treatments-xml/data/03/A2/C7/03A2C77C4627FF8AC79CB28D1447DB2A.xml
2024-06-21 12:22:17 +02:00

126 lines
16 KiB
XML

<document id="10A8F1E3DAB27C970182832475D62597" ID-DOI="10.11646/zootaxa.5365.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10146969" ID-ZooBank="1DC72714-D0E8-49D8-821D-03C6B2A7AE80" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1700197580075" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2023" docId="03A2C77C4627FF8AC79CB28D1447DB2A" docLanguage="en" docName="zootaxa.5365.1.1.pdf" docOrigin="Zootaxa 5365 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Porrhotegaeidae Colloff 2023, fam. nov." docType="treatment" docVersion="4" lastPageNumber="71" masterDocId="FF9BBF044661FFCCC70BB31F1724DE65" masterDocTitle="The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families" masterLastPageNumber="93" masterPageNumber="1" pageNumber="71" updateTime="1700839435638" updateUser="ExternalLinkService">
<mods:mods id="6925C9DCAD42FE26AA772354929C91F2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="18F85875FF00670D465F9F2FF7209667">
<mods:title id="65D44A7437747CCBA52F66DD69F8B966">The oribatid mite superfamily Eutegaeoidea (Acari, Oribatida), with descriptions of new taxa from Australia and New Caledonia and a re-assessment of genera and families</mods:title>
</mods:titleInfo>
<mods:name id="84EEDF3ABFBEAD0096E08CC440DFEC19" type="personal">
<mods:role id="7F357EE85BB9048D0CEA86E904D6A7A7">
<mods:roleTerm id="0297A824F9E1B16710CEF6E08FA8C00A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3CBFF14D6BEF9966B9D9C146EB5E98D4">Colloff, Matthew J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3112A573F18AD68EB5770E21E79FD28D">text</mods:typeOfResource>
<mods:relatedItem id="B8E099699D84B79746B0E3C02C28F980" type="host">
<mods:titleInfo id="820131BA657D39A9DAA7EDD78AB93926">
<mods:title id="ED5C365D1AE8CEA97138AAFD76259986">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A413FD6FA3C8A97C2209AAD9CF7CD630">
<mods:date id="1743AA5688A8B811F7B0BF8BAD6970BB">2023</mods:date>
<mods:detail id="E3A4BB005499E93733C705255D2F3ADD" type="pubDate">
<mods:number id="3F75E13F53435A73CFFF6A489FFD3687">2023-11-06</mods:number>
</mods:detail>
<mods:detail id="2E6E3261F04E9069B993F539127754A2" type="volume">
<mods:number id="34C10B7D5AB4EA8CD5BD58FABF9A4D5A">5365</mods:number>
</mods:detail>
<mods:detail id="A74DA52D15BB191E56F28E3A1B778702" type="issue">
<mods:number id="849535A2D0AE7AC24C130C360559C313">1</mods:number>
</mods:detail>
<mods:extent id="F71DAC3F3599ED1E9BE6EDD5FC5412C3" unit="page">
<mods:start id="3F3F86365E8BCA55D06AC78D6457673D">1</mods:start>
<mods:end id="C7A1F7D5483783532EEB32F19538EB7C">93</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="94C8AA74E7F185359AED1CF07054383B">
<mods:url id="158D9B368256DE4DAB41B41664C55C97">https://www.mapress.com/zt/article/download/zootaxa.5365.1.1/52220</mods:url>
</mods:location>
<mods:classification id="975E3B137F2C657D29E76C681D3AA84D">journal article</mods:classification>
<mods:identifier id="4695CCDA3E3BFCF5F093EEA58DFB1EA6" type="DOI">10.11646/zootaxa.5365.1.1</mods:identifier>
<mods:identifier id="B05222166037E78FB4ECF108BCA27B85" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="309B441B8DF50E54E0EBE7CD6695BE2A" type="Zenodo-Dep">10146969</mods:identifier>
<mods:identifier id="94B24A95078081D904DC34249E483F97" type="ZooBank">1DC72714-D0E8-49D8-821D-03C6B2A7AE80</mods:identifier>
</mods:mods>
<treatment id="03A2C77C4627FF8AC79CB28D1447DB2A" LSID="urn:lsid:plazi:treatment:03A2C77C4627FF8AC79CB28D1447DB2A" httpUri="http://treatment.plazi.org/id/03A2C77C4627FF8AC79CB28D1447DB2A" lastPageNumber="71" pageId="70" pageNumber="71">
<subSubSection id="C31125E14627FF8AC79CB28D16C4DFC8" box="[151,480,402,429]" pageId="70" pageNumber="71" type="nomenclature">
<paragraph id="8BB4766A4627FF8AC79CB28D16C4DFC8" blockId="70.[151,480,402,429]" box="[151,480,402,429]" pageId="70" pageNumber="71">
<heading id="D0FCC1064627FF8AC79CB28D16C4DFC8" bold="true" box="[151,480,402,429]" fontSize="11" level="2" pageId="70" pageNumber="71" reason="3">
<emphasis id="B97FAA784627FF8AC79CB28D16C4DFC8" bold="true" box="[151,480,402,429]" pageId="70" pageNumber="71">
<taxonomicName id="4C0B0DE94627FF8AC79CB28D1642DFC9" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[151,358,402,428]" family="Porrhotegaeidae" kingdom="Animalia" order="Oribatida" pageId="70" pageNumber="71" rank="family" status="fam. nov.">Porrhotegaeidae</taxonomicName>
<taxonomicNameLabel id="A24C17034627FF8AC666B28D16C4DFC8" box="[365,480,402,429]" pageId="70" pageNumber="71" rank="family">fam. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31125E14627FF8AC79CB2C71403DF8A" box="[151,807,471,496]" pageId="70" pageNumber="71" type="materials_examined">
<paragraph id="8BB4766A4627FF8AC79CB2C71403DF8A" blockId="70.[151,807,471,496]" box="[151,807,471,496]" pageId="70" pageNumber="71">
<typeStatus id="54B0C8C84627FF8AC79CB2C717EDDF95" box="[151,201,472,496]" pageId="70" pageNumber="71">Type</typeStatus>
genus:
<taxonomicName id="4C0B0DE94627FF8AC613B2C81407DF8A" authority="J. Balogh &amp; Mahunka, 1966, p. 557" authorityName="J. Balogh &amp; Mahunka" authorityPageNumber="557" authorityYear="1966" box="[280,803,471,495]" class="Arachnida" family="Eutegaeidae" genus="Porrhotegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B97FAA784627FF8AC613B2C8168ADF8A" box="[280,430,471,495]" italics="true" pageId="70" pageNumber="71">Porrhotegaeus</emphasis>
J.
<bibRefCitation id="EF9A0B9B4627FF8AC6C0B2C815F2DF8A" author="Balogh, J. &amp; Mahunka, S." box="[459,726,471,495]" pageId="70" pageNumber="71" pagination="553 - 568" refId="ref64344" refString="Balogh, J. &amp; Mahunka, S. (1966) New oribatids from Australian soils. Folia Entomologica Hungaricae, 19, 553 - 568." type="journal article" year="1966">Balogh &amp; Mahunka, 1966</bibRefCitation>
, p. 557
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31125E14627FF8AC79CB1021222DDA6" pageId="70" pageNumber="71" type="diagnosis">
<paragraph id="8BB4766A4627FF8AC79CB1021222DDA6" blockId="70.[151,1437,541,1359]" pageId="70" pageNumber="71">
<emphasis id="B97FAA784627FF8AC79CB1021636DC52" bold="true" box="[151,274,541,567]" pageId="70" pageNumber="71">Diagnosis.</emphasis>
Lamellae broad, well developed, ovoid, with free cusps bearing one or two teeth, lacking translamella. Bothridia free, not fused with lamellae. With long blade-like humeral processes extending from lateral margin of notogaster at widest part to point mid-way along lamellae, lateral margin reflexed ventrally; humeral processes lacking lobes posterior of the main anterior structure. Tutorium broad, with a well-developed, curved point. Prodorsum waisted posterior of bothridium. Notogaster U-shaped, surface flat or concave, never convex; margins elevated above level of central notogaster; with eight or nine pairs of long notogastral setae positioned marginally and pointed dorsally. Pedotectum I sub-rectangular; pedotectum II sub-rectangular or lobe-like; discidium pointed. Anterior margin of pedotectum I and epimere I forming straight or curved transverse tectum covering posterior mentum. Circumpedal carina and enantiophyses E4 present; with or without perigenital carinae. Six pairs of genital setae aligned longitudinally, three pairs of adanal setae. Anal plates lozenge-shaped. Pre-anal organ T-shaped. Nymphs with well-developed free lamellar cusps and lateral notogastral setae emerging from long, fine apophyses fused with gastronotum; lateral setae flanked by elongated, flat scales with strongly serrated margins.
</paragraph>
</subSubSection>
<subSubSection id="C31125E14627FF8AC7CCB0D21447DB2A" pageId="70" pageNumber="71" type="discussion">
<paragraph id="8BB4766A4627FF8AC7CCB0D21447DB2A" blockId="70.[151,1437,541,1359]" pageId="70" pageNumber="71">
<emphasis id="B97FAA784627FF8AC7CCB0D2161FDD82" bold="true" box="[199,315,973,999]" pageId="70" pageNumber="71">Remarks.</emphasis>
<taxonomicName id="4C0B0DE94627FF8AC64DB0D216D8DD82" authority="Colloff, 2023" authorityName="Colloff" authorityYear="2023" box="[326,508,973,999]" family="Porrhotegaeidae" pageId="70" pageNumber="83" rank="family" status="fam. nov.">Porrhotegaeidae</taxonomicName>
is established for
<taxonomicName id="4C0B0DE94627FF8AC5D3B0D2145ADD82" authorityName="J. Balogh &amp; Mahunka" authorityYear="1966" box="[728,894,973,999]" class="Arachnida" family="Eutegaeidae" genus="Porrhotegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B97FAA784627FF8AC5D3B0D2145ADD82" box="[728,894,973,999]" italics="true" pageId="70" pageNumber="71">Porrhotegaeus</emphasis>
</taxonomicName>
because the genus shares few character states with
<taxonomicName id="4C0B0DE94627FF8AC7D9B0EE166ADA6E" authorityName="J. Balogh" authorityYear="1970" box="[210,334,1009,1035]" class="Arachnida" family="Eutegaeidae" genus="Dudichella" kingdom="Animalia" order="Sarcoptiformes" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B97FAA784627FF8AC7D9B0EE166ADA6E" box="[210,334,1009,1035]" italics="true" pageId="70" pageNumber="71">Dudichella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B0DE94627FF8AC683B0ED16D8DA6E" authorityName=", Hammer" authorityYear="1966" box="[392,508,1010,1035]" class="Arachnida" family="Eutegaeidae" genus="Pterozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B97FAA784627FF8AC683B0ED16D8DA6E" box="[392,508,1010,1035]" italics="true" pageId="70" pageNumber="71">Pterozetes</emphasis>
</taxonomicName>
which
<bibRefCitation id="EF9A0B9B4627FF8AC55AB0ED1425DA6E" author="Luxton, M." box="[593,769,1009,1036]" pageId="70" pageNumber="71" pagination="71 - 91" refId="ref67435" refString="Luxton, M. (1988 a) A new mite superfamily (Acari: Cryptostigmata). Zoological Journal of the Linnean Society, 93, 71 - 91. https: // doi. org / 10.1111 / j. 1096 - 3642.1988. tb 01360. x" type="journal article" year="1988">Luxton (1988a)</bibRefCitation>
placed with
<taxonomicName id="4C0B0DE94627FF8AC49DB0EE1318DA6E" authorityName="J. Balogh &amp; Mahunka" authorityYear="1966" box="[918,1084,1009,1035]" class="Arachnida" family="Eutegaeidae" genus="Porrhotegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B97FAA784627FF8AC49DB0EE1318DA6E" box="[918,1084,1009,1035]" italics="true" pageId="70" pageNumber="71">Porrhotegaeus</emphasis>
</taxonomicName>
in the
<taxonomicName id="4C0B0DE94627FF8AC386B0EE123FDA6E" authorityName="Luxton" authorityYear="1988" box="[1165,1307,1009,1035]" class="Arachnida" family="Pterozetidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="70" pageNumber="71" phylum="Arthropoda" rank="family">Pterozetidae</taxonomicName>
. The genus clearly does not fit within
<taxonomicName id="4C0B0DE94627FF8AC6B6B70A156DDA4A" authorityName="J. Balogh" authorityYear="1965" box="[445,585,1045,1071]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="70" pageNumber="71" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
, where it was placed by J. and P.
<bibRefCitation id="EF9A0B9B4627FF8AC4B7B70A137ADA4A" author="Balogh, J. &amp; Balogh, P." box="[956,1118,1045,1071]" pageId="70" pageNumber="71" refId="ref64225" refString="Balogh, J. &amp; Balogh, P. (1992) The Oribatid Mites Genera of the World. 2 Vols. Akademiai Kiado, Budapest, 263 pp. &amp; 375 pp." type="book" year="1992">Balogh (1992)</bibRefCitation>
and
<bibRefCitation id="EF9A0B9B4627FF8AC39FB70A1214DA4A" author="Subias, L. S." box="[1172,1328,1045,1071]" pageId="70" pageNumber="71" pagination="3 - 305" refId="ref68838" refString="Subias, L. S. (2004) Listado sistematico sinonimico y biogeografico de los acaros oribatidos (Acariformes, Oribatida) del mundo (1758 - 2002). Graellsia, 60 (Supplement), 3 - 305. https: // doi. org / 10.3989 / graellsia. 2004. v 60. iExtra. 218" type="journal article" year="2004">Subías (2004)</bibRefCitation>
(
<tableCitation id="C68943D14627FF8AC235B70A12B5DA4A" box="[1342,1425,1045,1071]" captionStart="TABLE 1" captionStartId="5.[145,237,449,475]" captionTargetPageId="5" captionText="TABLE 1. Classifications of families and genera of Eutegaeoidea and Cepheoidea. Luxton (1988a) covers Eutegaeoidea only. Norton and Behan-Pelletier (2009) did not include genera under the family group names." httpUri="http://table.plazi.org/id/DF7426E24664FFC9C79AB2DE1502DF9A" pageId="70" pageNumber="71" tableUuid="DF7426E24664FFC9C79AB2DE1502DF9A">Table 1</tableCitation>
) although it shares with
<taxonomicName id="4C0B0DE94627FF8AC694B726150CDA36" authorityName="J. Balogh" authorityYear="1965" box="[415,552,1081,1107]" class="Arachnida" family="Eutegaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="70" pageNumber="71" phylum="Arthropoda" rank="family">Eutegaeidae</taxonomicName>
the prodorsum markedly waisted posterior of the bothridia. The establishment of a separate family for
<taxonomicName id="4C0B0DE94627FF8AC6ACB7421569DA12" authorityName="J. Balogh &amp; Mahunka" authorityYear="1966" box="[423,589,1117,1143]" class="Arachnida" family="Eutegaeidae" genus="Porrhotegaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="70" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="B97FAA784627FF8AC6ACB7421569DA12" box="[423,589,1117,1143]" italics="true" pageId="70" pageNumber="71">Porrhotegaeus</emphasis>
</taxonomicName>
is justified based on a series of autapomorphies including the broad, ovoid lamellae with pointed, free cusps; the flat or concave surface of the notogaster with a raised margin; the reflexed humeral processes and the long notogastral setae positioned marginally that are curved at their bases and pointed dorsally. Furthermore, the immatures are unique among those known in the
<taxonomicName id="4C0B0DE94627FF8AC310B7D61396DA86" authorityName="Balogh" authorityYear="1965" box="[1051,1202,1225,1251]" class="Pilidiophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteronemertea" pageId="70" pageNumber="8" phylum="Nemertea" rank="superFamily" superFamily="Eutegaeoidea">Eutegaeoidea</taxonomicName>
in having strongly-developed lamellar cusps projecting beyond the rostrum, and the notogastral setae
<emphasis id="B97FAA784627FF8AC346B7F1137DDB62" box="[1101,1113,1262,1287]" italics="true" pageId="70" pageNumber="71">c</emphasis>
<subScript id="178F742F4627FF8AC351B61F1346DB6B" attach="left" box="[1114,1122,1280,1294]" fontSize="6" pageId="70" pageNumber="71">3</subScript>
and those of the
<emphasis id="B97FAA784627FF8AC226B7F21211DB62" box="[1325,1333,1261,1287]" italics="true" pageId="70" pageNumber="71">l</emphasis>
,
<emphasis id="B97FAA784627FF8AC24EB7F21277DB62" box="[1349,1363,1261,1287]" italics="true" pageId="70" pageNumber="71">h</emphasis>
and
<emphasis id="B97FAA784627FF8AC285B7F112B8DB62" box="[1422,1436,1262,1287]" italics="true" pageId="70" pageNumber="71">p</emphasis>
series in the nymphs emerge from extremely long, fine apophyses that are part of the gastronotum and which are flanked by elongated, flat scales with strongly serrated margins.
</paragraph>
</subSubSection>
</treatment>
</document>