266 lines
32 KiB
XML
266 lines
32 KiB
XML
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<mods:title id="E9939F9CF94D3BC4A9F13EDC6EB0625E">A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths</mods:title>
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<mods:namePart id="507AA229307C33AC531A7D87099507BD">Schwarzhans, Werner</mods:namePart>
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<mods:affiliation id="852F359F582846CD1500028990C959F7">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, 2100 Copenhagen, Denmark</mods:affiliation>
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<mods:namePart id="199387F9E2B02897B142ECAE923B399C">Klots, Oleksandr</mods:namePart>
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<mods:namePart id="D86AF5DFB6E41C6F69D907BA2DBD5443">Ryabokon, Tamara</mods:namePart>
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<mods:namePart id="95B0342F2AC645738ABD1C1C79CE478B">Kovalchuk, Oleksandr</mods:namePart>
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<mods:title id="40CCED2B8695A2D3C80581DFFF9B499A">Swiss Journal of Palaeontology</mods:title>
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<mods:date id="640EF5137F128A7A6F87B8C0932D1566">2022</mods:date>
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<mods:number id="C7055B8D9F7B8C6807FD08F845EDB274">2022-11-17</mods:number>
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<mods:number id="F8ADF5F86F97852BECEFD68FC09FC857">141</mods:number>
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<heading id="D0C37B21FE43FFA8F872FEF0FBE5FEB2" bold="true" box="[813,1119,299,323]" fontSize="10" level="1" pageId="10" pageNumber="11" reason="1">
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<emphasis id="B940105FFE43FFA8F872FEF0FBE5FEB2" bold="true" box="[813,1119,299,323]" pageId="10" pageNumber="11">
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Genus
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<taxonomicName id="4C34B7CEFE43FFA8F822FEF0FBB7FEB2" authority="Schwarzhans & Klots & Ryabokon & Kovalchuk, 2022" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[893,1037,299,323]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="B940105FFE43FFA8F822FEF0FBB7FEB2" bold="true" box="[893,1037,299,323]" italics="true" pageId="10" pageNumber="11">Parenypnias</emphasis>
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<taxonomicNameLabel id="A273AD24FE43FFA8FF4CFEF0FBE5FEB2" box="[1043,1119,299,323]" pageId="10" pageNumber="11" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C32E9FC6FE43FFAEF872FEB7FDC4FEB2" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph id="8B8BCC4DFE43FFA8F872FEB7FB4AFE72" blockId="10.[813,1264,363,387]" box="[813,1264,363,387]" pageId="10" pageNumber="11">
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<emphasis id="B940105FFE43FFA8F872FEB7FC0AFE72" box="[813,944,364,387]" italics="true" pageId="10" pageNumber="11">
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<typeStatus id="548F72EFFE43FFA8F872FEB7FCDBFE72" box="[813,865,364,387]" pageId="10" pageNumber="11">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="4C34B7CEFE43FFA8F8E3FEB7FB0BFE72" authority="Schwarzhans & Klots & Ryabokon & Kovalchuk, 2022" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[956,1201,363,387]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="inauditus" status="sp. nov.">
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<emphasis id="B940105FFE43FFA8F8E3FEB7FB0BFE72" box="[956,1201,363,387]" italics="true" pageId="10" pageNumber="11">Parenypnias inauditus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A273AD24FE43FFA8FFE8FEB7FB4AFE72" box="[1207,1264,364,387]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8B8BCC4DFE43FFA8F872FE70FA91FDB2" blockId="10.[813,1437,427,579]" pageId="10" pageNumber="11">
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<emphasis id="B940105FFE43FFA8F872FE70FC24FE32" box="[813,926,427,451]" italics="true" pageId="10" pageNumber="11">Etymology</emphasis>
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Derived from the gobiosomatine genus
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<taxonomicName id="4C34B7CEFE43FFA8FE02FE77FCE1FE12" authorityName="Jordan & Evermann" authorityYear="1898" family="Gobiidae" genus="Enypnias" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
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<emphasis id="B940105FFE43FFA8FE02FE77FCE1FE12" italics="true" pageId="10" pageNumber="11">Enypnias</emphasis>
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</taxonomicName>
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<collectingCountry id="F3238CDDFE43FFA8F83BFE10FC16FE12" box="[868,940,459,483]" name="Jordan" pageId="10" pageNumber="11">Jordan</collectingCountry>
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& Evermann, 1898 because of some resemblance to otoliths of that genus, which, however, is regarded as a junior synonym of
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<taxonomicName id="4C34B7CEFE43FFA8FFCDFDD0FA26FDD2" authority="Girard, 1858" authorityName="Girard" authorityYear="1858" box="[1170,1436,523,547]" family="Gobiidae" genus="Gobiosoma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
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<emphasis id="B940105FFE43FFA8FFCDFDD0FAB6FDD2" box="[1170,1292,523,547]" italics="true" pageId="10" pageNumber="11">Gobiosoma</emphasis>
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Girard, 1858
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</taxonomicName>
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according to
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<bibRefCitation id="EFA5B1BCFE43FFA8F8E5FDF0FA9DFDB2" author="Tornabene, L. & Van Tassell, J. L." box="[954,1319,555,579]" pageId="10" pageNumber="11" refId="ref23817" refString="Tornabene, L., & Van Tassell, J. L. (2014). Redescription of the goby genus Gobiosoma (Teleostei: Gobiidae: Gobiosomatini), with the synonymy" type="book" year="2014">Tornabene and van Tassell (2014)</bibRefCitation>
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.
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||
</paragraph>
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||
<paragraph id="8B8BCC4DFE43FFA8F872FDB7FBDBFBD2" blockId="10.[813,1437,619,1859]" pageId="10" pageNumber="11">
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<emphasis id="B940105FFE43FFA8F872FDB7FC2DFD72" box="[813,919,620,643]" italics="true" pageId="10" pageNumber="11">Diagnosis</emphasis>
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An otolith of the family
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<taxonomicName id="4C34B7CEFE43FFA8FFEFFDB0FAAFFD72" authorityName="Cuvier" authorityYear="1816" box="[1200,1301,619,643]" family="Gobiidae" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
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, supposedly belonging to the Gobiosomatini, with the following combination of diagnostic characters. Otolith size small, up to slightly over
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<quantity id="4CCC61A8FE43FFA8F884FD10FB8BFD12" box="[987,1073,715,739]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="mm" value="1.5">1.5 mm</quantity>
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in length. Ventral rim of otolith straight, horizontal; anterior rim near vertical or with pointed preventral projection; anterior and posterior rims with deep incisions at anterior and posterior terminations of sulcus. Dorsal rim with distinct middorsal angle and relatively short, mildly to distinctly outward bent postdorsal projection. Inner and outer faces about equally convex or inner face more convex. Sulcus narrow, moderately short, oval, without ostial lobe or subcaudal iugum, inclined at 15°–18°. Ventral furrow following outline of ventral rim of otolith.
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</paragraph>
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<paragraph id="8B8BCC4DFE43FFA8F81DFBF7FA27FA32" blockId="10.[813,1437,619,1859]" pageId="10" pageNumber="11">
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<emphasis id="B940105FFE43FFA8F81DFBF7FC0FFBB2" box="[834,949,1068,1091]" italics="true" pageId="10" pageNumber="11">Discussion</emphasis>
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Te Gobiosomatini are a group of gobiid fishes that are today only known from the seas around the Americas, where they form one of the most diverse and species rich groups of the family (
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<bibRefCitation id="EFA5B1BCFE43FFA8FF91FB50FA37FB52" author="RUber, L. & Van Tassell, J. L. & Zardoya, R." box="[1230,1421,1163,1187]" pageId="10" pageNumber="11" pagination="1584 - 1598" refId="ref22835" refString="RUber, L., Van Tassell, J. L., & Zardoya, R. (2003). Rapid speciation and ecological divergence in the American seven-spined gobies (Gobiidae, Gobiosomatini) inferred from a molecular phylogeny. Evolution, 57 (7), 1584 - 1598." type="journal article" year="2003">Rüber et al., 2003</bibRefCitation>
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). Teir otoliths appear inconspicuous at first glance but are characterized by certain traits not usually found in combination in Atlanto-Mediterranean goby groups. Tese traits are the relatively narrow sulcus with or without ostial lobe, the lack of a subcaudal iugum, the outward bent postdorsal projection, and the inner face being as strongly convex as the outer face or more so. In that respect,
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<taxonomicName id="4C34B7CEFE43FFA8F8CFFA57FBACFA52" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[912,1046,1420,1443]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
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<emphasis id="B940105FFE43FFA8F8CFFA57FBACFA52" box="[912,1046,1420,1443]" italics="true" pageId="10" pageNumber="11">Parenypnias</emphasis>
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</taxonomicName>
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closely resembles otoliths of the
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<typeStatus id="548F72EFFE43FFA8F872FA77FCE6FA35" box="[813,860,1452,1476]" pageId="10" pageNumber="11">type</typeStatus>
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species of
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<taxonomicName id="4C34B7CEFE43FFA8F883FA77FBFAFA32" authorityName="Jordan & Evermann" authorityYear="1898" box="[988,1088,1452,1475]" family="Gobiidae" genus="Enypnias" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
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<emphasis id="B940105FFE43FFA8F883FA77FBFAFA32" box="[988,1088,1452,1475]" italics="true" pageId="10" pageNumber="11">Enypnias</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C34B7CEFE43FFA8FF0FFA77FA27FA32" authority="(Gunther, 1861)" baseAuthorityName="Gunther" baseAuthorityYear="1861" box="[1104,1437,1451,1475]" family="Gobiidae" genus="Enypnias" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="seminudus">
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<emphasis id="B940105FFE43FFA8FF0FFA77FB59FA32" box="[1104,1251,1451,1475]" italics="true" pageId="10" pageNumber="11">E. seminudus</emphasis>
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(Günther, 1861)
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</taxonomicName>
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</paragraph>
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<paragraph id="8B8BCC4DFE43FFA9F872FA10FD03F93B" blockId="10.[813,1437,619,1859]" lastBlockId="11.[151,775,1682,1898]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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(
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<figureCitation id="130FD0C8FE43FFA8F86BFA10FCC4FA12" box="[820,894,1483,1507]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: a–d Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.e–h Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (1616–1622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22′ N 87° 52′ W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="10" pageNumber="11">Fig. 5j</figureCitation>
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) in the outline of the otolith, the pronounced postdorsal projection, and the narrow sulcus with a low ostial lobe. It differs in the rather strongly reduced and small, nearly oval sulcus (vs. sole-shaped or somewhat smoothened sole-shaped) and the ventral furrow following the ventral rim of the otolith (vs. being more regularly curved and clipping the postventral angles/projections). Te otoliths of
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<taxonomicName id="4C34B7CEFE43FFA8F888F970FBEBF932" authorityName="Girard" authorityYear="1858" box="[983,1105,1707,1731]" family="Gobiidae" genus="Gobiosoma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
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<emphasis id="B940105FFE43FFA8F888F970FBEBF932" box="[983,1105,1707,1731]" italics="true" pageId="10" pageNumber="11">Gobiosoma</emphasis>
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</taxonomicName>
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with the
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<typeStatus id="548F72EFFE43FFA8FFE0F977FB54F935" box="[1215,1262,1708,1732]" pageId="10" pageNumber="11">type</typeStatus>
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species
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<taxonomicName id="4C34B7CEFE43FFA8FE12F977FC5DF912" authority="(Lacepede, 1800)" baseAuthorityName="Lacepede" baseAuthorityYear="1800" family="Gobiidae" genus="Gobiosoma" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="bosc">
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<emphasis id="B940105FFE43FFA8FE12F977FA26F932" box="[1357,1436,1707,1731]" italics="true" pageId="10" pageNumber="11">G. bosc</emphasis>
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(Lacépède, 1800)
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</taxonomicName>
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(
|
||
<figureCitation id="130FD0C8FE43FFA8F8A8F910FBF9F912" box="[1015,1091,1739,1763]" captionStart="Fig" captionStartId="11.[168,199,1466,1486]" captionTargetBox="[152,1438,237,1629]" captionTargetId="figure-126@11.[167,1422,253,1453]" captionTargetPageId="11" captionText="Fig. 5 Otoliths of Odondebuenia, Parenypnias and Bathygobius?: a–d Odondebuenia agiadiae Schwarzhans et al., 2020a, 2020b, a (reversed) Kozatskyi Yar,NMB P1219, b (reversed), d Kozatskyi Yar, NMNHU-P PI 2569, c (reversed) Mlyntsi, NMNHU-P PI 2570.e–h Parenypnias inauditus n. gen. et n. sp., f holotype, Kozatskyi Yar,NMNHU-P PI 2550,e, g (reversed) Shydlivshchyna, NMB P1207, h Žižkov 1 well (1616–1622 m), DGS MU-0395. i Parenypnias kiselevi n. gen. et n. sp., holotype, Mlyntsi,NMNHU-P PI 2574. j Enypnias seminudus (GÜnther, 1861), Recent,USNM 407784,13° 22′ N 87° 52′ W.k Gobiosoma bosc (Lacépède,1800), Recent (reversed), LACM coll. Fitch, off Rappahannock. l Bathygobius? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548" figureDoi="http://doi.org/10.5281/zenodo.11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="10" pageNumber="11">Fig. 5k</figureCitation>
|
||
) are more similar in respect to the shape and size of the sulcus but differ significantly in the reduced postdorsal projection and the postventral projection being significantly more pronounced than the postdorsal one. In this respect,
|
||
<taxonomicName id="4C34B7CEFE42FFA9F972F948FD09F95B" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[557,691,1683,1706]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
|
||
<emphasis id="B940105FFE42FFA9F972F948FD09F95B" box="[557,691,1683,1706]" italics="true" pageId="11" pageNumber="12">Parenypnias</emphasis>
|
||
</taxonomicName>
|
||
almost resembles a combination of the two morphotypes.
|
||
</paragraph>
|
||
<caption id="DF4B9CC5FE42FFA9FBF7FA61FF68F9A2" ID-DOI="http://doi.org/10.5281/zenodo.11999891" ID-Zenodo-Dep="11999891" httpUri="https://zenodo.org/record/11999891/files/figure.png" pageId="11" pageNumber="12" startId="11.[168,199,1466,1486]" targetBox="[152,1438,237,1629]" targetPageId="11" targetType="figure">
|
||
<paragraph id="8B8BCC4DFE42FFA9FBF7FA61FF68F9A2" blockId="11.[168,1421,1466,1619]" pageId="11" pageNumber="12">
|
||
<emphasis id="B940105FFE42FFA9FBF7FA61FF61FA3F" bold="true" box="[168,219,1466,1486]" pageId="11" pageNumber="12">Fig. 5</emphasis>
|
||
Otoliths of
|
||
<emphasis id="B940105FFE42FFA9FA1CFA61FD93FA3F" box="[323,553,1466,1486]" italics="true" pageId="11" pageNumber="12">
|
||
<taxonomicName id="4C34B7CEFE42FFA9FA1CFA61FE05FA3F" authorityName="de Buen" authorityYear="1930" box="[323,447,1466,1486]" family="Gobiidae" genus="Odondebuenia" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">Odondebuenia</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C34B7CEFE42FFA9FA99FA61FD93FA3F" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[454,553,1466,1486]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">Parenypnias</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C34B7CEFE42FFA9F90CFA61FD02FA3F" authorityName="Bleeker" authorityYear="1878" box="[595,696,1466,1486]" family="Gobiidae" genus="Bathygobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
|
||
<emphasis id="B940105FFE42FFA9F90CFA61FD02FA3F" box="[595,696,1466,1486]" italics="true" pageId="11" pageNumber="12">Bathygobius</emphasis>
|
||
</taxonomicName>
|
||
?:
|
||
<emphasis id="B940105FFE42FFA9F996FA61FD50FA3F" bold="true" box="[713,746,1466,1486]" pageId="11" pageNumber="12">a–d</emphasis>
|
||
<taxonomicName id="4C34B7CEFE42FFA9F9B0FA61FB28FA3F" authority="Schwarzhans et al., 2020 a" authorityName="Schwarzhans" authorityYear="2020" box="[751,1170,1466,1486]" family="Gobiidae" genus="Odondebuenia" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="agiadiae">
|
||
<emphasis id="B940105FFE42FFA9F9B0FA61FC0FFA3F" box="[751,949,1466,1486]" italics="true" pageId="11" pageNumber="12">Odondebuenia agiadiae</emphasis>
|
||
<bibRefCitation id="EFA5B1BCFE42FFA9F8E5FA61FB28FA3F" author="Schwarzhans, W. & Brzobohaty, R. & Radwanska, U." box="[954,1170,1466,1486]" pageId="11" pageNumber="12" refId="ref23388" refString="Schwarzhans, W., Brzobohaty, R., & Radwanska, U. (2020 a). Goby otoliths from the Badenian (middle Miocene) of the Central Paratethys from the Czech Republic, Slovakia and Poland: A baseline for the evolution of" type="book" year="2020">Schwarzhans et al., 2020a</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<bibRefCitation id="EFA5B1BCFE42FFA9FFC5FA61FB75FA3F" author="Schwarzhans, W. & Agiadi, K. & Carnevale, G." box="[1178,1231,1466,1486]" pageId="11" pageNumber="12" pagination="657 - 724" refId="ref23336" refString="Schwarzhans, W., Agiadi, K., & Carnevale, G. (2020 b). Late Miocene - early Pliocene evolution of Mediterranean gobies and their environmental and biogeographic significance. Rivista Italiana di Paleontologia e Stratigrafia, 126 (3), 657 - 724." type="journal article" year="2020">2020b</bibRefCitation>
|
||
,
|
||
<emphasis id="B940105FFE42FFA9FF88FA61FB58FA3F" bold="true" box="[1239,1250,1466,1486]" pageId="11" pageNumber="12">a</emphasis>
|
||
(reversed) Kozatskyi Yar,NMB P1219,
|
||
<emphasis id="B940105FFE42FFA9FA6BFA0FFEFAFA19" bold="true" box="[308,320,1492,1512]" pageId="11" pageNumber="12">b</emphasis>
|
||
(reversed),
|
||
<emphasis id="B940105FFE42FFA9FAFEFA0FFE17FA19" bold="true" box="[417,429,1492,1512]" pageId="11" pageNumber="12">d</emphasis>
|
||
Kozatskyi Yar, NMNHU-P PI 2569,
|
||
<emphasis id="B940105FFE42FFA9F998FA0FFD6BFA19" bold="true" box="[711,721,1492,1512]" pageId="11" pageNumber="12">c</emphasis>
|
||
(reversed) Mlyntsi, NMNHU-P PI 2570.
|
||
<emphasis id="B940105FFE42FFA9FF4AFA0FFB8DFA19" bold="true" box="[1045,1079,1492,1512]" pageId="11" pageNumber="12">e–h</emphasis>
|
||
<taxonomicName id="4C34B7CEFE42FFA9FF63FA0FFB55FA19" authority="Schwarzhans & Klots & Ryabokon & Kovalchuk, 2022" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[1084,1263,1492,1512]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="inauditus" status="n. gen. et n. sp.">
|
||
<emphasis id="B940105FFE42FFA9FF63FA0FFB55FA19" box="[1084,1263,1492,1512]" italics="true" pageId="11" pageNumber="12">Parenypnias inauditus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273AD24FE42FFA9FFABFA0FFACFFA19" box="[1268,1397,1492,1512]" pageId="11" pageNumber="12" rank="species">n. gen. et n. sp.</taxonomicNameLabel>
|
||
,
|
||
<emphasis id="B940105FFE42FFA9FE24FA0FFA38FA19" bold="true" box="[1403,1410,1492,1512]" pageId="11" pageNumber="12">f</emphasis>
|
||
holotype, Kozatskyi Yar,NMNHU-P PI 2550,
|
||
<emphasis id="B940105FFE42FFA9F94DFA34FDA7F9F2" bold="true" box="[530,541,1519,1539]" pageId="11" pageNumber="12">e</emphasis>
|
||
,
|
||
<emphasis id="B940105FFE42FFA9F979FA34FD88F9F2" bold="true" box="[550,562,1519,1539]" pageId="11" pageNumber="12">g</emphasis>
|
||
(reversed) Shydlivshchyna, NMB P1207,
|
||
<emphasis id="B940105FFE42FFA9F8DAFA34FC2BF9F2" bold="true" box="[901,913,1519,1539]" pageId="11" pageNumber="12">h</emphasis>
|
||
Žižkov 1 well (1616–1622 m), DGS MU-0395.
|
||
<emphasis id="B940105FFE42FFA9FE4FFA34FAAFF9F2" bold="true" box="[1296,1301,1519,1539]" pageId="11" pageNumber="12">i</emphasis>
|
||
<taxonomicName id="4C34B7CEFE42FFA9FE45FA34FF5BF9EF" authority="Schwarzhans & Klots & Ryabokon & Kovalchuk, 2022" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="kiselevi" status="n. gen. et n. sp.">
|
||
<emphasis id="B940105FFE42FFA9FE45FA34FF5BF9EF" italics="true" pageId="11" pageNumber="12">Parenypnias kiselevi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273AD24FE42FFA9FBB9F9D1FEDDF9EF" box="[230,359,1546,1566]" pageId="11" pageNumber="12" rank="species">n. gen. et n. sp.</taxonomicNameLabel>
|
||
, holotype, Mlyntsi,NMNHU-P PI 2574.
|
||
<emphasis id="B940105FFE42FFA9F9F6F9D1FD15F9EF" bold="true" box="[681,687,1546,1566]" pageId="11" pageNumber="12">j</emphasis>
|
||
<taxonomicName id="4C34B7CEFE42FFA9F9ECF9D1FCE2F9EF" baseAuthorityName="Gunther" baseAuthorityYear="1861" box="[691,856,1546,1566]" family="Gobiidae" genus="Enypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="seminudus">
|
||
<emphasis id="B940105FFE42FFA9F9ECF9D1FCE2F9EF" box="[691,856,1546,1566]" italics="true" pageId="11" pageNumber="12">Enypnias seminudus</emphasis>
|
||
</taxonomicName>
|
||
(GÜnther, 1861), Recent,USNM 407784,13° 22
|
||
<emphasis id="B940105FFE42FFA9FFBDF9D2FB5DF9EC" bold="true" box="[1250,1255,1545,1565]" italics="true" pageId="11" pageNumber="12">′</emphasis>
|
||
N 87° 52
|
||
<emphasis id="B940105FFE42FFA9FE6CF9D2FA82F9EC" bold="true" box="[1331,1336,1545,1565]" italics="true" pageId="11" pageNumber="12">′</emphasis>
|
||
W.
|
||
<emphasis id="B940105FFE42FFA9FE0AF9D1FADAF9EF" bold="true" box="[1365,1376,1546,1566]" pageId="11" pageNumber="12">k</emphasis>
|
||
<taxonomicName id="4C34B7CEFE42FFA9FBF7F9FFFE7DF9C9" authority="(Lacepede, 1800)" baseAuthorityName="Lacepede" baseAuthorityYear="1800" box="[168,455,1572,1592]" family="Gobiidae" genus="Gobiosoma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="bosc">
|
||
<emphasis id="B940105FFE42FFA9FBF7F9FFFE8BF9C9" box="[168,305,1572,1592]" italics="true" pageId="11" pageNumber="12">Gobiosoma bosc</emphasis>
|
||
(Lacépède,1800)
|
||
</taxonomicName>
|
||
, Recent (reversed), LACM coll. Fitch, off Rappahannock.
|
||
<emphasis id="B940105FFE42FFA9F8C2F9FFFC18F9C9" bold="true" box="[925,930,1572,1592]" pageId="11" pageNumber="12">l</emphasis>
|
||
<taxonomicName id="4C34B7CEFE42FFA9F8F8F9FFFBB6F9C9" authorityName="Bleeker" authorityYear="1878" box="[935,1036,1572,1592]" family="Gobiidae" genus="Bathygobius" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
|
||
<emphasis id="B940105FFE42FFA9F8F8F9FFFBB6F9C9" box="[935,1036,1572,1592]" italics="true" pageId="11" pageNumber="12">Bathygobius</emphasis>
|
||
</taxonomicName>
|
||
? sp. (reversed), Kozatskyi Yar, NMNHU-P PI 2548
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8BCC4DFE42FFA9FBF2F909FBDEF89B" blockId="11.[151,775,1682,1898]" lastBlockId="11.[813,1437,1682,1898]" pageId="11" pageNumber="12">
|
||
Te Gobiosomatini are considered to represent a group endemic to the tropical to subtropical shores of the Americas (
|
||
<bibRefCitation id="EFA5B1BCFE42FFA9FA4EF8C9FE61F8DB" author="RUber, L. & Van Tassell, J. L. & Zardoya, R." box="[273,475,1810,1834]" pageId="11" pageNumber="12" pagination="1584 - 1598" refId="ref22835" refString="RUber, L., Van Tassell, J. L., & Zardoya, R. (2003). Rapid speciation and ecological divergence in the American seven-spined gobies (Gobiidae, Gobiosomatini) inferred from a molecular phylogeny. Evolution, 57 (7), 1584 - 1598." type="journal article" year="2003">Rüber et al., 2003</bibRefCitation>
|
||
). Teir outgroup relationship is still elusive and has been discussed in Rüber et. al. (2003). Due to the occurrence of
|
||
<taxonomicName id="4C34B7CEFE42FFA9F960F888FD7FF89B" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[575,709,1875,1898]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
|
||
<emphasis id="B940105FFE42FFA9F960F888FD7FF89B" box="[575,709,1875,1898]" italics="true" pageId="11" pageNumber="12">Parenypnias</emphasis>
|
||
</taxonomicName>
|
||
away from the extant distribution range and some remaining morphological uncertainties, our allocation to the Gobiosomatini should be considered tentative. Interestingly, however, the extant Gobiosomatini of the genus
|
||
<taxonomicName id="4C34B7CEFE42FFA9FE02F928FCE1F8DB" authorityName="Jordan & Evermann" authorityYear="1898" family="Gobiidae" genus="Enypnias" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
|
||
<emphasis id="B940105FFE42FFA9FE02F928FCE1F8DB" italics="true" pageId="11" pageNumber="12">Enypnias</emphasis>
|
||
</taxonomicName>
|
||
are mostly associated with reef environments, while those of the genus
|
||
<taxonomicName id="4C34B7CEFE42FFA9FF57F8E9FB38F8BB" authorityName="Girard" authorityYear="1858" box="[1032,1154,1842,1866]" family="Gobiidae" genus="Gobiosoma" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
|
||
<emphasis id="B940105FFE42FFA9FF57F8E9FB38F8BB" box="[1032,1154,1842,1866]" italics="true" pageId="11" pageNumber="12">Gobiosoma</emphasis>
|
||
</taxonomicName>
|
||
occur in shallow coastal and estuarine environments.
|
||
</paragraph>
|
||
<paragraph id="8B8BCC4DFE45FFAEFBC8FF37FDC4FEB2" blockId="12.[151,775,235,323]" pageId="12" pageNumber="13">
|
||
<emphasis id="B940105FFE45FFAEFBC8FF37FF5EFEF2" box="[151,228,236,259]" italics="true" pageId="12" pageNumber="13">Species</emphasis>
|
||
Two species:
|
||
<taxonomicName id="4C34B7CEFE45FFAEFADDFF37FDBEFEF2" authority="Schwarzhans & Klots & Ryabokon & Kovalchuk, 2022" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[386,516,235,259]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="inauditus" status="n. sp.">
|
||
<emphasis id="B940105FFE45FFAEFADDFF37FDBEFEF2" box="[386,516,235,259]" italics="true" pageId="12" pageNumber="13">P. inauditus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273AD24FE45FFAEF952FF37FDF0FEF2" box="[525,586,236,259]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
|
||
, which is known from the late Badenian of the
|
||
<collectingCountry id="F3238CDDFE45FFAEFABFFED0FD31FED2" box="[480,651,267,291]" name="Czech Republic" pageId="12" pageNumber="13">Czech Republic</collectingCountry>
|
||
(as revised here) and
|
||
<collectingCountry id="F3238CDDFE45FFAEFA5BFEF0FEE4FEB2" box="[260,350,299,323]" name="Ukraine" pageId="12" pageNumber="13">Ukraine</collectingCountry>
|
||
, and
|
||
<taxonomicName id="4C34B7CEFE45FFAEFAC8FEF7FE44FEB2" authorityName="Schwarzhans & Klots & Ryabokon & Kovalchuk" authorityYear="2022" box="[407,510,299,323]" family="Gobiidae" genus="Parenypnias" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="kiselevi">
|
||
<emphasis id="B940105FFE45FFAEFAC8FEF7FE44FEB2" box="[407,510,299,323]" italics="true" pageId="12" pageNumber="13">P. kiselevi</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F3238CDDFE45FFAEF97FFEF0FDC1FEB2" box="[544,635,299,323]" name="Ukraine" pageId="12" pageNumber="13">Ukraine</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |