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<document id="ACC3A46C02F4A11ADEC370C3735217A7" ID-CLB-Dataset="171013" ID-DOI="10.5852/ejt.2022.843.1947" ID-GBIF-Dataset="044b60a6-0e4e-44be-b75a-614b58434dcc" ID-ISSN="2118-9773" ID-Zenodo-Dep="7222308" ID-ZooBank="C03BE897-EFE2-4CCD-913A-723792CDF050" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666126358981" checkinUser="felipe" docAuthor="Wood, Thomas J. &amp; Monfared, Alireza" docDate="2022" docId="03AB87A2FF86513DDA42D81CFCB5FE03" docLanguage="en" docName="ejt-843_1-136.pdf" docOrigin="European Journal of Taxonomy 843" docSource="http://dx.doi.org/10.5852/ejt.2022.843.1947" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Andrena (Micrandrena) elam Wood &amp; Monfared 2022, sp. nov." docType="treatment" docUuid="339546A8-6F7C-4250-8E6B-49017F4EB6A8" docUuidSource="ZooBank" docVersion="10" lastPageNumber="21" masterDocId="FF92FFDAFF945129D84DD914FFB0FFB4" masterDocTitle="A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species" masterLastPageNumber="136" masterPageNumber="1" pageNumber="19" updateTime="1699408809500" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2E59A08A99DAC0E971E2D6AFDB8BF4DE">A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="AABB293C9C1443C79362A913E313336D">Wood, Thomas J.</mods:namePart>
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<mods:affiliation id="23E0FBEF0A4CE38F3281E55E12587378">Laboratory of Zoology, University of Mons, 7000 Mons, Belgium.</mods:affiliation>
<mods:nameIdentifier id="727503BE71EE5E40A6FAE5126174D1AD" type="email">thomasjames.wood@umons.ac.be</mods:nameIdentifier>
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<mods:namePart id="F78777A62027EBB65425520927928308">Monfared, Alireza</mods:namePart>
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<mods:affiliation id="3B9DB695E3925248C4DE33B70B72E07A">Department of Plant Protection, Faculty of Agriculture, Yasouj University, Yasouj, Iran.</mods:affiliation>
<mods:nameIdentifier id="9A3320E9DD7D4569B14B8576F5141FFD" type="email">amonfared@yu.ac.ir</mods:nameIdentifier>
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<mods:title id="0C98D9A20F4B708CA956CE8AA53AC791">European Journal of Taxonomy</mods:title>
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<mods:date id="B521F97DB1A67F178E6880E3FB790828">2022</mods:date>
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<mods:number id="304016D73FAEECC300888046464B3B3D">2022-10-17</mods:number>
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<mods:number id="CB667647C59CC61EB263722E1A065209">843</mods:number>
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<mods:classification id="62337C2082489D2BFEFF0C2C4785F060">journal article</mods:classification>
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<treatment id="03AB87A2FF86513DDA42D81CFCB5FE03" ID-DOI="http://doi.org/10.5281/zenodo.7237387" ID-GBIF-Taxon="203125001" ID-Zenodo-Dep="7237387" LSID="urn:lsid:zoobank.org:act:339546A8-6F7C-4250-8E6B-49017F4EB6A8" httpUri="http://treatment.plazi.org/id/03AB87A2FF86513DDA42D81CFCB5FE03" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19">
<subSubSection id="C318653FFF86513BDA42D81CFB95FE97" box="[527,1061,264,291]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BBD36B4FF86513BDA42D81CFB95FE97" blockId="18.[374,1213,264,363]" box="[527,1061,264,291]" pageId="18" pageNumber="19">
<heading id="D0F581D8FF86513BDA42D81CFB95FE97" box="[527,1061,264,291]" centered="true" fontSize="11" level="2" pageId="18" pageNumber="19" reason="2">
<taxonomicName id="4C024D37FF86513BDA42D81CFC72FE97" authority="Wood" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[527,962,264,291]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="elam" status="sp. nov." subGenus="Micrandrena">
<emphasis id="B976EAA6FF86513BDA42D81CFCC6FE97" bold="true" box="[527,886,264,291]" pageId="18" pageNumber="19">
<emphasis id="B976EAA6FF86513BDA42D81CFDC9FE97" bold="true" box="[527,633,264,291]" italics="true" pageId="18" pageNumber="19">Andrena</emphasis>
(
<emphasis id="B976EAA6FF86513BDAC7D81CFC9AFE97" bold="true" box="[650,810,264,291]" italics="true" pageId="18" pageNumber="19">Micrandrena</emphasis>
)
<emphasis id="B976EAA6FF86513BDB76D81CFCC6FE97" bold="true" box="[827,886,264,291]" italics="true" pageId="18" pageNumber="19">elam</emphasis>
</emphasis>
Wood
</taxonomicName>
<taxonomicNameLabel id="A24557DDFF86513BDB87D81DFB95FE97" box="[970,1061,265,291]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C318653FFF86513BD93BD839FCEBFEDF" pageId="18" pageNumber="19" type="description">
<paragraph id="8BBD36B4FF86513BD93BD839FB0DFEFC" blockId="18.[374,1213,264,363]" box="[374,1213,301,328]" pageId="18" pageNumber="19">
<uri id="FF933AB6FF86513BD93BD839FB0DFEFC" box="[374,1213,301,328]" pageId="18" pageNumber="19">
urn:lsid:zoobank.org:act:
<uuid id="FFA40C61FF86513BDAD8D839FB0DFEFC" box="[661,1213,301,328]" pageId="18" pageNumber="19">339546A8-6F7C-4250-8E6B-49017F4EB6A8</uuid>
</uri>
</paragraph>
<paragraph id="8BBD36B4FF86513BDA95D845FCEBFEDF" blockId="18.[374,1213,264,363]" box="[728,859,337,363]" pageId="18" pageNumber="19">
<figureCitation id="13392A31FF86513BDA95D845FCEBFEDF" box="[728,859,337,363]" captionStart="Figs 41–44" captionStartId="19.[189,241,1999,2025]" captionTargetBox="[189,1398,1061,1964]" captionTargetId="figure-285@19.[189,1398,1061,1964]" captionTargetPageId="19" captionText="Figs 41–44. Andrena elam Wood sp. nov., ♀ (OÖLM). 41. Profile. 42. Face. 43. Dorsum. 44. Terga." figureDoi="http://doi.org/10.5281/zenodo.7222336" httpUri="https://zenodo.org/record/7222336/files/figure.png" pageId="18" pageNumber="19">Figs 41–44</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C318653FFF86513BD8F0D887FDECFCF1" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8BBD36B4FF86513BD8F0D887FE86FE19" blockId="18.[189,310,403,429]" box="[189,310,403,429]" pageId="18" pageNumber="19">
<heading id="D0F581D8FF86513BD8F0D887FE86FE19" bold="true" box="[189,310,403,429]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B976EAA6FF86513BD8F0D887FE86FE19" bold="true" box="[189,310,403,429]" pageId="18" pageNumber="19">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FF86513BD8F0D8D6FDECFCF1" blockId="18.[189,1401,450,837]" pageId="18" pageNumber="19">
<taxonomicName id="4C024D37FF86513BD8F0D8D6FED6FE68" authority="Wood &amp; Monfared, 2022" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[189,358,450,476]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="elam" status="sp. nov.">
<emphasis id="B976EAA6FF86513BD8F0D8D6FED6FE68" box="[189,358,450,476]" italics="true" pageId="18" pageNumber="19">Andrena elam</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557DDFF86513BD93FD8D7FE62FE69" box="[370,466,451,477]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
can be placed in the subgenus
<taxonomicName id="4C024D37FF86513BDB2DD8D6FB71FE69" authority="Ashmead, 1899" authorityName="Ashmead" authorityYear="1899" box="[864,1217,450,477]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="subGenus" subGenus="Micrandrena">
<emphasis id="B976EAA6FF86513BDB2DD8D6FC4AFE68" box="[864,1018,450,476]" italics="true" pageId="18" pageNumber="19">Micrandrena</emphasis>
Ashmead, 1899
</taxonomicName>
because of its small body size (
<figureCitation id="13392A31FF86513BD9C0D8F2FE56FDB5" box="[397,486,486,513]" captionStart="Figs 41–44" captionStartId="19.[189,241,1999,2025]" captionTargetBox="[189,1398,1061,1964]" captionTargetId="figure-285@19.[189,1398,1061,1964]" captionTargetPageId="19" captionText="Figs 41–44. Andrena elam Wood sp. nov., ♀ (OÖLM). 41. Profile. 42. Face. 43. Dorsum. 44. Terga." figureDoi="http://doi.org/10.5281/zenodo.7222336" httpUri="https://zenodo.org/record/7222336/files/figure.png" pageId="18" pageNumber="19">Fig. 41</figureCitation>
), black integument, and strongly rugose propodeal triangle. It has a domed clypeus with faint latitudinal striations (
<figureCitation id="13392A31FF86513BDADEDB1EFD5AFD91" box="[659,746,522,549]" captionStart="Figs 41–44" captionStartId="19.[189,241,1999,2025]" captionTargetBox="[189,1398,1061,1964]" captionTargetId="figure-285@19.[189,1398,1061,1964]" captionTargetPageId="19" captionText="Figs 41–44. Andrena elam Wood sp. nov., ♀ (OÖLM). 41. Profile. 42. Face. 43. Dorsum. 44. Terga." figureDoi="http://doi.org/10.5281/zenodo.7222336" httpUri="https://zenodo.org/record/7222336/files/figure.png" pageId="18" pageNumber="19">Fig. 42</figureCitation>
) (common in many Turkish
<taxonomicName id="4C024D37FF86513BDC71DB1EFB66FD90" authorityName="Ashmead" authorityYear="1899" box="[1084,1238,522,548]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="133" phylum="Arthropoda" rank="subGenus" subGenus="Micrandrena">
<emphasis id="B976EAA6FF86513BDC71DB1EFB66FD90" box="[1084,1238,522,548]" italics="true" pageId="18" pageNumber="19">Micrandrena</emphasis>
</taxonomicName>
and typically reminicient of
<taxonomicName id="4C024D37FF86513BD921DB3BFCACFDFD" authority="Noskiewicz, 1939" authorityName="Noskiewicz" authorityYear="1939" box="[364,796,558,585]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="simontornyella">
<emphasis id="B976EAA6FF86513BD921DB3BFD8DFDFC" box="[364,573,558,585]" italics="true" pageId="18" pageNumber="19">A. simontornyella</emphasis>
Noskiewicz, 1939
</taxonomicName>
), and shiny and densely punctate scutum (
<figureCitation id="13392A31FF86513BDD5CDB3AFAD9FDFD" box="[1297,1385,558,585]" captionStart="Figs 41–44" captionStartId="19.[189,241,1999,2025]" captionTargetBox="[189,1398,1061,1964]" captionTargetId="figure-285@19.[189,1398,1061,1964]" captionTargetPageId="19" captionText="Figs 41–44. Andrena elam Wood sp. nov., ♀ (OÖLM). 41. Profile. 42. Face. 43. Dorsum. 44. Terga." figureDoi="http://doi.org/10.5281/zenodo.7222336" httpUri="https://zenodo.org/record/7222336/files/figure.png" pageId="18" pageNumber="19">Fig. 43</figureCitation>
). This places it close to
<taxonomicName id="4C024D37FF86513BD995DB47FCA0FDD9" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" box="[472,784,594,621]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="puffina">
<emphasis id="B976EAA6FF86513BD995DB47FDFEFDD9" box="[472,590,595,621]" italics="true" pageId="18" pageNumber="19">A. puffina</emphasis>
<bibRefCitation id="EF934B45FF86513BDA17DB47FCA0FDD9" author="Warncke" box="[602,784,594,621]" firstAuthor="Warncke" pageId="18" pageNumber="19" pagination="29 - 102" refId="ref70340" refString="Warncke K. 1975. Die Sandbienen der Turkei (Hymenoptera, Apoidea, Andrena), Teil B. Mitteilungen der Munchner Entomologischen Gesellschaft 65: 29 - 102." type="journal article" year="1975">Warncke, 1975</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C024D37FF86513BDB1CDB47FB3AFDD9" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" box="[849,1162,594,621]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="stolida">
<emphasis id="B976EAA6FF86513BDB1CDB47FC76FDD9" box="[849,966,595,621]" italics="true" pageId="18" pageNumber="19">A. stolida</emphasis>
<bibRefCitation id="EF934B45FF86513BDB9CDB47FB3AFDD9" author="Warncke" box="[977,1162,594,621]" firstAuthor="Warncke" pageId="18" pageNumber="19" pagination="29 - 102" refId="ref70340" refString="Warncke K. 1975. Die Sandbienen der Turkei (Hymenoptera, Apoidea, Andrena), Teil B. Mitteilungen der Munchner Entomologischen Gesellschaft 65: 29 - 102." type="journal article" year="1975">Warncke, 1975</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C024D37FF86513BDCD4DB47FAF2FDD9" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[1177,1346,595,621]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="elam">
<emphasis id="B976EAA6FF86513BDCD4DB47FAF2FDD9" box="[1177,1346,595,621]" italics="true" pageId="18" pageNumber="19">Andrena elam</emphasis>
</taxonomicName>
can immediately be separated from both these taxa by the sculpturing of the terga which have extremely strong, raised microreticulation covering the entirety of T1 and the discs of T2–4 with large, dense punctures intermixed throughout, punctures particularly large and coarse laterally (
<figureCitation id="13392A31FF86513BDCECDBABFB4CFD6D" box="[1185,1276,703,729]" captionStart="Figs 41–44" captionStartId="19.[189,241,1999,2025]" captionTargetBox="[189,1398,1061,1964]" captionTargetId="figure-285@19.[189,1398,1061,1964]" captionTargetPageId="19" captionText="Figs 41–44. Andrena elam Wood sp. nov., ♀ (OÖLM). 41. Profile. 42. Face. 43. Dorsum. 44. Terga." figureDoi="http://doi.org/10.5281/zenodo.7222336" httpUri="https://zenodo.org/record/7222336/files/figure.png" pageId="18" pageNumber="19">Fig. 44</figureCitation>
).
<taxonomicName id="4C024D37FF86513BDD5FDBABFEBEFD49" authorityName="Warncke" authorityYear="1975" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="puffina">
<emphasis id="B976EAA6FF86513BDD5FDBABFEBEFD49" italics="true" pageId="18" pageNumber="19">Andrena puffina</emphasis>
</taxonomicName>
has the terga finely and subtly punctate, and
<taxonomicName id="4C024D37FF86513BDB6FDBF7FC23FD49" authorityName="Warncke" authorityYear="1975" box="[802,915,739,765]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="stolida">
<emphasis id="B976EAA6FF86513BDB6FDBF7FC23FD49" box="[802,915,739,765]" italics="true" pageId="18" pageNumber="19">A. stolida</emphasis>
</taxonomicName>
has the terga with fine punctures that are not clearly visible against the underlying shagreen, not becoming noticeably coarser laterally. See also
<taxonomicName id="4C024D37FF86513BD8F3DA3FFED4FCF1" authority="Wood &amp; Monfared, 2022" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[190,356,811,837]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="subviridula" status="sp. nov.">
<emphasis id="B976EAA6FF86513BD8F3DA3FFED4FCF1" box="[190,356,811,837]" italics="true" pageId="18" pageNumber="19">A. subviridula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557DDFF86513BD926DA3FFE75FCF1" box="[363,453,811,837]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
(see below).
</paragraph>
</subSubSection>
<subSubSection id="C318653FFF86513BD8F0DA79FD6BFC6E" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8BBD36B4FF86513BD8F0DA79FEF4FC33" blockId="18.[189,324,877,903]" box="[189,324,877,903]" pageId="18" pageNumber="19">
<heading id="D0F581D8FF86513BD8F0DA79FEF4FC33" bold="true" box="[189,324,877,903]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B976EAA6FF86513BD8F0DA79FEF4FC33" bold="true" box="[189,324,877,903]" pageId="18" pageNumber="19">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FF86513BD8F0DA88FD6BFC6E" blockId="18.[188,1398,924,987]" pageId="18" pageNumber="19">
Named after the ancient kingdom of Elam (ca 3000–500 BC, noun in apposition) that was largely found in the region that is now called
<collectingRegion id="49C6F856FF86513BDA60DAD4FD67FC6E" box="[557,727,960,986]" country="Iran" name="Ilam" pageId="18" pageNumber="19">Ilam Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C318653FFF86513BD8F0DD16FE15FADD" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="8BBD36B4FF86513BD8F0DD16FE1AFBA8" blockId="18.[189,426,1026,1052]" box="[189,426,1026,1052]" pageId="18" pageNumber="19">
<heading id="D0F581D8FF86513BD8F0DD16FE1AFBA8" bold="true" box="[189,426,1026,1052]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B976EAA6FF86513BD8F0DD16FE1AFBA8" bold="true" box="[189,426,1026,1052]" pageId="18" pageNumber="19">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FF86513BD8A7DD26FAC7FBC6" blockId="18.[189,1399,1074,1138]" pageId="18" pageNumber="19">
<materialsCitation id="3B6A3CE9FF86513BD8A7DD26FAC7FBC6" ID-GBIF-Occurrence="3944043714" collectingDate="2016-05-12" collectionCode="OOLM" collectorName="M. Kafka" country="Iran" elevation="1830" location="Dinar Gaouh" municipality="Abda Man" pageId="18" pageNumber="19" specimenCount="1" specimenCount-female="1" stateProvince="Ilam Province" typeStatus="holotype">
<emphasis id="B976EAA6FF86513BD8A7DD26FEECFBF8" bold="true" box="[234,348,1074,1100]" pageId="18" pageNumber="19">
<typeStatus id="54B98816FF86513BD8A7DD26FEECFBF8" box="[234,348,1074,1100]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3157624FF86513BD8F0DD42FEB5FBC4" box="[189,261,1110,1136]" name="Iran" pageId="18" pageNumber="19">IRAN</collectingCountry>
•
<specimenCount id="9D04FD3DFF86513BD956DD42FE82FBC4" box="[283,306,1110,1136]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
;
<collectingRegion id="49C6F856FF86513BD90DDD42FE57FBC4" box="[320,487,1110,1136]" country="Iran" name="Ilam" pageId="18" pageNumber="19">Ilam Province</collectingRegion>
,
<collectingMunicipality id="6BD9ACCEFF86513BD9BDDD42FDDFFBC4" box="[496,623,1110,1136]" pageId="18" pageNumber="19">Abda Man</collectingMunicipality>
,
<location id="8EDD606FFF86513BDA34DD42FCA4FBC4" LSID="urn:lsid:plazi:treatment:03AB87A2FF86513DDA42D81CFCB5FE03:8EDD606FFF86513BDA34DD42FCA4FBC4" box="[633,788,1110,1136]" country="Iran" municipality="Abda Man" name="Dinar Gaouh" pageId="18" pageNumber="19" stateProvince="Ilam Province">Dinar Gaouh</location>
;
<elevation id="002FD187FF86513BDB53DD42FC05FBC4" box="[798,949,1110,1136]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="18" pageNumber="19" unit="m" value="1830.0">
<quantity id="4CFA9B51FF86513BDB53DD42FCC6FBC4" box="[798,886,1110,1136]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="18" pageNumber="19" unit="m" value="1830.0">1830 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBC1074FF86513BDB8CDD42FBEFFBC4" box="[961,1119,1110,1136]" pageId="18" pageNumber="19" value="2016-05-12">
<collectingDate id="EFF8E99CFF86513BDB8CDD42FBEFFBC4" box="[961,1119,1110,1136]" pageId="18" pageNumber="19" value="2016-05-12">12 May 2016</collectingDate>
</date>
;
<collectorName id="26F75362FF86513BDC27DD42FB69FBC4" box="[1130,1241,1110,1136]" pageId="18" pageNumber="19">M. Kafka</collectorName>
leg.;
<collectionCode id="ED13AE71FF86513BDD54DD42FAC3FBC6" box="[1305,1395,1110,1138]" collectionName="OOLM" pageId="18" pageNumber="19">OÖLM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBD36B4FF86513BD8A7DD8CFE15FADD" blockId="18.[189,1399,1176,1385]" pageId="18" pageNumber="19">
<materialsCitation id="3B6A3CE9FF86513BD8A7DD8CFEA3FB49" ID-GBIF-Occurrence="3944043356" collectingDate="2016-05-12" collectionCode="OOLM" collectorName="M. Kafka" country="Iran" elevation="1830" location="Dinar Gaouh" municipality="Abda Man" pageId="18" pageNumber="19" specimenCount="9" specimenCount-female="9" stateProvince="Ilam Province" typeStatus="paratype">
<emphasis id="B976EAA6FF86513BD8A7DD8CFED7FB06" bold="true" box="[234,359,1176,1202]" pageId="18" pageNumber="19">
<typeStatus id="54B98816FF86513BD8A7DD8CFED7FB06" box="[234,359,1176,1202]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F3157624FF86513BD8F0DDA8FEB5FB62" box="[189,261,1212,1238]" name="Iran" pageId="18" pageNumber="19">IRAN</collectingCountry>
•
<specimenCount id="9D04FD3DFF86513BD96FDDA8FED7FB63" box="[290,359,1212,1239]" pageId="18" pageNumber="19" type="female">9 ♀♀</specimenCount>
;
<collectingRegion id="49C6F856FF86513BD93ADDA8FD91FB62" box="[375,545,1212,1238]" country="Iran" name="Ilam" pageId="18" pageNumber="19">Ilam Province</collectingRegion>
,
<collectingMunicipality id="6BD9ACCEFF86513BDA63DDA8FD00FB63" box="[558,688,1212,1239]" pageId="18" pageNumber="19">Abda Man</collectingMunicipality>
,
<location id="8EDD606FFF86513BDAF0DDA8FCEBFB62" LSID="urn:lsid:plazi:treatment:03AB87A2FF86513DDA42D81CFCB5FE03:8EDD606FFF86513BDAF0DDA8FCEBFB62" box="[701,859,1212,1238]" country="Iran" municipality="Abda Man" name="Dinar Gaouh" pageId="18" pageNumber="19" stateProvince="Ilam Province">Dinar Gaouh</location>
;
<elevation id="002FD187FF86513BDB24DDA8FBB6FB62" box="[873,1030,1212,1239]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="18" pageNumber="19" unit="m" value="1830.0">
<quantity id="4CFA9B51FF86513BDB24DDA8FC74FB63" box="[873,964,1212,1239]" metricMagnitude="3" metricUnit="m" metricValue="1.83" pageId="18" pageNumber="19" unit="m" value="1830.0">1830 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBC1074FF86513BDC58DDA8FB09FB62" box="[1045,1209,1212,1239]" pageId="18" pageNumber="19" value="2016-05-12">
<collectingDate id="EFF8E99CFF86513BDC58DDA8FB09FB62" box="[1045,1209,1212,1239]" pageId="18" pageNumber="19" value="2016-05-12">12 May 2016</collectingDate>
</date>
;
<collectorName id="26F75362FF86513BDC8ADDA9FA8AFB62" box="[1223,1338,1212,1239]" pageId="18" pageNumber="19">M. Kafka</collectorName>
leg.;
<collectionCode id="ED13AE71FF86513BD8F0DDF5FEA3FB49" box="[189,275,1249,1277]" collectionName="OOLM" pageId="18" pageNumber="19">OÖLM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B6A3CE9FF86513BD963DDF4FEA3FA95" ID-GBIF-Occurrence="3944043903" collectingDate="2016-05-11" collectionCode="OOLM" collectorName="M. Kafka" country="Iran" elevation="670" location="Dorud Lanjaban" pageId="18" pageNumber="19" specimenCount="16" specimenCount-female="16" stateProvince="Lorestan Province" typeStatus="paratype">
<specimenCount id="9D04FD3DFF86513BD963DDF4FE31FB4F" box="[302,385,1248,1275]" pageId="18" pageNumber="19" type="female">16 ♀♀</specimenCount>
;
<collectingRegion id="49C6F856FF86513BD9DCDDF5FDDBFB4E" box="[401,619,1248,1275]" country="Iran" name="Lorestan" pageId="18" pageNumber="19">Lorestan Province</collectingRegion>
,
<location id="8EDD606FFF86513BDA35DDF4FC88FB4E" LSID="urn:lsid:plazi:treatment:03AB87A2FF86513DDA42D81CFCB5FE03:8EDD606FFF86513BDA35DDF4FC88FB4E" box="[632,824,1248,1274]" country="Iran" name="Dorud Lanjaban" pageId="18" pageNumber="19" stateProvince="Lorestan Province">Dorud Lanjaban</location>
env.;
<elevation id="002FD187FF86513BDBCCDDF4FBBDFB4E" box="[897,1037,1248,1275]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="18" pageNumber="19" unit="m" value="670.0">
<quantity id="4CFA9B51FF86513BDBCCDDF4FC7CFB4F" box="[897,972,1248,1275]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="18" pageNumber="19" unit="m" value="670.0">670 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBC1074FF86513BDC56DDF4FB0CFB4E" box="[1051,1212,1248,1275]" pageId="18" pageNumber="19" value="2016-05-11">
<collectingDate id="EFF8E99CFF86513BDC56DDF4FB0CFB4E" box="[1051,1212,1248,1275]" pageId="18" pageNumber="19" value="2016-05-11">11 May 2016</collectingDate>
</date>
;
<collectorName id="26F75362FF86513BDC84DDF5FA8BFB4E" box="[1225,1339,1248,1275]" pageId="18" pageNumber="19">M. Kafka</collectorName>
leg.;
<collectionCode id="ED13AE71FF86513BD8F0DC11FEA3FA95" box="[189,275,1285,1313]" collectionName="OOLM" pageId="18" pageNumber="19">OÖLM</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B6A3CE9FF86513BD963DC10FC27FAAA" ID-GBIF-Occurrence="3944044025" box="[302,919,1284,1311]" collectingDate="2016-05-11" collectionCode="TJWC" collectorName="M. Kafka" country="Iran" elevation="670" location="Dorud Lanjaban" pageId="18" pageNumber="19" specimenCount="5" specimenCount-female="5" stateProvince="Lorestan" typeStatus="paratype">
<specimenCount id="9D04FD3DFF86513BD963DC10FEC2FAAB" box="[302,370,1284,1311]" pageId="18" pageNumber="19" type="female">5 ♀♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED13AE71FF86513BDB07DC10FC27FAAA" box="[842,919,1284,1310]" collectionName="TJWC" pageId="18" pageNumber="19">TJWC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B6A3CE9FF86513BDBFCDC10FE11FADD" ID-GBIF-Occurrence="3944043325" collectingDate="2016-05-04" collectionCode="OOLM" collectorName="M. Kafka" country="Iran" elevation="2030" location="Sarb-e Taveh" municipality="Yasuj" pageId="18" pageNumber="19" specimenCount="2" specimenCount-female="2" stateProvince="Fars Province" typeStatus="paratype">
<specimenCount id="9D04FD3DFF86513BDBFCDC10FC45FAAB" box="[945,1013,1284,1311]" pageId="18" pageNumber="19" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="49C6F856FF86513BDC49DC11FB17FAAA" box="[1028,1191,1284,1311]" country="Iran" name="Fars" pageId="18" pageNumber="19">Fars Province</collectingRegion>
[Kohgiluyeh and Boyer-Ahmad Province],
<collectingMunicipality id="6BD9ACCEFF86513BD9A6DC3CFD9DFAF6" box="[491,557,1320,1346]" pageId="18" pageNumber="19">Yasuj</collectingMunicipality>
[Yasouj],
<location id="8EDD606FFF86513BDAEBDC3CFCF0FAF6" LSID="urn:lsid:plazi:treatment:03AB87A2FF86513DDA42D81CFCB5FE03:8EDD606FFF86513BDAEBDC3CFCF0FAF6" box="[678,832,1320,1346]" country="Iran" municipality="Yasuj" name="Sarb-e Taveh" pageId="18" pageNumber="19" stateProvince="Fars Province">Sarb-e Taveh</location>
[Sarab-e Taveh];
<elevation id="002FD187FF86513BDC5FDC3CFB1BFAF6" box="[1042,1195,1320,1347]" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="18" pageNumber="19" unit="m" value="2030.0">
<quantity id="4CFA9B51FF86513BDC5FDC3CFBDBFAF7" box="[1042,1131,1320,1347]" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="18" pageNumber="19" unit="m" value="2030.0">2030 m</quantity>
a.s.l.
</elevation>
;
<date id="FFBC1074FF86513BDCF5DC3CFAF9FAF6" box="[1208,1353,1320,1347]" pageId="18" pageNumber="19" value="2016-05-04">
<collectingDate id="EFF8E99CFF86513BDCF5DC3CFAF9FAF6" box="[1208,1353,1320,1347]" pageId="18" pageNumber="19" value="2016-05-04">4 May 2016</collectingDate>
</date>
;
<collectorName id="26F75362FF86513BDD18DC3DFEB5FAD2" pageId="18" pageNumber="19">M. Kafka</collectorName>
leg.;
<collectionCode id="ED13AE71FF86513BD90ADC59FE11FADD" box="[327,417,1357,1385]" collectionName="OOLM" pageId="18" pageNumber="19">OÖLM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C318653FFF86513DD8F0DC9AFE89FEF2" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="description">
<paragraph id="8BBD36B4FF86513BD8F0DC9AFEFEFA1C" blockId="18.[189,334,1422,1448]" box="[189,334,1422,1448]" pageId="18" pageNumber="19">
<heading id="D0F581D8FF86513BD8F0DC9AFEFEFA1C" bold="true" box="[189,334,1422,1448]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="B976EAA6FF86513BD8F0DC9AFEFEFA1C" bold="true" box="[189,334,1422,1448]" pageId="18" pageNumber="19">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FF86513BD8A7DCAAFEF5FA6C" blockId="18.[189,767,1470,1533]" box="[234,325,1470,1496]" pageId="18" pageNumber="19">
<emphasis id="B976EAA6FF86513BD8A7DCAAFEF5FA6C" bold="true" box="[234,325,1470,1496]" pageId="18" pageNumber="19">Female</emphasis>
</paragraph>
<paragraph id="8BBD36B4FF86513BD8F0DCF6FD4FFA48" blockId="18.[189,767,1470,1533]" box="[189,767,1506,1533]" pageId="18" pageNumber="19">
MEASUREMENTS. Body length
<quantity id="4CFA9B51FF86513BDA5EDCF6FD3AFA48" box="[531,650,1506,1533]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="18" pageNumber="19" unit="mm" value="6.75" valueMax="7.0" valueMin="6.5">6.5–7 mm</quantity>
(
<figureCitation id="13392A31FF86513BDAD7DCF6FD42FA48" box="[666,754,1506,1532]" captionStart="Figs 41–44" captionStartId="19.[189,241,1999,2025]" captionTargetBox="[189,1398,1061,1964]" captionTargetId="figure-285@19.[189,1398,1061,1964]" captionTargetPageId="19" captionText="Figs 41–44. Andrena elam Wood sp. nov., ♀ (OÖLM). 41. Profile. 42. Face. 43. Dorsum. 44. Terga." figureDoi="http://doi.org/10.5281/zenodo.7222336" httpUri="https://zenodo.org/record/7222336/files/figure.png" pageId="18" pageNumber="19">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="8BBD36B4FF86513BD8F0DF31FD8BF8EB" blockId="18.[188,1400,1572,1887]" pageId="18" pageNumber="19">
HEAD. 1.2 times as wide as long. Clypeus dark, domed, punctate, punctures separated by 0.5–3 puncture diameters. Clypeus surface microreticulate, centrally and basally with weakly raised microreticulation giving impression of weak latitudinal striations, becoming obscure apically; underlying surface weakly shining (
<figureCitation id="13392A31FF86513BD96ADF84FE33F91E" box="[295,387,1680,1706]" captionStart="Figs 41–44" captionStartId="19.[189,241,1999,2025]" captionTargetBox="[189,1398,1061,1964]" captionTargetId="figure-285@19.[189,1398,1061,1964]" captionTargetPageId="19" captionText="Figs 41–44. Andrena elam Wood sp. nov., ♀ (OÖLM). 41. Profile. 42. Face. 43. Dorsum. 44. Terga." figureDoi="http://doi.org/10.5281/zenodo.7222336" httpUri="https://zenodo.org/record/7222336/files/figure.png" pageId="18" pageNumber="19">Fig. 42</figureCitation>
). Process of labrum narrow, weakly tongue-shaped, slightly wider than long, apical margin weakly rounded. Face, gena, vertex, and scape with short whitish to brownish hairs. Gena slightly exceeding width of compound eye; ocelloccipital distance slightly shorter than diameter of lateral ocellus. Foveae narrow, dorsally occupying ¼ distance between compound eye and lateral ocellus, subequal to width of flagellum, of uniform width throughout; foveae filled with light brownish hairs. Antennae dark, A3=A4+5.
</paragraph>
<paragraph id="8BBD36B4FF86513AD8F0DE93FD4FFDD4" blockId="18.[188,1399,1926,2025]" lastBlockId="19.[188,1399,265,608]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
MESOSOMA. Scutum and scutellum polished, smooth and shiny over majority of area, with shagreen in anterior 1/5 and laterally as scutum descends to pronotum and pronotal lobes, otherwise without shagreen (
<figureCitation id="13392A31FF86513BD888DEDAFEAFF85C" box="[197,287,1998,2024]" captionStart="Figs 41–44" captionStartId="19.[189,241,1999,2025]" captionTargetBox="[189,1398,1061,1964]" captionTargetId="figure-285@19.[189,1398,1061,1964]" captionTargetPageId="19" captionText="Figs 41–44. Andrena elam Wood sp. nov., ♀ (OÖLM). 41. Profile. 42. Face. 43. Dorsum. 44. Terga." figureDoi="http://doi.org/10.5281/zenodo.7222336" httpUri="https://zenodo.org/record/7222336/files/figure.png" pageId="18" pageNumber="19">Fig. 43</figureCitation>
). Scutum and scutellum regularly and clearly punctate, punctures separated by 1–2 puncture diameters. Pronotum without humeral angle. Mesepisternum and lateral and dorsolateral faces of propodeum with fine granular shagreen, weakly shining; mesespisternum anteriorly and dorsolateral faces of propodeum with fine network of raised rugosity. Propodeal triangle delineated laterally with fine carinae, internal surface with fine network of rugae, thus weakly differentiated from sculpture of dorsolateral faces of propodeum. Mesosoma with moderately long whitish hairs; propodeal corbiculae thin, composed of sparse whitish plumose hairs, internal surface with numerous simple white hairs. Legs uniformly dark, pubescence whitish to light brownish; tibial scopae whitish ventrally, light brownish dorsally; femoral scopae and flocculus white. Hind tarsal claws with inner tooth. Wings hyaline, stigma dark brown, venation dark orange to dark brown, nervulus interstitial to weakly antefurcal; first recurrent vein enters second submarginal cell at its middle.
</paragraph>
<paragraph id="8BBD36B4FF87513AD8F0DB9CFD64FC6A" blockId="19.[186,1400,647,990]" pageId="19" pageNumber="20">
METASOMA. Terga dark, apical margins of T1–4 slightly depressed, depression weakest on T1, strongest on T4, all marginal areas unicolourous black. Tergal discs with extremely dense raised microreticulation, tergal discs also densely punctate, punctures separated by 0.5–1 puncture diameter, punctures and microreticulation combining to
<taxonomicName id="4C024D37FF87513ADA7EDBE5FD06FCBF" box="[563,694,753,779]" form="dense" pageId="19" pageNumber="20" rank="form">form dense</taxonomicName>
network of bead-like punctures separated by slightly raised margins; integument dull (
<figureCitation id="13392A31FF87513ADA4BDA00FDD4FC9A" box="[518,612,788,814]" captionStart="Figs 41–44" captionStartId="19.[189,241,1999,2025]" captionTargetBox="[189,1398,1061,1964]" captionTargetId="figure-285@19.[189,1398,1061,1964]" captionTargetPageId="19" captionText="Figs 41–44. Andrena elam Wood sp. nov., ♀ (OÖLM). 41. Profile. 42. Face. 43. Dorsum. 44. Terga." figureDoi="http://doi.org/10.5281/zenodo.7222336" httpUri="https://zenodo.org/record/7222336/files/figure.png" pageId="19" pageNumber="20">Fig. 44</figureCitation>
). Entirety of T1 covered with dense microreticulation, discs of T2–4 with microreticulation, becoming progressively weaker, disc of T4 without bead-like punctures, punctures distinct, without raised margins. Marginal areas of T2–4 impunctate, without microreticulation, shagreened and weakly shining. T2–4 medially almost hairless, laterally with weak whitish apical hair fringes, obscure. Tergal hairbands absent. Apical fringe of T5 and hairs flanking pygidial plate dark brown; pygidial plate rounded triangular, flat.
</paragraph>
<caption id="DF7D663CFF87513AD8F0DEDBFAE0F85E" ID-DOI="http://doi.org/10.5281/zenodo.7222336" ID-Zenodo-Dep="7222336" box="[189,1360,1999,2028]" httpUri="https://zenodo.org/record/7222336/files/figure.png" pageId="19" pageNumber="20" startId="19.[189,241,1999,2025]" targetBox="[189,1398,1061,1964]" targetPageId="19">
<paragraph id="8BBD36B4FF87513AD8F0DEDBFAE0F85E" blockId="19.[189,1360,1999,2028]" box="[189,1360,1999,2028]" pageId="19" pageNumber="20">
<emphasis id="B976EAA6FF87513AD8F0DEDBFEF9F85D" bold="true" box="[189,329,1999,2025]" pageId="19" pageNumber="20">Figs 41–44.</emphasis>
<taxonomicName id="4C024D37FF87513AD91DDEDBFDF1F85D" authority="Wood" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[336,577,1999,2025]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="elam" status="sp. nov.">
<emphasis id="B976EAA6FF87513AD91DDEDBFE45F85D" box="[336,501,1999,2025]" italics="true" pageId="19" pageNumber="20">Andrena elam</emphasis>
Wood
</taxonomicName>
<taxonomicNameLabel id="A24557DDFF87513ADA05DEC4FD15F85E" box="[584,677,2000,2026]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
, ♀ (OÖLM).
<emphasis id="B976EAA6FF87513ADB0ADEDBFCD4F85D" bold="true" box="[839,868,1999,2025]" pageId="19" pageNumber="20">41</emphasis>
. Profile.
<emphasis id="B976EAA6FF87513ADB9DDEDBFC5DF85D" bold="true" box="[976,1005,1999,2025]" pageId="19" pageNumber="20">42</emphasis>
. Face.
<emphasis id="B976EAA6FF87513ADC0FDEDBFBEFF85D" bold="true" box="[1090,1119,1999,2025]" pageId="19" pageNumber="20">43</emphasis>
. Dorsum.
<emphasis id="B976EAA6FF87513ADC96DEDBFB48F85D" bold="true" box="[1243,1272,1999,2025]" pageId="19" pageNumber="20">44</emphasis>
. Terga.
</paragraph>
</caption>
<paragraph id="8BBD36B4FF80513DD8A7D81DFE9AFE97" blockId="20.[189,313,265,326]" box="[234,298,265,291]" pageId="20" pageNumber="21">
<emphasis id="B976EAA6FF80513DD8A7D81DFE9AFE97" bold="true" box="[234,298,265,291]" pageId="20" pageNumber="21">Male</emphasis>
</paragraph>
<paragraph id="8BBD36B4FF80513DD8F0D838FE89FEF2" blockId="20.[189,313,265,326]" box="[189,313,300,326]" pageId="20" pageNumber="21">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C318653FFF80513DD8F0D879FCB5FE03" box="[189,773,365,439]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BBD36B4FF80513DD8F0D879FEE6FE33" blockId="20.[189,342,365,391]" box="[189,342,365,391]" pageId="20" pageNumber="21">
<heading id="D0F581D8FF80513DD8F0D879FEE6FE33" bold="true" box="[189,342,365,391]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B976EAA6FF80513DD8F0D879FEE6FE33" bold="true" box="[189,342,365,391]" pageId="20" pageNumber="21">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FF80513DD8F0D888FCB5FE03" blockId="20.[189,773,412,439]" box="[189,773,412,439]" pageId="20" pageNumber="21">
Southern
<collectingCountry id="F3157624FF80513DD963D889FEEDFE03" box="[302,349,413,439]" name="Iran" pageId="20" pageNumber="21">Iran</collectingCountry>
(provinces of
<collectingRegion id="49C6F856FF80513DDA45D888FDF1FE02" box="[520,577,412,438]" country="Iran" name="Ilam" pageId="20" pageNumber="21">Ilam</collectingRegion>
,
<collectingRegion id="49C6F856FF80513DDA01D889FD06FE03" box="[588,694,413,439]" country="Iran" name="Lorestan" pageId="20" pageNumber="21">Lorestan</collectingRegion>
,
<collectingRegion id="49C6F856FF80513DDA8CD889FD48FE03" box="[705,760,413,439]" country="Iran" name="Fars" pageId="20" pageNumber="21">Fars</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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