160 lines
19 KiB
XML
160 lines
19 KiB
XML
<document id="3840B55C4A041F9D14B25E8E769A1BC7" ID-CLB-Dataset="46921" ID-DOI="10.5281/zenodo.199982" ID-GBIF-Dataset="558b2fa1-879c-44e4-956d-9abd9c9a249a" ID-ISSN="1175-5326" ID-Zenodo-Dep="199982" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460351134285" checkinUser="plazi" docAuthor="Wu, Min & Gao, Linhui" docDate="2010" docId="03A187F4FFB3FFDC9E8005D87C5FFD83" docLanguage="en" docName="zt02725p027.pdf" docOrigin="Zootaxa 2725" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pupopsis yuxu Wu & Gao, 2010, sp. nov." docType="treatment" docVersion="10" lastPageNumber="25" masterDocId="FF98FF8CFFA5FFC49E170677796BFFDC" masterDocTitle="A review of the genus Pupopsis Gredler, 1898 (Gastropoda: Stylommatophora: Enidae), with the descriptions of eight new species from China" masterLastPageNumber="27" masterPageNumber="1" pageNumber="23" updateTime="1698293683227" updateUser="ExternalLinkService">
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<mods:mods id="859D09F9D5FADC27D737958239F09AD6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D0B9FE868E9B81F1EA03F6D8AEF584EC">
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<mods:title id="BC4C5CB89C5B2C035416F3019B016E31">A review of the genus Pupopsis Gredler, 1898 (Gastropoda: Stylommatophora: Enidae), with the descriptions of eight new species from China</mods:title>
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</mods:titleInfo>
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<mods:name id="087E4F2808607CBDD8E50C33E27B8285" type="personal">
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<mods:roleTerm id="6435C33DE8954C930120FA2465956A80">Author</mods:roleTerm>
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<mods:namePart id="FA696195B7A838C5371671C4537575DB">Wu, Min</mods:namePart>
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<mods:name id="AD645A121174C7340AFD5920BC4107FC" type="personal">
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<mods:roleTerm id="135656046A5B215D3EB3666B2E9856B6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3E1E348A45909F96FB9F19AAB1D60C2A">Gao, Linhui</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="FDB7259E23C16AAA37989E034F2F01F1">text</mods:typeOfResource>
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<mods:title id="FDFA68202317C24A8224AF3A73F7B2EB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0C2569A43A8E706B3621578542051B3F">
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<mods:date id="ADA10AFD6E90FEC06E52F4F553F1BE1A">2010</mods:date>
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<mods:detail id="BD47267794A424C7CCF9E4C07F13FA4E" type="volume">
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<mods:number id="580F55B89AD38175CCE404CEA024629D">2725</mods:number>
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<mods:start id="CB3D4B3EBA65B7F69ADEECCB5902D575">1</mods:start>
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<mods:classification id="484F1DDE33D6921001E46E0B18AB5840">journal article</mods:classification>
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<mods:identifier id="555114A691DDFEF46A0707B344F0B4ED" type="CLB-Dataset">46921</mods:identifier>
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<mods:identifier id="122BF142369073174591E32EC53FDA2D" type="DOI">10.5281/zenodo.199982</mods:identifier>
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<mods:identifier id="6BDF72E57ADC64BAE475D2F3D7EF2D1F" type="GBIF-Dataset">558b2fa1-879c-44e4-956d-9abd9c9a249a</mods:identifier>
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<mods:identifier id="13E6399A3D974C19F1E4D449BC67EC09" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E883603E5B7D61F36C3DB47262611DA0" type="Zenodo-Dep">199982</mods:identifier>
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<treatment id="03A187F4FFB3FFDC9E8005D87C5FFD83" ID-DOI="http://doi.org/10.5281/zenodo.5625411" ID-GBIF-Taxon="119407885" ID-Zenodo-Dep="5625411" LSID="urn:lsid:plazi:treatment:03A187F4FFB3FFDC9E8005D87C5FFD83" httpUri="http://treatment.plazi.org/id/03A187F4FFB3FFDC9E8005D87C5FFD83" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
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<subSubSection id="C3126569FFB3FFD29E8005D878B3FC15" box="[151,472,943,969]" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph id="8BB736E2FFB3FFD29E8005D878B3FC15" blockId="22.[151,609,943,1002]" box="[151,472,943,969]" pageId="22" pageNumber="23">
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<heading id="D0FF818EFFB3FFD29E8005D878B3FC15" bold="true" box="[151,472,943,969]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
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<emphasis id="B97CEAF0FFB3FFD29E8005D878B3FC15" bold="true" box="[151,472,943,969]" pageId="22" pageNumber="23">
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13.
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<taxonomicName id="4C084D61FFB3FFD29ED405D8781FFC15" ID-CoL="7SX8D" box="[195,372,943,969]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="yuxu" status="sp. nov.">
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<emphasis id="B97CEAF0FFB3FFD29ED405D8781FFC15" bold="true" box="[195,372,943,969]" italics="true" pageId="22" pageNumber="23">Pupopsis yuxu</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24F578BFFB3FFD29F6D05D878B3FC15" box="[378,472,943,969]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126569FFB3FFD29E8005A57B0AFC36" box="[151,609,978,1002]" pageId="22" pageNumber="23" type="description">
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<paragraph id="8BB736E2FFB3FFD29E8005A57B0AFC36" blockId="22.[151,609,943,1002]" box="[151,609,978,1002]" pageId="22" pageNumber="23">
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<figureCitation id="13332A67FFB3FFD29E8005A57988FC36" box="[151,227,978,1002]" captionStart="FIGURE 1" captionStartId="4.[151,255,1432,1456]" captionTargetBox="[151,1430,346,1400]" captionTargetId="figure@4.[151,1436,303,1411]" captionTargetPageId="4" captionText="FIGURE 1. Distribution map of the genus Pupopsis. Red dot, Pupopsis dissociabilis (Sturany, 1900); blue dot, P. retrodens (Martens, 1879); blue square, P. maoxian sp. nov.; red square, P. rhodostoma sp. nov.; green dot, P. subpupopsis sp. nov.; green square, P. yengiawat sp. nov. Two grey dots each with two or five species as indicated in the boxes on the map. The species P. paraplesius (Sturany, 1900), P. polystreptus (Sturany, 1900) and P. zilchi sp. nov. are not mapped due to no precise locality information available." httpUri="https://zenodo.org/record/199983/files/figure.png" pageId="22" pageNumber="23">Figs. 1</figureCitation>
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,
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<figureCitation id="13332A67FFB3FFD29EE605A57995FC36" box="[241,254,978,1002]" captionStart="FIGURE 3" captionStartId="6.[151,255,1784,1808]" captionTargetBox="[172,1419,194,1772]" captionTargetId="figure@6.[172,1420,194,1774]" captionTargetPageId="6" captionText="FIGURE 3. Apertural view of Pupopsis species. A, Pupopsis pupopsis (Gredler, 1898), SMF- 42076; B, Pupopsis retrodens (Martens, 1879), HBUMM- 06460 – specimen 1; C-D, Pupopsis torquilla (Möllendorff, 1901). C, lectotype, SMF- 42082; D, HBUMM- 05584 – specimen 1; E, Pupopsis hendan sp. nov., holotype, HBUMM- 05592 – specimen 2; F, Pupopsis maoxian sp. nov., holotype, HBUMM- 01023 – specimen 1; G, Pupopsis rhodostoma sp. nov., holotype, HBUMM- 05872 – specimen 2; H, Pupopsis subpupopsis sp. nov., holotype, HBUMM- 05676 – specimen 1; I, Pupopsis subtorquilla sp. nov. A, holotype, HBUMM- 05429 – specimen 2; J, Pupopsis yengiawat sp. nov., holotype, HBUMM- 04439 – specimen 1; K, Pupopsis yuxu sp. nov., Holotype, HBUMM- 06089 – specimen 3; L, Pupopsis zilchi sp. nov., holotype, SMF- 334777." httpUri="https://zenodo.org/record/199985/files/figure.png" pageId="22" pageNumber="23">3</figureCitation>
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K, 5F, 11F–11G, 13;
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<tableCitation id="C68A0359FFB3FFD29FFF05A57B2CFC36" box="[488,583,978,1002]" captionStart="TABLE 2" captionStartId="3.[151,239,617,641]" captionTargetBox="[151,1435,700,1395]" captionTargetPageId="3" captionText="TABLE 2. Pupopsis species and their conchological measurements (data from Sturany, 1900; Möllendorff, 1901; Kobelt, 1902 and this study; *, calculated from the published data; /, data unavailable)." httpUri="http://table.plazi.org/id/DF77666AFFA6FFC79E80041E7D5DFD7C" pageId="22" pageNumber="23" tableUuid="DF77666AFFA6FFC79E80041E7D5DFD7C">Tables 2</tableCitation>
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,
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<tableCitation id="C68A0359FFB3FFD29C4405A57B0AFC36" box="[595,609,978,1002]" captionStart="TABLE 3" captionStartId="10.[151,239,908,932]" captionText="TABLE 3. Comparison of genital organs among Pupopsis species. + present / yes; - absent / no." pageId="22" pageNumber="23">3</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126569FFB3FFD29E80026C7B32FB3A" pageId="22" pageNumber="23" type="materials_examined">
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<paragraph id="8BB736E2FFB3FFD29E80026C7A49FB42" blockId="22.[151,1436,1050,1980]" pageId="22" pageNumber="23">
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<emphasis id="B97CEAF0FFB3FFD29E80026C782DFBEF" bold="true" box="[151,326,1050,1075]" pageId="22" pageNumber="23">
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<typeStatus id="54B38840FFB3FFD29E80026C79B9FBEF" box="[151,210,1051,1075]" pageId="22" pageNumber="23">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54B38840FFB3FFD29F58026D78D7FBEE" box="[335,444,1050,1074]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
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: HBUMM-06089–specimen 3 (fma); Yuxushan Hill,
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<quantity id="4CF09B07FFB3FFD29A0B026D7D0FFBEE" box="[1052,1124,1050,1074]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="23" unit="km" value="10.0">10 km</quantity>
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north of Wenxian, site No. 0 45, 1070 m a.s.l., 32°57ʹ35.5ʹ N, 104°40ʹ41.6ʹ E, Wenxian County, S. Gansu,
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<collectingCountry id="F31F7672FFB3FFD29A1A02497D38FB8A" box="[1037,1107,1086,1110]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
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,
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<date id="FFB61022FFB3FFD29A4802497D84FB8A" box="[1119,1263,1086,1110]" pageId="22" pageNumber="23" value="2006-09-27">2006–IX–27</date>
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, leg. M. Wu, J. Liu, W. Zheng & L. Gao.
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<typeStatus id="54B38840FFB3FFD29FF202147B3DFBA7" box="[485,598,1123,1147]" pageId="22" pageNumber="23" type="paratype">Paratypes</typeStatus>
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: HBUMM-06089–specimens 1, 2 (2 fma, shell partially removed for dissection), 4, 5 (2 fms), same collection data as
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<typeStatus id="54B38840FFB3FFD29CAE02F17A74FB42" box="[697,799,1158,1182]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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<paragraph id="8BB736E2FFB3FFD29EDE02DC7D1AFB1E" blockId="22.[151,1436,1050,1980]" box="[201,1137,1194,1219]" pageId="22" pageNumber="23">
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||
<emphasis id="B97CEAF0FFB3FFD29EDE02DC780EFB1F" bold="true" box="[201,357,1194,1219]" pageId="22" pageNumber="23">
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<typeStatus id="54B38840FFB3FFD29EDE02DC7869FB1F" box="[201,258,1195,1219]" pageId="22" pageNumber="23">Type</typeStatus>
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locality.
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</emphasis>
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Yuxushan Hill,
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<quantity id="4CF09B07FFB3FFD29C0A02DD7B0FFB1E" box="[541,612,1194,1218]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="22" pageNumber="23" unit="km" value="10.0">10 km</quantity>
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north of Wenxian, Wenxian County, S. Gansu.
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</paragraph>
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<paragraph id="8BB736E2FFB3FFD29EDE02B97B32FB3A" blockId="22.[151,1436,1050,1980]" box="[201,601,1230,1255]" pageId="22" pageNumber="23">
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||
<emphasis id="B97CEAF0FFB3FFD29EDE02B978D9FB3B" bold="true" box="[201,434,1230,1255]" pageId="22" pageNumber="23">Material examined.</emphasis>
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<typeStatus id="54B38840FFB3FFD29FAE02B87884FB3B" box="[441,495,1231,1255]" pageId="22" pageNumber="23">Type</typeStatus>
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material.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3126569FFB3FFD29EDE02857A4DFAD6" box="[201,806,1266,1291]" pageId="22" pageNumber="23" type="distribution">
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<paragraph id="8BB736E2FFB3FFD29EDE02857A4DFAD6" blockId="22.[151,1436,1050,1980]" box="[201,806,1266,1291]" pageId="22" pageNumber="23">
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<emphasis id="B97CEAF0FFB3FFD29EDE02857809FAD7" bold="true" box="[201,354,1266,1291]" pageId="22" pageNumber="23">Distribution.</emphasis>
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Gansu (only known from
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<typeStatus id="54B38840FFB3FFD29C9D02847BD1FAD7" box="[650,698,1267,1291]" pageId="22" pageNumber="23">type</typeStatus>
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locality).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126569FFB3FFD29EDE03607CF3FAF2" box="[201,1432,1302,1327]" pageId="22" pageNumber="23" type="etymology">
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<paragraph id="8BB736E2FFB3FFD29EDE03607CF3FAF2" blockId="22.[151,1436,1050,1980]" box="[201,1432,1302,1327]" pageId="22" pageNumber="23">
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<emphasis id="B97CEAF0FFB3FFD29EDE03607824FAF3" bold="true" box="[201,335,1303,1327]" pageId="22" pageNumber="23">Etymology.</emphasis>
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The new species is named after the
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<typeStatus id="54B38840FFB3FFD29CF203607A7EFAF3" box="[741,789,1303,1327]" pageId="22" pageNumber="23">type</typeStatus>
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locality “Yuxushan”. The epithet is a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126569FFB3FFDC9EDE034D7A95FDCB" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23" type="diagnosis">
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<paragraph id="8BB736E2FFB3FFD29EDE034D7A9DFA3E" blockId="22.[151,1436,1050,1980]" pageId="22" pageNumber="23">
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<emphasis id="B97CEAF0FFB3FFD29EDE034D7828FA8F" bold="true" box="[201,323,1338,1363]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
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Shell with 6.750–7.500 whorls. Height 6.2–6.7, diam. maj. 2.6–2.9. Parietal tooth present or absent. Columellar tooth invisible from aperture but expanded inwards. Palatal tooth ridge-like. Epiphallus forming a few loops. Flagellum present. Penial caecum present. Penis distally without V-shaped pilasters, with two thick, parallel pilasters connected at distal end. A-2 near A-3 internally with a ring of papillae. A-3 opening into A-2 without papilla. Bursa copulatrix duct proximally twisted. Diverticle present.
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</paragraph>
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<paragraph id="8BB736E2FFB3FFD29EDE03997A77F88C" blockId="22.[151,1436,1050,1980]" pageId="22" pageNumber="23">
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<emphasis id="B97CEAF0FFB3FFD29EDE03997860F9DB" bold="true" box="[201,267,1518,1543]" pageId="22" pageNumber="23">Shell.</emphasis>
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Cylindrical with conical top; apex not acuminate; dextral; thin-shelled; solid; translucent or semitranslucent; glossy; with 6.750–7.000–7.500 whorls. Whorls never spirally grooved; convex. Embryonic shell smooth; polished; with 1.500–1.650–1.750 whorls. Postnuclear whorls wrinkled. Suture simple. Last whorl gradually ascending towards aperture; rounded at periphery. Shell 6.2–6.5–6.7 high; 2.6–2.7–
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<quantity id="4CF09B07FFB3FFD29AC4002D7C7CF9AE" box="[1235,1303,1626,1650]" metricMagnitude="-2" metricUnit="m" metricValue="7.366" pageId="22" pageNumber="23" unit="in" value="2.9">2.9 in</quantity>
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diam. maj. Height/diam. maj. ratio 2.33–2.38–2.50. Aperture ovate; almost vertical; its insertions separated; 2.2–2.3–2.4 high, 1.8–1.8–1.9 wide. Ratio of shell height to aperture height 2.72–2.86–3.00. Palatal tooth ridge-like, extending inward for ca. 1/4–1/3 whorl. Palatal margin rounded. Peristome expanded; sharp. Parietal callus indistinct. Angular tubercle present. Parietal tooth present (in
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<typeStatus id="54B38840FFB3FFD29C6500877BB3F8D4" box="[626,728,1776,1800]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
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), or absent (in all
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<typeStatus id="54B38840FFB3FFD29DB500877D78F8D4" box="[930,1043,1776,1800]" pageId="22" pageNumber="23" type="paratype">paratypes</typeStatus>
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). Columellar margin reflexed; with one prominent tooth expanding inward but nearly invisible from aperture. Umbilicus a narrow slit. Shell, including upper whorls, uniformly corneous-brown; aperture white.
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</paragraph>
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<paragraph id="8BB736E2FFB3FFDC9EDE012B7B41FE13" blockId="22.[151,1436,1050,1980]" lastBlockId="24.[151,1437,151,607]" lastPageId="24" lastPageNumber="25" pageId="22" pageNumber="23">
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<emphasis id="B97CEAF0FFB3FFD29EDE012B7854F8A9" bold="true" box="[201,319,1884,1909]" pageId="22" pageNumber="23">Genitalia.</emphasis>
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Vas deferens long; of even thickness; entering epiphallus apically at an angle. Epiphallus long; cylindrical and of uniform thickness; forming a few loops. Epiphallic caecum apically blunt. Flagellum present; short; tubular; proximally not expanded; with pointed tip. Penis evenly thick; with pilasters fused at epiphallic pore forming velum; without V-shaped pilasters; the two thicker parallel pilasters distally connected. Penial caecum saclike; situated at boundary between epiphallus and penis. Penial verge absent. Penial appendix short; branched off from proximal portion of penis; divided into sections A-1+A-2, A-3 and A-4+A-5. A-1 short; without verge toward penial chamber; with longitudinal pilasters. A-2 internally with longitudinal pilasters. A-1 and A-2 fused. A-2 near A-3 internally with a ring of papillae. A-2 and A-3 not fused. A-3 opening into A-2 without papilla. Boundary between A-4 and A-5 indistinct. A-5 short; straight. Atrium with short atrial retractor. Free oviduct short; subequal to vagina in length. Bursa copulatrix duct long; proximally twisted. Bursa copulatrix well defined. Diverticle longer than reservoir; unexpanded. Bursa copulatrix and diverticle distinguishable. Diverticle diverging from bursa duct just proximally from the bursa.
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</paragraph>
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<caption id="DF77666AFFB2FFD39E80012A7C61F84B" httpUri="https://zenodo.org/record/199995/files/figure.png" pageId="23" pageNumber="24" targetBox="[185,1399,264,1846]" targetPageId="23">
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||
<paragraph id="8BB736E2FFB2FFD39E80012A7C61F84B" blockId="23.[151,1436,1885,1944]" pageId="23" pageNumber="24">
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||
<emphasis id="B97CEAF0FFB2FFD39E80012A7844F8AA" bold="true" box="[151,303,1885,1910]" pageId="23" pageNumber="24">FIGURE 13.</emphasis>
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<taxonomicName id="4C084D61FFB2FFD39F22012878B3F8AA" box="[309,472,1887,1910]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="yuxu" status="sp. nov.">
|
||
<emphasis id="B97CEAF0FFB2FFD39F22012878B3F8AA" box="[309,472,1887,1910]" italics="true" pageId="23" pageNumber="24">Pupopsis yuxu</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A24F578BFFB2FFD39FC801297B52F8AA" box="[479,569,1886,1910]" pageId="23" pageNumber="24" rank="species">
|
||
<emphasis id="B97CEAF0FFB2FFD39FC801297B52F8AA" bold="true" box="[479,569,1886,1910]" pageId="23" pageNumber="24">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, HBUMM-06089–specimen 2, paratype. A, genitalia, general view; B, distal part of penis, interior view. Arrows indicate the part that was pulled straight for preparing illustration.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB736E2FFBDFFDC9EDE07AA7A95FDCB" blockId="24.[151,1437,151,607]" pageId="24" pageNumber="25">
|
||
Measurement of genitalia. P: 1.1; Ep: 6.6; Fl: 0.4; VD: 5.7; Va: 0.9; FO: 0.7; BCD: 10.1; BC: 0.8; D: 4.1; A- 1+A-2: 0.6; A-3: 0.8; A-4+A-5: 3.6 (HBUMM-06089–specimen 2,
|
||
<typeStatus id="54B38840FFBDFFDC9D9B07887A99FDCB" box="[908,1010,511,535]" pageId="24" pageNumber="25" type="paratype">paratype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3126569FFBDFFDC9EDE04537C5FFD83" pageId="24" pageNumber="25" type="discussion">
|
||
<paragraph id="8BB736E2FFBDFFDC9EDE04537C5FFD83" blockId="24.[151,1437,151,607]" pageId="24" pageNumber="25">
|
||
<emphasis id="B97CEAF0FFBDFFDC9EDE04537856FDE0" bold="true" box="[201,317,548,572]" pageId="24" pageNumber="25">Remarks.</emphasis>
|
||
This species is quite distinct by being the smallest in the genus, by the combination of a columellar tooth and a weak or absent parietal tooth, and by two thick parallel pilasters in the distal part of the penis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |