treatments-xml/data/03/AF/87/03AF878FFFC7FFAB0C933F80FD54FB97.xml

<document id="0820305C1DDD3C639FF2A2E639E20B72" ID-CLB-Dataset="3646" ID-DOI="10.11646/zootaxa.5067.1.5" ID-GBIF-Dataset="74f3ce16-7e8f-420a-b76e-d306ea2a42ad" ID-ISSN="1175-5326" ID-Zenodo-Dep="5656552" ID-ZooBank="4763DE40-3A0D-4F23-9381-3C4754A59AE4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1636445280528" checkinUser="plazi" docAuthor="Nozaki, Takao &amp; Park, Sun-Jin" docDate="2021" docId="03AF878FFFC7FFAB0C933F80FD54FB97" docLanguage="en" docName="zootaxa.5067.1.5.pdf" docOrigin="Zootaxa 5067 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacophila kangae Park &amp; Nozaki 2021, sp. nov." docType="treatment" docVersion="8" lastPageNumber="99" masterDocId="FF96FFF7FFC6FFA90C043D33FFE8FF86" masterDocTitle="Two new species of the genus Rhyacophila Pictet (Trichoptera, Rhyacophilidae) from Korea and Japan" masterLastPageNumber="105" masterPageNumber="97" pageNumber="98" updateTime="1701999324714" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8E42F5FE6CC73A66A3452FFEF3C2019F">Two new species of the genus Rhyacophila Pictet (Trichoptera, Rhyacophilidae) from Korea and Japan</mods:title>
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<mods:namePart id="95BA14A38570741661F9768EEB3F11E6">Nozaki, Takao</mods:namePart>
<mods:affiliation id="70A6C3E7F2240EC9AD46696F28A382C1">3 - 16 - 15, Midorigaoka, Ninomiya-machi, Naka-gun, Kanagawa, 259 - 0132 Japan.</mods:affiliation>
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<mods:namePart id="EB4EFA21FFBEC0D2C1A0D2185A8B4023">Park, Sun-Jin</mods:namePart>
<mods:affiliation id="2228C0508D34CA80F82D4A3944FAE0AF">Department of Life Science, Kyonggi University, Suwon 16227, Korea.</mods:affiliation>
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<treatment id="03AF878FFFC7FFAB0C933F80FD54FB97" ID-DOI="http://doi.org/10.5281/zenodo.5698997" ID-GBIF-Taxon="190304773" ID-Zenodo-Dep="5698997" LSID="urn:lsid:plazi:treatment:03AF878FFFC7FFAB0C933F80FD54FB97" httpUri="http://treatment.plazi.org/id/03AF878FFFC7FFAB0C933F80FD54FB97" lastPageId="2" lastPageNumber="99" pageId="1" pageNumber="98">
<subSubSection id="C31C6512FFC7FFA80C933F80FD51FD4B" box="[151,697,691,718]" pageId="1" pageNumber="98" type="nomenclature">
<paragraph id="8BB93699FFC7FFA80C933F80FD51FD4B" blockId="1.[151,697,691,753]" box="[151,697,691,718]" pageId="1" pageNumber="98">
<heading id="D0F181F5FFC7FFA80C933F80FD51FD4B" bold="true" box="[151,697,691,718]" fontSize="11" level="1" pageId="1" pageNumber="98" reason="1">
<emphasis id="B972EA8BFFC7FFA80C933F80FD51FD4B" bold="true" box="[151,697,691,718]" pageId="1" pageNumber="98">
<taxonomicName id="4C064D1AFFC7FFA80C933F80FDBCFD4B" authority="Park &amp; Nozaki" authorityName="Park &amp; Nozaki" authorityYear="2021" box="[151,596,691,718]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="kangae" status="sp. nov.">
<emphasis id="B972EA8BFFC7FFA80C933F80FE78FD48" bold="true" box="[151,400,691,718]" italics="true" pageId="1" pageNumber="98">Rhyacophila kangae</emphasis>
Park &amp; Nozaki
</taxonomicName>
<taxonomicNameLabel id="A24157F0FFC7FFA80E5F3F80FD51FD4B" box="[603,697,691,717]" pageId="1" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31C6512FFC7FFA80C933FE4FCBCFCB5" pageId="1" pageNumber="98" type="description">
<paragraph id="8BB93699FFC7FFA80C933FE4FECFFD77" blockId="1.[151,697,691,753]" box="[151,295,727,753]" pageId="1" pageNumber="98">
(
<figureCitation id="133D2A1CFFC7FFA80C9B3FE4FF0EFD77" box="[159,230,727,753]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Figs 1</figureCitation>
, 
<figureCitation id="133D2A1CFFC7FFA80CF03FE4FEECFD77" box="[244,260,727,753]" captionStart="FIGURE 4–7" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 4–7. Rhyacophila spp., female. 4A–4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A–5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="1" pageNumber="98">4</figureCitation>
, 
<figureCitation id="133D2A1CFFC7FFA80D143FE4FEF7FD77" box="[272,287,727,753]" captionStart="FIGURE 9" captionStartId="7.[151,250,1110,1134]" captionTargetBox="[155,1428,281,1079]" captionTargetId="figure-64@7.[151,1436,273,1087]" captionTargetPageId="7" captionText="FIGURE 9. Distributions of 4 Rhyacophila species in East Asia. Distribution areas of R. confissa and R. vicina are cited from Ivanov (2011), Park &amp; Kong (2020), and Yang et al. (2016)." figureDoi="http://doi.org/10.5281/zenodo.5656565" httpUri="https://zenodo.org/record/5656565/files/figure.png" pageId="1" pageNumber="98">9</figureCitation>
)
</paragraph>
<paragraph id="8BB93699FFC7FFA80C933E2FFCBCFCB5" blockId="1.[151,852,796,820]" box="[151,852,796,820]" pageId="1" pageNumber="98">
<taxonomicName id="4C064D1AFFC7FFA80C933E2FFEB8FCB2" box="[151,336,796,820]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="undefined-3">
<emphasis id="B972EA8BFFC7FFA80C933E2FFEFFFCB2" box="[151,279,796,820]" italics="true" pageId="1" pageNumber="98">Rhyacophila</emphasis>
sp. 3
</taxonomicName>
: 
<bibRefCitation id="EF974B68FFC7FFA80D5F3E2FFE25FCB2" author="Kang, M. - S." box="[347,461,796,820]" pageId="1" pageNumber="98" refId="ref5287" refString="Kang, M. - S. (2020) The Korean caddisflies. Nature and Ecology, Seoul, 548 pp. [in Korean]" type="book" year="2020">Kang 2020</bibRefCitation>
, 232–233, male (photographs), 
<collectingCountry id="F3117609FFC7FFA80F0B3E2FFCB8FCB5" box="[783,848,796,819]" name="South Korea" pageId="1" pageNumber="98">Korea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C6512FFC7FFAB0C933E52FDFBFEE3" lastPageId="2" lastPageNumber="99" pageId="1" pageNumber="98" type="diagnosis">
<paragraph id="8BB93699FFC7FFA80C933E52FDDBFAAA" blockId="1.[151,1437,865,2008]" pageId="1" pageNumber="98">
<emphasis id="B972EA8BFFC7FFA80C933E52FEFAFCFD" bold="true" box="[151,274,865,891]" pageId="1" pageNumber="98">Diagnosis.</emphasis>
This species belongs to the 
<taxonomicName id="4C064D1AFFC7FFA80E573E51FC94FCFA" authorityName="Iwata" authorityYear="1927" box="[595,892,866,892]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="nigrocephala">
<emphasis id="B972EA8BFFC7FFA80E573E51FC94FCFA" box="[595,892,866,892]" italics="true" pageId="1" pageNumber="98">Rhyacophila nigrocephala</emphasis>
</taxonomicName>
Species Group (
<bibRefCitation id="EF974B68FFC7FFA8083E3E51FB5EFCFB" author="Ross, H. H." box="[1082,1206,866,893]" pageId="1" pageNumber="98" refId="ref5765" refString="Ross, H. H. (1956) Evolution and classification of the mountain caddisflies. The University of Illinois Press, Urbana, Illinois, 213 pp." type="book" year="1956">Ross 1956</bibRefCitation>
). The male genitalia are similar to those of 
<taxonomicName id="4C064D1AFFC7FFA80D963EB4FDE5FC26" authorityName="Botosaneanu" authorityYear="1970" box="[402,525,902,928]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="confissa">
<emphasis id="B972EA8BFFC7FFA80D963EB4FDE5FC26" box="[402,525,902,928]" italics="true" pageId="1" pageNumber="98">R. confissa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C064D1AFFC7FFA80E473EB4FD4EFC26" authorityName="Botosaneanu" authorityYear="1970" box="[579,678,903,928]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B972EA8BFFC7FFA80E473EB4FD4EFC26" box="[579,678,903,928]" italics="true" pageId="1" pageNumber="98">R. vicina</emphasis>
</taxonomicName>
, but the male of 
<taxonomicName id="4C064D1AFFC7FFA80F603EB4FC3CFC26" authority="Nozaki &amp; Park, 2021" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[868,980,902,928]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="kangae" status="sp. nov.">
<emphasis id="B972EA8BFFC7FFA80F603EB4FC3CFC26" box="[868,980,902,928]" italics="true" pageId="1" pageNumber="98">R. kangae</emphasis>
</taxonomicName>
<emphasis id="B972EA8BFFC7FFA80FDF3EB5FBDCFC26" bold="true" box="[987,1076,902,928]" pageId="1" pageNumber="98">
<taxonomicNameLabel id="A24157F0FFC7FFA80FDF3EB5FBDCFC26" box="[987,1076,902,928]" pageId="1" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguishable from those of the latter two species by the shape of the complex of preanal appendages and apicodorsal lobe of segment IX: In dorsal aspect, the two lobes are widely separated from each other, forming a large U-shape in 
<taxonomicName id="4C064D1AFFC7FFA808CC3EFCFAD1FC6E" authority="Nozaki &amp; Park, 2021" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[1224,1337,974,1000]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="kangae" status="sp. nov.">
<emphasis id="B972EA8BFFC7FFA808CC3EFCFAD1FC6E" box="[1224,1337,974,1000]" italics="true" pageId="1" pageNumber="98">R. kangae</emphasis>
</taxonomicName>
<emphasis id="B972EA8BFFC7FFA809453EFDFA74FC6E" bold="true" box="[1345,1436,974,1000]" pageId="1" pageNumber="98">
<taxonomicNameLabel id="A24157F0FFC7FFA809453EFDFA74FC6E" box="[1345,1436,974,1000]" pageId="1" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="133D2A1CFFC7FFA80CA43EC1FF1EFB8A" box="[160,246,1010,1036]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Fig. 1B</figureCitation>
), but they form a narrow slit in 
<taxonomicName id="4C064D1AFFC7FFA80E5A3EC0FD31FB8A" authorityName="Botosaneanu" authorityYear="1970" box="[606,729,1010,1036]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="confissa">
<emphasis id="B972EA8BFFC7FFA80E5A3EC0FD31FB8A" box="[606,729,1010,1036]" italics="true" pageId="1" pageNumber="98">R. confissa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C064D1AFFC7FFA80F143EC0FC9CFB8A" authorityName="Botosaneanu" authorityYear="1970" box="[784,884,1011,1036]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B972EA8BFFC7FFA80F143EC0FC9CFB8A" box="[784,884,1011,1036]" italics="true" pageId="1" pageNumber="98">R. vicina</emphasis>
</taxonomicName>
(
<figureCitation id="133D2A1CFFC7FFA80F803EC1FBFEFB8A" box="[900,1046,1010,1036]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Figs. 2B, 3B</figureCitation>
). Furthermore, the male of 
<taxonomicName id="4C064D1AFFC7FFA8094F3EC0FF57FBB6" authority="Nozaki &amp; Park, 2021" authorityName="Nozaki &amp; Park" authorityYear="2021" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="kangae" status="sp. nov.">
<emphasis id="B972EA8BFFC7FFA8094F3EC0FF57FBB6" italics="true" pageId="1" pageNumber="98">R. kangae</emphasis>
</taxonomicName>
<emphasis id="B972EA8BFFC7FFA80CCC3925FECBFBB6" bold="true" box="[200,291,1046,1072]" pageId="1" pageNumber="98">
<taxonomicNameLabel id="A24157F0FFC7FFA80CCC3925FECBFBB6" box="[200,291,1046,1072]" pageId="1" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
bears large compound eyes in proportion to its head width when compared with those of 
<taxonomicName id="4C064D1AFFC7FFA809243924FA74FBB6" authorityName="Botosaneanu" authorityYear="1970" box="[1312,1436,1046,1072]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="confissa">
<emphasis id="B972EA8BFFC7FFA809243924FA74FBB6" box="[1312,1436,1046,1072]" italics="true" pageId="1" pageNumber="98">R. confissa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C064D1AFFC7FFA80CC23908FEC1FBD2" authorityName="Botosaneanu" authorityYear="1970" box="[198,297,1083,1108]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B972EA8BFFC7FFA80CC23908FEC1FBD2" box="[198,297,1083,1108]" italics="true" pageId="1" pageNumber="98">R. vicina</emphasis>
</taxonomicName>
: 
<quantity id="4CFE9B7CFFC7FFA80D3C3909FE2CFBD2" box="[312,452,1082,1109]" metricMagnitude="-2" metricUnit="m" metricValue="1.0414" metricValueMax="1.143" metricValueMin="0.9398" pageId="1" pageNumber="98" unit="in" value="0.41" valueMax="0.45" valueMin="0.37">0.37–0.45 in</quantity>
<taxonomicName id="4C064D1AFFC7FFA80DCF3908FDD2FBD2" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[459,570,1082,1108]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="kangae">
<emphasis id="B972EA8BFFC7FFA80DCF3908FDD2FBD2" box="[459,570,1082,1108]" italics="true" pageId="1" pageNumber="98">R. kangae</emphasis>
</taxonomicName>
, but 
<quantity id="4CFE9B7CFFC7FFA80E763909FD17FBD2" box="[626,767,1082,1108]" metricMagnitude="-3" metricUnit="m" metricValue="6.858" metricValueMax="7.874" metricValueMin="5.842" pageId="1" pageNumber="98" unit="in" value="0.27" valueMax="0.31" valueMin="0.23">0.23–0.31 in</quantity>
<taxonomicName id="4C064D1AFFC7FFA80F013908FC68FBD2" authorityName="Botosaneanu" authorityYear="1970" box="[773,896,1082,1108]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="confissa">
<emphasis id="B972EA8BFFC7FFA80F013908FC68FBD2" box="[773,896,1082,1108]" italics="true" pageId="1" pageNumber="98">R. confissa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C064D1AFFC7FFA80FB13908FBF0FBD2" authorityName="Botosaneanu" authorityYear="1970" box="[949,1048,1083,1108]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B972EA8BFFC7FFA80FB13908FBF0FBD2" box="[949,1048,1083,1108]" italics="true" pageId="1" pageNumber="98">R. vicina</emphasis>
</taxonomicName>
(
<figureCitation id="133D2A1CFFC7FFA808233909FAE3FBD2" box="[1063,1291,1082,1108]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Figs. 1F a b, 2C, 3C</figureCitation>
). The female genitalia are similar to those of 
<taxonomicName id="4C064D1AFFC7FFA80E05396CFD95FBFE" authorityName="Botosaneanu" authorityYear="1970" box="[513,637,1118,1144]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="confissa">
<emphasis id="B972EA8BFFC7FFA80E05396CFD95FBFE" box="[513,637,1118,1144]" italics="true" pageId="1" pageNumber="98">R. confissa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C064D1AFFC7FFA80EB2396CFCF2FBFE" authorityName="Botosaneanu" authorityYear="1970" box="[694,794,1119,1144]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B972EA8BFFC7FFA80EB2396CFCF2FBFE" box="[694,794,1119,1144]" italics="true" pageId="1" pageNumber="98">R. vicina</emphasis>
</taxonomicName>
but can be distinguished by the shape of the vaginal apparatus: In ventral aspect, the posterior process of the vaginal apparatus is long and rectangular in 
<taxonomicName id="4C064D1AFFC7FFA8090439B0FA9AFB1A" authority="Nozaki &amp; Park, 2021" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[1280,1394,1154,1180]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="kangae" status="sp. nov.">
<emphasis id="B972EA8BFFC7FFA8090439B0FA9AFB1A" box="[1280,1394,1154,1180]" italics="true" pageId="1" pageNumber="98">R. kangae</emphasis>
</taxonomicName>
<emphasis id="B972EA8BFFC7FFA8097F39B1FF20FB46" bold="true" pageId="1" pageNumber="98">
<taxonomicNameLabel id="A24157F0FFC7FFA8097F39B1FF20FB46" pageId="1" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="133D2A1CFFC7FFA80CDC3995FEC7FB46" box="[216,303,1190,1216]" captionStart="FIGURE 4–7" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 4–7. Rhyacophila spp., female. 4A–4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A–5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="1" pageNumber="98">Fig. 4F</figureCitation>
), but that of 
<taxonomicName id="4C064D1AFFC7FFA80DC43994FDD4FB46" authorityName="Botosaneanu" authorityYear="1970" box="[448,572,1190,1216]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="confissa">
<emphasis id="B972EA8BFFC7FFA80DC43994FDD4FB46" box="[448,572,1190,1216]" italics="true" pageId="1" pageNumber="98">R. confissa</emphasis>
</taxonomicName>
bears an 8-shaped unpigmented part apicoventrally (
<figureCitation id="133D2A1CFFC7FFA808903995FB05FB47" box="[1172,1261,1190,1217]" captionStart="FIGURE 4–7" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 4–7. Rhyacophila spp., female. 4A–4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A–5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="1" pageNumber="98">Fig. 6B</figureCitation>
), and that of 
<taxonomicName id="4C064D1AFFC7FFA809803994FF33FB62" authorityName="Botosaneanu" authorityYear="1970" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B972EA8BFFC7FFA809803994FF33FB62" italics="true" pageId="1" pageNumber="98">R. vicina</emphasis>
</taxonomicName>
is constricted at 2/5 from the posterior apex (
<figureCitation id="133D2A1CFFC7FFA80EE039F9FCD6FB62" box="[740,830,1226,1252]" captionStart="FIGURE 4–7" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 4–7. Rhyacophila spp., female. 4A–4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A–5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="1" pageNumber="98">Fig. 7B</figureCitation>
). The male and female of 
<taxonomicName id="4C064D1AFFC7FFA8086D39F8FB32FB62" authority="Nozaki &amp; Park, 2021" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[1129,1242,1226,1252]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="kangae" status="sp. nov.">
<emphasis id="B972EA8BFFC7FFA8086D39F8FB32FB62" box="[1129,1242,1226,1252]" italics="true" pageId="1" pageNumber="98">R. kangae</emphasis>
</taxonomicName>
<emphasis id="B972EA8BFFC7FFA808E639F9FAD5FB62" bold="true" box="[1250,1341,1226,1252]" pageId="1" pageNumber="98">
<taxonomicNameLabel id="A24157F0FFC7FFA808E639F9FAD5FB62" box="[1250,1341,1226,1252]" pageId="1" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are also similar to those of a Japanese species, 
<taxonomicName id="4C064D1AFFC7FFA80E4C39DCFD0BFA8E" authority="Nozaki &amp; Park, 2021" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[584,739,1263,1288]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="98" phylum="Arthropoda" rank="species" species="yamamotoi" status="sp. nov.">
<emphasis id="B972EA8BFFC7FFA80E4C39DCFD0BFA8E" box="[584,739,1263,1288]" italics="true" pageId="1" pageNumber="98">R. yamamotoi</emphasis>
</taxonomicName>
<emphasis id="B972EA8BFFC7FFA80EEF39DDFCADFA8E" bold="true" box="[747,837,1262,1288]" pageId="1" pageNumber="98">
<taxonomicNameLabel id="A24157F0FFC7FFA80EEF39DDFCADFA8E" box="[747,837,1262,1288]" pageId="1" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but they are distinguished by the characters given in the diagnosis for that species, below.
</paragraph>
<paragraph id="8BB93699FFC7FFA80CC33806FC93FA1F" blockId="1.[151,1437,865,2008]" pageId="1" pageNumber="98">
<emphasis id="B972EA8BFFC7FFA80CC33806FEFBFAC9" bold="true" box="[199,275,1333,1359]" pageId="1" pageNumber="98">Adult.</emphasis>
Specimens in alcohol mostly dark brown, but tibiae and tarsi of all legs light brown. Forewings each 
<quantity id="4CFE9B7CFFC7FFA80C933869FED3FAF2" box="[151,315,1370,1397]" metricMagnitude="-2" metricUnit="m" metricValue="1.285" metricValueMax="1.35" metricValueMin="1.22" pageId="1" pageNumber="98" unit="mm" value="12.85" valueMax="13.5" valueMin="12.2">12.2–13.5 mm</quantity>
in male (n = 6), 
<quantity id="4CFE9B7CFFC7FFA80DFD3869FD6FFAF2" box="[505,647,1370,1396]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" metricValueMax="1.48" metricValueMin="1.3" pageId="1" pageNumber="98" unit="mm" value="13.9" valueMax="14.8" valueMin="13.0">13–14.8 mm</quantity>
in female (n = 2). Ratio of width of eye to distance between eyes (a/b in 
<figureCitation id="133D2A1CFFC7FFA80CB0384DFEEDFA1E" box="[180,261,1406,1432]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Fig. 1F</figureCitation>
) 
<quantity id="4CFE9B7CFFC7FFA80D11384DFE4AFA1E" box="[277,418,1406,1433]" metricMagnitude="-2" metricUnit="m" metricValue="1.0414" metricValueMax="1.143" metricValueMin="0.9398" pageId="1" pageNumber="98" unit="in" value="0.41" valueMax="0.45" valueMin="0.37">0.37–0.45 in</quantity>
male (n = 6), 
<quantity id="4CFE9B7CFFC7FFA80E45384DFD26FA1E" box="[577,718,1406,1433]" metricMagnitude="-3" metricUnit="m" metricValue="7.366" metricValueMax="8.382" metricValueMin="6.35" pageId="1" pageNumber="98" unit="in" value="0.29" valueMax="0.33" valueMin="0.25">0.25–0.33 in</quantity>
female (n = 6).
</paragraph>
<paragraph id="8BB93699FFC7FFA80CC33891FD35F8EA" blockId="1.[151,1437,865,2008]" pageId="1" pageNumber="98">
<emphasis id="B972EA8BFFC7FFA80CC33891FE91FA3D" bold="true" box="[199,377,1441,1468]" pageId="1" pageNumber="98">Male genitalia.</emphasis>
Segment IX (IX) longitudinally short in lateral aspect (
<figureCitation id="133D2A1CFFC7FFA80FEA3891FBA0FA3A" box="[1006,1096,1442,1468]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Fig. 1A</figureCitation>
), ventral half longer than dorsum; membranous along midline in dorsal aspect (
<figureCitation id="133D2A1CFFC7FFA80EC238F5FCF4FA66" box="[710,796,1478,1504]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Fig. 1B</figureCitation>
). Complex of pair of preanal appendages and apicodorsal lobe of segment IX (com.) bilobed from about basal 2/5, widely separated, large, U-shaped in dorsal aspect (
<figureCitation id="133D2A1CFFC7FFA8096B38D9FF51F9AE" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Fig. 1B</figureCitation>
); each lobe finger-like in lateral aspect (
<figureCitation id="133D2A1CFFC7FFA80E833B3DFD09F9AE" box="[647,737,1550,1576]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Fig. 1A</figureCitation>
), with 3 tiny teeth apicomesally (2 dorsal, 1 ventral, inset for 
<figureCitation id="133D2A1CFFC7FFA80C933B01FF07F9CA" box="[151,239,1586,1612]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Fig. 1A</figureCitation>
). Anal sclerites (a.s.) fused basally, bilobed apically, each round apex with minute black denticles dorsally. Apical band (a.b.) long, curved posterad, dorsal margin connected to base of anal sclerite; ventral margin connected to base of sagittal appendage (s.a.) of tergal band (
<figureCitation id="133D2A1CFFC7FFA80ECD3B49FCC9F912" box="[713,801,1658,1684]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Fig. 1A</figureCitation>
); sagittal appendage trapezoidal in dorsal and ventral aspects (
<figureCitation id="133D2A1CFFC7FFA80CDB3BADFEDEF93E" box="[223,310,1694,1720]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Fig. 1C</figureCitation>
). Basal segment of each inferior appendage (i.a.) thick in lateral aspect (
<figureCitation id="133D2A1CFFC7FFA8085C3BADFB59F93E" box="[1112,1201,1694,1720]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Fig. 1A</figureCitation>
); posteromesal angle triangular, directed mesad in dorsal aspect (
<figureCitation id="133D2A1CFFC7FFA80E7F3BF1FD39F95A" box="[635,721,1730,1756]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Fig. 1B</figureCitation>
); mesal face with ridged tendon (t.i.a.) along midline, extending to phallotheca (pha.) (
<figureCitation id="133D2A1CFFC7FFA80D903BD5FDCFF886" box="[404,551,1766,1792]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Figs 1A, 1D</figureCitation>
). Distal segment of each inferior appendage bilobed, dorsal lobe club-like in lateral aspect (
<figureCitation id="133D2A1CFFC7FFA80D3C3A39FE2BF8A2" box="[312,451,1802,1828]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Figs 1A, 1D</figureCitation>
), ventral lobe semicircular in lateral aspect, both lobes with minute spines apicomesally. In phallic apparatus (
<figureCitation id="133D2A1CFFC7FFA80D863A1DFE3FF8CE" box="[386,471,1838,1864]" captionStart="FIGURES 1–3" captionStartId="4.[151,264,1766,1790]" captionTargetBox="[151,1436,210,1734]" captionTargetId="figure-28@4.[151,1436,198,1740]" captionTargetPageId="4" captionText="FIGURES 1–3. Rhyacophila spp., male. 1A–1F, Rhyacophila kangae sp. nov. 1A–1E, genitalia: 1A, left lateral, apicomesal part of complex of preanal appendages and apicodorsal lobe of segment IX enlarged; 1B, dorsal; 1C, upper part, ventral; 1D, left inferior appendage, mesal; 1E, phallic apparatus, dorsal. 1F, head, dorsal (a, b: see text). 2A–2C, Rhyacophila confissa Botosaneanu 1970: 2A, genitalia, left lateral; 2B, same, dorsal, anal sclerite enlarged; 2C, head, dorsal. 3A–3C, Rhyacophila vicina Botosaneanu 1970: 3A, genitalia, left lateral; 3B, same, dorsal, anal sclerite enlarged; 3C, head, dorsal. Abbreviations: a.b. = apical band, ae. = aedeagus, a.s. = anal sclerite, com = complex of preanal appendages and apicodorsal lobe of segment IX, i.a. = inferior appendage (paired), para. = paramere (paired), pha. = phallotheca, s.a. = sagittal appendage, t.b. = tergal band, t.i.a., = tendon of inferior appendage (paired), IX = segment IX." figureDoi="http://doi.org/10.5281/zenodo.5656554" httpUri="https://zenodo.org/record/5656554/files/figure.png" pageId="1" pageNumber="98">Fig. 1E</figureCitation>
), aedeagus (ae.) bottle-shaped in dorsal and ventral aspects; parameres (para.) slender, club-like, apices curve laterad, and with tiny spines.
</paragraph>
<paragraph id="8BB93699FFC7FFAB0CC33A45FBD5FEC7" blockId="1.[151,1437,865,2008]" lastBlockId="2.[151,1437,151,1041]" lastPageId="2" lastPageNumber="99" pageId="1" pageNumber="98">
<emphasis id="B972EA8BFFC7FFA80CC33A45FE7BF809" bold="true" box="[199,403,1909,1936]" pageId="1" pageNumber="98">Female genitalia.</emphasis>
Segment VIII (VIII) annular, but semi-membranous along dorsal and ventral midlines, with pair of dorsal weakly sclerotized extensions (
<figureCitation id="133D2A1CFFC7FFA80E8B3AA9FD00F832" box="[655,744,1946,1972]" captionStart="FIGURE 4–7" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 4–7. Rhyacophila spp., female. 4A–4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A–5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="1" pageNumber="98">Fig. 4C</figureCitation>
); pair of apodemal rods reaching posterior end of segment VI. Segment IX (IX) membranous, with pair of strongly sclerotized bands dorsolaterally extending into segment VI as apodemal rods (
<figureCitation id="133D2A1CFFC4FFAB0D4D3DA4FE48FF37" box="[329,416,151,177]" captionStart="FIGURE 4–7" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 4–7. Rhyacophila spp., female. 4A–4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A–5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="2" pageNumber="99">Fig. 4B</figureCitation>
, ap. IX); fused with segment X (X) ventrally, pair of ventrolateral sclerotized bands extending to segment X (
<figureCitation id="133D2A1CFFC4FFAB0D6F3D88FE2DFF53" box="[363,453,187,213]" captionStart="FIGURE 4–7" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 4–7. Rhyacophila spp., female. 4A–4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A–5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="2" pageNumber="99">Fig. 4B</figureCitation>
). Segment X slender, with pair of strongly sclerotized bands dorsolaterally; fused with segment XI (XI) (
<figureCitation id="133D2A1CFFC4FFAB0D5A3DECFE5EFF7F" box="[350,438,223,249]" captionStart="FIGURE 4–7" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 4–7. Rhyacophila spp., female. 4A–4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A–5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="2" pageNumber="99">Fig. 4B</figureCitation>
). In vaginal apparatus, processus spermathecae (p.s.) claw-like in lateral aspect (
<figureCitation id="133D2A1CFFC4FFAB093D3DECFA67FF7F" box="[1337,1423,223,249]" captionStart="FIGURE 4–7" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 4–7. Rhyacophila spp., female. 4A–4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A–5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="2" pageNumber="99">Fig. 4E</figureCitation>
), semicircular in ventral aspect (
<figureCitation id="133D2A1CFFC4FFAB0DEF3C30FDA8FE9B" box="[491,576,259,285]" captionStart="FIGURE 4–7" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 4–7. Rhyacophila spp., female. 4A–4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A–5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="2" pageNumber="99">Fig. 4F</figureCitation>
); posterior process (p.p.) long rectangular in ventral aspect, more than 3 times longer than anterior width, face-down and saucer-shaped in lateral aspect (
<figureCitation id="133D2A1CFFC4FFAB0FDD3C14FBD9FEC7" box="[985,1073,295,321]" captionStart="FIGURE 4–7" captionStartId="5.[151,250,1490,1514]" captionTargetBox="[165,1423,190,1455]" captionTargetId="figure-19@5.[151,1436,181,1466]" captionTargetPageId="5" captionText="FIGURE 4–7. Rhyacophila spp., female. 4A–4F, Rhyacophila kangae sp. nov.: 4A, head, dorsal; 4B, abdominal segments VIII to XI, left lateral, 4C, abdominal segment VIII, dorsal; 4D, same, ventral; 4E, vaginal apparatus, left lateral; 4F, same, ventral. 5A–5E, Rhyacophila yamamotoi sp. nov.: 5A, head dorsal; 5B, abdominal segment VIII, left lateral; 5C, same, dorsal; 5D, same, ventral; 5E, vaginal apparatus, left lateral; 5F, same, ventral. 6A, 6B, Rhyacophila confissa Botosaneanu 1970: 6A, vaginal apparatus, left lateral; 6B, same, ventral. 7A, 7B, Rhyacophila vicina Botosaneanu 1970: 7A, vaginal apparatus, left lateral; 7B, same, ventral. Abbreviations: ap.VIII = apodemal rod of abdominal segment VIII (paired); ap.IX = apodemal rod of abdominal segment IX (paired); p.p. = posterior process; p.s. = processus spermathecae; VIII, IX, X, XI = abdominal segments VIII, IX, X, XI." figureDoi="http://doi.org/10.5281/zenodo.5656559" httpUri="https://zenodo.org/record/5656559/files/figure.png" pageId="2" pageNumber="99">Fig. 4E</figureCitation>
).
</paragraph>
<paragraph id="8BB93699FFC4FFAB0CC33C78FDFBFEE3" blockId="2.[151,1437,151,1041]" box="[199,531,331,357]" pageId="2" pageNumber="99">
<emphasis id="B972EA8BFFC4FFAB0CC33C78FE7EFEE3" bold="true" box="[199,406,331,357]" pageId="2" pageNumber="99">Immature stages.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31C6512FFC4FFAB0CC33C5CFCD9FDE7" pageId="2" pageNumber="99" type="materials_examined">
<paragraph id="8BB93699FFC4FFAB0CC33C5CFB4AFE2B" blockId="2.[151,1437,151,1041]" pageId="2" pageNumber="99">
<materialsCitation id="3B6E3CC4FFC4FFAB0CC33C5CFB4AFE2B" ID-GBIF-Occurrence="3400170321" collectingDate="2018-05-23" collectionCode="NIBR" country="South Korea" county="Gangwon-do" elevation="690" latitude="38.024" location="Male" longLatPrecision="70" longitude="128.472" municipality="Inje-gun" pageId="2" pageNumber="99" specimenCode="NIBR0000935610" specimenCount="1" typeStatus="holotype">
<emphasis id="B972EA8BFFC4FFAB0CC33C5CFED3FE0F" bold="true" box="[199,315,367,393]" pageId="2" pageNumber="99">
<typeStatus id="54BD883BFFC4FFAB0CC33C5CFEDEFE0F" box="[199,310,367,393]" pageId="2" pageNumber="99" type="holotype">Holotype</typeStatus>
.
</emphasis>
<location id="8ED96042FFC4FFAB0D403C5CFE96FE0F" LSID="urn:lsid:plazi:treatment:03AF878FFFC7FFAB0C933F80FD54FB97:8ED96042FFC4FFAB0D403C5CFE96FE0F" box="[324,382,367,393]" country="South Korea" county="Gangwon-do" latitude="38.024" longLatPrecision="70" longitude="128.472" municipality="Inje-gun" name="Male" pageId="2" pageNumber="99">Male</location>
(in alcohol), 
<location id="8ED96042FFC4FFAB0E1C3C5CFCF9FE0F" LSID="urn:lsid:plazi:treatment:03AF878FFFC7FFAB0C933F80FD54FB97:8ED96042FFC4FFAB0E1C3C5CFCF9FE0F" box="[536,785,367,393]" country="South Korea" county="Gangwon-do" latitude="38.024" longLatPrecision="70" longitude="128.472" municipality="Inje-gun" name="Bangtaecheon Stream" pageId="2" pageNumber="99">Bangtaecheon Stream</location>
, 
<location id="8ED96042FFC4FFAB0F1A3C5CFC7CFE0F" LSID="urn:lsid:plazi:treatment:03AF878FFFC7FFAB0C933F80FD54FB97:8ED96042FFC4FFAB0F1A3C5CFC7CFE0F" box="[798,916,367,393]" country="South Korea" county="Gangwon-do" latitude="38.024" longLatPrecision="70" longitude="128.472" municipality="Inje-gun" name="Jindong-ri" pageId="2" pageNumber="99">Jindong-ri</location>
, 
<location id="8ED96042FFC4FFAB0FA53C5CFBDDFE0F" LSID="urn:lsid:plazi:treatment:03AF878FFFC7FFAB0C933F80FD54FB97:8ED96042FFC4FFAB0FA53C5CFBDDFE0F" box="[929,1077,367,393]" country="South Korea" county="Gangwon-do" latitude="38.024" longLatPrecision="70" longitude="128.472" municipality="Inje-gun" name="Girin-myeon" pageId="2" pageNumber="99">Girin-myeon</location>
, 
<collectingMunicipality id="6BDDACE3FFC4FFAB08453C5CFB4BFE0F" box="[1089,1187,367,393]" pageId="2" pageNumber="99">Inje-gun</collectingMunicipality>
, 
<collectingCounty id="62D84E15FFC4FFAB08AB3C5CFAACFE0F" box="[1199,1348,367,393]" pageId="2" pageNumber="99">Gangwon-do</collectingCounty>
, 
<collectingCountry id="F3117609FFC4FFAB09553C5CFA70FE0F" box="[1361,1432,367,393]" name="South Korea" pageId="2" pageNumber="99">Korea</collectingCountry>
, 
<geoCoordinate id="EE32505EFFC4FFAB0C933CA0FEEEFE2B" box="[151,262,403,429]" degrees="38.024" direction="north" orientation="latitude" pageId="2" pageNumber="99" precision="55" value="38.024">38.024°N</geoCoordinate>
, 
<geoCoordinate id="EE32505EFFC4FFAB0D163CA0FE64FE2B" box="[274,396,403,429]" degrees="128.472" direction="east" orientation="longitude" pageId="2" pageNumber="99" precision="55" value="128.472">128.472°E</geoCoordinate>
, alt. 
<quantity id="4CFE9B7CFFC4FFAB0DC53CA0FDE2FE2B" box="[449,522,403,429]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="2" pageNumber="99" unit="m" value="690.0">
<elevation id="002BD1AAFFC4FFAB0DC53CA0FDE2FE2B" box="[449,522,403,429]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="2" pageNumber="99" unit="m" value="690.0">690 m</elevation>
</quantity>
, 
<date id="FFB81059FFC4FFAB0E123CA0FD6FFE2B" box="[534,647,403,429]" pageId="2" pageNumber="99" value="2018-05-23">
<collectingDate id="EFFCE9B1FFC4FFAB0E123CA0FD6FFE2B" box="[534,647,403,429]" pageId="2" pageNumber="99" value="2018-05-23">23.v.2018</collectingDate>
</date>
, MS. Kang (net sweeping) (
<specimenCode id="DBA09EE2FFC4FFAB0FC43CA1FB7FFE2B" box="[960,1175,402,429]" collectionCode="NIBR" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="2" pageNumber="99">NIBR0000935610</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BB93699FFC4FFAB0CC33C84FE92FD9C" blockId="2.[151,1437,151,1041]" pageId="2" pageNumber="99">
<materialsCitation id="3B6E3CC4FFC4FFAB0CC33C84FB2AFE57" ID-GBIF-Occurrence="3400170324" box="[199,1218,438,465]" collectingDate="2018-05-23" collectionCode="NIBR" country="South Korea" county="Gangwon-do" elevation="690" latitude="38.024" location="Male" longLatPrecision="70" longitude="128.472" municipality="Inje-gun" pageId="2" pageNumber="99" specimenCode="NIBR0000935608-0000935609" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B972EA8BFFC4FFAB0CC33C84FEAEFE57" bold="true" box="[199,326,439,465]" pageId="2" pageNumber="99">
<typeStatus id="54BD883BFFC4FFAB0CC33C84FEA9FE57" box="[199,321,439,465]" pageId="2" pageNumber="99" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D00FD10FFC4FFAB0D4A3C84FE74FE57" box="[334,412,439,465]" pageId="2" pageNumber="99" type="male">1 male</specimenCount>
, 
<specimenCount id="9D00FD10FFC4FFAB0DAC3C84FDE5FE57" box="[424,525,439,465]" pageId="2" pageNumber="99" type="female">1 female</specimenCount>
, same data as the holotype (
<specimenCode id="DBA09EE2FFC4FFAB0F4E3C85FB52FE57" box="[842,1210,438,465]" collectionCode="NIBR" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="2" pageNumber="99">NIBR0000935608–0000935609</specimenCode>
)
</materialsCitation>
; 
<materialsCitation id="3B6E3CC4FFC4FFAB08CA3C84FBD4FE73" ID-GBIF-Occurrence="3400170306" collectingDate="2014-06-10" collectionCode="T" country="South Korea" county="Gangwon-do" location="Samil River" municipality="Hwacheon-gun" pageId="2" pageNumber="99" specimenCount="5" specimenCount-female="1" specimenCount-male="4" typeStatus="paratype">
<specimenCount id="9D00FD10FFC4FFAB08CA3C84FACFFE57" box="[1230,1319,439,465]" pageId="2" pageNumber="99" type="male">4 males</specimenCount>
, 
<specimenCount id="9D00FD10FFC4FFAB09373C84FA70FE57" box="[1331,1432,439,465]" pageId="2" pageNumber="99" type="female">1 female</specimenCount>
, 
<location id="8ED96042FFC4FFAB0C933CE8FECDFE73" LSID="urn:lsid:plazi:treatment:03AF878FFFC7FFAB0C933F80FD54FB97:8ED96042FFC4FFAB0C933CE8FECDFE73" box="[151,293,475,501]" country="South Korea" county="Gangwon-do" municipality="Hwacheon-gun" name="Samil River" pageId="2" pageNumber="99">Samil River</location>
, 
<location id="8ED96042FFC4FFAB0D323CE8FE39FE73" LSID="urn:lsid:plazi:treatment:03AF878FFFC7FFAB0C933F80FD54FB97:8ED96042FFC4FFAB0D323CE8FE39FE73" box="[310,465,475,501]" country="South Korea" county="Gangwon-do" municipality="Hwacheon-gun" name="Sanae-myeon" pageId="2" pageNumber="99">Sanae-myeon</location>
, 
<collectingMunicipality id="6BDDACE3FFC4FFAB0DE63CE8FD78FE73" box="[482,656,475,501]" pageId="2" pageNumber="99">Hwacheon-gun</collectingMunicipality>
, 
<collectingCounty id="62D84E15FFC4FFAB0EA53CE8FCDEFE73" box="[673,822,475,501]" pageId="2" pageNumber="99">Gangwon-do</collectingCounty>
, 
<collectingCountry id="F3117609FFC4FFAB0F433CE8FC66FE73" box="[839,910,475,501]" name="South Korea" pageId="2" pageNumber="99">Korea</collectingCountry>
, 
<date id="FFB81059FFC4FFAB0F9B3CE8FBF1FE73" box="[927,1049,475,501]" pageId="2" pageNumber="99" value="2014-06-10">
<collectingDate id="EFFCE9B1FFC4FFAB0F9B3CE8FBF1FE73" box="[927,1049,475,501]" pageId="2" pageNumber="99" value="2014-06-10">10.vi.2014</collectingDate>
</date>
, 
<collectionCode id="ED17AE5CFFC4FFAB082E3CE8FBD4FE73" box="[1066,1084,475,501]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="99">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="3B6E3CC4FFC4FFAB08493CE8FE9EFD9C" ID-GBIF-Occurrence="3400170312" collectionCode="CBM-ZI" country="South Korea" pageId="2" pageNumber="99" specimenCode="CBM-ZI 0180236-0180240" specimenCount="1" typeStatus="paratype">
Ito (net sweeping) (
<specimenCode id="DBA09EE2FFC4FFAB093C3CE8FE87FD9C" collectionCode="CBM-ZI" pageId="2" pageNumber="99">CBM-ZI 0180236–0180240</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB93699FFC4FFAB0CC33F11FCD9FDE7" blockId="2.[151,1437,151,1041]" pageId="2" pageNumber="99">
<emphasis id="B972EA8BFFC4FFAB0CC33F11FDFAFDBA" bold="true" box="[199,530,546,572]" pageId="2" pageNumber="99">Other specimens examined.</emphasis>
<materialsCitation id="3B6E3CC4FFC4FFAB0E1E3F10FCC5FDE7" ID-GBIF-Occurrence="3400170319" collectingDate="2014-06-11" collectionCode="T, SJP" country="South Korea" location="Gyebangsan" municipality="Gangwon-do" pageId="2" pageNumber="99" specimenCount="12" specimenCount-female="8" specimenCount-male="4">
<collectingCountry id="F3117609FFC4FFAB0E1E3F10FD97FDBB" box="[538,639,547,573]" name="South Korea" pageId="2" pageNumber="99">KOREA</collectingCountry>
: 
<specimenCount id="9D00FD10FFC4FFAB0E8E3F10FD0CFDBB" box="[650,740,547,573]" pageId="2" pageNumber="99" type="male">2 males</specimenCount>
, 
<specimenCount id="9D00FD10FFC4FFAB0EF43F10FC88FDBB" box="[752,864,547,573]" pageId="2" pageNumber="99" type="female">4 females</specimenCount>
, 
<location id="8ED96042FFC4FFAB0F683F10FC11FDBB" LSID="urn:lsid:plazi:treatment:03AF878FFFC7FFAB0C933F80FD54FB97:8ED96042FFC4FFAB0F683F10FC11FDBB" box="[876,1017,547,573]" country="South Korea" municipality="Gangwon-do" name="Gyebangsan" pageId="2" pageNumber="99">Gyebangsan</location>
, 
<collectingMunicipality id="6BDDACE3FFC4FFAB08013F10FB72FDBB" box="[1029,1178,547,573]" pageId="2" pageNumber="99">Gangwon-do</collectingMunicipality>
, 
<date id="FFB81059FFC4FFAB08A23F10FAF7FDBB" box="[1190,1311,547,573]" pageId="2" pageNumber="99" value="2014-06-11">
<collectingDate id="EFFCE9B1FFC4FFAB08A23F10FAF7FDBB" box="[1190,1311,547,573]" pageId="2" pageNumber="99" value="2014-06-11">11.vi.2014</collectingDate>
</date>
, 
<collectionCode id="ED17AE5CFFC4FFAB09283F10FAD6FDBB" box="[1324,1342,547,573]" name="Tavera, Department of Geology and Geophysics" pageId="2" pageNumber="99">T</collectionCode>
. Ito (net sweeping) (
<specimenCount id="9D00FD10FFC4FFAB0D183F74FE81FDE7" box="[284,361,583,609]" pageId="2" pageNumber="99" type="male">1 male</specimenCount>
, 
<specimenCount id="9D00FD10FFC4FFAB0D703F74FE0DFDE7" box="[372,485,583,609]" pageId="2" pageNumber="99" type="female">2 females</specimenCount>
: 
<collectionCode id="ED17AE5CFFC4FFAB0DEB3F74FDF4FDE7" box="[495,540,583,609]" pageId="2" pageNumber="99">SJP</collectionCode>
; 
<specimenCount id="9D00FD10FFC4FFAB0E2C3F74FD9EFDE7" box="[552,630,583,609]" pageId="2" pageNumber="99" type="male">1 male</specimenCount>
, 
<specimenCount id="9D00FD10FFC4FFAB0E853F74FD1AFDE7" box="[641,754,583,609]" pageId="2" pageNumber="99" type="female">2 females</specimenCount>
: TN)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C6512FFC4FFAB0CC33F58FD1AFD2F" pageId="2" pageNumber="99" type="etymology">
<paragraph id="8BB93699FFC4FFAB0CC33F58FD1AFD2F" blockId="2.[151,1437,151,1041]" pageId="2" pageNumber="99">
<emphasis id="B972EA8BFFC4FFAB0CC33F58FEA6FD03" bold="true" box="[199,334,619,645]" pageId="2" pageNumber="99">Etymology.</emphasis>
This species is dedicated to Ms. Mi-Sook Kang, who gifted us valuable specimens and information concerning Korean caddisflies, including this species.
</paragraph>
</subSubSection>
<subSubSection id="C31C6512FFC4FFAB0CC33F81FC46FD4B" box="[199,942,690,717]" pageId="2" pageNumber="99" type="distribution">
<paragraph id="8BB93699FFC4FFAB0CC33F81FC46FD4B" blockId="2.[151,1437,151,1041]" box="[199,942,690,717]" pageId="2" pageNumber="99">
<emphasis id="B972EA8BFFC4FFAB0CC33F81FE89FD4A" bold="true" box="[199,353,690,716]" pageId="2" pageNumber="99">Distribution.</emphasis>
The Korean Peninsula (Gangwon-do, 
<collectingCountry id="F3117609FFC4FFAB0F143F80FC4BFD4B" box="[784,931,691,717]" name="South Korea" pageId="2" pageNumber="99">South Korea</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C31C6512FFC4FFAB0CC33FE4FD54FB97" pageId="2" pageNumber="99" type="discussion">
<paragraph id="8BB93699FFC4FFAB0CC33FE4FF08FC07" blockId="2.[151,1437,151,1041]" pageId="2" pageNumber="99">
<emphasis id="B972EA8BFFC4FFAB0CC33FE4FED3FD77" bold="true" box="[199,315,727,753]" pageId="2" pageNumber="99">Remarks.</emphasis>
This species was found in Gangwon-do, 
<collectingCountry id="F3117609FFC4FFAB0F083FE4FC73FD77" box="[780,923,727,753]" name="South Korea" pageId="2" pageNumber="99">South Korea</collectingCountry>
, where both 
<taxonomicName id="4C064D1AFFC4FFAB08283FE4FB4FFD77" authorityName="Botosaneanu" authorityYear="1970" box="[1068,1191,727,753]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="confissa">
<emphasis id="B972EA8BFFC4FFAB08283FE4FB4FFD77" box="[1068,1191,727,753]" italics="true" pageId="2" pageNumber="99">R. confissa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C064D1AFFC4FFAB08DA3FE4FAA9FD76" authorityName="Botosaneanu" authorityYear="1970" box="[1246,1345,727,752]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B972EA8BFFC4FFAB08DA3FE4FAA9FD76" box="[1246,1345,727,752]" italics="true" pageId="2" pageNumber="99">R. vicina</emphasis>
</taxonomicName>
are distributed (
<figureCitation id="133D2A1CFFC4FFAB0CF93FC8FEA9FC93" box="[253,321,763,789]" captionStart="FIGURE 9" captionStartId="7.[151,250,1110,1134]" captionTargetBox="[155,1428,281,1079]" captionTargetId="figure-64@7.[151,1436,273,1087]" captionTargetPageId="7" captionText="FIGURE 9. Distributions of 4 Rhyacophila species in East Asia. Distribution areas of R. confissa and R. vicina are cited from Ivanov (2011), Park &amp; Kong (2020), and Yang et al. (2016)." figureDoi="http://doi.org/10.5281/zenodo.5656565" httpUri="https://zenodo.org/record/5656565/files/figure.png" pageId="2" pageNumber="99">Fig. 9</figureCitation>
). One of the sites where 
<taxonomicName id="4C064D1AFFC4FFAB0E533FC8FD39FC93" authorityName="Botosaneanu" authorityYear="1970" box="[599,721,763,789]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="confissa">
<emphasis id="B972EA8BFFC4FFAB0E533FC8FD39FC93" box="[599,721,763,789]" italics="true" pageId="2" pageNumber="99">R. confissa</emphasis>
</taxonomicName>
was collected in this study is only 
<quantity id="4CFE9B7CFFC4FFAB08533FC8FB4DFC93" box="[1111,1189,763,789]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="99" unit="km" value="1.4">1.4 km</quantity>
from the 
<typeStatus id="54BD883BFFC4FFAB09153FC8FAA9FC93" box="[1297,1345,763,789]" pageId="2" pageNumber="99">type</typeStatus>
locality of 
<taxonomicName id="4C064D1AFFC4FFAB0CB33E2CFEC1FCBF" authority="Nozaki &amp; Park, 2021" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[183,297,799,825]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="kangae" status="sp. nov.">
<emphasis id="B972EA8BFFC4FFAB0CB33E2CFEC1FCBF" box="[183,297,799,825]" italics="true" pageId="2" pageNumber="99">R. kangae</emphasis>
</taxonomicName>
<emphasis id="B972EA8BFFC4FFAB0D353E2CFE65FCBF" bold="true" box="[305,397,799,825]" pageId="2" pageNumber="99">
<taxonomicNameLabel id="A24157F0FFC4FFAB0D353E2CFE65FCBF" box="[305,397,799,825]" pageId="2" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the same stream (Bangtaecheon Stream). Although the genitalic morphology of the new species is similar to those of 
<taxonomicName id="4C064D1AFFC4FFAB0DD83E70FDB0FCDB" authorityName="Botosaneanu" authorityYear="1970" box="[476,600,835,861]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="confissa">
<emphasis id="B972EA8BFFC4FFAB0DD83E70FDB0FCDB" box="[476,600,835,861]" italics="true" pageId="2" pageNumber="99">R. confissa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C064D1AFFC4FFAB0E8A3E70FD1AFCDA" authorityName="Botosaneanu" authorityYear="1970" box="[654,754,835,860]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B972EA8BFFC4FFAB0E8A3E70FD1AFCDA" box="[654,754,835,860]" italics="true" pageId="2" pageNumber="99">R. vicina</emphasis>
</taxonomicName>
, the differences in diagnostic characters described above are stable.
</paragraph>
<paragraph id="8BB93699FFC4FFAB0CC33EB8FD54FB97" blockId="2.[151,1437,151,1041]" pageId="2" pageNumber="99">
The male of 
<taxonomicName id="4C064D1AFFC4FFAB0D573EB8FE2BFC23" authority="Nozaki &amp; Park, 2021" authorityName="Nozaki &amp; Park" authorityYear="2021" box="[339,451,907,933]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="kangae" status="sp. nov.">
<emphasis id="B972EA8BFFC4FFAB0D573EB8FE2BFC23" box="[339,451,907,933]" italics="true" pageId="2" pageNumber="99">R. kangae</emphasis>
</taxonomicName>
<emphasis id="B972EA8BFFC4FFAB0DCD3EB8FDCAFC23" bold="true" box="[457,546,907,933]" pageId="2" pageNumber="99">
<taxonomicNameLabel id="A24157F0FFC4FFAB0DCD3EB8FDCAFC23" box="[457,546,907,933]" pageId="2" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
bears large compound eyes in proportion to its head width when compared with those of males of 
<taxonomicName id="4C064D1AFFC4FFAB0D593E9CFE3FFC4F" authorityName="Botosaneanu" authorityYear="1970" box="[349,471,943,969]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="confissa">
<emphasis id="B972EA8BFFC4FFAB0D593E9CFE3FFC4F" box="[349,471,943,969]" italics="true" pageId="2" pageNumber="99">R. confissa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C064D1AFFC4FFAB0E093E9CFD87FC4E" authorityName="Botosaneanu" authorityYear="1970" box="[525,623,943,968]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B972EA8BFFC4FFAB0E093E9CFD87FC4E" box="[525,623,943,968]" italics="true" pageId="2" pageNumber="99">R. vicina</emphasis>
</taxonomicName>
, and also with those of females of these three species. The sexual dimorphism in the size of compound eyes observed in this new species suggests that the mating behavior of this species may differ from that of 
<taxonomicName id="4C064D1AFFC4FFAB0DA43EC4FDF3FB97" authorityName="Botosaneanu" authorityYear="1970" box="[416,539,1015,1041]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="confissa">
<emphasis id="B972EA8BFFC4FFAB0DA43EC4FDF3FB97" box="[416,539,1015,1041]" italics="true" pageId="2" pageNumber="99">R. confissa</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C064D1AFFC4FFAB0E553EC4FD5CFB96" authorityName="Botosaneanu" authorityYear="1970" box="[593,692,1015,1040]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B972EA8BFFC4FFAB0E553EC4FD5CFB96" box="[593,692,1015,1040]" italics="true" pageId="2" pageNumber="99">R. vicina</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>