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<document id="CF08DDBA862D43D94F585BC6D095342B" ID-DOI="10.11646/zootaxa.3860.6.5" ID-GBIF-Dataset="fadaaee1-ecee-4e69-acdd-a8253a6af8a6" ID-ISSN="1175-5326" ID-Zenodo-Dep="228451" ID-ZooBank="BAFCC005-6DF2-458D-8C75-F261F27B429F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460908168503" checkinUser="plazi" docAuthor="Campos, Martha R. &amp; Magalhães, Célio" docDate="2014" docId="03AF87B82F2EF177FF06FE5FB69EF9EF" docLanguage="en" docName="zt03860p579.pdf" docOrigin="Zootaxa 3860 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eidocamptophallus chacei Pretzmann 1967" docType="treatment" docVersion="8" lastPageNumber="578" masterDocId="FF96FFC02F2BF170FF91FD1CB453FFE7" masterDocTitle="Colombiathelphusa, a new genus of freshwater crab from Colombia, and the first location record of Eidocamptophallus chacei (Pretzmann, 1967) (Crustacea: Decapoda: Pseudothelphusidae)" masterLastPageNumber="579" masterPageNumber="571" pageNumber="576" updateTime="1698360976663" updateUser="plazi">
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<mods:title id="6A6D4CAEA0120F88A92D9BD9B8F7C29D">Colombiathelphusa, a new genus of freshwater crab from Colombia, and the first location record of Eidocamptophallus chacei (Pretzmann, 1967) (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<mods:namePart id="1FCB3D4CF7A124BB3EE46FDB323D946F">Campos, Martha R.</mods:namePart>
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<mods:namePart id="6C7B336A4EE9323CDE9A3C1FF67C7923">Magalhães, Célio</mods:namePart>
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<mods:date id="ACBA36B153BC9C1193A412398F707B20">2014</mods:date>
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<treatment id="03AF87B82F2EF177FF06FE5FB69EF9EF" ID-DOI="http://doi.org/10.5281/zenodo.6130485" ID-GBIF-Taxon="119588229" ID-Zenodo-Dep="6130485" LSID="urn:lsid:plazi:treatment:03AF87B82F2EF177FF06FE5FB69EF9EF" httpUri="http://treatment.plazi.org/id/03AF87B82F2EF177FF06FE5FB69EF9EF" lastPageId="7" lastPageNumber="578" pageId="5" pageNumber="576">
<subSubSection id="C31C65252F2EF175FF06FE5FB4B8FC67" pageId="5" pageNumber="576" type="nomenclature">
<paragraph id="8BB936AE2F2EF175FF06FE5FB6FFFCBA" blockId="5.[151,684,835,896]" box="[151,684,835,861]" pageId="5" pageNumber="576">
<heading id="D0F181C22F2EF175FF06FE5FB6FFFCBA" bold="true" box="[151,684,835,861]" fontSize="10" level="2" pageId="5" pageNumber="576" reason="2">
<taxonomicName id="4C064D2D2F2EF175FF06FE5FB6FFFCBA" ID-CoL="38VD5" authority="Pretzmann, 1967" authorityName="Pretzmann" authorityYear="1967" box="[151,684,835,861]" class="Malacostraca" family="Pseudothelphusidae" genus="Eidocamptophallus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2EF175FF06FE5FB6FFFCBA" bold="true" box="[151,684,835,861]" pageId="5" pageNumber="576">
<emphasis id="B972EABC2F2EF175FF06FE5FB59AFCBB" bold="true" box="[151,457,835,860]" italics="true" pageId="5" pageNumber="576">Eidocamptophallus chacei</emphasis>
(
<bibRefCitation id="EF974B5F2F2EF175FE46FE58B6F7FCBA" author="Pretzmann" box="[471,676,836,861]" pageId="5" pageNumber="576" refString="Pretzmann, G. (1967) Neue Sudamerikanische Susswasserkrabben der Gattung Pseudothelphusa. Entomologisches Nachrichtenblatt, Wien, 15, 1 - 15." type="journal article" year="1967">Pretzmann, 1967</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB936AE2F2EF175FF06FE7BB4B8FC67" blockId="5.[151,684,835,896]" box="[151,235,871,896]" pageId="5" pageNumber="576">
(
<figureCitation id="133D2A2B2F2EF175FF0EFE7BB4B1FC67" box="[159,226,871,896]" captionStart="FIGURE 3" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[245,1352,194,1892]" captionTargetId="figure@6.[234,1353,193,1903]" captionTargetPageId="6" captionText="FIGURE 3. Eidocamptophallus chacei (Pretzmann, 1967), male, ICN-MHN-CR 2754. A, habitus; B, left first gonopod, caudal view; C, same, cephalic view; D, same, mesial view; E, same, detail of apex, latero-cephalic view; F, left third maxilliped, external view. 1, mesial process; 2, distal lobe of caudal surface; 3, marginal process; 4, marginal tooth; 5, lateral lobe; 6, cephalic lobe; 7, cephalic processes; 8, spermatic channel and field of apical spines." httpUri="https://zenodo.org/record/228454/files/figure.png" pageId="5" pageNumber="576">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31C65252F2EF175FF06FEADB751FBC5" pageId="5" pageNumber="576" type="reference_group">
<paragraph id="8BB936AE2F2EF175FF06FEADB773FC21" blockId="5.[151,1302,944,1058]" box="[151,800,944,966]" pageId="5" pageNumber="576">
<treatmentCitationGroup id="AB1611802F2EF175FF06FEADB773FC21" box="[151,800,944,966]" pageId="5" pageNumber="576">
<taxonomicName id="4C064D2D2F2EF175FF06FEADB74FFC21" ID-CoL="4M3FJ" authority="Pretzmann, 1967: 24" authorityName="Pretzmann" authorityPageNumber="24" authorityYear="1967" box="[151,796,944,966]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="chacei" subGenus="Potamocarcinus">
<emphasis id="B972EABC2F2EF175FF06FEADB613FC21" box="[151,576,944,966]" italics="true" pageId="5" pageNumber="576">Potamocarcinus (Potamocarcinus) chacei</emphasis>
<treatmentCitation id="0AA710BF2F2EF175FDD6FEADB74FFC21" author="Pretzmann" box="[583,796,944,966]" page="24" pageId="5" pageNumber="576" year="1967">
<bibRefCitation id="EF974B5F2F2EF175FDD6FEADB6A5FC21" author="Pretzmann" box="[583,758,944,966]" pageId="5" pageNumber="576" refString="Pretzmann, G. (1967) Neue Sudamerikanische Susswasserkrabben der Gattung Pseudothelphusa. Entomologisches Nachrichtenblatt, Wien, 15, 1 - 15." type="journal article" year="1967">Pretzmann, 1967</bibRefCitation>
: 24
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB936AE2F2EF175FF06FECCB6ACFBE4" blockId="5.[151,1302,944,1058]" pageId="5" pageNumber="576">
<treatmentCitationGroup id="AB1611802F2EF175FF06FECCB6ACFBE4" pageId="5" pageNumber="576">
<taxonomicName id="4C064D2D2F2EF175FF06FECCB664FC03" ID-CoL="4M3FJ" box="[151,567,974,997]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="chacei" subGenus="Megathelphusa">
<emphasis id="B972EABC2F2EF175FF06FECCB664FC03" box="[151,567,974,997]" italics="true" pageId="5" pageNumber="576">Potamocarcinus (Megathelphusa) chacei</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AA710BF2F2EF175FDC5FED3B770FC03" author="Pretzmann" box="[596,803,974,996]" page="20" pageId="5" pageNumber="576" year="1971">
<bibRefCitation id="EF974B5F2F2EF175FDC5FED3B753FC03" author="Pretzmann" box="[596,768,974,996]" pageId="5" pageNumber="576" refString="Pretzmann, G. (1971) Fortschritte in der Klassifizierung der Pseudothelphusidae. Anzeiger der Mathematisch Naturwissenschaftliche der Osterreichischen Akademie der Wissenschaften, 179 (1 - 4), 14 - 24. [(1)]" type="journal article" year="1971">Pretzmann 1971</bibRefCitation>
: 20
</treatmentCitation>
.—
<treatmentCitation id="0AA710BF2F2EF175FCD1FED2B043FC03" author="Pretzmann" box="[832,1040,974,996]" page="73" pageId="5" pageNumber="576" year="1972">
<bibRefCitation id="EF974B5F2F2EF175FCD1FED2B7B8FC03" author="Pretzmann" box="[832,1003,974,996]" pageId="5" pageNumber="576" refString="Pretzmann, G. (1972) Die Pseudothelphusidae (Crustacea Brachyura). Zoologica, 42 (120), 1 - 182. [pt. 1]" type="journal article" year="1972">Pretzmann 1972</bibRefCitation>
: 73
</treatmentCitation>
, figs. 390392, 402404.
<taxonomicName id="4C064D2D2F2EF175FF06FEF2B5D7FBE4" ID-CoL="4M3FJ" box="[151,388,1005,1027]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2EF175FF06FEF2B5D7FBE4" box="[151,388,1005,1027]" italics="true" pageId="5" pageNumber="576">Potamocarcinus chacei</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AA710BF2F2EF175FE33FEF1B628FBE4" author="Rodriguez" box="[418,635,1005,1027]" page="120" pageId="5" pageNumber="576" year="1982">
<bibRefCitation id="EF974B5F2F2EF175FE33FEF1B61AFBE4" author="Rodriguez" box="[418,585,1005,1027]" pageId="5" pageNumber="576" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Familie des Pseudothelphusidae. Vol. 22. Faune Tropicale. Paris, ORSTOM, 223 pp." type="book" year="1982">Rodríguez 1982</bibRefCitation>
: 120
</treatmentCitation>
, figs. 78, 79.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB936AE2F2EF175FF06F910B751FBC5" blockId="5.[151,1302,944,1058]" box="[151,770,1036,1058]" pageId="5" pageNumber="576">
<treatmentCitationGroup id="AB1611802F2EF175FF06F910B751FBC5" box="[151,770,1036,1058]" pageId="5" pageNumber="576">
<taxonomicName id="4C064D2D2F2EF175FF06F910B5F7FBC5" ID-CoL="38VD5" box="[151,420,1036,1058]" class="Malacostraca" family="Pseudothelphusidae" genus="Eidocamptophallus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2EF175FF06F910B5F7FBC5" box="[151,420,1036,1058]" italics="true" pageId="5" pageNumber="576">Eidocamptophallus chacei</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AA710BF2F2EF175FE50F910B6ADFBC5" author="Rodriguez" box="[449,766,1036,1058]" page="191" pageId="5" pageNumber="576" year="1989">
<bibRefCitation id="EF974B5F2F2EF175FE50F910B69FFBC5" author="Rodriguez" box="[449,716,1036,1058]" pageId="5" pageNumber="576" refString="Rodriguez, G. &amp; Hobbs, H. H. Jr. (1989) A new cavernicolous crab, Zilchia falcata, from Guatemala, with notes on the genera of the Potamocarcinini (Crustacea: Decapoda: Pseudothelphusidae). Bulletin du Museum national d'Histoire naturelle (Paris), serie 4, section A, 1, 183 - 192." type="journal article" year="1989">Rodríguez &amp; Hobbs 1989</bibRefCitation>
: 191
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31C65252F2EF175FF06F952B637FAFC" pageId="5" pageNumber="576" type="materials_examined">
<paragraph id="8BB936AE2F2EF175FF06F952B733FB48" blockId="5.[151,1436,1102,1991]" pageId="5" pageNumber="576">
<emphasis id="B972EABC2F2EF175FF06F952B555FB80" bold="true" box="[151,262,1102,1127]" pageId="5" pageNumber="576">Material.</emphasis>
Male, cl
<quantity id="4CFE9B4B2F2EF175FEEDF952B5B5FB81" box="[380,486,1102,1127]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="5" pageNumber="576" unit="mm" value="14.5">14.5 mm</quantity>
, cb
<quantity id="4CFE9B4B2F2EF175FD8AF953B6D0FB81" box="[539,643,1103,1127]" metricMagnitude="-2" metricUnit="m" metricValue="2.23" pageId="5" pageNumber="576" unit="mm" value="22.3">22.3 mm</quantity>
(ICN-MHN-CR 2754); Municipio Alto Baudó, Resguardo Indígena Nauca, Comunidad Puerto Alegre, trail to La Verrugosa stream, elevation
<quantity id="4CFE9B4B2F2EF175FC79F96FB060FB6D" box="[1000,1075,1139,1163]" metricMagnitude="2" metricUnit="m" metricValue="1.22" pageId="5" pageNumber="576" unit="m" value="122.0">122 m</quantity>
, Chocó Department,
<collectingCountry id="F311763E2F2EF175FAB6F96EB1CBFB6C" box="[1319,1432,1138,1163]" name="Colombia" pageId="5" pageNumber="576">Colombia</collectingCountry>
,
<geoCoordinate id="EE3250692F2EF175FF06F98AB56AFB49" box="[151,313,1174,1199]" direction="north" orientation="latitude" pageId="5" pageNumber="576" precision="1" value="5.613325">5º3647.97” N</geoCoordinate>
<geoCoordinate id="EE3250692F2EF175FED1F98AB5ABFB49" box="[320,504,1174,1199]" direction="west" orientation="longitude" pageId="5" pageNumber="576" precision="1" value="-77.06571">77º0356.55” W</geoCoordinate>
,
<date id="FFB8106E2F2EF175FD92F98BB6DEFB48" box="[515,653,1175,1199]" pageId="5" pageNumber="576" value="2014-01-29">29 Jan 2014</date>
, leg. G. F. Medina.
</paragraph>
<paragraph id="8BB936AE2F2EF175FF56F9A6B637FAFC" blockId="5.[151,1436,1102,1991]" pageId="5" pageNumber="576">
<emphasis id="B972EABC2F2EF175FF56F9A6B66FFB34" bold="true" box="[199,572,1210,1235]" pageId="5" pageNumber="576">Additional material examined.</emphasis>
<taxonomicName id="4C064D2D2F2EF175FDD6F9A6B052FB34" authority="Smalley, 1965: 1" authorityName="Smalley" authorityPageNumber="1" authorityYear="1965" box="[583,1025,1210,1235]" class="Malacostraca" family="Pseudothelphusidae" genus="Camptophallus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="botti">
<emphasis id="B972EABC2F2EF175FDD6F9A6B762FB34" box="[583,817,1210,1235]" italics="true" pageId="5" pageNumber="576">Camptophallus botti</emphasis>
Smalley, 1965:
<specimenCount id="9D00FD272F2EF175FC62F9A7B052FB34" box="[1011,1025,1211,1235]" pageId="5" pageNumber="576" type="male">1</specimenCount>
</taxonomicName>
male, cl
<quantity id="4CFE9B4B2F2EF175FBE4F9A7B08CFB35" box="[1141,1247,1211,1235]" metricMagnitude="-2" metricUnit="m" metricValue="1.29" pageId="5" pageNumber="576" unit="mm" value="12.9">12.9 mm</quantity>
, cb 19.0 mm,
<specimenCount id="9D00FD272F2EF175FA1FF9A7B4B5FB10" pageId="5" pageNumber="576" type="female">1 female</specimenCount>
, cl
<quantity id="4CFE9B4B2F2EF175FE89F9C3B5D7FB11" box="[280,388,1247,1271]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="5" pageNumber="576" unit="mm" value="14.3">14.3 mm</quantity>
, cb
<quantity id="4CFE9B4B2F2EF175FE2CF9C2B675FB11" box="[445,550,1246,1271]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" pageId="5" pageNumber="576" unit="mm" value="22.5">22.5 mm</quantity>
(
<collectionCode id="ED17AE6B2F2EF175FDABF9C3B628FB10" box="[570,635,1247,1271]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="576">INPA</collectionCode>
1489),
<collectingCountry id="F311763E2F2EF175FD4CF9C2B700FB10" box="[733,851,1246,1271]" name="Nicaragua" pageId="5" pageNumber="576">Nicaragua</collectingCountry>
, Jinotega Department, Cerro Kilabé,
<geoCoordinate id="EE3250692F2EF175FA80F9C3B1CFFB11" box="[1297,1436,1247,1271]" direction="north" orientation="latitude" pageId="5" pageNumber="576" precision="9" value="13.5685">13°34.11' N</geoCoordinate>
<geoCoordinate id="EE3250692F2EF175FF06F81EB577FAFD" box="[151,292,1282,1307]" direction="west" orientation="longitude" pageId="5" pageNumber="576" precision="9" value="-85.69683">85°41.81' W</geoCoordinate>
,
<date id="FFB8106E2F2EF175FEBEF81EB5E6FAFC" box="[303,437,1282,1307]" pageId="5" pageNumber="576" value="1998-08-06">6 Aug 1998</date>
, leg. G. Köhler.
</paragraph>
</subSubSection>
<subSubSection id="C31C65252F2EF175FF56F83AB7FBF92C" pageId="5" pageNumber="576" type="description">
<paragraph id="8BB936AE2F2EF175FF56F83AB7FBF92C" blockId="5.[151,1436,1102,1991]" pageId="5" pageNumber="576">
<emphasis id="B972EABC2F2EF175FF56F83AB6E3FAD8" bold="true" box="[199,688,1318,1343]" pageId="5" pageNumber="576">Redescription of the male first gonopod.</emphasis>
(
<figureCitation id="133D2A2B2F2EF175FD53F83AB755FAD8" box="[706,774,1318,1343]" captionStart="FIGURE 3" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[245,1352,194,1892]" captionTargetId="figure@6.[234,1353,193,1903]" captionTargetPageId="6" captionText="FIGURE 3. Eidocamptophallus chacei (Pretzmann, 1967), male, ICN-MHN-CR 2754. A, habitus; B, left first gonopod, caudal view; C, same, cephalic view; D, same, mesial view; E, same, detail of apex, latero-cephalic view; F, left third maxilliped, external view. 1, mesial process; 2, distal lobe of caudal surface; 3, marginal process; 4, marginal tooth; 5, lateral lobe; 6, cephalic lobe; 7, cephalic processes; 8, spermatic channel and field of apical spines." httpUri="https://zenodo.org/record/228454/files/figure.png" pageId="5" pageNumber="576">Fig. 3</figureCitation>
A-F). Male first gonopod straight, narrower in mid-distal portion, wider in mid-proximal portion (
<figureCitation id="133D2A2B2F2EF175FDFCF856B6E0FA84" box="[621,691,1354,1379]" captionStart="FIGURE 3" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[245,1352,194,1892]" captionTargetId="figure@6.[234,1353,193,1903]" captionTargetPageId="6" captionText="FIGURE 3. Eidocamptophallus chacei (Pretzmann, 1967), male, ICN-MHN-CR 2754. A, habitus; B, left first gonopod, caudal view; C, same, cephalic view; D, same, mesial view; E, same, detail of apex, latero-cephalic view; F, left third maxilliped, external view. 1, mesial process; 2, distal lobe of caudal surface; 3, marginal process; 4, marginal tooth; 5, lateral lobe; 6, cephalic lobe; 7, cephalic processes; 8, spermatic channel and field of apical spines." httpUri="https://zenodo.org/record/228454/files/figure.png" pageId="5" pageNumber="576">Fig. 3</figureCitation>
B); mesial border straight, with large subdistal tooth-like mesial process; caudal surface smooth, distal border as rounded lobe fringed with minute spinules (
<figureCitation id="133D2A2B2F2EF175FB57F872B158FA60" box="[1222,1291,1390,1415]" captionStart="FIGURE 3" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[245,1352,194,1892]" captionTargetId="figure@6.[234,1353,193,1903]" captionTargetPageId="6" captionText="FIGURE 3. Eidocamptophallus chacei (Pretzmann, 1967), male, ICN-MHN-CR 2754. A, habitus; B, left first gonopod, caudal view; C, same, cephalic view; D, same, mesial view; E, same, detail of apex, latero-cephalic view; F, left third maxilliped, external view. 1, mesial process; 2, distal lobe of caudal surface; 3, marginal process; 4, marginal tooth; 5, lateral lobe; 6, cephalic lobe; 7, cephalic processes; 8, spermatic channel and field of apical spines." httpUri="https://zenodo.org/record/228454/files/figure.png" pageId="5" pageNumber="576">Fig. 3</figureCitation>
B); marginal suture nearly straight, forming rounded marginal process distally, fringed with spinules, followed by acute marginal tooth projected cephalically; proximal portion of marginal suture forming rounded lobe with conspicuous long setae (
<figureCitation id="133D2A2B2F2EF175FE8DF8C6B531FA14" box="[284,354,1498,1523]" captionStart="FIGURE 3" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[245,1352,194,1892]" captionTargetId="figure@6.[234,1353,193,1903]" captionTargetPageId="6" captionText="FIGURE 3. Eidocamptophallus chacei (Pretzmann, 1967), male, ICN-MHN-CR 2754. A, habitus; B, left first gonopod, caudal view; C, same, cephalic view; D, same, mesial view; E, same, detail of apex, latero-cephalic view; F, left third maxilliped, external view. 1, mesial process; 2, distal lobe of caudal surface; 3, marginal process; 4, marginal tooth; 5, lateral lobe; 6, cephalic lobe; 7, cephalic processes; 8, spermatic channel and field of apical spines." httpUri="https://zenodo.org/record/228454/files/figure.png" pageId="5" pageNumber="576">Fig. 3</figureCitation>
B); lateral border with mid-distal portion straight, strong lobe in mid-proximal portion, nearly rectangular in caudal view, external margin covered with rows of conspicuous setae, which extend to cephalic surface (
<figureCitation id="133D2A2B2F2EF175FF68FB3EB56EF9DC" box="[249,317,1570,1595]" captionStart="FIGURE 3" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[245,1352,194,1892]" captionTargetId="figure@6.[234,1353,193,1903]" captionTargetPageId="6" captionText="FIGURE 3. Eidocamptophallus chacei (Pretzmann, 1967), male, ICN-MHN-CR 2754. A, habitus; B, left first gonopod, caudal view; C, same, cephalic view; D, same, mesial view; E, same, detail of apex, latero-cephalic view; F, left third maxilliped, external view. 1, mesial process; 2, distal lobe of caudal surface; 3, marginal process; 4, marginal tooth; 5, lateral lobe; 6, cephalic lobe; 7, cephalic processes; 8, spermatic channel and field of apical spines." httpUri="https://zenodo.org/record/228454/files/figure.png" pageId="5" pageNumber="576">Fig. 3</figureCitation>
B - D); distal cephalic lobe sinuous, its basal surface with some spinules, cephalic surface with 3 tooth-like processes: mesial tooth the largest and attached to mesial process basally, median tooth slightly smaller, lateral tooth smallest, bulbous; apex narrow, rather compressed between caudal lobe and distal cephalic lobe, slightly directed cephalically; caudal, cephalic lobes forming long slit where funnel shaped spermatic channel and reduced, not discernible field of apical spines are located (
<figureCitation id="133D2A2B2F2EF175FCB2FBAEB736F92C" box="[803,869,1714,1739]" captionStart="FIGURE 3" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[245,1352,194,1892]" captionTargetId="figure@6.[234,1353,193,1903]" captionTargetPageId="6" captionText="FIGURE 3. Eidocamptophallus chacei (Pretzmann, 1967), male, ICN-MHN-CR 2754. A, habitus; B, left first gonopod, caudal view; C, same, cephalic view; D, same, mesial view; E, same, detail of apex, latero-cephalic view; F, left third maxilliped, external view. 1, mesial process; 2, distal lobe of caudal surface; 3, marginal process; 4, marginal tooth; 5, lateral lobe; 6, cephalic lobe; 7, cephalic processes; 8, spermatic channel and field of apical spines." httpUri="https://zenodo.org/record/228454/files/figure.png" pageId="5" pageNumber="576">Fig. 3</figureCitation>
C, E).
</paragraph>
</subSubSection>
<subSubSection id="C31C65252F2EF177FF56FBCAB69EF9EF" lastPageId="7" lastPageNumber="578" pageId="5" pageNumber="576" type="discussion">
<paragraph id="8BB936AE2F2EF175FF56FBCAB4A3F820" blockId="5.[151,1436,1102,1991]" pageId="5" pageNumber="576">
<emphasis id="B972EABC2F2EF175FF56FBCAB568F908" bold="true" box="[199,315,1750,1775]" pageId="5" pageNumber="576">Remarks.</emphasis>
<bibRefCitation id="EF974B5F2F2EF175FED3FBCBB658F908" author="Pretzmann" box="[322,523,1750,1775]" pageId="5" pageNumber="576" refString="Pretzmann, G. (1967) Neue Sudamerikanische Susswasserkrabben der Gattung Pseudothelphusa. Entomologisches Nachrichtenblatt, Wien, 15, 1 - 15." type="journal article" year="1967">Pretzmann (1967)</bibRefCitation>
described this species as
<taxonomicName id="4C064D2D2F2EF175FCBBFBCBB152F908" box="[810,1281,1750,1775]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="chacei" subGenus="Potamocarcinus">
<emphasis id="B972EABC2F2EF175FCBBFBCBB152F908" box="[810,1281,1750,1775]" italics="true" pageId="5" pageNumber="576">Potamocarcinus (Potamocarcinus) chacei</emphasis>
</taxonomicName>
from a single male, of unknown location, donated to the Natural History Museum, London, by the Royal Haslar Hospital, and designed as the
<typeStatus id="54BD880C2F2EF175FEC4FA03B5D6F8D0" box="[341,389,1823,1847]" pageId="5" pageNumber="576">type</typeStatus>
under the number BM 1855.61. Afterwards,
<bibRefCitation id="EF974B5F2F2EF175FC0AFA03B03BF8D0" author="Pretzmann" box="[923,1128,1823,1847]" pageId="5" pageNumber="576" refString="Pretzmann, G. (1971) Fortschritte in der Klassifizierung der Pseudothelphusidae. Anzeiger der Mathematisch Naturwissenschaftliche der Osterreichischen Akademie der Wissenschaften, 179 (1 - 4), 14 - 24. [(1)]" type="journal article" year="1971">Pretzmann (1971)</bibRefCitation>
rearranged this species as
<emphasis id="B972EABC2F2EF175FF06FA58B620F8BC" box="[151,627,1858,1883]" italics="true" pageId="5" pageNumber="576">
<taxonomicName id="4C064D2D2F2EF175FF06FA58B63CF8BC" box="[151,623,1858,1883]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="576" phylum="Arthropoda" rank="species" species="chacei" subGenus="Megathelphusa">Potamocarcinus (Megathelphusa) chacei</taxonomicName>
,
</emphasis>
and later redescribed the specimen and included some photos of the carapace in dorsal, frontal and ventral views, the third maxilliped, and the male first gonopod in cephalic view (
<bibRefCitation id="EF974B5F2F2EF175FF0FFA97B50EF844" author="Pretzmann" box="[158,349,1931,1955]" pageId="5" pageNumber="576" refString="Pretzmann, G. (1972) Die Pseudothelphusidae (Crustacea Brachyura). Zoologica, 42 (120), 1 - 182. [pt. 1]" type="journal article" year="1972">Pretzmann 1972</bibRefCitation>
). He then informed the provenance of the species as &quot;presumably
<collectingCountry id="F311763E2F2EF175FBC6FA96B14FF844" box="[1111,1308,1930,1955]" name="Belize" pageId="5" pageNumber="576">British Honduras</collectingCountry>
&quot; (currently
<collectingCountry id="F311763E2F2EF175FF06FAB2B4B7F820" box="[151,228,1966,1991]" name="Belize" pageId="5" pageNumber="576">Belize</collectingCountry>
).
</paragraph>
<caption id="DF7966262F2DF176FF06FA9FB7BBF812" httpUri="https://zenodo.org/record/228454/files/figure.png" pageId="6" pageNumber="577" targetBox="[245,1352,194,1892]" targetPageId="6">
<paragraph id="8BB936AE2F2DF176FF06FA9FB7BBF812" blockId="6.[151,1436,1923,2037]" pageId="6" pageNumber="577">
<emphasis id="B972EABC2F2DF176FF06FA9FB55FF87F" bold="true" box="[151,268,1923,1945]" pageId="6" pageNumber="577">FIGURE 3</emphasis>
.
<taxonomicName id="4C064D2D2F2DF176FE89FA9FB6BAF87E" authority="(Pretzmann, 1967)" authorityName="Pretzmann" authorityYear="1967" baseAuthorityName="Pretzmann" baseAuthorityYear="1967" box="[280,745,1923,1945]" class="Malacostraca" family="Pseudothelphusidae" genus="Eidocamptophallus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="577" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2DF176FE89FA9FB676F87E" box="[280,549,1923,1945]" italics="true" pageId="6" pageNumber="577">Eidocamptophallus chacei</emphasis>
(Pretzmann, 1967)
</taxonomicName>
, male, ICN-MHN-CR 2754. A, habitus; B, left first gonopod, caudal view; C, same, cephalic view; D, same, mesial view; E, same, detail of apex, latero-cephalic view; F, left third maxilliped, external view. 1, mesial process; 2, distal lobe of caudal surface; 3, marginal process; 4, marginal tooth; 5, lateral lobe; 6, cephalic lobe; 7, cephalic processes; 8, spermatic channel and field of apical spines.
</paragraph>
</caption>
<paragraph id="8BB936AE2F2CF177FF56FD8BB0DFFE4A" blockId="7.[151,1437,151,1544]" pageId="7" pageNumber="578">
<bibRefCitation id="EF974B5F2F2CF177FF56FD8BB5C0FF57" author="Rodriguez" box="[199,403,151,176]" pageId="7" pageNumber="578" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Familie des Pseudothelphusidae. Vol. 22. Faune Tropicale. Paris, ORSTOM, 223 pp." type="book" year="1982">Rodríguez (1982)</bibRefCitation>
identified the species as
<taxonomicName id="4C064D2D2F2CF177FD57FD85B780FF57" box="[710,979,151,176]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2CF177FD57FD85B780FF57" box="[710,979,151,176]" italics="true" pageId="7" pageNumber="578">Potamocarcinus chacei</emphasis>
</taxonomicName>
and described and illustrated the
<typeStatus id="54BD880C2F2CF177FAFDFD84B1CFFF57" box="[1388,1436,152,176]" pageId="7" pageNumber="578">type</typeStatus>
specimen. Judging from the illustrations, the gonopod of the specimen described herein is very similar to that of the
<typeStatus id="54BD880C2F2CF177FF06FDFCB494FF1F" box="[151,199,224,248]" pageId="7" pageNumber="578">type</typeStatus>
specimen of
<taxonomicName id="4C064D2D2F2CF177FECFFDFDB590FF1F" box="[350,451,223,248]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2CF177FECFFDFDB590FF1F" box="[350,451,223,248]" italics="true" pageId="7" pageNumber="578">P. chacei</emphasis>
</taxonomicName>
, but a few differences can be noticed: (a) the lateral lobe in the
<typeStatus id="54BD880C2F2CF177FB03FDFCB091FF1F" box="[1170,1218,224,248]" pageId="7" pageNumber="578">type</typeStatus>
specimen is a little more prominent and subtriangular in caudal view, whereas it is less prominent and nearly rectangular in the present specimen; (b) the marginal tooth is a little more prominent and rounded in the
<typeStatus id="54BD880C2F2CF177FB9CFC34B013FEA7" box="[1037,1088,296,320]" pageId="7" pageNumber="578">type</typeStatus>
, but rather sharp in the present specimen; (c) both specimens have three subdistal tooth-like cephalic processes, but the median tooth is more separated from the mesial one in the
<typeStatus id="54BD880C2F2CF177FDACFC6CB63EFE6F" box="[573,621,368,392]" pageId="7" pageNumber="578">type</typeStatus>
specimen, closer to each other in the present specimen; in addition, the lateral tooth of the cephalic processes is acute but small and blunt in the present specimen.
</paragraph>
<paragraph id="8BB936AE2F2CF177FF56FCABB6F7FC0A" blockId="7.[151,1437,151,1544]" pageId="7" pageNumber="578">
<bibRefCitation id="EF974B5F2F2CF177FF56FCABB660FE37" author="Rodriguez" box="[199,563,439,464]" pageId="7" pageNumber="578" refString="Rodriguez, G. &amp; Hobbs, H. H. Jr. (1989) A new cavernicolous crab, Zilchia falcata, from Guatemala, with notes on the genera of the Potamocarcinini (Crustacea: Decapoda: Pseudothelphusidae). Bulletin du Museum national d'Histoire naturelle (Paris), serie 4, section A, 1, 183 - 192." type="journal article" year="1989">Rodríguez &amp; Hobbs (1989: 191)</bibRefCitation>
recognized that the affinities of
<taxonomicName id="4C064D2D2F2CF177FC0FFCA5B0F6FE37" box="[926,1189,439,464]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2CF177FC0FFCA5B0F6FE37" box="[926,1189,439,464]" italics="true" pageId="7" pageNumber="578">Potamocarcinus chacei</emphasis>
</taxonomicName>
with Potamocarcinini were not clear and removed it from this tribe, proposing a new genus,
<taxonomicName id="4C064D2D2F2CF177FC5AFCC0B481FDFF" authority="Rodriguez &amp; Hobbs, 1989" authorityName="Rodriguez &amp; Hobbs" authorityYear="1989" class="Malacostraca" family="Pseudothelphusidae" genus="Eidocamptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B972EABC2F2CF177FC5AFCC0B0F6FE12" box="[971,1189,476,501]" italics="true" pageId="7" pageNumber="578">Eidocamptophallus</emphasis>
<bibRefCitation id="EF974B5F2F2CF177FB3FFCC0B481FDFF" author="Rodriguez" pageId="7" pageNumber="578" refString="Rodriguez, G. &amp; Hobbs, H. H. Jr. (1989) A new cavernicolous crab, Zilchia falcata, from Guatemala, with notes on the genera of the Potamocarcinini (Crustacea: Decapoda: Pseudothelphusidae). Bulletin du Museum national d'Histoire naturelle (Paris), serie 4, section A, 1, 183 - 192." type="journal article" year="1989">Rodríguez &amp; Hobbs, 1989</bibRefCitation>
</taxonomicName>
, after considering that this taxon would be closer to
<taxonomicName id="4C064D2D2F2CF177FC8DFCE3B0FFFDFF" authority="Smalley, 1965" authorityName="Smalley" authorityYear="1965" box="[796,1196,511,536]" class="Malacostraca" family="Pseudothelphusidae" genus="Camptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="botti">
<emphasis id="B972EABC2F2CF177FC8DFCE3B052FDFF" box="[796,1025,511,536]" italics="true" pageId="7" pageNumber="578">Camptophallus botti</emphasis>
Smalley, 1965
</taxonomicName>
, as far as the features of their gonopods. A comparison with
<taxonomicName id="4C064D2D2F2CF177FDDFFF39B6F0FDDA" box="[590,675,548,573]" class="Malacostraca" family="Pseudothelphusidae" genus="Camptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="botti">
<emphasis id="B972EABC2F2CF177FDDFFF39B6F0FDDA" box="[590,675,548,573]" italics="true" pageId="7" pageNumber="578">C. botti</emphasis>
</taxonomicName>
showed that the resemblances between their gonopods were not as evident as given by
<bibRefCitation id="EF974B5F2F2CF177FEEBFF5BB6E1FD87" author="Rodriguez" box="[378,690,583,608]" pageId="7" pageNumber="578" refString="Rodriguez, G. &amp; Hobbs, H. H. Jr. (1989) A new cavernicolous crab, Zilchia falcata, from Guatemala, with notes on the genera of the Potamocarcinini (Crustacea: Decapoda: Pseudothelphusidae). Bulletin du Museum national d'Histoire naturelle (Paris), serie 4, section A, 1, 183 - 192." type="journal article" year="1989">Rodríguez &amp; Hobbs (1989)</bibRefCitation>
. The gonopods of
<taxonomicName id="4C064D2D2F2CF177FC15FF55B7BEFD87" box="[900,1005,583,608]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2CF177FC15FF55B7BEFD87" box="[900,1005,583,608]" italics="true" pageId="7" pageNumber="578">E. chacei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C064D2D2F2CF177FBB4FF54B028FD87" box="[1061,1147,583,608]" class="Malacostraca" family="Pseudothelphusidae" genus="Camptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="botti">
<emphasis id="B972EABC2F2CF177FBB4FF54B028FD87" box="[1061,1147,583,608]" italics="true" pageId="7" pageNumber="578">C. botti</emphasis>
</taxonomicName>
differ on (a) the situation of the marginal process: broadly rounded, simple, directed distally in
<taxonomicName id="4C064D2D2F2CF177FC37FF71B05CFD62" box="[934,1039,620,645]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2CF177FC37FF71B05CFD62" box="[934,1039,620,645]" italics="true" pageId="7" pageNumber="578">E. chacei</emphasis>
</taxonomicName>
, rather narrow and produced to the mesial side into a denticulate lobe that almost exceed the length of the mesial process in
<taxonomicName id="4C064D2D2F2CF177FBFEFF8CB090FD4F" box="[1135,1219,655,680]" class="Malacostraca" family="Pseudothelphusidae" genus="Camptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="botti">
<emphasis id="B972EABC2F2CF177FBFEFF8CB090FD4F" box="[1135,1219,655,680]" italics="true" pageId="7" pageNumber="578">C. botti</emphasis>
</taxonomicName>
; (b) mesial process more developed and simple in
<taxonomicName id="4C064D2D2F2CF177FD93FFA9B63DFD2A" box="[514,622,692,717]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2CF177FD93FFA9B63DFD2A" box="[514,622,692,717]" italics="true" pageId="7" pageNumber="578">E. chacei</emphasis>
</taxonomicName>
, shorter and clearly bilobed (in caudal view) in
<taxonomicName id="4C064D2D2F2CF177FB3BFFA9B151FD2A" box="[1194,1282,692,717]" class="Malacostraca" family="Pseudothelphusidae" genus="Camptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="botti">
<emphasis id="B972EABC2F2CF177FB3BFFA9B151FD2A" box="[1194,1282,692,717]" italics="true" pageId="7" pageNumber="578">C. botti</emphasis>
</taxonomicName>
; (c) cephalic process has 3 tooth-like, well developed cephalic processes in
<taxonomicName id="4C064D2D2F2CF177FCC4FFC5B7EEFD17" box="[853,957,727,752]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2CF177FCC4FFC5B7EEFD17" box="[853,957,727,752]" italics="true" pageId="7" pageNumber="578">E. chacei</emphasis>
</taxonomicName>
, only a short, blunt process in
<taxonomicName id="4C064D2D2F2CF177FA89FFC4B13FFD17" box="[1304,1388,727,752]" class="Malacostraca" family="Pseudothelphusidae" genus="Camptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="botti">
<emphasis id="B972EABC2F2CF177FA89FFC4B13FFD17" box="[1304,1388,727,752]" italics="true" pageId="7" pageNumber="578">C. botti</emphasis>
</taxonomicName>
; (d) apex is very narrow and rather compressed between the caudal and distal cephalic lobes, with the apical field of setae not discernible in
<taxonomicName id="4C064D2D2F2CF177FE0FFE3DB655FCDF" box="[414,518,799,824]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2CF177FE0FFE3DB655FCDF" box="[414,518,799,824]" italics="true" pageId="7" pageNumber="578">E. chacei</emphasis>
</taxonomicName>
, wider, oblong, with apical field of setae distinct in
<taxonomicName id="4C064D2D2F2CF177FBDBFE3CB0CEFCDF" box="[1098,1181,799,824]" class="Malacostraca" family="Pseudothelphusidae" genus="Camptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="botti">
<emphasis id="B972EABC2F2CF177FBDBFE3CB0CEFCDF" box="[1098,1181,799,824]" italics="true" pageId="7" pageNumber="578">C. botti</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF974B5F2F2CF177FB3DFE03B1CFFCDF" author="Rodriguez" box="[1196,1436,799,824]" pageId="7" pageNumber="578" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Familie des Pseudothelphusidae. Vol. 22. Faune Tropicale. Paris, ORSTOM, 223 pp." type="book" year="1982">Rodríguez (1982: 91)</bibRefCitation>
considered that
<taxonomicName id="4C064D2D2F2CF177FEC3FE58B5AEFCBA" box="[338,509,836,861]" class="Malacostraca" family="Pseudothelphusidae" genus="Camptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B972EABC2F2CF177FEC3FE58B5AEFCBA" box="[338,509,836,861]" italics="true" pageId="7" pageNumber="578">Camptophallus</emphasis>
</taxonomicName>
might be an intermediate form between
<taxonomicName id="4C064D2D2F2CF177FC76FE58B0D4FCBA" box="[999,1159,836,861]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B972EABC2F2CF177FC76FE58B0D4FCBA" box="[999,1159,836,861]" italics="true" pageId="7" pageNumber="578">Ptychophallus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C064D2D2F2CF177FB59FE58B481FC67" authority="Bott, 1967" authorityName="Bott" authorityYear="1967" class="Malacostraca" family="Pseudothelphusidae" genus="Elsalvadoria" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B972EABC2F2CF177FB59FE58B10AFCBA" box="[1224,1369,836,861]" italics="true" pageId="7" pageNumber="578">Elsalvadoria</emphasis>
Bott, 1967
</taxonomicName>
, all groups that seem to have evolved in Central
<collectingCountry id="F311763E2F2CF177FC8FFE7BB72DFC67" box="[798,894,871,896]" name="United States of America" pageId="7" pageNumber="578">America</collectingCountry>
(
<taxonomicName id="4C064D2D2F2CF177FC03FE7BB070FC67" box="[914,1059,871,896]" class="Malacostraca" family="Pseudothelphusidae" genus="Elsalvadoria" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B972EABC2F2CF177FC03FE7BB070FC67" box="[914,1059,871,896]" italics="true" pageId="7" pageNumber="578">Elsalvadoria</emphasis>
</taxonomicName>
in
<collectingCountry id="F311763E2F2CF177FBDEFE7BB098FC67" box="[1103,1227,871,896]" name="Guatemala" pageId="7" pageNumber="578">Guatemala</collectingCountry>
,
<collectingCountry id="F311763E2F2CF177FB4AFE7BB137FC67" box="[1243,1380,871,896]" name="El Salvador" pageId="7" pageNumber="578">El Salvador</collectingCountry>
, and
<collectingCountry id="F311763E2F2CF177FF06FE90B55BFC42" box="[151,264,908,933]" name="Honduras" pageId="7" pageNumber="578">Honduras</collectingCountry>
;
<taxonomicName id="4C064D2D2F2CF177FE83FE90B5EEFC42" box="[274,445,908,933]" class="Malacostraca" family="Pseudothelphusidae" genus="Camptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B972EABC2F2CF177FE83FE90B5EEFC42" box="[274,445,908,933]" italics="true" pageId="7" pageNumber="578">Camptophallus</emphasis>
</taxonomicName>
in
<collectingCountry id="F311763E2F2CF177FE71FE90B604FC42" box="[480,599,908,933]" name="Nicaragua" pageId="7" pageNumber="578">Nicaragua</collectingCountry>
; and
<taxonomicName id="4C064D2D2F2CF177FD01FE90B763FC42" box="[656,816,908,933]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B972EABC2F2CF177FD01FE90B763FC42" box="[656,816,908,933]" italics="true" pageId="7" pageNumber="578">Ptychophallus</emphasis>
</taxonomicName>
in
<collectingCountry id="F311763E2F2CF177FCC2FE90B79EFC42" box="[851,973,908,933]" name="Costa Rica" pageId="7" pageNumber="578">Costa Rica</collectingCountry>
and
<collectingCountry id="F311763E2F2CF177FB92FE90B032FC43" box="[1027,1121,908,932]" name="Panama" pageId="7" pageNumber="578">Panama</collectingCountry>
). It therefore seems unlikely that, due to their geographic separation and the morphological features mentioned above,
<taxonomicName id="4C064D2D2F2CF177FB2DFEB3B13BFC2F" box="[1212,1384,943,968]" class="Malacostraca" family="Pseudothelphusidae" genus="Camptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B972EABC2F2CF177FB2DFEB3B13BFC2F" box="[1212,1384,943,968]" italics="true" pageId="7" pageNumber="578">Camptophallus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C064D2D2F2CF177FF06FEC8B522FC0A" box="[151,369,980,1005]" class="Malacostraca" family="Pseudothelphusidae" genus="Eidocamptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B972EABC2F2CF177FF06FEC8B522FC0A" box="[151,369,980,1005]" italics="true" pageId="7" pageNumber="578">Eidocamptophallus</emphasis>
</taxonomicName>
are phylogenetically close.
</paragraph>
<paragraph id="8BB936AE2F2CF177FF56FEEBB69EF9EF" blockId="7.[151,1437,151,1544]" pageId="7" pageNumber="578">
The position of
<taxonomicName id="4C064D2D2F2CF177FE11FEEBB6FFFBF7" box="[384,684,1015,1040]" class="Malacostraca" family="Pseudothelphusidae" genus="Eidocamptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2CF177FE11FEEBB6FFFBF7" box="[384,684,1015,1040]" italics="true" pageId="7" pageNumber="578">Eidocamptophallus chacei</emphasis>
</taxonomicName>
among the
<taxonomicName id="4C064D2D2F2CF177FCA6FEEBB046FBF7" box="[823,1045,1015,1040]" class="Malacostraca" family="Pseudothelphusidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="subFamily" subFamily="Pseudothelphusinae">Pseudothelphusinae</taxonomicName>
is somewhat difficult to evaluate.
<bibRefCitation id="EF974B5F2F2CF177FF06F900B59FFBD3" author="Rodriguez" box="[151,460,1052,1077]" pageId="7" pageNumber="578" refString="Rodriguez, G. &amp; Hobbs, H. H. Jr. (1989) A new cavernicolous crab, Zilchia falcata, from Guatemala, with notes on the genera of the Potamocarcinini (Crustacea: Decapoda: Pseudothelphusidae). Bulletin du Museum national d'Histoire naturelle (Paris), serie 4, section A, 1, 183 - 192." type="journal article" year="1989">Rodríguez &amp; Hobbs (1989)</bibRefCitation>
did not clearly indicate in which tribe this species was to be included, but commented that without knowing the whereabouts of the
<typeStatus id="54BD880C2F2CF177FD2EF95CB6BCFBBF" box="[703,751,1088,1112]" pageId="7" pageNumber="578">type</typeStatus>
locality the assessment of its affinities was complicated. Rodríguez (1992: 184) just listed the species among the Pseudothelphusini with no further argument. The set of characters diagnostic of this tribe (see
<bibRefCitation id="EF974B5F2F2CF177FDD6F99BB729FB47" author="Villalobos" box="[583,890,1159,1184]" pageId="7" pageNumber="578" refString="Villalobos, J. L. &amp; Alvarez, F. (2010) Phylogenetic analysis of the Mexican freshwater crabs of the tribe Pseudothelphusini (Decapoda: Brachyura: Pseudothelphusidae). Zoological Journal of the Linnean Society, 160, 457 - 481. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2009.00606. x" type="journal article" year="2010">Villalobos &amp; Álvarez 2010</bibRefCitation>
: 472-473), however, are absent in
<taxonomicName id="4C064D2D2F2CF177FB6CF995B136FB47" box="[1277,1381,1159,1184]" class="Malacostraca" family="Pseudothelphusidae" genus="Eidocamptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2CF177FB6CF995B136FB47" box="[1277,1381,1159,1184]" italics="true" pageId="7" pageNumber="578">E. chacei</emphasis>
</taxonomicName>
, and its distribution is not compatible with the Pseudothelphusini, a group endemic to
<collectingCountry id="F311763E2F2CF177FBD1F9B0B0C6FB22" box="[1088,1173,1196,1221]" name="Mexico" pageId="7" pageNumber="578">Mexico</collectingCountry>
(
<bibRefCitation id="EF974B5F2F2CF177FB37F9B0B487FB0F" author="Villalobos" pageId="7" pageNumber="578" refString="Villalobos, J. L. &amp; Alvarez, F. (2010) Phylogenetic analysis of the Mexican freshwater crabs of the tribe Pseudothelphusini (Decapoda: Brachyura: Pseudothelphusidae). Zoological Journal of the Linnean Society, 160, 457 - 481. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2009.00606. x" type="journal article" year="2010">Villalobos &amp; Álvarez 2010</bibRefCitation>
). One could infer that
<bibRefCitation id="EF974B5F2F2CF177FE5FF9D3B751FB0F" author="Rodriguez" box="[462,770,1231,1256]" pageId="7" pageNumber="578" refString="Rodriguez, G. &amp; Hobbs, H. H. Jr. (1989) A new cavernicolous crab, Zilchia falcata, from Guatemala, with notes on the genera of the Potamocarcinini (Crustacea: Decapoda: Pseudothelphusidae). Bulletin du Museum national d'Histoire naturelle (Paris), serie 4, section A, 1, 183 - 192." type="journal article" year="1989">Rodríguez &amp; Hobbs (1989)</bibRefCitation>
made an implicit suggestion that
<taxonomicName id="4C064D2D2F2CF177FBEDF9CDB0B6FB0F" box="[1148,1253,1231,1256]" class="Malacostraca" family="Pseudothelphusidae" genus="Eidocamptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2CF177FBEDF9CDB0B6FB0F" box="[1148,1253,1231,1256]" italics="true" pageId="7" pageNumber="578">E. chacei</emphasis>
</taxonomicName>
could be placed in Hypolobocerini when they pointed out the similarities with
<taxonomicName id="4C064D2D2F2CF177FCC4F9E9B7FAFAEA" box="[853,937,1268,1293]" class="Malacostraca" family="Pseudothelphusidae" genus="Camptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="botti">
<emphasis id="B972EABC2F2CF177FCC4F9E9B7FAFAEA" box="[853,937,1268,1293]" italics="true" pageId="7" pageNumber="578">C. botti</emphasis>
</taxonomicName>
. Although the features usually associated to Hypolobocerini are not easily discernible in the gonopod of
<taxonomicName id="4C064D2D2F2CF177FCADF805B7F7FAD7" box="[828,932,1303,1328]" class="Malacostraca" family="Pseudothelphusidae" genus="Eidocamptophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="chacei">
<emphasis id="B972EABC2F2CF177FCADF805B7F7FAD7" box="[828,932,1303,1328]" italics="true" pageId="7" pageNumber="578">E. chacei</emphasis>
</taxonomicName>
, we believe that it should be included in this tribe because of its presence within the distributional range of the tribe, and due to some morphological features (the rounded outer border of merus of third maxilliped, gonopod with a lateral lobe, and its apex with an apical field of spines in a distal position). Some features of the gonopod might be evidence of a possible closer relationship between this genus and other genera of Hypolobocerini such as
<taxonomicName id="4C064D2D2F2CF177FC61F8BBB0C3FA27" box="[1008,1168,1447,1472]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="B972EABC2F2CF177FC61F8BBB0C3FA27" box="[1008,1168,1447,1472]" italics="true" pageId="7" pageNumber="578">Ptychophallus</emphasis>
</taxonomicName>
(well developed mesial and lateral processes, and apex directed cephalically), and
<taxonomicName id="4C064D2D2F2CF177FCAFF8D0B049FA02" box="[830,1050,1484,1509]" class="Malacostraca" family="Pseudothelphusidae" genus="Colombiathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B972EABC2F2CF177FCAFF8D0B049FA02" box="[830,1050,1484,1509]" italics="true" pageId="7" pageNumber="578">Colombiathelphusa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24157C72F2CF177FBB2F8D0B0DCFA02" box="[1059,1167,1484,1509]" pageId="7" pageNumber="578" rank="genus">
<emphasis id="B972EABC2F2CF177FBB2F8D0B0DCFA02" bold="true" box="[1059,1167,1484,1509]" pageId="7" pageNumber="578">gen. nov.</emphasis>
</taxonomicNameLabel>
(presence of tooth-like cephalic processes adjacent to the mesial process).
</paragraph>
</subSubSection>
</treatment>
</document>