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<document id="53CA5E7DBB1AC0F5C0A9336A9E8B7F55" ID-CLB-Dataset="21205" ID-DOI="10.5281/zenodo.3985368" ID-GBIF-Dataset="efec1c9a-9bf5-4599-bfd4-b932b9d34026" ID-ISSN="0253-116X" ID-Zenodo-Dep="3985368" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597412148234" checkinUser="valdenar" docAuthor="Assing, Volker" docDate="2018" docId="03A187CAFFC294291FEAFB6593A9FC51" docLanguage="en" docName="Linzer.biol.Beitr.50.1.0149-0182.pdf.imf" docOrigin="Linzer biologische Beiträge 50 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Quedius (Raphirus) josue" docType="treatment" docVersion="8" lastPageNumber="177" masterDocId="FF98FFB2FFD894351F58FFC89247FFBA" masterDocTitle="On the taxonomy and zoogeography of some West Palaearctic Quedius species, with a focus on the East Mediterranean and the species allied to Quedius umbrinus and Q. nivicola (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="182" masterPageNumber="149" pageNumber="175" updateTime="1698844552053" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="052BCA974A9C108CBDABC4B83EB1B867" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="8281122F444B3419B9A5BE68E7430F53">On the taxonomy and zoogeography of some West Palaearctic Quedius species, with a focus on the East Mediterranean and the species allied to Quedius umbrinus and Q. nivicola (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="276ABC9D1A114DE46EDADD730326EF3B">Assing, Volker</mods:namePart>
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<mods:title id="6BD279A7D85CF4056CD6F887B6187D15">Linzer biologische Beiträge</mods:title>
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<mods:date id="9123C0731DD5D52B06A9E47E2A428D0F">2018</mods:date>
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<mods:number id="8973C393942039A786678C7791278FC1">2018-07-27</mods:number>
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<mods:number id="BBD5FDFB299563F73EA92CFBF392C3A9">50</mods:number>
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<mods:identifier id="41A3A20B748E7A1488980A4BD1FAC264" type="CLB-Dataset">21205</mods:identifier>
<mods:identifier id="972B95E836EBCE9580C3D74B92F3BED7" type="DOI">10.5281/zenodo.3985368</mods:identifier>
<mods:identifier id="6C9CC92D0772622F2E047C29318ED9A0" type="GBIF-Dataset">efec1c9a-9bf5-4599-bfd4-b932b9d34026</mods:identifier>
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<treatment id="03A187CAFFC294291FEAFB6593A9FC51" ID-DOI="http://doi.org/10.5281/zenodo.4334287" ID-GBIF-Taxon="167288476" ID-Zenodo-Dep="4334287" LSID="urn:lsid:plazi:treatment:03A187CAFFC294291FEAFB6593A9FC51" httpUri="http://treatment.plazi.org/id/03A187CAFFC294291FEAFB6593A9FC51" lastPageId="28" lastPageNumber="177" pageId="26" pageNumber="175">
<subSubSection id="C3126557FFC2942F1FEAFB65916CFB7E" box="[178,811,1197,1220]" pageId="26" pageNumber="175" type="nomenclature">
<paragraph id="8BB736DCFFC2942F1FEAFB65916CFB7E" blockId="26.[178,811,1197,1220]" box="[178,811,1197,1220]" pageId="26" pageNumber="175">
<heading id="D0FF81B0FFC2942F1FEAFB65916CFB7E" box="[178,811,1197,1220]" fontSize="8" level="2" pageId="26" pageNumber="175" reason="2">
<emphasis id="B97CEACEFFC2942F1FEAFB65905DFB78" bold="true" box="[178,538,1197,1219]" pageId="26" pageNumber="175">
<taxonomicName id="4C084D5FFFC2942F1FEAFB6593F8FB79" ID-CoL="4R366" authority="SAULCY, 1865" box="[178,447,1197,1219]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="175" phylum="Arthropoda" rank="species" species="josue" subGenus="Raphirus">
<emphasis id="B97CEACEFFC2942F1FEAFB65934EFB79" bold="true" box="[178,265,1197,1219]" italics="true" pageId="26" pageNumber="175">Quedius</emphasis>
(
<emphasis id="B97CEACEFFC2942F1E40FB65933EFB79" bold="true" box="[280,377,1197,1219]" italics="true" pageId="26" pageNumber="175">Raphirus</emphasis>
)
<emphasis id="B97CEACEFFC2942F1ED0FB6693F8FB79" bold="true" box="[392,447,1198,1219]" italics="true" pageId="26" pageNumber="175">josue</emphasis>
</taxonomicName>
<bibRefCitation id="EF994B2DFFC2942F1E9DFB65905DFB78" author="SAULCY F" box="[453,538,1197,1219]" pageId="26" pageNumber="175" pagination="629 - 660" refId="ref21389" refString="SAULCY F. H. C. DE (1865): Descriptions des especes nouvelle de coleopteres recueillies en Syrie, en Egypte et en Palestine, pendant les mois d'octobre 1863 a janvier 1864, par M. de Saulcy, senateur, membre de l'institut. 2 e partie. - Annales de la Societe de Entomologique de France (4) 4: [1864]: 629 - 660." type="journal article" year="1865">SAULCY</bibRefCitation>
</emphasis>
,
<emphasis id="B97CEACEFFC2942F1D7EFB66901EFB79" bold="true" box="[550,601,1198,1219]" pageId="26" pageNumber="175">1865</emphasis>
(
<figureCitation id="13332A59FFC2942F1D3EFB669090FB7E" box="[614,727,1198,1220]" captionStart-0="Figs 53-62" captionStart-1="Figs 63-68" captionStartId-0="22.[178,218,1506,1528]" captionStartId-1="23.[178,218,906,928]" captionTargetBox-0="[179,1076,224,1480]" captionTargetBox-1="[183,1080,224,879]" captionTargetId-0="figure@22.[178,1085,212,1497]" captionTargetId-1="figure@23.[178,1085,212,897]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="Figs 53-62: Quedius josue from Lebanon (53), Israel (54-56), and Cyprus (57-62): (53-54, 57, 60) aedeagus in lateral view; (55, 58, 61) median lobe of aedeagus in ventral view; (56, 59, 62) paramere. Scale bar: 0.5 mm." captionText-1="Figs 63-68: Quedius josue from Syria (63-65) and Q. petraensis, holotype (66-68; figures in transparent light): (63) median lobe of aedeagus in lateral view; (64, 67) median lobe of aedeagus in ventral view; (65, 68) paramere; (66) aedeagus in lateral view. Scale bar: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3985386" figureDoi-1="http://doi.org/10.5281/zenodo.3985388" httpUri-0="https://zenodo.org/record/3985386/files/figure.png" httpUri-1="https://zenodo.org/record/3985388/files/figure.png" pageId="26" pageNumber="175">Figs 53-65</figureCitation>
,
<figureCitation id="13332A59FFC2942F1DB9FB669163FB7E" box="[737,804,1198,1220]" captionStart="Map 4" captionStartId="27.[178,223,871,892]" captionTargetBox="[178,1085,212,861]" captionTargetId="figure@27.[178,1085,212,861]" captionTargetPageId="27" captionText="Map 4: Distributions of Quedius nivicola (white circles; revised records), Q. ortrudae (black circles; revised records), and Q. josue (black triangles: revised records; white triangles literature records) in the East Mediterranean." figureDoi="http://doi.org/10.5281/zenodo.3985390" httpUri="https://zenodo.org/record/3985390/files/figure.png" pageId="26" pageNumber="175">Map 4</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3126557FFC2942F1FEAFB11967AFA92" pageId="26" pageNumber="175" type="reference_group">
<paragraph id="8BB736DCFFC2942F1FEAFB1193B6FB55" blockId="26.[178,1086,1241,1541]" box="[178,497,1241,1263]" pageId="26" pageNumber="175">
<treatmentCitationGroup id="AB1811F2FFC2942F1FEAFB1193B6FB55" box="[178,497,1241,1263]" pageId="26" pageNumber="175">
<treatmentCitation id="0AA910CDFFC2942F1FEAFB1193AAFB55" author="SAULCY F" box="[178,493,1241,1263]" page="636" pageId="26" pageNumber="175" year="1865">
<taxonomicName id="4C084D5FFFC2942F1FEAFB1193AAFB55" ID-CoL="4R366" authority="SAULCY, 1865: 636" authorityName="Saulcy" authorityYear="1865" box="[178,493,1241,1263]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="species" species="josue">
<emphasis id="B97CEACEFFC2942F1FEAFB119374FB55" box="[178,307,1241,1263]" italics="true" pageId="26" pageNumber="175">Quedius josue</emphasis>
<bibRefCitation id="EF994B2DFFC2942F1E61FB1193AAFB55" author="SAULCY F" box="[313,493,1241,1263]" pageId="26" pageNumber="175" pagination="629 - 660" refId="ref21389" refString="SAULCY F. H. C. DE (1865): Descriptions des especes nouvelle de coleopteres recueillies en Syrie, en Egypte et en Palestine, pendant les mois d'octobre 1863 a janvier 1864, par M. de Saulcy, senateur, membre de l'institut. 2 e partie. - Annales de la Societe de Entomologique de France (4) 4: [1864]: 629 - 660." type="journal article" year="1865">SAULCY, 1865: 636</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFC2942F1FEAFB3B9160FAB3" blockId="26.[178,1086,1241,1541]" box="[178,807,1267,1289]" pageId="26" pageNumber="175">
<treatmentCitationGroup id="AB1811F2FFC2942F1FEAFB3B9160FAB3" box="[178,807,1267,1289]" pageId="26" pageNumber="175">
<emphasis id="B97CEACEFFC2942F1FEAFB3B9314FAB2" box="[178,339,1267,1288]" italics="true" pageId="26" pageNumber="175">
<taxonomicName id="4C084D5FFFC2942F1FEAFB3B92B9FAB2" ID-CoL="8VS2M" box="[178,254,1267,1288]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
troodites
</emphasis>
<treatmentCitation id="0AA910CDFFC2942F1E01FB3C93B6FAB3" author="FAGEL G" box="[345,497,1267,1289]" page="7" pageId="26" pageNumber="175" year="1968">
<bibRefCitation id="EF994B2DFFC2942F1E01FB3C93B6FAB3" author="FAGEL G" box="[345,497,1267,1289]" pageId="26" pageNumber="175" pagination="1 - 13" refId="ref20496" refString="FAGEL G. (1968 a): Contribution a la connaissance des Staphylinidae CI. - Remarques diverses sur quelques Quedius de Mediterranee orientale. - B ulletin Institut Royale des Sciences Naturelles de Belgique 44 (12): 1 - 13." type="journal article" year="1968">FAGEL, 1968a: 7</bibRefCitation>
</treatmentCitation>
ff.; synonymy by ASSING (2004).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFC2942F1FEAFADB967AFA92" blockId="26.[178,1086,1241,1541]" box="[178,1085,1298,1320]" pageId="26" pageNumber="175">
T y p e m a t e r i a l e x a m i n e d:
<typeStatus id="54B3887EFFC2942F1D03FADB90EFFA92" box="[603,680,1299,1320]" pageId="26" pageNumber="175" type="neotype">Neotype</typeStatus>
<specimenCount id="9D0EFD55FFC2942F1DEBFADB9081FA92" box="[691,710,1299,1320]" pageId="26" pageNumber="175" type="male">♂</specimenCount>
, present designation: &quot;
<collectingCountry id="F31F764CFFC2942F1CFBFADA91B1FA9D" box="[931,1014,1298,1319]" name="Israel" pageId="26" pageNumber="175">ISRAEL</collectingCountry>
, Upper
</paragraph>
</subSubSection>
<subSubSection id="C3126557FFC294291FEAFAE1906EFCD2" lastPageId="28" lastPageNumber="177" pageId="26" pageNumber="175" type="description">
<paragraph id="8BB736DCFFC2942F1FEAFAE1967AFA85" blockId="26.[178,1086,1241,1541]" box="[178,1085,1321,1343]" pageId="26" pageNumber="175">
Galilee, near Meron, &quot;Appleplot&quot;,
<date id="FFB6101CFFC2942F1EAAFAE190CFFA85" box="[498,648,1321,1343]" pageId="26" pageNumber="175" value="2007-05-15" valueMax="2007-06-05" valueMin="2007-05-15">15.V.-5.VI.2007</date>
(31), N33.00.37 E35.24.13, leg. J. Buse, flight
</paragraph>
<paragraph id="8BB736DCFFC2942F1FEAFA889101FAEC" blockId="26.[178,1086,1241,1541]" box="[178,838,1343,1366]" pageId="26" pageNumber="175">
trap /
<typeStatus id="54B3887EFFC2942F1FBFFA889307FAEF" box="[231,320,1344,1365]" pageId="26" pageNumber="175" type="neotype">Neotypus</typeStatus>
<specimenCount id="9D0EFD55FFC2942F1E18FA889308FAEF" box="[320,335,1344,1365]" pageId="26" pageNumber="175" type="male">♂</specimenCount>
<taxonomicName id="4C084D5FFFC2942F1E0CFAF7905CFAEF" authority="Saulcy" authorityName="Saulcy" box="[340,539,1343,1365]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="175" phylum="Arthropoda" rank="species" species="josue">
<emphasis id="B97CEACEFFC2942F1E0CFAF79391FAEF" box="[340,470,1343,1365]" italics="true" pageId="26" pageNumber="175">Quedius josue</emphasis>
Saulcy
</taxonomicName>
, desig. V.
<bibRefCitation id="EF994B2DFFC2942F1D23FAF790A8FAEF" author="ASSING V &amp; SCHULKE M &amp; BRACHAT V" box="[635,751,1343,1365]" pageId="26" pageNumber="175" pagination="1" refId="ref19985" refString="ASSING V., SCHULKE M., BRACHAT V. &amp; H. MEYBOHM (in press): On the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera). - Contributions to En tomology 68 (1) (2018)." type="journal article" year="2018">Assing 2018</bibRefCitation>
&quot; (MNB).
</paragraph>
<paragraph id="8BB736DCFFC2942F1FEAFA97919DF9BF" blockId="26.[178,1086,1241,1541]" pageId="26" pageNumber="175">
<materialsCitation id="3B603C81FFC2942F1FEAFA979028FA1F" ID-GBIF-Occurrence="2846796351" collectingDate="1996-04-28" collectingDateMax="2016-05" collectingDateMin="1996-04-28" collectorName="Sprick &amp; Reuter" country="Syria" county="Samaan" elevation="900" latitude="33.483334" location="Simeons Monastery" longLatPrecision="1204" longitude="35.8" municipality="Qualaat" pageId="26" pageNumber="175" specimenCount="6" specimenCount-female="1" specimenCount-male="5">
A d d i t i o n a l m a t e r i a l e x a m i n e d:
<collectingCountry id="F31F764CFFC2942F1DEEFA9790A0FACF" box="[694,743,1375,1397]" name="Syria" pageId="26" pageNumber="175">Syria</collectingCountry>
:
<specimenCount id="9D0EFD55FFC2942F1DA9FA979159FACE" box="[753,798,1375,1396]" pageId="26" pageNumber="175" type="male">3♂♂</specimenCount>
,
<specimenCount id="9D0EFD55FFC2942F1C71FA979102FACD" box="[809,837,1375,1399]" pageId="26" pageNumber="175" type="female">1♀</specimenCount>
,
<collectingCounty id="62D64E50FFC2942F1C08FA9791D0FACE" box="[848,919,1375,1396]" pageId="26" pageNumber="175">Samaan</collectingCounty>
,
<collectingMunicipality id="6BD3ACA6FFC2942F1CFAFA9791AFFACF" box="[930,1000,1375,1397]" pageId="26" pageNumber="175">Qualaat</collectingMunicipality>
,
<location id="8ED76007FFC2942F1CAAFA979379FA37" LSID="urn:lsid:plazi:treatment:03A187CAFFC294291FEAFB6593A9FC51:8ED76007FFC2942F1CAAFA979379FA37" country="Syria" county="Samaan" latitude="33.483334" longLatPrecision="1204" longitude="35.8" municipality="Qualaat" name="Simeons Monastery" pageId="26" pageNumber="175">Simeons Monastery</location>
,
<date id="FFB6101CFFC2942F1E1FFABF93EBFA37" box="[327,428,1399,1421]" pageId="26" pageNumber="175" value="1996-04-28">
<collectingDate id="EFF2E9F4FFC2942F1E1FFABF93EBFA37" box="[327,428,1399,1421]" pageId="26" pageNumber="175" value="1996-04-28">28.IV.1996</collectingDate>
</date>
, leg.
<collectorName id="26FD530AFFC2942F1E82FABF9054FA37" box="[474,531,1399,1421]" pageId="26" pageNumber="175">Sprick</collectorName>
(cAss).
<collectingCountry id="F31F764CFFC2942F1D04FABF90EBFA37" box="[604,684,1399,1421]" name="Lebanon" pageId="26" pageNumber="175">Lebanon</collectingCountry>
:
<specimenCount id="9D0EFD55FFC2942F1DECFABF90A5FA36" box="[692,738,1399,1420]" pageId="26" pageNumber="175" type="male">2♂♂</specimenCount>
,
<location id="8ED76007FFC2942F1DB3FABF917FFA37" LSID="urn:lsid:plazi:treatment:03A187CAFFC294291FEAFB6593A9FC51:8ED76007FFC2942F1DB3FABF917FFA37" box="[747,824,1399,1421]" country="Syria" county="Samaan" latitude="33.483334" longLatPrecision="1204" longitude="35.8" municipality="Qualaat" name="Rachaya" pageId="26" pageNumber="175">Rachaya</location>
,
<location id="8ED76007FFC2942F1C19FAB091D0FA37" LSID="urn:lsid:plazi:treatment:03A187CAFFC294291FEAFB6593A9FC51:8ED76007FFC2942F1C19FAB091D0FA37" box="[833,919,1400,1421]" country="Syria" county="Samaan" latitude="33.483334" longLatPrecision="1204" longitude="35.8" municipality="Qualaat" name="Tannoura" pageId="26" pageNumber="175">Tannoura</location>
,
<geoCoordinate id="EE3C501BFFC2942F1CC7FABF91AEFA36" box="[927,1001,1399,1420]" degrees="33" direction="north" minutes="29" orientation="latitude" pageId="26" pageNumber="175" precision="925" value="33.483334">33°29'N</geoCoordinate>
,
<geoCoordinate id="EE3C501BFFC2942F1CAAFABF967EFA37" box="[1010,1081,1399,1421]" degrees="35" direction="east" minutes="48" orientation="longitude" pageId="26" pageNumber="175" precision="925" value="35.8">35°48'E</geoCoordinate>
,
<quantity id="4CF09B39FFC2942F1F87FA47935DFA1F" box="[223,282,1423,1445]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="26" pageNumber="175" unit="m" value="900.0">
<elevation id="0025D1EFFFC2942F1F87FA47935DFA1F" box="[223,282,1423,1445]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="26" pageNumber="175" unit="m" value="900.0">900 m</elevation>
</quantity>
, oak forest,
<date id="FFB6101CFFC2942F1EDEFA47938EFA1F" box="[390,457,1423,1445]" pageId="26" pageNumber="175" value="2016-05">
<collectingDate id="EFF2E9F4FFC2942F1EDEFA47938EFA1F" box="[390,457,1423,1445]" pageId="26" pageNumber="175" value="2016-05">V.2016</collectingDate>
</date>
, leg.
<collectorName id="26FD530AFFC2942F1EA1FA589074FA1F" box="[505,563,1424,1445]" pageId="26" pageNumber="175">Reuter</collectorName>
(cFel)
</materialsCitation>
;
<materialsCitation id="3B603C81FFC2942F1D21FA479037FA07" ID-GBIF-Occurrence="2846796396" collectingDate="2016-11-18" collectingDateMax="2016-12-16" collectingDateMin="2016-11-18" collectorName="Reuter" country="Syria" elevation="990" latitude="33.966667" location="Rayfoun" longLatPrecision="1202" longitude="35.7" pageId="26" pageNumber="175" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D0EFD55FFC2942F1D21FA4790D2FA1E" box="[633,661,1423,1444]" pageId="26" pageNumber="175" type="male">1♂</specimenCount>
,
<location id="8ED76007FFC2942F1DF8FA4790AAFA1F" LSID="urn:lsid:plazi:treatment:03A187CAFFC294291FEAFB6593A9FC51:8ED76007FFC2942F1DF8FA4790AAFA1F" box="[672,749,1423,1445]" country="Syria" latitude="33.966667" longLatPrecision="1202" longitude="35.7" name="Rayfoun" pageId="26" pageNumber="175">Rayfoun</location>
,
<geoCoordinate id="EE3C501BFFC2942F1DAFFA479106FA1F" box="[759,833,1423,1445]" degrees="33" direction="north" minutes="58" orientation="latitude" pageId="26" pageNumber="175" precision="925" value="33.966667">33°58'N</geoCoordinate>
,
<geoCoordinate id="EE3C501BFFC2942F1C13FA4791D4FA1F" box="[843,915,1423,1445]" degrees="35" direction="east" minutes="42" orientation="longitude" pageId="26" pageNumber="175" precision="925" value="35.7">35°42'E</geoCoordinate>
,
<quantity id="4CF09B39FFC2942F1CC4FA479191FA1F" box="[924,982,1423,1445]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="26" pageNumber="175" unit="m" value="990.0">
<elevation id="0025D1EFFFC2942F1CC4FA479191FA1F" box="[924,982,1423,1445]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="26" pageNumber="175" unit="m" value="990.0">990 m</elevation>
</quantity>
, mixed oak forest,
<date id="FFB6101CFFC2942F1E45FA6F938FFA07" box="[285,456,1447,1469]" pageId="26" pageNumber="175" value="2016-11-18" valueMax="2016-12-16" valueMin="2016-11-18">
<collectingDate id="EFF2E9F4FFC2942F1E45FA6F938FFA07" box="[285,456,1447,1469]" pageId="26" pageNumber="175" value="2016-11-18" valueMax="2016-12-16" valueMin="2016-11-18">18.XI.-16.XII.2016</collectingDate>
</date>
, leg.
<collectorName id="26FD530AFFC2942F1EA1FA609074FA07" box="[505,563,1448,1469]" pageId="26" pageNumber="175">Reuter</collectorName>
(cFel)
</materialsCitation>
;
<materialsCitation id="3B603C81FFC2942F1D22FA6F9192F9BF" ID-GBIF-Occurrence="2846796325" collectingDate="2009-04-30" collectingDateMax="2012-05-20" collectingDateMin="2009-04-30" collectionCode="MNB" collectorName="Reuter &amp; Buse &amp; Pavlicek &amp; Stavros &amp; Psokas &amp; Wolf" country="Israel" county="Fnaideq" elevation="1350" latitude="32.75" location="Mount Carmel" longLatPrecision="1209" longitude="35.016666" municipality="Djebel Qammouaa" pageId="26" pageNumber="175" specimenCount="8" specimenCount-female="1" specimenCount-male="1" stateProvince="Pafos">
<specimenCount id="9D0EFD55FFC2942F1D22FA6F90D1FA06" box="[634,662,1447,1468]" pageId="26" pageNumber="175" type="male">1♂</specimenCount>
,
<locationDeviation id="9A052FFEFFC2942F1DFAFA6F9172FA07" box="[674,821,1447,1469]" location="Tripoli" pageId="26" pageNumber="175">
<quantity id="4CF09B39FFC2942F1DFAFA6F909CFA07" box="[674,731,1447,1469]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="26" pageNumber="175" unit="km" value="28.0">28 km</quantity>
E Tripoli
</locationDeviation>
,
<collectingCounty id="62D64E50FFC2942F1C67FA6F91C0FA07" box="[831,903,1447,1469]" pageId="26" pageNumber="175">Fnaideq</collectingCounty>
,
<collectingMunicipality id="6BD3ACA6FFC2942F1CC9FA6F967DFA06" box="[913,1082,1447,1469]" pageId="26" pageNumber="175">Djebel Qammouaa</collectingMunicipality>
,
<quantity id="4CF09B39FFC2942F1F87FA77931FFA6F" box="[223,344,1471,1493]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="26" pageNumber="175" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">
<elevation id="0025D1EFFFC2942F1F87FA77931FFA6F" box="[223,344,1471,1493]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="26" pageNumber="175" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">1300-1400 m</elevation>
</quantity>
, coniferous forest,
<date id="FFB6101CFFC2942F1D5AFA779018FA6E" box="[514,607,1471,1492]" pageId="26" pageNumber="175" value="2012-05-20">
<collectingDate id="EFF2E9F4FFC2942F1D5AFA779018FA6E" box="[514,607,1471,1492]" pageId="26" pageNumber="175" value="2012-05-20">20.V.2012</collectingDate>
</date>
, leg.
<collectorName id="26FD530AFFC2942F1DC8FA08908DFA6F" box="[656,714,1472,1493]" pageId="26" pageNumber="175">Reuter</collectorName>
(cAss).
<collectingCountry id="F31F764CFFC2942F1C4EFA77910EFA6F" box="[790,841,1471,1493]" name="Israel" pageId="26" pageNumber="175">Israel</collectingCountry>
:
<specimenCount id="9D0EFD55FFC2942F1C0AFA779129FA6D" box="[850,878,1471,1495]" pageId="26" pageNumber="175" type="female">1♀</specimenCount>
, Haifa,
<location id="8ED76007FFC2942F1CEEFA08967DFA6F" LSID="urn:lsid:plazi:treatment:03A187CAFFC294291FEAFB6593A9FC51:8ED76007FFC2942F1CEEFA08967DFA6F" box="[950,1082,1471,1493]" country="Israel" county="Fnaideq" latitude="32.75" longLatPrecision="1209" longitude="35.016666" municipality="Djebel Qammouaa" name="Mount Carmel" pageId="26" pageNumber="175" stateProvince="Pafos">Mount Carmel</location>
,
<location id="8ED76007FFC2942F1F87FA1F9334FA57" LSID="urn:lsid:plazi:treatment:03A187CAFFC294291FEAFB6593A9FC51:8ED76007FFC2942F1F87FA1F9334FA57" box="[223,371,1495,1517]" country="Israel" county="Fnaideq" latitude="32.75" longLatPrecision="1209" longitude="35.016666" municipality="Djebel Qammouaa" name="Ya'ar Ha'arbaim" pageId="26" pageNumber="175" stateProvince="Pafos">Ya'ar Ha'arbaim</location>
,
<geoCoordinate id="EE3C501BFFC2942F1ED8FA1F938CFA57" box="[384,459,1495,1517]" degrees="32" direction="north" minutes="45" orientation="latitude" pageId="26" pageNumber="175" precision="925" value="32.75">32°45'N</geoCoordinate>
,
<geoCoordinate id="EE3C501BFFC2942F1E8FFA1F9059FA57" box="[471,542,1495,1517]" degrees="35" direction="east" minutes="01" orientation="longitude" pageId="26" pageNumber="175" precision="925" value="35.016666">35°01'E</geoCoordinate>
,
<date id="FFB6101CFFC2942F1D74FA1F90D5FA56" box="[556,658,1495,1516]" pageId="26" pageNumber="175" value="2009-04-30">
<collectingDate id="EFF2E9F4FFC2942F1D74FA1F90D5FA56" box="[556,658,1495,1516]" pageId="26" pageNumber="175" value="2009-04-30">30.IV.2009</collectingDate>
</date>
, leg.
<collectorName id="26FD530AFFC2942F1D90FA1090B3FA57" box="[712,756,1496,1517]" pageId="26" pageNumber="175">Buse</collectorName>
&amp;
<collectorName id="26FD530AFFC2942F1C41FA1F9123FA57" box="[793,868,1495,1517]" pageId="26" pageNumber="175">Pavlicek</collectorName>
(cFel).
<collectingCountry id="F31F764CFFC2942F1CE9FA1F91B3FA56" box="[945,1012,1495,1516]" name="Cyprus" pageId="26" pageNumber="175">Cyprus</collectingCountry>
:
<specimenCount id="9D0EFD55FFC2942F1B58FA1F967DFA57" box="[1024,1082,1495,1517]" pageId="26" pageNumber="175" type="generic">6 exs.</specimenCount>
,
<collectingRegion id="49CCF83EFFC2942F1F87FA279356F9BF" box="[223,273,1519,1541]" country="Cyprus" name="Pafos" pageId="26" pageNumber="175">Pafos</collectingRegion>
,
<location id="8ED76007FFC2942F1E42FA2793D4F9BF" LSID="urn:lsid:plazi:treatment:03A187CAFFC294291FEAFB6593A9FC51:8ED76007FFC2942F1E42FA2793D4F9BF" box="[282,403,1519,1541]" country="Israel" county="Fnaideq" latitude="32.75" longLatPrecision="1209" longitude="35.016666" municipality="Djebel Qammouaa" name="Troodos Mts." pageId="26" pageNumber="175" stateProvince="Pafos">Troodos Mts.</location>
,
<collectorName id="26FD530AFFC2942F1EC2FA27939CF9BE" box="[410,475,1519,1540]" pageId="26" pageNumber="175">Stavros</collectorName>
tis
<collectorName id="26FD530AFFC2942F1EA2FA27907EF9BF" box="[506,569,1519,1541]" pageId="26" pageNumber="175">Psokas</collectorName>
,
<quantity id="4CF09B39FFC2942F1D19FA2790E5F9BF" box="[577,674,1519,1541]" metricMagnitude="2" metricUnit="m" metricValue="8.75" metricValueMax="9.5" metricValueMin="8.0" pageId="26" pageNumber="175" unit="m" value="875.0" valueMax="950.0" valueMin="800.0">
<elevation id="0025D1EFFFC2942F1D19FA2790E5F9BF" box="[577,674,1519,1541]" metricMagnitude="2" metricUnit="m" metricValue="8.75" metricValueMax="9.5" metricValueMin="8.0" pageId="26" pageNumber="175" unit="m" value="875.0" valueMax="950.0" valueMin="800.0">800-950 m</elevation>
</quantity>
, 12+
<date id="FFB6101CFFC2942F1D95FA279175F9BE" box="[717,818,1519,1540]" pageId="26" pageNumber="175" value="2010-04-18">
<collectingDate id="EFF2E9F4FFC2942F1D95FA279175F9BE" box="[717,818,1519,1540]" pageId="26" pageNumber="175" value="2010-04-18">18.IV.2010</collectingDate>
</date>
, leg.
<collectorName id="26FD530AFFC2942F1C38FA2791CAF9BF" box="[864,909,1519,1541]" pageId="26" pageNumber="175">Wolf</collectorName>
(
<collectionCode id="ED19AE19FFC2942F1CC1FA279197F9BF" box="[921,976,1519,1541]" pageId="26" pageNumber="175">MNB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="DF776654FFC3942E1FEAFCAF93B4FC10" ID-DOI="http://doi.org/10.5281/zenodo.3985390" ID-Zenodo-Dep="3985390" httpUri="https://zenodo.org/record/3985390/files/figure.png" pageId="27" pageNumber="176" startId="27.[178,223,871,892]" targetBox="[178,1085,212,861]" targetPageId="27">
<paragraph id="8BB736DCFFC3942E1FEAFCAF93B4FC10" blockId="27.[178,1085,870,938]" pageId="27" pageNumber="176">
<emphasis id="B97CEACEFFC3942E1FEAFCAF92B1FCC1" bold="true" box="[178,246,870,892]" pageId="27" pageNumber="176">Map 4</emphasis>
: Distributions of
<taxonomicName id="4C084D5FFFC3942E1EFEFCAE9001FCC1" authorityName="Kiesenwetter" authorityYear="1858" box="[422,582,870,891]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="176" phylum="Arthropoda" rank="species" species="nivicola">
<emphasis id="B97CEACEFFC3942E1EFEFCAE9001FCC1" box="[422,582,870,891]" italics="true" pageId="27" pageNumber="176">Quedius nivicola</emphasis>
</taxonomicName>
(white circles; revised records),
<taxonomicName id="4C084D5FFFC3942E1CD1FCAF91BEFCC1" authorityName="Korge" authorityYear="1971" box="[905,1017,870,892]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="176" phylum="Arthropoda" rank="species" species="ortrudae">
<emphasis id="B97CEACEFFC3942E1CD1FCAF91BEFCC1" box="[905,1017,870,892]" italics="true" pageId="27" pageNumber="176">Q. ortrudae</emphasis>
</taxonomicName>
(black circles; revised records), and
<taxonomicName id="4C084D5FFFC3942E1E90FCB69050FC29" box="[456,535,894,915]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="176" phylum="Arthropoda" rank="species" species="josue">
<emphasis id="B97CEACEFFC3942E1E90FCB69050FC29" box="[456,535,894,915]" italics="true" pageId="27" pageNumber="176">Q. josue</emphasis>
</taxonomicName>
(black triangles: revised records; white triangles literature records) in the East Mediterranean.
</paragraph>
</caption>
<paragraph id="8BB736DCFFC3942E1FEAFC01933BF99C" blockId="27.[178,1085,969,1574]" pageId="27" pageNumber="176">
C o m m e n t:
<taxonomicName id="4C084D5FFFC3942E1E3CFC0193B1FC65" box="[356,502,969,991]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="176" phylum="Arthropoda" rank="species" species="josue">
<emphasis id="B97CEACEFFC3942E1E3CFC0193B1FC65" box="[356,502,969,991]" italics="true" pageId="27" pageNumber="176">Quedius josue</emphasis>
</taxonomicName>
was originally described based on a unique female from &quot;Arag-el-Emir&quot; (
<bibRefCitation id="EF994B2DFFC3942E1E07FC2093B7FC47" author="SAULCY F" box="[351,496,999,1021]" pageId="27" pageNumber="176" pagination="629 - 660" refId="ref21389" refString="SAULCY F. H. C. DE (1865): Descriptions des especes nouvelle de coleopteres recueillies en Syrie, en Egypte et en Palestine, pendant les mois d'octobre 1863 a janvier 1864, par M. de Saulcy, senateur, membre de l'institut. 2 e partie. - Annales de la Societe de Entomologique de France (4) 4: [1864]: 629 - 660." type="journal article" year="1865">SAULCY 1865</bibRefCitation>
), today
<collectingCountry id="F31F764CFFC3942E1D1CFC20902AFC47" box="[580,621,1000,1021]" name="Iraq" pageId="27" pageNumber="176">Iraq</collectingCountry>
al-Amir [
<geoCoordinate id="EE3C501BFFC3942E1D8BFC2F916CFC47" box="[723,811,999,1021]" degrees="31" direction="north" minutes="55" orientation="latitude" pageId="27" pageNumber="176" precision="925" value="31.916666">31°55'N</geoCoordinate>
, 35°45'] in
<collectingCountry id="F31F764CFFC3942E1CFDFC2091ABFC44" box="[933,1004,1000,1022]" name="Jordan" pageId="27" pageNumber="176">Jordan</collectingCountry>
. According to the description, the apical antennomeres are &quot;d'un roux foncé&quot;, the pronotum has the &quot;bord latéral et postérieur étroitement d'un roux foncé&quot;, and the elytra are &quot;testacées,..., avec un grande tache triangulaire commune d'un brun noir entourant l'écusson, s'étendant depuis la base&quot;. The species was subsequently reported from
<collectingCountry id="F31F764CFFC3942E1FEAFB49934AFB2D" box="[178,269,1153,1175]" name="Lebanon" pageId="27" pageNumber="176">Lebanon</collectingCountry>
,
<collectingCountry id="F31F764CFFC3942E1E42FB499324FB2C" box="[282,355,1153,1174]" name="Cyprus" pageId="27" pageNumber="176">Cyprus</collectingCountry>
(as
<emphasis id="B97CEACEFFC3942E1ECBFB499048FB2C" box="[403,527,1153,1174]" italics="true" pageId="27" pageNumber="176">Q. troodites</emphasis>
),
<collectingCountry id="F31F764CFFC3942E1D7FFB499025FB2D" box="[551,610,1153,1175]" name="Israel" pageId="27" pageNumber="176">Israel</collectingCountry>
,
<collectingCountry id="F31F764CFFC3942E1D37FB4990FCFB2D" box="[623,699,1153,1175]" name="Turkey" pageId="27" pageNumber="176">Turkey</collectingCountry>
, and
<collectingCountry id="F31F764CFFC3942E1DADFB499159FB2C" box="[757,798,1153,1174]" name="Iraq" pageId="27" pageNumber="176">Iraq</collectingCountry>
by
<bibRefCitation id="EF994B2DFFC3942E1C11FB4991A0FB2D" author="FAGEL G" box="[841,999,1153,1175]" pageId="27" pageNumber="176" pagination="1 - 13" refId="ref20496" refString="FAGEL G. (1968 a): Contribution a la connaissance des Staphylinidae CI. - Remarques diverses sur quelques Quedius de Mediterranee orientale. - B ulletin Institut Royale des Sciences Naturelles de Belgique 44 (12): 1 - 13." type="journal article" year="1968">FAGEL (1968a)</bibRefCitation>
,
<bibRefCitation id="EF994B2DFFC3942E1CACFB499344FB0C" author="KORGE H" pageId="27" pageNumber="176" pagination="1 - 68" refId="ref20958" refString="KORGE H. (1971 a): Beitrage zur Kenntnis der Koleopterenfauna Kleinasiens. - Annotationes Zoologicae et Botanicae 67: 1 - 68." type="journal article" year="1971">KORGE (1971a)</bibRefCitation>
, ASSING &amp; WUNDERLE (2001), and ASSING (2004). When treating the species of the
<taxonomicName id="4C084D5FFFC3942E1F83FB779316FB69" authorityName="Kiesenwetter" authorityYear="1858" box="[219,337,1214,1236]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="176" phylum="Arthropoda" rank="species" species="nivicola">
<emphasis id="B97CEACEFFC3942E1F83FB779316FB69" box="[219,337,1214,1236]" italics="true" pageId="27" pageNumber="176">Q. nivicola</emphasis>
</taxonomicName>
group,
<bibRefCitation id="EF994B2DFFC3942E1EFFFB779025FB6E" author="FAGEL G" box="[423,610,1214,1236]" pageId="27" pageNumber="176" pagination="1 - 13" refId="ref20496" refString="FAGEL G. (1968 a): Contribution a la connaissance des Staphylinidae CI. - Remarques diverses sur quelques Quedius de Mediterranee orientale. - B ulletin Institut Royale des Sciences Naturelles de Belgique 44 (12): 1 - 13." type="journal article" year="1968">FAGEL (1968a: 2)</bibRefCitation>
stated that the types of the
<taxonomicName id="4C084D5FFFC3942E1CCDFB7691AEFB69" box="[917,1001,1214,1235]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="B97CEACEFFC3942E1CCDFB7691AEFB69" box="[917,1001,1214,1235]" italics="true" pageId="27" pageNumber="176">Quedius</emphasis>
</taxonomicName>
species described by Saulcy were &quot;introuvables au Muséum d'Histoire naturelle, de Paris&quot;, so that they, like the type material of so many other
<taxonomicName id="4C084D5FFFC3942E1D96FB34911EFAA8" box="[718,857,1276,1298]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="176" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
species described by Saulcy, must be regarded as lost. In view of the similarity of the species allied to
<taxonomicName id="4C084D5FFFC3942E1B7CFAD392A0FAF5" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="176" phylum="Arthropoda" rank="species" species="josue">
<emphasis id="B97CEACEFFC3942E1B7CFAD392A0FAF5" italics="true" pageId="27" pageNumber="176">Q. josue</emphasis>
</taxonomicName>
, of the frequent previous confusion of these species, and of the fact that they have partly sympatric distributions, the designation of a
<typeStatus id="54B3887EFFC3942E1D8CFA909163FAD7" box="[724,804,1368,1389]" pageId="27" pageNumber="176" type="neotype">neotype</typeStatus>
is indispensable to unambiguously define the species. Based on the original description and the distributions of species of the
<taxonomicName id="4C084D5FFFC3942E1E11FA5D93FAFA10" authorityName="Kiesenwetter" authorityYear="1858" box="[329,445,1429,1450]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="176" phylum="Arthropoda" rank="species" species="nivicola">
<emphasis id="B97CEACEFFC3942E1E11FA5D93FAFA10" box="[329,445,1429,1450]" italics="true" pageId="27" pageNumber="176">Q. nivicola</emphasis>
</taxonomicName>
group, three species would qualify as suitable candidates:
<taxonomicName id="4C084D5FFFC3942E1B7CFA5D935DFA73" authorityName="Fagel" authorityYear="1968" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="176" phylum="Arthropoda" rank="species" species="petraensis">
<emphasis id="B97CEACEFFC3942E1B7CFA5D935DFA73" italics="true" pageId="27" pageNumber="176">Q. petraensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C084D5FFFC3942E1E73FA7C93A7FA73" authorityName="Fagel" authorityYear="1968" box="[299,480,1460,1481]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="176" phylum="Arthropoda" rank="species" species="problematicus">
<emphasis id="B97CEACEFFC3942E1E73FA7C93A7FA73" box="[299,480,1460,1481]" italics="true" pageId="27" pageNumber="176">Q. problematicus</emphasis>
</taxonomicName>
, and the species previously interpreted as
<taxonomicName id="4C084D5FFFC3942E1CF4FA7C9642FA73" box="[940,1029,1460,1481]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="176" phylum="Arthropoda" rank="species" species="josue">
<emphasis id="B97CEACEFFC3942E1CF4FA7C9642FA73" box="[940,1029,1460,1481]" italics="true" pageId="27" pageNumber="176">Q. josue</emphasis>
</taxonomicName>
(and illustrated as such) by
<bibRefCitation id="EF994B2DFFC3942E1EC6FA1B907CFA52" author="FAGEL G" box="[414,571,1490,1512]" pageId="27" pageNumber="176" pagination="1 - 13" refId="ref20496" refString="FAGEL G. (1968 a): Contribution a la connaissance des Staphylinidae CI. - Remarques diverses sur quelques Quedius de Mediterranee orientale. - B ulletin Institut Royale des Sciences Naturelles de Belgique 44 (12): 1 - 13." type="journal article" year="1968">FAGEL (1968a)</bibRefCitation>
. In the interest of the stability of nomenclature, a male belonging to the latter species is designated as the
<typeStatus id="54B3887EFFC3942E1C5CFA399110F9BC" box="[772,855,1521,1542]" pageId="27" pageNumber="176" type="neotype">neotype</typeStatus>
, thus avoiding further taxonomic changes.
</paragraph>
<paragraph id="8BB736DCFFC494291FEAFF2C903DFE29" blockId="28.[178,1085,228,403]" pageId="28" pageNumber="177">
The coloration of
<taxonomicName id="4C084D5FFFC494291E2BFF2C938DFF43" box="[371,458,228,249]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="josue">
<emphasis id="B97CEACEFFC494291E2BFF2C938DFF43" box="[371,458,228,249]" italics="true" pageId="28" pageNumber="177">Q. josue</emphasis>
</taxonomicName>
is subject to rather pronounced intraspecific variation. The pronotum may be of uniformly dark-brown to blackish coloration or it may have the margins narrowly reddish or yellowish (especially in material from
<collectingCountry id="F31F764CFFC494291C2BFEE99186FE8D" box="[883,961,289,311]" name="Cyprus" pageId="28" pageNumber="177">Cyprus</collectingCountry>
). The elytra may be uniformly yellowish to pale reddish, or they may be more or less extensively and more or less distinctly infuscate near the scutellum and along the suture. The aedeagus and its variation are illustrated in
<figureCitation id="13332A59FFC494291D5FFEB59031FE29" box="[519,630,381,403]" captionStart-0="Figs 53-62" captionStart-1="Figs 63-68" captionStartId-0="22.[178,218,1506,1528]" captionStartId-1="23.[178,218,906,928]" captionTargetBox-0="[179,1076,224,1480]" captionTargetBox-1="[183,1080,224,879]" captionTargetId-0="figure@22.[178,1085,212,1497]" captionTargetId-1="figure@23.[178,1085,212,897]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="Figs 53-62: Quedius josue from Lebanon (53), Israel (54-56), and Cyprus (57-62): (53-54, 57, 60) aedeagus in lateral view; (55, 58, 61) median lobe of aedeagus in ventral view; (56, 59, 62) paramere. Scale bar: 0.5 mm." captionText-1="Figs 63-68: Quedius josue from Syria (63-65) and Q. petraensis, holotype (66-68; figures in transparent light): (63) median lobe of aedeagus in lateral view; (64, 67) median lobe of aedeagus in ventral view; (65, 68) paramere; (66) aedeagus in lateral view. Scale bar: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3985386" figureDoi-1="http://doi.org/10.5281/zenodo.3985388" httpUri-0="https://zenodo.org/record/3985386/files/figure.png" httpUri-1="https://zenodo.org/record/3985388/files/figure.png" pageId="28" pageNumber="177">Figs 53-65</figureCitation>
.
</paragraph>
<paragraph id="8BB736DCFFC494291FEAFE6C91FEFE62" blockId="28.[178,1085,420,472]" pageId="28" pageNumber="177">
<taxonomicName id="4C084D5FFFC494291FEAFE6C9304FE03" box="[178,323,420,441]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="josue">
<emphasis id="B97CEACEFFC494291FEAFE6C9304FE03" box="[178,323,420,441]" italics="true" pageId="28" pageNumber="177">Quedius josue</emphasis>
</taxonomicName>
is distinguished from the sympatric or geographically close species of the
<taxonomicName id="4C084D5FFFC494291FEAFE0B9364FE6D" authorityName="Kiesenwetter" authorityYear="1858" box="[178,291,450,472]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="nivicola">
<emphasis id="B97CEACEFFC494291FEAFE0B9364FE6D" box="[178,291,450,472]" italics="true" pageId="28" pageNumber="177">Q. nivicola</emphasis>
</taxonomicName>
group (
<taxonomicName id="4C084D5FFFC494291E2BFE0B93BDFE62" authorityName="Fagel" authorityYear="1968" box="[371,506,451,472]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="petraensis">
<emphasis id="B97CEACEFFC494291E2BFE0B93BDFE62" box="[371,506,451,472]" italics="true" pageId="28" pageNumber="177">Q. petraensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C084D5FFFC494291D5EFE0B90F1FE6D" authorityName="Fagel" authorityYear="1968" box="[518,694,450,472]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="problematicus">
<emphasis id="B97CEACEFFC494291D5EFE0B90F1FE6D" box="[518,694,450,472]" italics="true" pageId="28" pageNumber="177">Q. problematicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C084D5FFFC494291D9BFE0B917CFE6D" authorityName="Korge" authorityYear="1971" box="[707,827,450,472]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="ortrudae">
<emphasis id="B97CEACEFFC494291D9BFE0B917CFE6D" box="[707,827,450,472]" italics="true" pageId="28" pageNumber="177">Q. ortrudae</emphasis>
</taxonomicName>
) as follows:
</paragraph>
<paragraph id="8BB736DCFFC494291FEAFE2190B6FDA4" blockId="28.[178,1085,489,542]" pageId="28" pageNumber="177">
from
<taxonomicName id="4C084D5FFFC494291FB4FE219331FE45" authorityName="Fagel" authorityYear="1968" box="[236,374,489,511]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="petraensis">
<emphasis id="B97CEACEFFC494291FB4FE219331FE45" box="[236,374,489,511]" italics="true" pageId="28" pageNumber="177">Q. petraensis</emphasis>
</taxonomicName>
by the different shapes of the median lobe and the paramere of the aedeagus, as well as by the paler coloration of the elytra;
</paragraph>
<paragraph id="8BB736DCFFC494291FEAFDE690DBFD1A" blockId="28.[178,1085,558,672]" pageId="28" pageNumber="177">
from
<taxonomicName id="4C084D5FFFC494291FB6FDE793E4FDF9" authorityName="Fagel" authorityYear="1968" box="[238,419,558,580]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="problematicus">
<emphasis id="B97CEACEFFC494291FB6FDE793E4FDF9" box="[238,419,558,580]" italics="true" pageId="28" pageNumber="177">Q. problematicus</emphasis>
</taxonomicName>
by the different shape of the apex of the median lobe of the aedeagus both in lateral and in ventral view, as well as by the darker coloration (
<taxonomicName id="4C084D5FFFC494291B7DFD859304FD3B" authorityName="Fagel" authorityYear="1968" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="problematicus">
<emphasis id="B97CEACEFFC494291B7DFD859304FD3B" italics="true" pageId="28" pageNumber="177">Q. problematicus</emphasis>
</taxonomicName>
: pronotum usually more extensively, or even completely reddish or yellowish) of the slightly less slender pronotum;
</paragraph>
<paragraph id="8BB736DCFFC494291FEAFD7990CDFCBE" blockId="28.[178,1085,689,772]" pageId="28" pageNumber="177">
from
<taxonomicName id="4C084D5FFFC494291FB4FD799320FD7C" authorityName="Korge" authorityYear="1971" box="[236,359,689,710]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="ortrudae">
<emphasis id="B97CEACEFFC494291FB4FD799320FD7C" box="[236,359,689,710]" italics="true" pageId="28" pageNumber="177">Q. ortrudae</emphasis>
</taxonomicName>
by a slightly smaller aedeagus with the apical portion of the median lobe of slightly different shape, and by the paler coloration of the elytra (
<taxonomicName id="4C084D5FFFC494291CE3FD189672FD5F" authorityName="Korge" authorityYear="1971" box="[955,1077,720,741]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="ortrudae">
<emphasis id="B97CEACEFFC494291CE3FD189672FD5F" box="[955,1077,720,741]" italics="true" pageId="28" pageNumber="177">Q. ortrudae</emphasis>
</taxonomicName>
: elytra reddish to dark reddish, or even darker).
</paragraph>
<paragraph id="8BB736DCFFC494291FEAFCDD906EFCD2" blockId="28.[178,1085,789,872]" pageId="28" pageNumber="177">
The known distribution of
<taxonomicName id="4C084D5FFFC494291E90FCDD9059FC91" box="[456,542,789,811]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="josue">
<emphasis id="B97CEACEFFC494291E90FCDD9059FC91" box="[456,542,789,811]" italics="true" pageId="28" pageNumber="177">Q. josue</emphasis>
</taxonomicName>
includes
<collectingCountry id="F31F764CFFC494291DDAFCDD908CFC90" box="[642,715,789,810]" name="Cyprus" pageId="28" pageNumber="177">Cyprus</collectingCountry>
and the Middle East from
<collectingCountry id="F31F764CFFC494291CBCFCDD967AFC91" box="[996,1085,789,811]" name="Lebanon" pageId="28" pageNumber="177">Lebanon</collectingCountry>
and North
<collectingCountry id="F31F764CFFC494291E73FCFC9327FCF0" box="[299,352,820,842]" name="Syria" pageId="28" pageNumber="177">Syria</collectingCountry>
(first record) to
<collectingCountry id="F31F764CFFC494291D47FCFC9010FCF0" box="[543,599,820,842]" name="Israel" pageId="28" pageNumber="177">Israel</collectingCountry>
and
<collectingCountry id="F31F764CFFC494291DCDFCFC909FFCF0" box="[661,728,820,842]" name="Jordan" pageId="28" pageNumber="177">Jordan</collectingCountry>
(
<figureCitation id="13332A59FFC494291DB4FCFC9170FCF0" box="[748,823,820,842]" captionStart="Map 4" captionStartId="27.[178,223,871,892]" captionTargetBox="[178,1085,212,861]" captionTargetId="figure@27.[178,1085,212,861]" captionTargetPageId="27" captionText="Map 4: Distributions of Quedius nivicola (white circles; revised records), Q. ortrudae (black circles; revised records), and Q. josue (black triangles: revised records; white triangles literature records) in the East Mediterranean." figureDoi="http://doi.org/10.5281/zenodo.3985390" httpUri="https://zenodo.org/record/3985390/files/figure.png" pageId="28" pageNumber="177">Map 4</figureCitation>
). Previous records from
<collectingCountry id="F31F764CFFC494291FEAFC9A92BCFCD2" box="[178,251,850,872]" name="Turkey" pageId="28" pageNumber="177">Turkey</collectingCountry>
and
<collectingCountry id="F31F764CFFC494291E75FC9B9311FCD2" box="[301,342,851,872]" name="Iraq" pageId="28" pageNumber="177">Iraq</collectingCountry>
refer to
<taxonomicName id="4C084D5FFFC494291EF4FC9B9064FCDD" authorityName="Korge" authorityYear="1971" box="[428,547,850,872]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="ortrudae">
<emphasis id="B97CEACEFFC494291EF4FC9B9064FCDD" box="[428,547,850,872]" italics="true" pageId="28" pageNumber="177">Q. ortrudae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3126557FFC494291FEAFCB193A9FC51" pageId="28" pageNumber="177" type="materials_examined">
<paragraph id="8BB736DCFFC494291FEAFCB193A9FC51" blockId="28.[178,1085,889,1003]" pageId="28" pageNumber="177">
Like
<taxonomicName id="4C084D5FFFC494291FB2FCB193DAFC34" authorityName="Fagel" authorityYear="1968" box="[234,413,889,910]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="problematicus">
<emphasis id="B97CEACEFFC494291FB2FCB193DAFC34" box="[234,413,889,910]" italics="true" pageId="28" pageNumber="177">Q. problematicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C084D5FFFC494291EF5FCB19042FC35" box="[429,517,889,911]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="177" phylum="Arthropoda" rank="species" species="josue">
<emphasis id="B97CEACEFFC494291EF5FCB19042FC35" box="[429,517,889,911]" italics="true" pageId="28" pageNumber="177">Q. josue</emphasis>
</taxonomicName>
is epigeic, as can be inferred from the habitat details reported by
<bibRefCitation id="EF994B2DFFC494291E75FC509382FC14" author="FAGEL G" box="[301,453,920,942]" pageId="28" pageNumber="177" pagination="1 - 13" refId="ref20496" refString="FAGEL G. (1968 a): Contribution a la connaissance des Staphylinidae CI. - Remarques diverses sur quelques Quedius de Mediterranee orientale. - B ulletin Institut Royale des Sciences Naturelles de Belgique 44 (12): 1 - 13." type="journal article" year="1968">FAGEL (1968a)</bibRefCitation>
and ASSING &amp; WUNDERLE (2001), by the fact that most of the examined material was sifted from leaf litter, and by the long series of specimens collected on various occasions.
</paragraph>
</subSubSection>
</treatment>
</document>
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