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<document id="CE18F5FD2C48296DE74E0D185277DE9B" ID-DOI="10.1649/0010-065X-73.3.687" ID-ISSN="1938-4394" ID-Zenodo-Dep="5402498" ID-ZooBank="urn:lsid:zoobank.org:pub:6044D8D0-770C-4904-90CF-E0184D4F3BD5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630337023334" checkinUser="valdenar" docAuthor="Delgado, Leonardo &amp; Mora-Aguilar, Eder F." docDate="2019" docId="03A66664FF86FF88A885FF21FE2EFBB1" docLanguage="en" docName="0010-065X-73.3.687.pdf" docOrigin="The Coleopterists Bulletin 73 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-73.3.687" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Verodes chimalapensis Delgado and Mora-Aguilar 2019, new species" docType="treatment" docUuid="8CD13586-6C8D-47F5-A68C-3AFC9D899F90" docUuidSource="ZooBank" docVersion="4" lastPageNumber="690" masterDocId="FF9F1E1CFF84FF8CA835FFCEFF9BFF93" masterDocTitle="A New Mexican Species and Distributional Comments of Verodes Casey (Coleoptera: Zopheridae: Zopherinae)" masterLastPageNumber="692" masterPageNumber="687" pageNumber="688" updateTime="1699234001971" updateUser="plazi">
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<mods:title id="F3C162C265BBE93796E70A88EA3661B0">A New Mexican Species and Distributional Comments of Verodes Casey (Coleoptera: Zopheridae: Zopherinae)</mods:title>
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<mods:namePart id="C91648E8D92E850C06F425D132D7CAEF">Delgado, Leonardo</mods:namePart>
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<mods:namePart id="CDA31AE3920FE487B58D7776B7A0BD57">Mora-Aguilar, Eder F.</mods:namePart>
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<mods:typeOfResource id="BF495BD1ABD06C752CDFE4675196A28E">text</mods:typeOfResource>
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<mods:title id="C52074B0D308A5621B1B20D4E76EDC74">The Coleopterists Bulletin</mods:title>
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<mods:date id="359F2989145615B7C3926F9F539FF367">2019</mods:date>
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<mods:number id="E7C2BD271BB5D19EB30EF4854B4758D9">2019-09-22</mods:number>
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<mods:classification id="B04CF91F79C65C6119DFD1FAE9871069">journal article</mods:classification>
<mods:identifier id="3971817D2CC45408D05720C716897007" type="DOI">10.1649/0010-065X-73.3.687</mods:identifier>
<mods:identifier id="5D84A0470EC992C15B3AD463FBBAAFDE" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03A66664FF86FF88A885FF21FE2EFBB1" LSID="urn:lsid:zoobank.org:act:8CD13586-6C8D-47F5-A68C-3AFC9D899F90" httpUri="http://treatment.plazi.org/id/03A66664FF86FF88A885FF21FE2EFBB1" lastPageId="4" lastPageNumber="690" pageId="2" pageNumber="688">
<subSubSection id="C31584F9FF86FF8EA885FF21FE57FEFC" pageId="2" pageNumber="688" type="nomenclature">
<paragraph id="8BB0D772FF86FF8EA885FF21FE69FE8C" blockId="2.[171,618,239,367]" pageId="2" pageNumber="688">
<heading id="D0F8601EFF86FF8EA885FF21FE69FE8C" centered="true" fontSize="9" level="2" pageId="2" pageNumber="688" reason="2">
<emphasis id="B97B0B60FF86FF8EA885FF21FE69FE8C" bold="true" pageId="2" pageNumber="688">
<taxonomicName id="4C0FACF1FF86FF8EA885FF21FE74FE97" authority="Delgado" authorityName="Delgado and Mora-Aguilar" authorityYear="2019" box="[176,495,239,260]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="688" phylum="Arthropoda" rank="species" species="chimalapensis">
<emphasis id="B97B0B60FF86FF8EA885FF21FE0FFE97" bold="true" box="[176,404,239,260]" italics="true" pageId="2" pageNumber="688">Verodes chimalapensis</emphasis>
Delgado
</taxonomicName>
and Mora- Aguilar,
<taxonomicNameLabel id="A248B61BFF86FF8EA949FEC4FE69FE8C" box="[380,498,266,287]" pageId="2" pageNumber="688" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB0D772FF86FF8EA8E1FEEAFE57FEFC" blockId="2.[171,618,239,367]" pageId="2" pageNumber="688">
Zoobank.org/
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urn:lsid:zoobank.org:act:
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</uri>
(
<figureCitation id="1334CBF7FF86FF8EA965FE94FE5EFEFC" box="[336,453,345,367]" captionStart="Figs" captionStartId="3.[197,236,1611,1630]" captionTargetBox="[177,1188,197,1593]" captionTargetId="figure-1@3.[175,1190,194,1595]" captionTargetPageId="3" captionText="Figs. 17. Verodes species. 1) V. chimalapensis, dorsal habitus; 2) V. chimalapensis, lateral habitus; 3) V. chimalapensis, dorsal surface of pronotum; 4) V. asperatus oblique habitus; 5) V. scabrosus, oblique habitus; 6) V. scabrosus, apical two-fifths of left elytron; 7) V. chimalapensis, apical two-fifths of left elytron." figureDoi="http://doi.org/10.5281/zenodo.5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="2" pageNumber="688">Figs. 13, 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31584F9FF86FF8EA89AFEB3FDC1FD12" pageId="2" pageNumber="688" type="materials_examined">
<paragraph id="8BB0D772FF86FF8EA89AFEB3FDC1FD12" blockId="2.[151,639,380,1730]" pageId="2" pageNumber="688">
<emphasis id="B97B0B60FF86FF8EA89AFEB3FED3FE01" bold="true" box="[175,328,381,402]" pageId="2" pageNumber="688">Type Material.</emphasis>
<materialsCitation id="3B67DD2FFF86FF8EA964FEB3FEE2FE6F" collectingDate="2013-06-07" collectorName="L. Delgado &amp; E. Mora-Aguilar" county="San Miguel Chimalapa" elevation="1600" latitude="16.6786" location="El Gringo" longLatPrecision="7" longitude="-94.2627" municipality="El Reten" pageId="2" pageNumber="688" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
<typeStatus id="54B469D0FF86FF8EA964FEB3FE32FE01" box="[337,425,381,402]" pageId="2" pageNumber="688" type="holotype">Holotype</typeStatus>
male labeled “MEX- ICO:
<collectingRegion id="49CB1990FF86FF8EA8E7FE59FE80FE3F" box="[210,283,407,428]" country="Mexico" name="Oaxaca" pageId="2" pageNumber="688">Oaxaca</collectingRegion>
,
<collectingCounty id="62D1AFFEFF86FF8EA91FFE59FD89FE3F" box="[298,530,407,428]" pageId="2" pageNumber="688">San Miguel Chimalapa</collectingCounty>
,
<collectingMunicipality id="6BD44D08FF86FF8EAA15FE59FDE1FE3F" box="[544,634,407,428]" pageId="2" pageNumber="688">El Retén</collectingMunicipality>
,
<location id="8ED081A9FF86FF8EA8A2FE7CFF60FE54" LSID="urn:lsid:plazi:treatment:03A66664FF86FF88A885FF21FE2EFBB1:8ED081A9FF86FF8EA8A2FE7CFF60FE54" box="[151,251,434,455]" county="San Miguel Chimalapa" latitude="16.6786" longLatPrecision="7" longitude="-94.2627" municipality="El Reten" name="El Gringo" pageId="2" pageNumber="688" stateProvince="Oaxaca">El Gringo</location>
,
<date id="FFB1F1B2FF86FF8EA93DFE7CFEF5FE54" box="[264,366,434,455]" pageId="2" pageNumber="688" value="2013-06-07">
<collectingDate id="EFF5085AFF86FF8EA93DFE7CFEF5FE54" box="[264,366,434,455]" pageId="2" pageNumber="688" value="2013-06-07">7-VI-2013</collectingDate>
</date>
,
<geoCoordinate id="EE3BB1B5FF86FF8EA94EFE7CFE75FE54" box="[379,494,434,455]" degrees="16.6786" direction="north" orientation="latitude" pageId="2" pageNumber="688" precision="5" value="16.6786">16.6786° N</geoCoordinate>
, -
<geoCoordinate id="EE3BB1B5FF86FF8EAA37FE7CFDE1FE54" box="[514,634,434,455]" degrees="94.2627" direction="west" orientation="longitude" pageId="2" pageNumber="688" precision="5" value="-94.2627">94.2627° W</geoCoordinate>
,
<quantity id="4CF77A97FF86FF8EA8A2FE02FF79FE72" box="[151,226,460,481]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="688" unit="m" value="1600.0">
<elevation id="00223041FF86FF8EA8A2FE02FF79FE72" box="[151,226,460,481]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="688" unit="m" value="1600.0">1,600m</elevation>
</quantity>
, Colecta directa en troncos,
<collectorName id="26FAB2A4FF86FF8EAA3DFE02FDE1FE72" box="[520,634,460,481]" pageId="2" pageNumber="688">L. Delgado</collectorName>
,
<collectorName id="26FAB2A4FF86FF8EA8A2FE29FEA7FE6F" box="[151,316,487,508]" pageId="2" pageNumber="688">E. Mora-Aguilar</collectorName>
cols”
</materialsCitation>
.
<materialsCitation id="3B67DD2FFF86FF8EA9B8FE29FF49FD12" collectingDate="2017-07-26" collectionCode="IEXA" collectorName="E. Arriaga-Varela" country="Mexico" county="San Miguel Chimalapa" elevation="1800" latitude="16.6637" location="Camino" longLatPrecision="7" longitude="-94.2335" municipality="El Reten" pageId="2" pageNumber="688" specimenCount="2" stateProvince="Oaxaca" typeStatus="paratype">
<typeStatus id="54B469D0FF86FF8EA9B8FE29FE7BFE6F" box="[397,480,487,508]" pageId="2" pageNumber="688" type="paratype">Paratype</typeStatus>
female labeled “
<collectingCountry id="F31897E2FF86FF8EA897FDCFFE98FD85" box="[162,259,513,534]" name="Mexico" pageId="2" pageNumber="688">MEXICO</collectingCountry>
:
<collectingRegion id="49CB1990FF86FF8EA926FDCFFEC7FD85" box="[275,348,513,534]" country="Mexico" name="Oaxaca" pageId="2" pageNumber="688">Oaxaca</collectingRegion>
,
<collectingCounty id="62D1AFFEFF86FF8EA958FDCFFDC3FD85" box="[365,600,513,534]" pageId="2" pageNumber="688">San Miguel Chimalapa</collectingCounty>
,
<collectingMunicipality id="6BD44D08FF86FF8EAA5CFDCFFF4AFDA2" pageId="2" pageNumber="688">El Retén</collectingMunicipality>
,
<location id="8ED081A9FF86FF8EA8E8FDD2FEB3FDA2" LSID="urn:lsid:plazi:treatment:03A66664FF86FF88A885FF21FE2EFBB1:8ED081A9FF86FF8EA8E8FDD2FEB3FDA2" box="[221,296,540,561]" country="Mexico" county="San Miguel Chimalapa" latitude="16.6637" longLatPrecision="7" longitude="-94.2335" municipality="El Reten" name="Camino" pageId="2" pageNumber="688" stateProvince="Oaxaca">Camino</location>
a la Torre,
<date id="FFB1F1B2FF86FF8EA995FDD2FD81FDA2" box="[416,538,540,561]" pageId="2" pageNumber="688" value="2017-07-26">
<collectingDate id="EFF5085AFF86FF8EA995FDD2FD81FDA2" box="[416,538,540,561]" pageId="2" pageNumber="688" value="2017-07-26">26-VII-2017</collectingDate>
</date>
,
<geoCoordinate id="EE3BB1B5FF86FF8EAA13FDD2FF31FDDF" degrees="16.6637" direction="north" orientation="latitude" pageId="2" pageNumber="688" precision="5" value="16.6637">16.6637° N</geoCoordinate>
, -
<geoCoordinate id="EE3BB1B5FF86FF8EA8F7FDF9FEA5FDDF" box="[194,318,567,588]" degrees="94.2335" direction="west" orientation="longitude" pageId="2" pageNumber="688" precision="5" value="-94.2335">94.2335° W</geoCoordinate>
,
<quantity id="4CF77A97FF86FF8EA97BFDF9FE09FDDF" box="[334,402,567,588]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="688" unit="m" value="1800.0">
<elevation id="00223041FF86FF8EA97BFDF9FE09FDDF" box="[334,402,567,588]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="688" unit="m" value="1800.0">1800m</elevation>
</quantity>
, en suelo (muerto),
<collectorName id="26FAB2A4FF86FF8EAA5CFDF9FEB8FDF5" pageId="2" pageNumber="688">E. Arriaga-Varela</collectorName>
col”. The
<typeStatus id="54B469D0FF86FF8EA9A7FD9FFE7EFDF5" box="[402,485,593,614]" pageId="2" pageNumber="688" type="holotype">holotype</typeStatus>
is deposited in
<collectionCode id="ED1E4FB7FF86FF8EA8A2FDA2FF49FD12" box="[151,210,620,641]" pageId="2" pageNumber="688">IEXA</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B67DD2FFF86FF8EA8EBFDA2FDCDFD12" box="[222,598,620,641]" collectionCode="CNIN" country="Mexico" pageId="2" pageNumber="688" specimenCount="1" stateProvince="Oaxaca" typeStatus="paratype">
and the
<typeStatus id="54B469D0FF86FF8EA919FDA2FEE6FD12" box="[300,381,620,641]" pageId="2" pageNumber="688" type="paratype">paratype</typeStatus>
is deposited in
<collectionCode id="ED1E4FB7FF86FF8EAA2CFDA2FDCDFD12" box="[537,598,620,641]" country="Mexico" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="2" pageNumber="688">CNIN</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31584F9FF86FF8EA89AFD48FB91FF62" pageId="2" pageNumber="688" type="description">
<paragraph id="8BB0D772FF86FF8EA89AFD48FE98F9AE" blockId="2.[151,639,380,1730]" pageId="2" pageNumber="688">
<emphasis id="B97B0B60FF86FF8EA89AFD48FEB1FD08" bold="true" box="[175,298,646,667]" pageId="2" pageNumber="688">Description.</emphasis>
<typeStatus id="54B469D0FF86FF8EA904FD48FE12FD08" box="[305,393,646,667]" pageId="2" pageNumber="688" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="1334CBF7FF86FF8EA9F9FD48FDB3FD08" box="[460,552,646,668]" captionStart="Figs" captionStartId="3.[197,236,1611,1630]" captionTargetBox="[177,1188,197,1593]" captionTargetId="figure-1@3.[175,1190,194,1595]" captionTargetPageId="3" captionText="Figs. 17. Verodes species. 1) V. chimalapensis, dorsal habitus; 2) V. chimalapensis, lateral habitus; 3) V. chimalapensis, dorsal surface of pronotum; 4) V. asperatus oblique habitus; 5) V. scabrosus, oblique habitus; 6) V. scabrosus, apical two-fifths of left elytron; 7) V. chimalapensis, apical two-fifths of left elytron." figureDoi="http://doi.org/10.5281/zenodo.5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="2" pageNumber="688">Figs. 12</figureCitation>
). Length
<quantity id="4CF77A97FF86FF8EA8A2FD6FFF6BFD25" box="[151,240,673,694]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="2" pageNumber="688" unit="mm" value="16.5">16.5 mm</quantity>
; width
<quantity id="4CF77A97FF86FF8EA902FD6FFE18FD25" box="[311,387,673,694]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="2" pageNumber="688" unit="mm" value="6.6">6.6 mm</quantity>
. Dorsal color dark brown, with irregular and small areas light brown. Dorsal surface of pronotum and elytra with dense setae between tubercles (
<figureCitation id="1334CBF7FF86FF8EA97AFD3EFE12FC96" box="[335,393,752,773]" captionStart="Figs" captionStartId="3.[197,236,1611,1630]" captionTargetBox="[177,1188,197,1593]" captionTargetId="figure-1@3.[175,1190,194,1595]" captionTargetPageId="3" captionText="Figs. 17. Verodes species. 1) V. chimalapensis, dorsal habitus; 2) V. chimalapensis, lateral habitus; 3) V. chimalapensis, dorsal surface of pronotum; 4) V. asperatus oblique habitus; 5) V. scabrosus, oblique habitus; 6) V. scabrosus, apical two-fifths of left elytron; 7) V. chimalapensis, apical two-fifths of left elytron." figureDoi="http://doi.org/10.5281/zenodo.5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="2" pageNumber="688">Fig. 3</figureCitation>
), setae tan or light brown inserted on cuticular surface. Dorsum with tubercles almost uniformly distributed on ridges, nodules, and depressions. Ventral surface tuberculate, setose.
<emphasis id="B97B0B60FF86FF8EA8A2FC95FF4EFCE3" bold="true" box="[151,213,859,880]" pageId="2" pageNumber="688">Head:</emphasis>
Clypeus and frons almost flat, supra-antennal lobes prominent. Antennae 10-segmented with weak, 2-segmented club. Mandibles with membranous prostheca. Apical maxillary palpomere with truncate apex.
<emphasis id="B97B0B60FF86FF8EA91BFC0BFE00FC49" bold="true" box="[302,411,965,986]" pageId="2" pageNumber="688">Pronotum:</emphasis>
Nearly square, lateral margins prominent, without lobes or denticles, and rounded anteriorly, posterior angles strongly recurved in lateral view. Dorsum with 2 median lateral ridges enclosing a central depression, 1 lateral depression on each side, and 1 middle basal ridge. Hhypomeron without antennal cavity.
<emphasis id="B97B0B60FF86FF8EAA27FBAAFDE5FBEA" bold="true" box="[530,638,1124,1145]" pageId="2" pageNumber="688">Scutellum:</emphasis>
Not visible.
<emphasis id="B97B0B60FF86FF8EA922FBB1FEC5FB07" bold="true" box="[279,350,1151,1172]" pageId="2" pageNumber="688">Elytra:</emphasis>
Humeral umbone large and robust. Epipleural ridge absent. Third elytral interval with inconspicuous ridge at base. Fifth elytral interval with 2 disconnected nodules, one on basal third short, not reaching base of elytra; second one (posterior) settled near 6
<superScript id="7C7A7A3AFF86FF8EA9B7FB30FE0BFA9F" attach="left" box="[386,400,1278,1292]" fontSize="6" pageId="2" pageNumber="688">th</superScript>
interval, at level of sternites 12. Apical 2/5 of each elytron with 3 projecting, conical nodules, subapical one very large and robust, one closest to apex smaller than others, outermost moderate in size, and placed on 9
<superScript id="7C7A7A3AFF86FF8EAA05FAA7FDA5FAE4" attach="left" box="[560,574,1385,1399]" fontSize="6" pageId="2" pageNumber="688">th</superScript>
elytral interval (
<figureCitation id="1334CBF7FF86FF8EA8DAFA47FEB6FA0D" box="[239,301,1417,1438]" captionStart="Figs" captionStartId="3.[197,236,1611,1630]" captionTargetBox="[177,1188,197,1593]" captionTargetId="figure-1@3.[175,1190,194,1595]" captionTargetPageId="3" captionText="Figs. 17. Verodes species. 1) V. chimalapensis, dorsal habitus; 2) V. chimalapensis, lateral habitus; 3) V. chimalapensis, dorsal surface of pronotum; 4) V. asperatus oblique habitus; 5) V. scabrosus, oblique habitus; 6) V. scabrosus, apical two-fifths of left elytron; 7) V. chimalapensis, apical two-fifths of left elytron." figureDoi="http://doi.org/10.5281/zenodo.5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="2" pageNumber="688">Fig. 7</figureCitation>
). Apical margin weakly serrate at sides, with apex shallowly emarginated, rounded.
<emphasis id="B97B0B60FF86FF8EA8A2FA70FF79FA40" bold="true" box="[151,226,1470,1491]" pageId="2" pageNumber="688">Venter:</emphasis>
Prosternum flattened, not grooved. Metepisternum tuberculate, not foveolate.
<emphasis id="B97B0B60FF86FF8EAA3DFA16FDA5FA7E" bold="true" box="[520,574,1496,1517]" pageId="2" pageNumber="688">Legs:</emphasis>
Meso and metafemora with shiny nodules on lower bor- der of base. Last tarsomere on all legs without empodium.
</paragraph>
<paragraph id="8BB0D772FF86FF8EA89AF98DFB91FF62" blockId="2.[151,639,380,1730]" lastBlockId="2.[685,1172,194,961]" pageId="2" pageNumber="688">
<emphasis id="B97B0B60FF86FF8EA89AF98DFE8AF9CB" bold="true" box="[175,273,1603,1624]" pageId="2" pageNumber="688">
<typeStatus id="54B469D0FF86FF8EA89AF98DFE96F9CB" box="[175,269,1603,1624]" pageId="2" pageNumber="688" type="paratype">Paratype</typeStatus>
.
</emphasis>
<specimenCount id="9D091CFBFF86FF8EA92DF98DFEA2F9CB" box="[280,313,1602,1624]" pageId="2" pageNumber="688" type="female">1 ♀</specimenCount>
. Length
<quantity id="4CF77A97FF86FF8EA9BAF98DFE73F9CB" box="[399,488,1603,1624]" metricMagnitude="-2" metricUnit="m" metricValue="1.83" pageId="2" pageNumber="688" unit="mm" value="18.3">18.3 mm</quantity>
; width
<quantity id="4CF77A97FF86FF8EAA1BF98DFDE0F9CB" box="[558,635,1603,1624]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="688" unit="mm" value="7.5">7.5 mm</quantity>
. Specimen is damaged, and all legs are missing. It differs from the
<typeStatus id="54B469D0FF86FF8EA975F9B6FE08F91E" box="[320,403,1656,1677]" pageId="2" pageNumber="688" type="holotype">holotype</typeStatus>
by the uniformly light brown color, pronotal tubercles absent from intervening areas, basal ridge on third elytral interval distinct, and basal nodule on fifth elytral interval lower and a little longer posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C31584F9FF86FF8EAAF0FF39FC57FC52" pageId="2" pageNumber="688" type="etymology">
<paragraph id="8BB0D772FF86FF8EAAF0FF39FB95FEC8" blockId="2.[685,1172,194,961]" pageId="2" pageNumber="688">
<emphasis id="B97B0B60FF86FF8EAAF0FF39FCA3FE9F" bold="true" box="[709,824,247,268]" pageId="2" pageNumber="688">Etymology.</emphasis>
The specific epithet is derived from Chimalapas, the region of the state of
<collectingRegion id="49CB1990FF86FF8EAC3BFEDFFBCEFEB5" box="[1038,1109,273,294]" country="Mexico" name="Oaxaca" pageId="2" pageNumber="688">Oaxaca</collectingRegion>
where this species was collected, combined with the Latin suffix
<emphasis id="B97B0B60FF86FF8EAACDFE89FCBCFECF" box="[760,807,327,348]" italics="true" pageId="2" pageNumber="688">ensis</emphasis>
, meaning belonging to.
</paragraph>
<paragraph id="8BB0D772FF86FF8EAAF0FEAFFCDEFC1A" blockId="2.[685,1172,194,961]" pageId="2" pageNumber="688">
<emphasis id="B97B0B60FF86FF8EAAF0FEAFFC06FEE5" bold="true" box="[709,925,353,374]" pageId="2" pageNumber="688">Taxonomic Remarks.</emphasis>
<taxonomicName id="4C0FACF1FF86FF8EAB91FEAEFBE5FEE5" authorityName="Delgado and Mora-Aguilar" authorityYear="2019" box="[932,1150,352,374]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="688" phylum="Arthropoda" rank="species" species="chimalapensis">
<emphasis id="B97B0B60FF86FF8EAB91FEAEFBE5FEE5" box="[932,1150,352,374]" italics="true" pageId="2" pageNumber="688">Verodes chimalapensis</emphasis>
</taxonomicName>
is distinguished from the other species of the genus by the following combination of characters: dorsal surface of pronotum with dense setae among tubercles (
<figureCitation id="1334CBF7FF86FF8EAAC9FE05FCAFFE73" box="[764,820,459,480]" captionStart="Figs" captionStartId="3.[197,236,1611,1630]" captionTargetBox="[177,1188,197,1593]" captionTargetId="figure-1@3.[175,1190,194,1595]" captionTargetPageId="3" captionText="Figs. 17. Verodes species. 1) V. chimalapensis, dorsal habitus; 2) V. chimalapensis, lateral habitus; 3) V. chimalapensis, dorsal surface of pronotum; 4) V. asperatus oblique habitus; 5) V. scabrosus, oblique habitus; 6) V. scabrosus, apical two-fifths of left elytron; 7) V. chimalapensis, apical two-fifths of left elytron." figureDoi="http://doi.org/10.5281/zenodo.5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="2" pageNumber="688">Fig. 3</figureCitation>
); nodule on the basal third of the fifth elytral interval not reaching the elytral base (
<figureCitation id="1334CBF7FF86FF8EAC7BFE28FB1FFE68" box="[1102,1156,486,507]" captionStart="Figs" captionStartId="3.[197,236,1611,1630]" captionTargetBox="[177,1188,197,1593]" captionTargetId="figure-1@3.[175,1190,194,1595]" captionTargetPageId="3" captionText="Figs. 17. Verodes species. 1) V. chimalapensis, dorsal habitus; 2) V. chimalapensis, lateral habitus; 3) V. chimalapensis, dorsal surface of pronotum; 4) V. asperatus oblique habitus; 5) V. scabrosus, oblique habitus; 6) V. scabrosus, apical two-fifths of left elytron; 7) V. chimalapensis, apical two-fifths of left elytron." figureDoi="http://doi.org/10.5281/zenodo.5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="2" pageNumber="688">Fig. 1</figureCitation>
); fifth elytral interval with disconnected nodules (
<figureCitation id="1334CBF7FF86FF8EAA81FDD5FC89FDA3" box="[692,786,538,560]" captionStart="Figs" captionStartId="3.[197,236,1611,1630]" captionTargetBox="[177,1188,197,1593]" captionTargetId="figure-1@3.[175,1190,194,1595]" captionTargetPageId="3" captionText="Figs. 17. Verodes species. 1) V. chimalapensis, dorsal habitus; 2) V. chimalapensis, lateral habitus; 3) V. chimalapensis, dorsal surface of pronotum; 4) V. asperatus oblique habitus; 5) V. scabrosus, oblique habitus; 6) V. scabrosus, apical two-fifths of left elytron; 7) V. chimalapensis, apical two-fifths of left elytron." figureDoi="http://doi.org/10.5281/zenodo.5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="2" pageNumber="688">Figs. 12</figureCitation>
); apical two-fifths of elytron with three large projecting conical nodules (
<figureCitation id="1334CBF7FF86FF8EABD8FDFBFBB3FDD9" box="[1005,1064,565,586]" captionStart="Figs" captionStartId="3.[197,236,1611,1630]" captionTargetBox="[177,1188,197,1593]" captionTargetId="figure-1@3.[175,1190,194,1595]" captionTargetPageId="3" captionText="Figs. 17. Verodes species. 1) V. chimalapensis, dorsal habitus; 2) V. chimalapensis, lateral habitus; 3) V. chimalapensis, dorsal surface of pronotum; 4) V. asperatus oblique habitus; 5) V. scabrosus, oblique habitus; 6) V. scabrosus, apical two-fifths of left elytron; 7) V. chimalapensis, apical two-fifths of left elytron." figureDoi="http://doi.org/10.5281/zenodo.5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="2" pageNumber="688">Fig. 7</figureCitation>
); and sides of apical margin of elytra weakly serrate (
<figureCitation id="1334CBF7FF86FF8EAC7EFD9EFB1EFDF6" box="[1099,1157,592,613]" captionStart="Figs" captionStartId="3.[197,236,1611,1630]" captionTargetBox="[177,1188,197,1593]" captionTargetId="figure-1@3.[175,1190,194,1595]" captionTargetPageId="3" captionText="Figs. 17. Verodes species. 1) V. chimalapensis, dorsal habitus; 2) V. chimalapensis, lateral habitus; 3) V. chimalapensis, dorsal surface of pronotum; 4) V. asperatus oblique habitus; 5) V. scabrosus, oblique habitus; 6) V. scabrosus, apical two-fifths of left elytron; 7) V. chimalapensis, apical two-fifths of left elytron." figureDoi="http://doi.org/10.5281/zenodo.5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="2" pageNumber="688">Fig. 1</figureCitation>
). The new species seems to be most similar to
<taxonomicName id="4C0FACF1FF86FF8EAC7CFDA4FC14FD09" authority="(Champion)" baseAuthorityName="Champion" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="688" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="B97B0B60FF86FF8EAC7CFDA4FC92FD09" italics="true" pageId="2" pageNumber="688">Verodes asperatus</emphasis>
(Champion)
</taxonomicName>
and
<taxonomicName id="4C0FACF1FF86FF8EABEDFD4BFD63FD26" authority="(Solier)" baseAuthorityName="Solier" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="688" phylum="Arthropoda" rank="species" species="scabrosus">
<emphasis id="B97B0B60FF86FF8EABEDFD4BFB08FD08" box="[984,1171,645,667]" italics="true" pageId="2" pageNumber="688">Verodes scabrosus</emphasis>
(Solier)
</taxonomicName>
. From the former, it is distinguished by the presence of a lateral subapical nodule on the ninth elytral interval (on the seventh elytral interval in
<taxonomicName id="4C0FACF1FF86FF8EAA98FD3EFCBAFC96" box="[685,801,752,773]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="688" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="B97B0B60FF86FF8EAA98FD3EFCBAFC96" box="[685,801,752,773]" italics="true" pageId="2" pageNumber="688">V. asperatus</emphasis>
</taxonomicName>
) and the weakly serrate sides (strongly serrate in
<taxonomicName id="4C0FACF1FF86FF8EAB3FFCC4FCE6FC8C" box="[778,893,778,799]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="688" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="B97B0B60FF86FF8EAB3FFCC4FCE6FC8C" box="[778,893,778,799]" italics="true" pageId="2" pageNumber="688">V. asperatus</emphasis>
</taxonomicName>
). From the latter species, it is distinguished by the presence of three subapical, conical nodules on the apical two-fifths each elytron, as opposed to four subapical, rounded nodules in
<taxonomicName id="4C0FACF1FF86FF8EAAF2FCBAFCA4FC19" box="[711,831,884,906]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="688" phylum="Arthropoda" rank="species" species="scabrosus">
<emphasis id="B97B0B60FF86FF8EAAF2FCBAFCA4FC19" box="[711,831,884,906]" italics="true" pageId="2" pageNumber="688">V. scabrosus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB0D772FF86FF8EAAF0FC5FFC57FC52" blockId="2.[685,1172,194,961]" pageId="2" pageNumber="688">We propose the following modifications to the key in Foley and Ivie (2008):</paragraph>
</subSubSection>
<subSubSection id="C31584F9FF86FF8EAA9BFC16FB17FB41" pageId="2" pageNumber="688" type="key">
<keyStep id="30FBCFD7FF86FF8EAA9BFC16FB0FFB34" pageId="2" pageNumber="688">
<key id="FF9E8043FF86FF8EAA9BFC16FB0FFB34" pageId="2" pageNumber="688">
<paragraph id="8BB0D772FF86FF8EAA9BFC16FB0FFBB1" blockId="2.[686,1172,984,1191]" pageId="2" pageNumber="688">
<keyLead id="30FE7447FF86FF8EAA9BFC16FB0FFBB1" pageId="2" pageNumber="688">1. Dorsal surface of pronotum with all setae singly inserted at the base of tubercles, tubercles on pronotum uniformly distributed ................. 2</keyLead>
</paragraph>
<paragraph id="8BB0D772FF86FF8EAA85FBE9FB0FFB34" blockId="2.[686,1172,984,1191]" pageId="2" pageNumber="688">
<keyLead id="30FE7447FF86FF8EAA85FBE9FB0FFB34" pageId="2" pageNumber="688">- Dorsal surface of pronotum with dense setae between tubercles inserted directly on cuticular surface, tubercles on pronotum usually denser on ridges, and usually absent in intervening areas .................................................... 5</keyLead>
</paragraph>
</key>
</keyStep>
<paragraph id="8BB0D772FF86FF8EAA9BFB73FB17FB41" blockId="2.[686,1164,1213,1234]" box="[686,1164,1213,1234]" pageId="2" pageNumber="688">And replace couplets 6 and 7 with the following:</paragraph>
</subSubSection>
<subSubSection id="C31584F9FF86FF88AA81FB27FDE5FEE5" lastPageId="4" lastPageNumber="690" pageId="2" pageNumber="688" type="description">
<paragraph id="8BB0D772FF86FF8EAA81FB27FB0FFA10" blockId="2.[692,1173,1257,1730]" pageId="2" pageNumber="688">
6. Fifth elytral interval with arcuate ridge from level just behind mesocoxal cavities to the suture between ventrites 2 and 3; apical third of elytron with 3 moderate size nodules subequal in size, nodule near apex slightly smaller than subapical nodule ...........
<taxonomicName id="4C0FACF1FF86FF8EABE1FAA0FB0FFA10" authority="(Say)" baseAuthorityName="Say" baseAuthorityYear="1835" box="[980,1172,1390,1411]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="688" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B97B0B60FF86FF8EABE1FAA0FB0FFA10" bold="true" box="[980,1172,1390,1411]" pageId="2" pageNumber="688">
<emphasis id="B97B0B60FF86FF8EABE1FAA0FBCCFA10" bold="true" box="[980,1111,1390,1411]" italics="true" pageId="2" pageNumber="688">V. inaequalis</emphasis>
(Say)
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BB0D772FF86FF8EAA83FA47FB0FF9B1" blockId="2.[692,1173,1257,1730]" pageId="2" pageNumber="688">- Fifth elytral interval with disconnected nodules from level of midpoint of metasternum to suture between ventrites 1 and 2 or 2 and 3; apical third of elytron with 2 or 3 large, projecting, conical nodules, lateral subapical nodule very large, much larger than nodule near apex ............ 7</paragraph>
<paragraph id="8BB0D772FF86FF8EAA81F9E6FB08F91E" blockId="2.[692,1173,1257,1730]" pageId="2" pageNumber="688">
7. Apical 2/5 of elytron with a lateral nodule on 7
<superScript id="7C7A7A3AFF86FF8EACB3F9EDFB0FF9A2" attach="none" box="[1158,1172,1571,1585]" fontSize="6" pageId="2" pageNumber="688">th</superScript>
interval, but without nodule on 9
<superScript id="7C7A7A3AFF86FF8EAC33F9F3FB8FF9D8" attach="left" box="[1030,1044,1597,1611]" fontSize="6" pageId="2" pageNumber="688">th</superScript>
interval (
<figureCitation id="1334CBF7FF86FF8EAC58F98DFD7AF9E1" captionStart="Figs" captionStartId="3.[197,236,1611,1630]" captionTargetBox="[177,1188,197,1593]" captionTargetId="figure-1@3.[175,1190,194,1595]" captionTargetPageId="3" captionText="Figs. 17. Verodes species. 1) V. chimalapensis, dorsal habitus; 2) V. chimalapensis, lateral habitus; 3) V. chimalapensis, dorsal surface of pronotum; 4) V. asperatus oblique habitus; 5) V. scabrosus, oblique habitus; 6) V. scabrosus, apical two-fifths of left elytron; 7) V. chimalapensis, apical two-fifths of left elytron." figureDoi="http://doi.org/10.5281/zenodo.5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="2" pageNumber="688">Fig. 4</figureCitation>
). Sides of apical margin of elytra strongly serrate ................
<taxonomicName id="4C0FACF1FF86FF8EABA3F9B9FB08F91E" authority="(Champion)" baseAuthorityName="Champion" box="[918,1171,1655,1677]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="688" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="B97B0B60FF86FF8EABA3F9B9FB08F91E" bold="true" box="[918,1171,1655,1677]" pageId="2" pageNumber="688">
<emphasis id="B97B0B60FF86FF8EABA3F9B9FB89F91F" bold="true" box="[918,1042,1655,1676]" italics="true" pageId="2" pageNumber="688">V. asperatus</emphasis>
(Champion)
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BB0D772FF86FF8EAA83F95CFB08F925" blockId="2.[692,1173,1257,1730]" pageId="2" pageNumber="688">
- Apical 2/5 of elytron without a lateral nodule on 7
<superScript id="7C7A7A3AFF86FF8EAACBF966FC97F925" attach="left" box="[766,780,1704,1718]" fontSize="6" pageId="2" pageNumber="688">th</superScript>
interval, but with a lateral nodule on 9
<superScript id="7C7A7A3AFF86FF8EACB0F966FB08F925" attach="none" box="[1157,1171,1704,1718]" fontSize="6" pageId="2" pageNumber="688">th</superScript>
</paragraph>
<caption id="DF7087FAFF87FF8FA8F0F985FC78F91D" ID-DOI="http://doi.org/10.5281/zenodo.5402500" ID-Zenodo-Dep="5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="3" pageNumber="689" startId="3.[197,236,1611,1630]" targetBox="[177,1188,197,1593]" targetPageId="3">
<paragraph id="8BB0D772FF87FF8FA8F0F985FC78F91D" blockId="3.[173,1194,1610,1678]" pageId="3" pageNumber="689">
<emphasis id="B97B0B60FF87FF8FA8F0F985FEBAF9CD" bold="true" box="[197,289,1611,1630]" pageId="3" pageNumber="689">Figs. 17.</emphasis>
<taxonomicName id="4C0FACF1FF87FF8FA903F984FE46F9CD" box="[310,477,1610,1630]" class="Insecta" family="Zopheridae" genus="Verodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="689" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="B97B0B60FF87FF8FA903F984FEE3F9CE" box="[310,376,1610,1629]" italics="true" pageId="3" pageNumber="689">Verodes</emphasis>
species.
<emphasis id="B97B0B60FF87FF8FA9E6F985FE46F9CD" bold="true" box="[467,477,1611,1630]" pageId="3" pageNumber="689">1</emphasis>
</taxonomicName>
<emphasis id="B97B0B60FF87FF8FA9EBF985FE7EF9CD" bold="true" box="[478,485,1611,1630]" pageId="3" pageNumber="689">)</emphasis>
<taxonomicName id="4C0FACF1FF87FF8FA9C7F985FD1DF9CE" authorityName="Delgado and Mora-Aguilar" authorityYear="2019" box="[498,646,1610,1629]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="689" phylum="Arthropoda" rank="species" species="chimalapensis">
<emphasis id="B97B0B60FF87FF8FA9C7F985FD1DF9CE" box="[498,646,1610,1629]" italics="true" pageId="3" pageNumber="689">V. chimalapensis</emphasis>
</taxonomicName>
, dorsal habitus;
<emphasis id="B97B0B60FF87FF8FAB12F985FCA3F9CD" bold="true" box="[807,824,1611,1630]" pageId="3" pageNumber="689">2)</emphasis>
<taxonomicName id="4C0FACF1FF87FF8FAB70F985FC42F9CE" authorityName="Delgado and Mora-Aguilar" authorityYear="2019" box="[837,985,1610,1629]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="689" phylum="Arthropoda" rank="species" species="chimalapensis">
<emphasis id="B97B0B60FF87FF8FAB70F985FC42F9CE" box="[837,985,1610,1629]" italics="true" pageId="3" pageNumber="689">V. chimalapensis</emphasis>
</taxonomicName>
, lateral habitus;
<emphasis id="B97B0B60FF87FF8FAC4EF985FB17F9CD" bold="true" box="[1147,1164,1611,1630]" pageId="3" pageNumber="689">3)</emphasis>
<taxonomicName id="4C0FACF1FF87FF8FACACF985FEBEF9E6" authorityName="Delgado and Mora-Aguilar" authorityYear="2019" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="689" phylum="Arthropoda" rank="species" species="chimalapensis">
<emphasis id="B97B0B60FF87FF8FACACF985FEBEF9E6" italics="true" pageId="3" pageNumber="689">V. chimalapensis</emphasis>
</taxonomicName>
, dorsal surface of pronotum;
<emphasis id="B97B0B60FF87FF8FAA04F9ADFDD8F9E5" bold="true" box="[561,579,1635,1654]" pageId="3" pageNumber="689">4)</emphasis>
<taxonomicName id="4C0FACF1FF87FF8FAA79F9ADFD2CF9E6" box="[588,695,1635,1653]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="689" phylum="Arthropoda" rank="species" species="asperatus">
<emphasis id="B97B0B60FF87FF8FAA79F9ADFD2CF9E6" box="[588,695,1635,1653]" italics="true" pageId="3" pageNumber="689">V. asperatus</emphasis>
</taxonomicName>
oblique habitus;
<emphasis id="B97B0B60FF87FF8FAB60F9ADFCFDF9E5" bold="true" box="[853,870,1635,1654]" pageId="3" pageNumber="689">5)</emphasis>
<taxonomicName id="4C0FACF1FF87FF8FAB5AF9ADFC47F9E6" box="[879,988,1634,1653]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="689" phylum="Arthropoda" rank="species" species="scabrosus">
<emphasis id="B97B0B60FF87FF8FAB5AF9ADFC47F9E6" box="[879,988,1634,1653]" italics="true" pageId="3" pageNumber="689">V. scabrosus</emphasis>
</taxonomicName>
, oblique habitus;
<emphasis id="B97B0B60FF87FF8FAC4AF9ADFB0BF9E5" bold="true" box="[1151,1168,1635,1654]" pageId="3" pageNumber="689">6)</emphasis>
<emphasis id="B97B0B60FF87FF8FACACF9ADFE9CF91E" italics="true" pageId="3" pageNumber="689">
<taxonomicName id="4C0FACF1FF87FF8FACACF9ADFE98F91E" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="689" phylum="Arthropoda" rank="species" species="scabrosus">V. scabrosus</taxonomicName>
,
</emphasis>
apical two-fifths of left elytron;
<emphasis id="B97B0B60FF87FF8FAA14F9B5FDA9F91D" bold="true" box="[545,562,1659,1678]" pageId="3" pageNumber="689">7)</emphasis>
<taxonomicName id="4C0FACF1FF87FF8FAA0CF9B5FD5CF91E" authorityName="Delgado and Mora-Aguilar" authorityYear="2019" box="[569,711,1658,1677]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="689" phylum="Arthropoda" rank="species" species="chimalapensis">
<emphasis id="B97B0B60FF87FF8FAA0CF9B5FD5CF91E" box="[569,711,1658,1677]" italics="true" pageId="3" pageNumber="689">V. chimalapensis</emphasis>
</taxonomicName>
, apical two-fifths of left elytron.
</paragraph>
</caption>
<paragraph id="8BB0D772FF80FF88A888FF0CFDE5FF44" blockId="4.[152,638,194,374]" box="[189,638,194,215]" pageId="4" pageNumber="690">
interval (
<figureCitation id="1334CBF7FF80FF88A92BFF0CFEF8FF44" box="[286,355,194,215]" captionStart="Figs" captionStartId="3.[197,236,1611,1630]" captionTargetBox="[177,1188,197,1593]" captionTargetId="figure-1@3.[175,1190,194,1595]" captionTargetPageId="3" captionText="Figs. 17. Verodes species. 1) V. chimalapensis, dorsal habitus; 2) V. chimalapensis, lateral habitus; 3) V. chimalapensis, dorsal surface of pronotum; 4) V. asperatus oblique habitus; 5) V. scabrosus, oblique habitus; 6) V. scabrosus, apical two-fifths of left elytron; 7) V. chimalapensis, apical two-fifths of left elytron." figureDoi="http://doi.org/10.5281/zenodo.5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="4" pageNumber="690">Fig. 5</figureCitation>
). Apical margin of elytra
</paragraph>
<paragraph id="8BB0D772FF80FF88A888FF12FDE6FE9F" blockId="4.[152,638,194,374]" pageId="4" pageNumber="690">
weakly serrate .................................... 7
<emphasis id="B97B0B60FF80FF88AA4DFF17FDE6FF75" bold="true" box="[632,637,217,230]" italics="true" pageId="4" pageNumber="690">
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</emphasis>
7
<emphasis id="B97B0B60FF80FF88A891FF3AFF32FE92" bold="true" box="[164,169,244,257]" italics="true" pageId="4" pageNumber="690">
<superScript id="7C7A7A3AFF80FF88A891FF3AFF32FE92" attach="none" box="[164,169,244,257]" fontSize="6" pageId="4" pageNumber="690"></superScript>
</emphasis>
. Apical 2/5 of elytron with 4 tubercles (
<figureCitation id="1334CBF7FF80FF88AA14FF39FDC3FE9F" box="[545,600,247,268]" captionStart="Figs" captionStartId="3.[197,236,1611,1630]" captionTargetBox="[177,1188,197,1593]" captionTargetId="figure-1@3.[175,1190,194,1595]" captionTargetPageId="3" captionText="Figs. 17. Verodes species. 1) V. chimalapensis, dorsal habitus; 2) V. chimalapensis, lateral habitus; 3) V. chimalapensis, dorsal surface of pronotum; 4) V. asperatus oblique habitus; 5) V. scabrosus, oblique habitus; 6) V. scabrosus, apical two-fifths of left elytron; 7) V. chimalapensis, apical two-fifths of left elytron." figureDoi="http://doi.org/10.5281/zenodo.5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="4" pageNumber="690">Fig. 6</figureCitation>
)...
</paragraph>
<paragraph id="8BB0D772FF80FF88A888FEDFFDE5FED2" blockId="4.[152,638,194,374]" pageId="4" pageNumber="690">
.....................................
<taxonomicName id="4C0FACF1FF80FF88A98FFEDFFDE6FEB5" authority="(Solier)" baseAuthorityName="Solier" box="[442,637,273,294]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="690" phylum="Arthropoda" rank="species" species="scabrosus">
<emphasis id="B97B0B60FF80FF88A98FFEDFFDE6FEB5" bold="true" box="[442,637,273,294]" pageId="4" pageNumber="690">
<emphasis id="B97B0B60FF80FF88A98FFEDFFDAAFEB5" bold="true" box="[442,561,273,294]" italics="true" pageId="4" pageNumber="690">V. scabrosus</emphasis>
(Solier)
</emphasis>
</taxonomicName>
- Apical 2/5 of elytron with 3 tubercles
</paragraph>
<paragraph id="8BB0D772FF80FF88A888FE88FDE5FEC8" blockId="4.[152,638,194,374]" box="[189,638,326,347]" pageId="4" pageNumber="690">
(
<figureCitation id="1334CBF7FF80FF88A8F1FE88FE99FEC8" box="[196,258,326,347]" captionStart="Figs" captionStartId="3.[197,236,1611,1630]" captionTargetBox="[177,1188,197,1593]" captionTargetId="figure-1@3.[175,1190,194,1595]" captionTargetPageId="3" captionText="Figs. 17. Verodes species. 1) V. chimalapensis, dorsal habitus; 2) V. chimalapensis, lateral habitus; 3) V. chimalapensis, dorsal surface of pronotum; 4) V. asperatus oblique habitus; 5) V. scabrosus, oblique habitus; 6) V. scabrosus, apical two-fifths of left elytron; 7) V. chimalapensis, apical two-fifths of left elytron." figureDoi="http://doi.org/10.5281/zenodo.5402500" httpUri="https://zenodo.org/record/5402500/files/figure.png" pageId="4" pageNumber="690">Fig. 7</figureCitation>
) ..............
<taxonomicName id="4C0FACF1FF80FF88A9BCFE88FDE5FEC8" authority="Delgado" authorityName="Delgado and Mora-Aguilar" authorityYear="2019" box="[393,638,326,347]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="690" phylum="Arthropoda" rank="species" species="chimalapensis">
<emphasis id="B97B0B60FF80FF88A9BCFE88FDE5FEC8" bold="true" box="[393,638,326,347]" pageId="4" pageNumber="690">
<emphasis id="B97B0B60FF80FF88A9BCFE88FDB3FEC8" bold="true" box="[393,552,326,347]" italics="true" pageId="4" pageNumber="690">V. chimalapensis</emphasis>
Delgado
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BB0D772FF80FF88A994FEAFFDE5FEE5" blockId="4.[152,638,194,374]" box="[417,638,353,374]" pageId="4" pageNumber="690">
<emphasis id="B97B0B60FF80FF88A994FEAFFDE5FEE5" bold="true" box="[417,638,353,374]" pageId="4" pageNumber="690">
and Mora,
<taxonomicNameLabel id="A248B61BFF80FF88AA3AFEAFFDE5FEE5" box="[527,638,353,374]" pageId="4" pageNumber="690" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31584F9FF80FF88A89AFE43FE06FC13" pageId="4" pageNumber="690" type="distribution">
<paragraph id="8BB0D772FF80FF88A89AFE43FF6DFD55" blockId="4.[151,638,396,976]" pageId="4" pageNumber="690">
<emphasis id="B97B0B60FF80FF88A89AFE43FEABFE31" bold="true" box="[175,304,397,418]" pageId="4" pageNumber="690">Distribution.</emphasis>
<taxonomicName id="4C0FACF1FF80FF88A971FE42FD84FE31" authorityName="Delgado and Mora-Aguilar" authorityYear="2019" box="[324,543,396,418]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="690" phylum="Arthropoda" rank="species" species="chimalapensis">
<emphasis id="B97B0B60FF80FF88A971FE42FD84FE31" box="[324,543,396,418]" italics="true" pageId="4" pageNumber="690">Verodes chimalapensis</emphasis>
</taxonomicName>
is only known from the
<typeStatus id="54B469D0FF80FF88A90BFE69FEFEFE2F" box="[318,357,423,444]" pageId="4" pageNumber="690">type</typeStatus>
locality, near the Isthmus of Tehuantepec in the undisturbed forests of the mountainous areas of El Retén in the Chimalapas region, at
<quantity id="4CF77A97FF80FF88A8A2FE39FEBDFD9F" box="[151,294,503,525]" metricMagnitude="3" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="4" pageNumber="690" unit="m" value="1700.0" valueMax="1800.0" valueMin="1600.0">
<geoCoordinate id="EE3BB1B5FF80FF88A8A2FE39FF51FD9F" box="[151,202,503,524]" degrees="1,600" direction="north" orientation="latitude" pageId="4" pageNumber="690" precision="55" value="1.6">1,600</geoCoordinate>
<geoCoordinate id="EE3BB1B5FF80FF88A8FEFE39FE90FD9F" box="[203,267,503,525]" degrees="1,800" direction="west" orientation="longitude" pageId="4" pageNumber="690" precision="55" value="-1.8">1,800</geoCoordinate>
m
</quantity>
altitude. The
<typeStatus id="54B469D0FF80FF88A984FE39FE43FD9F" box="[433,472,503,524]" pageId="4" pageNumber="690">type</typeStatus>
locality and surrounding areas present a rugged topography with ravines and hills with moist cloud forests ranging from about
<quantity id="4CF77A97FF80FF88A93EFD89FEC7FDCF" box="[267,348,583,604]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="4" pageNumber="690" unit="m" value="1450.0">1,450 m</quantity>
to
<quantity id="4CF77A97FF80FF88A94BFD89FE54FDCF" box="[382,463,583,604]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="690" unit="m" value="1800.0">1,800 m</quantity>
in elevation. The cloud forests of the Chimalapas region are isolated from the cloud forests of the Meseta Central de
<collectingRegion id="49CB1990FF80FF88A8A2FD59FF7EFD3F" box="[151,229,663,684]" country="Mexico" name="Chiapas" pageId="4" pageNumber="690">Chiapas</collectingRegion>
and the Sierra Madre de
<collectingRegion id="49CB1990FF80FF88A9E8FD59FDB0FD3F" box="[477,555,663,684]" country="Mexico" name="Chiapas" pageId="4" pageNumber="690">Chiapas</collectingRegion>
by drier lowlands.
</paragraph>
<paragraph id="8BB0D772FF80FF88A89AFD02FE06FC13" blockId="4.[151,638,396,976]" pageId="4" pageNumber="690">
As in the case of
<taxonomicName id="4C0FACF1FF80FF88A97BFD05FEA5FD68" authority="Delgado and Mora-Aguilar" authorityName="Delgado and Mora-Aguilar" class="Insecta" family="Passalidae" genus="Proculus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="690" phylum="Arthropoda" rank="species" species="reyescastilloi">
<emphasis id="B97B0B60FF80FF88A97BFD05FDB1FD72" box="[334,554,715,737]" italics="true" pageId="4" pageNumber="690">Proculus reyescastilloi</emphasis>
Delgado and Mora-Aguilar
</taxonomicName>
(
<taxonomicName id="4C0FACF1FF80FF88A97CFD28FE28FD68" authorityName="Linnaeus" authorityYear="1758" box="[329,435,742,763]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="690" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C0FACF1FF80FF88A98FFD28FD86FD68" authorityName="Leach" authorityYear="1815" box="[442,541,742,763]" class="Insecta" family="Passalidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="690" phylum="Arthropoda" rank="family">Passalidae</taxonomicName>
), a species also endemic to the Chimalapas region (Delgado and Mora-Aguilar 2014),
<taxonomicName id="4C0FACF1FF80FF88A95AFCD5FD89FCA3" authorityName="Delgado and Mora-Aguilar" authorityYear="2019" box="[367,530,794,816]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="690" phylum="Arthropoda" rank="species" species="chimalapensis">
<emphasis id="B97B0B60FF80FF88A95AFCD5FD89FCA3" box="[367,530,794,816]" italics="true" pageId="4" pageNumber="690">V. chimalapensis</emphasis>
</taxonomicName>
shows an isolated distribution and flightlessness, which suggests that this species is restricted to the mid-elevation cloud forests of this region.
</paragraph>
</subSubSection>
<subSubSection id="C31584F9FF80FF88A89AFC48FE2EFBB1" pageId="4" pageNumber="690" type="biology_ecology">
<paragraph id="8BB0D772FF80FF88A89AFC48FEA2FC43" blockId="4.[151,638,396,976]" pageId="4" pageNumber="690">
<emphasis id="B97B0B60FF80FF88A89AFC48FE9AFC08" bold="true" box="[175,257,902,923]" pageId="4" pageNumber="690">Biology.</emphasis>
One specimen of
<taxonomicName id="4C0FACF1FF80FF88A987FC48FDC9FC08" authorityName="Delgado and Mora-Aguilar" authorityYear="2019" box="[434,594,901,923]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="690" phylum="Arthropoda" rank="species" species="chimalapensis">
<emphasis id="B97B0B60FF80FF88A987FC48FDC9FC08" box="[434,594,901,923]" italics="true" pageId="4" pageNumber="690">V. chimalapensis</emphasis>
</taxonomicName>
was found under the bark of a large log, and another was dead on the soil.
</paragraph>
<paragraph id="8BB0D772FF80FF88A88CFC3AFE2EFBB1" blockId="4.[185,604,1012,1060]" pageId="4" pageNumber="690">
<emphasis id="B97B0B60FF80FF88A88CFC3AFDC7FB9B" bold="true" box="[185,604,1012,1034]" pageId="4" pageNumber="690">DISTRIBUTIONAL COMMENTS ON SPECIES OF</emphasis>
<taxonomicName id="4C0FACF1FF80FF88A955FBC1FE2EFBB1" box="[352,437,1039,1060]" class="Insecta" family="Zopheridae" genus="Verodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="690" phylum="Arthropoda" rank="genus">
<emphasis id="B97B0B60FF80FF88A955FBC1FE2EFBB1" bold="true" box="[352,437,1039,1060]" italics="true" pageId="4" pageNumber="690">VERODES</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>