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<document id="2AE56966B8278F9D86B0E2FB796F4ECB" ID-CLB-Dataset="6498" ID-DOI="10.11646/zootaxa.4966.1.1" ID-GBIF-Dataset="e3a4e21d-e4fd-4d82-a596-de175e91f976" ID-ISSN="1175-5326" ID-Zenodo-Dep="4729328" ID-ZooBank="BD4EE68E-893C-47FC-91CE-418F4A6FE46C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619760982532" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2021" docId="03AF8785A37E2765FF04256E1B27FD15" docLanguage="en" docName="zootaxa.4966.1.1.pdf" docOrigin="Zootaxa 4966 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hlavaciellus concavus Jałoszyński 2021, sp. n." docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="FF96FFFDA379276CFF9323341A13FFE4" masterDocTitle="Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae)" masterLastPageNumber="15" masterPageNumber="1" pageNumber="8" updateTime="1698938032643" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C6D0C5FC24D09196E26E48FEA459F38C">Six new species of the Oriental genus Hlavaciellus Jałoszyński (Coleoptera Staphylinidae, Scydmaeninae)</mods:title>
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<treatment id="03AF8785A37E2765FF04256E1B27FD15" ID-DOI="http://doi.org/10.5281/zenodo.4791059" ID-GBIF-Taxon="182387821" ID-Zenodo-Dep="4791059" LSID="urn:lsid:plazi:treatment:03AF8785A37E2765FF04256E1B27FD15" httpUri="http://treatment.plazi.org/id/03AF8785A37E2765FF04256E1B27FD15" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
<subSubSection id="C31C6518A37E276BFF04256E1BE3F991" box="[151,496,1626,1653]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BB93693A37E276BFF04256E1BE3F991" blockId="7.[151,496,1626,1689]" box="[151,496,1626,1653]" pageId="7" pageNumber="8">
<heading id="D0F181FFA37E276BFF04256E1BE3F991" bold="true" box="[151,496,1626,1653]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<emphasis id="B972EA81A37E276BFF04256E1BE3F991" bold="true" box="[151,496,1626,1653]" pageId="7" pageNumber="8">
<taxonomicName id="4C064D10A37E276BFF04256E1BB5F991" ID-CoL="8Y4JN" authority="Jałoszyński, 2021" authorityName="Jałoszyński" authorityYear="2021" box="[151,422,1626,1653]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="concavus" status="sp. nov.">
<emphasis id="B972EA81A37E276BFF04256E1BB5F991" bold="true" box="[151,422,1626,1653]" italics="true" pageId="7" pageNumber="8">Hlavaciellus concavus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24157FAA37E276BFE3D256F1BE3F991" box="[430,496,1627,1653]" pageId="7" pageNumber="8" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C31C6518A37E276BFF04254B1B42F97D" box="[151,337,1662,1689]" pageId="7" pageNumber="8" type="description">
<paragraph id="8BB93693A37E276BFF04254B1B42F97D" blockId="7.[151,496,1626,1689]" box="[151,337,1662,1689]" pageId="7" pageNumber="8">
(
<figureCitation id="133D2A16A37E276BFF0C254B1B33F97D" box="[159,288,1663,1689]" captionStart="FIGURES 1720" captionStartId="8.[151,264,1491,1515]" captionTargetBox="[161,1426,197,1459]" captionTargetId="figure-19@8.[151,1436,181,1467]" captionTargetPageId="8" captionText="FIGURES 1720. Hlavaciellus concavus sp. n., male. Dorsal habitus (17); head in anterior view (18); aedeagus in ventral (19) and lateral (20) views." figureDoi="http://doi.org/10.5281/zenodo.4729338" httpUri="https://zenodo.org/record/4729338/files/figure.png" pageId="7" pageNumber="8">Figs 1720</figureCitation>
,
<figureCitation id="133D2A16A37E276BFEBF254A1B5AF97D" box="[300,329,1662,1689]" captionStart="FIGURE 25" captionStartId="11.[151,250,1285,1309]" captionTargetBox="[151,1436,237,1261]" captionTargetId="figure-43@11.[151,1436,237,1261]" captionTargetPageId="11" captionText="FIGURE 25. Distribution of nominal species of Hlavaciellus." figureDoi="http://doi.org/10.5281/zenodo.4729342" httpUri="https://zenodo.org/record/4729342/files/figure.png" pageId="7" pageNumber="8">25</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31C6518A37E276BFF0425F3192BF8CD" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8BB93693A37E276BFF0425F3192BF8CD" blockId="7.[151,1437,1734,1977]" pageId="7" pageNumber="8">
<emphasis id="B972EA81A37E276BFF0425F31B5BF904" bold="true" box="[151,328,1734,1761]" pageId="7" pageNumber="8">Type material.</emphasis>
<materialsCitation id="3B6E3CCEA37E276BFEC125F3192BF8CD" ID-GBIF-Occurrence="3118551302" collectionCode="MHNG" collectorName="P. Jaloszynski" country="Indonesia" elevation="1850" location="Mt Kerinci" pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<emphasis id="B972EA81A37E276BFEC125F318EDF904" bold="true" box="[338,766,1734,1761]" pageId="7" pageNumber="8">
<typeStatus id="54BD8831A37E276BFEC125F31BD0F905" box="[338,451,1735,1761]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3117603A37E276BFE4025F21863F904" box="[467,624,1734,1760]" name="Indonesia" pageId="7" pageNumber="8">INDONESIA</collectingCountry>
(
<collectingRegion id="49C2F871A37E276BFD1125F218E2F904" box="[642,753,1734,1760]" country="Indonesia" name="Sumatera Utara" pageId="7" pageNumber="8">Sumatra</collectingRegion>
):
</emphasis>
<specimenCount id="9D00FD1AA37E276BFC9B25F3190CF905" box="[776,799,1735,1761]" pageId="7" pageNumber="8" type="male"></specimenCount>
, two labels: “
<collectingRegion id="49C2F871A37E276BFC2C25F31E5FF905" box="[959,1100,1735,1761]" country="Indonesia" name="Sumatera Utara" pageId="7" pageNumber="8">SUMATRA</collectingRegion>
:
<collectingRegion id="49C2F871A37E276BFBC925F31E8EF905" box="[1114,1181,1735,1761]" country="Indonesia" name="Jambi" pageId="7" pageNumber="8">Jambi</collectingRegion>
/
<location id="8ED96048A37E276BFB2A25F31F28F905" LSID="urn:lsid:plazi:treatment:03AF8785A37E2765FF04256E1B27FD15:8ED96048A37E276BFB2A25F31F28F905" box="[1209,1339,1735,1761]" country="Indonesia" name="Mt Kerinci" pageId="7" pageNumber="8" stateProvince="Sumatera Utara">Mt Kerinci</location>
, 1750- /
<quantity id="4CFE9B76A37E276BFF0425DE1AE3F8E1" box="[151,240,1770,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="7" pageNumber="8" unit="m" value="1850.0">
<elevation id="002BD1A0A37E276BFF0425DE1AE3F8E1" box="[151,240,1770,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="7" pageNumber="8" unit="m" value="1850.0">1850 m</elevation>
</quantity>
,
<date id="FFB81053A37E276BFF6D25DF1BACF8E1" box="[254,447,1771,1797]" pageId="7" pageNumber="8" value="1989-11-11" valueMax="1989-11-12" valueMin="1989-11-11">11-12.XI. / 1989</date>
, Agosti, Löbl / Burckhardt #11” [white, printed], “
<taxonomicName id="4C064D10A37E276BFB9625DF1ED4F8E1" authorityName="Jaloszynski" authorityYear="2006" box="[1029,1223,1771,1797]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA81A37E276BFB9625DF1ED4F8E1" box="[1029,1223,1771,1797]" italics="true" pageId="7" pageNumber="8">HLAVACIELLUS</emphasis>
</taxonomicName>
/
<taxonomicName id="4C064D10A37E276BFB7225DF1F5AF8E0" authorityName="Jałoszyński" authorityYear="2021" box="[1249,1353,1771,1796]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="concavus">
<emphasis id="B972EA81A37E276BFB7225DF1F5AF8E0" box="[1249,1353,1771,1796]" italics="true" pageId="7" pageNumber="8">concavus</emphasis>
</taxonomicName>
m. /
<collectorName id="26F35345A37E276BFA1A25DF1B08F8CD" pageId="7" pageNumber="8">P. Jałoszyński</collectorName>
, 2021 / HOLOTYPUS” [red, printed] (
<collectionCode id="ED17AE56A37E276BFD43243B193FF8CD" box="[720,812,1807,1833]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="7" pageNumber="8" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31C6518A37E276BFF5424061F89F85D" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BB93693A37E276BFF5424061F89F85D" blockId="7.[151,1437,1734,1977]" pageId="7" pageNumber="8">
<emphasis id="B972EA81A37E276BFF5424061B51F8A8" bold="true" box="[199,322,1842,1868]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
Body very large, exceeding
<quantity id="4CFE9B76A37E276BFD19240718F1F8A9" box="[650,738,1843,1869]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="8" unit="mm" value="2.2">2.2 mm</quantity>
in length; head in males with large and deep median impression between eyes filled with short and dense setae directed toward its center; each of antennomeres 310 elongate; aedeagus in ventral view moderately stout, 3 times as long as broad and distinctly narrowing both toward apex and base; parameres with thin subapical setae and conspicuously broadened and then rapidly narrowed apical portions.
</paragraph>
</subSubSection>
<caption id="DF79661BA3712764FF0426E71B68F9EB" ID-DOI="http://doi.org/10.5281/zenodo.4729338" ID-Zenodo-Dep="4729338" httpUri="https://zenodo.org/record/4729338/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,264,1491,1515]" targetBox="[161,1426,197,1459]" targetPageId="8">
<paragraph id="8BB93693A3712764FF0426E71B68F9EB" blockId="8.[151,1436,1491,1552]" pageId="8" pageNumber="9">
<emphasis id="B972EA81A3712764FF0426E71B47FA0F" bold="true" box="[151,340,1491,1515]" pageId="8" pageNumber="9">FIGURES 1720.</emphasis>
<taxonomicName id="4C064D10A3712764FECA26E7182DFA0F" authority="Jałoszyński, 2021" authorityName="Jałoszyński" authorityYear="2021" box="[345,574,1491,1515]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="concavus" status="sp. nov.">
<emphasis id="B972EA81A3712764FECA26E7182DFA0F" box="[345,574,1491,1515]" italics="true" pageId="8" pageNumber="9">Hlavaciellus concavus</emphasis>
</taxonomicName>
<emphasis id="B972EA81A3712764FDD726E0186FFA08" bold="true" box="[580,636,1492,1516]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="A24157FAA3712764FDD726E0186FFA08" box="[580,636,1492,1516]" pageId="8" pageNumber="9" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male. Dorsal habitus (17); head in anterior view (18); aedeagus in ventral (19) and lateral (20) views.
</paragraph>
</caption>
<subSubSection id="C31C6518A3712765FF54250F1B8DFDFD" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
<paragraph id="8BB93693A3712764FF54250F1B95F99C" blockId="8.[151,1437,1595,2037]" pageId="8" pageNumber="9">
<emphasis id="B972EA81A3712764FF54250F1B4AF9B1" bold="true" box="[199,345,1595,1621]" pageId="8" pageNumber="9">Description.</emphasis>
Body of male (
<figureCitation id="133D2A16A3712764FD96250F1845F9B1" box="[517,598,1595,1621]" captionStart="FIGURES 1720" captionStartId="8.[151,264,1491,1515]" captionTargetBox="[161,1426,197,1459]" captionTargetId="figure-19@8.[151,1436,181,1467]" captionTargetPageId="8" captionText="FIGURES 1720. Hlavaciellus concavus sp. n., male. Dorsal habitus (17); head in anterior view (18); aedeagus in ventral (19) and lateral (20) views." figureDoi="http://doi.org/10.5281/zenodo.4729338" httpUri="https://zenodo.org/record/4729338/files/figure.png" pageId="8" pageNumber="9">Fig. 17</figureCitation>
) stout, strongly convex, uniformly light brown; setae distinctly lighter than cuticle; BL
<quantity id="4CFE9B76A3712764FE89256A1B92F99C" box="[282,385,1630,1656]" metricMagnitude="-3" metricUnit="m" metricValue="2.23" pageId="8" pageNumber="9" unit="mm" value="2.23">2.23 mm</quantity>
.
</paragraph>
<paragraph id="8BB93693A3712764FF5425B419C9F8AC" blockId="8.[151,1437,1595,2037]" pageId="8" pageNumber="9">
Head (
<figureCitation id="133D2A16A3712764FE8125B41B70F97E" box="[274,355,1664,1690]" captionStart="FIGURES 1720" captionStartId="8.[151,264,1491,1515]" captionTargetBox="[161,1426,197,1459]" captionTargetId="figure-19@8.[151,1436,181,1467]" captionTargetPageId="8" captionText="FIGURES 1720. Hlavaciellus concavus sp. n., male. Dorsal habitus (17); head in anterior view (18); aedeagus in ventral (19) and lateral (20) views." figureDoi="http://doi.org/10.5281/zenodo.4729338" httpUri="https://zenodo.org/record/4729338/files/figure.png" pageId="8" pageNumber="9">Fig. 18</figureCitation>
) broadest at large, strongly convex and coarsely faceted eyes, HL
<quantity id="4CFE9B76A3712764FBD925B41EA3F97F" box="[1098,1200,1664,1691]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="mm" value="0.3">0.30 mm</quantity>
, HW
<quantity id="4CFE9B76A3712764FB6325B41F44F97F" box="[1264,1367,1664,1691]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="9" unit="mm" value="0.6">0.60 mm</quantity>
; frons and vertex with large and deep circular median impression. Punctures on frons in front of impression fine and inconspicuous; setae between supraantennal tubercles and on sides of impression long, moderately dense and suberect, impression densely filled with short, recumbent setae directed toward its center. Antennae slender, AnL
<quantity id="4CFE9B76A3712764FAA325DC1F8BF8E7" box="[1328,1432,1768,1795]" metricMagnitude="-3" metricUnit="m" metricValue="1.63" pageId="8" pageNumber="9" unit="mm" value="1.63">1.63 mm</quantity>
, BL/AnL 1.37; scape indistinctly elongate, pedicel as long as broad, antennomeres 310 each elongate (4 and 5 weakly so), 11 slightly shorter than 910 combined, 2.2 ×as long as broad.
</paragraph>
<paragraph id="8BB93693A3712764FF54246519F5F811" blockId="8.[151,1437,1595,2037]" pageId="8" pageNumber="9">
Pronotum subrectangular, equally broad at base and at anterior third; PL
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, PW
<quantity id="4CFE9B76A3712764FB0724641EE9F88F" box="[1172,1274,1872,1899]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="8" pageNumber="9" unit="mm" value="0.93">0.93 mm</quantity>
, PI 0.73; anterior margin weakly rounded; sides with finely crenulate lateral carinae, in posterior 2/3 distinctly concave; posterior corners sharp-angled but with blunt tips; posterior margin shallowly bisinuate; pronotal disc strongly convex, with area at each posterior corner distinctly impressed; antebasal pits slightly closer to posterior than to lateral margins. Punctures on pronotum fine and inconspicuous; setae long, dense, suberect.
</paragraph>
<paragraph id="8BB93693A3702765FF5423A31B3CFEF9" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="10">
Elytra together oval, broadest distinctly in front of middle; EL
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, EW
<quantity id="4CFE9B76A3702765FBD323A31EBBFF55" box="[1088,1192,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="9" pageNumber="10" unit="mm" value="1.08">1.08 mm</quantity>
, EI 1.16; subhumeral lines sharply marked, each developed as step-wise border between elevated humeral region and median area, as long as 0.3 EL; sides of elytra strongly rounded; elytral apices separately rounded. Punctures and setae similar to those on pronotum.
</paragraph>
<paragraph id="8BB93693A3702765FF5422131840FEA5" blockId="9.[151,1437,150,753]" box="[199,595,295,321]" pageId="9" pageNumber="10">Legs long and slender, unmodified.</paragraph>
<paragraph id="8BB93693A3702765FF54227F1955FE11" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="10">
Aedeagus (
<figureCitation id="133D2A16A3702765FED5227F1BD4FE81" box="[326,455,331,357]" captionStart="FIGURES 1720" captionStartId="8.[151,264,1491,1515]" captionTargetBox="[161,1426,197,1459]" captionTargetId="figure-19@8.[151,1436,181,1467]" captionTargetPageId="8" captionText="FIGURES 1720. Hlavaciellus concavus sp. n., male. Dorsal habitus (17); head in anterior view (18); aedeagus in ventral (19) and lateral (20) views." figureDoi="http://doi.org/10.5281/zenodo.4729338" httpUri="https://zenodo.org/record/4729338/files/figure.png" pageId="9" pageNumber="10">Figs 1920</figureCitation>
) elongate but not very slender; AeL
<quantity id="4CFE9B76A3702765FCFF227E19C7FE81" box="[876,980,330,357]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="10" unit="mm" value="0.45">0.45 mm</quantity>
; median lobe in ventral view 3 times as long as broad, broadest near sub-basal fourth and with broadly rounded sides near base, but lateral margins straight in distal 2/3; endophallic structures distinct, in ventral view appearing as two elongate elements, but in lateral view one tubular structure can be seen bent at sharp angle near base of median lobe; parameres with thin subapical setae, each with broadened and then rapidly narrowed distal region.
</paragraph>
<paragraph id="8BB93693A3702765FF5422CB1B8DFDFD" blockId="9.[151,1437,150,753]" box="[199,414,511,537]" pageId="9" pageNumber="10">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C31C6518A3702765FF5421161826FDD9" box="[199,565,546,573]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BB93693A3702765FF5421161826FDD9" blockId="9.[151,1437,150,753]" box="[199,565,546,573]" pageId="9" pageNumber="10">
<emphasis id="B972EA81A3702765FF5421161B4AFDD8" bold="true" box="[199,345,546,572]" pageId="9" pageNumber="10">Distribution</emphasis>
.
<collectingRegion id="49C2F871A3702765FEF421171BD7FDD9" box="[359,452,547,573]" country="Indonesia" name="Sumatera Utara" pageId="9" pageNumber="10">Sumatra</collectingRegion>
(
<figureCitation id="133D2A16A3702765FE472117183BFDD9" box="[468,552,546,573]" captionStart="FIGURE 25" captionStartId="11.[151,250,1285,1309]" captionTargetBox="[151,1436,237,1261]" captionTargetId="figure-43@11.[151,1436,237,1261]" captionTargetPageId="11" captionText="FIGURE 25. Distribution of nominal species of Hlavaciellus." figureDoi="http://doi.org/10.5281/zenodo.4729342" httpUri="https://zenodo.org/record/4729342/files/figure.png" pageId="9" pageNumber="10">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31C6518A3702765FF5421731B33FD61" pageId="9" pageNumber="10" type="etymology">
<paragraph id="8BB93693A3702765FF5421731B33FD61" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="10">
<emphasis id="B972EA81A3702765FF5421731B5DFD85" bold="true" box="[199,334,583,609]" pageId="9" pageNumber="10">Etymology.</emphasis>
<emphasis id="B972EA81A3702765FEC721731BD1FD85" box="[340,450,583,609]" italics="true" pageId="9" pageNumber="10">Concavus</emphasis>
is a Latin adjective meaning “hollow, concave” and refers to the large median concavity on the head.
</paragraph>
</subSubSection>
<subSubSection id="C31C6518A3702765FF5421BB1B27FD15" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BB93693A3702765FF5421BB1B27FD15" blockId="9.[151,1437,150,753]" pageId="9" pageNumber="10">
<emphasis id="B972EA81A3702765FF5421BB1B28FD4D" bold="true" box="[199,315,655,681]" pageId="9" pageNumber="10">Remarks.</emphasis>
The conspicuous round impression densely filled with short setae on the head is a unique apomorphy of this species; in addition,
<taxonomicName id="4C064D10A3702765FE422187184FFD28" authorityName="Jałoszyński" authorityYear="2021" box="[465,604,691,716]" class="Insecta" family="Staphylinidae" genus="Hlavaciellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="concavus">
<emphasis id="B972EA81A3702765FE422187184FFD28" box="[465,604,691,716]" italics="true" pageId="9" pageNumber="10">H. concavus</emphasis>
</taxonomicName>
has the largest adults of all known members of this genus, exceeding
<quantity id="4CFE9B76A3702765FAEA21871AD0FD15" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="10" unit="mm" value="2.2">2.2 mm</quantity>
in length.
</paragraph>
</subSubSection>
</treatment>
</document>