215 lines
26 KiB
XML
215 lines
26 KiB
XML
<document id="728519C7EC9E050EFD780E269ABA439D" ID-CLB-Dataset="46921" ID-DOI="10.5281/zenodo.199982" ID-GBIF-Dataset="558b2fa1-879c-44e4-956d-9abd9c9a249a" ID-ISSN="1175-5326" ID-Zenodo-Dep="199982" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460351134285" checkinUser="plazi" docAuthor="Wu, Min & Gao, Linhui" docDate="2010" docId="03A187F4FFB6FFD29E8000707ABBFC80" docLanguage="en" docName="zt02725p027.pdf" docOrigin="Zootaxa 2725" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pupopsis yengiawat Wu & Gao, 2010, sp. nov." docType="treatment" docVersion="10" lastPageNumber="23" masterDocId="FF98FF8CFFA5FFC49E170677796BFFDC" masterDocTitle="A review of the genus Pupopsis Gredler, 1898 (Gastropoda: Stylommatophora: Enidae), with the descriptions of eight new species from China" masterLastPageNumber="27" masterPageNumber="1" pageNumber="20" updateTime="1698293683227" updateUser="ExternalLinkService">
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<mods:mods id="1F1040A00E5A54AA647F3F4C5C6D00CE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="F37F1BB0F73D773C59A2CD3A4B172FF6">
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<mods:title id="907FD76FE6939CC9ED2AE57DE068D13C">A review of the genus Pupopsis Gredler, 1898 (Gastropoda: Stylommatophora: Enidae), with the descriptions of eight new species from China</mods:title>
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</mods:titleInfo>
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<mods:name id="B233B464DEF1F9BBB0DD3AC7659C9D50" type="personal">
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<mods:roleTerm id="9258F24AB8B8D4D9FD364434EBA1D662">Author</mods:roleTerm>
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<mods:namePart id="4946D83B6929CA5F6C4005B7D7F564A6">Wu, Min</mods:namePart>
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<mods:name id="6AA77097C121C168485368EA524F2FBF" type="personal">
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<mods:roleTerm id="0FA283F88BFD40955E4E88FE3C6CE567">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9E51CB37AC01434B305438F7D5A7A61C">Gao, Linhui</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E4E141B1DB7BE2ACEF60F1EC299CA756">text</mods:typeOfResource>
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<mods:title id="0AD907ED642C8CBF8579AD051B53F34A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B75A186977AFC40CE08A16BD825A4DBB">
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<mods:date id="2839E59A9D96918D75372BDAECBC9311">2010</mods:date>
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<mods:detail id="41CBB5DCFCC1AE4D75070D65106F280D" type="volume">
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<mods:number id="88F286CE6CFD872F6F502D3467E86246">2725</mods:number>
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<mods:start id="DDBF2167566C003964A6429BAD57F0E3">1</mods:start>
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<mods:end id="DA1BC983E54F8416E05830987A300A22">27</mods:end>
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<mods:classification id="3F69C2FA9A7341771A880F5D467364D3">journal article</mods:classification>
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<mods:identifier id="45CDB75AFCB4371F546BAA48D106E9F9" type="CLB-Dataset">46921</mods:identifier>
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<mods:identifier id="819E884F7F95E1A36406F143DA3EC403" type="DOI">10.5281/zenodo.199982</mods:identifier>
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<mods:identifier id="55ECB3430117DA5DE1FEBB2CCDD6E4B9" type="GBIF-Dataset">558b2fa1-879c-44e4-956d-9abd9c9a249a</mods:identifier>
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<mods:identifier id="2499ED1F08A004FE935581C9AA3234A3" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B7E98E06173CC4EE2E741A2FD1F91518" type="Zenodo-Dep">199982</mods:identifier>
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<treatment id="03A187F4FFB6FFD29E8000707ABBFC80" ID-DOI="http://doi.org/10.5281/zenodo.5778010" ID-GBIF-Taxon="119407880" ID-Zenodo-Dep="5778010" LSID="urn:lsid:plazi:treatment:03A187F4FFB6FFD29E8000707ABBFC80" httpUri="http://treatment.plazi.org/id/03A187F4FFB6FFD29E8000707ABBFC80" lastPageId="22" lastPageNumber="23" pageId="19" pageNumber="20">
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<subSubSection id="C3126569FFB6FFD79E8000707B7DF9FD" box="[151,534,1543,1569]" pageId="19" pageNumber="20" type="nomenclature">
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<paragraph id="8BB736E2FFB6FFD79E8000707B7DF9FD" blockId="19.[151,609,1543,1602]" box="[151,534,1543,1569]" pageId="19" pageNumber="20">
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<heading id="D0FF818EFFB6FFD79E8000707B7DF9FD" bold="true" box="[151,534,1543,1569]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
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||
<emphasis id="B97CEAF0FFB6FFD79E8000707B7DF9FD" bold="true" box="[151,534,1543,1569]" pageId="19" pageNumber="20">
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12.
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<taxonomicName id="4C084D61FFB6FFD79ED4007078DAF9FD" ID-CoL="7SX8C" box="[195,433,1543,1569]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="yengiawat" status="sp. nov.">
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<emphasis id="B97CEAF0FFB6FFD79ED4007078DAF9FD" bold="true" box="[195,433,1543,1569]" italics="true" pageId="19" pageNumber="20">Pupopsis yengiawat</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24F578BFFB6FFD79FAE00707B7DF9FD" box="[441,534,1543,1569]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126569FFB6FFD79E80005D7B0AF99E" box="[151,609,1578,1602]" pageId="19" pageNumber="20" type="description">
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||
<paragraph id="8BB736E2FFB6FFD79E80005D7B0AF99E" blockId="19.[151,609,1543,1602]" box="[151,609,1578,1602]" pageId="19" pageNumber="20">
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<figureCitation id="13332A67FFB6FFD79E80005D7988F99E" box="[151,227,1578,1602]" captionStart="FIGURE 1" captionStartId="4.[151,255,1432,1456]" captionTargetBox="[151,1430,346,1400]" captionTargetId="figure@4.[151,1436,303,1411]" captionTargetPageId="4" captionText="FIGURE 1. Distribution map of the genus Pupopsis. Red dot, Pupopsis dissociabilis (Sturany, 1900); blue dot, P. retrodens (Martens, 1879); blue square, P. maoxian sp. nov.; red square, P. rhodostoma sp. nov.; green dot, P. subpupopsis sp. nov.; green square, P. yengiawat sp. nov. Two grey dots each with two or five species as indicated in the boxes on the map. The species P. paraplesius (Sturany, 1900), P. polystreptus (Sturany, 1900) and P. zilchi sp. nov. are not mapped due to no precise locality information available." httpUri="https://zenodo.org/record/199983/files/figure.png" pageId="19" pageNumber="20">Figs. 1</figureCitation>
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,
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<figureCitation id="13332A67FFB6FFD79EE6005D7994F99E" box="[241,255,1578,1602]" captionStart="FIGURE 3" captionStartId="6.[151,255,1784,1808]" captionTargetBox="[172,1419,194,1772]" captionTargetId="figure@6.[172,1420,194,1774]" captionTargetPageId="6" captionText="FIGURE 3. Apertural view of Pupopsis species. A, Pupopsis pupopsis (Gredler, 1898), SMF- 42076; B, Pupopsis retrodens (Martens, 1879), HBUMM- 06460 – specimen 1; C-D, Pupopsis torquilla (Möllendorff, 1901). C, lectotype, SMF- 42082; D, HBUMM- 05584 – specimen 1; E, Pupopsis hendan sp. nov., holotype, HBUMM- 05592 – specimen 2; F, Pupopsis maoxian sp. nov., holotype, HBUMM- 01023 – specimen 1; G, Pupopsis rhodostoma sp. nov., holotype, HBUMM- 05872 – specimen 2; H, Pupopsis subpupopsis sp. nov., holotype, HBUMM- 05676 – specimen 1; I, Pupopsis subtorquilla sp. nov. A, holotype, HBUMM- 05429 – specimen 2; J, Pupopsis yengiawat sp. nov., holotype, HBUMM- 04439 – specimen 1; K, Pupopsis yuxu sp. nov., Holotype, HBUMM- 06089 – specimen 3; L, Pupopsis zilchi sp. nov., holotype, SMF- 334777." httpUri="https://zenodo.org/record/199985/files/figure.png" pageId="19" pageNumber="20">3</figureCitation>
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J, 5E, 11D–11E, 12;
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<tableCitation id="C68A0359FFB6FFD79FF0005D7B2CF99E" box="[487,583,1578,1602]" captionStart="TABLE 2" captionStartId="3.[151,239,617,641]" captionTargetBox="[151,1435,700,1395]" captionTargetPageId="3" captionText="TABLE 2. Pupopsis species and their conchological measurements (data from Sturany, 1900; Möllendorff, 1901; Kobelt, 1902 and this study; *, calculated from the published data; /, data unavailable)." httpUri="http://table.plazi.org/id/DF77666AFFA6FFC79E80041E7D5DFD7C" pageId="19" pageNumber="20" tableUuid="DF77666AFFA6FFC79E80041E7D5DFD7C">Tables 2</tableCitation>
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,
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<tableCitation id="C68A0359FFB6FFD79C44005D7B0AF99E" box="[595,609,1578,1602]" captionStart="TABLE 3" captionStartId="10.[151,239,908,932]" captionText="TABLE 3. Comparison of genital organs among Pupopsis species. + present / yes; - absent / no." pageId="19" pageNumber="20">3</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126569FFB6FFD79E8000047B32F8C6" pageId="19" pageNumber="20" type="materials_examined">
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<paragraph id="8BB736E2FFB6FFD79E800004786BF90E" blockId="19.[151,1437,1650,2034]" pageId="19" pageNumber="20">
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<emphasis id="B97CEAF0FFB6FFD79E8000047823F957" bold="true" box="[151,328,1650,1675]" pageId="19" pageNumber="20">
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<typeStatus id="54B38840FFB6FFD79E80000479B9F957" box="[151,210,1651,1675]" pageId="19" pageNumber="20">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54B38840FFB6FFD79F45000578D4F956" box="[338,447,1650,1674]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
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: HBUMM-04439–specimen 1 (fma);
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<taxonomicName id="4C084D61FFB6FFD79D6400057A86F956" box="[883,1005,1650,1674]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="22" phylum="Mollusca" rank="species" species="yengiawat" status="sp. nov.">Yengiawat</taxonomicName>
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, 41°24ʹ N, 79°24ʹE, Xinjiang,
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<collectingCountry id="F31F7672FFB6FFD79B4500057CF3F956" box="[1362,1432,1650,1674]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
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,
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<date id="FFB61022FFB6FFD79E8000E1784CF972" box="[151,295,1686,1710]" pageId="19" pageNumber="20" value="2005-06-30">2005–VI–30</date>
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, leg. A. Kanatbaytegi.
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<typeStatus id="54B38840FFB6FFD79C3800E07BCBF973" box="[559,672,1687,1711]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="ED19AE27FFB6FFD79CBA00E07A71F972" box="[685,794,1687,1710]" pageId="19" pageNumber="20">HBUMM</collectionCode>
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-04439–specimens 2–8 (7 fms), collection data same as
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<typeStatus id="54B38840FFB6FFD79E8000CD7996F90E" box="[151,253,1722,1746]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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<paragraph id="8BB736E2FFB6FFD79EDE00A87B3FF92A" blockId="19.[151,1437,1650,2034]" box="[201,596,1758,1783]" pageId="19" pageNumber="20">
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<emphasis id="B97CEAF0FFB6FFD79EDE00A8780EF92B" bold="true" box="[201,357,1758,1783]" pageId="19" pageNumber="20">
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<typeStatus id="54B38840FFB6FFD79EDE00A87869F92B" box="[201,258,1759,1783]" pageId="19" pageNumber="20">Type</typeStatus>
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locality.
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</emphasis>
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<taxonomicName id="4C084D61FFB6FFD79F7900A9788CF92A" box="[366,487,1758,1782]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="22" phylum="Mollusca" rank="species" species="yengiawat" status="sp. nov.">Yengiawat</taxonomicName>
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, Xinjiang
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</paragraph>
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<paragraph id="8BB736E2FFB6FFD79EDE01757B32F8C6" blockId="19.[151,1437,1650,2034]" box="[201,601,1794,1819]" pageId="19" pageNumber="20">
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<emphasis id="B97CEAF0FFB6FFD79EDE017578D9F8C7" bold="true" box="[201,434,1794,1819]" pageId="19" pageNumber="20">Material examined.</emphasis>
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<typeStatus id="54B38840FFB6FFD79FAE01747884F8C7" box="[441,495,1795,1819]" pageId="19" pageNumber="20">Type</typeStatus>
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material.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126569FFB6FFD79EDE01517A01F8E2" box="[201,874,1830,1855]" pageId="19" pageNumber="20" type="distribution">
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<paragraph id="8BB736E2FFB6FFD79EDE01517A01F8E2" blockId="19.[151,1437,1650,2034]" box="[201,874,1830,1855]" pageId="19" pageNumber="20">
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<emphasis id="B97CEAF0FFB6FFD79EDE01517809F8E3" bold="true" box="[201,354,1830,1855]" pageId="19" pageNumber="20">Distribution.</emphasis>
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Xinjiang (only known from the
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<typeStatus id="54B38840FFB6FFD79CD901507B95F8E3" box="[718,766,1831,1855]" pageId="19" pageNumber="20">type</typeStatus>
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locality).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126569FFB6FFD79EDE013C7CF6F8BE" box="[201,1437,1866,1891]" pageId="19" pageNumber="20" type="etymology">
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<paragraph id="8BB736E2FFB6FFD79EDE013C7CF6F8BE" blockId="19.[151,1437,1650,2034]" box="[201,1437,1866,1891]" pageId="19" pageNumber="20">
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<emphasis id="B97CEAF0FFB6FFD79EDE013C7824F8BF" bold="true" box="[201,335,1867,1891]" pageId="19" pageNumber="20">Etymology.</emphasis>
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The new species is named after the
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<typeStatus id="54B38840FFB6FFD79CF3013C7A7FF8BF" box="[740,788,1867,1891]" pageId="19" pageNumber="20">type</typeStatus>
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locality “
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<taxonomicName id="4C084D61FFB6FFD79D97013D7A94F8BE" box="[896,1023,1866,1890]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="19" pageNumber="22" phylum="Mollusca" rank="species" species="yengiawat" status="sp. nov.">Yengiawat</taxonomicName>
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.” The epithet is a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126569FFB6FFD29EDE01197816FD58" lastPageId="22" lastPageNumber="23" pageId="19" pageNumber="20" type="diagnosis">
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<paragraph id="8BB736E2FFB6FFD79EDE01197DD4F82E" blockId="19.[151,1437,1650,2034]" pageId="19" pageNumber="20">
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<emphasis id="B97CEAF0FFB6FFD79EDE01197828F85B" bold="true" box="[201,323,1902,1927]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
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Shell with 6.125–6.750 whorls. Height 8.0–9.4, diam. maj. 4.1–4.7. Parietal tooth distinct, indistinct or absent. Columellar tooth absent. No tooth on palatal wall. Epiphallus straight. Flagellum absent. Penial caecum absent. Penis distally with one V-shaped pilaster. A-2 near A-3 internally without a ring of papillae. A-3 opening into A-2 without papilla. Bursa copulatrix duct proximally straight. Diverticle present.
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</paragraph>
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<caption id="DF77666AFFB1FFD09E8001407B1AF813" httpUri="https://zenodo.org/record/199993/files/figure.png" pageId="20" pageNumber="21" targetBox="[151,1436,194,1826]" targetPageId="20">
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<paragraph id="8BB736E2FFB1FFD09E8001407B1AF813" blockId="20.[151,1436,1847,1999]" pageId="20" pageNumber="21">
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<emphasis id="B97CEAF0FFB1FFD09E800140784CF893" bold="true" box="[151,295,1847,1871]" pageId="20" pageNumber="21">FIGURE 11.</emphasis>
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A–C,
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<taxonomicName id="4C084D61FFB1FFD09F66014F7B37F893" box="[369,604,1848,1871]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="subtorquilla" status="sp. nov.">
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<emphasis id="B97CEAF0FFB1FFD09F66014F7B37F893" box="[369,604,1848,1871]" italics="true" pageId="20" pageNumber="21">Pupopsis subtorquilla</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24F578BFFB1FFD09C7201407BD6F893" box="[613,701,1847,1871]" pageId="20" pageNumber="21" rank="species">
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<emphasis id="B97CEAF0FFB1FFD09C7201407BD6F893" bold="true" box="[613,701,1847,1871]" pageId="20" pageNumber="21">sp. nov.</emphasis>
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</taxonomicNameLabel>
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A, holotype, HBUMM-05429–specimen 2; B, paratype, HBUMM- 05429–specimen 1; C, paraytpe, HBUMM-00349–specimen 2. D–E,
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<taxonomicName id="4C084D61FFB1FFD09DA2012F7DF8F8B3" box="[949,1171,1880,1903]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="yengiawat" status="sp. nov.">
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<emphasis id="B97CEAF0FFB1FFD09DA2012F7DF8F8B3" box="[949,1171,1880,1903]" italics="true" pageId="20" pageNumber="21">Pupopsis yengiawat</emphasis>
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</taxonomicName>
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<emphasis id="B97CEAF0FFB1FFD09AB501207C6BF8B3" bold="true" box="[1186,1280,1879,1903]" pageId="20" pageNumber="21">
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||
<taxonomicNameLabel id="A24F578BFFB1FFD09AB501207C6BF8B3" box="[1186,1280,1879,1903]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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D, holotype, HBUMM-04439–specimen 1; E, paratype, HBUMM-04439–specimen 6. F–G,
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<taxonomicName id="4C084D61FFB1FFD09A1F010F7DC3F853" box="[1032,1192,1912,1935]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="yuxu" status="sp. nov.">
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<emphasis id="B97CEAF0FFB1FFD09A1F010F7DC3F853" box="[1032,1192,1912,1935]" italics="true" pageId="20" pageNumber="21">Pupopsis yuxu</emphasis>
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</taxonomicName>
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||
<emphasis id="B97CEAF0FFB1FFD09AA401007C66F853" bold="true" box="[1203,1293,1911,1935]" pageId="20" pageNumber="21">
|
||
<taxonomicNameLabel id="A24F578BFFB1FFD09AA401007C66F853" box="[1203,1293,1911,1935]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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F, holotype, HBUMM-06089–specimen 3; G, paratype, HBUMM-06089–specimen 2. H–I,
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<taxonomicName id="4C084D61FFB1FFD09DE901EF7DCEF873" box="[1022,1189,1944,1967]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="zilchi" status="sp. nov.">
|
||
<emphasis id="B97CEAF0FFB1FFD09DE901EF7DCEF873" box="[1022,1189,1944,1967]" italics="true" pageId="20" pageNumber="21">Pupopsis zilchi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97CEAF0FFB1FFD09AB801E07C62F873" bold="true" box="[1199,1289,1943,1967]" pageId="20" pageNumber="21">
|
||
<taxonomicNameLabel id="A24F578BFFB1FFD09AB801E07C62F873" box="[1199,1289,1943,1967]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
H, holotype, SMF-specimen; I, paratype, SMF-specimen.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF77666AFFB0FFD19E80000D78D3F90E" httpUri="https://zenodo.org/record/199994/files/figure.png" pageId="21" pageNumber="22" targetBox="[181,1411,219,1657]" targetPageId="21">
|
||
<paragraph id="8BB736E2FFB0FFD19E80000D78D3F90E" blockId="21.[151,1436,1658,1746]" pageId="21" pageNumber="22">
|
||
<emphasis id="B97CEAF0FFB0FFD19E80000D7841F94E" bold="true" box="[151,298,1658,1682]" pageId="21" pageNumber="22">FIGURE 12.</emphasis>
|
||
<taxonomicName id="4C084D61FFB0FFD19F23000C7B65F94E" box="[308,526,1659,1682]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="yengiawat" status="sp. nov.">
|
||
<emphasis id="B97CEAF0FFB0FFD19F23000C7B65F94E" box="[308,526,1659,1682]" italics="true" pageId="21" pageNumber="22">Pupopsis yengiawat</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24F578BFFB0FFD19C00000D7B1BF94E" box="[535,624,1658,1682]" pageId="21" pageNumber="22" rank="species">
|
||
<emphasis id="B97CEAF0FFB0FFD19C00000D7B1BF94E" bold="true" box="[535,624,1658,1682]" pageId="21" pageNumber="22">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, HBUMM-04439–specimen 1, holotype. A, vagina and its distal part; B, penial appendage; C, the genital section from penis to vas deferens. Arrow indicates the absence of flagellum; D, distal part of penis, interior view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB736E2FFB0FFD29EDE00897C71FF2B" blockId="21.[151,1436,1790,2031]" lastBlockId="22.[151,1437,151,860]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
|
||
<emphasis id="B97CEAF0FFB0FFD19EDE00897860F8CB" bold="true" box="[201,267,1790,1815]" pageId="21" pageNumber="22">Shell.</emphasis>
|
||
Cylindrical with conical top; apex not acuminate; dextral; thin-shelled; solid; dull; with 6.125–6.446– 6.750 whorls. Whorls with indistinct spiral grooves; somewhat convex. Embryonic shell smooth; polished; with 1.500–1.554–1.625 whorls. Postnuclear whorls wrinkled. Suture simple. Last whorl gradually ascending toward aperture; rounded at periphery. Shell 8.0–8.8–9.4 high; 4.1–4.4–
|
||
<quantity id="4CF09B07FFB0FFD19D6E011C7AD0F85F" box="[889,955,1899,1923]" metricMagnitude="-1" metricUnit="m" metricValue="1.1938" pageId="21" pageNumber="22" unit="in" value="4.7">4.7 in</quantity>
|
||
diam. maj. Height/diam. maj. ratio 1.90– 2.01–2.08. Aperture subovate; almost vertical; its insertions connected by a distinct parietal callus; with or without teeth; 3.1–3.5–3.9 high, 2.6–2.9–3.2 wide. Ratio of shell height to aperture height 2.36–2.57–2.72. Angular tubercle fused with aperture insertion. Parietal tooth clearly present, or a more or less distinct, low thickening, or absent. Palatal tooth absent. Palatal margin rounded. Peristome broadly and abruptly reflexed; outer edge sharp; with a strong, white callus within. Palatal wall toothless. Columellar margin reflexed; simple. Columellar tooth absent. Umbilicus narrowly open. Shell, including upper whorls, uniformly dirty white; aperture white.
|
||
</paragraph>
|
||
<paragraph id="8BB736E2FFB3FFD29EDE07747816FD58" blockId="22.[151,1437,151,860]" pageId="22" pageNumber="23">
|
||
<emphasis id="B97CEAF0FFB3FFD29EDE07747854FEC0" bold="true" box="[201,319,259,284]" pageId="22" pageNumber="23">Genitalia.</emphasis>
|
||
Vas deferens entering epiphallus apically. Epiphallus moderately long; cylindrical and of uniform thickness; straight. Epiphallic caecum apically blunt. Flagellum wanting (
|
||
<figureCitation id="13332A67FFB3FFD29DF607507D5DFEE3" box="[993,1078,295,319]" captionStart="FIGURE 12" captionStartId="21.[151,255,1658,1682]" captionTargetBox="[181,1411,219,1657]" captionTargetId="figure@21.[151,1435,194,1663]" captionTargetPageId="21" captionText="FIGURE 12. Pupopsis yengiawat sp. nov., HBUMM- 04439 – specimen 1, holotype. A, vagina and its distal part; B, penial appendage; C, the genital section from penis to vas deferens. Arrow indicates the absence of flagellum; D, distal part of penis, interior view." httpUri="https://zenodo.org/record/199994/files/figure.png" pageId="22" pageNumber="23">Fig. 12</figureCitation>
|
||
, arrow). Penial caecum absent. Penis with terminal entrance of epiphallus; evenly thick; thin-walled; with pilasters fused at epiphallic pore forming velum; with one V-shaped pilaster. V-shaped pilaster with proximal free end approaching at penial retractor insertion. A large papilla developed by fused distal pilasters forming V-shaped structure present. Penial appendix moderately long; divided into sections A-1+A-2, A-3 and A-4+A-5. A-1 short. A-1 and A-2 with evenly arranged transversal folds. A-1 and A-2 fused. A-2 near A-3 internally without a ring of papillae. A-2 and A-3 not fused. A-3 opening into A-2 without papilla. Boundary between A-4 and A-5 indistinct. A-5 short; straight. Free oviduct moderately long. Bursa copulatrix duct moderately long; proximally straight. Bursa copulatrix with short duct; well defined. Diverticle longer than reservoir. Diverticle diverging from bursa duct closer to reservoir than to duct’s proximal end.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3126569FFB3FFD29EDE04E77ABBFC80" pageId="22" pageNumber="23" type="discussion">
|
||
<paragraph id="8BB736E2FFB3FFD29EDE04E77ABBFC80" blockId="22.[151,1437,151,860]" pageId="22" pageNumber="23">
|
||
<emphasis id="B97CEAF0FFB3FFD29EDE04E77856FD74" bold="true" box="[201,317,656,680]" pageId="22" pageNumber="23">Remarks.</emphasis>
|
||
This new species is quite unique among the known
|
||
<taxonomicName id="4C084D61FFB3FFD29D8D04E67D6BFD74" box="[922,1024,657,680]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="20" phylum="Mollusca" rank="species" species="pupopsis">
|
||
<emphasis id="B97CEAF0FFB3FFD29D8D04E67D6BFD74" box="[922,1024,657,680]" italics="true" pageId="22" pageNumber="23">Pupopsis</emphasis>
|
||
</taxonomicName>
|
||
species. It is included in this genus because its aperture is usually furnished with a weak tooth. The species has neither a columellar tooth nor a palatal tooth. Even the weak parietal tooth is not always present in the observed
|
||
<typeStatus id="54B38840FFB3FFD29DFA04AF7D76FD2C" box="[1005,1053,728,752]" pageId="22" pageNumber="23">type</typeStatus>
|
||
specimens. The shell of the new species resembles in overall appearance the toothless genus
|
||
<taxonomicName id="4C084D61FFB3FFD29D2D048B7DFBFCC8" authority="Westerlund, 1887" authorityName="Westerlund" authorityYear="1887" box="[826,1168,764,788]" class="Gastropoda" family="Enidae" genus="Subzebrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97CEAF0FFB3FFD29D2D048B7AA8FCC8" box="[826,963,764,788]" italics="true" pageId="22" pageNumber="23">Subzebrinus</emphasis>
|
||
Westerlund, 1887
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B97CEAF0FFB3FFD29AB7048A7DB5FCC8" box="[1184,1246,765,788]" italics="true" pageId="22" pageNumber="23">sensu</emphasis>
|
||
<bibRefCitation id="EF994B13FFB3FFD29AF1048B7CF3FCC8" author="Schileyko" box="[1254,1432,764,788]" pageId="22" pageNumber="23" refString="Schileyko, A. A. (1998) Treatise on recent terrestrial pulmonate molluscs. Part. 2. Gastrocoptidae, Hypselostomatidae, Vertiginidae, Truncatellinidae, Pachnodidae, Enidae, Sagdidae. Ruthenica, Supplement 2, 129 - 261." type="journal article" year="1998">Schileyko 1998</bibRefCitation>
|
||
, =
|
||
<taxonomicName id="4C084D61FFB3FFD29EB805577A16FCEB" authority="Westerlund, 1887 sensu Bank & Neubert 1998" authorityName="Westerlund, 1887 sensu Bank & Neubert" authorityYear="1998" box="[175,893,799,824]" class="Gastropoda" family="Enidae" genus="Pseudonapaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97CEAF0FFB3FFD29EB805577834FCE4" box="[175,351,800,824]" italics="true" pageId="22" pageNumber="23">Pseudonapaeus</emphasis>
|
||
Westerlund, 1887
|
||
<emphasis id="B97CEAF0FFB3FFD29C2105567B1FFCE4" box="[566,628,801,824]" italics="true" pageId="22" pageNumber="23">sensu</emphasis>
|
||
<bibRefCitation id="EF994B13FFB3FFD29C6B05687A16FCEB" author="Bank" box="[636,893,799,823]" pageId="22" pageNumber="23" refString="Bank, R. A. & Neubert, E. (1998) Notes on Buliminidae, 5: on the systematic position of Arabian Buliminidae (Gastropoda Pulmonata), with the description of a new genus. Basteria, 61, 73 - 84." type="journal article" year="1998">Bank & Neubert 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
). However,
|
||
<taxonomicName id="4C084D61FFB3FFD29A1105577DE4FCE4" box="[1030,1167,800,824]" class="Gastropoda" family="Enidae" genus="Subzebrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97CEAF0FFB3FFD29A1105577DE4FCE4" box="[1030,1167,800,824]" italics="true" pageId="22" pageNumber="23">Subzebrinus</emphasis>
|
||
</taxonomicName>
|
||
has neither a flagellum nor a diverticle and shows a completely different interior penial structure.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |