1162 lines
167 KiB
XML
1162 lines
167 KiB
XML
<document id="7EA2DEE9A7FE02DF1DF1D96A9063F2CB" ID-CLB-Dataset="226117" ID-DOI="10.11646/zootaxa.5228.3.3" ID-GBIF-Dataset="98820bb9-0993-425a-8698-910ed6d4c5c6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7532705" ID-ZooBank="42773A43-8AC3-43AF-A110-DFA5E7EC43B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1673572147550" checkinUser="plazi" docAuthor="Scarsbrook, Lachie, Walton, Kerry, Rawlence, Nicolas J. & Hitchmough, Rodney A." docDate="2023" docId="03AF87A5D949FFE26BC8FE8EFED9FB0B" docLanguage="en" docName="zootaxa.5228.3.3.pdf" docOrigin="Zootaxa 5228 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hoplodactylus tohu Scarsbrook & Walton & Rawlence & Hitchmough 2023, n. sp." docType="treatment" docUuid="1BA7530E-52D7-44F7-9FC1-61C9A4A4B7AA" docUuidSource="ZooBank" docVersion="7" lastPageNumber="279" masterDocId="FF96FFDDD94EFFEE6B5FFFA8FFE9FFE2" masterDocTitle="Revision of the New Zealand gecko genus Hoplodactylus, with the description of a new species" masterLastPageNumber="291" masterPageNumber="267" pageNumber="274" updateTime="1699473923421" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="22A0E762D68CDD9CBE9D61C1859D8574">Revision of the New Zealand gecko genus Hoplodactylus, with the description of a new species</mods:title>
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<mods:namePart id="CF5549EF257E37853A595EA6047519A9">Scarsbrook, Lachie</mods:namePart>
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<mods:affiliation id="B20EDDF33E8502C158785EFA5AFFC8CE">Otago Palaeogenetics Laboratory, Department of Zoology, University of Otago, Dunedin 9016, New Zealand; walton. kerry @ gmail. com; https: // orcid. org / 0000 - 0001 - 7341 - 5715</mods:affiliation>
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<mods:nameIdentifier id="DD5321BF3F7FB423BB91E0E36D49F317" type="email">walton.kerry@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5A6FC9720AF910A2587819E613F8D610">Walton, Kerry</mods:namePart>
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<mods:affiliation id="ED04B489B0C2524B1A86875391B1DC8D">Otago Palaeogenetics Laboratory, Department of Zoology, University of Otago, Dunedin 9016, New Zealand; walton. kerry @ gmail. com; https: // orcid. org / 0000 - 0001 - 7341 - 5715</mods:affiliation>
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<mods:namePart id="5825FBA35DF1EFEBAB18003B77D61AE8">Rawlence, Nicolas J.</mods:namePart>
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<mods:affiliation id="1AC0DE92E374F86751838CE97F575053">Otago Palaeogenetics Laboratory, Department of Zoology, University of Otago, Dunedin 9016, New Zealand; walton. kerry @ gmail. com; https: // orcid. org / 0000 - 0001 - 7341 - 5715</mods:affiliation>
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<mods:namePart id="0B2F8B8EF94381D9634BBCA839253B5F">Hitchmough, Rodney A.</mods:namePart>
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<mods:affiliation id="853C5F15A549A0301AD21A38271FC065">Department of Conservation, Wellington 6011, New Zealand.</mods:affiliation>
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<mods:title id="8725DED4F27C559E5A7E3A75B3CD0CCC">Zootaxa</mods:title>
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<mods:date id="C1395E9F3A16A24F0294B1CD1F44966B">2023</mods:date>
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<treatment id="03AF87A5D949FFE26BC8FE8EFED9FB0B" ID-DOI="http://doi.org/10.5281/zenodo.7542115" ID-GBIF-Taxon="208978147" ID-Zenodo-Dep="7542115" LSID="urn:lsid:zoobank.org:act:1BA7530E-52D7-44F7-9FC1-61C9A4A4B7AA" httpUri="http://treatment.plazi.org/id/03AF87A5D949FFE26BC8FE8EFED9FB0B" lastPageId="12" lastPageNumber="279" pageId="7" pageNumber="274">
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<taxonomicName id="4C064D30D949FFE96BC8FE8EFE6DFEA3" authority="Scarsbrook & Walton & Rawlence & Hitchmough, 2023" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[151,388,294,321]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="tohu" status="sp. nov.">
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<emphasis id="B972EAA1D949FFE96BC8FE8EFE6DFEA3" bold="true" box="[151,388,294,321]" italics="true" pageId="7" pageNumber="274">Hoplodactylus tohu</emphasis>
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<taxonomicNameLabel id="A24157DAD949FFE96AD3FE8FFE27FEA3" box="[396,462,295,321]" pageId="7" pageNumber="274" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C31C6538D949FFE96BC8FEE3FA66FE45" pageId="7" pageNumber="274" type="description">
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<paragraph id="8BB936B3D949FFE96BC8FEE3FCF5FE87" blockId="7.[151,796,294,357]" box="[151,796,330,357]" pageId="7" pageNumber="274">
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<figureCitation id="133D2A36D949FFE96BC8FEE3FEFEFE87" box="[151,279,331,357]" captionStart="FIGURE 2" captionStartId="8.[151,250,1877,1901]" captionTargetBox="[163,1423,191,1843]" captionTargetId="figure-26@8.[153,1434,181,1853]" captionTargetPageId="8" captionText="FIGURE 2. Hoplodactylus (Fitzinger, 1843) sp.—dorsal and ventral body views (left-right). A, Hoplodactylus tohu n. sp., holotype, Middle Trios Island, Marlborough Sounds, RE.000503. B, Hoplodactylus duvaucelii Duméril & Bibron, 1836, topotype, Muriwhenua Island, Hen and Chickens Islands, RE.003491. Automontage images generated by Jean-Claude Stahl (NMNZ). Scale bar = 20 mm." figureDoi="http://doi.org/10.5281/zenodo.7532711" httpUri="https://zenodo.org/record/7532711/files/figure.png" pageId="7" pageNumber="274">Figures 2A</figureCitation>
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,
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<figureCitation id="133D2A36D949FFE96A7DFEE3FE80FE87" box="[290,361,331,357]" captionStart="FIGURE 3" captionStartId="9.[151,250,815,839]" captionTargetBox="[151,1435,185,785]" captionTargetId="figure-20@9.[151,1435,181,791]" captionTargetPageId="9" captionText="FIGURE 3. Details of: A-D, Hoplodactylus tohu n. sp., holotype (RE.000503); and E-H, Hoplodactylus duvaucelii Duméril & Bibron, 1836, topotype (RE.003491): A, E, right-lateral view of head; B, F, oblique right-lateroventral view of head, with diagnostic characters (a) pronounced brillar fold, and (b) abrupt size decrease at the 5th infralabial scale; C, G, ventral view of the right manus showing subdigital lamellae, and highlighting the less emergent claw on the first digit in H. duvaucelii; D, H, ventral view of the right pes showing subdigital lamellae. Automontage images generated by Jean-Claude Stahl (NMNZ). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7532713" httpUri="https://zenodo.org/record/7532713/files/figure.png" pageId="7" pageNumber="274">3A–D</figureCitation>
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,
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<figureCitation id="133D2A36D949FFE96A2AFEE3FE52FE87" box="[373,443,331,357]" captionStart="FIGURE 4" captionStartId="10.[151,250,1519,1543]" captionTargetBox="[151,1435,181,1496]" captionTargetId="figure-26@10.[151,1435,181,1496]" captionTargetPageId="10" captionText="FIGURE 4. Living individuals of Hoplodactylus species. A–C, Hoplodactylus tohu n. sp.; D–F, H. duvaucelii (Duméril & Bibron 1836). A, Brothers Islands (Cook Strait); B–C, Sentinel Rock (Marlborough Sounds). D–F, Korapuki Island (Mercury Islands). The first and second rows show sub-adult and adult representatives of each species respectively. Photographs: Nicholas Harker (Auckland, New Zealand)." figureDoi="http://doi.org/10.5281/zenodo.7532715" httpUri="https://zenodo.org/record/7532715/files/figure.png" pageId="7" pageNumber="274">4A–C</figureCitation>
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; Supplementary Figures 5A–C.
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</paragraph>
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ZooBank registration of
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<taxonomicName id="4C064D30D949FFE96ACEFE27FDB1FE45" authority="Scarsbrook & Walton & Rawlence & Hitchmough, 2023" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[401,600,399,423]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="tohu" status="sp. nov.">
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<emphasis id="B972EAA1D949FFE96ACEFE27FDB1FE45" box="[401,600,399,423]" italics="true" pageId="7" pageNumber="274">Hoplodactylus tohu</emphasis>
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<taxonomicNameLabel id="A24157DAD949FFE96901FE38FD71FE45" box="[606,664,400,423]" pageId="7" pageNumber="274" rank="species">n. sp.</taxonomicNameLabel>
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:
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urn:lsid:zoobank.org:pub:
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.
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<paragraph id="8BB936B3D949FFE96BC8FE7AFCFEFE08" blockId="7.[151,1437,466,1165]" box="[151,791,466,490]" pageId="7" pageNumber="274">
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<taxonomicName id="4C064D30D949FFE96BC8FE7AFEB2FE08" box="[151,347,466,490]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="pacificus">
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<emphasis id="B972EAA1D949FFE96BC8FE7AFEB2FE08" box="[151,347,466,490]" italics="true" pageId="7" pageNumber="274">Naultinus pacificus</emphasis>
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</taxonomicName>
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.–
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<treatmentCitation id="0AA710A2D949FFE96A2AFE7AFDFDFE08" author="Gray, J. E." box="[373,532,466,490]" page="203" pageId="7" pageNumber="274" year="1843">
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<bibRefCitation id="EF974B42D949FFE96A2AFE7AFDFDFE08" author="Gray, J. E." box="[373,532,466,490]" pageId="7" pageNumber="274" refId="ref15992" refString="Gray, J. E. (1843) Descriptions of the reptiles and amphibians hitherto observed in New Zealand. In: Dieffenbach, E. (Ed.), Travels in New Zealand with contributions to the geography, geology, botany, and natural history of that country. Vol. 2. Murray, London, pp. 202 - 206." type="book" year="1843">Gray 1843: 203</bibRefCitation>
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(in part, not Gray, 1842).
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</paragraph>
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<paragraph id="8BB936B3D949FFE96BC8FE58FA9DFDA4" blockId="7.[151,1437,466,1165]" pageId="7" pageNumber="274">
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<treatmentCitationGroup id="AB16119DD949FFE96BC8FE58FA9DFDA4" pageId="7" pageNumber="274">
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<taxonomicName id="4C064D30D949FFE96BC8FE58FE74FDEA" baseAuthorityName="Dumeril & Bibron" baseAuthorityYear="1836" box="[151,413,496,520]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="duvaucelii">
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<emphasis id="B972EAA1D949FFE96BC8FE58FE74FDEA" box="[151,413,496,520]" italics="true" pageId="7" pageNumber="274">Hoplodactylus duvaucelii</emphasis>
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</taxonomicName>
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[sic]
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<treatmentCitation id="0AA710A2D949FFE96A84FE59FD7EFDEB" author="McCann, C." box="[475,663,496,521]" page="39" pageId="7" pageNumber="274" year="1955">
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<bibRefCitation id="EF974B42D949FFE96A84FE59FD7EFDEB" author="McCann, C." box="[475,663,496,521]" pageId="7" pageNumber="274" pagination="1 - 127" refId="ref19359" refString="McCann, C. (1955) The lizards of New Zealand. Dominion Museum Bulletin, 17, 1 - 127." type="journal article" year="1955">McCann 1955: 39</bibRefCitation>
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</treatmentCitation>
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, fig. 3, pl. 4;
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<treatmentCitation id="0AA710A2D949FFE96876FE59FC1BFDEA" author="McCann, C." box="[809,1010,496,521]" page="46" pageId="7" pageNumber="274" year="1956">
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<bibRefCitation id="EF974B42D949FFE96876FE59FC1BFDEA" author="McCann, C." box="[809,1010,496,521]" pageId="7" pageNumber="274" pagination="45 - 51" refId="ref19382" refString="McCann, C. (1956 a) Keys to the lizards of New Zealand. Tuatara, 6, 45 - 51." type="journal article" year="1956">McCann 1956a: 46</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE968A0FE59FB20FDEA" author="McCann, C." box="[1023,1225,496,521]" page="15" pageId="7" pageNumber="274" year="1956">
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<bibRefCitation id="EF974B42D949FFE968A0FE59FB20FDEA" author="McCann, C." box="[1023,1225,496,521]" pageId="7" pageNumber="274" pagination="15 - 16" refId="ref19406" refString="McCann, C. (1956 b) The distribution of New Zealand reptiles. Proceedings of the New Zealand Ecological Society, 4, 15 - 16." type="journal article" year="1956">McCann 1956b: 15</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE96F89FE58FA71FDEB" author="Bustard, H." box="[1238,1432,496,521]" page="218" pageId="7" pageNumber="274" year="1963">
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<bibRefCitation id="EF974B42D949FFE96F89FE58FA71FDEB" author="Bustard, H." box="[1238,1432,496,521]" pageId="7" pageNumber="274" pagination="217 - 218" refId="ref13681" refString="Bustard, H. (1963) Gecko behavioural trait: tongue wiping spectacle. Herpetologica, 19, 217 - 218. https: // doi. org / 10.2307 / 1441366" type="journal article" year="1963">Bustard 1963: 218</bibRefCitation>
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</treatmentCitation>
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;
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||
<treatmentCitation id="0AA710A2D949FFE96B98FDA7FE9CFDC5" author="Sharell, R." box="[199,373,527,551]" page="49" pageId="7" pageNumber="274" year="1966">
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<bibRefCitation id="EF974B42D949FFE96B98FDA7FE9CFDC5" author="Sharell, R." box="[199,373,527,551]" pageId="7" pageNumber="274" refId="ref21876" refString="Sharell, R. (1966) The tuatara, lizards and frogs of New Zealand. Collins, London, 94 pp." type="book" year="1966">Sharell 1966: 49</bibRefCitation>
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</treatmentCitation>
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, pls. 28–31;
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<treatmentCitation id="0AA710A2D949FFE96AA3FDA7FD27FDC5" author="Thoresen, A. C." box="[508,718,527,551]" page="197" pageId="7" pageNumber="274" year="1967">
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<bibRefCitation id="EF974B42D949FFE96AA3FDA7FD27FDC5" author="Thoresen, A. C." box="[508,718,527,551]" pageId="7" pageNumber="274" pagination="182 - 200" refId="ref22705" refString="Thoresen, A. C. (1967) Ecological observations on Stanley and Green Islands, Mercury Group. Notornis, 14, 182 - 200." type="journal article" year="1967">Thoresen 1967: 197</bibRefCitation>
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||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96985FDA7FC42FDC5" author="Whitaker, A. H." box="[730,939,527,551]" page="122" pageId="7" pageNumber="274" year="1973">
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||
<bibRefCitation id="EF974B42D949FFE96985FDA7FC42FDC5" author="Whitaker, A. H." box="[730,939,527,551]" pageId="7" pageNumber="274" pagination="121 - 130" refId="ref24261" refString="Whitaker, A. H. (1973) Lizard populations on islands with and without Polynesian rats Rattus exulans. Proceedings of the New Zealand Ecological Society, 20, 121 - 130." type="journal article" year="1973">Whitaker 1973: 122</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE968E7FDB8FB2BFDC5" author="Domrow, R. & Heath, A. C. G. & Kennedy, C." box="[952,1218,527,551]" page="295" pageId="7" pageNumber="274" year="1980">
|
||
<bibRefCitation id="EF974B42D949FFE968E7FDB8FB2BFDC5" author="Domrow, R. & Heath, A. C. G. & Kennedy, C." box="[952,1218,527,551]" pageId="7" pageNumber="274" pagination="291 - 297" refId="ref14623" refString="Domrow, R., Heath, A. C. G. & Kennedy, C. (1980) Two new species of Ophionyssus (Acari: Dermanyssidae) from New Zealand lizards. New Zealand Journal of Zoology, 7, 291 - 297. https: // doi. org / 10.1080 / 03014223.1980.10423790" type="journal article" year="1980">
|
||
Domrow
|
||
<emphasis id="B972EAA1D949FFE96F45FDB8FBB9FDC5" box="[1050,1104,527,551]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
1980: 295
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96F90FDA7FA71FDC5" author="Barwick, R. E." box="[1231,1432,527,551]" page="377" pageId="7" pageNumber="274" year="1982">
|
||
<bibRefCitation id="EF974B42D949FFE96F90FDA7FA71FDC5" author="Barwick, R. E." box="[1231,1432,527,551]" pageId="7" pageNumber="274" pagination="377 - 390" refId="ref12563" refString="Barwick, R. E. (1982) The growth and ecology of the gecko Hoplodactylus duvaucelii at The Brothers islands. In: Newman, D. G. (Ed.), New Zealand Herpetology. New Zealand Wildlife Service Occasional Publication, Wellington, pp. 377 - 390." type="book chapter" year="1982">Barwick 1982: 377</bibRefCitation>
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||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96B98FD86FE8FFDA4" author="Bauer, A. M." box="[199,358,558,582]" page="90" pageId="7" pageNumber="274" year="1985">
|
||
<bibRefCitation id="EF974B42D949FFE96B98FD86FE8FFDA4" author="Bauer, A. M." box="[199,358,558,582]" pageId="7" pageNumber="274" pagination="81 - 94" refId="ref12648" refString="Bauer, A. M. (1985) Notes on the taxonomy, morphology and behaviour of Rhacodactylus chahoua (Bavay) (Reptilia: Gekkonidae). Bonner Zoologische Beitrage, 36, 81 - 94." type="journal article" year="1985">Bauer 1985: 90</bibRefCitation>
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||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96A2EFD86FD75FDA4" author="Halliday, T. R. & Verrell, P. A." box="[369,668,558,582]" page="260" pageId="7" pageNumber="274" year="1988">
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||
<bibRefCitation id="EF974B42D949FFE96A2EFD86FD75FDA4" author="Halliday, T. R. & Verrell, P. A." box="[369,668,558,582]" pageId="7" pageNumber="274" pagination="253 - 265" refId="ref16300" refString="Halliday, T. R. & Verrell, P. A. (1988) Body size and age in amphibians and reptiles. Journal of Herpetology, 22, 253 - 265. https: // doi. org / 10.2307 / 1564148" type="journal article" year="1988">Halliday & Verrell 1988: 260</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE969F9FD86FC52FDA4" author="Wilson, K. J. & Freeman, A." box="[678,955,558,582]" page="1" pageId="7" pageNumber="274" year="1993">
|
||
<bibRefCitation id="EF974B42D949FFE969F9FD86FC52FDA4" author="Wilson, K. J. & Freeman, A." box="[678,955,558,582]" pageId="7" pageNumber="274" refId="ref24423" refString="Wilson, K. J. & Freeman, A. (1993) A preliminary investigation of the impact of burrow-breeding seabirds on vegetation, insects and reptiles. Lincoln University, Christchurch, 30 pp." type="book" year="1993">Wilson & Freeman 1993: 1</bibRefCitation>
|
||
</treatmentCitation>
|
||
– all in part (not
|
||
<bibRefCitation id="EF974B42D949FFE96F35FD86FA80FDA4" author="Dumeril, A. M. C. & Bibron, G." box="[1130,1385,558,582]" pageId="7" pageNumber="274" refId="ref14713" refString="Dumeril, A. M. C. & Bibron, G. (1836) Erpetologie generale, ou, Histoire naturelle complete des reptiles. Vol. 3. Roret, Paris, 531 pp." type="book" year="1836">Duméril & Bibron, 1836</bibRefCitation>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB936B3D949FFE96BC8FDE4FC28FBAD" blockId="7.[151,1437,466,1165]" pageId="7" pageNumber="274">
|
||
<treatmentCitationGroup id="AB16119DD949FFE96BC8FDE4FC28FBAD" pageId="7" pageNumber="274">
|
||
<taxonomicName id="4C064D30D949FFE96BC8FDE4FE72FD86" baseAuthorityName="Dumeril & Bibron" baseAuthorityYear="1836" box="[151,411,588,612]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D949FFE96BC8FDE4FE72FD86" box="[151,411,588,612]" italics="true" pageId="7" pageNumber="274">Hoplodactylus duvaucelii</emphasis>
|
||
</taxonomicName>
|
||
.–
|
||
<treatmentCitation id="0AA710A2D949FFE96AEBFDE4FD80FD87" author="Holder, L. A." box="[436,617,588,613]" page="302" pageId="7" pageNumber="274" year="1960">
|
||
<bibRefCitation id="EF974B42D949FFE96AEBFDE4FD80FD87" author="Holder, L. A." box="[436,617,588,613]" pageId="7" pageNumber="274" pagination="300 - 335" refId="ref17817" refString="Holder, L. A. (1960) The comparative morphology of the axial skeleton in the Australian Gekkonidae. Zoological Journal of the Linnean Society, 44, 300 - 335. https: // doi. org / 10.1111 / j. 1096 - 3642.1960. tb 01617. x" type="journal article" year="1960">Holder 1960: 302</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE9692CFDE4FCF7FD86" author="Kluge, A." box="[627,798,588,612]" page="25" pageId="7" pageNumber="274" year="1967">
|
||
<bibRefCitation id="EF974B42D949FFE9692CFDE4FCF7FD86" author="Kluge, A." box="[627,798,588,612]" pageId="7" pageNumber="274" pagination="1 - 60" refId="ref18358" refString="Kluge, A. (1967 a) Higher taxonomic categories of gekkonid lizards and their evolution. Bulletin of the American Museum of Natural History, 135, 1 - 60." type="journal article" year="1967">Kluge 1967a: 25</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96876FDE4FC06FD87" author="Kluge, A." box="[809,1007,588,613]" page="1013" pageId="7" pageNumber="274" year="1967">
|
||
<bibRefCitation id="EF974B42D949FFE96876FDE4FC06FD87" author="Kluge, A." box="[809,1007,588,613]" pageId="7" pageNumber="274" pagination="1007 - 1108" refId="ref18391" refString="Kluge, A. (1967 b) Systematics, phylogeny, and zoogeography of the lizard genus Diplodactylus Gray (Gekkonidae). Australian Journal of Zoology, 15, 1007 - 1108. https: // doi. org / 10.1071 / ZO 9671007" type="journal article" year="1967">Kluge 1967b: 1013</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE968A6FDE5FB03FD87" author="Werner, Y. L. & Whitaker, A. H. & Whitaker, A. H." box="[1017,1258,588,613]" page="378" pageId="7" pageNumber="274" year="1978">
|
||
<bibRefCitation id="EF974B42D949FFE968A6FDE5FB03FD87" author="Werner, Y. L. & Whitaker, A. H. & Whitaker, A. H." box="[1017,1258,588,613]" pageId="7" pageNumber="274" pagination="375 - 393" refId="ref24037" refString="Werner, Y. L., Whitaker, A. H. & Whitaker, A. H. (1978) Observations and comments on the body temperatures of some New Zealand reptiles. New Zealand Journal of Zoology, 5, 375 - 393. https: // doi. org / 10.1080 / 03014223.1978.10428324" type="journal article" year="1978">
|
||
Werner
|
||
<emphasis id="B972EAA1D949FFE96F16FDE5FB95FD86" box="[1097,1148,588,612]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
1978: 378
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;7
|
||
<treatmentCitation id="0AA710A2D949FFE96E5EFDE4FF3DFD61" author="Kennedy, C. M." page="1" pageId="7" pageNumber="274" year="1979">
|
||
<bibRefCitation id="EF974B42D949FFE96E5EFDE4FF3DFD61" author="Kennedy, C. M." pageId="7" pageNumber="274" refId="ref18280" refString="Kennedy, C. M. (1979) Observations on a haemogregarine and mites from the gecko Hoplodactylus duvaucelii (Reptilia: Lacertilia), from the Trios Islands. MSc thesis, University of Canterbury, Christchurch. [unknown pagination]" type="book" year="1979">Kennedy 1979: 1</bibRefCitation>
|
||
</treatmentCitation>
|
||
; Bauer 1986: 9;
|
||
<treatmentCitation id="0AA710A2D949FFE96A21FDC3FDAFFD61" author="Worthy, T. H." box="[382,582,619,643]" page="416" pageId="7" pageNumber="274" year="1987">
|
||
<bibRefCitation id="EF974B42D949FFE96A21FDC3FDAFFD61" author="Worthy, T. H." box="[382,582,619,643]" pageId="7" pageNumber="274" pagination="409 - 420" refId="ref25088" refString="Worthy, T. H. (1987 b) Palaeoecological information concerning members of the frog genus leiopelma: leiopelmatidae in New Zealand. Journal of the Royal Society of New Zealand, 17, 409 - 420. https: // doi. org / 10.1080 / 03036758.1987.10426482" type="journal article" year="1987">Worthy 1987b: 416</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE9690EFDC4FD14FD61" author="Bauer, A." box="[593,765,619,643]" page="108" pageId="7" pageNumber="274" year="1990">
|
||
<bibRefCitation id="EF974B42D949FFE9690EFDC4FD14FD61" author="Bauer, A." box="[593,765,619,643]" pageId="7" pageNumber="274" pagination="1 - 218" refId="ref12618" refString="Bauer, A. (1990) Phylogenetic systematics and biogeography of the Carphodactylini (Reptilia: Gekkonidae). Bonn Zoological Bulletin, 30, 1 - 218." type="journal article" year="1990">Bauer 1990: 108</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96858FDC3FBF5FD61" author="Thompson, M. B. & Daugherty, C. H. & Cree, A. & French, D. C. & Gillingham, J. C. & Barwick, R. E." box="[775,1052,619,643]" page="123" pageId="7" pageNumber="274" year="1992">
|
||
<bibRefCitation id="EF974B42D949FFE96858FDC3FBF5FD61" author="Thompson, M. B. & Daugherty, C. H. & Cree, A. & French, D. C. & Gillingham, J. C. & Barwick, R. E." box="[775,1052,619,643]" pageId="7" pageNumber="274" pagination="123 - 130" refId="ref22613" refString="Thompson, M. B., Daugherty, C. H., Cree, A., French, D. C., Gillingham, J. C. & Barwick, R. E. (1992) Status and longevity of the tuatara, Sphenodon guntheri, and Duvaucel's gecko, Hoplodactylus duvaucelii, on North Brother Island, New Zealand. Journal of the Royal Society of New Zealand, 22, 123 - 130. https: // doi. org / 10.1080 / 03036758.1992.10420810" type="journal article" year="1992">
|
||
Thompson
|
||
<emphasis id="B972EAA1D949FFE96825FDC4FC44FD61" box="[890,941,619,643]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
1992: 123
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96F78FDC3FAD2FD61" author="Daugherty, C. H. & Gibbs, G. W. & Hitchmough, R. A." box="[1063,1339,619,643]" page="439" pageId="7" pageNumber="274" year="1993">
|
||
<bibRefCitation id="EF974B42D949FFE96F78FDC3FAD2FD61" author="Daugherty, C. H. & Gibbs, G. W. & Hitchmough, R. A." box="[1063,1339,619,643]" pageId="7" pageNumber="274" pagination="437 - 442" refId="ref14317" refString="Daugherty, C. H., Gibbs, G. W. & Hitchmough, R. A. (1993) Mega-island or micro-continent? New Zealand and its fauna. Trends in Ecology and Evolution, 8, 437 - 442. https: // doi. org / 10.1016 / 0169 - 5347 (93) 90006 - B" type="journal article" year="1993">
|
||
Daugherty
|
||
<emphasis id="B972EAA1D949FFE96FC7FDC4FB25FD61" box="[1176,1228,619,643]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
1993: 439
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96E19FDC4FE9AFD40" author="Bauer, A. M. & Henle, K." page="139" pageId="7" pageNumber="274" year="1994">
|
||
<bibRefCitation id="EF974B42D949FFE96E19FDC4FE9AFD40" author="Bauer, A. M. & Henle, K." pageId="7" pageNumber="274" refId="ref12876" refString="Bauer, A. M. & Henle, K. (1994) Das Tierreich. Teilband 109: Familia Gekkonidae (Reptilia, Sauria). Walter de Gruyter, Berlin, 319 pp. https: // doi. org / 10.1515 / 9783110885958" type="book" year="1994">Bauer & Henle 1994: 139</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96A21FD22FDF2FD40" author="Cree, A." box="[382,539,650,674]" page="352" pageId="7" pageNumber="274" year="1994">
|
||
<bibRefCitation id="EF974B42D949FFE96A21FD22FDF2FD40" author="Cree, A." box="[382,539,650,674]" pageId="7" pageNumber="274" pagination="351 - 372" refId="ref14207" refString="Cree, A. (1994) Low annual reproductive output in female reptiles from New Zealand. New Zealand Journal of Zoology, 21, 351 - 372. https: // doi. org / 10.1080 / 03014223.1994.9518005" type="journal article" year="1994">Cree 1994: 352</bibRefCitation>
|
||
</treatmentCitation>
|
||
Daugherty
|
||
<emphasis id="B972EAA1D949FFE969CDFD23FD2FFD40" box="[658,710,650,674]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
1994: 318;
|
||
<treatmentCitation id="0AA710A2D949FFE96860FD22FB97FD40" author="Towns, D. R. & Daugherty, C. H." box="[831,1150,650,674]" page="327" pageId="7" pageNumber="274" year="1994">
|
||
<bibRefCitation id="EF974B42D949FFE96860FD22FB97FD40" author="Towns, D. R. & Daugherty, C. H." box="[831,1150,650,674]" pageId="7" pageNumber="274" pagination="325 - 339" refId="ref23424" refString="Towns, D. R. & Daugherty, C. H. (1994) Patterns of range contractions and extinctions in the New Zealand herpetofauna following human colonisation. New Zealand Journal of Zoology, 21, 325 - 339. https: // doi. org / 10.1080 / 03014223.1994.9518003" type="journal article" year="1994">Towns & Daugherty 1994: 327</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96FD7FD22FF19FD22" author="Worthy, T. H. & Holdaway, R. N." page="326" pageId="7" pageNumber="274" year="1994">
|
||
<bibRefCitation id="EF974B42D949FFE96FD7FD22FF19FD22" author="Worthy, T. H. & Holdaway, R. N." pageId="7" pageNumber="274" pagination="297 - 391" refId="ref25352" refString="Worthy, T. H. & Holdaway, R. N. (1994) Quaternary fossil faunas from caves in Takaka Valley and on Takaka Hill, northwest Nelson, South Island, New Zealand. Journal of the Royal Society of New Zealand, 24, 297 - 391. https: // doi. org / 10.1080 / 03014223.1994.9517474" type="journal article" year="1994">Worthy & Holdaway 1994: 326</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96BA4FD01FE27FD22" author="East, K. T. & East, M. R. & Daugherty, C. H." box="[251,462,680,704]" page="256" pageId="7" pageNumber="274" year="1995">
|
||
<bibRefCitation id="EF974B42D949FFE96BA4FD01FE27FD22" author="East, K. T. & East, M. R. & Daugherty, C. H." box="[251,462,680,704]" pageId="7" pageNumber="274" pagination="249 - 261" refId="ref14842" refString="East, K. T., East, M. R. & Daugherty, C. H. (1995) Ecological restoration and habitat relationships of reptiles on Stephens Island, New Zealand. New Zealand Journal of Zoology, 22, 249 - 261. https: // doi. org / 10.1080 / 03014223.1995.9518040" type="journal article" year="1995">
|
||
East
|
||
<emphasis id="B972EAA1D949FFE96A73FD01FE89FD22" box="[300,352,680,704]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
1995: 256
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96A87FD00FCF2FD22" author="Worthy, T. H. & Holdaway, R. N." box="[472,795,680,705]" page="350" pageId="7" pageNumber="274" year="1995">
|
||
<bibRefCitation id="EF974B42D949FFE96A87FD00FCF2FD22" author="Worthy, T. H. & Holdaway, R. N." box="[472,795,680,705]" pageId="7" pageNumber="274" pagination="333 - 370" refId="ref25415" refString="Worthy, T. H. & Holdaway, R. N. (1995) Quaternary fossil faunas from caves on Mt Cookson, North Canterbury, South Island, New Zealand. Journal of the Royal Society of New Zealand, 25, 333 - 370. https: // doi. org / 10.1080 / 03014223.1995.9517494" type="journal article" year="1995">Worthy & Holdaway 1995: 350</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE9687AFD00FB81FD23" author="Worthy, T. H. & Holdaway, R. N." box="[805,1128,680,705]" page="314" pageId="7" pageNumber="274" year="1996">
|
||
<bibRefCitation id="EF974B42D949FFE9687AFD00FB81FD23" author="Worthy, T. H. & Holdaway, R. N." box="[805,1128,680,705]" pageId="7" pageNumber="274" pagination="275 - 361" refId="ref25474" refString="Worthy, T. H. & Holdaway, R. N. (1996) Quaternary fossil faunas, overlapping taphonomies, and palaeofaunal reconstruction in North Canterbury, South Island, New Zealand. Journal of the Royal Society of New Zealand, 26, 275 - 361. https: // doi. org / 10.1080 / 03014223.1996.9517514" type="journal article" year="1996">Worthy & Holdaway 1996: 314</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96F2DFD00FAADFD23" author="Hitchmough, R. A." box="[1138,1348,680,705]" page="1" pageId="7" pageNumber="274" year="1997">
|
||
<bibRefCitation id="EF974B42D949FFE96F2DFD00FAADFD23" author="Hitchmough, R. A." box="[1138,1348,680,705]" pageId="7" pageNumber="274" refId="ref17230" refString="Hitchmough, R. A. (1997) A systematic revision of the New Zealand Gekkonidae. PhD thesis, Victoria University of Wellington, Wellington, 374 pp." type="book" year="1997">Hitchmough 1997: 1</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96E0EFD00FECDFD3D" author="Worthy, T. H." page="93" pageId="7" pageNumber="274" year="1997">
|
||
<bibRefCitation id="EF974B42D949FFE96E0EFD00FECDFD3D" author="Worthy, T. H." pageId="7" pageNumber="274" pagination="67 - 162" refId="ref25192" refString="Worthy, T. H. (1997) Quaternary fossil fauna of South Canterbury, South Island, New Zealand. Journal of the Royal Society of New Zealand, 27, 67 - 162. https: // doi. org / 10.1080 / 03014223.1997.9517528" type="journal article" year="1997">Worthy 1997: 93</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96A6FFD60FE38FD3D" author="Bauer, A. M." box="[304,465,711,735]" page="43" pageId="7" pageNumber="274" year="1998">
|
||
<bibRefCitation id="EF974B42D949FFE96A6FFD60FE38FD3D" author="Bauer, A. M." box="[304,465,711,735]" pageId="7" pageNumber="274" pagination="41 - 58" refId="ref12776" refString="Bauer, A. M. (1998) Morphology of the adhesive tail tips of Carphodactyline geckos (Reptilia: Diplodactylidae). Journal of Morphology, 235, 41 - 58. https: // doi. org / 10.1002 / (SICI) 1097 - 4687 (199801) 235: 1" type="journal article" year="1998">Bauer 1998: 43</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96A83FD6FFD38FD3D" author="Girling, J. E. & Cree, A. & Guillette, L. J." box="[476,721,711,735]" page="139" pageId="7" pageNumber="274" year="1998">
|
||
<bibRefCitation id="EF974B42D949FFE96A83FD6FFD38FD3D" author="Girling, J. E. & Cree, A. & Guillette, L. J." box="[476,721,711,735]" pageId="7" pageNumber="274" pagination="139 - 154" refId="ref15740" refString="Girling, J. E., Cree, A. & Guillette, L. J. (1998) Oviducal structure in four species of gekkonid lizard differing in parity mode and eggshell structure. Reproduction, Fertility and Development, 10, 139 - 154. https: // doi. org / 10.1071 / R 97056" type="journal article" year="1998">
|
||
Girling
|
||
<emphasis id="B972EAA1D949FFE96972FD60FD88FD3D" box="[557,609,711,735]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
1998: 139
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 4;
|
||
<treatmentCitation id="0AA710A2D949FFE9687FFD6FFC34FD3D" author="Worthy, T. H." box="[800,989,711,735]" page="448" pageId="7" pageNumber="274" year="1998">
|
||
<bibRefCitation id="EF974B42D949FFE9687FFD6FFC34FD3D" author="Worthy, T. H." box="[800,989,711,735]" pageId="7" pageNumber="274" pagination="421 - 521" refId="ref25239" refString="Worthy, T. H. (1998) Quaternary fossil faunas of Otago, South Island, New Zealand. Journal of the Royal Society of New Zealand, 28, 421 - 521. https: // doi. org / 10.1080 / 03014223.1998.9517573" type="journal article" year="1998">Worthy 1998: 448</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE968B7FD6FFB06FD3D" author="Bannock, C. & Whitaker, A. & Hickling, G." box="[1000,1263,711,735]" page="102" pageId="7" pageNumber="274" year="1999">
|
||
<bibRefCitation id="EF974B42D949FFE968B7FD6FFB06FD3D" author="Bannock, C. & Whitaker, A. & Hickling, G." box="[1000,1263,711,735]" pageId="7" pageNumber="274" pagination="101 - 103" refId="ref12222" refString="Bannock, C., Whitaker, A. & Hickling, G. (1999) Extreme longevity of the common gecko (Hoplodactylus maculatus) on Motunau Island, Canterbury, New Zealand. New Zealand Journal of Ecology, 23, 101 - 103." type="journal article" year="1999">
|
||
Bannock
|
||
<emphasis id="B972EAA1D949FFE96F15FD60FB97FD3D" box="[1098,1150,711,735]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
1999: 102
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96FA5FD6FFA71FD3D" author="Lukis, K." box="[1274,1432,711,735]" page="12" pageId="7" pageNumber="274" year="1999">
|
||
<bibRefCitation id="EF974B42D949FFE96FA5FD6FFA71FD3D" author="Lukis, K." box="[1274,1432,711,735]" pageId="7" pageNumber="274" refId="ref19065" refString="Lukis, K. (1999) Survey of the lizard fauna of Quail Island, Lyttelton Harbour, Canterbury. Lincoln University Wildlife Management Report. No. 23. Lincoln University, Christchurch, 16 pp." type="book" year="1999">Lukis 1999: 12</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96B98FD4EFE6DFD1C" author="Flannagan, H. J." box="[199,388,742,766]" page="4" pageId="7" pageNumber="274" year="2000">
|
||
<bibRefCitation id="EF974B42D949FFE96B98FD4EFE6DFD1C" author="Flannagan, H. J." box="[199,388,742,766]" pageId="7" pageNumber="274" refId="ref15212" refString="Flannagan, H. J. (2000) Conservation biology of the goldstripe gecko (Hoplodactylus chrysosireticus) and interactions with Duvaucel's gecko (Hoplodactylus duvaucelii) on Mana Island, Cook Strait, New Zealand. MSc thesis, Massey University, Auckland, 116 pp." type="book" year="2000">Flannagan 2000: 4</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96AD1FD4EFDFEFD1C" author="Jones, N." box="[398,535,742,766]" page="1" pageId="7" pageNumber="274" year="2000">
|
||
<bibRefCitation id="EF974B42D949FFE96AD1FD4EFDFEFD1C" author="Jones, N." box="[398,535,742,766]" pageId="7" pageNumber="274" refId="ref18148" refString="Jones, N. (2000) Establishment, dispersal and population viability of translocated Duvaucel's gecko (Hoplodactylus duvaucelii) on Mana Island. MSc thesis, Victoria University of Wellington, Wellington, 79 pp." type="book" year="2000">Jones 2000: 1</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 2.9;
|
||
<treatmentCitation id="0AA710A2D949FFE9692AFD4EFC7DFD1C" author="Towns, D. R. & Ferreira, S. M." box="[629,916,742,766]" page="219" pageId="7" pageNumber="274" year="2001">
|
||
<bibRefCitation id="EF974B42D949FFE9692AFD4EFC7DFD1C" author="Towns, D. R. & Ferreira, S. M." box="[629,916,742,766]" pageId="7" pageNumber="274" pagination="211 - 222" refId="ref23547" refString="Towns, D. R. & Ferreira, S. M. (2001) Conservation of New Zealand lizards (Lacertilia: Scincidae) by translocation of small populations. Biological Conservation, 98, 211 - 222. https: // doi. org / 10.1016 / S 0006 - 3207 (00) 00156 - 7" type="journal article" year="2001">Towns & Ferreira 2001: 219</bibRefCitation>
|
||
</treatmentCitation>
|
||
; Towns
|
||
<emphasis id="B972EAA1D949FFE968BAFD4FFBFEFD1C" box="[997,1047,742,766]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
2001: 4;
|
||
<treatmentCitation id="0AA710A2D949FFE96F2BFD4EFAA4FD1C" author="Seligmann, H." box="[1140,1357,742,766]" page="277" pageId="7" pageNumber="274" year="2002">
|
||
<bibRefCitation id="EF974B42D949FFE96F2BFD4EFAA4FD1C" author="Seligmann, H." box="[1140,1357,742,766]" pageId="7" pageNumber="274" pagination="277 - 283" refId="ref21780" refString="Seligmann, H. (2002) Behavioural and morphological asymmetries in hindlimbs of Hoplodactylus duvaucelii (Lacertilia: Gekkonomorpha: Gekkota: Diplodactylinae). Laterality, 7 (3), 277 - 283. https: // doi. org / 10.1080 / 13576500244000067" type="journal article" year="2002">Seligmann 2002: 277</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96E09FD4EFE9DFCFF" author="Whitaker, T. & Tocher, M. & Blair, T." page="1" pageId="7" pageNumber="274" year="2002">
|
||
<bibRefCitation id="EF974B42D949FFE96E09FD4EFE9DFCFF" author="Whitaker, T. & Tocher, M. & Blair, T." pageId="7" pageNumber="274" refId="ref24341" refString="Whitaker, T., Tocher, M. & Blair, T. (2002) Conservation of lizards in Otago Conservancy 2002 - 2007. Department of Conservation, Wellington, 96 pp." type="book" year="2002">
|
||
Whitaker
|
||
<emphasis id="B972EAA1D949FFE96BB1FCADFEC8FCFE" box="[238,289,772,796]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
2002: 1
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96ADDFCADFDA1FCFE" author="Hay, J. M. & Daugherty, C. H. & Cree, A. & Maxson, L. R." box="[386,584,772,797]" page="16" pageId="7" pageNumber="274" year="2003">
|
||
<bibRefCitation id="EF974B42D949FFE96ADDFCADFDA1FCFE" author="Hay, J. M. & Daugherty, C. H. & Cree, A. & Maxson, L. R." box="[386,584,772,797]" pageId="7" pageNumber="274" pagination="1 - 19" refId="ref16854" refString="Hay, J. M., Daugherty, C. H., Cree, A. & Maxson, L. R. (2003) Low genetic divergence obscures phylogeny among populations of Sphenodon, remnant of an ancient reptile lineage. Molecular Phylogenetics and Evolution, 29, 1 - 19. https: // doi. org / 10.1016 / S 1055 - 7903 (03) 00091 - 5" type="journal article" year="2003">
|
||
Hay
|
||
<emphasis id="B972EAA1D949FFE96AEDFCADFE0FFCFE" box="[434,486,772,796]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
2003: 16
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE9690DFCACFD01FCFF" author="Todd, A. C." box="[594,744,772,797]" page="17" pageId="7" pageNumber="274" year="2003">
|
||
<bibRefCitation id="EF974B42D949FFE9690DFCACFD01FCFF" author="Todd, A. C." box="[594,744,772,797]" pageId="7" pageNumber="274" refId="ref22867" refString="Todd, A. C. (2003) Mating strategies and sperm competition in New Zealand geckos (Family Gekkonidae). PhD, University of Otago, Dunedin, 192 pp." type="book" year="2003">Todd 2003: 17</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE969ACFCACFBE1FCFF" author="Seligmann, H. & Beiles, A. & Werner, Y. L." box="[755,1032,772,797]" page="130" pageId="7" pageNumber="274" year="2003">
|
||
<bibRefCitation id="EF974B42D949FFE969ACFCACFBE1FCFF" author="Seligmann, H. & Beiles, A. & Werner, Y. L." box="[755,1032,772,797]" pageId="7" pageNumber="274" pagination="129 - 144" refId="ref21827" refString="Seligmann, H., Beiles, A. & Werner, Y. L. (2003) More injuries in left-footed individual lizards and Sphenodon. Journal of Zoology, 260, 129 - 144. https: // doi. org / 10.1017 / S 0952836903003558" type="journal article" year="2003">
|
||
Seligmann
|
||
<emphasis id="B972EAA1D949FFE96839FCADFC70FCFE" box="[870,921,772,796]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
2003: 130
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96F4CFCACFB5EFCFF" author="Holmes, K." box="[1043,1207,772,797]" page="4" pageId="7" pageNumber="274" year="2004">
|
||
<bibRefCitation id="EF974B42D949FFE96F4CFCACFB5EFCFF" author="Holmes, K." box="[1043,1207,772,797]" pageId="7" pageNumber="274" refId="ref17910" refString="Holmes, K. (2004) The female reproductive cycle of a viviparous skink, Oligosoma maccanni, in a subalpine environment. MSc thesis, University of Otago, Dunedin, 119 pp." type="book" year="2004">Holmes 2004: 4</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96F9DFCACFA89FCFF" author="Naish, D." box="[1218,1376,772,797]" page="18" pageId="7" pageNumber="274" year="2004">
|
||
<bibRefCitation id="EF974B42D949FFE96F9DFCACFA89FCFF" author="Naish, D." box="[1218,1376,772,797]" pageId="7" pageNumber="274" pagination="17 - 21" refId="ref20286" refString="Naish, D. (2004) New Zealand's giant gecko: a review of current knowledge of Hoplodactylus delcourti and the kawekaweau of legend. The Cryptozoology Review, 4, 17 - 21." type="journal article" year="2004">Naish 2004: 18</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96E34FCADFE6AFCD9" author="Hare, K. & Cree, A." page="137" pageId="7" pageNumber="274" year="2005">
|
||
<bibRefCitation id="EF974B42D949FFE96E34FCADFE6AFCD9" author="Hare, K. & Cree, A." pageId="7" pageNumber="274" pagination="137 - 142" refId="ref16468" refString="Hare, K. & Cree, A. (2005) Natural history of Hoplodactylus stephensi (Reptilia: Gekkonidae) on Stephens Island, Cook Strait, New Zealand. New Zealand Journal of Ecology, 29, 137 - 142." type="journal article" year="2005">Hare & Cree 2005: 137</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96AD2FC8BFD08FCD9" author="Armstrong, D. P. & Davidson, R." box="[397,737,803,827]" page="74" pageId="7" pageNumber="274" year="2006">
|
||
<bibRefCitation id="EF974B42D949FFE96AD2FC8BFD08FCD9" author="Armstrong, D. P. & Davidson, R." box="[397,737,803,827]" pageId="7" pageNumber="274" pagination="73 - 85" refId="ref11704" refString="Armstrong, D. P. & Davidson, R. (2006) Developing population models for guiding reintroductions of extirpated bird species back to the New Zealand mainland. New Zealand Journal of Ecology, 30, 73 - 85. https: // doi. org / 10.1016 / 0006 - 3207 (94) 00038 - R" type="journal article" year="2006">Armstrong & Davidson 2006: 74</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE969B2FC8CFC54FCD9" author="Hare, K. & Hoare, J. & Hitchmough, R." box="[749,957,803,827]" page="89" pageId="7" pageNumber="274" year="2007">
|
||
<bibRefCitation id="EF974B42D949FFE969B2FC8CFC54FCD9" author="Hare, K. & Hoare, J. & Hitchmough, R." box="[749,957,803,827]" pageId="7" pageNumber="274" pagination="81 - 93" refId="ref16512" refString="Hare, K., Hoare, J. & Hitchmough, R. (2007) Investigating natural population dynamics of Naultinus manukanus to inform conservation management of New Zealand's cryptic diurnal geckos. Journal of Herpetology, 41, 81 - 93. https: // doi. org / 10.1670 / 0022 - 1511 (2007) 41 [81: INPDON] 2.0. CO; 2" type="journal article" year="2007">
|
||
Hare
|
||
<emphasis id="B972EAA1D949FFE9687AFC8CFCB0FCD9" box="[805,857,803,827]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
2007: 89
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96897FC8BFB84FCD9" author="Nielsen, S. V." box="[968,1133,803,827]" page="5" pageId="7" pageNumber="274" year="2008">
|
||
<bibRefCitation id="EF974B42D949FFE96897FC8BFB84FCD9" author="Nielsen, S. V." box="[968,1133,803,827]" pageId="7" pageNumber="274" refId="ref20587" refString="Nielsen, S. V. (2008) Molecular systematics of the geckos of New Zealand. MSc thesis, Villanova University, Villanova, 83 pp." type="book" year="2008">Nielsen 2008: 5</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96F26FC8BFA71FCD9" author="Kelly, D. & Sullivan, J. J." box="[1145,1432,803,827]" page="208" pageId="7" pageNumber="274" year="2010">
|
||
<bibRefCitation id="EF974B42D949FFE96F26FC8BFA71FCD9" author="Kelly, D. & Sullivan, J. J." box="[1145,1432,803,827]" pageId="7" pageNumber="274" pagination="207 - 217" refId="ref18231" refString="Kelly, D. & Sullivan, J. J. (2010) Life histories, dispersal, invasions, and global change: progress and prospects in New Zealand ecology, 1989 - 2029. New Zealand Journal of Ecology, 34, 207 - 217." type="journal article" year="2010">Kelly & Sullivan 2010: 208</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96B98FCEAFE9FFCB8" author="Miskelly, C." box="[199,374,834,858]" page="3" pageId="7" pageNumber="274" year="2010">
|
||
<bibRefCitation id="EF974B42D949FFE96B98FCEAFE9FFCB8" author="Miskelly, C." box="[199,374,834,858]" pageId="7" pageNumber="274" refId="ref19699" refString="Miskelly, C. (2010) Mana Island ecological restoration plan review. Department of Conservation, Wellington, 54 pp." type="book" year="2010">Miskelly 2010: 3</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96ADEFCEAFDF7FCB8" author="Wilson, J." box="[385,542,834,858]" page="6" pageId="7" pageNumber="274" year="2010">
|
||
<bibRefCitation id="EF974B42D949FFE96ADEFCEAFDF7FCB8" author="Wilson, J." box="[385,542,834,858]" pageId="7" pageNumber="274" refId="ref24377" refString="Wilson, J. (2010) Population viability and resource competition on North Brother Island: conservation implications for tuatara (Sphenodon punctatus) and Duvaucel's gecko (Hoplodactylus duvaucelii). MSc Thesis, Victoria University of Wellington, Wellington, 106 pp." type="book" year="2010">Wilson 2010: 6</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96976FCEAFCC5FCB8" author="Frank, H. & Wilson, D." box="[553,812,834,858]" page="16" pageId="7" pageNumber="274" year="2011">
|
||
<bibRefCitation id="EF974B42D949FFE96976FCEAFCC5FCB8" author="Frank, H. & Wilson, D." box="[553,812,834,858]" pageId="7" pageNumber="274" pagination="15 - 28" refId="ref15293" refString="Frank, H. & Wilson, D. (2011) Distribution, status and conservation measures for lizards in limestone areas of South Canterbury, New Zealand. New Zealand Journal of Zoology, 38, 15 - 28. https: // doi. org / 10.1080 / 03014223.2010.520326" type="journal article" year="2011">Frank & Wilson 2011: 16</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96868FCEAFBF7FCB8" author="Nielsen, S. V & Bauer, A. M. & Jackman, T. R. & Hitchmough, R. A. & Daugherty, C. H." box="[823,1054,834,858]" page="17" pageId="7" pageNumber="274" year="2011">
|
||
<bibRefCitation id="EF974B42D949FFE96868FCEAFBF7FCB8" author="Nielsen, S. V & Bauer, A. M. & Jackman, T. R. & Hitchmough, R. A. & Daugherty, C. H." box="[823,1054,834,858]" pageId="7" pageNumber="274" pagination="1 - 22" refId="ref20616" refString="Nielsen, S. V, Bauer, A. M., Jackman, T. R., Hitchmough, R. A. & Daugherty, C. H. (2011) New Zealand geckos (Diplodactylidae): Cryptic diversity in a post-Gondwanan lineage with trans-Tasman affinities. Molecular Phylogenetics and Evolution, 59, 1 - 22. https: // doi. org / 10.1016 / j. ympev. 2010.12.007" type="journal article" year="2011">
|
||
Nielsen
|
||
<emphasis id="B972EAA1D949FFE968D5FCEBFC57FCB8" box="[906,958,834,858]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
2011: 17
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96F77FCEAFAFCFCB8" author="Bell, T. P. & Herbert, S. M." box="[1064,1301,834,858]" page="8" pageId="7" pageNumber="274" year="2012">
|
||
<bibRefCitation id="EF974B42D949FFE96F77FCEAFAFCFCB8" author="Bell, T. P. & Herbert, S. M." box="[1064,1301,834,858]" pageId="7" pageNumber="274" pagination="1 - 35" refId="ref13033" refString="Bell, T. P. & Herbert, S. M. (2012) Following up Duvaucel's gecko (Hoplodactylus duvaucelii) on Mana Island. Post-translocation monitoring 11 - 14 years on. In: Sanctuaries of New Zealand Workshop. EcoGecko Consultants, Inglewood, pp. 1 - 35." type="book chapter" year="2012">Bell & Herbert 2012: 8</bibRefCitation>
|
||
</treatmentCitation>
|
||
; Str̂ckens
|
||
<emphasis id="B972EAA1D949FFE96ED5FCEBFF0BFC9A" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
2012: 542;
|
||
<treatmentCitation id="0AA710A2D949FFE96A03FCC8FD4DFC9A" author="Garcia-Porta, J. & Ord, T. J." box="[348,676,864,889]" page="2667" pageId="7" pageNumber="274" year="2013">
|
||
<bibRefCitation id="EF974B42D949FFE96A03FCC8FD4DFC9A" author="Garcia-Porta, J. & Ord, T. J." box="[348,676,864,889]" pageId="7" pageNumber="274" pagination="2662 - 2680" refId="ref15381" refString="Garcia-Porta, J. & Ord, T. J. (2013) Key innovations and island colonization as engines of evolutionary diversification: a comparative test with the Australasian diplodactyloid geckos. Journal of Evolutionary Biology, 26, 2662 - 2680. https: // doi. org / 10.1111 / jeb. 12261" type="journal article" year="2013">Garcia-Porta & Ord 2013: 2667</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE969F1FCC8FC23FC9B" author="Hitchmough, R. & Anderson, P. & Barr, B. & Monks, J. & Lettink, M. & Reardon, J. & Tocher, M. & Whitaker, T." box="[686,970,864,889]" page="10" pageId="7" pageNumber="274" year="2013">
|
||
<bibRefCitation id="EF974B42D949FFE969F1FCC8FC23FC9B" author="Hitchmough, R. & Anderson, P. & Barr, B. & Monks, J. & Lettink, M. & Reardon, J. & Tocher, M. & Whitaker, T." box="[686,970,864,889]" pageId="7" pageNumber="274" pagination="1 - 16" refId="ref17157" refString="Hitchmough, R., Anderson, P., Barr, B., Monks, J., Lettink, M., Reardon, J., Tocher, M. & Whitaker, T. (2013) Conservation status of New Zealand reptiles. 2012. In: New Zealand Threat Classification. Series 2. Department of Conservation, Wellington, pp. 1 - 16." type="book chapter" year="2013">
|
||
Hitchmough
|
||
<emphasis id="B972EAA1D949FFE9686CFCC9FC8EFC9A" box="[819,871,864,888]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
2013: 10
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE9688AFCC8FBBCFC9B" author="Bell, T." box="[981,1109,864,889]" page="8" pageId="7" pageNumber="274" year="2014">
|
||
<bibRefCitation id="EF974B42D949FFE9688AFCC8FBBCFC9B" author="Bell, T." box="[981,1109,864,889]" pageId="7" pageNumber="274" refId="ref12979" refString="Bell, T. (2014) Standardized common names for New Zealand reptiles. BioGecko, Wellington, 11 pp." type="book" year="2014">Bell 2014: 8</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
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Romijn
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2014: 111
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;
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;
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<treatmentCitation id="0AA710A2D949FFE96AE6FCD7FD86FC75" author="Mockett, S." box="[441,623,895,919]" page="73" pageId="7" pageNumber="274" year="2015">
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;
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<treatmentCitation id="0AA710A2D949FFE96925FCD7FCD9FC75" author="Chapple, D. G." box="[634,816,895,919]" page="4" pageId="7" pageNumber="274" year="2016">
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<bibRefCitation id="EF974B42D949FFE96925FCD7FCD9FC75" author="Chapple, D. G." box="[634,816,895,919]" pageId="7" pageNumber="274" pagination="361 - 375" refId="ref13916" refString="Chapple, D. G. (2016 b) The future of New Zealand lizard research. In: Chapple, D. G. (Ed.), New Zealand Lizards. Springer, Cham, pp. 361 - 375. https: // doi. org / 10.1007 / 978 - 3 - 319 - 41674 - 8 _ 14" type="book chapter" year="2016">Chapple 2016b: 4</bibRefCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE96864FCD7FB74FC75" author="Hitchmough, R. A. & Patterson, G. B. & Chapple, D. G." box="[827,1181,895,919]" page="116" pageId="7" pageNumber="274" year="2016">
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<bibRefCitation id="EF974B42D949FFE96864FCD7FB74FC75" author="Hitchmough, R. A. & Patterson, G. B. & Chapple, D. G." box="[827,1181,895,919]" pageId="7" pageNumber="274" pagination="87 - 108" refId="ref17547" refString="Hitchmough, R. A., Patterson, G. B. & Chapple, D. G. (2016 b) Putting a name to diversity: Taxonomy of the New Zealand lizard fauna. In: Chapple, D. G. (Ed.), New Zealand Lizards. Springer, Cham, pp. 87 - 108. https: // doi. org / 10.1007 / 978 - 3 - 319 - 41674 - 8 _ 4" type="book chapter" year="2016">Chapple & Hitchmough 2016: 116</bibRefCitation>
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; Cree & Hare 2016: 174; Hare
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2016: 140; Hare &
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<treatmentCitation id="0AA710A2D949FFE96AA8FC36FD7DFC54" author="Cree, A. & Hare, K. M." box="[503,660,926,950]" page="246" pageId="7" pageNumber="274" year="2016">
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<bibRefCitation id="EF974B42D949FFE96AA8FC36FD7DFC54" author="Cree, A. & Hare, K. M." box="[503,660,926,950]" pageId="7" pageNumber="274" pagination="169 - 206" refId="ref14247" refString="Cree, A. & Hare, K. M. (2016) Reproduction and life history of New Zealand lizards. In: Chapple, D. G. (Ed.), New Zealand Lizards. Springer, Cham, pp. 169 - 206. https: // doi. org / 10.1007 / 978 - 3 - 319 - 41674 - 8 _ 7" type="book chapter" year="2016">Cree 2016: 246</bibRefCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE969C1FC36FC5AFC54" author="Hitchmough, R. A. & Barr, B. & Lettnik, M. & Monks, J. M. & Reardon, J. T. & Tocher, M. & van Winkel, D. & Rolfe, J." box="[670,947,926,950]" page="9" pageId="7" pageNumber="274" year="2016">
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<bibRefCitation id="EF974B42D949FFE969C1FC36FC5AFC54" author="Hitchmough, R. A. & Barr, B. & Lettnik, M. & Monks, J. M. & Reardon, J. T. & Tocher, M. & van Winkel, D. & Rolfe, J." box="[670,947,926,950]" pageId="7" pageNumber="274" refId="ref17341" refString="Hitchmough, R. A., Barr, B., Lettnik, M., Monks, J. M., Reardon, J. T., Tocher, M., van Winkel, D. & Rolfe, J. (2016 a) Conservation status of New Zealand reptiles. 2015. Department of Conservation, Wellington, 18 pp." type="book" year="2016">
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Hitchmough
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<emphasis id="B972EAA1D949FFE9687DFC37FCBDFC54" box="[802,852,926,950]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
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2016a: 9
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</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE968E2FC36FB09FC54" author="Hitchmough, R. A. & Patterson, G. B. & Chapple, D. G." box="[957,1248,926,950]" page="89" pageId="7" pageNumber="274" year="2016">
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<bibRefCitation id="EF974B42D949FFE968E2FC36FB09FC54" author="Hitchmough, R. A. & Patterson, G. B. & Chapple, D. G." box="[957,1248,926,950]" pageId="7" pageNumber="274" pagination="87 - 108" refId="ref17547" refString="Hitchmough, R. A., Patterson, G. B. & Chapple, D. G. (2016 b) Putting a name to diversity: Taxonomy of the New Zealand lizard fauna. In: Chapple, D. G. (Ed.), New Zealand Lizards. Springer, Cham, pp. 87 - 108. https: // doi. org / 10.1007 / 978 - 3 - 319 - 41674 - 8 _ 4" type="book chapter" year="2016">
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Hitchmough
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<emphasis id="B972EAA1D949FFE96F1EFC37FB9AFC54" box="[1089,1139,926,950]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
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2016b: 89
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</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE96FB5FC36FEB7FC36" author="Morgan-Richards, M. & Hinlo, A. R. & Smuts-Kennedy, C. & Innes, J. & Ji, W. & Barry, M. & Brunton, D. & Hitchmough, R. A." page="77" pageId="7" pageNumber="274" year="2016">
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<bibRefCitation id="EF974B42D949FFE96FB5FC36FEB7FC36" author="Morgan-Richards, M. & Hinlo, A. R. & Smuts-Kennedy, C. & Innes, J. & Ji, W. & Barry, M. & Brunton, D. & Hitchmough, R. A." pageId="7" pageNumber="274" pagination="77 - 86" refId="ref20012" refString="Morgan-Richards, M., Hinlo, A. R., Smuts-Kennedy, C., Innes, J., Ji, W., Barry, M., Brunton, D. & Hitchmough, R. A. (2016) Identification of a rare gecko from North Island New Zealand, and genetic assessment of its probable origin: A novel mainland conservation priority? Journal of Herpetology, 50, 77 - 86. https: // doi. org / 10.1670 / 13 - 128" type="journal article" year="2016">
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Morgan-Richards
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2016: 77
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</bibRefCitation>
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</treatmentCitation>
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, fig. 2;
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<treatmentCitation id="0AA710A2D949FFE96AF2FC14FD01FC36" author="Romijn, R. L. & Hartley, S." box="[429,744,956,980]" page="196" pageId="7" pageNumber="274" year="2016">
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<bibRefCitation id="EF974B42D949FFE96AF2FC14FD01FC36" author="Romijn, R. L. & Hartley, S." box="[429,744,956,980]" pageId="7" pageNumber="274" pagination="191 - 210" refId="ref21339" refString="Romijn, R. L. & Hartley, S. (2016) Trends in lizard translocations in New Zealand between 1988 and 2013. New Zealand Journal of Zoology, 43, 191 - 210. https: // doi. org / 10.1080 / 03014223.2016.1146311" type="journal article" year="2016">Romijn and Hartley 2016: 196</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE969ADFC15FC04FC37" author="Towns, D. R. & Hitchmough, R. A. & Perrott, J." box="[754,1005,956,981]" page="308" pageId="7" pageNumber="274" year="2016">
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<bibRefCitation id="EF974B42D949FFE969ADFC15FC04FC37" author="Towns, D. R. & Hitchmough, R. A. & Perrott, J." box="[754,1005,956,981]" pageId="7" pageNumber="274" pagination="293 - 320" refId="ref23637" refString="Towns, D. R., Hitchmough, R. A. & Perrott, J. (2016 b) Conservation of New Zealand lizards: A fauna not forgotten but undervalued? In: Chapple, D. G. (Ed.), New Zealand Lizards. Springer, Cham, pp. 293 - 320. https: // doi. org / 10.1007 / 978 - 3 - 319 - 41674 - 8 _ 11" type="book chapter" year="2016">
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Towns
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2016b: 308
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE968A8FC14FB4FFC37" author="Worthy, T. H." box="[1015,1190,956,981]" page="71" pageId="7" pageNumber="274" year="2016">
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<bibRefCitation id="EF974B42D949FFE968A8FC14FB4FFC37" author="Worthy, T. H." box="[1015,1190,956,981]" pageId="7" pageNumber="274" pagination="65 - 86" refId="ref25285" refString="Worthy, T. H. (2016) A review of the fossil record of New Zealand lizards. In: Chapple, D. G. (Ed.), New Zealand Lizards. Springer, Cham, pp. 65 - 86. https: // doi. org / 10.1007 / 978 - 3 - 319 - 41674 - 8 _ 3" type="book chapter" year="2016">Worthy 2016: 71</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE96FEDFC14FF0AFC11" author="Bell, T. P. & Herbert, S. M." page="38" pageId="7" pageNumber="274" year="2017">
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<bibRefCitation id="EF974B42D949FFE96FEDFC14FF0AFC11" author="Bell, T. P. & Herbert, S. M." pageId="7" pageNumber="274" pagination="37 - 46" refId="ref13088" refString="Bell, T. P. & Herbert, S. M. (2017) Establishment of a self-sustaining population of a long-lived, slow-breeding gecko species (Diplodactylidae: Hoplodactylus duvaucelii) evident 15 years after translocation. Journal of Herpetology, 51, 37 - 46. https: // doi. org / 10.1670 / 15 - 106" type="journal article" year="2017">Bell and Herbert 2017: 38</bibRefCitation>
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</treatmentCitation>
|
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;
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<treatmentCitation id="0AA710A2D949FFE96BB0FC74FE4BFC11" author="Bowers, L." box="[239,418,987,1011]" page="64" pageId="7" pageNumber="274" year="2017">
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<bibRefCitation id="EF974B42D949FFE96BB0FC74FE4BFC11" author="Bowers, L." box="[239,418,987,1011]" pageId="7" pageNumber="274" refId="ref13472" refString="Bowers, L. (2017) Change over time on Mana Island. MA thesis, University of Otago, Dunedin, 90 pp." type="book" year="2017">Bowers 2017: 64</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE96AF1FC74FD7DFC11" author="Knox, C. D. & Jarvie, S. & Easton, L. J. & Monks, J. M." box="[430,660,987,1011]" page="490" pageId="7" pageNumber="274" year="2017">
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<bibRefCitation id="EF974B42D949FFE96AF1FC74FD7DFC11" author="Knox, C. D. & Jarvie, S. & Easton, L. J. & Monks, J. M." box="[430,660,987,1011]" pageId="7" pageNumber="274" pagination="490 - 496" refId="ref18515" refString="Knox, C. D., Jarvie, S., Easton, L. J. & Monks, J. M. (2017) Soft-release, but not cool winter temperatures, reduces post-translocation dispersal of jewelled geckos. Journal of Herpetology, 51, 490 - 496. https: // doi. org / 10.1670 / 16 - 078" type="journal article" year="2017">
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Knox
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2017: 490
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE969C0FC73FC41FC11" author="Lozito, T. P. & Tuan, R. S." box="[671,936,987,1011]" page="148" pageId="7" pageNumber="274" year="2017">
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<bibRefCitation id="EF974B42D949FFE969C0FC73FC41FC11" author="Lozito, T. P. & Tuan, R. S." box="[671,936,987,1011]" pageId="7" pageNumber="274" pagination="145 - 154" refId="ref18978" refString="Lozito, T. P. & Tuan, R. S. (2017) Lizard tail regeneration as an instructive model of enhanced healing capabilities in an adult amniote. Connective Tissue Research, 58, 145 - 154. https: // doi. org / 10.1080 / 03008207.2016.1215444" type="journal article" year="2017">Lozito & Tuan 2017: 148</bibRefCitation>
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</treatmentCitation>
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, fig. 2
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<treatmentCitation id="0AA710A2D949FFE968ACFC73FB7AFC11" author="Sion, G." box="[1011,1171,987,1011]" page="131" pageId="7" pageNumber="274" year="2017">
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<bibRefCitation id="EF974B42D949FFE968ACFC73FB7AFC11" author="Sion, G." box="[1011,1171,987,1011]" pageId="7" pageNumber="274" pagination="129 - 151" refId="ref22029" refString="Sion, G. (2017) Foot-preference underlies bite-scar asymmetry in the gecko Ptyodactylus guttatus. Laterality, Series 1, 23, 129 - 151. https: // doi. org / 10.1080 / 1357650 X. 2017.1322097" type="journal article" year="2017">Sion 2017: 131</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE96FC0FC73FA71FC11" author="Stancher, G. & Sovrano, V. A. & Vallortigara, G." box="[1183,1432,987,1011]" page="36" pageId="7" pageNumber="274" year="2018">
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<bibRefCitation id="EF974B42D949FFE96FC0FC73FA71FC11" author="Stancher, G. & Sovrano, V. A. & Vallortigara, G." box="[1183,1432,987,1011]" pageId="7" pageNumber="274" pagination="33 - 56" refId="ref22300" refString="Stancher, G., Sovrano, V. A. & Vallortigara, G. (2018) Motor asymmetries in fishes, amphibians, and reptiles. Progress in Brain Research, 238, 33 - 56. https: // doi. org / 10.1016 / BS. PBR. 2018.06.002" type="journal article" year="2018">
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Stancher
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2018: 36
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</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE96B98FC52FE08FBF0" author="van Winkel, D. & Baling, M. & Hitchmough, R." box="[199,481,1018,1042]" page="114" pageId="7" pageNumber="274" year="2018">
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<bibRefCitation id="EF974B42D949FFE96B98FC52FE08FBF0" author="van Winkel, D. & Baling, M. & Hitchmough, R." box="[199,481,1018,1042]" pageId="7" pageNumber="274" refId="ref24596" refString="van Winkel, D., Baling, M. & Hitchmough, R. (2018) Reptiles and amphibians of New Zealand. Auckland University Press, Auckland, 712 pp." type="book" year="2018">
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van Winkel
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2018: 114
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</bibRefCitation>
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</treatmentCitation>
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, pls. 31, 40, 46, 128;
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<treatmentCitation id="0AA710A2D949FFE969E4FC52FC5BFBF0" author="Skipwith, P. L. & Bi, K. & Oliver, P. M." box="[699,946,1018,1042]" page="10" pageId="7" pageNumber="274" year="2019">
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<bibRefCitation id="EF974B42D949FFE969E4FC52FC5BFBF0" author="Skipwith, P. L. & Bi, K. & Oliver, P. M." box="[699,946,1018,1042]" pageId="7" pageNumber="274" pagination="106589" refId="ref22070" refString="Skipwith, P. L., Bi, K. & Oliver, P. M. (2019) Relicts and radiations: phylogenomics of an Australasian lizard clade with east Gondwanan origins (Gekkota: Diplodactyloidea). Molecular Phylogenetics and Evolution, 140, 106589. https: // doi. org / 10.1016 / j. ympev. 2019.106589" type="journal article" year="2019">
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Skipwith
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<emphasis id="B972EAA1D949FFE96842FC53FCB9FBF0" box="[797,848,1018,1042]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
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2019: 10
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</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE968E3FC52FB27FBF0" author="Florence-Bennett, B. J." box="[956,1230,1018,1042]" page="13" pageId="7" pageNumber="274" year="2020">
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE96F86FC52FF19FBD3" author="Glynne, E. & Daza, J. D. & Bauer, A. M." page="804" pageId="7" pageNumber="274" year="2020">
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<bibRefCitation id="EF974B42D949FFE96F86FC52FF19FBD3" author="Glynne, E. & Daza, J. D. & Bauer, A. M." pageId="7" pageNumber="274" pagination="801 - 813" refId="ref15838" refString="Glynne, E., Daza, J. D. & Bauer, A. M. (2020) Surface sculpturing in the skull of gecko lizards (Squamata: Gekkota). Biological Journal of the Linnean Society, 131, 801 - 813. https: // doi. org / 10.1093 / biolinnean / blaa 144" type="journal article" year="2020">
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Glynne
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2020: 804
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</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE96BA4FBB0FE45FBD3" author="Herbert, S. M." box="[251,428,1048,1073]" page="12" pageId="7" pageNumber="274" year="2020">
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<bibRefCitation id="EF974B42D949FFE96BA4FBB0FE45FBD3" author="Herbert, S. M." box="[251,428,1048,1073]" pageId="7" pageNumber="274" refId="ref17069" refString="Herbert, S. M. (2020) Is habitat enhancement a viable strategy for conserving New Zealand's endemic lizards? PhD thesis, Victoria University of Wellington, Wellington, 274 pp." type="book" year="2020">Herbert 2020: 12</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0AA710A2D949FFE96AE8FBB0FD7FFBD3" author="Price, S. J. & Grayson, K. L. & Gartrell, B. D. & Nelson, N. J." box="[439,662,1048,1073]" page="231" pageId="7" pageNumber="274" year="2020">
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<bibRefCitation id="EF974B42D949FFE96AE8FBB0FD7FFBD3" author="Price, S. J. & Grayson, K. L. & Gartrell, B. D. & Nelson, N. J." box="[439,662,1048,1073]" pageId="7" pageNumber="274" pagination="222 - 233" refId="ref21104" refString="Price, S. J., Grayson, K. L., Gartrell, B. D. & Nelson, N. J. (2020) Survival and growth of tuatara Sphenodon punctatus following translocation from the Cook Strait to warmer locations in their historic range. Oryx, 54, 222 - 233. https: // doi. org / 10.1017 / S 003060531800008 X" type="journal article" year="2020">
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Price
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</bibRefCitation>
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</treatmentCitation>
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; Scarsbrook 2021: 19, figs. 1.5, 2.1, 3.1, 3.3, 4.1;
|
||
<treatmentCitation id="0AA710A2D949FFE96FCAFBB0FA71FBD3" author="Scarsbrook, L. & Sherratt, E. & Hitchmough, R. A. & Rawlence, N. J." box="[1173,1432,1048,1073]" page="2" pageId="7" pageNumber="274" year="2021">
|
||
<bibRefCitation id="EF974B42D949FFE96FCAFBB0FA71FBD3" author="Scarsbrook, L. & Sherratt, E. & Hitchmough, R. A. & Rawlence, N. J." box="[1173,1432,1048,1073]" pageId="7" pageNumber="274" pagination="67" refId="ref21569" refString="Scarsbrook, L., Sherratt, E., Hitchmough, R. A. & Rawlence, N. J. (2021) Skeletal variation in extant species enables systematic identification of New Zealand's large, subfossil diplodactylids. BMC ecology and evolution, 21, 67. https: // doi. org / 10.1186 / s 12862 - 021 - 01808 - 7" type="journal article" year="2021">
|
||
Scarsbrook
|
||
<emphasis id="B972EAA1D949FFE96E51FBB1FAABFBD2" box="[1294,1346,1048,1072]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
2021: 2
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA710A2D949FFE96B98FB9FFE2EFBAD" author="Scarsbrook, L. & Verry, A. J. F. & Walton, K. & Hitchmough, R. A. & Rawlence, N. J." box="[199,455,1079,1103]" page="3" pageId="7" pageNumber="274" year="2022">
|
||
<bibRefCitation id="EF974B42D949FFE96B98FB9FFE2EFBAD" author="Scarsbrook, L. & Verry, A. J. F. & Walton, K. & Hitchmough, R. A. & Rawlence, N. J." box="[199,455,1079,1103]" pageId="7" pageNumber="274" pagination="1 - 21" refId="ref21637" refString="Scarsbrook, L., Verry, A. J. F., Walton, K., Hitchmough, R. A. & Rawlence, N. J. (2022) Ancient mitochondrial genomes recovered from small vertebrate bones through minimally destructive DNA extraction: phylogeography of the New Zealand gecko genus Hoplodactylus. Molecular Ecology, 1 - 21. [early view] https: // doi. org / 10.1111 / mec. 16434" type="book chapter" year="2022">
|
||
Scarsbrook
|
||
<emphasis id="B972EAA1D949FFE96A60FB90FE9AFBAD" box="[319,371,1079,1103]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
2022: 3
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 1 – all in part (not
|
||
<bibRefCitation id="EF974B42D949FFE969E8FB9FFC5FFBAD" author="Dumeril, A. M. C. & Bibron, G." box="[695,950,1079,1103]" pageId="7" pageNumber="274" refId="ref14713" refString="Dumeril, A. M. C. & Bibron, G. (1836) Erpetologie generale, ou, Histoire naturelle complete des reptiles. Vol. 3. Roret, Paris, 531 pp." type="book" year="1836">Duméril & Bibron, 1836</bibRefCitation>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB936B3D949FFE96BC8FBFEFBEEFB8C" blockId="7.[151,1437,466,1165]" box="[151,1031,1110,1134]" pageId="7" pageNumber="274">
|
||
<treatmentCitationGroup id="AB16119DD949FFE96BC8FBFEFBEEFB8C" box="[151,1031,1110,1134]" pageId="7" pageNumber="274">
|
||
<taxonomicName id="4C064D30D949FFE96BC8FBFEFE83FB8C" box="[151,362,1110,1134]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D949FFE96BC8FBFEFE83FB8C" box="[151,362,1110,1134]" italics="true" pageId="7" pageNumber="274">Naultinus duvaucelii</emphasis>
|
||
</taxonomicName>
|
||
.–
|
||
<treatmentCitation id="0AA710A2D949FFE96ADCFBFEFD93FB8C" author="Chrapliwy, P. S. & Smith, H. M. & Grant, C." box="[387,634,1110,1134]" page="6" pageId="7" pageNumber="274" year="1961">
|
||
<bibRefCitation id="EF974B42D949FFE96ADCFBFEFD93FB8C" author="Chrapliwy, P. S. & Smith, H. M. & Grant, C." box="[387,634,1110,1134]" pageId="7" pageNumber="274" pagination="5 - 12" refId="ref14090" refString="Chrapliwy, P. S., Smith, H. M. & Grant, C. (1961) The systematic status of the Geckonid lizard genera Gehyra, Peropus, Hoplodactylus and Naultinus. Herpetologica, 17, 5 - 12." type="journal article" year="1961">
|
||
Chrapliwy
|
||
<emphasis id="B972EAA1D949FFE96AABFBFFFDCEFB8C" box="[500,551,1110,1134]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
1961: 6
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
(in part, not
|
||
<bibRefCitation id="EF974B42D949FFE969A2FBFEFC15FB8C" author="Dumeril, A. M. C. & Bibron, G." box="[765,1020,1110,1134]" pageId="7" pageNumber="274" refId="ref14713" refString="Dumeril, A. M. C. & Bibron, G. (1836) Erpetologie generale, ou, Histoire naturelle complete des reptiles. Vol. 3. Roret, Paris, 531 pp." type="book" year="1836">Duméril & Bibron, 1836</bibRefCitation>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BB936B3D949FFE96BC8FBDCFC3FFB6E" blockId="7.[151,1437,466,1165]" box="[151,982,1140,1165]" pageId="7" pageNumber="274">
|
||
<treatmentCitationGroup id="AB16119DD949FFE96BC8FBDCFC3FFB6E" box="[151,982,1140,1165]" pageId="7" pageNumber="274">
|
||
<taxonomicName id="4C064D30D949FFE96BC8FBDCFE66FB6E" authorityName="Jewell" authorityYear="2008" box="[151,399,1140,1164]" class="Reptilia" family="Diplodactylidae" genus="Woodworthia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D949FFE96BC8FBDCFE66FB6E" box="[151,399,1140,1164]" italics="true" pageId="7" pageNumber="274">Woodworthia duvaucelii</emphasis>
|
||
</taxonomicName>
|
||
.–
|
||
<treatmentCitation id="0AA710A2D949FFE96AF7FBDCFDA1FB6E" author="Jewell, T." box="[424,584,1140,1165]" page="50" pageId="7" pageNumber="274" year="2008">
|
||
<bibRefCitation id="EF974B42D949FFE96AF7FBDCFDA1FB6E" author="Jewell, T." box="[424,584,1140,1165]" pageId="7" pageNumber="274" refId="ref17998" refString="Jewell, T. (2008) A photographic guide to reptiles and amphibians of New Zealand. New Holland Publishers Ltd, Auckland, 143 pp." type="book" year="2008">Jewell 2008: 50</bibRefCitation>
|
||
</treatmentCitation>
|
||
(in part, not
|
||
<bibRefCitation id="EF974B42D949FFE96993FBDCFC22FB6E" author="Dumeril, A. M. C. & Bibron, G." box="[716,971,1140,1165]" pageId="7" pageNumber="274" refId="ref14713" refString="Dumeril, A. M. C. & Bibron, G. (1836) Erpetologie generale, ou, Histoire naturelle complete des reptiles. Vol. 3. Roret, Paris, 531 pp." type="book" year="1836">Duméril & Bibron, 1836</bibRefCitation>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31C6538D949FFE96BC8FB13FBDAF9FB" pageId="7" pageNumber="274" type="materials_examined">
|
||
<paragraph id="8BB936B3D949FFE96BC8FB13FF01FAA3" blockId="7.[151,1437,1210,2029]" pageId="7" pageNumber="274">
|
||
<emphasis id="B972EAA1D949FFE96BC8FB13FE88FB36" bold="true" box="[151,353,1210,1237]" pageId="7" pageNumber="274">Type material.—</emphasis>
|
||
<materialsCitation id="3B6E3CEED949FFE96A3FFB13FE63FB1B" ID-GBIF-Occurrence="4015527303" collectingDate="1950-02" collectorName="Y. M. McCann" country="New Zealand" location="Middle Trios Island" pageId="7" pageNumber="274" specimenCount="1" stateProvince="Marlborough" typeStatus="holotype">
|
||
<emphasis id="B972EAA1D949FFE96A3FFB13FE2EFB37" box="[352,455,1211,1237]" italics="true" pageId="7" pageNumber="274">
|
||
<typeStatus id="54BD8811D949FFE96A3FFB13FE2EFB37" box="[352,455,1211,1237]" pageId="7" pageNumber="274" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<collectingCountry id="F3117623D949FFE96A8AFB13FD87FB37" box="[469,622,1211,1237]" name="New Zealand" pageId="7" pageNumber="274">New Zealand</collectingCountry>
|
||
,
|
||
<collectingRegion id="49C2F851D949FFE96927FB13FCE5FB37" box="[632,780,1211,1237]" country="New Zealand" name="Marlborough" pageId="7" pageNumber="274">Marlborough</collectingRegion>
|
||
Sounds,
|
||
<location id="8ED96068D949FFE9682CFB13FBB8FB37" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE9682CFB13FBB8FB37" box="[883,1105,1211,1237]" country="New Zealand" name="Middle Trios Island" pageId="7" pageNumber="274" stateProvince="Marlborough">Middle Trios Island</location>
|
||
, male,
|
||
<collectorName id="26F35365D949FFE96FC0FB13FAAEFB37" box="[1183,1351,1211,1237]" pageId="7" pageNumber="274">Y. M. McCann</collectorName>
|
||
,
|
||
<date id="FFB81073D949FFE96E0DFB13FF14FB1B" pageId="7" pageNumber="274" value="1950-02">
|
||
<collectingDate id="EFFCE99BD949FFE96E0DFB13FF14FB1B" pageId="7" pageNumber="274" value="1950-02">February 1950</collectingDate>
|
||
</date>
|
||
, RE.000503
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B6E3CEED949FFE96ACAFB77FF0DFAA3" ID-GBIF-Occurrence="4015527302" collectingDate="1988-11-22" collectorName="C. H. Daugherty" country="New Zealand" latitude="-40.842167" location="Middle Trios Island" longLatPrecision="11" longitude="174.0" municipality="Sounds" pageId="7" pageNumber="274" specimenCount="1" stateProvince="Marlborough" typeStatus="paratype">
|
||
<emphasis id="B972EAA1D949FFE96ACAFB77FDEEFB1A" box="[405,519,1247,1272]" italics="true" pageId="7" pageNumber="274">
|
||
<typeStatus id="54BD8811D949FFE96ACAFB77FDEEFB1A" box="[405,519,1247,1272]" pageId="7" pageNumber="274" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<collectingCountry id="F3117623D949FFE9694AFB77FD47FB1B" box="[533,686,1247,1273]" name="New Zealand" pageId="7" pageNumber="274">New Zealand</collectingCountry>
|
||
,
|
||
<collectingRegion id="49C2F851D949FFE969E6FB77FCA4FB1B" box="[697,845,1247,1273]" country="New Zealand" name="Marlborough" pageId="7" pageNumber="274">Marlborough</collectingRegion>
|
||
<collectingMunicipality id="6BDDACC9D949FFE9680BFB77FC40FB1B" box="[852,937,1247,1273]" pageId="7" pageNumber="274">Sounds</collectingMunicipality>
|
||
,
|
||
<location id="8ED96068D949FFE968EBFB77FB7DFB1B" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE968EBFB77FB7DFB1B" box="[948,1172,1247,1273]" country="New Zealand" latitude="-40.842167" longLatPrecision="11" longitude="174.0" municipality="Sounds" name="Middle Trios Island" pageId="7" pageNumber="274" stateProvince="Marlborough">Middle Trios Island</location>
|
||
,
|
||
<geoCoordinate id="EE325074D949FFE96FC0FB48FACAFB1B" box="[1183,1315,1247,1275]" degrees="40" direction="south" minutes="50.53" orientation="latitude" pageId="7" pageNumber="274" precision="9" value="-40.842167">40°50.53′ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE325074D949FFE96E6FFB48FF43FAFF" degrees="174" direction="east" minutes="0.00" orientation="longitude" pageId="7" pageNumber="274" precision="9" value="174.0">174°0.00′ E</geoCoordinate>
|
||
, both male,
|
||
<collectorName id="26F35365D949FFE96A6CFAABFE04FAFF" box="[307,493,1283,1309]" pageId="7" pageNumber="274">C. H. Daugherty</collectorName>
|
||
,
|
||
<date id="FFB81073D949FFE96AA6FAABFD3DFAFF" box="[505,724,1283,1309]" pageId="7" pageNumber="274" value="1988-11-22">
|
||
<collectingDate id="EFFCE99BD949FFE96AA6FAABFD3DFAFF" box="[505,724,1283,1309]" pageId="7" pageNumber="274" value="1988-11-22">22 November 1988</collectingDate>
|
||
</date>
|
||
, RE.006685, RE.006686 (tissue clips: FT2047, FT2048 respectively)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB936B3D949FFE96B98FAE2FBDAF9FB" blockId="7.[151,1437,1210,2029]" pageId="7" pageNumber="274">
|
||
<emphasis id="B972EAA1D949FFE96B98FAE2FE26FA86" bold="true" box="[199,463,1354,1380]" pageId="7" pageNumber="274">Material examined.—</emphasis>
|
||
<materialsCitation id="3B6E3CEED949FFE96A91FAE3FBEAFA87" ID-GBIF-Occurrence="4015527305" box="[462,1027,1354,1381]" collectionCode="OMVT" location="Stephens Island" municipality="The" pageId="7" pageNumber="274" specimenCode="OMVT949" specimenCount="1">
|
||
<collectingMunicipality id="6BDDACC9D949FFE96A91FAE3FE13FA87" box="[462,506,1355,1381]" pageId="7" pageNumber="274">The</collectingMunicipality>
|
||
<typeStatus id="54BD8811D949FFE9695CFAE3FDDAFA87" box="[515,563,1355,1381]" pageId="7" pageNumber="274">type</typeStatus>
|
||
material.
|
||
<emphasis id="B972EAA1D949FFE969F7FAE2FC8EFA87" bold="true" box="[680,871,1354,1381]" pageId="7" pageNumber="274">
|
||
<location id="8ED96068D949FFE969F7FAE2FC8EFA87" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE969F7FAE2FC8EFA87" box="[680,871,1354,1381]" municipality="The" name="Stephens Island" pageId="7" pageNumber="274">Stephens Island</location>
|
||
</emphasis>
|
||
(
|
||
<specimenCode id="DBA09EC8D949FFE96827FAE3FC12FA87" box="[888,1019,1355,1381]" collectionCode="OMVT" pageId="7" pageNumber="274">OMVT949</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B6E3CEED949FFE96F51FAE2FBC6F9FB" ID-GBIF-Occurrence="4015527304" collectionCode="OMVT" county="Trios Islands" location="South Trios Island" municipality="Middle Trios Island" pageId="7" pageNumber="274" specimenCode="OMVT719, OMVT807, OMVT3331, OMVT3332, OMVT3333" specimenCount="1" stateProvince="Canterbury">
|
||
<emphasis id="B972EAA1D949FFE96F51FAE2FB42FA87" bold="true" box="[1038,1195,1354,1381]" pageId="7" pageNumber="274">
|
||
<collectingCounty id="62D84E3FD949FFE96F51FAE2FB42FA87" box="[1038,1195,1354,1381]" pageId="7" pageNumber="274">Trios Islands</collectingCounty>
|
||
</emphasis>
|
||
:
|
||
<collectingMunicipality id="6BDDACC9D949FFE96FE4FAE3FA75FA87" box="[1211,1436,1355,1381]" pageId="7" pageNumber="274">Middle Trios Island</collectingMunicipality>
|
||
(RE.000505,FT2046);
|
||
<location id="8ED96068D949FFE96AC9FAC7FDB2FA6B" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE96AC9FAC7FDB2FA6B" box="[406,603,1391,1417]" county="Trios Islands" municipality="Middle Trios Island" name="South Trios Island" pageId="7" pageNumber="274" stateProvince="Canterbury">South Trios Island</location>
|
||
(RE.006687).
|
||
<emphasis id="B972EAA1D949FFE969A4FAC6FC75FA6B" bold="true" box="[763,924,1390,1417]" pageId="7" pageNumber="274">
|
||
<location id="8ED96068D949FFE969A4FAC6FC75FA6B" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE969A4FAC6FC75FA6B" box="[763,924,1390,1417]" county="Trios Islands" municipality="Middle Trios Island" name="Sentinel Rock" pageId="7" pageNumber="274" stateProvince="Canterbury">Sentinel Rock</location>
|
||
</emphasis>
|
||
(RE.000948).
|
||
<emphasis id="B972EAA1D949FFE96F64FAC6FAE8FA6B" bold="true" box="[1083,1281,1390,1417]" pageId="7" pageNumber="274">
|
||
<location id="8ED96068D949FFE96F64FAC6FAE8FA6B" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE96F64FAC6FAE8FA6B" box="[1083,1281,1390,1417]" county="Trios Islands" municipality="Middle Trios Island" name="Chetwode Island" pageId="7" pageNumber="274" stateProvince="Canterbury">Chetwode Island</location>
|
||
</emphasis>
|
||
(RE.000949).
|
||
<emphasis id="B972EAA1D949FFE96BC8FA3BFE83FA4F" bold="true" box="[151,362,1427,1453]" pageId="7" pageNumber="274">
|
||
<location id="8ED96068D949FFE96BC8FA3BFE8CFA4F" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE96BC8FA3BFE8CFA4F" box="[151,357,1427,1453]" county="Trios Islands" municipality="Middle Trios Island" name="Brothers Islands" pageId="7" pageNumber="274" stateProvince="Canterbury">Brothers Islands</location>
|
||
:
|
||
</emphasis>
|
||
<location id="8ED96068D949FFE96A2BFA3BFD8DFA4F" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE96A2BFA3BFD8DFA4F" box="[372,612,1427,1453]" county="Trios Islands" municipality="Middle Trios Island" name="North Brother Island" pageId="7" pageNumber="274" stateProvince="Canterbury">North Brother Island</location>
|
||
(RE.002561, RE.005496, RE.006509, RE.006510, RE.007265, FT277, FT278).
|
||
<emphasis id="B972EAA1D949FFE96BA6FA1FFD4EFA32" bold="true" box="[249,679,1462,1489]" pageId="7" pageNumber="274">
|
||
<location id="8ED96068D949FFE96BA6FA1FFDFAFA33" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE96BA6FA1FFDFAFA33" box="[249,531,1462,1489]" county="Trios Islands" municipality="Middle Trios Island" name="Northwest South Island" pageId="7" pageNumber="274" stateProvince="Canterbury">Northwest South Island</location>
|
||
(mainland):
|
||
</emphasis>
|
||
<location id="8ED96068D949FFE969F1FA1FFC8EFA33" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE969F1FA1FFC8EFA33" box="[686,871,1463,1489]" county="Trios Islands" municipality="Middle Trios Island" name="Gouland Downs" pageId="7" pageNumber="274" stateProvince="Canterbury">Gouland Downs</location>
|
||
,
|
||
<location id="8ED96068D949FFE9682DFA1FFC34FA33" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE9682DFA1FFC34FA33" box="[882,989,1463,1489]" county="Trios Islands" municipality="Middle Trios Island" name="Holocene" pageId="7" pageNumber="274" stateProvince="Canterbury">Holocene</location>
|
||
fossil (S.38813.2).
|
||
<emphasis id="B972EAA1D949FFE96FE6FA1EFF37FA16" bold="true" pageId="7" pageNumber="274">
|
||
<collectingRegion id="49C2F851D949FFE96FE6FA1EFAAEFA32" box="[1209,1351,1462,1488]" country="New Zealand" name="Canterbury" pageId="7" pageNumber="274">Canterbury</collectingRegion>
|
||
(mainland):
|
||
</emphasis>
|
||
<location id="8ED96068D949FFE96BBBFA73FED6FA17" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE96BBBFA73FED6FA17" box="[228,319,1499,1525]" county="Trios Islands" municipality="Middle Trios Island" name="Waikari" pageId="7" pageNumber="274" stateProvince="Canterbury">Waikari</location>
|
||
,
|
||
<location id="8ED96068D949FFE96A17FA73FE5AFA17" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE96A17FA73FE5AFA17" box="[328,435,1499,1525]" county="Trios Islands" municipality="Middle Trios Island" name="Holocene" pageId="7" pageNumber="274" stateProvince="Canterbury">Holocene</location>
|
||
fossil (S.33501, S.33703.1, S.33703.7, S.33703.10, S.33703.11);
|
||
<location id="8ED96068D949FFE96FD3FA73FB0CFA17" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE96FD3FA73FB0CFA17" box="[1164,1253,1499,1525]" county="Trios Islands" municipality="Middle Trios Island" name="Waitaki" pageId="7" pageNumber="274" stateProvince="Canterbury">Waitaki</location>
|
||
,
|
||
<location id="8ED96068D949FFE96FB0FA73FAB3FA17" LSID="urn:lsid:plazi:treatment:03AF87A5D949FFE26BC8FE8EFED9FB0B:8ED96068D949FFE96FB0FA73FAB3FA17" box="[1263,1370,1499,1525]" county="Trios Islands" municipality="Middle Trios Island" name="Holocene" pageId="7" pageNumber="274" stateProvince="Canterbury">Holocene</location>
|
||
fossil (
|
||
<specimenCode id="DBA09EC8D949FFE96BC0FA57FEC9F9FB" box="[159,288,1535,1561]" collectionCode="OMVT" pageId="7" pageNumber="274">OMVT719</specimenCode>
|
||
a,
|
||
<specimenCode id="DBA09EC8D949FFE96A68FA57FE5EF9FB" box="[311,439,1535,1561]" collectionCode="OMVT" pageId="7" pageNumber="274">OMVT807</specimenCode>
|
||
a,
|
||
<specimenCode id="DBA09EC8D949FFE96A91FA57FDA7F9FB" box="[462,590,1535,1561]" collectionCode="OMVT" pageId="7" pageNumber="274">OMVT807</specimenCode>
|
||
b,
|
||
<specimenCode id="DBA09EC8D949FFE96938FA57FD1AF9FB" box="[615,755,1535,1561]" collectionCode="OMVT" pageId="7" pageNumber="274">OMVT3331</specimenCode>
|
||
,
|
||
<specimenCode id="DBA09EC8D949FFE969A0FA57FC65F9FB" box="[767,908,1535,1561]" collectionCode="OMVT" pageId="7" pageNumber="274">OMVT3332</specimenCode>
|
||
,
|
||
<specimenCode id="DBA09EC8D949FFE968C7FA57FBCEF9FB" box="[920,1063,1535,1561]" collectionCode="OMVT" pageId="7" pageNumber="274">OMVT3333</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31C6538D949FFE96B98F98AFC87F80F" pageId="7" pageNumber="274" type="diagnosis">
|
||
<paragraph id="8BB936B3D949FFE96B98F98AFC87F80F" blockId="7.[151,1437,1210,2029]" pageId="7" pageNumber="274">
|
||
<emphasis id="B972EAA1D949FFE96B98F98AFEB4F9DE" bold="true" box="[199,349,1570,1596]" pageId="7" pageNumber="274">Diagnosis.—</emphasis>
|
||
<taxonomicName id="4C064D30D949FFE96A02F98BFDD3F9DF" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[349,570,1571,1597]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="tohu">
|
||
<emphasis id="B972EAA1D949FFE96A02F98BFDD3F9DF" box="[349,570,1571,1597]" italics="true" pageId="7" pageNumber="274">Hoplodactylus tohu</emphasis>
|
||
</taxonomicName>
|
||
may be distinguished from its only congener,
|
||
<taxonomicName id="4C064D30D949FFE96F1BF98BFB32F9DF" authorityName="Jewell" authorityYear="2008" box="[1092,1243,1571,1597]" class="Reptilia" family="Diplodactylidae" genus="Woodworthia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D949FFE96F1BF98BFB32F9DF" box="[1092,1243,1571,1597]" italics="true" pageId="7" pageNumber="274">H. duvaucelii</emphasis>
|
||
</taxonomicName>
|
||
, by several characters:
|
||
<taxonomicName id="4C064D30D949FFE96BB7F9EFFED4F983" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[232,317,1607,1633]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="tohu">
|
||
<emphasis id="B972EAA1D949FFE96BB7F9EFFED4F983" box="[232,317,1607,1633]" italics="true" pageId="7" pageNumber="274">H. tohu</emphasis>
|
||
</taxonomicName>
|
||
(generally) does not attain as great a size at maturity (Supp.
|
||
<figureCitation id="133D2A36D949FFE968B0F9EFFBD9F983" box="[1007,1072,1607,1633]" captionStart="FIGURE 1" captionStartId="1.[151,250,1679,1703]" captionTargetBox="[159,1427,179,1653]" captionTargetId="figure-20@1.[159,1427,179,1653]" captionTargetPageId="1" captionText="FIGURE 1. Maps of New Zealand showing distribution of Hoplodactylus duvaucelii (Duméril & Bibron 1836) (left) and H. tohu n. sp. (right), with inter- and intraspecific phylogenetic distance highlighted in a simplified mitochondrial genome phylogeny (bottom; adapted from Scarsbrook et al. 2022). Extant relict (pink) and translocated (purple) populations, historic (yellow) and Holocene subfossil records (blue) are indicated (Supp. Tables 1–2), with triangles denoting localities included in genetic analyses (see phylogeny). Abbreviations for referenced localities are as follows: Cape Maria van Dieman (CM), Poor Knights Islands (PK), Hen and Chickens Islands (HC), Great Barrier Island (GB), Maungatautari Ecosanctuary (ME) and Mana Island (MI)." figureDoi="http://doi.org/10.5281/zenodo.7532707" httpUri="https://zenodo.org/record/7532707/files/figure.png" pageId="7" pageNumber="274">Fig. 1</figureCitation>
|
||
), has a more pronounced brillar fold (
|
||
<figureCitation id="133D2A36D949FFE96B8CF9C3FE6EF967" box="[211,391,1643,1669]" captionStart="FIGURE 3" captionStartId="9.[151,250,815,839]" captionTargetBox="[151,1435,185,785]" captionTargetId="figure-20@9.[151,1435,181,791]" captionTargetPageId="9" captionText="FIGURE 3. Details of: A-D, Hoplodactylus tohu n. sp., holotype (RE.000503); and E-H, Hoplodactylus duvaucelii Duméril & Bibron, 1836, topotype (RE.003491): A, E, right-lateral view of head; B, F, oblique right-lateroventral view of head, with diagnostic characters (a) pronounced brillar fold, and (b) abrupt size decrease at the 5th infralabial scale; C, G, ventral view of the right manus showing subdigital lamellae, and highlighting the less emergent claw on the first digit in H. duvaucelii; D, H, ventral view of the right pes showing subdigital lamellae. Automontage images generated by Jean-Claude Stahl (NMNZ). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7532713" httpUri="https://zenodo.org/record/7532713/files/figure.png" pageId="7" pageNumber="274">Fig. 3A–B, E–F</figureCitation>
|
||
,
|
||
<figureCitation id="133D2A36D949FFE96ACDF9C3FE3BF967" box="[402,466,1643,1669]" captionStart="FIGURE 4" captionStartId="10.[151,250,1519,1543]" captionTargetBox="[151,1435,181,1496]" captionTargetId="figure-26@10.[151,1435,181,1496]" captionTargetPageId="10" captionText="FIGURE 4. Living individuals of Hoplodactylus species. A–C, Hoplodactylus tohu n. sp.; D–F, H. duvaucelii (Duméril & Bibron 1836). A, Brothers Islands (Cook Strait); B–C, Sentinel Rock (Marlborough Sounds). D–F, Korapuki Island (Mercury Islands). The first and second rows show sub-adult and adult representatives of each species respectively. Photographs: Nicholas Harker (Auckland, New Zealand)." figureDoi="http://doi.org/10.5281/zenodo.7532715" httpUri="https://zenodo.org/record/7532715/files/figure.png" pageId="7" pageNumber="274">4C, F</figureCitation>
|
||
), and less often bears a median cleft in the mental scale (Supp. Table 3). An abrupt size decrease at the 5
|
||
<superScript id="7C739BFBD949FFE96A0DF926FEB6F97E" attach="left" box="[338,351,1678,1692]" fontSize="6" pageId="7" pageNumber="274">th</superScript>
|
||
infralabial scale characterizing
|
||
<taxonomicName id="4C064D30D949FFE9699AF927FCF3F94B" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[709,794,1679,1705]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="tohu">
|
||
<emphasis id="B972EAA1D949FFE9699AF927FCF3F94B" box="[709,794,1679,1705]" italics="true" pageId="7" pageNumber="274">H. tohu</emphasis>
|
||
</taxonomicName>
|
||
is far less common in
|
||
<taxonomicName id="4C064D30D949FFE96F46F927FB58F94B" authorityName="Jewell" authorityYear="2008" box="[1049,1201,1679,1705]" class="Reptilia" family="Diplodactylidae" genus="Woodworthia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D949FFE96F46F927FB58F94B" box="[1049,1201,1679,1705]" italics="true" pageId="7" pageNumber="274">H. duvaucelii</emphasis>
|
||
</taxonomicName>
|
||
, with most individuals of the latter examined exhibiting a gradual size decrease in infralabial scales (
|
||
<figureCitation id="133D2A36D949FFE96F41F91BFB7AF92F" box="[1054,1171,1715,1741]" captionStart="FIGURE 3" captionStartId="9.[151,250,815,839]" captionTargetBox="[151,1435,185,785]" captionTargetId="figure-20@9.[151,1435,181,791]" captionTargetPageId="9" captionText="FIGURE 3. Details of: A-D, Hoplodactylus tohu n. sp., holotype (RE.000503); and E-H, Hoplodactylus duvaucelii Duméril & Bibron, 1836, topotype (RE.003491): A, E, right-lateral view of head; B, F, oblique right-lateroventral view of head, with diagnostic characters (a) pronounced brillar fold, and (b) abrupt size decrease at the 5th infralabial scale; C, G, ventral view of the right manus showing subdigital lamellae, and highlighting the less emergent claw on the first digit in H. duvaucelii; D, H, ventral view of the right pes showing subdigital lamellae. Automontage images generated by Jean-Claude Stahl (NMNZ). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7532713" httpUri="https://zenodo.org/record/7532713/files/figure.png" pageId="7" pageNumber="274">Fig. 3B, F</figureCitation>
|
||
).
|
||
<taxonomicName id="4C064D30D949FFE96FF8F91BFB13F92F" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[1191,1274,1715,1741]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="tohu">
|
||
<emphasis id="B972EAA1D949FFE96FF8F91BFB13F92F" box="[1191,1274,1715,1741]" italics="true" pageId="7" pageNumber="274">H. tohu</emphasis>
|
||
</taxonomicName>
|
||
further differs from
|
||
<taxonomicName id="4C064D30D949FFE96B8AF97FFE85F913" authorityName="Jewell" authorityYear="2008" box="[213,364,1751,1777]" class="Reptilia" family="Diplodactylidae" genus="Woodworthia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D949FFE96B8AF97FFE85F913" box="[213,364,1751,1777]" italics="true" pageId="7" pageNumber="274">H. duvaucelii</emphasis>
|
||
</taxonomicName>
|
||
in generally having fewer subdigital lamellae on all digits of both the right manus and pes (
|
||
<figureCitation id="133D2A36D949FFE96E30F97FFECBF8F7" captionStart="FIGURE 3" captionStartId="9.[151,250,815,839]" captionTargetBox="[151,1435,185,785]" captionTargetId="figure-20@9.[151,1435,181,791]" captionTargetPageId="9" captionText="FIGURE 3. Details of: A-D, Hoplodactylus tohu n. sp., holotype (RE.000503); and E-H, Hoplodactylus duvaucelii Duméril & Bibron, 1836, topotype (RE.003491): A, E, right-lateral view of head; B, F, oblique right-lateroventral view of head, with diagnostic characters (a) pronounced brillar fold, and (b) abrupt size decrease at the 5th infralabial scale; C, G, ventral view of the right manus showing subdigital lamellae, and highlighting the less emergent claw on the first digit in H. duvaucelii; D, H, ventral view of the right pes showing subdigital lamellae. Automontage images generated by Jean-Claude Stahl (NMNZ). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7532713" httpUri="https://zenodo.org/record/7532713/files/figure.png" pageId="7" pageNumber="274">Fig. 3C–D, G–H</figureCitation>
|
||
, Supp.
|
||
<figureCitation id="133D2A36D949FFE96A27F953FE55F8F7" box="[376,444,1787,1813]" captionStart="FIGURE 3" captionStartId="9.[151,250,815,839]" captionTargetBox="[151,1435,185,785]" captionTargetId="figure-20@9.[151,1435,181,791]" captionTargetPageId="9" captionText="FIGURE 3. Details of: A-D, Hoplodactylus tohu n. sp., holotype (RE.000503); and E-H, Hoplodactylus duvaucelii Duméril & Bibron, 1836, topotype (RE.003491): A, E, right-lateral view of head; B, F, oblique right-lateroventral view of head, with diagnostic characters (a) pronounced brillar fold, and (b) abrupt size decrease at the 5th infralabial scale; C, G, ventral view of the right manus showing subdigital lamellae, and highlighting the less emergent claw on the first digit in H. duvaucelii; D, H, ventral view of the right pes showing subdigital lamellae. Automontage images generated by Jean-Claude Stahl (NMNZ). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7532713" httpUri="https://zenodo.org/record/7532713/files/figure.png" pageId="7" pageNumber="274">Fig. 3</figureCitation>
|
||
). The first digit of the right manus in
|
||
<taxonomicName id="4C064D30D949FFE96832F953FBECF8F7" authorityName="Jewell" authorityYear="2008" box="[877,1029,1787,1813]" class="Reptilia" family="Diplodactylidae" genus="Woodworthia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D949FFE96832F953FBECF8F7" box="[877,1029,1787,1813]" italics="true" pageId="7" pageNumber="274">H. duvaucelii</emphasis>
|
||
</taxonomicName>
|
||
differs in having a consistently less emergent claw and, usually, a comparatively bulbous distal end (
|
||
<figureCitation id="133D2A36D949FFE96837F8B7FC0BF8DB" box="[872,994,1823,1849]" captionStart="FIGURE 3" captionStartId="9.[151,250,815,839]" captionTargetBox="[151,1435,185,785]" captionTargetId="figure-20@9.[151,1435,181,791]" captionTargetPageId="9" captionText="FIGURE 3. Details of: A-D, Hoplodactylus tohu n. sp., holotype (RE.000503); and E-H, Hoplodactylus duvaucelii Duméril & Bibron, 1836, topotype (RE.003491): A, E, right-lateral view of head; B, F, oblique right-lateroventral view of head, with diagnostic characters (a) pronounced brillar fold, and (b) abrupt size decrease at the 5th infralabial scale; C, G, ventral view of the right manus showing subdigital lamellae, and highlighting the less emergent claw on the first digit in H. duvaucelii; D, H, ventral view of the right pes showing subdigital lamellae. Automontage images generated by Jean-Claude Stahl (NMNZ). Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7532713" httpUri="https://zenodo.org/record/7532713/files/figure.png" pageId="7" pageNumber="274">Fig. 3C, G</figureCitation>
|
||
). Previously reported differences (
|
||
<bibRefCitation id="EF974B42D949FFE96E3DF8B7FE47F8BF" author="Morgan-Richards, M. & Hinlo, A. R. & Smuts-Kennedy, C. & Innes, J. & Ji, W. & Barry, M. & Brunton, D. & Hitchmough, R. A." pageId="7" pageNumber="274" pagination="77 - 86" refId="ref20012" refString="Morgan-Richards, M., Hinlo, A. R., Smuts-Kennedy, C., Innes, J., Ji, W., Barry, M., Brunton, D. & Hitchmough, R. A. (2016) Identification of a rare gecko from North Island New Zealand, and genetic assessment of its probable origin: A novel mainland conservation priority? Journal of Herpetology, 50, 77 - 86. https: // doi. org / 10.1670 / 13 - 128" type="journal article" year="2016">
|
||
Morgan-Richards
|
||
<emphasis id="B972EAA1D949FFE96A6BF8EBFE84F8BF" box="[308,365,1859,1885]" italics="true" pageId="7" pageNumber="274">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
) in the coloration and extent of patterning between these taxa (
|
||
<figureCitation id="133D2A36D949FFE96F2AF8EBFB50F8BF" box="[1141,1209,1859,1885]" captionStart="FIGURE 4" captionStartId="10.[151,250,1519,1543]" captionTargetBox="[151,1435,181,1496]" captionTargetId="figure-26@10.[151,1435,181,1496]" captionTargetPageId="10" captionText="FIGURE 4. Living individuals of Hoplodactylus species. A–C, Hoplodactylus tohu n. sp.; D–F, H. duvaucelii (Duméril & Bibron 1836). A, Brothers Islands (Cook Strait); B–C, Sentinel Rock (Marlborough Sounds). D–F, Korapuki Island (Mercury Islands). The first and second rows show sub-adult and adult representatives of each species respectively. Photographs: Nicholas Harker (Auckland, New Zealand)." figureDoi="http://doi.org/10.5281/zenodo.7532715" httpUri="https://zenodo.org/record/7532715/files/figure.png" pageId="7" pageNumber="274">Fig. 4</figureCitation>
|
||
) are generalizations and can be misleading given fluid overlap between the two species. Dorsal body coloration and patterning in
|
||
<taxonomicName id="4C064D30D949FFE96E16F8CFFA75F863" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[1353,1436,1895,1921]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="tohu">
|
||
<emphasis id="B972EAA1D949FFE96E16F8CFFA75F863" box="[1353,1436,1895,1921]" italics="true" pageId="7" pageNumber="274">H. tohu</emphasis>
|
||
</taxonomicName>
|
||
often resembles those of young
|
||
<taxonomicName id="4C064D30D949FFE96AA3F823FD7AF847" authorityName="Jewell" authorityYear="2008" box="[508,659,1931,1957]" class="Reptilia" family="Diplodactylidae" genus="Woodworthia" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="274" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D949FFE96AA3F823FD7AF847" box="[508,659,1931,1957]" italics="true" pageId="7" pageNumber="274">H. duvaucelii</emphasis>
|
||
</taxonomicName>
|
||
in being relatively more strongly contrasting (
|
||
<figureCitation id="133D2A36D949FFE96FC4F823FABEF847" box="[1179,1367,1931,1957]" captionStart="FIGURE 4" captionStartId="10.[151,250,1519,1543]" captionTargetBox="[151,1435,181,1496]" captionTargetId="figure-26@10.[151,1435,181,1496]" captionTargetPageId="10" captionText="FIGURE 4. Living individuals of Hoplodactylus species. A–C, Hoplodactylus tohu n. sp.; D–F, H. duvaucelii (Duméril & Bibron 1836). A, Brothers Islands (Cook Strait); B–C, Sentinel Rock (Marlborough Sounds). D–F, Korapuki Island (Mercury Islands). The first and second rows show sub-adult and adult representatives of each species respectively. Photographs: Nicholas Harker (Auckland, New Zealand)." figureDoi="http://doi.org/10.5281/zenodo.7532715" httpUri="https://zenodo.org/record/7532715/files/figure.png" pageId="7" pageNumber="274">Fig. 4A–B, D–E</figureCitation>
|
||
), with patterning generally becoming weaker at maturity. However, considerable variation in coloration and pattern (see Supp. Fig. 5) was observed throughout ontogeny in both species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF79663BD946FFE66BC8F8FDFEB7F83B" ID-DOI="http://doi.org/10.5281/zenodo.7532711" ID-Zenodo-Dep="7532711" httpUri="https://zenodo.org/record/7532711/files/figure.png" pageId="8" pageNumber="275" startId="8.[151,250,1877,1901]" targetBox="[163,1423,191,1843]" targetPageId="8">
|
||
<paragraph id="8BB936B3D946FFE66BC8F8FDFEB7F83B" blockId="8.[151,1437,1877,2010]" pageId="8" pageNumber="275">
|
||
<emphasis id="B972EAA1D946FFE66BC8F8FDFEFBF88F" bold="true" box="[151,274,1877,1901]" pageId="8" pageNumber="275">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C064D30D946FFE66A48F8FEFD90F88C" box="[279,633,1878,1902]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="275" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B972EAA1D946FFE66A48F8FEFE42F88C" box="[279,427,1878,1902]" italics="true" pageId="8" pageNumber="275">Hoplodactylus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF974B42D946FFE66AE8F8FEFDBAF88C" author="Fitzinger, L." box="[439,595,1878,1902]" pageId="8" pageNumber="275" refId="ref15174" refString="Fitzinger, L. (1843) Systema reptilium, fasciculus primus: Amblyglossae. Braumuller et Seidel, Vindobonae, 106 pp. https: // doi. org / 10.5962 / bhl. title. 4694" type="book" year="1843">Fitzinger, 1843</bibRefCitation>
|
||
) sp.
|
||
</taxonomicName>
|
||
—dorsal and ventral body views (left-right).
|
||
<emphasis id="B972EAA1D946FFE66F62F8FDFBBCF88F" bold="true" box="[1085,1109,1877,1901]" pageId="8" pageNumber="275">A,</emphasis>
|
||
<taxonomicName id="4C064D30D946FFE66F04F8FEFAC8F88C" authority="Scarsbrook & Walton & Rawlence & Hitchmough, 2023" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[1115,1313,1878,1902]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="275" phylum="Chordata" rank="species" species="tohu" status="sp. nov.">
|
||
<emphasis id="B972EAA1D946FFE66F04F8FEFAC8F88C" box="[1115,1313,1878,1902]" italics="true" pageId="8" pageNumber="275">Hoplodactylus tohu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24157DAD946FFE66E79F8FEFAB4F88F" box="[1318,1373,1878,1901]" pageId="8" pageNumber="275" rank="species">n. sp.</taxonomicNameLabel>
|
||
, holotype, Middle Trios Island, Marlborough Sounds, RE.000503.
|
||
<emphasis id="B972EAA1D946FFE66854F8D2FCC8F870" bold="true" box="[779,801,1914,1938]" pageId="8" pageNumber="275">B,</emphasis>
|
||
<taxonomicName id="4C064D30D946FFE66876F8D2FADBF873" authority="Dumeril & Bibron, 1836" authorityName="Dumeril & Bibron" authorityYear="1836" box="[809,1330,1913,1938]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="275" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D946FFE66876F8D2FBC4F870" box="[809,1069,1914,1938]" italics="true" pageId="8" pageNumber="275">Hoplodactylus duvaucelii</emphasis>
|
||
<bibRefCitation id="EF974B42D946FFE66F6BF8D2FADBF873" author="Dumeril, A. M. C. & Bibron, G." box="[1076,1330,1913,1938]" pageId="8" pageNumber="275" refId="ref14713" refString="Dumeril, A. M. C. & Bibron, G. (1836) Erpetologie generale, ou, Histoire naturelle complete des reptiles. Vol. 3. Roret, Paris, 531 pp." type="book" year="1836">Duméril & Bibron, 1836</bibRefCitation>
|
||
</taxonomicName>
|
||
, topotype, Muriwhenua Island, Hen and Chickens Islands, RE.003491. Automontage images generated by Jean-Claude Stahl (NMNZ). Scale bar = 20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF79663BD947FFE76BC8FC87FEC8FC19" ID-DOI="http://doi.org/10.5281/zenodo.7532713" ID-Zenodo-Dep="7532713" httpUri="https://zenodo.org/record/7532713/files/figure.png" pageId="9" pageNumber="276" startId="9.[151,250,815,839]" targetBox="[151,1435,185,785]" targetPageId="9">
|
||
<paragraph id="8BB936B3D947FFE76BC8FC87FEC8FC19" blockId="9.[151,1436,815,1020]" pageId="9" pageNumber="276">
|
||
<emphasis id="B972EAA1D947FFE76BC8FC87FEFDFCA5" bold="true" box="[151,276,815,839]" pageId="9" pageNumber="276">FIGURE 3.</emphasis>
|
||
Details of:
|
||
<emphasis id="B972EAA1D947FFE76ACFFC87FE2AFCA5" bold="true" box="[400,451,815,839]" pageId="9" pageNumber="276">A-D,</emphasis>
|
||
<taxonomicName id="4C064D30D947FFE76A94FC98FD7AFCAA" authority="Scarsbrook & Walton & Rawlence & Hitchmough, 2023" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[459,659,816,840]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="276" phylum="Chordata" rank="species" species="tohu" status="sp. nov.">
|
||
<emphasis id="B972EAA1D947FFE76A94FC98FD7AFCAA" box="[459,659,816,840]" italics="true" pageId="9" pageNumber="276">Hoplodactylus tohu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24157DAD947FFE769C4FC98FD3CFCA5" box="[667,725,816,839]" pageId="9" pageNumber="276" rank="species">n. sp.</taxonomicNameLabel>
|
||
, holotype (RE.000503); and
|
||
<emphasis id="B972EAA1D947FFE76F5FFC98FBDAFCAA" bold="true" box="[1024,1075,816,840]" pageId="9" pageNumber="276">E-H,</emphasis>
|
||
<taxonomicName id="4C064D30D947FFE76F64FC98FED5FC89" authority="Dumeril & Bibron, 1836" authorityName="Dumeril & Bibron" authorityYear="1836" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="276" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D947FFE76F64FC98FAA8FCAA" box="[1083,1345,816,840]" italics="true" pageId="9" pageNumber="276">Hoplodactylus duvaucelii</emphasis>
|
||
<bibRefCitation id="EF974B42D947FFE76E17FC98FED5FC89" author="Dumeril, A. M. C. & Bibron, G." pageId="9" pageNumber="276" refId="ref14713" refString="Dumeril, A. M. C. & Bibron, G. (1836) Erpetologie generale, ou, Histoire naturelle complete des reptiles. Vol. 3. Roret, Paris, 531 pp." type="book" year="1836">Duméril & Bibron, 1836</bibRefCitation>
|
||
</taxonomicName>
|
||
, topotype (RE.003491):
|
||
<emphasis id="B972EAA1D947FFE76964FCFBFD9BFC8E" bold="true" box="[571,626,851,876]" pageId="9" pageNumber="276">A, E,</emphasis>
|
||
right-lateral view of head;
|
||
<emphasis id="B972EAA1D947FFE768D1FCFCFC29FC8E" bold="true" box="[910,960,852,876]" pageId="9" pageNumber="276">B, F,</emphasis>
|
||
oblique right-lateroventral view of head, with diagnostic characters (a) pronounced brillar fold, and (b) abrupt size decrease at the 5
|
||
<superScript id="7C739BFBD947FFE768A0FCD0FBE3FC64" attach="left" box="[1023,1034,888,902]" fontSize="6" pageId="9" pageNumber="276">th</superScript>
|
||
infralabial scale;
|
||
<emphasis id="B972EAA1D947FFE76F9EFCDFFB12FC6D" bold="true" box="[1217,1275,887,911]" pageId="9" pageNumber="276">C, G,</emphasis>
|
||
ventral view of the right manus showing subdigital lamellae, and highlighting the less emergent claw on the first digit in
|
||
<taxonomicName id="4C064D30D947FFE76F94FC34FABCFC56" authorityName="Dumeril & Bibron" authorityYear="1836" box="[1227,1365,924,948]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="276" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D947FFE76F94FC34FABCFC56" box="[1227,1365,924,948]" italics="true" pageId="9" pageNumber="276">H. duvaucelii</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B972EAA1D947FFE76E3CFC34FA75FC56" bold="true" box="[1379,1436,924,948]" pageId="9" pageNumber="276">D, H,</emphasis>
|
||
ventral view of the right pes showing subdigital lamellae. Automontage images generated by Jean-Claude Stahl (NMNZ). Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31C6538D947FFE56B98FB81FAEDFC0F" lastPageId="11" lastPageNumber="278" pageId="9" pageNumber="276" type="description">
|
||
<paragraph id="8BB936B3D947FFE56B98FB81FAEDFC0F" blockId="9.[151,1437,1065,2028]" lastBlockId="11.[151,1437,151,2013]" lastPageId="11" lastPageNumber="278" pageId="9" pageNumber="276">
|
||
<emphasis id="B972EAA1D947FFE76B98FB81FE9DFBA1" bold="true" box="[199,372,1065,1091]" pageId="9" pageNumber="276">Description.—</emphasis>
|
||
Coloration on dorsal surface grey to grey-brown, sometimes with olive and dark brown blotches; patterning of well-developed, roughly symmetrical transverse bands from nape to tail base centered along spine, resembling chevrons or paired diamonds, less defined on tail and generally less defined on older individuals; often bearing irregular series of longitudinal rows of pale grey or white spots dorsolaterally, extending onto limbs; ventral surface buff, with occasional light brown speckling, speckles less frequent on head. Mouth lining and tongue pink. Body moderately large (SVL:
|
||
<quantity id="4CFE9B56D947FFE76AB8FB76FD73FB1A" box="[487,666,1246,1273]" metricMagnitude="-2" metricUnit="m" metricValue="9.845" metricValueMax="11.57" metricValueMin="8.12" pageId="9" pageNumber="276" unit="mm" value="98.45" valueMax="115.7" valueMin="81.2">81.2–115.7 mm</quantity>
|
||
), robust, stout (TrK:
|
||
<quantity id="4CFE9B56D947FFE768DFFB76FBCFFB1A" box="[896,1062,1246,1273]" metricMagnitude="-2" metricUnit="m" metricValue="3.7849999999999993" metricValueMax="5.029999999999999" metricValueMin="2.54" pageId="9" pageNumber="276" unit="mm" value="37.849999999999994" valueMax="50.3" valueMin="25.4">25.4–50.3 mm</quantity>
|
||
). Head large (HL:
|
||
<quantity id="4CFE9B56D947FFE76FACFB76FA71FB1A" box="[1267,1432,1246,1272]" metricMagnitude="-2" metricUnit="m" metricValue="2.995" metricValueMax="3.38" metricValueMin="2.61" pageId="9" pageNumber="276" unit="mm" value="29.95" valueMax="33.8" valueMin="26.1">26.1–33.8 mm</quantity>
|
||
; HW:
|
||
<quantity id="4CFE9B56D947FFE76B8BFAAAFE8FFAFE" box="[212,358,1282,1308]" metricMagnitude="-2" metricUnit="m" metricValue="2.2649999999999997" metricValueMax="2.7" metricValueMin="1.83" pageId="9" pageNumber="276" unit="mm" value="22.65" valueMax="27.0" valueMin="18.3">18.3–27 mm</quantity>
|
||
; HH:
|
||
<quantity id="4CFE9B56D947FFE76AF4FAAAFDD4FAFE" box="[427,573,1282,1308]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" metricValueMax="1.6" metricValueMin="1.22" pageId="9" pageNumber="276" unit="mm" value="14.1" valueMax="16.0" valueMin="12.2">12.2–16 mm</quantity>
|
||
), robust, subtriangular; inflated laterally between posterior edge of orbit and ear opening (EE:
|
||
<quantity id="4CFE9B56D947FFE76A00FA8EFE11FAA2" box="[351,504,1318,1344]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" metricValueMax="11.8" metricValueMin="7.7" pageId="9" pageNumber="276" unit="mm" value="9.75" valueMax="11.8" valueMin="7.7">7.7–11.8 mm</quantity>
|
||
), narrowing towards craniovertebral junction; neck clearly demarcated. Snout oviform (EN:
|
||
<quantity id="4CFE9B56D947FFE76A4DFAE2FE43FA86" box="[274,426,1354,1380]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="11.1" metricValueMin="7.4" pageId="9" pageNumber="276" unit="mm" value="9.25" valueMax="11.1" valueMin="7.4">7.4–11.1 mm</quantity>
|
||
); slight indentation in medial nasal region, often blunt along anteriormost margin between nares (IN:
|
||
<quantity id="4CFE9B56D947FFE76A4CFAC6FE62FA6A" box="[275,395,1390,1417]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" metricValueMax="5.8" metricValueMin="4.0" pageId="9" pageNumber="276" unit="mm" value="4.9" valueMax="5.8" valueMin="4.0">4–5.8 mm</quantity>
|
||
). Dorsum of occiput/nape covered in small granular scales that abruptly increase in size at level of anterior edge of orbit towards anterior margin of snout (2–3 times diameter of occipital granules); one row of enlarged, oval scales posterior to internasal(s) and dorsal to supralabials, broader than high; twice the diameter of adjacent loreal scales. Eyes approximately one fifth head length; varying in shade from pale olive-buff to dark greens or browns; pupils lenticular with weakly crenulated margin. Supraciliary scales elongate, conical, directed increasingly posterior posteriorly; brillar fold very pronounced; frontal narrowing anteriorly (IO:
|
||
<quantity id="4CFE9B56D947FFE76FA9F98AFA79F9DE" box="[1270,1424,1570,1596]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" metricValueMax="11.8" metricValueMin="7.8" pageId="9" pageNumber="276" unit="mm" value="9.8" valueMax="11.8" valueMin="7.8">7.8–11.8 mm</quantity>
|
||
), supraocular portion deeply furrowed. Ear opening moderate (EL:
|
||
<quantity id="4CFE9B56D947FFE7682DF9EEFC0EF982" box="[882,999,1606,1632]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="4.1" metricValueMin="2.0" pageId="9" pageNumber="276" unit="mm" value="3.05" valueMax="4.1" valueMin="2.0">2–4.1 mm</quantity>
|
||
), ovoid, twice as high as wide; oriented obliquely (widest posterodorsally to anteroventrally), skin fold covering dorsal limit. Rostral subpentagonal, much broader than high; contacted dorsally by 2 enlarged, oval supranasals and 1–3 smaller (homogenous), round internasals; medial rostral crease extending ventrally from upper margin ½ to ¾ length of rostral; usually terminating diffusely, but sometimes as an ovoid crease. Nares rounded, situated anterolaterally; bordered by rostral, supranasal, 3–5 small postnasals and first supralabial. Supralabials rectangular, rounded dorsally; slightly higher than broad; numbering 13–16 per side; gradually decreasing in size posteriorly; final 3–4 more narrowly convex; supralabials terminating beneath the posterior orbital margin (with 11 or 12 beneath orbit midline); posteriormost twice the size of loreal scales. Mental trapezoidal to subtriangular, with longest face along jaw margin, narrowing posteriorly; mental crease usually absent, extending ½ length of scale where present; mental shorter than laterally adjacent first infralabials; contacted posteriorly by 1 large or 2 smaller hexagonal postmentals, which separate infralabials. Infralabials numbering 9–14 per side; from the snout, infralabials 1–4 on each side are quadrate, more rounded ventrally, higher than broad; infralabial 5 usually marks commencement of a significant and discrete infralabial size decrease, sometimes on one side only, with subsequent infralabials becoming progressively smaller and increasingly circular in shape, terminating in-line with the posterior orbital margin. Anterior infralabials and postmental(s) bordered posteriorly by series of enlarged irregular chin shields; rows indistinct; anteriormost approximately ¾ diameter of postmental, with gradual transition to very small, rounded throat granules posteriorly. Dorsal scales small, mostly homogenous in size, bead-like, apically flattened, partially overlain; indistinct from scales of nape. Ventral scales roughly twice diameter of dorsal scales, circular, flattened, subimbricate, slightly enlarged in precloacal region, extending in rows onto thighs where they form a subtriangular patch; transition to granules at throat abrupt. Skin folds extending ventrolaterally along trunk; anteroposterior folds above fore- and hindlimb insertion. Limbs relatively short and robust, hindlimbs longer; scales on forelimb dorsum larger than dorsal body scales, subimbricate distally; ventral forelimb scales smaller than those dorsally; clear transition to enlarged scales of palm which resemble ventral body scales. Scales on preaxial surface of thigh enlarged, up to three times diameter of dorsal body scales at knee; circular, subimbricate; gradually decreasing in size both posteriorly and distally (along shin) to smaller granules; transition to scales of soles indistinct (resembling ventral body scales). 5–6 rows of precloacal pores in males, anteriormost 2 rows extending distally just over halfway along hind thigh; absent in females. Digits broad, with dilated pads on digits II–V that rapidly transition into slender distal extension, mostly of constant width but becoming narrowly tapered near distal end; digit I smallest on all feet, digit IV longest on manus, digit V longest on pes; angle at rest between digits IV and V greatest, greater on pes than manus; digits II–V of pes and I–IV of manus very weakly webbed; dorsal scales on digits large, especially on first digit; all digits bear recurved, mostly exposed claws. Basalmost 1–2 subdigital lamellae sometimes fragmented; unfragmented subdigital lamellae curved outwards, usually smoothly but sometimes more sharply curved medially; lamellar counts of right manus: 6–7 (I), 9–11 (II), 12–14 (III), 12–15 (IV), 9–11 (V), and pes: 6–9 (I), 10–13 (II), 14–16 (III), 14–18 (IV), 11–16 (V) digits. Tail (original) stout, shorter than SVL, gradually tapered to end, roughly circular in cross-section; often lost through autotomy and then regenerated. Regenerated tail demarcated by abrupt decrease in scale size and anteroposterior striations from point of detachment distally. Base of tail distinctly swollen (TW: 8.7–14.7), most notable in males at cloacal spurs, with enlarged hexagonal scales on underside roughly twice size of more anterior ventral scales. Caudal scales usually arranged in discrete, irregular rows, generally decreasing in size distally from the septum; autotomy septa visibly marked by one, sometimes two rows of large or smaller scales, separated by 9–11 scale rows; dorsal caudal scales approximately 1.5 times size of dorsal body scales, demarcating tail base; highly variable in both size and shape, circular to rounded rectangular; ventral caudal scales enlarged medially, twice as large as dorsal; rectangular to hexagonal, higher than broad, subimbricate. Cloacal spurs consisting of a set of greatly enlarged, conical scales, with most acute point orientated posterodorsally, situated adjacent to cloaca (laterally); often asymmetric in number: 2–5 (L) and 1–4 (R), first (largest) roughly twice size of ventral scales, decreasing in size posteriorly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF79663BD944FFE46BC8FA47FE1AF996" ID-DOI="http://doi.org/10.5281/zenodo.7532715" ID-Zenodo-Dep="7532715" httpUri="https://zenodo.org/record/7532715/files/figure.png" pageId="10" pageNumber="277" startId="10.[151,250,1519,1543]" targetBox="[151,1435,181,1496]" targetPageId="10">
|
||
<paragraph id="8BB936B3D944FFE46BC8FA47FE1AF996" blockId="10.[151,1437,1519,1652]" pageId="10" pageNumber="277">
|
||
<emphasis id="B972EAA1D944FFE46BC8FA47FEFCF9E5" bold="true" box="[151,277,1519,1543]" pageId="10" pageNumber="277">FIGURE 4.</emphasis>
|
||
Living individuals of
|
||
<taxonomicName id="4C064D30D944FFE4695CFA58FCE2F9E5" box="[515,779,1519,1544]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="277" phylum="Chordata" rank="species" species="undefined-a">
|
||
<emphasis id="B972EAA1D944FFE4695CFA58FD7EF9EA" box="[515,663,1520,1544]" italics="true" pageId="10" pageNumber="277">Hoplodactylus</emphasis>
|
||
species.
|
||
<emphasis id="B972EAA1D944FFE469A7FA47FCE2F9E5" bold="true" box="[760,779,1519,1543]" pageId="10" pageNumber="277">A</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B972EAA1D944FFE46855FA47FCC6F9E5" bold="true" box="[778,815,1519,1543]" pageId="10" pageNumber="277">–C,</emphasis>
|
||
<taxonomicName id="4C064D30D944FFE46867FA58FBEBF9EA" authority="Scarsbrook & Walton & Rawlence & Hitchmough, 2023" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[824,1026,1520,1544]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="277" phylum="Chordata" rank="species" species="tohu" status="sp. nov.">
|
||
<emphasis id="B972EAA1D944FFE46867FA58FBEBF9EA" box="[824,1026,1520,1544]" italics="true" pageId="10" pageNumber="277">Hoplodactylus tohu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24157DAD944FFE46F54FA58FBAFF9E5" box="[1035,1094,1520,1543]" pageId="10" pageNumber="277" rank="species">n. sp.</taxonomicNameLabel>
|
||
;
|
||
<emphasis id="B972EAA1D944FFE46F0CFA58FB6FF9EA" bold="true" box="[1107,1158,1520,1544]" pageId="10" pageNumber="277">D–F,</emphasis>
|
||
<taxonomicName id="4C064D30D944FFE46FD0FA58FECAF9C9" authority="(Dumeril & Bibron 1836)" baseAuthorityName="Dumeril & Bibron" baseAuthorityYear="1836" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="277" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D944FFE46FD0FA58FAF3F9EA" box="[1167,1306,1520,1544]" italics="true" pageId="10" pageNumber="277">H. duvaucelii</emphasis>
|
||
(
|
||
<bibRefCitation id="EF974B42D944FFE46E76FA58FEF5F9C9" author="Dumeril, A. M. C. & Bibron, G." pageId="10" pageNumber="277" refId="ref14713" refString="Dumeril, A. M. C. & Bibron, G. (1836) Erpetologie generale, ou, Histoire naturelle complete des reptiles. Vol. 3. Roret, Paris, 531 pp." type="book" year="1836">Duméril & Bibron 1836</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B972EAA1D944FFE46A72F9BBFEACF9C9" bold="true" box="[301,325,1555,1579]" pageId="10" pageNumber="277">A,</emphasis>
|
||
Brothers Islands (Cook Strait);
|
||
<emphasis id="B972EAA1D944FFE469D1F9BBFD2DF9C9" bold="true" box="[654,708,1555,1579]" pageId="10" pageNumber="277">B–C,</emphasis>
|
||
Sentinel Rock (Marlborough Sounds).
|
||
<emphasis id="B972EAA1D944FFE46F08F9BCFB63F9CE" bold="true" box="[1111,1162,1556,1580]" pageId="10" pageNumber="277">D–F,</emphasis>
|
||
Korapuki Island (Mercury Islands). The first and second rows show sub-adult and adult representatives of each species respectively. Photographs: Nicholas Harker (Auckland, New Zealand).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31C6538D945FFE56B98FC5EFC91FBD7" pageId="11" pageNumber="278" type="distribution">
|
||
<paragraph id="8BB936B3D945FFE56B98FC5EFC91FBD7" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="278">
|
||
<emphasis id="B972EAA1D945FFE56B98FC5EFE95FBF2" bold="true" box="[199,380,1014,1040]" pageId="11" pageNumber="278">Distribution.—</emphasis>
|
||
<collectingCountry id="F3117623D945FFE56A23FC5FFDF0FBF3" box="[380,537,1015,1041]" name="New Zealand" pageId="11" pageNumber="278">New Zealand</collectingCountry>
|
||
: formerly throughout the South Island (Holocene); presently restricted to some islands in the outer
|
||
<collectingRegion id="49C2F851D945FFE56A2CFBB3FDEEFBD7" box="[371,519,1051,1077]" country="New Zealand" name="Marlborough" pageId="11" pageNumber="278">Marlborough</collectingRegion>
|
||
Sounds and Cook Strait (
|
||
<figureCitation id="133D2A36D945FFE56879FBB3FC8EFBD7" box="[806,871,1051,1077]" captionStart="FIGURE 1" captionStartId="1.[151,250,1679,1703]" captionTargetBox="[159,1427,179,1653]" captionTargetId="figure-20@1.[159,1427,179,1653]" captionTargetPageId="1" captionText="FIGURE 1. Maps of New Zealand showing distribution of Hoplodactylus duvaucelii (Duméril & Bibron 1836) (left) and H. tohu n. sp. (right), with inter- and intraspecific phylogenetic distance highlighted in a simplified mitochondrial genome phylogeny (bottom; adapted from Scarsbrook et al. 2022). Extant relict (pink) and translocated (purple) populations, historic (yellow) and Holocene subfossil records (blue) are indicated (Supp. Tables 1–2), with triangles denoting localities included in genetic analyses (see phylogeny). Abbreviations for referenced localities are as follows: Cape Maria van Dieman (CM), Poor Knights Islands (PK), Hen and Chickens Islands (HC), Great Barrier Island (GB), Maungatautari Ecosanctuary (ME) and Mana Island (MI)." figureDoi="http://doi.org/10.5281/zenodo.7532707" httpUri="https://zenodo.org/record/7532707/files/figure.png" pageId="11" pageNumber="278">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31C6538D945FFE26B98FB97FB70FBD7" lastPageId="12" lastPageNumber="279" pageId="11" pageNumber="278" type="discussion">
|
||
<paragraph id="8BB936B3D945FFE56B98FB97FD44F8E7" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="278">
|
||
<emphasis id="B972EAA1D945FFE56B98FB97FEBEFBBB" bold="true" box="[199,343,1087,1113]" pageId="11" pageNumber="278">Remarks.—</emphasis>
|
||
<taxonomicName id="4C064D30D945FFE56A08FB97FDDAFBBB" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[343,563,1087,1113]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="278" phylum="Chordata" rank="species" species="tohu">
|
||
<emphasis id="B972EAA1D945FFE56A08FB97FDDAFBBB" box="[343,563,1087,1113]" italics="true" pageId="11" pageNumber="278">Hoplodactylus tohu</emphasis>
|
||
</taxonomicName>
|
||
has been recognized as distinct from
|
||
<taxonomicName id="4C064D30D945FFE56889FB97FB84FBBB" baseAuthorityName="Dumeril & Bibron" baseAuthorityYear="1836" box="[982,1133,1087,1113]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="278" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D945FFE56889FB97FB84FBBB" box="[982,1133,1087,1113]" italics="true" pageId="11" pageNumber="278">H. duvaucelii</emphasis>
|
||
</taxonomicName>
|
||
for several years (MorganRichards
|
||
<emphasis id="B972EAA1D945FFE56A5EFBCBFED0FB9F" box="[257,313,1123,1149]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
2016;
|
||
<bibRefCitation id="EF974B42D945FFE56ADAFBCBFD76FB9F" author="Hitchmough, R. A. & Barr, B. & Knox, C. & Lettink, M. & Monks, J. M. & Patterson, G. B. & Reardon, J. T. & van Winkel, D. & Rolfe, J. & Michel, P." box="[389,671,1123,1149]" pageId="11" pageNumber="278" refId="ref17261" refString="Hitchmough, R. A., Barr, B., Knox, C., Lettink, M., Monks, J. M., Patterson, G. B., Reardon, J. T., van Winkel, D., Rolfe, J. & Michel, P. (2021 a) Conservation status of New Zealand reptiles. 2021. Department of Conservation, Wellington, 27 pp." type="book" year="2021">
|
||
Hitchmough
|
||
<emphasis id="B972EAA1D945FFE56947FBCBFDB9FB9F" box="[536,592,1123,1149]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
2021a
|
||
</bibRefCitation>
|
||
). However, it has been unclear what rank to apply to this taxon.
|
||
<collectingCountry id="F3117623D945FFE56E38FBCBFF18FB43" name="New Zealand" pageId="11" pageNumber="278">New Zealand</collectingCountry>
|
||
diplodactylids frequently have highly conserved morphologies, and often greater intra- than inter-specific morphological variation (
|
||
<bibRefCitation id="EF974B42D945FFE56AEAFB03FD3EFB27" author="Hitchmough, R. A. & Patterson, G. B. & Chapple, D. G." box="[437,727,1195,1221]" pageId="11" pageNumber="278" pagination="87 - 108" refId="ref17547" refString="Hitchmough, R. A., Patterson, G. B. & Chapple, D. G. (2016 b) Putting a name to diversity: Taxonomy of the New Zealand lizard fauna. In: Chapple, D. G. (Ed.), New Zealand Lizards. Springer, Cham, pp. 87 - 108. https: // doi. org / 10.1007 / 978 - 3 - 319 - 41674 - 8 _ 4" type="book chapter" year="2016">
|
||
Hitchmough
|
||
<emphasis id="B972EAA1D945FFE56914FB03FD6CFB27" box="[587,645,1195,1221]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
2016b
|
||
</bibRefCitation>
|
||
). With both
|
||
<taxonomicName id="4C064D30D945FFE5683CFB03FB89FB27" box="[867,1120,1195,1221]" pageId="11" pageNumber="278">
|
||
<emphasis id="B972EAA1D945FFE5683CFB03FBEFFB27" box="[867,1030,1195,1221]" italics="true" pageId="11" pageNumber="278">Hoplodactylus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
recognized here having undergone recent major range contractions resulting in significant declines in morphological and genetic diversity (
|
||
<bibRefCitation id="EF974B42D945FFE56BC0FB5BFE4AFAEF" author="Scarsbrook, L. & Sherratt, E. & Hitchmough, R. A. & Rawlence, N. J." box="[159,419,1267,1293]" pageId="11" pageNumber="278" pagination="67" refId="ref21569" refString="Scarsbrook, L., Sherratt, E., Hitchmough, R. A. & Rawlence, N. J. (2021) Skeletal variation in extant species enables systematic identification of New Zealand's large, subfossil diplodactylids. BMC ecology and evolution, 21, 67. https: // doi. org / 10.1186 / s 12862 - 021 - 01808 - 7" type="journal article" year="2021">
|
||
Scarsbrook
|
||
<emphasis id="B972EAA1D945FFE56A78FB5BFE89FAEF" box="[295,352,1267,1293]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF974B42D945FFE56AEFFB5BFD5AFAEF" author="Scarsbrook, L. & Verry, A. J. F. & Walton, K. & Hitchmough, R. A. & Rawlence, N. J." box="[432,691,1267,1293]" pageId="11" pageNumber="278" pagination="1 - 21" refId="ref21637" refString="Scarsbrook, L., Verry, A. J. F., Walton, K., Hitchmough, R. A. & Rawlence, N. J. (2022) Ancient mitochondrial genomes recovered from small vertebrate bones through minimally destructive DNA extraction: phylogeography of the New Zealand gecko genus Hoplodactylus. Molecular Ecology, 1 - 21. [early view] https: // doi. org / 10.1111 / mec. 16434" type="book chapter" year="2022">
|
||
Scarsbrook
|
||
<emphasis id="B972EAA1D945FFE56969FB5BFD86FAEF" box="[566,623,1267,1293]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
), it is difficult to weight subtle character differences as these may have arisen, or increased in prevalence, recently, through chance, in relictual populations, and therefore not reflect deeper evolutionary divergences. That these taxa have been reported to produce viable cross-bred offspring in captive populations (
|
||
<bibRefCitation id="EF974B42D945FFE56A03FAF7FD4FFA9B" author="Morgan-Richards, M. & Hinlo, A. R. & Smuts-Kennedy, C. & Innes, J. & Ji, W. & Barry, M. & Brunton, D. & Hitchmough, R. A." box="[348,678,1375,1401]" pageId="11" pageNumber="278" pagination="77 - 86" refId="ref20012" refString="Morgan-Richards, M., Hinlo, A. R., Smuts-Kennedy, C., Innes, J., Ji, W., Barry, M., Brunton, D. & Hitchmough, R. A. (2016) Identification of a rare gecko from North Island New Zealand, and genetic assessment of its probable origin: A novel mainland conservation priority? Journal of Herpetology, 50, 77 - 86. https: // doi. org / 10.1670 / 13 - 128" type="journal article" year="2016">
|
||
Morgan-Richards
|
||
<emphasis id="B972EAA1D945FFE56974FAF7FD8DFA9B" box="[555,612,1375,1401]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
) fails to meet the criteria of the Biological Species Concept (
|
||
<bibRefCitation id="EF974B42D945FFE56E01FAF7FF3AFA7F" author="Mayr, E." pageId="11" pageNumber="278" refId="ref19250" refString="Mayr, E. (1942) Systematics and the Origin of Species. Columbia University, New York, New York, 367 pp." type="book" year="1942">Mayr 1942</bibRefCitation>
|
||
). Reproductive isolation in the wild cannot be tested given their allopatric distributions. However, several pairings of lizard species that are widely recognized to be distinct (
|
||
<bibRefCitation id="EF974B42D945FFE56824FA0FFBB2FA23" author="Brennan, I. G. & Bauer, A. M. & Jackman, T. R." box="[891,1115,1447,1473]" pageId="11" pageNumber="278" pagination="577 - 590" refId="ref13498" refString="Brennan, I. G., Bauer, A. M. & Jackman, T. R. (2016) Mitochondrial introgression via ancient hybridization, and systematics of the Australian endemic pygopodid gecko genus Delma. Molecular Phylogenetics and Evolution, 94, 577 - 590. https: // doi. org / 10.1016 / j. ympev. 2015.10.005" type="journal article" year="2016">
|
||
Brennan
|
||
<emphasis id="B972EAA1D945FFE568BDFA0FFBF0FA23" box="[994,1049,1447,1473]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF974B42D945FFE56F3AFA0FFAA6FA23" author="Leache, A. D. & Cole, C. J." box="[1125,1359,1447,1473]" pageId="11" pageNumber="278" pagination="1035 - 1054" refId="ref18580" refString="Leache, A. D. & Cole, C. J. (2006) Hybridization between multiple fence lizard lineages in an ecotone: locally discordant variation in mitochondrial DNA, chromosomes, and morphology. Molecular Ecology, 16, 1035 - 1054. https: // doi. org / 10.1111 / j. 1365 - 294 X. 2006.03194. x" type="journal article" year="2006">Leaché & Cole 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF974B42D945FFE56E06FA0FFEF1FA07" author="Olave, M. & Martinez, L. E. & Avila, L. J. & Sites, J. W. & Morando, M." pageId="11" pageNumber="278" pagination="381 - 391" refId="ref20717" refString="Olave, M., Martinez, L. E., Avila, L. J., Sites, J. W. & Morando, M. (2011) Evidence of hybridization in the Argentinean lizards Liolaemus gracilis and Liolaemus bibronii (Iguania: Liolaemini): an integrative approach based on genes and morphology. Molecular Phylogenetics and Evolution, 61, 381 - 391. https: // doi. org / 10.1016 / j. ympev. 2011.07.006" type="journal article" year="2011">
|
||
Olave
|
||
<emphasis id="B972EAA1D945FFE56BC8FA63FF3AFA07" box="[151,211,1483,1509]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
), including some other
|
||
<collectingCountry id="F3117623D945FFE56972FA63FD2EFA07" box="[557,711,1483,1509]" name="New Zealand" pageId="11" pageNumber="278">New Zealand</collectingCountry>
|
||
diplodactylids (e.g.,
|
||
<taxonomicName id="4C064D30D945FFE568E1FA63FBBCFA07" box="[958,1109,1483,1509]" class="Reptilia" family="Diplodactylidae" genus="Naultinus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="278" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B972EAA1D945FFE568E1FA63FBC2FA07" box="[958,1067,1483,1509]" italics="true" pageId="11" pageNumber="278">Naultinus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EF974B42D945FFE56F36FA63FA79FA07" author="Hitchmough, R. A. & Patterson, G. B. & Chapple, D. G." box="[1129,1424,1483,1509]" pageId="11" pageNumber="278" pagination="87 - 108" refId="ref17547" refString="Hitchmough, R. A., Patterson, G. B. & Chapple, D. G. (2016 b) Putting a name to diversity: Taxonomy of the New Zealand lizard fauna. In: Chapple, D. G. (Ed.), New Zealand Lizards. Springer, Cham, pp. 87 - 108. https: // doi. org / 10.1007 / 978 - 3 - 319 - 41674 - 8 _ 4" type="book chapter" year="2016">
|
||
Hitchmough
|
||
<emphasis id="B972EAA1D945FFE56E5FFA63FAD5FA07" box="[1280,1340,1483,1509]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
2016b
|
||
</bibRefCitation>
|
||
), have been shown to similarly produce viable hybrid offspring. Intergeneric hybrids have even been reported (RH pers. obs). Genetic distance (i.e., ~2.2% and ~4.0% across the mitochondrial genome and ND2 respectively) and estimated timing of lineage divergence of ~4.51 mya (
|
||
<bibRefCitation id="EF974B42D945FFE569A9F99FFC1FF9B3" author="Scarsbrook, L. & Verry, A. J. F. & Walton, K. & Hitchmough, R. A. & Rawlence, N. J." box="[758,1014,1591,1617]" pageId="11" pageNumber="278" pagination="1 - 21" refId="ref21637" refString="Scarsbrook, L., Verry, A. J. F., Walton, K., Hitchmough, R. A. & Rawlence, N. J. (2022) Ancient mitochondrial genomes recovered from small vertebrate bones through minimally destructive DNA extraction: phylogeography of the New Zealand gecko genus Hoplodactylus. Molecular Ecology, 1 - 21. [early view] https: // doi. org / 10.1111 / mec. 16434" type="book chapter" year="2022">
|
||
Scarsbrook
|
||
<emphasis id="B972EAA1D945FFE56822F99FFC5CF9B3" box="[893,949,1591,1617]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
) between the two
|
||
<taxonomicName id="4C064D30D945FFE56F9CF99FFF2AF997" pageId="11" pageNumber="278">
|
||
<emphasis id="B972EAA1D945FFE56F9CF99FFA8FF9B3" box="[1219,1382,1591,1617]" italics="true" pageId="11" pageNumber="278">Hoplodactylus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
recognized here are comparable with those of other recognized diplodactylid species pairings (e.g., 3.8% ND2 divergence between
|
||
<taxonomicName id="4C064D30D945FFE56A24F9D7FCBBF97B" authority="Whitaker, 1985" authorityName="Whitaker" authorityYear="1985" box="[379,850,1662,1689]" class="Reptilia" family="Diplodactylidae" genus="Mokopirirakau" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="278" phylum="Chordata" rank="species" species="kahutarae">
|
||
<emphasis id="B972EAA1D945FFE56A24F9D7FD74F97B" box="[379,669,1663,1689]" italics="true" pageId="11" pageNumber="278">Mokopirirakau kahutarae</emphasis>
|
||
Whitaker, 1985
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C064D30D945FFE568D4F9D7FAD0F97B" authority="Gray, 1845" authorityName="Gray" authorityYear="1845" box="[907,1337,1662,1689]" class="Reptilia" family="Diplodactylidae" genus="Mokopirirakau" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="278" phylum="Chordata" rank="species" species="granulatus">
|
||
<emphasis id="B972EAA1D945FFE568D4F9D7FB5DF97B" box="[907,1204,1663,1689]" italics="true" pageId="11" pageNumber="278">Mokopirirakau granulatus</emphasis>
|
||
<bibRefCitation id="EF974B42D945FFE56FE5F9D7FAD0F97B" author="Gray, J. E." box="[1210,1337,1662,1689]" pageId="11" pageNumber="278" refId="ref16059" refString="Gray, J. E. (1845) Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of the British Museum, London, 289 pp." type="book" year="1845">Gray, 1845</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EF974B42D945FFE56E1BF9D7FF11F95F" author="Knox, C. & Hitchmough, R. A. & Nielsen, S. V. & Jewell, T. & Bell, T. P." pageId="11" pageNumber="278" pagination="140 - 156" refId="ref18437" refString="Knox, C., Hitchmough, R. A., Nielsen, S. V., Jewell, T. & Bell, T. P. (2021) A new, enigmatic species of black-eyed gecko (Reptilia: Diplodactylidae: Mokopirirakau) from North Otago, New Zealand. Zootaxa, 4964 (1), 140 - 156. https: // doi. org / 10.11646 / zootaxa. 4964.1.7" type="journal article" year="2021">
|
||
Knox
|
||
<emphasis id="B972EAA1D945FFE56ED7F9D7FF44F95F" italics="true" pageId="11" pageNumber="278">et al</emphasis>
|
||
. 2021
|
||
</bibRefCitation>
|
||
). Their discrete (allopatric) distributions reflect commonly observed biogeographic patterns in other taxa (
|
||
<bibRefCitation id="EF974B42D945FFE56BC0F96FFE81F903" author="Baker, A. J. & Daugherty, C. H. & Colbourne, R. & Mclennan, J. L." box="[159,360,1734,1761]" pageId="11" pageNumber="278" pagination="8254 - 8258" refId="ref12017" refString="Baker, A. J., Daugherty, C. H., Colbourne, R. & Mclennan, J. L. (1995) Flightless brown kiwis of New Zealand possess extremely subdivided population structure and cryptic species like small mammals. Proceedings of the National Academy of Sciences of the United States of America, 92, 8254 - 8258. https: // doi. org / 10.1073 / pnas. 92.18.8254" type="journal article" year="1995">
|
||
Baker
|
||
<emphasis id="B972EAA1D945FFE56BB4F96FFECCF903" box="[235,293,1735,1761]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
1995
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF974B42D945FFE56A2AF96FFDABF903" author="Efford, M. & Howitt, R. & Gleeson, D." box="[373,578,1735,1761]" pageId="11" pageNumber="278" pagination="445 - 456" refId="ref14972" refString="Efford, M., Howitt, R. & Gleeson, D. (2002) Phylogenetic relationships of Wainuia (Mollusca: Pulmonata) - Biogeography and conservation implications. Journal of the Royal Society of New Zealand, 32, 445 - 456. https: // doi. org / 10.1080 / 03014223.2002.9517703" type="journal article" year="2002">
|
||
Efford
|
||
<emphasis id="B972EAA1D945FFE56A9AF96FFE16F903" box="[453,511,1735,1761]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
2002
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF974B42D945FFE56911F96FFCC7F903" author="Greaves, S. N. J. & Chapple, D. G. & Gleeson, D. M. & Daugherty, C. H. & Ritchie, P. A." box="[590,814,1735,1761]" pageId="11" pageNumber="278" pagination="729 - 739" refId="ref16093" refString="Greaves, S. N. J., Chapple, D. G., Gleeson, D. M., Daugherty, C. H. & Ritchie, P. A. (2007) Phylogeography of the spotted skink (Oligosoma lineoocellatum) and green skink (O. chloronoton) species complex (Lacertilia: Scincidae) in New Zealand reveals pre-Pleistocene divergence. Molecular Phylogenetics and Evolution, 45, 729 - 739. https: // doi. org / 10.1016 / j. ympev. 2007.06.008" type="journal article" year="2007">
|
||
Greaves
|
||
<emphasis id="B972EAA1D945FFE569EDF96FFD02F903" box="[690,747,1735,1761]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF974B42D945FFE56864F96FFBFCF903" author="Liggins, L. & Chapple, D. G. & Daugherty, C. H. & Ritchie, P. A." box="[827,1045,1735,1761]" pageId="11" pageNumber="278" pagination="3668 - 3683" refId="ref18853" refString="Liggins, L., Chapple, D. G., Daugherty, C. H. & Ritchie, P. A. (2008) A SINE of restricted gene flow across the Alpine Fault: phylogeography of the New Zealand common skink (Oligosoma nigriplantare polychroma). Molecular Ecology, 17, 3668 - 3683. https: // doi. org / 10.1111 / j. 1365 - 294 X. 2008.03864. x" type="journal article" year="2008">
|
||
Liggins
|
||
<emphasis id="B972EAA1D945FFE568C6F96FFC3BF903" box="[921,978,1735,1761]" italics="true" pageId="11" pageNumber="278">et al.</emphasis>
|
||
2008
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF974B42D945FFE56F7DF96FFB40F903" author="Lloyd, B. D." box="[1058,1193,1735,1761]" pageId="11" pageNumber="278" pagination="460 - 476" refId="ref18933" refString="Lloyd, B. D. (2003) Intraspecific phylogeny of the New Zealand short-tailed bat Mystacina tuberculata inferred from multiple mitochondrial gene sequences. Systematic Biology, 52, 460 - 476. https: // doi. org / 10.1080 / 10635150390218187" type="journal article" year="2003">Lloyd 2003</bibRefCitation>
|
||
), which further influences our treatment of species-level distinction.
|
||
</paragraph>
|
||
<paragraph id="8BB936B3D945FFE26B98F8A7FEB0FEFF" blockId="11.[151,1437,151,2013]" lastBlockId="12.[151,1437,151,1257]" lastPageId="12" lastPageNumber="279" pageId="11" pageNumber="278">
|
||
Reports of interbreeding between
|
||
<taxonomicName id="4C064D30D945FFE56960F8A7FD7BF8CB" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[575,658,1807,1833]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="278" phylum="Chordata" rank="species" species="tohu">
|
||
<emphasis id="B972EAA1D945FFE56960F8A7FD7BF8CB" box="[575,658,1807,1833]" italics="true" pageId="11" pageNumber="278">H. tohu</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C064D30D945FFE5699AF8A7FCB2F8CB" baseAuthorityName="Dumeril & Bibron" baseAuthorityYear="1836" box="[709,859,1807,1833]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="278" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D945FFE5699AF8A7FCB2F8CB" box="[709,859,1807,1833]" italics="true" pageId="11" pageNumber="278">H. duvaucelii</emphasis>
|
||
</taxonomicName>
|
||
in captivity makes careful management of both species essential to maintain species/genetic integrity. Individuals sourced for reintroduction or translocation purposes need to be of the correct species, and sourced from moderately large and stable populations (e.g. North Brother Island;
|
||
<bibRefCitation id="EF974B42D945FFE56BB6F8D3FE90F877" author="Wilson, J." box="[233,377,1915,1941]" pageId="11" pageNumber="278" refId="ref24377" refString="Wilson, J. (2010) Population viability and resource competition on North Brother Island: conservation implications for tuatara (Sphenodon punctatus) and Duvaucel's gecko (Hoplodactylus duvaucelii). MSc Thesis, Victoria University of Wellington, Wellington, 106 pp." type="book" year="2010">Wilson 2010</bibRefCitation>
|
||
) to minimise impact on the source population. Further, source populations should be proximate (where practicable) to the site of translocation to ensure preservation of regional adaptations; a consideration more applicable to
|
||
<taxonomicName id="4C064D30D945FFE56A72F86BFE2DF83F" baseAuthorityName="Dumeril & Bibron" baseAuthorityYear="1836" box="[301,452,1987,2013]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="278" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D945FFE56A72F86BFE2DF83F" box="[301,452,1987,2013]" italics="true" pageId="11" pageNumber="278">H. duvaucelii</emphasis>
|
||
</taxonomicName>
|
||
as extant populations are more numerous and occupy more ecologically varied habitats. It seems probable, for example, that
|
||
<taxonomicName id="4C064D30D942FFE26969FF3FFD27FF53" baseAuthorityName="Dumeril & Bibron" baseAuthorityYear="1836" box="[566,718,151,177]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="279" phylum="Chordata" rank="species" species="duvaucelii">
|
||
<emphasis id="B972EAA1D942FFE26969FF3FFD27FF53" box="[566,718,151,177]" italics="true" pageId="12" pageNumber="279">H. duvaucelii</emphasis>
|
||
</taxonomicName>
|
||
rather than
|
||
<taxonomicName id="4C064D30D942FFE26808FF3FFC45FF53" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[855,940,151,177]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="279" phylum="Chordata" rank="species" species="tohu">
|
||
<emphasis id="B972EAA1D942FFE26808FF3FFC45FF53" box="[855,940,151,177]" italics="true" pageId="12" pageNumber="279">H. tohu</emphasis>
|
||
</taxonomicName>
|
||
, would have naturally occurred at Mana Island, off the southern North Island. However, the latter was translocated there from North Brother Island in 1998 through the release of 40 individuals (
|
||
<bibRefCitation id="EF974B42D942FFE26966FF77FD09FF1B" author="Miskelly, C." box="[569,736,223,249]" pageId="12" pageNumber="279" refId="ref19699" refString="Miskelly, C. (2010) Mana Island ecological restoration plan review. Department of Conservation, Wellington, 54 pp." type="book" year="2010">Miskelly 2010</bibRefCitation>
|
||
), with a self-sustaining population reported 15-years later (
|
||
<bibRefCitation id="EF974B42D942FFE26E33FF77FEA4FEFF" author="Bell, T. P. & Herbert, S. M." pageId="12" pageNumber="279" pagination="37 - 46" refId="ref13088" refString="Bell, T. P. & Herbert, S. M. (2017) Establishment of a self-sustaining population of a long-lived, slow-breeding gecko species (Diplodactylidae: Hoplodactylus duvaucelii) evident 15 years after translocation. Journal of Herpetology, 51, 37 - 46. https: // doi. org / 10.1670 / 15 - 106" type="journal article" year="2017">Bell & Herbert 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB936B3D942FFE26B98FE8FFCD7FCDB" blockId="12.[151,1437,151,1257]" pageId="12" pageNumber="279">
|
||
<taxonomicName id="4C064D30D942FFE26B98FE8FFEF6FEA3" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[199,287,295,321]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="279" phylum="Chordata" rank="species" species="tohu">
|
||
<emphasis id="B972EAA1D942FFE26B98FE8FFEF6FEA3" box="[199,287,295,321]" italics="true" pageId="12" pageNumber="279">H. tohu</emphasis>
|
||
</taxonomicName>
|
||
has been recognized as a distinct management unit by the Department of Conservation since 2021 (
|
||
<bibRefCitation id="EF974B42D942FFE26BC0FEE3FE2BFE87" author="Hitchmough, R. A. & Barr, B. & Knox, C. & Lettink, M. & Monks, J. M. & Patterson, G. B. & Reardon, J. T. & van Winkel, D. & Rolfe, J. & Michel, P." box="[159,450,331,357]" pageId="12" pageNumber="279" refId="ref17261" refString="Hitchmough, R. A., Barr, B., Knox, C., Lettink, M., Monks, J. M., Patterson, G. B., Reardon, J. T., van Winkel, D., Rolfe, J. & Michel, P. (2021 a) Conservation status of New Zealand reptiles. 2021. Department of Conservation, Wellington, 27 pp." type="book" year="2021">
|
||
Hitchmough
|
||
<emphasis id="B972EAA1D942FFE26A69FEE3FE98FE87" box="[310,369,331,357]" italics="true" pageId="12" pageNumber="279">et al.</emphasis>
|
||
2021a
|
||
</bibRefCitation>
|
||
), listed as “nationally increasing” with the ‘Conservation Dependant’ and ‘Range Restricted’ qualifiers.
|
||
<taxonomicName id="4C064D30D942FFE26A2CFEC7FE20FE6B" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[371,457,367,393]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="279" phylum="Chordata" rank="species" species="tohu">
|
||
<emphasis id="B972EAA1D942FFE26A2CFEC7FE20FE6B" box="[371,457,367,393]" italics="true" pageId="12" pageNumber="279">H. tohu</emphasis>
|
||
</taxonomicName>
|
||
has a severely restricted distribution comprising a few islands (
|
||
<figureCitation id="133D2A36D942FFE26FFEFEC7FB0FFE6B" box="[1185,1254,367,393]" captionStart="FIGURE 1" captionStartId="1.[151,250,1679,1703]" captionTargetBox="[159,1427,179,1653]" captionTargetId="figure-20@1.[159,1427,179,1653]" captionTargetPageId="1" captionText="FIGURE 1. Maps of New Zealand showing distribution of Hoplodactylus duvaucelii (Duméril & Bibron 1836) (left) and H. tohu n. sp. (right), with inter- and intraspecific phylogenetic distance highlighted in a simplified mitochondrial genome phylogeny (bottom; adapted from Scarsbrook et al. 2022). Extant relict (pink) and translocated (purple) populations, historic (yellow) and Holocene subfossil records (blue) are indicated (Supp. Tables 1–2), with triangles denoting localities included in genetic analyses (see phylogeny). Abbreviations for referenced localities are as follows: Cape Maria van Dieman (CM), Poor Knights Islands (PK), Hen and Chickens Islands (HC), Great Barrier Island (GB), Maungatautari Ecosanctuary (ME) and Mana Island (MI)." figureDoi="http://doi.org/10.5281/zenodo.7532707" httpUri="https://zenodo.org/record/7532707/files/figure.png" pageId="12" pageNumber="279">Fig. 1</figureCitation>
|
||
; Supp.
|
||
<tableCitation id="C6840308D942FFE26E60FEC7FA78FE6B" box="[1343,1425,367,393]" captionStart="TABLE 1" captionStartId="3.[151,239,151,175]" captionTargetId="graphics-154@3.[151,1436,465,1997]" captionTargetPageId="3" captionText="TABLE 1. Measurements (mm), scalation counts and proportional values for mature specimens of Hoplodactylus duvaucelii (n = 10) and H. tohu n. sp. (n = 11), (see Supp. Table 3 for individual values and Supp. Fig. 1–3 for plots). Abbreviations are as follows: snout-vent length (SVL); trunk length (TrK); head length (HL); head width (HW); head height (HH); interorbital distance (IO); eye to nares distance (EN); internarial distance (IN); eye to ear distance (EE); ear length (EL); supralabials to eye (SuLE); infralabials (InL); internasals (IntN); nasal border (NB); postmentals (PstM); cloacal spurs (CS); precloacal pores (PcP); caudal annuli (CA); subdigital lamellae of manual (m.SDL) and pedal (p.SDL) digits. Left (L) and right (R) are abbreviated at times and digits 1-5 are referred to as I-V." httpUri="http://table.plazi.org/id/DF79663BD94DFFED6BC8FF3FFBEDFE65" pageId="12" pageNumber="279" tableUuid="DF79663BD94DFFED6BC8FF3FFBEDFE65">Table 1</tableCitation>
|
||
) with highly anthropogenically modified habitats. The estimated total population size of the largest extant population, on the Brothers Islands, is between 583–677 (
|
||
<bibRefCitation id="EF974B42D942FFE26997FE1FFCB4FE33" author="Wilson, J." box="[712,861,439,465]" pageId="12" pageNumber="279" refId="ref24377" refString="Wilson, J. (2010) Population viability and resource competition on North Brother Island: conservation implications for tuatara (Sphenodon punctatus) and Duvaucel's gecko (Hoplodactylus duvaucelii). MSc Thesis, Victoria University of Wellington, Wellington, 106 pp." type="book" year="2010">Wilson 2010</bibRefCitation>
|
||
). Based on this, the suggested IUCN Red List status of
|
||
<taxonomicName id="4C064D30D942FFE26BEAFE73FEE0FE17" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[181,265,475,501]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="279" phylum="Chordata" rank="species" species="tohu">
|
||
<emphasis id="B972EAA1D942FFE26BEAFE73FEE0FE17" box="[181,265,475,501]" italics="true" pageId="12" pageNumber="279">H. tohu</emphasis>
|
||
</taxonomicName>
|
||
is ‘Critically Endangered A1 (a, b, c, e)’: population reduction of>90% observed, estimated, inferred, or suspected in the past where the causes of the reduction are clearly reversible AND understood AND have ceased. Occurrence of
|
||
<taxonomicName id="4C064D30D942FFE26A61FD8BFE7AFDDF" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[318,403,547,573]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="279" phylum="Chordata" rank="species" species="tohu">
|
||
<emphasis id="B972EAA1D942FFE26A61FD8BFE7AFDDF" box="[318,403,547,573]" italics="true" pageId="12" pageNumber="279">H. tohu</emphasis>
|
||
</taxonomicName>
|
||
on several managed predator-free islands that already have conservation management strategies in place does at least render moderate security to this species from localized disturbances such as fire or predator incursions. However, additional protection through translocations to fenced and more distant mainland sanctuaries (e.g. Orokonui Ecosanctuary near Dunedin, and the Mokomoko Dryland Sanctuary in Central
|
||
<collectingRegion id="49C2F851D942FFE26E11FD27FA7DFD4B" box="[1358,1428,655,681]" country="New Zealand" name="Otago" pageId="12" pageNumber="279">Otago</collectingRegion>
|
||
) could be beneficial, given the increased vulnerability of small and sparsely forested islands to the effects of climate change (e.g. sea-level rise, coastal erosion, storm intensity;
|
||
<bibRefCitation id="EF974B42D942FFE2686AFD7FFBA5FD13" author="Macinnis-Ng, C. & Mcintosh, A. R. & Monks, J. M. & Waipara, N. & White, R. S. A. & Boudjelas, S. & Clark, C. D. & Clearwater, M. J. & Curran, T. J. & Dickinson, K. J. M. & Nelson, N. & Perry, G. L. W. & Richardson, S. J. & Stanley, M. C. & Peltzer, D. A." box="[821,1100,727,753]" pageId="12" pageNumber="279" pagination="216 - 224" refId="ref19104" refString="Macinnis-Ng, C., Mcintosh, A. R., Monks, J. M., Waipara, N., White, R. S. A., Boudjelas, S., Clark, C. D., Clearwater, M. J., Curran, T. J., Dickinson, K. J. M., Nelson, N., Perry, G. L. W., Richardson, S. J., Stanley, M. C. & Peltzer, D. A. (2021) Climatechange impacts exacerbate conservation threats in island systems: New Zealand as a case study. Frontiers in Ecology and the Environment, 19, 216 - 224. https: // doi. org / 10.1002 / FEE. 2285" type="journal article" year="2021">
|
||
Macinnis-Ng
|
||
<emphasis id="B972EAA1D942FFE2688EFD7FFBE0FD13" box="[977,1033,727,753]" italics="true" pageId="12" pageNumber="279">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
). Such translocation may also facilitate the re-establishment of ‘lost’ ecological interactions and morphological diversity (e.g.
|
||
<bibRefCitation id="EF974B42D942FFE26F83FD53FF39FCDB" author="Scarsbrook, L. & Sherratt, E. & Hitchmough, R. A. & Rawlence, N. J." pageId="12" pageNumber="279" pagination="67" refId="ref21569" refString="Scarsbrook, L., Sherratt, E., Hitchmough, R. A. & Rawlence, N. J. (2021) Skeletal variation in extant species enables systematic identification of New Zealand's large, subfossil diplodactylids. BMC ecology and evolution, 21, 67. https: // doi. org / 10.1186 / s 12862 - 021 - 01808 - 7" type="journal article" year="2021">
|
||
Scarsbrook
|
||
<emphasis id="B972EAA1D942FFE26E3DFD53FA74FCF7" box="[1378,1437,763,789]" italics="true" pageId="12" pageNumber="279">et al.</emphasis>
|
||
2021
|
||
</bibRefCitation>
|
||
) specific to mainland and densely forested ecosystems.
|
||
</paragraph>
|
||
<paragraph id="8BB936B3D942FFE26B98FCEBFB70FBD7" blockId="12.[151,1437,151,1257]" pageId="12" pageNumber="279">
|
||
There is evidence of genetic sub-structuring (
|
||
<bibRefCitation id="EF974B42D942FFE269E6FCEBFC5CFCBF" author="Scarsbrook, L. & Verry, A. J. F. & Walton, K. & Hitchmough, R. A. & Rawlence, N. J." box="[697,949,835,861]" pageId="12" pageNumber="279" pagination="1 - 21" refId="ref21637" refString="Scarsbrook, L., Verry, A. J. F., Walton, K., Hitchmough, R. A. & Rawlence, N. J. (2022) Ancient mitochondrial genomes recovered from small vertebrate bones through minimally destructive DNA extraction: phylogeography of the New Zealand gecko genus Hoplodactylus. Molecular Ecology, 1 - 21. [early view] https: // doi. org / 10.1111 / mec. 16434" type="book chapter" year="2022">
|
||
Scarsbrook
|
||
<emphasis id="B972EAA1D942FFE26860FCEBFC9FFCBF" box="[831,886,835,861]" italics="true" pageId="12" pageNumber="279">et al.</emphasis>
|
||
2022
|
||
</bibRefCitation>
|
||
) within the distribution of
|
||
<taxonomicName id="4C064D30D942FFE26F84FCEBFAC7FCBF" authorityName="Scarsbrook & Walton & Rawlence & Hitchmough" authorityYear="2023" box="[1243,1326,835,861]" class="Reptilia" family="Diplodactylidae" genus="Hoplodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="279" phylum="Chordata" rank="species" species="tohu">
|
||
<emphasis id="B972EAA1D942FFE26F84FCEBFAC7FCBF" box="[1243,1326,835,861]" italics="true" pageId="12" pageNumber="279">H. tohu</emphasis>
|
||
</taxonomicName>
|
||
, although this has been greatly reduced with the extinction of mainland South Island populations. Careful management, and possible further research should consider if individual populations are experiencing inbreeding depression and might benefit from genetic rescue – establishment of ‘new’ populations through interbreeding of individuals from different populations to increase genetic ‘fitness’ (i.e., adaptability). Conversely, continued maintenance of discrete lineages may also be appropriate if those lineages reflect local habitat adaptations or if pooled populations would comprise too few individuals to maintain the resulting genetic diversity due to genetic drift.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31C6538D942FFE26B98FB97FED9FB0B" pageId="12" pageNumber="279" type="etymology">
|
||
<paragraph id="8BB936B3D942FFE26B98FB97FED9FB0B" blockId="12.[151,1437,151,1257]" pageId="12" pageNumber="279">
|
||
<emphasis id="B972EAA1D942FFE26B98FB97FE80FBBB" bold="true" box="[199,361,1087,1113]" pageId="12" pageNumber="279">Etymology.—</emphasis>
|
||
The specific epithet was proposed by Dr Sharon Barcello-Gemmel, Rangatira of Te Ātiawa o Te Waka-a-Māui Trust, in recognition of the tupuna [ancestor] Tohu Kākahi. Tohu Kākahi was one of the two Parihaka prophets with whakapapa [genealogical] connections to Te Ātiawa – the iwi [tribe] with mana whenua [authority] over Ngāwhatu Kai Ponu [The Brothers], where the largest extant population occurs. Name used as noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |