate id="EFFCE35AFFE0FFE0FE72FEB6FD22ED2C" box="[427,653,295,320]" pageId="17" pageNumber="18" value="1986-11-21">November 21, 1986</collectingDate>
</date>
, leg.
<collectorName id="26F359A4FFE0FFE0FD10FEB6FD57ED2C" box="[713,760,295,320]" pageId="17" pageNumber="18">Suh</collectorName>
</materialsCitation>
.
<materialsCitation id="3B6E362FFFE0FFE0FCDBFEB6FAC3ED2C" ID-GBIF-Occurrence="1291639650" box="[770,1388,295,320]" collectionCode="NNM" collectorName="Christa Deeleman" location="Djojosudharmo" pageId="17" pageNumber="18" specimenCount="1" stateProvince="JAVA">
<location id="8ED96AA9FFE0FFE0FCDBFEB6FC1CED2C" LSID="urn:lsid:plazi:treatment:03AF8D64FFE1FFE8FF4EF957FB88EE83:8ED96AA9FFE0FFE0FCDBFEB6FC1CED2C" box="[770,947,295,320]" name="Djojosudharmo" pageId="17" pageNumber="18" stateProvince="JAVA">Djojosudharmo</location>
, coll.
<collectorName id="26F359A4FFE0FFE0FC2EFEB6FB6CED2C" box="[1015,1219,295,320]" pageId="17" pageNumber="18">Christa Deeleman</collectorName>
,
<collectionCode id="ED17A4B7FFE0FFE0FB17FEB9FAA0ED53" box="[1230,1295,296,319]" country="Netherlands" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="18">NNM</collectionCode>
Leiden.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF796CFAFFE0FFE0FF4EF8E8FF76EBA0" httpUri="https://zenodo.org/record/262282/files/figure.png" pageId="17" pageNumber="18" targetBox="[151,1436,373,1884]" targetPageId="17">
<paragraph id="8BB93C72FFE0FFE0FF4EF8E8FF76EBA0" blockId="17.[151,1436,1913,1996]" pageId="17" pageNumber="18">
<emphasis id="B972E060FFE0FFE0FF4EF8E8FEBCEBE3" bold="true" box="[151,275,1913,1935]" pageId="17" pageNumber="18">FIGURE 9.</emphasis>
<taxonomicName id="4C0647F1FFE0FFE0FEC2F8E8FDA6EBE3" box="[283,521,1913,1935]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ligulatus" status="sp. nov.">
<emphasis id="B972E060FFE0FFE0FEC2F8E8FDA6EBE3" box="[283,521,1913,1935]" italics="true" pageId="17" pageNumber="18">Ischnothyreus ligulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415D1BFFE0FFE0FDC9F8EBFDCEEBE3" box="[528,609,1914,1935]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
Male: A, habitus, dorsal view; B, habitus, lateral view; C, habitus, ventral view; D, carapace, frontal view; E, sternum, ventral view; F, opisthosoma, dorsal view; G, opisthosoma, ventral view. Scale bar: A = 1 mm.
</paragraph>
</caption>
<caption id="DF796CFAFFE3FFE3FF4EF8CBFD4DEBC1" httpUri="https://zenodo.org/record/262283/files/figure.png" pageId="18" pageNumber="19" targetBox="[180,1395,197,1856]" targetPageId="18">
<paragraph id="8BB93C72FFE3FFE3FF4EF8CBFD4DEBC1" blockId="18.[151,1436,1882,1965]" pageId="18" pageNumber="19">
<emphasis id="B972E060FFE3FFE3FF4EF8CBFE8FEB1C" bold="true" box="[151,288,1882,1904]" pageId="18" pageNumber="19">FIGURE 10.</emphasis>
<taxonomicName id="4C0647F1FFE3FFE3FEFEF8CBFDBCEB1C" box="[295,531,1882,1904]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ligulatus" status="sp. nov.">Ischnothyreus ligulatus</taxonomicName>
<taxonomicNameLabel id="A2415D1BFFE3FFE3FDC3F8CAFDC6EB1C" box="[538,617,1883,1904]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
Male: A, left palp, prolateral view; B, H, left palp, retrolate¬ral view; C, D, left embolus, prolateral view; E, F, left embolus, prolateral view; G, left palp, prolateral view (mirrored image). rl retrolateral lobe; ssa, spoon-shaped appendage. Scale bars: A, G = 0.2 mm.
</paragraph>
</caption>
<caption id="DF796CFAFFE2FFE2FF4EF99FFC8BEA2F" httpUri="https://zenodo.org/record/262284/files/figure.png" pageId="19" pageNumber="20" targetBox="[307,1275,198,1523]" targetPageId="19">
<paragraph id="8BB93C72FFE2FFE2FF4EF99FFC8BEA2F" blockId="19.[151,1436,1550,1603]" pageId="19" pageNumber="20">
<emphasis id="B972E060FFE2FFE2FF4EF99FFEB1EA48" bold="true" box="[151,286,1550,1572]" pageId="19" pageNumber="20">FIGURE 11.</emphasis>
<taxonomicName id="4C0647F1FFE2FFE2FEFDF99FFDBEEA48" box="[292,529,1550,1572]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ligulatus" status="sp. nov.">
<emphasis id="B972E060FFE2FFE2FEFDF99FFDBEEA48" box="[292,529,1550,1572]" italics="true" pageId="19" pageNumber="20">Ischnothyreus ligulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415D1BFFE2FFE2FDCEF99EFDC8EA48" box="[535,615,1551,1572]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
Male: A, C, left chelicera, anterior view; B, right chelicera, anterior view; D, left chelicera, posterior view.
<emphasis id="B972E060FFE2FFE2FE44F9BCFE04EA2F" bold="true" box="[413,427,1581,1603]" pageId="19" pageNumber="20">p</emphasis>
, process. Scale bars: A, C = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C31C6FF9FFE2FFE2FF1EF9FEFB0DEAC0" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8BB93C72FFE2FFE2FF1EF9FEFB0DEAC0" blockId="19.[151,1436,1647,1996]" pageId="19" pageNumber="20">
<emphasis id="B972E060FFE2FFE2FF1EF9FEFEE1EAE4" bold="true" box="[199,334,1647,1672]" pageId="19" pageNumber="20">Etymology.</emphasis>
The specific name is an adjective and refers to the spoon-like structures on the base of the male cheliceral fang and the tip of the embolus. It is derived from the latin word for spoon, ligula.
</paragraph>
</subSubSection>
<subSubSection id="C31C6FF9FFE2FFE2FF1EF926FBC8EBA0" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BB93C72FFE2FFE2FF1EF926FD5BEB50" blockId="19.[151,1436,1647,1996]" pageId="19" pageNumber="20">
<emphasis id="B972E060FFE2FFE2FF1EF926FEEEEABC" bold="true" box="[199,321,1719,1744]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
Males of
<taxonomicName id="4C0647F1FFE2FFE2FE63F929FD99EABC" box="[442,566,1719,1744]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ligulatus" status="sp. nov.">
<emphasis id="B972E060FFE2FFE2FE63F929FD99EABC" box="[442,566,1719,1744]" italics="true" pageId="19" pageNumber="20">I. ligulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415D1BFFE2FFE2FD99F926FD33EAA3" box="[576,668,1719,1744]" pageId="19" pageNumber="20" rank="species">
<emphasis id="B972E060FFE2FFE2FD99F926FD3AEABC" bold="true" box="[576,661,1719,1744]" pageId="19" pageNumber="20">sp. nov</emphasis>
.
</taxonomicNameLabel>
resemble
<taxonomicName id="4C0647F1FFE2FFE2FCCDF929FB18EABC" authority="Kranz-Baltensperger, 2011" authorityName="Kranz-Baltensperger" authorityYear="2011" box="[788,1207,1719,1744]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="florifer">
<emphasis id="B972E060FFE2FFE2FCCDF929FCD1EABC" box="[788,894,1719,1744]" italics="true" pageId="19" pageNumber="20">I. florifer</emphasis>
Kranz-Baltensperger, 2011
</taxonomicName>
males as they have very similar pedipalps with a spoon-shaped, less sclerotized appendage on the embolic tip (
<emphasis id="B972E060FFE2FFE2FB46F94AFB6CEA98" bold="true" box="[1183,1219,1755,1780]" pageId="19" pageNumber="20">ssa</emphasis>
,
<figureCitation id="133D20F7FFE2FFE2FB08F94AFA83EA98" box="[1233,1324,1755,1780]" captionStart="FIGURE 10" captionStartId="18.[151,250,1882,1904]" captionTargetBox="[180,1395,197,1856]" captionTargetId="figure@18.[180,1406,193,1861]" captionTargetPageId="18" captionText="FIGURE 10. Ischnothyreus ligulatus sp. nov. Male: A, left palp, prolateral view; B, H, left palp, retrolate ¬ ral view; C, D, left embolus, prolateral view; E, F, left embolus, prolateral view; G, left palp, prolateral view (mirrored image). rl retrolateral lobe; ssa, spoon-shaped appendage. Scale bars: A, G = 0.2 mm." httpUri="https://zenodo.org/record/262283/files/figure.png" pageId="19" pageNumber="20">Figs. 10</figureCitation>
E, F, 12E, F). They can be seperated by the absence of the spoon-shaped process on the cheliceral fang in
<taxonomicName id="4C0647F1FFE2FFE2FB1DF891FA85EB74" box="[1220,1322,1791,1816]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="florifer">
<emphasis id="B972E060FFE2FFE2FB1DF891FA85EB74" box="[1220,1322,1791,1816]" italics="true" pageId="19" pageNumber="20">I. florifer</emphasis>
</taxonomicName>
. The bulb has a single large ventral protuberance (
<figureCitation id="133D20F7FFE2FFE2FD8CF8B2FD00EB50" box="[597,687,1827,1852]" captionStart="FIGURE 10" captionStartId="18.[151,250,1882,1904]" captionTargetBox="[180,1395,197,1856]" captionTargetId="figure@18.[180,1406,193,1861]" captionTargetPageId="18" captionText="FIGURE 10. Ischnothyreus ligulatus sp. nov. Male: A, left palp, prolateral view; B, H, left palp, retrolate ¬ ral view; C, D, left embolus, prolateral view; E, F, left embolus, prolateral view; G, left palp, prolateral view (mirrored image). rl retrolateral lobe; ssa, spoon-shaped appendage. Scale bars: A, G = 0.2 mm." httpUri="https://zenodo.org/record/262283/files/figure.png" pageId="19" pageNumber="20">Figs. 10</figureCitation>
G, H).
</paragraph>
<paragraph id="8BB93C72FFE2FFE2FF1EF8D6FBC8EBA0" blockId="19.[151,1436,1647,1996]" pageId="19" pageNumber="20">
Females of
<taxonomicName id="4C0647F1FFE2FFE2FE96F8D9FE64EB0C" box="[335,459,1863,1888]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ligulatus" status="sp. nov.">
<emphasis id="B972E060FFE2FFE2FE96F8D9FE64EB0C" box="[335,459,1863,1888]" italics="true" pageId="19" pageNumber="20">I. ligulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415D1BFFE2FFE2FE0DF8D6FD9FEB33" box="[468,560,1863,1888]" pageId="19" pageNumber="20" rank="species">
<emphasis id="B972E060FFE2FFE2FE0DF8D6FD85EB0C" bold="true" box="[468,554,1863,1888]" pageId="19" pageNumber="20">sp. nov</emphasis>
.
</taxonomicNameLabel>
resemble
<taxonomicName id="4C0647F1FFE2FFE2FD70F8D9FBFEEB0C" authority="Edwards &amp; Harvey, 2014" authorityName="Edwards &amp; Harvey" authorityYear="2014" box="[681,1105,1863,1888]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="barratus">
<emphasis id="B972E060FFE2FFE2FD70F8D9FC8CEB0C" box="[681,803,1863,1888]" italics="true" pageId="19" pageNumber="20">I. barratus</emphasis>
Edwards &amp;
<bibRefCitation id="EF974183FFE2FFE2FC61F8D9FBFEEB0C" author="Harvey" box="[952,1105,1864,1888]" pageId="19" pageNumber="20" refString="Edward, K. L. &amp; Harvey, M. S. (2014) Australian goblin spiders of the genus Ischnothyreus (Araneae, Oonopidae). Bulletin of the American Museum of Natural History, 389, 1 - 144. http: // dx. doi. org / 10.1206 / 865.1" type="journal article" year="2014">Harvey, 2014</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C0647F1FFE2FFE2FB54F8D9FE9AEBE8" authority="Edwards &amp; Harvey, 2014" authorityName="Edwards &amp; Harvey" authorityYear="2014" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="B972E060FFE2FFE2FB54F8D9FABEEB0C" box="[1165,1297,1863,1888]" italics="true" pageId="19" pageNumber="20">I. monteithi</emphasis>
Edwards &amp;
<bibRefCitation id="EF974183FFE2FFE2FF4EF8FDFE9AEBE8" author="Harvey" box="[151,309,1900,1924]" pageId="19" pageNumber="20" refString="Edward, K. L. &amp; Harvey, M. S. (2014) Australian goblin spiders of the genus Ischnothyreus (Araneae, Oonopidae). Bulletin of the American Museum of Natural History, 389, 1 - 144. http: // dx. doi. org / 10.1206 / 865.1" type="journal article" year="2014">Harvey, 2014</bibRefCitation>
</taxonomicName>
with small lateral sclerites flanking the postepigastric scutum (
<emphasis id="B972E060FFE2FFE2FBE4F8FAFBFFEBE8" bold="true" box="[1085,1104,1899,1924]" pageId="19" pageNumber="20">ls</emphasis>
,
<figureCitation id="133D20F7FFE2FFE2FBBCF8FAFB14EBE8" box="[1125,1211,1899,1924]" captionStart="FIGURE 15" captionStartId="23.[151,250,1377,1399]" captionTargetBox="[182,1406,189,1354]" captionTargetId="figure@23.[182,1406,189,1355]" captionTargetPageId="23" captionText="FIGURE 15. Ischnothyreus ligulatus sp. nov.: A, B, C, E, female genital area, ventral view; D, dorsal view. a, apodeme; ls, lateral sclerite; nsp, nail-shaped projection; o, opening of apodeme; s, stigma; wt, winding tube. Scale bars: A, E = 0.2 mm; C = 0.1 mm." httpUri="https://zenodo.org/record/262288/files/figure.png" pageId="19" pageNumber="20">Fig. 15</figureCitation>
A) and a roundish appearance of the winding tube. But the flanking apodemes are different and the nail-shaped process on the postepigastric scutum (
<emphasis id="B972E060FFE2FFE2FE42F822FE17EBA0" bold="true" box="[411,440,1971,1996]" pageId="19" pageNumber="20">wt</emphasis>
,
<emphasis id="B972E060FFE2FFE2FE1FF822FE7BEBA0" bold="true" box="[454,468,1971,1996]" pageId="19" pageNumber="20">a</emphasis>
,
<emphasis id="B972E060FFE2FFE2FE3BF822FDA3EBA0" bold="true" box="[482,524,1971,1996]" pageId="19" pageNumber="20">nsp</emphasis>
,
<figureCitation id="133D20F7FFE2FFE2FDC3F822FDDDEBA0" box="[538,626,1971,1996]" captionStart="FIGURE 15" captionStartId="23.[151,250,1377,1399]" captionTargetBox="[182,1406,189,1354]" captionTargetId="figure@23.[182,1406,189,1355]" captionTargetPageId="23" captionText="FIGURE 15. Ischnothyreus ligulatus sp. nov.: A, B, C, E, female genital area, ventral view; D, dorsal view. a, apodeme; ls, lateral sclerite; nsp, nail-shaped projection; o, opening of apodeme; s, stigma; wt, winding tube. Scale bars: A, E = 0.2 mm; C = 0.1 mm." httpUri="https://zenodo.org/record/262288/files/figure.png" pageId="19" pageNumber="20">Figs. 15</figureCitation>
AE, 16EG) is lacking in the other species.
</paragraph>
</subSubSection>
<caption id="DF796CFAFFE5FFE5FF4EF9F9FEA7EAB6" httpUri="https://zenodo.org/record/262285/files/figure.png" pageId="20" pageNumber="21" targetBox="[151,1436,183,1618]" targetPageId="20">
<paragraph id="8BB93C72FFE5FFE5FF4EF9F9FEA7EAB6" blockId="20.[151,1436,1640,1754]" pageId="20" pageNumber="21">
<emphasis id="B972E060FFE5FFE5FF4EF9F9FE8EEA12" bold="true" box="[151,289,1640,1662]" pageId="20" pageNumber="21">FIGURE 12.</emphasis>
<taxonomicName id="4C0647F1FFE5FFE5FEF1F9F9FDB8EA12" box="[296,535,1640,1662]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="ligulatus" status="sp. nov.">
<emphasis id="B972E060FFE5FFE5FEF1F9F9FDB8EA12" box="[296,535,1640,1662]" italics="true" pageId="20" pageNumber="21">Ischnothyreus ligulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415D1BFFE5FFE5FDC7F9F8FDDFEA12" box="[542,624,1641,1662]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
Male, SEM images: A, carapace, dorsal view; B, mouthparts, ventral view; C, palp, retrolateral view; D, palp, prolateral view; E, embolus, retrolateral view; F, embolus, prolateral view.
<emphasis id="B972E060FFE5FFE5FAF8F916FA96EAF1" bold="true" box="[1313,1337,1671,1693]" pageId="20" pageNumber="21">eo</emphasis>
, embolic opening;
<emphasis id="B972E060FFE5FFE5FF2FF937FEA7EAD0" bold="true" box="[246,264,1702,1724]" pageId="20" pageNumber="21">rl</emphasis>
, retrolateral lobe;
<emphasis id="B972E060FFE5FFE5FE1DF937FE4BEAD0" bold="true" box="[452,484,1702,1724]" pageId="20" pageNumber="21">ssa</emphasis>
, spoon-shaped appendage. Scale bars: A = 0.5 mm, B = 0.2 mm, C, D = 0.1 mm, E = 20 µm, F = 10 µm.
</paragraph>
</caption>
<subSubSection id="C31C6FF9FFE5FFE8FF1EF893FD45EEA1" lastPageId="25" lastPageNumber="26" pageId="20" pageNumber="21" type="description">
<paragraph id="8BB93C72FFE5FFE5FF1EF893FE3DEB52" blockId="20.[151,1436,1794,2035]" pageId="20" pageNumber="21">
<emphasis id="B972E060FFE5FFE5FF1EF893FE0FEB77" bold="true" box="[199,416,1794,1819]" pageId="20" pageNumber="21">Description. Male</emphasis>
: (PBI_OON00031890): Total length 1.65; carapace length: 0.79; carapace width: 0.65; abdomen length; 0.86.
</paragraph>
<paragraph id="8BB93C72FFE5FFE4FF1EF8DBFA33EB9A" blockId="20.[151,1436,1794,2035]" lastBlockId="21.[151,1436,1868,2038]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">CEPHALOTHORAX: Carapace pale orange, ovoid in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to between 0.5 and 0.75 times its maximum width, surface of elevated portion of pars cephalica smooth, sides smooth; lateral margin straight, smooth, present in one row. Clypeus margin unmodified, straight in front view, vertical in lateral view, high, ALE separated from edge of carapace by their radius or more; setae dark, needle-like. Eyes six, well developed, ALE largest, ALE circular, PME squared, PLE oval; posterior eye row straight from above, procurved from front; ALE touching, ALE-PLE separated by less than ALE radius, PME touching throughout most of their length, PLE-PME touching. Sternum as long as wide, pale orange, uniform, surface smooth, microsculpture absent, lateral margin without infra-coxal grooves, distance between coxae approximately equal, extensions of pre-coxal triangles present, lateral margins unmodified; setae dark, needle-like, densest laterally, originating from surface. Mouthparts: Chelicerae, endites and labium pale orange.</paragraph>
<caption id="DF796CFAFFE4FFE4FF4EF925FD84EB49" httpUri="https://zenodo.org/record/262286/files/figure.png" pageId="21" pageNumber="22" targetBox="[151,1436,193,1698]" targetPageId="21">
<paragraph id="8BB93C72FFE4FFE4FF4EF925FD84EB49" blockId="21.[151,1436,1716,1829]" pageId="21" pageNumber="22">
<emphasis id="B972E060FFE4FFE4FF4EF925FE8EEAA6" bold="true" box="[151,289,1716,1738]" pageId="21" pageNumber="22">FIGURE 13.</emphasis>
<taxonomicName id="4C0647F1FFE4FFE4FEF0F925FDB7EAA6" box="[297,536,1716,1738]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ligulatus" status="sp. nov.">
<emphasis id="B972E060FFE4FFE4FEF0F925FDB7EAA6" box="[297,536,1716,1738]" italics="true" pageId="21" pageNumber="22">Ischnothyreus ligulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415D1BFFE4FFE4FDF8F924FDDCEAA6" box="[545,627,1717,1738]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
Male: A, left endite, ventral view; B, chelicerae, ventral view; C, base of left cheliceral fang, anteroventral view; D, base of left cheliceral fang, ventral view; E, sperm pore, ventral view; F, claws of leg III; G, tarsal organ of leg III; H, surface of carapace cuticle, median part of pars cephalica.
<emphasis id="B972E060FFE4FFE4FBDCF960FBBCEB6B" bold="true" box="[1029,1043,1777,1799]" pageId="21" pageNumber="22">p</emphasis>
, process. Scale bars: A, B, D = 50 µm; C, E, H = 20 µm; F = 10 µm; G = 2 µm.
</paragraph>
</caption>
<caption id="DF796CFAFFE7FFE7FF4EF8C7FD28EBC6" httpUri="https://zenodo.org/record/262287/files/figure.png" pageId="22" pageNumber="23" targetBox="[151,1436,193,1856]" targetPageId="22">
<paragraph id="8BB93C72FFE7FFE7FF4EF8C7FD28EBC6" blockId="22.[151,1436,1878,1962]" pageId="22" pageNumber="23">
<emphasis id="B972E060FFE7FFE7FF4EF8C7FE8EEB00" bold="true" box="[151,289,1878,1900]" pageId="22" pageNumber="23">FIGURE 14.</emphasis>
<taxonomicName id="4C0647F1FFE7FFE7FEF0F8C7FDB7EB00" box="[297,536,1878,1900]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ligulatus" status="sp. nov.">
<emphasis id="B972E060FFE7FFE7FEF0F8C7FDB7EB00" box="[297,536,1878,1900]" italics="true" pageId="22" pageNumber="23">Ischnothyreus ligulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415D1BFFE7FFE7FDF9F8C6FDDDEB00" box="[544,626,1879,1900]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
Female: A, habitus, dorsal view; B, habitus, ventral view; C carapace, frontal view; D, habitus, lateral view; E, opisthosoma, dorsal view; F, carapace, dorsal view; G, opisthosoma, lateral view; H, opisthosoma, ventral view. Scale bar: A = 1 mm.
</paragraph>
</caption>
<caption id="DF796CFAFFE6FFE6FF4EFAF0FEAAE9D9" httpUri="https://zenodo.org/record/262288/files/figure.png" pageId="23" pageNumber="24" targetBox="[182,1406,189,1354]" targetPageId="23">
<paragraph id="8BB93C72FFE6FFE6FF4EFAF0FEAAE9D9" blockId="23.[151,1435,1377,1461]" pageId="23" pageNumber="24">
<emphasis id="B972E060FFE6FFE6FF4EFAF0FE8EE91B" bold="true" box="[151,289,1377,1399]" pageId="23" pageNumber="24">FIGURE 15.</emphasis>
<taxonomicName id="4C0647F1FFE6FFE6FEF1FAF0FDB9E91B" box="[296,534,1377,1399]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="ligulatus" status="sp. nov.">
<emphasis id="B972E060FFE6FFE6FEF1FAF0FDB9E91B" box="[296,534,1377,1399]" italics="true" pageId="23" pageNumber="24">Ischnothyreus ligulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415D1BFFE6FFE6FDC6FAF3FDDBE91B" box="[543,628,1378,1399]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
: A, B, C, E, female genital area, ventral view; D, dorsal view.
<emphasis id="B972E060FFE6FFE6FAD9FAF0FAA3E91B" bold="true" box="[1280,1292,1377,1399]" pageId="23" pageNumber="24">a</emphasis>
, apodeme;
<emphasis id="B972E060FFE6FFE6FA5DFAF0FA3AE91B" bold="true" box="[1412,1429,1377,1399]" pageId="23" pageNumber="24">ls</emphasis>
, lateral sclerite;
<emphasis id="B972E060FFE6FFE6FEEDFA11FEF5E9FA" bold="true" box="[308,346,1408,1430]" pageId="23" pageNumber="24">nsp</emphasis>
, nail-shaped projection;
<emphasis id="B972E060FFE6FFE6FD8FFA11FDCDE9FA" bold="true" box="[598,610,1408,1430]" pageId="23" pageNumber="24">o</emphasis>
, opening of apodeme;
<emphasis id="B972E060FFE6FFE6FC94FA11FCF8E9FA" bold="true" box="[845,855,1408,1430]" pageId="23" pageNumber="24">s</emphasis>
, stigma;
<emphasis id="B972E060FFE6FFE6FC6CFA11FC60E9FA" bold="true" box="[949,975,1408,1430]" pageId="23" pageNumber="24">wt</emphasis>
, winding tube. Scale bars: A, E = 0.2 mm; C = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BB93C72FFE6FFE6FF4EFA4FFEEEEAA3" blockId="23.[151,1437,1502,2031]" pageId="23" pageNumber="24">
Chelicerae straight, anterior face unmodified; fangs directed posteriorly, shape normal, fang base with spoonshaped process (
<emphasis id="B972E060FFE6FFE6FE8BF992FECEEA70" bold="true" box="[338,353,1539,1564]" pageId="23" pageNumber="24">p</emphasis>
,
<figureCitation id="133D20F7FFE6FFE6FEA8F992FE64EA77" box="[369,459,1539,1564]" captionStart="FIGURE 11" captionStartId="19.[151,250,1550,1572]" captionTargetBox="[307,1275,198,1523]" captionTargetId="figure@19.[297,1290,193,1529]" captionTargetPageId="19" captionText="FIGURE 11. Ischnothyreus ligulatus sp. nov. Male: A, C, left chelicera, anterior view; B, right chelicera, anterior view; D, left chelicera, posterior view. p, process. Scale bars: A, C = 0.1 mm." httpUri="https://zenodo.org/record/262284/files/figure.png" pageId="23" pageNumber="24">Figs. 11</figureCitation>
C, D, 13BD); tip unmodified; setae light, needle-like, evenly scattered; paturon inner margin with scattered setae. Labium rectangular, not fused to sternum, anterior margin not indented at middle, same as sternum in sclerotization; with 35 setae on anterior margin, subdistal portion with unmodified setae. Endites distally not excavated, serrula absent, anteromedian tip with one strong, tooth-like projection, posteromedian part unmodified, same as sternum in sclerotization, with anterolateral, strongly sclerotized toothlike projection.
</paragraph>
<paragraph id="8BB93C72FFE6FFE6FF1EF94AFB91EB83" blockId="23.[151,1437,1502,2031]" pageId="23" pageNumber="24">ABDOMEN: cylindrical, rounded posteriorly; dorsum soft portions white, without color pattern. Posterior spiracles not connected by groove. Scutum not extending far dorsal of pedicel, cuticular outgrowths near pedicel absent. Dorsal scutum weakly sclerotized, yellow-brown, without color pattern, covering more than 3/4 of abdomen, more than 1/2 to most of abdomen width, not fused to epigastric scutum. Epigastric scutum weakly sclerotized, smooth, surrounding pedicel, not protruding. Postepigastric scutum weakly sclerotized, pale orange, long, almost rectangular, covering about 1/2 of abdominal length, not fused to epigastric scutum. Dorsum setae dark, needle-like. Epigastric area setae uniform, dark, needle-like. Postepigastric area setae dark, needle-like. Spinneret scutum with fringe of needle-like setae. Interscutal membrane with setae.</paragraph>
<caption id="DF796CFAFFE9FFE9FF4EF91CFD82EA93" httpUri="https://zenodo.org/record/262289/files/figure.png" pageId="24" pageNumber="25" targetBox="[174,1416,189,1655]" targetPageId="24">
<paragraph id="8BB93C72FFE9FFE9FF4EF91CFD82EA93" blockId="24.[151,1436,1677,1791]" pageId="24" pageNumber="25">
<emphasis id="B972E060FFE9FFE9FF4EF91CFE8CEACF" bold="true" box="[151,291,1677,1699]" pageId="24" pageNumber="25">FIGURE 16.</emphasis>
<taxonomicName id="4C0647F1FFE9FFE9FEF5F91CFDB2EACF" box="[300,541,1677,1699]" class="Arachnida" family="Oonopidae" genus="Ischnothyreus" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="ligulatus" status="sp. nov.">
<emphasis id="B972E060FFE9FFE9FEF5F91CFDB2EACF" box="[300,541,1677,1699]" italics="true" pageId="24" pageNumber="25">Ischnothyreus ligulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2415D1BFFE9FFE9FDFFF91FFDD6EACF" box="[550,633,1678,1699]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
Female: A, eyes, dorsal view; B, right endite with serrula, ventral view; C, sternum, ventral view; D, mouthparts, ventral view; E, postepigastric scutum, ventral view; F, row of pores, ventral view; G, nail-shaped process, ventral view.
<emphasis id="B972E060FFE9FFE9FE2DF95BFDB5EA8C" bold="true" box="[500,538,1738,1760]" pageId="24" pageNumber="25">nsp</emphasis>
, nail-shaped projection;
<emphasis id="B972E060FFE9FFE9FCCDF95BFC8FEA8C" bold="true" box="[788,800,1738,1760]" pageId="24" pageNumber="25">o</emphasis>
, opening of apodeme;
<emphasis id="B972E060FFE9FFE9FBD0F95BFB8BEA8C" bold="true" box="[1033,1060,1738,1760]" pageId="24" pageNumber="25">po</emphasis>
, pores. Scale bars: A, C = 0.2 mm; B = 50 µm; D, E = 0.1 mm; F, G = 20 µm.
</paragraph>
</caption>
<paragraph id="8BB93C72FFE9FFE9FF1EF8B6FE92EBEB" blockId="24.[151,1437,1830,2035]" pageId="24" pageNumber="25">LEGS: yellow, without color pattern; femur IV same size as femora I-III, patella plus tibia I near as long as carapace, tibia I unmodified. Leg spination: femora: I p0-1-1; II p0-0-1; tibiae: I, II p2-1-1; r2-1-1; metatarsi: I, II p1-1-0; r1-1-0.</paragraph>
<paragraph id="8BB93C72FFE9FFE9FF1EF802FBFDEB9F" blockId="24.[151,1437,1830,2035]" pageId="24" pageNumber="25">
GENITALIA: Epigastric region with sperm pore large, oval, situated at level of anterior spiracles, unmodified; without setae. Bulb with one large ventral protuberance, retrolateral lobe simple. Tip of bulb slightly elongated, spoon-like, with spoon-shaped less-sclerotized appendage (
<emphasis id="B972E060FFE9FFE9FCE9F84BFCFBEB9F" bold="true" box="[816,852,2010,2035]" pageId="24" pageNumber="25">ssa</emphasis>
,
<figureCitation id="133D20F7FFE9FFE9FCBBF84BFC13EB9F" box="[866,956,2010,2035]" captionStart="FIGURE 10" captionStartId="18.[151,250,1882,1904]" captionTargetBox="[180,1395,197,1856]" captionTargetId="figure@18.[180,1406,193,1861]" captionTargetPageId="18" captionText="FIGURE 10. Ischnothyreus ligulatus sp. nov. Male: A, left palp, prolateral view; B, H, left palp, retrolate ¬ ral view; C, D, left embolus, prolateral view; E, F, left embolus, prolateral view; G, left palp, prolateral view (mirrored image). rl retrolateral lobe; ssa, spoon-shaped appendage. Scale bars: A, G = 0.2 mm." httpUri="https://zenodo.org/record/262283/files/figure.png" pageId="24" pageNumber="25">Figs. 10</figureCitation>
E, F, 12E, F).
</paragraph>
<paragraph id="8BB93C72FFE8FFE8FF1EFF06FE8BECB8" blockId="25.[151,1437,151,752]" pageId="25" pageNumber="26">
<emphasis id="B972E060FFE8FFE8FF1EFF06FEB1ECDC" bold="true" box="[199,286,151,176]" pageId="25" pageNumber="26">Female</emphasis>
: (PBI_OON00031890). Total length 1.71 carapace length: 0.79; carapace width: 0.70; abdomen length; 0.98.
</paragraph>
<paragraph id="8BB93C72FFE8FFE8FF1EFF71FE53EC94" blockId="25.[151,1437,151,752]" box="[199,508,223,248]" pageId="25" pageNumber="26">As in male except as noted.</paragraph>
<paragraph id="8BB93C72FFE8FFE8FF1EFE95FE8FED2C" blockId="25.[151,1437,151,752]" pageId="25" pageNumber="26">CEPHALOTHORAX: Female palp claws absent; spines formula: femur p1-1-1. Serrula present. Endites unmodified.</paragraph>
<paragraph id="8BB93C72FFE8FFE8FF1EFEDDFE7AEE51" blockId="25.[151,1437,151,752]" pageId="25" pageNumber="26">ABDOMEN: ovoid, pointed posteriorly; dorsum soft portions yellow-brown. Dorsal scutum orange-brown, covering about 1/2 of abdomen. Epigastric scutum small, lateral sclerites present. Postepigastric scutum yellowbrown, widely hexagonal, only around epigastric furrow, with short posteriorly directed lateral apodemes, characteristic posteriorly pointing nail-shaped projection on ventral surface; may be hardly visible because situated beneath darkly sclerotized winding tube. LEGS: pale orange; patella plus tibia I longer than carapace. Leg spination: femora: I p0-1-1; II p0-0-1; tibiae: I p2-1-1; r2-1-1; II p2-1-1; r2-2-0; metatarsi: I, II p1-1-0; r1-1-0. Proclaws inner face smooth.</paragraph>
<paragraph id="8BB93C72FFE8FFE8FF1EFDD9FD45EEA1" blockId="25.[151,1437,151,752]" pageId="25" pageNumber="26">
GENITALIA: strongly sclerotized winding tube with globular appearance, apodemes lateral of winding tube with openings to surface of postepigastric scutum. Postepigastric scutum with central posteriorly pointing nailshaped process (
<emphasis id="B972E060FFE8FFE8FE80FD1EFED8EEC4" bold="true" box="[345,375,655,680]" pageId="25" pageNumber="26">wt</emphasis>
,
<emphasis id="B972E060FFE8FFE8FE50FD1EFE38EEC4" bold="true" box="[393,407,655,680]" pageId="25" pageNumber="26">a</emphasis>
,
<emphasis id="B972E060FFE8FFE8FE73FD1EFE7BEEC4" bold="true" box="[426,468,655,680]" pageId="25" pageNumber="26">nsp</emphasis>
,
<figureCitation id="133D20F7FFE8FFE8FE01FD1EFD97EEC4" box="[472,568,655,680]" captionStart="FIGURE 15" captionStartId="23.[151,250,1377,1399]" captionTargetBox="[182,1406,189,1354]" captionTargetId="figure@23.[182,1406,189,1355]" captionTargetPageId="23" captionText="FIGURE 15. Ischnothyreus ligulatus sp. nov.: A, B, C, E, female genital area, ventral view; D, dorsal view. a, apodeme; ls, lateral sclerite; nsp, nail-shaped projection; o, opening of apodeme; s, stigma; wt, winding tube. Scale bars: A, E = 0.2 mm; C = 0.1 mm." httpUri="https://zenodo.org/record/262288/files/figure.png" pageId="25" pageNumber="26">Figs. 15</figureCitation>
AE, 16EG). Row of small pores from anterior middle of postepigastric scutum to base of nail-shaped process (
<emphasis id="B972E060FFE8FFE8FD96FD25FDC2EEA1" bold="true" box="[591,621,692,717]" pageId="25" pageNumber="26">po</emphasis>
,
<figureCitation id="133D20F7FFE8FFE8FDA2FD25FD64EEA1" box="[635,715,692,717]" captionStart="FIGURE 16" captionStartId="24.[151,250,1677,1699]" captionTargetBox="[174,1416,189,1655]" captionTargetId="figure@24.[174,1418,189,1659]" captionTargetPageId="24" captionText="FIGURE 16. Ischnothyreus ligulatus sp. nov. Female: A, eyes, dorsal view; B, right endite with serrula, ventral view; C, sternum, ventral view; D, mouthparts, ventral view; E, postepigastric scutum, ventral view; F, row of pores, ventral view; G, nail-shaped process, ventral view. nsp, nail-shaped projection; o, opening of apodeme; po, pores. Scale bars: A, C = 0.2 mm; B = 50 µm; D, E = 0.1 mm; F, G = 20 µm." httpUri="https://zenodo.org/record/262289/files/figure.png" pageId="25" pageNumber="26">Fig. 16</figureCitation>
F).
</paragraph>
</subSubSection>
<subSubSection id="C31C6FF9FFE8FFE8FF1EFD46FB88EE83" box="[199,1063,727,752]" pageId="25" pageNumber="26" type="distribution">
<paragraph id="8BB93C72FFE8FFE8FF1EFD46FB88EE83" blockId="25.[151,1437,151,752]" box="[199,1063,727,752]" pageId="25" pageNumber="26">
<emphasis id="B972E060FFE8FFE8FF1EFD46FECFEE9C" bold="true" box="[199,352,727,752]" pageId="25" pageNumber="26">Distribution.</emphasis>
Only known from Ujung Kulon National Park in western
<collectingRegion id="49C2F290FFE8FFE8FC34FD49FB8CEE83" box="[1005,1059,728,751]" country="Indonesia" name="Jawa Timur" pageId="25" pageNumber="26">Java</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>