treatments-xml/data/03/AF/87/03AF87A5D94AFFEA6BC8FE1FFCA4F899.xml
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<mods:title id="1D1A5F5DB50BE0B67F34C72DF4A92770">Revision of the New Zealand gecko genus Hoplodactylus, with the description of a new species</mods:title>
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<mods:affiliation id="1482812CFF895084D6C868A9B9B1DFE6">Otago Palaeogenetics Laboratory, Department of Zoology, University of Otago, Dunedin 9016, New Zealand; walton. kerry @ gmail. com; https: // orcid. org / 0000 - 0001 - 7341 - 5715</mods:affiliation>
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<mods:namePart id="62A899086472E98D9050FEBF1A7B239C">Rawlence, Nicolas J.</mods:namePart>
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<mods:affiliation id="670FACEC3A1C02B4854DACBD87EE674E">Otago Palaeogenetics Laboratory, Department of Zoology, University of Otago, Dunedin 9016, New Zealand; walton. kerry @ gmail. com; https: // orcid. org / 0000 - 0001 - 7341 - 5715</mods:affiliation>
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Genus:
<taxonomicName id="4C064D30D94AFFEA6BA7FE1FFD8EFE33" authority="Fitzinger, 1843" authorityName="Fitzinger" authorityYear="1843" box="[248,615,439,466]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6BA7FE1FFE4FFE30" bold="true" box="[248,422,439,466]" italics="true" pageId="4" pageNumber="271">Hoplodactylus</emphasis>
<bibRefCitation id="EF974B42D94AFFEA6AF1FE1FFD8EFE33" author="Fitzinger, L." box="[430,615,439,465]" pageId="4" pageNumber="271" refId="ref15174" refString="Fitzinger, L. (1843) Systema reptilium, fasciculus primus: Amblyglossae. Braumuller et Seidel, Vindobonae, 106 pp. https: // doi. org / 10.5962 / bhl. title. 4694" type="book" year="1843">Fitzinger, 1843</bibRefCitation>
</taxonomicName>
</emphasis>
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<subSubSection id="C31C6538D94AFFEA6BC8FE54FDEFFDF6" box="[151,518,508,532]" pageId="4" pageNumber="271" type="reference_group">
<treatmentCitation id="0AA710A2D94AFFEA6BC8FE54FDEBFDF6" author="Fitzinger, L." box="[151,514,508,532]" page="100" pageId="4" pageNumber="271" year="1843">
<taxonomicName id="4C064D30D94AFFEA6BC8FE54FDEBFDF6" authority="Fitzinger, 1843: 100" authorityName="Fitzinger" authorityPageNumber="100" authorityYear="1843" box="[151,514,508,532]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6BC8FE54FEC2FDF6" box="[151,299,508,532]" italics="true" pageId="4" pageNumber="271">Hoplodactylus</emphasis>
<bibRefCitation id="EF974B42D94AFFEA6A6EFE54FDEBFDF6" author="Fitzinger, L." box="[305,514,508,532]" pageId="4" pageNumber="271" refId="ref15174" refString="Fitzinger, L. (1843) Systema reptilium, fasciculus primus: Amblyglossae. Braumuller et Seidel, Vindobonae, 106 pp. https: // doi. org / 10.5962 / bhl. title. 4694" type="book" year="1843">Fitzinger, 1843: 100</bibRefCitation>
</taxonomicName>
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.
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<subSubSection id="C31C6538D94AFFEA6954FE54FA75FDF6" box="[523,1436,507,532]" pageId="4" pageNumber="271" type="type_taxon">
<typeStatus id="54BD8811D94AFFEA6954FE54FDD4FDF6" box="[523,573,508,532]" pageId="4" pageNumber="271">Type</typeStatus>
species (by monotypy):
<treatmentCitation id="0AA710A2D94AFFEA6867FE54FA71FDF6" author="Dumeril, A. M. C. &amp; Bibron, G." box="[824,1432,507,532]" page="312" pageId="4" pageNumber="271" year="1836">
<taxonomicName id="4C064D30D94AFFEA6867FE54FA71FDF6" authority="Dumeril &amp; Bibron, 1836: 312" authorityName="Dumeril &amp; Bibron" authorityPageNumber="312" authorityYear="1836" box="[824,1432,507,532]" class="Reptilia" family="Diplodactylidae" genus="Platydactyle" isUncertain="true" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="species" species="duvaucelii">
<emphasis id="B972EAA1D94AFFEA6867FE54FC5EFDF6" box="[824,951,508,532]" italics="true" pageId="4" pageNumber="271">Platydactyle</emphasis>
[sic.]
<emphasis id="B972EAA1D94AFFEA68A8FE54FB88FDF6" box="[1015,1121,508,532]" italics="true" pageId="4" pageNumber="271">duvaucelii</emphasis>
<bibRefCitation id="EF974B42D94AFFEA6F38FE54FA71FDF6" author="Dumeril, A. M. C. &amp; Bibron, G." box="[1127,1432,507,532]" pageId="4" pageNumber="271" refId="ref14713" refString="Dumeril, A. M. C. &amp; Bibron, G. (1836) Erpetologie generale, ou, Histoire naturelle complete des reptiles. Vol. 3. Roret, Paris, 531 pp." type="book" year="1836">Duméril &amp; Bibron, 1836: 312</bibRefCitation>
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;
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</paragraph>
<paragraph id="8BB936B3D94AFFEA6B98FDB3FA75FDB3" blockId="4.[151,1436,507,746]" pageId="4" pageNumber="271">
extant, North Island and some adjacent islands,
<collectingCountry id="F3117623D94AFFEA69F3FDB3FCDEFDD0" box="[684,823,538,562]" name="New Zealand" pageId="4" pageNumber="271">New Zealand</collectingCountry>
.
<subSubSection id="C31C6538D94AFFEA6BC8FD91FA75FDB3" box="[151,1436,569,593]" pageId="4" pageNumber="271" type="reference_group">
<taxonomicName id="4C064D30D94AFFEA6BC8FD91FE39FDB3" authority="Gray, 1845: 160" authorityName="Gray" authorityPageNumber="160" authorityYear="1845" box="[151,464,569,593]" class="Reptilia" family="Diplodactylidae" genus="Pentadactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6BC8FD91FECEFDB3" box="[151,295,569,593]" italics="true" pageId="4" pageNumber="271">Pentadactylus</emphasis>
<bibRefCitation id="EF974B42D94AFFEA6A73FD91FE39FDB3" author="Gray, J. E." box="[300,464,569,593]" pageId="4" pageNumber="271" refId="ref16059" refString="Gray, J. E. (1845) Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of the British Museum, London, 289 pp." type="book" year="1845">Gray, 1845: 160</bibRefCitation>
</taxonomicName>
.
<typeStatus id="54BD8811D94AFFEA6A87FD91FDE3FDB3" box="[472,522,569,593]" pageId="4" pageNumber="271">Type</typeStatus>
species (by monotypy):
<taxonomicName id="4C064D30D94AFFEA685FFD91FAE6FDB3" authority="(Dumeril &amp; Bibron, 1836)" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1836" box="[768,1295,569,593]" class="Reptilia" family="Diplodactylidae" genus="Pentadactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="species" species="duvaucelii">
<emphasis id="B972EAA1D94AFFEA685FFD91FC16FDB3" box="[768,1023,569,593]" italics="true" pageId="4" pageNumber="271">Pentadactylus duvaucelii</emphasis>
(
<bibRefCitation id="EF974B42D94AFFEA6F55FD91FAEEFDB3" author="Dumeril, A. M. C. &amp; Bibron, G." box="[1034,1287,569,593]" pageId="4" pageNumber="271" refId="ref14713" refString="Dumeril, A. M. C. &amp; Bibron, G. (1836) Erpetologie generale, ou, Histoire naturelle complete des reptiles. Vol. 3. Roret, Paris, 531 pp." type="book" year="1836">Duméril &amp; Bibron, 1836</bibRefCitation>
)
</taxonomicName>
; extant, North
</subSubSection>
</paragraph>
<subSubSection id="C31C6538D94AFFEA6B98FDF0FD55FD08" pageId="4" pageNumber="271" type="description">
<paragraph id="8BB936B3D94AFFEA6B98FDF0FD55FD08" blockId="4.[151,1436,507,746]" pageId="4" pageNumber="271">
Island and some adjacent islands,
<collectingCountry id="F3117623D94AFFEA697FFDF0FD42FD92" box="[544,683,600,624]" name="New Zealand" pageId="4" pageNumber="271">New Zealand</collectingCountry>
.
<taxonomicName id="4C064D30D94AFFEA6BC8FDDEFEC8FD6C" box="[151,289,630,654]" class="Reptilia" family="Diplodactylidae" genus="Platydactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6BC8FDDEFEC8FD6C" box="[151,289,630,654]" italics="true" pageId="4" pageNumber="271">Platydactylus</emphasis>
</taxonomicName>
of some authors, in part, see below (not Goldfuss, 1820).
<taxonomicName id="4C064D30D94AFFEA6BC8FD3DFF13FD4F" box="[151,250,661,685]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6BC8FD3DFF13FD4F" box="[151,250,661,685]" italics="true" pageId="4" pageNumber="271">Naultinus</emphasis>
</taxonomicName>
of some authors, in part, see below (not Gray, 1842).
<taxonomicName id="4C064D30D94AFFEA6BC8FD1CFE43FD29" authority="Guibe 1954" authorityName="Guibe" authorityYear="1954" box="[151,426,691,716]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6BC8FD1CFEC5FD2E" box="[151,300,692,716]" italics="true" pageId="4" pageNumber="271">Rhacodactylus</emphasis>
<bibRefCitation id="EF974B42D94AFFEA6A6CFD1CFE43FD29" author="Guibe, J." box="[307,426,691,716]" pageId="4" pageNumber="271" refId="ref16188" refString="Guibe, J. (1954) Catalogue des types de lezards du Museum National D'Historie Naturelle. Colas, Paris, 119 pp." type="book" year="1954">Guibé 1954</bibRefCitation>
</taxonomicName>
(not Bocage, 1873).
<taxonomicName id="4C064D30D94AFFEA6BC8FD7AFE76FD08" authority="Jewell 2008" authorityName="Jewell" authorityYear="2008" box="[151,415,722,746]" class="Reptilia" family="Diplodactylidae" genus="Woodworthia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6BC8FD7AFEF6FD08" box="[151,287,722,746]" italics="true" pageId="4" pageNumber="271">Woodworthia</emphasis>
<bibRefCitation id="EF974B42D94AFFEA6A7AFD7AFE76FD08" author="Jewell, T." box="[293,415,722,746]" pageId="4" pageNumber="271" refId="ref17998" refString="Jewell, T. (2008) A photographic guide to reptiles and amphibians of New Zealand. New Holland Publishers Ltd, Auckland, 143 pp." type="book" year="2008">
Jewell
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in part (not Garman, 1901).
</paragraph>
</subSubSection>
<subSubSection id="C31C6538D94AFFEA6BC8FCB1FF19FC24" pageId="4" pageNumber="271" type="diagnosis">
<paragraph id="8BB936B3D94AFFEA6BC8FCB1FF19FC24" blockId="4.[151,1437,793,1915]" pageId="4" pageNumber="271">
<emphasis id="B972EAA1D94AFFEA6BC8FCB1FEC4FCD1" bold="true" box="[151,301,793,819]" pageId="4" pageNumber="271">Diagnosis.—</emphasis>
<taxonomicName id="4C064D30D94AFFEA6A72FCB1FE39FCD1" authorityName="Fitzinger" authorityYear="1843" box="[301,464,793,819]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6A72FCB1FE39FCD1" box="[301,464,793,819]" italics="true" pageId="4" pageNumber="271">Hoplodactylus</emphasis>
</taxonomicName>
(
<emphasis id="B972EAA1D94AFFEA6ABEFCB2FD86FCD1" box="[481,623,794,819]" italics="true" pageId="4" pageNumber="271">
<taxonomicName id="4C064D30D94AFFEA6ABEFCB2FDF6FCD1" box="[481,543,794,819]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="species" species="sensu">sensu</taxonomicName>
stricto
</emphasis>
) species differ from most other diplodactylid taxa in the combination of relatively large body size (SVL:
<quantity id="4CFE9B56D94AFFEA695EFC95FD61FCBA" box="[513,648,829,856]" metricMagnitude="-1" metricUnit="m" metricValue="1.275" metricValueMax="1.6" metricValueMin="0.95" pageId="4" pageNumber="271" unit="mm" value="127.5" valueMax="160.0" valueMin="95.0">95160 mm</quantity>
) and head dimensions at maturity; subdigital pads&gt;2/3
<superScript id="7C739BFBD94AFFEA6FB1FC95FAEBFCA9" attach="left" box="[1262,1282,829,843]" fontSize="6" pageId="4" pageNumber="271">rds</superScript>
of toe length; 1421 subdigital lamellae on underside of digit IV of pes; distal-most 1/3
<superScript id="7C739BFBD94AFFEA68AAFCCAFBEAFC92" attach="left" box="[1013,1027,866,880]" fontSize="6" pageId="4" pageNumber="271">rd</superScript>
of subdigital lamellae distinctively chevron-shaped, with angle of curvature of subsequent lamellae increasing proximally; ventral skin speckled and opaque.
</paragraph>
</subSubSection>
<subSubSection id="C31C6538D94AFFEA6B98FC78FCA4F899" pageId="4" pageNumber="271" type="discussion">
<paragraph id="8BB936B3D94AFFEA6B98FC78FCE5F9B5" blockId="4.[151,1437,793,1915]" pageId="4" pageNumber="271">
<emphasis id="B972EAA1D94AFFEA6B98FC78FEBEFC08" bold="true" box="[199,343,976,1002]" pageId="4" pageNumber="271">Remarks.—</emphasis>
<taxonomicName id="4C064D30D94AFFEA6A08FC78FD47FC08" authority="Fitzinger, 1843" authorityName="Fitzinger" authorityYear="1843" box="[343,686,976,1002]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6A08FC78FE15FC08" box="[343,508,976,1002]" italics="true" pageId="4" pageNumber="271">Rhacodactylus</emphasis>
<bibRefCitation id="EF974B42D94AFFEA695CFC78FD47FC08" author="Fitzinger, L." box="[515,686,976,1002]" pageId="4" pageNumber="271" refId="ref15174" refString="Fitzinger, L. (1843) Systema reptilium, fasciculus primus: Amblyglossae. Braumuller et Seidel, Vindobonae, 106 pp. https: // doi. org / 10.5962 / bhl. title. 4694" type="book" year="1843">Fitzinger, 1843</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54BD8811D94AFFEA69E2FC78FD06FC08" box="[701,751,976,1002]" pageId="4" pageNumber="271">type</typeStatus>
species by monotypy:
<taxonomicName id="4C064D30D94AFFEA68ADFC78FA7DFC08" authority="Cuvier, 1829" authorityName="Cuvier" authorityYear="1829" box="[1010,1428,976,1002]" class="Reptilia" family="Diplodactylidae" genus="Ascalabotes" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="species" species="leachianus">
<emphasis id="B972EAA1D94AFFEA68ADFC78FB13FC08" box="[1010,1274,976,1002]" italics="true" pageId="4" pageNumber="271">Ascalabotes leachianus</emphasis>
Cuvier, 1829
</taxonomicName>
) was introduced as an unspecified subdivision of
<taxonomicName id="4C064D30D94AFFEA69EEFC5CFCBDFBEC" authorityName="Fitzinger" authorityYear="1843" box="[689,852,1012,1038]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA69EEFC5CFCBDFBEC" box="[689,852,1012,1038]" italics="true" pageId="4" pageNumber="271">Hoplodactylus</emphasis>
</taxonomicName>
distinct from
<taxonomicName id="4C064D30D94AFFEA68AFFC5CFB7AFBEC" authorityName="Fitzinger" authorityYear="1843" box="[1008,1171,1012,1038]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA68AFFC5CFB7AFBEC" box="[1008,1171,1012,1038]" italics="true" pageId="4" pageNumber="271">Hoplodactylus</emphasis>
</taxonomicName>
(
<emphasis id="B972EAA1D94AFFEA6FFCFC5DFB34FBEC" box="[1187,1245,1013,1038]" italics="true" pageId="4" pageNumber="271">s. str.</emphasis>
)
<bibRefCitation id="EF974B42D94AFFEA6FB3FC5CFA71FBEC" author="Fitzinger, L." box="[1260,1432,1012,1038]" pageId="4" pageNumber="271" refId="ref15174" refString="Fitzinger, L. (1843) Systema reptilium, fasciculus primus: Amblyglossae. Braumuller et Seidel, Vindobonae, 106 pp. https: // doi. org / 10.5962 / bhl. title. 4694" type="book" year="1843">Fitzinger, 1843</bibRefCitation>
. Genus-level taxonomy in
<collectingCountry id="F3117623D94AFFEA6AE8FBB1FDA5FBD1" box="[439,588,1049,1075]" name="New Zealand" pageId="4" pageNumber="271">New Zealand</collectingCountry>
diplodactylids, especially application of
<taxonomicName id="4C064D30D94AFFEA6F4AFBB1FAC6FBD1" box="[1045,1327,1049,1075]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="subSpecies" species="sensu" subSpecies="latu">
<emphasis id="B972EAA1D94AFFEA6F4AFBB1FAC6FBD1" box="[1045,1327,1049,1075]" italics="true" pageId="4" pageNumber="271">Hoplodactylus sensu latu</emphasis>
</taxonomicName>
, has been highly unstable (see
<bibRefCitation id="EF974B42D94AFFEA6ADCFB95FD8FFBB5" author="Fitzinger, L." box="[387,614,1085,1111]" pageId="4" pageNumber="271" refId="ref15174" refString="Fitzinger, L. (1843) Systema reptilium, fasciculus primus: Amblyglossae. Braumuller et Seidel, Vindobonae, 106 pp. https: // doi. org / 10.5962 / bhl. title. 4694" type="book" year="1843">Fitzinger 1843: 100</bibRefCitation>
;
<bibRefCitation id="EF974B42D94AFFEA692CFB95FCC0FBB5" author="Gray, J. E." box="[627,809,1085,1111]" pageId="4" pageNumber="271" refId="ref15992" refString="Gray, J. E. (1843) Descriptions of the reptiles and amphibians hitherto observed in New Zealand. In: Dieffenbach, E. (Ed.), Travels in New Zealand with contributions to the geography, geology, botany, and natural history of that country. Vol. 2. Murray, London, pp. 202 - 206." type="book" year="1843">Gray 1843: 203</bibRefCitation>
;
<bibRefCitation id="EF974B42D94AFFEA6869FB95FBC0FBB5" author="Boulenger, G. A." box="[822,1065,1085,1112]" pageId="4" pageNumber="271" refId="ref13365" refString="Boulenger, G. A. (1885) Catalogue of the lizards in the British Museum (Natural History). Vol. 1. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. Taylor and Francis, London, 527 pp." type="book" year="1885">Boulenger 1885: 171</bibRefCitation>
;
<bibRefCitation id="EF974B42D94AFFEA6F69FB95FAA1FBB5" author="Chrapliwy, P. S. &amp; Smith, H. M. &amp; Grant, C." box="[1078,1352,1085,1111]" pageId="4" pageNumber="271" pagination="5 - 12" refId="ref14090" refString="Chrapliwy, P. S., Smith, H. M. &amp; Grant, C. (1961) The systematic status of the Geckonid lizard genera Gehyra, Peropus, Hoplodactylus and Naultinus. Herpetologica, 17, 5 - 12." type="journal article" year="1961">
Chrapliwy
<emphasis id="B972EAA1D94AFFEA6FEAFB96FB06FBB5" box="[1205,1263,1085,1111]" italics="true" pageId="4" pageNumber="271">et al.</emphasis>
1961:6
</bibRefCitation>
). Revision by
<bibRefCitation id="EF974B42D94AFFEA6BAFFBCAFE3FFB9E" author="Nielsen, S. V &amp; Bauer, A. M. &amp; Jackman, T. R. &amp; Hitchmough, R. A. &amp; Daugherty, C. H." box="[240,470,1122,1148]" pageId="4" pageNumber="271" pagination="1 - 22" refId="ref20616" refString="Nielsen, S. V, Bauer, A. M., Jackman, T. R., Hitchmough, R. A. &amp; Daugherty, C. H. (2011) New Zealand geckos (Diplodactylidae): Cryptic diversity in a post-Gondwanan lineage with trans-Tasman affinities. Molecular Phylogenetics and Evolution, 59, 1 - 22. https: // doi. org / 10.1016 / j. ympev. 2010.12.007" type="journal article" year="2011">
Nielsen
<emphasis id="B972EAA1D94AFFEA6A12FBCBFE6FFB9E" box="[333,390,1122,1148]" italics="true" pageId="4" pageNumber="271">et al.</emphasis>
(2011)
</bibRefCitation>
considered
<taxonomicName id="4C064D30D94AFFEA6901FBCAFCE8FB9E" authorityName="Fitzinger" authorityYear="1843" box="[606,769,1122,1148]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6901FBCAFCE8FB9E" box="[606,769,1122,1148]" italics="true" pageId="4" pageNumber="271">Hoplodactylus</emphasis>
</taxonomicName>
to form the sister clade to
<taxonomicName id="4C064D30D94AFFEA6F70FBCAFA84FB9E" authority="Garman, 1901" authorityName="Garman" authorityYear="1901" box="[1071,1389,1122,1148]" class="Reptilia" family="Diplodactylidae" genus="Woodworthia" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6F70FBCAFB2CFB9E" box="[1071,1221,1122,1148]" italics="true" pageId="4" pageNumber="271">Woodworthia</emphasis>
Garman, 1901
</taxonomicName>
and comprise two species:
<taxonomicName id="4C064D30D94AFFEA6ACDFB2FFD93FB42" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1836" box="[402,634,1158,1184]" class="Reptilia" family="Diplodactylidae" genus="Pentadactylus" isSensuStricto="true" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="species" species="duvaucelii">
<emphasis id="B972EAA1D94AFFEA6ACDFB2FFE47FB42" box="[402,430,1159,1184]" italics="true" pageId="4" pageNumber="271">H.</emphasis>
(
<emphasis id="B972EAA1D94AFFEA6AE3FB2FFE1FFB42" box="[444,502,1159,1184]" italics="true" pageId="4" pageNumber="271">s. str.</emphasis>
)
<emphasis id="B972EAA1D94AFFEA695AFB2EFD93FB42" box="[517,634,1158,1184]" italics="true" pageId="4" pageNumber="271">duvaucelii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C064D30D94AFFEA69F2FB2FFB29FB42" authority="Bauer &amp; Russell, 1986" authorityName="Bauer &amp; Russell" authorityYear="1986" box="[685,1216,1158,1185]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" isSensuLato="true" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="species" species="delcourti">
<emphasis id="B972EAA1D94AFFEA69F2FB2FFD20FB42" box="[685,713,1159,1184]" italics="true" pageId="4" pageNumber="271">H.</emphasis>
(
<emphasis id="B972EAA1D94AFFEA6987FB2FFCA1FB42" box="[728,840,1158,1184]" italics="true" pageId="4" pageNumber="271">sensu lato</emphasis>
)
<emphasis id="B972EAA1D94AFFEA6808FB2EFC55FB42" box="[855,956,1158,1184]" italics="true" pageId="4" pageNumber="271">delcourti</emphasis>
<bibRefCitation id="EF974B42D94AFFEA689DFB2FFB29FB42" author="Bauer, A. M. &amp; Russell, A. P." box="[962,1216,1158,1185]" pageId="4" pageNumber="271" pagination="26 - 30" refId="ref12831" refString="Bauer, A. M. &amp; Russell, A. P. (1986) The giant gecko Hoplodactylus delcourti and its relations to gigantism and insular endemism in the Gekkonidae. Bulletin of Chicago Herpetological Society, 26, 26 - 30." type="journal article" year="1986">Bauer &amp; Russell, 1986</bibRefCitation>
</taxonomicName>
. The relationship of
<taxonomicName id="4C064D30D94AFFEA6BC8FB04FDCAFB27" authority="Bauer &amp; Russell, 1986" authorityName="Bauer &amp; Russell" authorityYear="1986" box="[151,547,1195,1221]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="species" species="delcourti">
<emphasis id="B972EAA1D94AFFEA6BC8FB04FEF7FB27" box="[151,286,1195,1221]" italics="true" pageId="4" pageNumber="271">H. delcourti</emphasis>
<bibRefCitation id="EF974B42D94AFFEA6A7AFB03FDCAFB27" author="Bauer, A. M. &amp; Russell, A. P." box="[293,547,1195,1221]" pageId="4" pageNumber="271" pagination="26 - 30" refId="ref12831" refString="Bauer, A. M. &amp; Russell, A. P. (1986) The giant gecko Hoplodactylus delcourti and its relations to gigantism and insular endemism in the Gekkonidae. Bulletin of Chicago Herpetological Society, 26, 26 - 30." type="journal article" year="1986">Bauer &amp; Russell, 1986</bibRefCitation>
</taxonomicName>
to
<taxonomicName id="4C064D30D94AFFEA6918FB03FD03FB27" authorityName="Fitzinger" authorityYear="1843" box="[583,746,1195,1221]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6918FB03FD03FB27" box="[583,746,1195,1221]" italics="true" pageId="4" pageNumber="271">Hoplodactylus</emphasis>
</taxonomicName>
(
<emphasis id="B972EAA1D94AFFEA69A5FB04FCDCFB27" box="[762,821,1196,1221]" italics="true" pageId="4" pageNumber="271">s. str.</emphasis>
) species is unclear; and is presently being revised (Heinicke et al. in review).
<taxonomicName id="4C064D30D94AFFEA6AFEFB78FDC1FB0B" authorityName="Bauer &amp; Russell" authorityYear="1986" box="[417,552,1231,1257]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="species" species="delcourti">
<emphasis id="B972EAA1D94AFFEA6AFEFB78FDC1FB0B" box="[417,552,1231,1257]" italics="true" pageId="4" pageNumber="271">H. delcourti</emphasis>
</taxonomicName>
is by far the largest (SVL:
<quantity id="4CFE9B56D94AFFEA6805FB67FC50FB08" box="[858,953,1231,1258]" metricMagnitude="-1" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="271" unit="mm" value="370.0">370 mm</quantity>
, n = 1) of all diplodactylid taxa and further differs from
<taxonomicName id="4C064D30D94AFFEA6A7BFB5CFE2EFAEC" authorityName="Fitzinger" authorityYear="1843" box="[292,455,1268,1294]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6A7BFB5CFE2EFAEC" box="[292,455,1268,1294]" italics="true" pageId="4" pageNumber="271">Hoplodactylus</emphasis>
</taxonomicName>
(
<emphasis id="B972EAA1D94AFFEA6A88FB5DFDFBFAEC" box="[471,530,1269,1294]" italics="true" pageId="4" pageNumber="271">s. str.</emphasis>
) species in having higher subdigital lamellae counts on both the right pes: 12 (I), 20 (II), 20 (III), 23 (IV), 15 (V); and manus: 13 (I), 21 (II), 20 (III), 25 (IV), 11 (V), 35 additional rows of preanal pores (totaling 812, arranged in a subtriangular patch) and dorsal patterning of longitudinal stripes (
<bibRefCitation id="EF974B42D94AFFEA6E7AFA95FEC5FA99" author="Bauer, A. M. &amp; Russell, A. P." pageId="4" pageNumber="271" pagination="26 - 30" refId="ref12831" refString="Bauer, A. M. &amp; Russell, A. P. (1986) The giant gecko Hoplodactylus delcourti and its relations to gigantism and insular endemism in the Gekkonidae. Bulletin of Chicago Herpetological Society, 26, 26 - 30." type="journal article" year="1986">Bauer and Russell 1986</bibRefCitation>
).
<taxonomicName id="4C064D30D94AFFEA6A60FACAFE2CFA99" authorityName="Bauer &amp; Russell" authorityYear="1986" box="[319,453,1377,1403]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="species" species="delcourti">
<emphasis id="B972EAA1D94AFFEA6A60FACAFE2CFA99" box="[319,453,1377,1403]" italics="true" pageId="4" pageNumber="271">H. delcourti</emphasis>
</taxonomicName>
is presumed to be extinct. It is known from a single poorly preserved specimen (MNHN 1985-38) lacking locality information. A
<collectingCountry id="F3117623D94AFFEA693CFA2FFD10FA42" box="[611,761,1414,1441]" name="New Zealand" pageId="4" pageNumber="271">New Zealand</collectingCountry>
provenance was partially inferred (
<bibRefCitation id="EF974B42D94AFFEA6FD9FA2EFA7DFA42" author="Bauer, A. M. &amp; Russell, A. P." box="[1158,1428,1414,1440]" pageId="4" pageNumber="271" pagination="26 - 30" refId="ref12831" refString="Bauer, A. M. &amp; Russell, A. P. (1986) The giant gecko Hoplodactylus delcourti and its relations to gigantism and insular endemism in the Gekkonidae. Bulletin of Chicago Herpetological Society, 26, 26 - 30." type="journal article" year="1986">Bauer and Russell 1986</bibRefCitation>
) due to historical anecdotes of kawekaweau (very large lizards; Mair 1872). More recently, it has been posited that
<taxonomicName id="4C064D30D94AFFEA6BC8FA78FEC8FA0B" authorityName="Bauer &amp; Russell" authorityYear="1986" box="[151,289,1487,1513]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="species" species="delcourti">
<emphasis id="B972EAA1D94AFFEA6BC8FA78FEC8FA0B" box="[151,289,1487,1513]" italics="true" pageId="4" pageNumber="271">H. delcourti</emphasis>
</taxonomicName>
is unlikely to originate from
<collectingCountry id="F3117623D94AFFEA6924FA78FCFFFA0B" box="[635,790,1487,1514]" name="New Zealand" pageId="4" pageNumber="271">New Zealand</collectingCountry>
, with a New Caledonian origin inferred (
<bibRefCitation id="EF974B42D94AFFEA6FAAFA67FA78FA0B" author="Worthy, T. H." box="[1269,1425,1487,1513]" pageId="4" pageNumber="271" pagination="65 - 86" refId="ref25285" refString="Worthy, T. H. (2016) A review of the fossil record of New Zealand lizards. In: Chapple, D. G. (Ed.), New Zealand Lizards. Springer, Cham, pp. 65 - 86. https: // doi. org / 10.1007 / 978 - 3 - 319 - 41674 - 8 _ 3" type="book chapter" year="2016">Worthy 2016</bibRefCitation>
). However, its distribution remains unclear, but likely derives from the southwest Pacific. We consider
<taxonomicName id="4C064D30D94AFFEA6E4BFA5CFA75F9EC" authorityName="Bauer &amp; Russell" authorityYear="1986" box="[1300,1436,1524,1550]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="species" species="delcourti">
<emphasis id="B972EAA1D94AFFEA6E4BFA5CFA75F9EC" box="[1300,1436,1524,1550]" italics="true" pageId="4" pageNumber="271">H. delcourti</emphasis>
</taxonomicName>
is highly unlikely to be congeneric with
<taxonomicName id="4C064D30D94AFFEA6908F9B1FD07F9D0" baseAuthorityName="Dumeril &amp; Bibron" baseAuthorityYear="1836" box="[599,750,1560,1586]" class="Reptilia" family="Diplodactylidae" genus="Pentadactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="species" species="duvaucelii">
<emphasis id="B972EAA1D94AFFEA6908F9B1FD07F9D0" box="[599,750,1560,1586]" italics="true" pageId="4" pageNumber="271">H. duvaucelii</emphasis>
</taxonomicName>
. However, we leave this taxon in
<taxonomicName id="4C064D30D94AFFEA6F39F9B0FA69F9D0" box="[1126,1408,1560,1586]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="subSpecies" species="sensu" subSpecies="latu">
<emphasis id="B972EAA1D94AFFEA6F39F9B0FA69F9D0" box="[1126,1408,1560,1586]" italics="true" pageId="4" pageNumber="271">Hoplodactylus sensu latu</emphasis>
</taxonomicName>
in the interest of taxonomic stability pending further work.
</paragraph>
<paragraph id="8BB936B3D94AFFEA6B98F9C9FCA4F899" blockId="4.[151,1437,793,1915]" pageId="4" pageNumber="271">
<taxonomicName id="4C064D30D94AFFEA6B98F9C9FE83F999" authorityName="Fitzinger" authorityYear="1843" box="[199,362,1633,1659]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6B98F9C9FE83F999" box="[199,362,1633,1659]" italics="true" pageId="4" pageNumber="271">Hoplodactylus</emphasis>
</taxonomicName>
(
<emphasis id="B972EAA1D94AFFEA6A22F9CAFE42F999" box="[381,427,1634,1659]" italics="true" pageId="4" pageNumber="271">s. s.</emphasis>
) species presently inhabit lowland coastal forest and scrubland including flax, mānuka, kawakawa and pōhutukawa (
<bibRefCitation id="EF974B42D94AFFEA6A89F92EFDB9F943" author="Hard, G. C." box="[470,592,1670,1697]" pageId="4" pageNumber="271" pagination="143 - 146" refId="ref16345" refString="Hard, G. C. (1954) Notes on reptilia of islands visited by Field Club - 1953 - 1954. Tane, 6, 143 - 146." type="journal article" year="1954">Hard 1954</bibRefCitation>
;
<bibRefCitation id="EF974B42D94AFFEA6905F92EFD0DF942" author="Towns, D. R." box="[602,740,1670,1696]" pageId="4" pageNumber="271" pagination="91 - 96" refId="ref23124" refString="Towns, D. R. (1971) The lizards of the islands visited by Field Club 1953 - 1954. A revision with some additions 1969 - 1970. Tane, 17, 91 - 96." type="journal article" year="1971">Towns 1971</bibRefCitation>
;
<bibRefCitation id="EF974B42D94AFFEA69B1F92EFC7DF942" author="Whitaker, A. H." box="[750,916,1670,1696]" pageId="4" pageNumber="271" pagination="623 - 651" refId="ref24229" refString="Whitaker, A. H. (1968) The lizards of the Poor Knights Islands, New Zealand. New Zealand Journal of Science, 11, 623 - 651." type="journal article" year="1968">Whitaker 1968</bibRefCitation>
), taking refuge in tree hollows, beneath ground cover, and within deep crevices and burrows (such as of tuatara and petrels on rocky cliffs;
<bibRefCitation id="EF974B42D94AFFEA6FFFF902FAAAF927" author="McCann, C." box="[1184,1347,1706,1733]" pageId="4" pageNumber="271" pagination="1 - 127" refId="ref19359" refString="McCann, C. (1955) The lizards of New Zealand. Dominion Museum Bulletin, 17, 1 - 127." type="journal article" year="1955">McCann 1955</bibRefCitation>
;
<bibRefCitation id="EF974B42D94AFFEA6E11F902FF14F90B" author="Whitaker, A. H." pageId="4" pageNumber="271" pagination="623 - 651" refId="ref24229" refString="Whitaker, A. H. (1968) The lizards of the Poor Knights Islands, New Zealand. New Zealand Journal of Science, 11, 623 - 651." type="journal article" year="1968">Whitaker 1968</bibRefCitation>
;
<bibRefCitation id="EF974B42D94AFFEA6A56F967FDE5F90B" author="van Winkel, D. &amp; Baling, M. &amp; Hitchmough, R." box="[265,524,1743,1769]" pageId="4" pageNumber="271" refId="ref24596" refString="van Winkel, D., Baling, M. &amp; Hitchmough, R. (2018) Reptiles and amphibians of New Zealand. Auckland University Press, Auckland, 712 pp." type="book" year="2018">
van Winkel
<emphasis id="B972EAA1D94AFFEA6ACFF978FE21F90B" box="[400,456,1743,1769]" italics="true" pageId="4" pageNumber="271">et al.</emphasis>
2018
</bibRefCitation>
). Conversely, extinct mainland
<taxonomicName id="4C064D30D94AFFEA6833F967FBE6F90B" authorityName="Fitzinger" authorityYear="1843" box="[876,1039,1743,1769]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA6833F967FBE6F90B" box="[876,1039,1743,1769]" italics="true" pageId="4" pageNumber="271">Hoplodactylus</emphasis>
</taxonomicName>
populations likely inhabited mixed podocarp-broadleaf forests prior to widespread anthropogenic landscape modification (Worthy 1987;
<bibRefCitation id="EF974B42D94AFFEA6E4DF95BFF10F8D0" author="McWethy, D. B. &amp; Wilmshurst, J. M. &amp; Whitlock, C. &amp; Wood, J. R. &amp; McGlone, M. S." pageId="4" pageNumber="271" pagination="111328" refId="ref19478" refString="McWethy, D. B., Wilmshurst, J. M., Whitlock, C., Wood, J. R. &amp; McGlone, M. S. (2014). A high-resolution chronology of rapid forest transitions following Polynesian arrival in New Zealand. PLoS ONE, 9 (11), e 111328. https: // doi. org / 10.1371 / journal. pone. 0111328" type="journal article" year="2014">
McWethy
<emphasis id="B972EAA1D94AFFEA6ED7F95CFF5DF8D0" italics="true" pageId="4" pageNumber="271">et al.</emphasis>
2014
</bibRefCitation>
).
<taxonomicName id="4C064D30D94AFFEA6A53F8B0FDE1F8D0" box="[268,520,1816,1842]" pageId="4" pageNumber="271">
<emphasis id="B972EAA1D94AFFEA6A53F8B0FE46F8D0" box="[268,431,1816,1842]" italics="true" pageId="4" pageNumber="271">Hoplodactylus</emphasis>
species
</taxonomicName>
are extremely long-lived, with wild and captive-reared individuals recorded over 50 years old (Cree &amp; Hare 2016).
<taxonomicName id="4C064D30D94AFFEA694FF894FD5AF8B4" authorityName="Fitzinger" authorityYear="1843" box="[528,691,1852,1878]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="271" phylum="Chordata" rank="genus">
<emphasis id="B972EAA1D94AFFEA694FF894FD5AF8B4" box="[528,691,1852,1878]" italics="true" pageId="4" pageNumber="271">Hoplodactylus</emphasis>
</taxonomicName>
is the only
<collectingCountry id="F3117623D94AFFEA686CF895FC21F8B4" box="[819,968,1852,1879]" name="New Zealand" pageId="4" pageNumber="271">New Zealand</collectingCountry>
diplodactylid genus in which saurophagy, including cannibalism, has been documented (
<bibRefCitation id="EF974B42D94AFFEA69C2F8C9FCA8F899" author="Barwick, R. E." box="[669,833,1889,1915]" pageId="4" pageNumber="271" pagination="377 - 390" refId="ref12563" refString="Barwick, R. E. (1982) The growth and ecology of the gecko Hoplodactylus duvaucelii at The Brothers islands. In: Newman, D. G. (Ed.), New Zealand Herpetology. New Zealand Wildlife Service Occasional Publication, Wellington, pp. 377 - 390." type="book chapter" year="1982">Barwick 1982</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>