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<document id="70712F8DEFAA46FA812754F506DC67E5" ID-CLB-Dataset="46921" ID-DOI="10.5281/zenodo.199982" ID-GBIF-Dataset="558b2fa1-879c-44e4-956d-9abd9c9a249a" ID-ISSN="1175-5326" ID-Zenodo-Dep="199982" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460351134285" checkinUser="plazi" docAuthor="Wu, Min &amp; Gao, Linhui" docDate="2010" docId="03A187F4FFAFFFC89E8003357B1BFB34" docLanguage="en" docName="zt02725p027.pdf" docOrigin="Zootaxa 2725" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pupopsis retrodens Martens 1879" docType="treatment" docVersion="10" lastPageNumber="13" masterDocId="FF98FF8CFFA5FFC49E170677796BFFDC" masterDocTitle="A review of the genus Pupopsis Gredler, 1898 (Gastropoda: Stylommatophora: Enidae), with the descriptions of eight new species from China" masterLastPageNumber="27" masterPageNumber="1" pageNumber="11" updateTime="1698293683227" updateUser="ExternalLinkService">
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<mods:title id="40EBB90C01BD9FD24CD21FE021FF36C9">A review of the genus Pupopsis Gredler, 1898 (Gastropoda: Stylommatophora: Enidae), with the descriptions of eight new species from China</mods:title>
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<mods:namePart id="412F1229E5B9CF1EEB9FB6F0FF1F038C">Wu, Min</mods:namePart>
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<mods:namePart id="0B7BE6C16E49D1E719ABF5CE5C8408E3">Gao, Linhui</mods:namePart>
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<mods:date id="544F599A055A16F8C648C40EBF04BD3A">2010</mods:date>
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<treatment id="03A187F4FFAFFFC89E8003357B1BFB34" ID-DOI="http://doi.org/10.5281/zenodo.5625380" ID-GBIF-Taxon="119407886" ID-Zenodo-Dep="5625380" LSID="urn:lsid:plazi:treatment:03A187F4FFAFFFC89E8003357B1BFB34" httpUri="http://treatment.plazi.org/id/03A187F4FFAFFFC89E8003357B1BFB34" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
<subSubSection id="C3126569FFAFFFCE9E8003357B03FA80" box="[151,616,1346,1372]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BB736E2FFAFFFCE9E8003357B03FA80" blockId="10.[151,616,1346,1405]" box="[151,616,1346,1372]" pageId="10" pageNumber="11">
<heading id="D0FF818EFFAFFFCE9E8003357B03FA80" bold="true" box="[151,616,1346,1372]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="B97CEAF0FFAFFFCE9E8003357B03FA80" bold="true" box="[151,616,1346,1372]" pageId="10" pageNumber="11">
5.
<taxonomicName id="4C084D61FFAFFFCE9EA303357B03FA80" ID-CoL="7SX86" authority="Martens, 1879" authorityName="Martens" authorityYear="1879" box="[180,616,1346,1372]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="retrodens">
<emphasis id="B97CEAF0FFAFFFCE9EA3033578F0FA80" bold="true" box="[180,411,1346,1372]" italics="true" pageId="10" pageNumber="11">Pupopsis retrodens</emphasis>
(
<bibRefCitation id="EF994B13FFAFFFCE9FBD03357B0BFA80" author="Martens" box="[426,608,1346,1372]" pageId="10" pageNumber="11" refString="Martens, E. von (1879) Mittelasiatische Land- und Susswasserschnecken. Sitzungs-Berichte der Gesellschaft Naturforschender Freunde zu Berlin, 1879, 122 - 126." type="journal article" year="1879">Martens, 1879</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3126569FFAFFFCE9E8003127B56FAA1" box="[151,573,1381,1405]" pageId="10" pageNumber="11" type="description">
<paragraph id="8BB736E2FFAFFFCE9E8003127B56FAA1" blockId="10.[151,616,1346,1405]" box="[151,573,1381,1405]" pageId="10" pageNumber="11">
<figureCitation id="13332A67FFAFFFCE9E8003127988FAA1" box="[151,227,1381,1405]" captionStart="FIGURE 1" captionStartId="4.[151,255,1432,1456]" captionTargetBox="[151,1430,346,1400]" captionTargetId="figure@4.[151,1436,303,1411]" captionTargetPageId="4" captionText="FIGURE 1. Distribution map of the genus Pupopsis. Red dot, Pupopsis dissociabilis (Sturany, 1900); blue dot, P. retrodens (Martens, 1879); blue square, P. maoxian sp. nov.; red square, P. rhodostoma sp. nov.; green dot, P. subpupopsis sp. nov.; green square, P. yengiawat sp. nov. Two grey dots each with two or five species as indicated in the boxes on the map. The species P. paraplesius (Sturany, 1900), P. polystreptus (Sturany, 1900) and P. zilchi sp. nov. are not mapped due to no precise locality information available." httpUri="https://zenodo.org/record/199983/files/figure.png" pageId="10" pageNumber="11">Figs. 1</figureCitation>
,
<figureCitation id="13332A67FFAFFFCE9EE603127994FAA1" box="[241,255,1381,1405]" captionStart="FIGURE 3" captionStartId="6.[151,255,1784,1808]" captionTargetBox="[172,1419,194,1772]" captionTargetId="figure@6.[172,1420,194,1774]" captionTargetPageId="6" captionText="FIGURE 3. Apertural view of Pupopsis species. A, Pupopsis pupopsis (Gredler, 1898), SMF- 42076; B, Pupopsis retrodens (Martens, 1879), HBUMM- 06460 specimen 1; C-D, Pupopsis torquilla (Möllendorff, 1901). C, lectotype, SMF- 42082; D, HBUMM- 05584 specimen 1; E, Pupopsis hendan sp. nov., holotype, HBUMM- 05592 specimen 2; F, Pupopsis maoxian sp. nov., holotype, HBUMM- 01023 specimen 1; G, Pupopsis rhodostoma sp. nov., holotype, HBUMM- 05872 specimen 2; H, Pupopsis subpupopsis sp. nov., holotype, HBUMM- 05676 specimen 1; I, Pupopsis subtorquilla sp. nov. A, holotype, HBUMM- 05429 specimen 2; J, Pupopsis yengiawat sp. nov., holotype, HBUMM- 04439 specimen 1; K, Pupopsis yuxu sp. nov., Holotype, HBUMM- 06089 specimen 3; L, Pupopsis zilchi sp. nov., holotype, SMF- 334777." httpUri="https://zenodo.org/record/199985/files/figure.png" pageId="10" pageNumber="11">3</figureCitation>
B, 4F4G, 5B, 7;
<tableCitation id="C68A0359FFAFFFCE9FD403127B48FAA1" box="[451,547,1381,1405]" captionStart="TABLE 2" captionStartId="3.[151,239,617,641]" captionTargetBox="[151,1435,700,1395]" captionTargetPageId="3" captionText="TABLE 2. Pupopsis species and their conchological measurements (data from Sturany, 1900; Möllendorff, 1901; Kobelt, 1902 and this study; *, calculated from the published data; /, data unavailable)." httpUri="http://table.plazi.org/id/DF77666AFFA6FFC79E80041E7D5DFD7C" pageId="10" pageNumber="11" tableUuid="DF77666AFFA6FFC79E80041E7D5DFD7C">Tables 2</tableCitation>
,
<tableCitation id="C68A0359FFAFFFCE9C3803127B56FAA1" box="[559,573,1381,1405]" captionStart="TABLE 3" captionStartId="10.[151,239,908,932]" captionText="TABLE 3. Comparison of genital organs among Pupopsis species. + present / yes; - absent / no." pageId="10" pageNumber="11">3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3126569FFAFFFCE9E8003DA7BE7F980" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="8BB736E2FFAFFFCE9E8003DA7B5DFA1F" blockId="10.[151,940,1453,1629]" box="[151,566,1453,1475]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB1811CCFFAFFFCE9E8003DA7B5DFA1F" box="[151,566,1453,1475]" pageId="10" pageNumber="11">
<taxonomicName id="4C084D61FFAFFFCE9E8003DA7B59FA1F" ID-CoL="7QFLT" authority="Martens, 1879: 126" authorityName="Martens" authorityPageNumber="126" authorityYear="1879" box="[151,562,1453,1475]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="retrodens">
<emphasis id="B97CEAF0FFAFFFCE9E8003DA7808FA1F" box="[151,355,1453,1475]" italics="true" pageId="10" pageNumber="11">Buliminus retrodens</emphasis>
<treatmentCitation id="0AA910F3FFAFFFCE9F7E03D97B59FA1F" author="Martens" box="[361,562,1453,1475]" page="126" pageId="10" pageNumber="11" year="1879">
<bibRefCitation id="EF994B13FFAFFFCE9F7E03D97894FA1F" author="Martens" box="[361,511,1454,1475]" pageId="10" pageNumber="11" refString="Martens, E. von (1879) Mittelasiatische Land- und Susswasserschnecken. Sitzungs-Berichte der Gesellschaft Naturforschender Freunde zu Berlin, 1879, 122 - 126." type="journal article" year="1879">Martens, 1879</bibRefCitation>
: 126
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736E2FFAFFFCE9E8003BB7B8DF9DC" blockId="10.[151,940,1453,1629]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB1811CCFFAFFFCE9E8003BB7B8DF9DC" pageId="10" pageNumber="11">
<emphasis id="B97CEAF0FFAFFFCE9E8003BB7812FA3E" box="[151,377,1484,1506]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C084D61FFAFFFCE9E8003BB7995FA3E" ID-CoL="7NVC2" box="[151,254,1484,1506]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">Buliminus</taxonomicName>
retrodens—
</emphasis>
<treatmentCitation id="0AA910F3FFAFFFCE9F6E03BA7B5EFA3D" author="Martens" box="[377,565,1484,1506]" page="30" pageId="10" pageNumber="11" year="1881">
<bibRefCitation id="EF994B13FFAFFFCE9F6E03BA7B64FA3D" author="Martens" box="[377,527,1484,1506]" pageId="10" pageNumber="11" refString="Martens, E. von (1881) Conchologische Mittheilungen als Fortsetzung der Novitates conchologicae, 1, 1 - 101, pls. 1 - 18." type="journal article" year="1881">Martens, 1881</bibRefCitation>
: 30
</treatmentCitation>
, pl. 6, figs. 15-18.
<taxonomicName id="4C084D61FFAFFFCE9E80039D7808F9DC" box="[151,355,1514,1536]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="retrodens">
<emphasis id="B97CEAF0FFAFFFCE9E80039D7808F9DC" box="[151,355,1514,1536]" italics="true" pageId="10" pageNumber="11">Buliminus retrodens</emphasis>
</taxonomicName>
<treatmentCitation id="0AA910F3FFAFFFCE9F69039C7B40F9DC" author="Martens" box="[382,555,1514,1536]" page="26" pageId="10" pageNumber="11" year="1882">
<bibRefCitation id="EF994B13FFAFFFCE9F69039C7B6DF9DC" author="Martens" box="[382,518,1515,1536]" pageId="10" pageNumber="11" refString="Martens, E. von (1882) Uber Centralasiatische Mollusken. Memoires de l'Academie Imperiale des Sciences de St. - Petersbourg, 30, 1 - 65, pls. 1 - 5." type="journal article" year="1882">Martens1882</bibRefCitation>
: 26
</treatmentCitation>
, pl. 3, figs. 10, 11.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736E2FFAFFFCE9E80007E7BE7F980" blockId="10.[151,940,1453,1629]" pageId="10" pageNumber="11">
<taxonomicName id="4C084D61FFAFFFCE9E80007E788CF9C3" box="[151,487,1545,1567]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="retrodens" subGenus="Chondrula">
<emphasis id="B97CEAF0FFAFFFCE9E80007E7995F9C3" box="[151,254,1545,1567]" italics="true" pageId="10" pageNumber="11">Buliminus</emphasis>
(
<emphasis id="B97CEAF0FFAFFFCE9F1B007E7811F9C3" box="[268,378,1545,1567]" italics="true" pageId="10" pageNumber="11">Chondrula</emphasis>
)
<emphasis id="B97CEAF0FFAFFFCE9F9E007E788CF9C3" box="[393,487,1545,1567]" italics="true" pageId="10" pageNumber="11">retrodens</emphasis>
</taxonomicName>
—Kobelt, 1889: 44, pl. 101, fig. 587.
<taxonomicName id="4C084D61FFAFFFCE9E80005F7B65F9E2" box="[151,526,1576,1598]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="retrodens" subGenus="Chondrulopsis">
<emphasis id="B97CEAF0FFAFFFCE9E80005F7995F9E2" box="[151,254,1576,1598]" italics="true" pageId="10" pageNumber="11">Buliminus</emphasis>
(
<emphasis id="B97CEAF0FFAFFFCE9F1B005F78CAF9E2" box="[268,417,1576,1598]" italics="true" pageId="10" pageNumber="11">Chondrulopsis</emphasis>
)
<emphasis id="B97CEAF0FFAFFFCE9FA7005F7B65F9E2" box="[432,526,1576,1598]" italics="true" pageId="10" pageNumber="11">retrodens</emphasis>
</taxonomicName>
—Kobelt, 1902: 514, pl. 82, figs. 1920.
<emphasis id="B97CEAF0FFAFFFCE9E80003F7804F981" box="[151,367,1607,1629]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C084D61FFAFFFCE9E80003F799FF981" box="[151,244,1608,1629]" class="Gastropoda" family="Enidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="20" phylum="Mollusca" rank="species" species="pupopsis">Pupopsis</taxonomicName>
retrodens—
</emphasis>
<bibRefCitation id="EF994B13FFAFFFCE9F78003078B2F980" author="Yen" box="[367,473,1607,1628]" pageId="10" pageNumber="11" refString="Yen, T. C. (1939) Die chinesischen Land- und Sußwasser-Gastropoden des Natur-Museums Senckenberg. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 444, 1 - 234, pls. 1 - 16." type="journal article" year="1939">Yen, 1939</bibRefCitation>
: 90, pl. 8, fig. 42.
</paragraph>
</subSubSection>
<subSubSection id="C3126569FFAFFFCE9E8000FD78B9F839" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BB736E2FFAFFFCE9E8000FD7860F919" blockId="10.[151,1436,1673,2022]" pageId="10" pageNumber="11">
<emphasis id="B97CEAF0FFAFFFCE9E8000FD782DF97E" bold="true" box="[151,326,1673,1698]" pageId="10" pageNumber="11">
<typeStatus id="54B38840FFAFFFCE9E8000FD79BBF97E" box="[151,208,1674,1698]" pageId="10" pageNumber="11">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54B38840FFAFFFCE9F4400FD78A3F97E" box="[339,456,1674,1698]" pageId="10" pageNumber="11" type="lectotype">Lectotype</typeStatus>
:
<collectionCode id="ED19AE27FFAFFFCE9FC200FD7B79F97D" box="[469,530,1674,1697]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="10" pageNumber="11">ZMB</collectionCode>
Moll.-34878a; 15
<typeStatus id="54B38840FFAFFFCE9CF800FE7AFDF97D" box="[751,918,1673,1697]" pageId="10" pageNumber="11" type="paralectotype">paralectotypes</typeStatus>
:
<collectionCode id="ED19AE27FFAFFFCE9DB300FD7A8BF97D" box="[932,992,1674,1697]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="10" pageNumber="11">ZMB</collectionCode>
Moll.-34878b (
<bibRefCitation id="EF994B13FFAFFFCE9A8F00FE7C0BF97D" author="Kilias" box="[1176,1376,1673,1697]" pageId="10" pageNumber="11" refString="Kilias, R. (1971) Die Typen und Typoide der Mollusken-Sammlung des Zoologischen Museums Berlin (6). II. Euthyneura, Stylommatophora, Orthurethra, Pupillacea (Enidae). Mitteilungen aus dem Zoologischen Museum in Berlin, 47, 215 - 238." type="journal article" year="1971">Kilias 1971: 230</bibRefCitation>
); not examined.
</paragraph>
<paragraph id="8BB736E2FFAFFFCE9EDE00A57D15F935" blockId="10.[151,1436,1673,2022]" box="[201,1150,1745,1770]" pageId="10" pageNumber="11">
<emphasis id="B97CEAF0FFAFFFCE9EDE00A5780EF936" bold="true" box="[201,357,1745,1770]" pageId="10" pageNumber="11">
<typeStatus id="54B38840FFAFFFCE9EDE00A57869F936" box="[201,258,1746,1770]" pageId="10" pageNumber="11">Type</typeStatus>
locality.
</emphasis>
Near the Ili river and at Kuldscha (= Gulja in new spelling), Xinjiang.
</paragraph>
<paragraph id="8BB736E2FFAFFFCE9EDE008278B9F839" blockId="10.[151,1436,1673,2022]" pageId="10" pageNumber="11">
<emphasis id="B97CEAF0FFAFFFCE9EDE008278D2F8D2" bold="true" box="[201,441,1781,1806]" pageId="10" pageNumber="11">Material examined.</emphasis>
HBUMM-06460: 6 fma, 7 fms,
<specimenCount id="9D0EFD6BFFAFFFCE9D4700827AADF8D1" box="[848,966,1781,1805]" pageId="10" pageNumber="11" type="juvenile">1 juvenile</specimenCount>
with animal; Gulja, 44°0ʹ N, 81°36ʹE, Xinjiang,
<date id="FFB61022FFAFFFCE9F1C016E78C5F8ED" box="[267,430,1817,1841]" pageId="10" pageNumber="11" value="2006-08-14">2006VIII14</date>
, leg. A. Kanatbaytegi. HBUMM-05208: 18 fms, Gulja Forestry Centre,
<quantity id="4CF09B07FFAFFFCE9AEA016E7C3DF8ED" box="[1277,1366,1817,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.736" pageId="10" pageNumber="11" unit="m" value="1736.0">1736 m</quantity>
a.s.l., 44°09ʹ23.1ʹ N, 81°44ʹ18.0ʹ E, Gulja, Xinjiang,
<date id="FFB61022FFAFFFCE9CB8014A7A2BF889" box="[687,832,1853,1877]" pageId="10" pageNumber="11" value="2002-06-26">2002VI26</date>
, leg. A. Kanatbaytegi. HBUMM-04345: 5 fms with dried animals,
<specimenCount id="9D0EFD6BFFAFFFCE9F54011678DEF8A5" box="[323,437,1889,1913]" pageId="10" pageNumber="11" type="juvenile">1 juvenile</specimenCount>
with animal, Qapqal Xibe Zizhixian, 43°48ʹN, 81°6ʹE, Xinjiang,
<date id="FFB61022FFAFFFCE9AB801167C54F8A5" box="[1199,1343,1889,1913]" pageId="10" pageNumber="11" value="2005-04-20">2005IV20</date>
, leg. A. Kanatbaytegi. HBUMM-05868: 5 fms, Guozigou, Gulja, 44°0ʹN, 81°36ʹE, Xinjiang,
<date id="FFB61022FFAFFFCE9AD201F27C47F841" box="[1221,1324,1925,1949]" bridgedPair="-" pageId="10" pageNumber="11" value="2002-06">2002VI</date>
, leg. A. Kanatbaytegi. HBUMM-04542: 2 fma, Ji'ergelang Village, 44°00ʹ24.5ʹ N, 81°31ʹ12.6ʹ E, Gulja, Xinjiang,
<date id="FFB61022FFAFFFCE9B4101DE79BCF839" pageId="10" pageNumber="11" value="2002-05-25">2002 V25</date>
, leg. A. Kanatbaytegi.
</paragraph>
</subSubSection>
<caption id="DF77666AFFAEFFCF9E80015C78B7F8BF" httpUri="https://zenodo.org/record/199989/files/figure.png" pageId="11" pageNumber="12" targetBox="[157,1433,213,1834]" targetPageId="11">
<paragraph id="8BB736E2FFAEFFCF9E80015C78B7F8BF" blockId="11.[151,1436,1835,1891]" pageId="11" pageNumber="12">
<emphasis id="B97CEAF0FFAEFFCF9E80015C7874F89F" bold="true" box="[151,287,1835,1859]" pageId="11" pageNumber="12">FIGURE 7.</emphasis>
<taxonomicName id="4C084D61FFAEFFCF9F3B015B7BD4F89F" authority="Martens, 1879" authorityName="Martens" authorityYear="1879" box="[300,703,1836,1859]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="retrodens">
<emphasis id="B97CEAF0FFAEFFCF9F3B015B7895F89F" box="[300,510,1836,1859]" italics="true" pageId="11" pageNumber="12">Pupopsis retrodens</emphasis>
(Martens, 1879)
</taxonomicName>
, HBUMM-04542specimen 1. General view of genitalia. Arrow indicates very short flagellum.
</paragraph>
</caption>
<subSubSection id="C3126569FFAEFFCF9EDE01F878B8F874" box="[201,467,1935,1960]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8BB736E2FFAEFFCF9EDE01F878B8F874" blockId="11.[151,1437,1935,2033]" box="[201,467,1935,1960]" pageId="11" pageNumber="12">
<emphasis id="B97CEAF0FFAEFFCF9EDE01F87809F874" bold="true" box="[201,354,1935,1960]" pageId="11" pageNumber="12">Distribution.</emphasis>
Xinjiang.
</paragraph>
</subSubSection>
<subSubSection id="C3126569FFAEFFC89EDE01C37A6EFB84" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="diagnosis">
<paragraph id="8BB736E2FFAEFFC89EDE01C37984FF2B" blockId="11.[151,1437,1935,2033]" lastBlockId="12.[151,1437,151,1256]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="B97CEAF0FFAEFFCF9EDE01C37828F811" bold="true" box="[201,323,1972,1997]" pageId="11" pageNumber="12">Diagnosis.</emphasis>
Shell with 6.3757.500 whorls. Height 8.511.1, diam. maj. 3.75.0. Parietal tooth present. Columella with one blunt tooth. Palatal ridge-like tooth indistinct, very obtuse. Epiphallus straight. Flagellum indistinct. Penial caecum absent. Penis distally with one V-shaped pilaster. A-2 near A-3 internally with a ring of papillae. A-3 opening into A-2 by a long needlelike papilla. Bursa copulatrix duct proximally straight. Diverticle present.
</paragraph>
<paragraph id="8BB736E2FFA9FFC89EDE07747AC7FDE0" blockId="12.[151,1437,151,1256]" pageId="12" pageNumber="13">
<emphasis id="B97CEAF0FFA9FFC89EDE0774788FFEC0" bold="true" box="[201,484,259,284]" pageId="12" pageNumber="13">Re-description of shell.</emphasis>
Elongate-ovate; apex not acuminate; dextral; thin-shelled; solid; semitranslucent; glossy; with 6.3756.8997.500 whorls. Whorls with fine spiral grooves; convex. Embryonic shell smooth; polished; with 1.5001.7111.875 whorls. Postnuclear whorls wrinkled. Suture simple. Last whorl almost straight in front; rounded at periphery. Shell 8.510.011.1 high; 3.74.45.0 in diam. maj. Height/diam. maj. ratio 2.05 2.272.55. Aperture ear-shaped; vertical; its insertions separated; 3.13.74.3 high, 2.42.83.3 wide. Ratio of shell height to aperture height 2.372.672.93. Palatal margin rounded. Peristome slightly expanded; sharp. Parietal callus indistinct; with angular tubercle. Parietal tooth present. Palatal wall with an indistinct and obtuse, ridge-like tooth. Columellar margin reflexed; with one blunt tooth. Umbilicus narrowly open. Shell, including upper whorls, uniformly corneous-brown; aperture of the same colour.
</paragraph>
<paragraph id="8BB736E2FFA9FFC89EDE04307816FBCC" blockId="12.[151,1437,151,1256]" pageId="12" pageNumber="13">
<emphasis id="B97CEAF0FFA9FFC89EDE04307854FDBC" bold="true" box="[201,319,583,608]" pageId="12" pageNumber="13">Genitalia.</emphasis>
Vas deferens long; of even thickness; entering epiphallus apically at some angle. Epiphallus moderately long; cylindrical and uniformly thick; straight. Epiphallic caecum apically blunt. Flagellum indistinct (
<figureCitation id="13332A67FFA9FFC89E8804F87983FD7B" box="[159,232,655,679]" captionStart="FIGURE 7" captionStartId="11.[151,255,1835,1859]" captionTargetBox="[157,1433,213,1834]" captionTargetId="figure@11.[150,1436,152,1858]" captionTargetPageId="11" captionText="FIGURE 7. Pupopsis retrodens (Martens, 1879), HBUMM- 04542 specimen 1. General view of genitalia. Arrow indicates very short flagellum." httpUri="https://zenodo.org/record/199989/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
, arrow). Penis evenly thick; without penial caecum; with terminal entrance of epiphallus; thin-walled. Pilasters fused at epiphallic pore. With one V-shaped pilaster; without papilla developed by fused distal pilasters forming V-shaped structure. V-shaped pilaster with proximal free end approaching at penial retractor insertion. Penial verge absent. Penial appendix short; branched off from proximal portion of penis; divided into sections A- 1+A-2, A-3 and A-4+A-5. A-1 long; without verge toward penial chamber. A-1 and A-2 fused; internally with longitudinal pilasters. A-2 near A-3 internally with a ring of papillae. A-2 and A-3 not fused. A-3 opening into A-2 by needlelike long papilla. Boundary between A-4 and A-5 indistinct. A-5 short; straight. Free oviduct moderately long; subequal to vagina in length. Vagina moderately long; not swollen; straight. Bursa copulatrix duct moderately long; proximally straight. Bursa copulatrix well defined. Diverticle present; longer than reservoir; unexpanded. Bursa copulatrix and diverticle distinguishable; diverticle diverging from bursa duct closer to reservoir than to ducts proximal end.
</paragraph>
<paragraph id="8BB736E2FFA9FFC89EDE026A7A6EFB84" blockId="12.[151,1437,151,1256]" pageId="12" pageNumber="13">Measurement of genitalia. P: 4; Ep: 9.4; VD: 12.6; Va: 2.1; FO: 2.3; BCD: 6.7; BC: 1.7; D: 4.7; A-1+A-2: 3.6; A-3: 1.0; A-4+A-5: 7.6 (HBUMM-06460specimen 1).</paragraph>
</subSubSection>
<subSubSection id="C3126569FFA9FFC89EDE02137B1BFB34" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BB736E2FFA9FFC89EDE02137B1BFB34" blockId="12.[151,1437,151,1256]" pageId="12" pageNumber="13">
<emphasis id="B97CEAF0FFA9FFC89EDE02137856FBA0" bold="true" box="[201,317,1124,1148]" pageId="12" pageNumber="13">Remarks.</emphasis>
Notably, the flagellum is indistinct or missing in all anatomically known enid species distributed in Xinjiang. In addition to
<emphasis id="B97CEAF0FFA9FFC89FB202FE7B4CFB7C" box="[421,551,1160,1184]" italics="true" pageId="12" pageNumber="13">P. re t ro d e n s</emphasis>
, the species with this feature are
<taxonomicName id="4C084D61FFA9FFC89DB602FE7DE9FB7C" box="[929,1154,1161,1184]" class="Gastropoda" family="Enidae" genus="Pupopsis" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="yengiawat" status="sp. nov.">
<emphasis id="B97CEAF0FFA9FFC89DB602FE7DE9FB7C" box="[929,1154,1161,1184]" italics="true" pageId="12" pageNumber="13">Pupopsis yengiawat</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24F578BFFA9FFC89A9E02FF7D88FB7C" box="[1161,1251,1160,1184]" pageId="12" pageNumber="13" rank="species">
<emphasis id="B97CEAF0FFA9FFC89A9E02FF7D88FB7C" bold="true" box="[1161,1251,1160,1184]" pageId="12" pageNumber="13">sp. nov.</emphasis>
</taxonomicNameLabel>
(see below) and
<taxonomicName id="4C084D61FFA9FFC89E8002DB7B21FB18" authority="Martens, 1881" authorityName="Martens" authorityYear="1881" box="[151,586,1196,1220]" class="Gastropoda" family="Enidae" genus="Subzebrinus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="labiellus">
<emphasis id="B97CEAF0FFA9FFC89E8002DB78E2FB18" box="[151,393,1196,1220]" italics="true" pageId="12" pageNumber="13">Subzebrinus labiellus</emphasis>
(
<bibRefCitation id="EF994B13FFA9FFC89F8E02DB7B2AFB18" author="Martens" box="[409,577,1196,1220]" pageId="12" pageNumber="13" refString="Martens, E. von (1881) Conchologische Mittheilungen als Fortsetzung der Novitates conchologicae, 1, 1 - 101, pls. 1 - 18." type="journal article" year="1881">Martens, 1881</bibRefCitation>
)
</taxonomicName>
, which is widely distributed in SE
<collectingCountry id="F31F7672FFA9FFC89DCC02DB7D37FB18" box="[987,1116,1196,1220]" name="Kazakhstan" pageId="12" pageNumber="13">Kasakhstan</collectingCountry>
and neighbouring Xinjiang,
<collectingCountry id="F31F7672FFA9FFC89E8002A779B1FB34" box="[151,218,1232,1256]" name="China" pageId="12" pageNumber="13">China</collectingCountry>
(
<bibRefCitation id="EF994B13FFA9FFC89EFF02A77B0FFB34" author="Schileyko" box="[232,612,1232,1256]" pageId="12" pageNumber="13" refString="Schileyko, A. A. (1998) Treatise on recent terrestrial pulmonate molluscs. Part. 2. Gastrocoptidae, Hypselostomatidae, Vertiginidae, Truncatellinidae, Pachnodidae, Enidae, Sagdidae. Ruthenica, Supplement 2, 129 - 261." type="journal article" year="1998">Schileyko 1998; Wu unpublished</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>