198 lines
40 KiB
XML
198 lines
40 KiB
XML
<document id="B3429F6C84D6721AA41DCC56E70D18DD" ID-DOI="10.11646/zootaxa.5415.1.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="10692812" ID-ZooBank="27930661-26F3-40F0-859B-C5287F0ABE8F" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708602940988" checkinUser="plazi" docAuthor="Bosselaers, Jan" docDate="2024" docId="03AB87A7E73BFF8DFF07F8BFC18AF9B1" docLanguage="en" docName="zootaxa.5415.1.7.pdf" docOrigin="Zootaxa 5415 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5415.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coryssiphus Simon 1903" docType="treatment" docVersion="2" lastPageNumber="155" masterDocId="FF92FFDFE73AFF8FFF90FFC5C04CFFE0" masterDocTitle="A revision of the genus Coryssiphus Simon, 1903, with its transfer to Systariinae (Araneae: Miturgidae)" masterLastPageNumber="168" masterPageNumber="153" pageNumber="154" updateTime="1708603480815" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1B154E510A2D01F7E99D51F338293704">A revision of the genus Coryssiphus Simon, 1903, with its transfer to Systariinae (Araneae: Miturgidae)</mods:title>
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<mods:namePart id="EF0BA1F1B45609A1D934D38B60827CD0">Bosselaers, Jan</mods:namePart>
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<mods:title id="9F257BEEDD3992CC4F0F2D8351EC12BC">Zootaxa</mods:title>
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<mods:date id="BD6712AFE68C9BF0935577FA6D003F62">2024</mods:date>
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<treatment id="03AB87A7E73BFF8DFF07F8BFC18AF9B1" ID-DOI="http://doi.org/10.5281/zenodo.10692797" ID-Zenodo-Dep="10692797" LSID="urn:lsid:plazi:treatment:03AB87A7E73BFF8DFF07F8BFC18AF9B1" httpUri="http://treatment.plazi.org/id/03AB87A7E73BFF8DFF07F8BFC18AF9B1" lastPageId="2" lastPageNumber="155" pageId="1" pageNumber="154">
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<subSubSection id="C318653AE73BFF8EFF07F8BFC24EF874" box="[151,514,1914,1941]" pageId="1" pageNumber="154" type="nomenclature">
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<paragraph id="8BBD36B1E73BFF8EFF07F8BFC24EF874" blockId="1.[151,514,1914,1941]" box="[151,514,1914,1941]" pageId="1" pageNumber="154">
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<heading id="D0F581DDE73BFF8EFF07F8BFC24EF874" bold="true" box="[151,514,1914,1941]" fontSize="11" level="1" pageId="1" pageNumber="154" reason="1">
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<emphasis id="B976EAA3E73BFF8EFF07F8BFC24EF874" bold="true" box="[151,514,1914,1941]" pageId="1" pageNumber="154">
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<taxonomicName id="4C024D32E73BFF8EFF07F8BFC1B0F874" ID-CoL="62NZM" authority="Simon, 1903: 31" authorityName="Simon" authorityPageNumber="31" authorityYear="1903" box="[151,508,1914,1941]" class="Arachnida" family="Liocranidae" genus="Coryssiphus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="154" phylum="Arthropoda" rank="genus">
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<emphasis id="B976EAA3E73BFF8EFF07F8BFC167F875" bold="true" box="[151,299,1914,1941]" italics="true" pageId="1" pageNumber="154">Coryssiphus</emphasis>
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Simon, 1903: 31
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</taxonomicName>
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.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C318653AE73BFF8EFF07F806C4F9F83D" box="[151,1205,1986,2013]" pageId="1" pageNumber="154" type="type_taxon">
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<paragraph id="8BBD36B1E73BFF8EFF07F806C4F9F83D" blockId="1.[151,1205,1986,2013]" box="[151,1205,1986,2013]" pageId="1" pageNumber="154">
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<emphasis id="B976EAA3E73BFF8EFF07F806C179F83C" bold="true" box="[151,309,1986,2013]" pageId="1" pageNumber="154">
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<typeStatus id="54B98813E73BFF8EFF07F806C09EF83D" box="[151,210,1987,2013]" pageId="1" pageNumber="154">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C024D32E73BFF8EFEACF806C29FF83D" ID-CoL="YSTH" authority="Simon, 1903" authorityName="Simon" authorityYear="1903" box="[316,723,1987,2013]" class="Arachnida" family="Liocranidae" genus="Coryssiphus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="154" phylum="Arthropoda" rank="species" species="praeustus">
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<emphasis id="B976EAA3E73BFF8EFEACF806C276F83C" box="[316,570,1987,2013]" italics="true" pageId="1" pageNumber="154">Coryssiphus praeustus</emphasis>
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Simon, 1903
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</taxonomicName>
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, by subsequent designation (
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<bibRefCitation id="EF934B40E73BFF8EFB84F806C4E1F83D" author="Simon, E." box="[1044,1197,1987,2013]" pageId="1" pageNumber="154" pagination="669 - 1080" refId="ref8691" refString="Simon, E. (1903 b) n. k. In: Histoire naturelle des Araignees. Tome 2. Quatrieme fascicule. Roret, Paris, pp. 669 - 1080." type="book chapter" year="1903">Simon 1903b</bibRefCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C318653AE738FF8DFF57FF53C11FFE31" pageId="2" pageNumber="155" type="diagnosis">
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<paragraph id="8BBD36B1E738FF8DFF57FF53C11FFE31" blockId="2.[151,1437,150,1617]" pageId="2" pageNumber="155">
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<emphasis id="B976EAA3E738FF8DFF57FF53C10EFF50" bold="true" box="[199,322,150,176]" pageId="2" pageNumber="155">Diagnosis.</emphasis>
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<taxonomicName id="4C024D32E738FF8DFEDAFF52C198FF51" authorityName="Simon" authorityYear="1903" box="[330,468,151,177]" class="Arachnida" family="Liocranidae" genus="Coryssiphus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
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<emphasis id="B976EAA3E738FF8DFEDAFF52C198FF51" box="[330,468,151,177]" italics="true" pageId="2" pageNumber="155">Coryssiphus</emphasis>
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</taxonomicName>
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is vaguely similar to
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<taxonomicName id="4C024D32E738FF8DFD5FFF52C3B5FF51" authority="Bosselaers, 2009" authorityName="Bosselaers" authorityYear="2009" box="[719,1017,151,177]" class="Arachnida" family="Liocranidae" genus="Arabelia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
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<emphasis id="B976EAA3E738FF8DFD5FFF52C37DFF51" box="[719,817,151,177]" italics="true" pageId="2" pageNumber="155">Arabelia</emphasis>
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<bibRefCitation id="EF934B40E738FF8DFCAAFF52C3B5FF51" author="Bosselaers, J." box="[826,1017,151,177]" pageId="2" pageNumber="155" pagination="37 - 55" refId="ref7195" refString="Bosselaers, J. (2009) Studies in Liocranidae (Araneae): redescriptions and transfers in Apostenus Westring and Brachyanillus Simon, as well as description of a new genus. Zootaxa, 2141 (1), 37 - 55. https: // doi. org / 10.11646 / zootaxa. 2141.1.3" type="journal article" year="2009">Bosselaers, 2009</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C024D32E738FF8DFBA3FF52C501FF51" authority="Banks, 1904" authorityName="Banks" authorityYear="1904" box="[1075,1357,151,177]" class="Arachnida" family="Liocranidae" genus="Drassinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
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<emphasis id="B976EAA3E738FF8DFBA3FF52C4FAFF51" box="[1075,1206,151,177]" italics="true" pageId="2" pageNumber="155">Drassinella</emphasis>
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Banks, 1904
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</taxonomicName>
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by the absence of C, MA and other tegular apophyses in the male palp (
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<figureCitation id="13392A34E738FF8DFCF5FF7EC391FF35" box="[869,989,187,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[203,1378,187,1905]" captionTargetId="figure-15@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Coryssiphus praeustus, male holotype (A, B, F), female RMCA 131898A (C), male NMSA 18885 (D, E). A. Palp, ventral; B. Palp, retrolateral; C. Epigyne; D. Spinnerets, ventral; E. Chelicerae, ventrolateral; F. Eye region, frontal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10692820" httpUri="https://zenodo.org/record/10692820/files/figure.png" pageId="2" pageNumber="155">Fig. 4A, B</figureCitation>
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). For the same reason, it differs from all other
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<taxonomicName id="4C024D32E738FF8DFF46FF1AC113FF19" authorityName="Simon" authorityYear="1897" box="[214,351,223,249]" class="Arachnida" family="Liocranidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="155" phylum="Arthropoda" rank="family">Liocranidae</taxonomicName>
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. Moreover, it differs from
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<taxonomicName id="4C024D32E738FF8DFD10FF1AC3EDFF19" authority="L. Koch, 1866" authorityName="L. Koch" authorityYear="1866" box="[640,929,223,249]" class="Arachnida" family="Liocranidae" genus="Liocranum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
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<emphasis id="B976EAA3E738FF8DFD10FF1AC2B7FF18" box="[640,763,223,248]" italics="true" pageId="2" pageNumber="155">Liocranum</emphasis>
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L. Koch, 1866
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</taxonomicName>
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,
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<taxonomicName id="4C024D32E738FF8DFC3AFF1AC4FAFF19" authority="Simon, 1897" authorityName="Simon" authorityYear="1897" box="[938,1206,223,249]" class="Arachnida" family="Liocranidae" genus="Mesiotelus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
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<emphasis id="B976EAA3E738FF8DFC3AFF1AC46FFF19" box="[938,1059,223,249]" italics="true" pageId="2" pageNumber="155">Mesiotelus</emphasis>
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Simon, 1897
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</taxonomicName>
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and
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<taxonomicName id="4C024D32E738FF8DFB79FF1AC1ECFEFD" authority="Ubick & Platnick, 1991" authorityName="Ubick & Platnick" authorityYear="1991" class="Arachnida" family="Liocranidae" genus="Hesperocranum" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
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<emphasis id="B976EAA3E738FF8DFB79FF1AC5D0FF18" box="[1257,1436,223,248]" italics="true" pageId="2" pageNumber="155">Hesperocranum</emphasis>
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<bibRefCitation id="EF934B40E738FF8DFF07FEC6C1ECFEFD" author="Ubick, D. & Platnick, N. I." box="[151,416,259,285]" pageId="2" pageNumber="155" pagination="1 - 12" refId="ref8726" refString="Ubick, D. & Platnick, N. I. (1991) On Hesperocranum, a new spider genus from Western North America (Araneae, Liocranidae). American Museum Novitates, 3019, 1 - 12." type="journal article" year="1991">Ubick & Platnick, 1991</bibRefCitation>
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</taxonomicName>
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by the parallel tapeta in the PME (
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<figureCitation id="13392A34E738FF8DFCB3FEC6C336FEFD" box="[803,890,259,285]" captionStart="FIGURE 3" captionStartId="6.[151,250,1835,1859]" captionTargetBox="[151,1435,181,1811]" captionTargetId="figure-25@6.[151,1435,181,1811]" captionTargetPageId="6" captionText="FIGURE 3. Coryssiphus praeustus, male holotype (A–C, E, I), male NMSA 18885 (D, F–H). A. Dorsal; B. Ventral; C. Left palp, left to right: prolateral, ventral, retrolateral; D. Eye region, do; E. Label; F. Endites, ventral; G. Spinnerets, caudal; H. Spinnerets, ventrocaudal; I Leg IV, tarsal claw. Scale bars: A, B, F: 1 mm; C, D, G, H: 250 µm; I: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10692818" httpUri="https://zenodo.org/record/10692818/files/figure.png" pageId="2" pageNumber="155">Fig. 3D</figureCitation>
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) and the absence of ventrolateral erectile bristles on the anterior legs (
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<bibRefCitation id="EF934B40E738FF8DFE13FEE2C2DCFEA1" author="Ubick, D. & Platnick, N. I." box="[387,656,295,321]" pageId="2" pageNumber="155" pagination="1 - 12" refId="ref8726" refString="Ubick, D. & Platnick, N. I. (1991) On Hesperocranum, a new spider genus from Western North America (Araneae, Liocranidae). American Museum Novitates, 3019, 1 - 12." type="journal article" year="1991">Ubick & Platnick 1991</bibRefCitation>
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).
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<taxonomicName id="4C024D32E738FF8DFD35FEE2C363FEA1" authorityName="Simon" authorityYear="1903" box="[677,815,295,321]" class="Arachnida" family="Liocranidae" genus="Coryssiphus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
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<emphasis id="B976EAA3E738FF8DFD35FEE2C363FEA1" box="[677,815,295,321]" italics="true" pageId="2" pageNumber="155">Coryssiphus</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4C024D32E738FF8DFC57FEE2C465FEA1" authorityName="Bosselaers" authorityYear="2009" box="[967,1065,295,321]" class="Arachnida" family="Liocranidae" genus="Arabelia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
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<emphasis id="B976EAA3E738FF8DFC57FEE2C465FEA1" box="[967,1065,295,321]" italics="true" pageId="2" pageNumber="155">Arabelia</emphasis>
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||
</taxonomicName>
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and
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<taxonomicName id="4C024D32E738FF8DFBF3FEE2C4AAFEA1" authorityName="Banks" authorityYear="1904" box="[1123,1254,295,321]" class="Arachnida" family="Liocranidae" genus="Drassinella" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B976EAA3E738FF8DFBF3FEE2C4AAFEA1" box="[1123,1254,295,321]" italics="true" pageId="2" pageNumber="155">Drassinella</emphasis>
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</taxonomicName>
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by the shape of the male palpal bulbus, which is circular in ventral view instead of oval, by the long and basally inserted embolus, and by the short and simple ID. It differs from
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<taxonomicName id="4C024D32E738FF8DFD27FEAAC3E1FE69" authority="Simon, 1897" authorityName="Simon" authorityYear="1897" box="[695,941,367,393]" class="Arachnida" family="Miturgidae" genus="Systaria" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="155" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B976EAA3E738FF8DFD27FEAAC35FFE69" box="[695,787,367,393]" italics="true" pageId="2" pageNumber="155">Systaria</emphasis>
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Simon, 1897
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</taxonomicName>
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(
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<taxonomicName id="4C024D32E738FF8DFC2EFEAAC40CFE69" authorityName="Simon" authorityYear="1886" box="[958,1088,367,393]" class="Arachnida" family="Miturgidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="155" phylum="Arthropoda" rank="family">Miturgidae</taxonomicName>
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: Systariinae) by the somewhat smaller AME, stronger vsp on ti and mt I and II, trochanters that are only weakly notched, and a vulva with anterior CO (
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<figureCitation id="13392A34E738FF8DFF5DFE72C10BFE31" box="[205,327,439,465]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1892]" captionTargetBox="[159,1429,619,1843]" captionTargetId="figure-136@9.[151,1435,614,1844]" captionTargetPageId="9" captionText="FIGURE 6. A, B. Coryssiphus praeustus, female RMCA 131898 (A), female RMCA 247653 (B, left), female RMCA 131872 (B, right). C–E. C. cinerascens, female RMCA 177006 (C, left), C. unicolor, holotype (C, right), male holotype (D), female RMCA 131824 (e). A. Vulva, dorsal; B, C. Epigynes. D. palp, ventral (left) and retrolateral (right). E. Vulva, dorsal. Scale bars: A, E: 100 µm; B, C: 250 µm; D: 0.5 mm. Abbreviations: ID, Insemination duct; Sss, Sickle-shaped sclerite; ST1, Spermatheca 1; ST2, Spermatheca 2 (“bursa”)." figureDoi="http://doi.org/10.5281/zenodo.10692826" httpUri="https://zenodo.org/record/10692826/files/figure.png" pageId="2" pageNumber="155">Fig. 6A, E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C318653AE738FF8DFF57FE1FC18AF9B1" pageId="2" pageNumber="155" type="description">
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||
<paragraph id="8BBD36B1E738FF8DFF57FE1FC18AF9B1" blockId="2.[151,1437,150,1617]" pageId="2" pageNumber="155">
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||
<emphasis id="B976EAA3E738FF8DFF57FE1FC115FE14" bold="true" box="[199,345,474,500]" pageId="2" pageNumber="155">Description.</emphasis>
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Small to medium sized spiders (
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||
<quantity id="4CFA9B54E738FF8DFD5CFE1EC362FE15" box="[716,814,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="2" pageNumber="155" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3–7 mm</quantity>
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), females 20–50% larger than males. Carapace orange-brown, smooth and shiny, with thin brown lining, abruptly narrowed towards the cephalic region, somewhat darker around eye area (
|
||
<figureCitation id="13392A34E738FF8DFEC5FDE6C1E1FDDD" box="[341,429,547,573]" captionStart="FIGURE 3" captionStartId="6.[151,250,1835,1859]" captionTargetBox="[151,1435,181,1811]" captionTargetId="figure-25@6.[151,1435,181,1811]" captionTargetPageId="6" captionText="FIGURE 3. Coryssiphus praeustus, male holotype (A–C, E, I), male NMSA 18885 (D, F–H). A. Dorsal; B. Ventral; C. Left palp, left to right: prolateral, ventral, retrolateral; D. Eye region, do; E. Label; F. Endites, ventral; G. Spinnerets, caudal; H. Spinnerets, ventrocaudal; I Leg IV, tarsal claw. Scale bars: A, B, F: 1 mm; C, D, G, H: 250 µm; I: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10692818" httpUri="https://zenodo.org/record/10692818/files/figure.png" pageId="2" pageNumber="155">Fig. 3A</figureCitation>
|
||
). Width of cephalic region about half the width of carapace (
|
||
<figureCitation id="13392A34E738FF8DFBD9FDE6C4E9FDDD" box="[1097,1189,547,573]" captionStart="FIGURE 3" captionStartId="6.[151,250,1835,1859]" captionTargetBox="[151,1435,181,1811]" captionTargetId="figure-25@6.[151,1435,181,1811]" captionTargetPageId="6" captionText="FIGURE 3. Coryssiphus praeustus, male holotype (A–C, E, I), male NMSA 18885 (D, F–H). A. Dorsal; B. Ventral; C. Left palp, left to right: prolateral, ventral, retrolateral; D. Eye region, do; E. Label; F. Endites, ventral; G. Spinnerets, caudal; H. Spinnerets, ventrocaudal; I Leg IV, tarsal claw. Scale bars: A, B, F: 1 mm; C, D, G, H: 250 µm; I: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10692818" httpUri="https://zenodo.org/record/10692818/files/figure.png" pageId="2" pageNumber="155">Figs 3A</figureCitation>
|
||
,
|
||
<figureCitation id="13392A34E738FF8DFB20FDE6C499FDDD" box="[1200,1237,547,573]" captionStart="FIGURE 8" captionStartId="11.[151,250,1250,1274]" captionTargetBox="[151,1435,181,1226]" captionTargetId="figure-15@11.[151,1435,181,1226]" captionTargetPageId="11" captionText="FIGURE 8. Coryssiphus cinerascens, female, C. unicolor holotype (A, B, D, E top, F), RMCA 177006 (C, E bottom right), RMCA 131824 (E bottom left, G). A. Habitus, dorsal; B, Habitus, ventral; C. Spinnerets, ventral; D. Same, caudal; E. Epigynes. F. Label; G. Vulva, dorsal. Scale bars: A, B: 2 mm; C–E: 250 µm; G: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.10692830" httpUri="https://zenodo.org/record/10692830/files/figure.png" pageId="2" pageNumber="155">8A</figureCitation>
|
||
). Carapace gently sloping upwards from front towards fovea, then steeply sloping down towards petiolus. Fovea pronounced and narrow, dark brown. Faint, darker streaks radiating from fovea towards intercoxal spaces. Clypeus vertical, similar in size to diameter of AME. All eyes ringed with black, subequal in size (
|
||
<figureCitation id="13392A34E738FF8DFC4FFD4AC477FD49" box="[991,1083,655,681]" captionStart="FIGURE 3" captionStartId="6.[151,250,1835,1859]" captionTargetBox="[151,1435,181,1811]" captionTargetId="figure-25@6.[151,1435,181,1811]" captionTargetPageId="6" captionText="FIGURE 3. Coryssiphus praeustus, male holotype (A–C, E, I), male NMSA 18885 (D, F–H). A. Dorsal; B. Ventral; C. Left palp, left to right: prolateral, ventral, retrolateral; D. Eye region, do; E. Label; F. Endites, ventral; G. Spinnerets, caudal; H. Spinnerets, ventrocaudal; I Leg IV, tarsal claw. Scale bars: A, B, F: 1 mm; C, D, G, H: 250 µm; I: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10692818" httpUri="https://zenodo.org/record/10692818/files/figure.png" pageId="2" pageNumber="155">Fig. 3D</figureCitation>
|
||
). AME black, other eyes pearl. PME with parallel tapetum. ALE close to PLE, separated by less than one eye diameter. AER slightly recurved from above, slightly procurved from front. PER procurved dorsally and frontally (
|
||
<figureCitation id="13392A34E738FF8DFBA1FD12C4C3FD11" box="[1073,1167,727,753]" captionStart="FIGURE 3" captionStartId="6.[151,250,1835,1859]" captionTargetBox="[151,1435,181,1811]" captionTargetId="figure-25@6.[151,1435,181,1811]" captionTargetPageId="6" captionText="FIGURE 3. Coryssiphus praeustus, male holotype (A–C, E, I), male NMSA 18885 (D, F–H). A. Dorsal; B. Ventral; C. Left palp, left to right: prolateral, ventral, retrolateral; D. Eye region, do; E. Label; F. Endites, ventral; G. Spinnerets, caudal; H. Spinnerets, ventrocaudal; I Leg IV, tarsal claw. Scale bars: A, B, F: 1 mm; C, D, G, H: 250 µm; I: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10692818" httpUri="https://zenodo.org/record/10692818/files/figure.png" pageId="2" pageNumber="155">Figs 3D</figureCitation>
|
||
,
|
||
<figureCitation id="13392A34E738FF8DFB0BFD12C4F0FD11" box="[1179,1212,727,753]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[203,1378,187,1905]" captionTargetId="figure-15@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Coryssiphus praeustus, male holotype (A, B, F), female RMCA 131898A (C), male NMSA 18885 (D, E). A. Palp, ventral; B. Palp, retrolateral; C. Epigyne; D. Spinnerets, ventral; E. Chelicerae, ventrolateral; F. Eye region, frontal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10692820" httpUri="https://zenodo.org/record/10692820/files/figure.png" pageId="2" pageNumber="155">4F</figureCitation>
|
||
). MOQ trapezoidal, narrower anteriorly, distance between anterior and posterior end larger than distance between PME. Chelicerae orange-brown, smooth, with darker fangs. Paturon with two rl and three pl teeth, the smallest pl tooth positioned basally (
|
||
<figureCitation id="13392A34E738FF8DFF66FC86C11EFCBD" box="[246,338,835,861]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[203,1378,187,1905]" captionTargetId="figure-15@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Coryssiphus praeustus, male holotype (A, B, F), female RMCA 131898A (C), male NMSA 18885 (D, E). A. Palp, ventral; B. Palp, retrolateral; C. Epigyne; D. Spinnerets, ventral; E. Chelicerae, ventrolateral; F. Eye region, frontal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10692820" httpUri="https://zenodo.org/record/10692820/files/figure.png" pageId="2" pageNumber="155">Figs 4E</figureCitation>
|
||
,
|
||
<figureCitation id="13392A34E738FF8DFECEFC86C1C8FCBD" box="[350,388,835,861]" captionStart="FIGURE 7" captionStartId="10.[151,250,1895,1919]" captionTargetBox="[151,1435,417,1871]" captionTargetId="figure-136@10.[151,1435,417,1871]" captionTargetPageId="10" captionText="FIGURE 7. Coryssiphus praeustus, female RMCA 131898 (A, B, D, E), RMCA 131898a, b, c (C, top left, middle, right), RMCA 247653 (C, bottom left), RMCA 131872 (C, bottom middle), NCA 2011/886 (C, bottom right). A. Dorsal; B. ventral; C. Epigynes; D. Right chelicera, ventral; E. Vulva, dorsal. Scale bars: A, B: 1 mm; C–E: 250 µm." figureDoi="http://doi.org/10.5281/zenodo.10692828" httpUri="https://zenodo.org/record/10692828/files/figure.png" pageId="2" pageNumber="155">7D</figureCitation>
|
||
). Labium subrectangular with thickened frontal rim, as long as wide. Endites orange-yellow, without oblique depression or lateral notch, but with marked thin, diagonal groove (
|
||
<figureCitation id="13392A34E738FF8DFBF1FCA2C4F5FC61" box="[1121,1209,871,897]" captionStart="FIGURE 3" captionStartId="6.[151,250,1835,1859]" captionTargetBox="[151,1435,181,1811]" captionTargetId="figure-25@6.[151,1435,181,1811]" captionTargetPageId="6" captionText="FIGURE 3. Coryssiphus praeustus, male holotype (A–C, E, I), male NMSA 18885 (D, F–H). A. Dorsal; B. Ventral; C. Left palp, left to right: prolateral, ventral, retrolateral; D. Eye region, do; E. Label; F. Endites, ventral; G. Spinnerets, caudal; H. Spinnerets, ventrocaudal; I Leg IV, tarsal claw. Scale bars: A, B, F: 1 mm; C, D, G, H: 250 µm; I: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10692818" httpUri="https://zenodo.org/record/10692818/files/figure.png" pageId="2" pageNumber="155">Fig. 3F</figureCitation>
|
||
). Sparse apical hair tuft and very thin serrula present. Sternum shield-shaped, longer than wide or as long as wide, yellow, smooth, not rebordered, with somewhat thicker brown border. No PLB, PCT or ICS. LOP subrectangular, consisting of two consecutive plates connected by membrane. PSP arrow-shaped, reddish-brown. Abdomen without fringe of frontal setae, do grey with median white stripe in anterior half (
|
||
<figureCitation id="13392A34E738FF8DFCCEFC32C3F7FBF1" box="[862,955,1015,1041]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[161,1426,179,1884]" captionTargetId="figure-25@4.[161,1426,179,1917]" captionTargetPageId="4" captionText="FIGURE 1. Coryssiphus spp., male habitus. A. C. praeustus, NMSA 18885. B. C. cinerascens, RMCA 174995. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10692814" httpUri="https://zenodo.org/record/10692814/files/figure.png" pageId="2" pageNumber="155">Figs 1A</figureCitation>
|
||
,
|
||
<figureCitation id="13392A34E738FF8DFC57FC32C3A1FBF1" box="[967,1005,1015,1041]" captionStart="FIGURE 3" captionStartId="6.[151,250,1835,1859]" captionTargetBox="[151,1435,181,1811]" captionTargetId="figure-25@6.[151,1435,181,1811]" captionTargetPageId="6" captionText="FIGURE 3. Coryssiphus praeustus, male holotype (A–C, E, I), male NMSA 18885 (D, F–H). A. Dorsal; B. Ventral; C. Left palp, left to right: prolateral, ventral, retrolateral; D. Eye region, do; E. Label; F. Endites, ventral; G. Spinnerets, caudal; H. Spinnerets, ventrocaudal; I Leg IV, tarsal claw. Scale bars: A, B, F: 1 mm; C, D, G, H: 250 µm; I: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10692818" httpUri="https://zenodo.org/record/10692818/files/figure.png" pageId="2" pageNumber="155">3A</figureCitation>
|
||
). Two light brown oval anterior sigilla and two dark elongate posterior sigilla. Venter creamy-white (
|
||
<figureCitation id="13392A34E738FF8DFCCAFBDEC3FBFBD5" box="[858,951,1051,1077]" captionStart="FIGURE 3" captionStartId="6.[151,250,1835,1859]" captionTargetBox="[151,1435,181,1811]" captionTargetId="figure-25@6.[151,1435,181,1811]" captionTargetPageId="6" captionText="FIGURE 3. Coryssiphus praeustus, male holotype (A–C, E, I), male NMSA 18885 (D, F–H). A. Dorsal; B. Ventral; C. Left palp, left to right: prolateral, ventral, retrolateral; D. Eye region, do; E. Label; F. Endites, ventral; G. Spinnerets, caudal; H. Spinnerets, ventrocaudal; I Leg IV, tarsal claw. Scale bars: A, B, F: 1 mm; C, D, G, H: 250 µm; I: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10692818" httpUri="https://zenodo.org/record/10692818/files/figure.png" pageId="2" pageNumber="155">Figs 3B</figureCitation>
|
||
,
|
||
<figureCitation id="13392A34E738FF8DFC55FBDEC3A5FBD5" box="[965,1001,1051,1077]" captionStart="FIGURE 7" captionStartId="10.[151,250,1895,1919]" captionTargetBox="[151,1435,417,1871]" captionTargetId="figure-136@10.[151,1435,417,1871]" captionTargetPageId="10" captionText="FIGURE 7. Coryssiphus praeustus, female RMCA 131898 (A, B, D, E), RMCA 131898a, b, c (C, top left, middle, right), RMCA 247653 (C, bottom left), RMCA 131872 (C, bottom middle), NCA 2011/886 (C, bottom right). A. Dorsal; B. ventral; C. Epigynes; D. Right chelicera, ventral; E. Vulva, dorsal. Scale bars: A, B: 1 mm; C–E: 250 µm." figureDoi="http://doi.org/10.5281/zenodo.10692828" httpUri="https://zenodo.org/record/10692828/files/figure.png" pageId="2" pageNumber="155">7B</figureCitation>
|
||
). ALS rather short and subcylindrical, with rounded tip, separated by half their diameter. PMS small, conical in males and subcylindrical in females. PLS relatively large, with conical apical segment (
|
||
<figureCitation id="13392A34E738FF8DFD02FBA6C35DFB9D" box="[658,785,1123,1149]" captionStart="FIGURE 3" captionStartId="6.[151,250,1835,1859]" captionTargetBox="[151,1435,181,1811]" captionTargetId="figure-25@6.[151,1435,181,1811]" captionTargetPageId="6" captionText="FIGURE 3. Coryssiphus praeustus, male holotype (A–C, E, I), male NMSA 18885 (D, F–H). A. Dorsal; B. Ventral; C. Left palp, left to right: prolateral, ventral, retrolateral; D. Eye region, do; E. Label; F. Endites, ventral; G. Spinnerets, caudal; H. Spinnerets, ventrocaudal; I Leg IV, tarsal claw. Scale bars: A, B, F: 1 mm; C, D, G, H: 250 µm; I: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10692818" httpUri="https://zenodo.org/record/10692818/files/figure.png" pageId="2" pageNumber="155">Figs 3G, H</figureCitation>
|
||
,
|
||
<figureCitation id="13392A34E738FF8DFC8CFBA6C30DFB9D" box="[796,833,1123,1149]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[203,1378,187,1905]" captionTargetId="figure-15@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Coryssiphus praeustus, male holotype (A, B, F), female RMCA 131898A (C), male NMSA 18885 (D, E). A. Palp, ventral; B. Palp, retrolateral; C. Epigyne; D. Spinnerets, ventral; E. Chelicerae, ventrolateral; F. Eye region, frontal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10692820" httpUri="https://zenodo.org/record/10692820/files/figure.png" pageId="2" pageNumber="155">4D</figureCitation>
|
||
,
|
||
<figureCitation id="13392A34E738FF8DFCDCFBA6C3DDFB9D" box="[844,913,1123,1149]" captionStart="FIGURE 8" captionStartId="11.[151,250,1250,1274]" captionTargetBox="[151,1435,181,1226]" captionTargetId="figure-15@11.[151,1435,181,1226]" captionTargetPageId="11" captionText="FIGURE 8. Coryssiphus cinerascens, female, C. unicolor holotype (A, B, D, E top, F), RMCA 177006 (C, E bottom right), RMCA 131824 (E bottom left, G). A. Habitus, dorsal; B, Habitus, ventral; C. Spinnerets, ventral; D. Same, caudal; E. Epigynes. F. Label; G. Vulva, dorsal. Scale bars: A, B: 2 mm; C–E: 250 µm; G: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.10692830" httpUri="https://zenodo.org/record/10692830/files/figure.png" pageId="2" pageNumber="155">8C, D</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF934B40E738FF8DFC0DFBA6C48DFB9D" author="Deeleman-Reinhold, C. L." box="[925,1217,1123,1149]" pageId="2" pageNumber="155" refId="ref7508" refString="Deeleman-Reinhold, C. L. (2001) Forest spiders of South East Asia: with a revision of the sac and ground spiders (Araneae: Clubionidae, Corinnidae, Liocranidae, Gnaphosidae, Prodidomidae and Trochanterriidae [sic]). Brill Leiden, 591 pp. https: // doi. org / 10.1163 / 9789004475588" type="book" year="2001">Deeleman-Reinhold 2001</bibRefCitation>
|
||
: figs 230, 231, 234, 235). PMS and PLS with large cylindrical gland spigots in females.Anal tubercle a blunt triangle with fringe of long hairs. RH large, oval, hyaline (
|
||
<figureCitation id="13392A34E738FF8DFE64FB6EC203FB25" box="[500,591,1194,1221]" captionStart="FIGURE 5" captionStartId="8.[151,250,1639,1663]" captionTargetBox="[151,1435,181,1615]" captionTargetId="figure-25@8.[151,1435,181,1615]" captionTargetPageId="8" captionText="FIGURE 5. Coryssiphus cinerascens, male holotype (A–C, E), male RMCA 174990 (D). A. Dorsal view; B. Ventral view; C. Left palp, left to right: prolateral, ventral, retrolateral; D. Palp and retrocoxal hymen (arrow); E. Label. Scale bars: A, B: 1 mm; C, D: 250 µm." figureDoi="http://doi.org/10.5281/zenodo.10692822" httpUri="https://zenodo.org/record/10692822/files/figure.png" pageId="2" pageNumber="155">Fig. 5D</figureCitation>
|
||
). Legs yellow, without feathery hairs. Leg formula I IV II III in males and IV I II III in females. Trochanter notch shallow. Patellar indentation (
|
||
<bibRefCitation id="EF934B40E738FF8DFC32FB0AC4FCFB09" author="Ledoux, J. - C. & Canard, A." box="[930,1200,1231,1257]" pageId="2" pageNumber="155" refId="ref8257" refString="Ledoux, J. - C. & Canard, A. (1991) Initiation a l'etude systematique des araignees. Aramon, J. - C. Ledoux, imprimeur & editeur, Domazan, 66 pp." type="book" year="1991">Ledoux & Canard 1991</bibRefCitation>
|
||
) not very wide, ¼ to ½ l of pa. Tibia I with 6–7 vsp, ti II with 5–6 vsp, in both cases sometimes slightly irregular, e.g. 6 on pl side and 7 on rl side. Metatarsi I and II with two vsp, seldom three (
|
||
<figureCitation id="13392A34E738FF8DFC81FAD2C31AFAD1" box="[785,854,1303,1329]" captionStart="FIGURE 2" captionStartId="5.[151,250,1449,1473]" captionTargetBox="[160,1420,194,1416]" captionTargetId="figure-15@5.[151,1435,181,1425]" captionTargetPageId="5" captionText="FIGURE 2. Leg spination schematics. A. Legend; B. Coryssiphus praeustus, male holotype; C. C. praeustus, female, RMCA 131898; D. Same, RMCA 247653; E. C. cinerascens, male holotype; F. C. unicolor, female holotype = C. cinerascens; G. C. cinerascens, RMCA 131824." figureDoi="http://doi.org/10.5281/zenodo.10692816" httpUri="https://zenodo.org/record/10692816/files/figure.png" pageId="2" pageNumber="155">Fig. 2</figureCitation>
|
||
). Femora III and IV with one do terminal spine pair. No preening brushes on mt III and IV, ta IV straight, ti III and IV with three vsp, mt III and IV with one vts. All ta with dense claw tuft, tarsal claws with three (leg I) or four (leg IV) teeth (
|
||
<figureCitation id="13392A34E738FF8DFC51FA9AC442FA99" box="[961,1038,1375,1401]" captionStart="FIGURE 3" captionStartId="6.[151,250,1835,1859]" captionTargetBox="[151,1435,181,1811]" captionTargetId="figure-25@6.[151,1435,181,1811]" captionTargetPageId="6" captionText="FIGURE 3. Coryssiphus praeustus, male holotype (A–C, E, I), male NMSA 18885 (D, F–H). A. Dorsal; B. Ventral; C. Left palp, left to right: prolateral, ventral, retrolateral; D. Eye region, do; E. Label; F. Endites, ventral; G. Spinnerets, caudal; H. Spinnerets, ventrocaudal; I Leg IV, tarsal claw. Scale bars: A, B, F: 1 mm; C, D, G, H: 250 µm; I: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.10692818" httpUri="https://zenodo.org/record/10692818/files/figure.png" pageId="2" pageNumber="155">Fig. 3I</figureCitation>
|
||
). Male palp with cylindrical ti, RTA an elongated triangle with sharp apical hook directed inwards (
|
||
<figureCitation id="13392A34E738FF8DFCC8FA46C39AFA7D" box="[856,982,1411,1437]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[203,1378,187,1905]" captionTargetId="figure-15@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Coryssiphus praeustus, male holotype (A, B, F), female RMCA 131898A (C), male NMSA 18885 (D, E). A. Palp, ventral; B. Palp, retrolateral; C. Epigyne; D. Spinnerets, ventral; E. Chelicerae, ventrolateral; F. Eye region, frontal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10692820" httpUri="https://zenodo.org/record/10692820/files/figure.png" pageId="2" pageNumber="155">Figs 4A, B</figureCitation>
|
||
,
|
||
<figureCitation id="13392A34E738FF8DFC71FA46C44BFA7D" box="[993,1031,1411,1437]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1892]" captionTargetBox="[159,1429,619,1843]" captionTargetId="figure-136@9.[151,1435,614,1844]" captionTargetPageId="9" captionText="FIGURE 6. A, B. Coryssiphus praeustus, female RMCA 131898 (A), female RMCA 247653 (B, left), female RMCA 131872 (B, right). C–E. C. cinerascens, female RMCA 177006 (C, left), C. unicolor, holotype (C, right), male holotype (D), female RMCA 131824 (e). A. Vulva, dorsal; B, C. Epigynes. D. palp, ventral (left) and retrolateral (right). E. Vulva, dorsal. Scale bars: A, E: 100 µm; B, C: 250 µm; D: 0.5 mm. Abbreviations: ID, Insemination duct; Sss, Sickle-shaped sclerite; ST1, Spermatheca 1; ST2, Spermatheca 2 (“bursa”)." figureDoi="http://doi.org/10.5281/zenodo.10692826" httpUri="https://zenodo.org/record/10692826/files/figure.png" pageId="2" pageNumber="155">6D</figureCitation>
|
||
). Bulbus simple, a flattened circular disc without C or MA. Embolus sickle-shaped, implanted basally, running along pl edge of bulbus, ending in sharp thin point anteriorly (
|
||
<figureCitation id="13392A34E738FF8DFE01FA0EC1BEFA05" box="[401,498,1483,1509]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[203,1378,187,1905]" captionTargetId="figure-15@7.[203,1385,181,1919]" captionTargetPageId="7" captionText="FIGURE 4. Coryssiphus praeustus, male holotype (A, B, F), female RMCA 131898A (C), male NMSA 18885 (D, E). A. Palp, ventral; B. Palp, retrolateral; C. Epigyne; D. Spinnerets, ventral; E. Chelicerae, ventrolateral; F. Eye region, frontal. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10692820" httpUri="https://zenodo.org/record/10692820/files/figure.png" pageId="2" pageNumber="155">Figs 4A</figureCitation>
|
||
,
|
||
<figureCitation id="13392A34E738FF8DFD91FA0EC26BFA05" box="[513,551,1483,1509]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1892]" captionTargetBox="[159,1429,619,1843]" captionTargetId="figure-136@9.[151,1435,614,1844]" captionTargetPageId="9" captionText="FIGURE 6. A, B. Coryssiphus praeustus, female RMCA 131898 (A), female RMCA 247653 (B, left), female RMCA 131872 (B, right). C–E. C. cinerascens, female RMCA 177006 (C, left), C. unicolor, holotype (C, right), male holotype (D), female RMCA 131824 (e). A. Vulva, dorsal; B, C. Epigynes. D. palp, ventral (left) and retrolateral (right). E. Vulva, dorsal. Scale bars: A, E: 100 µm; B, C: 250 µm; D: 0.5 mm. Abbreviations: ID, Insemination duct; Sss, Sickle-shaped sclerite; ST1, Spermatheca 1; ST2, Spermatheca 2 (“bursa”)." figureDoi="http://doi.org/10.5281/zenodo.10692826" httpUri="https://zenodo.org/record/10692826/files/figure.png" pageId="2" pageNumber="155">6D</figureCitation>
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). Cymbium elongate-oval, with blunt triangular tip. Epigyne with elongated depression flanked by two longitudinal sclerotised rims (
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<figureCitation id="13392A34E738FF8DFCB2FA2AC3D3F9E9" box="[802,927,1519,1545]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1892]" captionTargetBox="[159,1429,619,1843]" captionTargetId="figure-136@9.[151,1435,614,1844]" captionTargetPageId="9" captionText="FIGURE 6. A, B. Coryssiphus praeustus, female RMCA 131898 (A), female RMCA 247653 (B, left), female RMCA 131872 (B, right). C–E. C. cinerascens, female RMCA 177006 (C, left), C. unicolor, holotype (C, right), male holotype (D), female RMCA 131824 (e). A. Vulva, dorsal; B, C. Epigynes. D. palp, ventral (left) and retrolateral (right). E. Vulva, dorsal. Scale bars: A, E: 100 µm; B, C: 250 µm; D: 0.5 mm. Abbreviations: ID, Insemination duct; Sss, Sickle-shaped sclerite; ST1, Spermatheca 1; ST2, Spermatheca 2 (“bursa”)." figureDoi="http://doi.org/10.5281/zenodo.10692826" httpUri="https://zenodo.org/record/10692826/files/figure.png" pageId="2" pageNumber="155">Fig. 6B, C</figureCitation>
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). Copulatory openings in two inconspicuous anterior grooves. Vulva with elongate-oval anterior ST2, connected by short wide canal to smaller, subglobular posterior ST1 (
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<figureCitation id="13392A34E738FF8DFED0F9F2C1F5F9B1" box="[320,441,1591,1617]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1892]" captionTargetBox="[159,1429,619,1843]" captionTargetId="figure-136@9.[151,1435,614,1844]" captionTargetPageId="9" captionText="FIGURE 6. A, B. Coryssiphus praeustus, female RMCA 131898 (A), female RMCA 247653 (B, left), female RMCA 131872 (B, right). C–E. C. cinerascens, female RMCA 177006 (C, left), C. unicolor, holotype (C, right), male holotype (D), female RMCA 131824 (e). A. Vulva, dorsal; B, C. Epigynes. D. palp, ventral (left) and retrolateral (right). E. Vulva, dorsal. Scale bars: A, E: 100 µm; B, C: 250 µm; D: 0.5 mm. Abbreviations: ID, Insemination duct; Sss, Sickle-shaped sclerite; ST1, Spermatheca 1; ST2, Spermatheca 2 (“bursa”)." figureDoi="http://doi.org/10.5281/zenodo.10692826" httpUri="https://zenodo.org/record/10692826/files/figure.png" pageId="2" pageNumber="155">Fig. 6A, E</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |