treatments-xml/data/03/99/87/039987FCFFF65171FF6FF2861D03E3FE.xml

<document id="BE9FA7EAE091F6D9158621695B23D644" ID-DOI="10.5281/zenodo.215101" ID-GBIF-Dataset="b87bd712-6625-473a-8cce-db82db29dece" ID-ISSN="1175-5326" ID-Zenodo-Dep="215101" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460483662183" checkinUser="plazi" docAuthor="Rowley, Jodi J. L., Hoang, Huy Duc, Dau, Vinh Quang, Le, Duong Thi Thuy &amp; Cao, Trung Tien" docDate="2012" docId="039987FCFFF65171FF6FF2861D03E3FE" docLanguage="en" docName="zt03321p068.pdf" docOrigin="Zootaxa 3321" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptolalax firthi Rowley, Hoang, Dau, Le &amp; Cao, 2012, sp. nov." docType="treatment" docVersion="9" lastPageNumber="65" masterDocId="FFA0FF84FFF45178FFF8F7201803E446" masterDocTitle="A new species of Leptolalax (Anura: Megophryidae) from central Vietnam" masterLastPageNumber="68" masterPageNumber="56" pageNumber="58" updateTime="1698308328421" updateUser="plazi">
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<mods:title id="132F43F3A86EB68D327DB61C391D8E26">A new species of Leptolalax (Anura: Megophryidae) from central Vietnam</mods:title>
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<mods:namePart id="27680E98AB6DEF0E8EA3B326C3B3DB53">Rowley, Jodi J. L.</mods:namePart>
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<mods:namePart id="7698B127651DE7D4E1EC9C6A7F806195">Hoang, Huy Duc</mods:namePart>
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<mods:namePart id="BDF8533B328786FE72EF2B2E3EAB3D15">Dau, Vinh Quang</mods:namePart>
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<mods:namePart id="50DE56B58B61F4C5CCF5C00F5DC0009D">Le, Duong Thi Thuy</mods:namePart>
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<mods:namePart id="3D94E493A37B919E1871AAEFD1BBABDD">Cao, Trung Tien</mods:namePart>
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<treatment id="039987FCFFF65171FF6FF2861D03E3FE" ID-DOI="http://doi.org/10.5281/zenodo.5684267" ID-GBIF-Taxon="119517825" ID-Zenodo-Dep="5684267" LSID="urn:lsid:plazi:treatment:039987FCFFF65171FF6FF2861D03E3FE" httpUri="http://treatment.plazi.org/id/039987FCFFF65171FF6FF2861D03E3FE" lastPageId="9" lastPageNumber="65" pageId="2" pageNumber="58">
<subSubSection id="C32A6561FFF6517AFF6FF28619C2E186" box="[151,449,1446,1472]" pageId="2" pageNumber="58" type="nomenclature">
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<emphasis id="B944EAF8FFF6517AFF6FF28619C2E186" bold="true" box="[151,449,1446,1472]" pageId="2" pageNumber="58">
<taxonomicName id="4C304D69FFF6517AFF6FF286195EE186" ID-CoL="3TQPN" box="[151,349,1446,1472]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="2" pageNumber="58" phylum="Chordata" rank="species" species="firthi" status="sp. nov.">
<emphasis id="B944EAF8FFF6517AFF6FF286195EE186" bold="true" box="[151,349,1446,1472]" italics="true" pageId="2" pageNumber="58">Leptolalax firthi</emphasis>
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</emphasis>
</heading>
</paragraph>
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: 
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R 176524, adult male, calling on fern 
<quantity id="4CC89B0FFFF6517AFCE9F2CD1B50E243" box="[785,851,1517,1541]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="58" unit="m" value="0.1">0.1 m</quantity>
from slightly disturbed, 
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wide, rocky stream with small cascades in montane evergreen forest in Ngoc Linh Nature Reserve, Kon Tum Province, 
<collectingCountry id="F327767AFFF6517AFAC5F1311D9FE26F" box="[1341,1436,1553,1577]" name="Vietnam" pageId="2" pageNumber="58">Vietnam</collectingCountry>
(
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, 
<geoCoordinate id="EE04502DFFF6517AFED7F11519BFE20B" box="[303,444,1589,1613]" direction="east" orientation="longitude" pageId="2" pageNumber="58" precision="5" value="107.8587">107.8587º E</geoCoordinate>
, 
<quantity id="4CC89B0FFFF6517AFE31F1151A22E20B" box="[457,545,1589,1613]" metricMagnitude="3" metricUnit="m" metricValue="1.716" pageId="2" pageNumber="58" unit="m" value="1716.0">1716 m</quantity>
, 
<figureCitation id="130B2A6FFFF6517AFDD6F1151A9BE20B" box="[558,664,1589,1613]" captionStart="FIGURE 1" captionStartId="2.[151,250,1343,1366]" captionTargetBox="[153,1433,197,1320]" captionTargetId="figure@2.[151,1436,195,1322]" captionTargetPageId="2" captionText="FIGURE 1. Type locality (black star) and collection localities (black circles) of Leptolalax firthi sp. nov." httpUri="https://zenodo.org/record/215102/files/figure.png" pageId="2" pageNumber="58">Figures 1</figureCitation>
, 
<figureCitation id="130B2A6FFFF6517AFD5EF1151AB7E20B" box="[678,692,1589,1613]" captionStart="FIGURE 2" captionStartId="3.[151,250,1921,1944]" captionTargetBox="[151,1410,949,1900]" captionTargetId="figure@3.[151,1436,949,1901]" captionTargetPageId="3" captionText="FIGURE 2. Adult male holotype (AMS R 176524) of L. firthi sp. nov. in life. (A) in situ, frontodorsolateral view; (B) dorsolateral view; (C) dorsal view; (D) ventral view." httpUri="https://zenodo.org/record/215103/files/figure.png" pageId="2" pageNumber="58">2</figureCitation>
&amp; 
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). Collected at 20:00 h on 
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by Jodi J. L. Rowley, Huy 
<collectingCountry id="F327767AFFF6517AFF36F17A18FFE237" box="[206,252,1626,1649]" name="Germany" pageId="2" pageNumber="58">Duc</collectingCountry>
Hoang, Duong Thi Thuy Le, and Vinh Quang Dau.
</paragraph>
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<typeStatus id="548B8848FFF6517AFF31F15D1942E2D3" box="[201,321,1661,1685]" pageId="2" pageNumber="58" type="paratype">
<emphasis id="B944EAF8FFF6517AFF31F15D1942E2D3" bold="true" box="[201,321,1661,1685]" pageId="2" pageNumber="58">Paratypes</emphasis>
</typeStatus>
: 
<collectionCode id="ED21AE2FFFF6517AFEA9F15D198DE2D3" box="[337,398,1661,1685]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176525 (adult male) calling on rock at 20:05 h, and 
<collectionCode id="ED21AE2FFFF6517AFBF5F15D1C46E2D3" box="[1037,1093,1661,1685]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="2" pageNumber="58">UNS</collectionCode>
00460/ 
<collectionCode id="ED21AE2FFFF6517AFB65F15D1CDBE2D3" box="[1181,1240,1661,1685]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176526 (adult male), collected at same geographic locality and date as 
<typeStatus id="548B8848FFF6517AFCE7F1811B86E2FF" box="[799,901,1697,1721]" pageId="2" pageNumber="58" type="holotype">holotype</typeStatus>
. 
<collectionCode id="ED21AE2FFFF6517AFC69F1811BCAE2FF" box="[913,969,1697,1721]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="2" pageNumber="58">UNS</collectionCode>
00461/ 
<collectionCode id="ED21AE2FFFF6517AFBD8F1811C5FE2FF" box="[1056,1116,1697,1721]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176527 (adult male) collected on tree root 
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from 
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wide, shallow rocky stream at 21:30 h on 
<date id="FF8E102AFFF6517AFC12F1E51C7BE29B" box="[1002,1144,1733,1757]" pageId="2" pageNumber="58" value="2010-04-04">4 April 2010</date>
at same geographic locality as 
<typeStatus id="548B8848FFF6517AFF27F1C91946E347" box="[223,325,1769,1793]" pageId="2" pageNumber="58" type="holotype">holotype</typeStatus>
. 
<collectionCode id="ED21AE2FFFF6517AFEA8F1C9198EE347" box="[336,397,1769,1793]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176513 (adult male), calling in shallow water under 
<quantity id="4CC89B0FFFF6517AFBEAF1C91C56E347" box="[1042,1109,1769,1793]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="58" unit="m" value="0.4">0.4 m</quantity>
diameter rock; 
<collectionCode id="ED21AE2FFFF6517AFAF4F1C91D47E347" box="[1292,1348,1769,1793]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="2" pageNumber="58">UNS</collectionCode>
00456/ 
<collectionCode id="ED21AE2FFFF6517AFF6FF02D18D7E363" box="[151,212,1805,1829]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176514 (adult male) calling on gravel in stream bed; and 
<collectionCode id="ED21AE2FFFF6517AFC70F02D1BC6E363" box="[904,965,1805,1829]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176515 (adult female, gravid); all collected at ~19:00 h on 
<date id="FF8E102AFFF6517AFE72F0111A35E30F" box="[394,566,1841,1865]" pageId="2" pageNumber="58" value="2010-03-27">
27 March 
<quantity id="4CC89B0FFFF6517AFE06F0111A35E30F" box="[510,566,1841,1865]" metricMagnitude="1" metricUnit="m" metricValue="5.1053999999999995" pageId="2" pageNumber="58" unit="in" value="2010.0">2010</quantity>
</date>
in a 
<quantity id="4CC89B0FFFF6517AFD95F0111AADE30F" box="[621,686,1841,1865]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="58" unit="m" value="1.5">1.5 m</quantity>
wide stream with medium flow and low gradient in montane evergreen forest in Ngoc Linh Nature Reserve, Kon Tum Province, 
<collectingCountry id="F327767AFFF6517AFCA5F0751BBFE32B" box="[861,956,1877,1901]" name="Vietnam" pageId="2" pageNumber="58">Vietnam</collectingCountry>
(
<geoCoordinate id="EE04502DFFF6517AFC32F0751C4EE32B" box="[970,1101,1877,1901]" direction="north" orientation="latitude" pageId="2" pageNumber="58" precision="5" value="15.0323">15.0323º N</geoCoordinate>
, 
<geoCoordinate id="EE04502DFFF6517AFBA1F0751CE7E32B" box="[1113,1252,1877,1901]" direction="east" orientation="longitude" pageId="2" pageNumber="58" precision="5" value="107.8153">107.8153º E</geoCoordinate>
, 
<quantity id="4CC89B0FFFF6517AFB08F0751D44E32B" box="[1264,1351,1877,1901]" metricMagnitude="3" metricUnit="m" metricValue="1.722" pageId="2" pageNumber="58" unit="m" value="1722.0">1722 m</quantity>
, 
<figureCitation id="130B2A6FFFF6517AFAABF07518A7E3D7" captionStart="FIGURE 1" captionStartId="2.[151,250,1343,1366]" captionTargetBox="[153,1433,197,1320]" captionTargetId="figure@2.[151,1436,195,1322]" captionTargetPageId="2" captionText="FIGURE 1. Type locality (black star) and collection localities (black circles) of Leptolalax firthi sp. nov." httpUri="https://zenodo.org/record/215102/files/figure.png" pageId="2" pageNumber="58">Figure 1</figureCitation>
). 
<collectionCode id="ED21AE2FFFF6517AFF45F059190FE3D7" box="[189,268,1913,1937]" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="2" pageNumber="58">NCSM</collectionCode>
78995, 
<collectionCode id="ED21AE2FFFF6517AFE93F05919ABE3D7" box="[363,424,1913,1937]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176503–176507, 
<collectionCode id="ED21AE2FFFF6517AFD68F0591ACBE3D7" box="[656,712,1913,1937]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="2" pageNumber="58">UNS</collectionCode>
00453/ 
<collectionCode id="ED21AE2FFFF6517AFCE7F0591B58E3D7" box="[799,859,1913,1937]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176509, 
<collectionCode id="ED21AE2FFFF6517AFC1AF0591C19E3D7" box="[994,1050,1913,1937]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="2" pageNumber="58">UNS</collectionCode>
00454/ 
<collectionCode id="ED21AE2FFFF6517AFB89F0591CAFE3D7" box="[1137,1196,1913,1937]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176510, and 
<collectionCode id="ED21AE2FFFF6517AFA9CF0591D9FE3D7" box="[1380,1436,1913,1937]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="2" pageNumber="58">UNS</collectionCode>
00455/ 
<collectionCode id="ED21AE2FFFF6517AFF1DF0BD1922E3F3" box="[229,289,1949,1973]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176511 (
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males) and 
<collectionCode id="ED21AE2FFFF6517AFD83F0BD1AB4E3F3" box="[635,695,1949,1973]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176500, 
<collectionCode id="ED21AE2FFFF6517AFCC2F0BD1B71E3F3" box="[826,882,1949,1973]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="2" pageNumber="58">UNS</collectionCode>
00452/ 
<collectionCode id="ED21AE2FFFF6517AFC30F0BD1C00E3F3" box="[968,1027,1949,1973]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176501, 
<collectionCode id="ED21AE2FFFF6517AFB7EF0BD1CC0E3F3" box="[1158,1219,1949,1973]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176502, 
<collectionCode id="ED21AE2FFFF6517AFABDF0BD1D81E3F3" box="[1349,1410,1949,1973]" pageId="2" pageNumber="58">AMS</collectionCode>
R 176508, and 
<collectionCode id="ED21AE2FFFF6517AFED2F0E11964E39F" box="[298,359,1985,2009]" pageId="2" pageNumber="58">AMS</collectionCode>
R 1765012 (
<specimenCount id="9D36FD63FFF6517AFE03F0E11A4AE39F" box="[507,585,1985,2009]" pageId="2" pageNumber="58" type="adult">5 adult</specimenCount>
females) collected at night in a large rocky stream with no riparian vegetation (due to storm damage) in montane evergreen forest in Ngoc Linh Nature Reserve, Kon Tum Province, 
<collectingCountry id="F327767AFFF7517BFAC5F7B71D9FE4E9" box="[1341,1436,151,175]" name="Vietnam" pageId="3" pageNumber="59">Vietnam</collectingCountry>
(
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, 
<geoCoordinate id="EE04502DFFF7517BFED7F79C19BFE492" box="[303,444,188,212]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="5" value="107.8165">107.8165º E</geoCoordinate>
, 
<quantity id="4CC89B0FFFF7517BFE32F79C1A21E492" box="[458,546,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.494" pageId="3" pageNumber="59" unit="m" value="1494.0">1494 m</quantity>
, 
<figureCitation id="130B2A6FFFF7517BFDD7F79C1A8FE492" box="[559,652,188,212]" captionStart="FIGURE 1" captionStartId="2.[151,250,1343,1366]" captionTargetBox="[153,1433,197,1320]" captionTargetId="figure@2.[151,1436,195,1322]" captionTargetPageId="2" captionText="FIGURE 1. Type locality (black star) and collection localities (black circles) of Leptolalax firthi sp. nov." httpUri="https://zenodo.org/record/215102/files/figure.png" pageId="3" pageNumber="59">Figure 1</figureCitation>
) on 
<date id="FF8E102AFFF7517BFD3CF79C1B76E492" box="[708,885,188,212]" pageId="3" pageNumber="59" value="2010-03-26">26 March 2010</date>
. 
<collectionCode id="ED21AE2FFFF7517BFC79F79C1BBDE492" box="[897,958,188,212]" pageId="3" pageNumber="59">AMS</collectionCode>
R 176517, 
<collectionCode id="ED21AE2FFFF7517BFBBCF79C1C90E492" box="[1092,1171,188,212]" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="3" pageNumber="59">NCSM</collectionCode>
78996–78997, 
<collectionCode id="ED21AE2FFFF7517BFABCF79C1D82E492" box="[1348,1409,188,212]" pageId="3" pageNumber="59">AMS</collectionCode>
R 176518 (
<specimenCount id="9D36FD63FFF7517BFF04F7FF194FE4B1" box="[252,332,223,247]" pageId="3" pageNumber="59" type="adult">4 adult</specimenCount>
males) and 
<collectionCode id="ED21AE2FFFF7517BFE24F7FF1A17E4B1" box="[476,532,223,247]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="3" pageNumber="59">UNS</collectionCode>
00457/ 
<collectionCode id="ED21AE2FFFF7517BFD94F7FF1AABE4B1" box="[620,680,223,247]" pageId="3" pageNumber="59">AMS</collectionCode>
R 176516 (adult female), collected at night in large rocky stream with no riparian vegetation (due to storm damage) in montane evergreen forest in Ngoc Linh Nature Reserve, Kon Tum Province, 
<collectingCountry id="F327767AFFF7517BFEBCF60719A0E579" box="[324,419,295,319]" name="Vietnam" pageId="3" pageNumber="59">Vietnam</collectingCountry>
(
<geoCoordinate id="EE04502DFFF7517BFE49F6071A37E579" box="[433,564,295,319]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="5" value="15.0407">15.0407º N</geoCoordinate>
, 
<geoCoordinate id="EE04502DFFF7517BFDB8F6071ACFE579" box="[576,716,295,319]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="5" value="107.8165">107.8165º E</geoCoordinate>
, 
<quantity id="4CC89B0FFFF7517BFD20F6071B2CE579" box="[728,815,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.494" pageId="3" pageNumber="59" unit="m" value="1494.0">1494 m</quantity>
, 
<figureCitation id="130B2A6FFFF7517BFCC3F6071B94E579" box="[827,919,295,319]" captionStart="FIGURE 1" captionStartId="2.[151,250,1343,1366]" captionTargetBox="[153,1433,197,1320]" captionTargetId="figure@2.[151,1436,195,1322]" captionTargetPageId="2" captionText="FIGURE 1. Type locality (black star) and collection localities (black circles) of Leptolalax firthi sp. nov." httpUri="https://zenodo.org/record/215102/files/figure.png" pageId="3" pageNumber="59">Figure 1</figureCitation>
) on 
<date id="FF8E102AFFF7517BFC34F6071C7FE579" box="[972,1148,295,319]" pageId="3" pageNumber="59" value="2010-03-28">28 March 2010</date>
. 
<collectionCode id="ED21AE2FFFF7517BFB7EF6071CC0E579" box="[1158,1219,295,319]" pageId="3" pageNumber="59">AMS</collectionCode>
R 176520, 
<collectionCode id="ED21AE2FFFF7517BFABDF6071D81E579" box="[1349,1410,295,319]" pageId="3" pageNumber="59">AMS</collectionCode>
R 176522 (
<specimenCount id="9D36FD63FFF7517BFF03F66C194AE522" box="[251,329,332,356]" pageId="3" pageNumber="59" type="adult">2 adult</specimenCount>
males, calling) and 
<collectionCode id="ED21AE2FFFF7517BFDD6F66C1A65E522" box="[558,614,332,356]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="3" pageNumber="59">UNS</collectionCode>
00458/ 
<collectionCode id="ED21AE2FFFF7517BFD44F66C1AFBE522" box="[700,760,332,356]" pageId="3" pageNumber="59">AMS</collectionCode>
R 176519, 
<collectionCode id="ED21AE2FFFF7517BFC84F66C1BBAE522" box="[892,953,332,356]" pageId="3" pageNumber="59">AMS</collectionCode>
R 176521, 
<collectionCode id="ED21AE2FFFF7517BFBC5F66C1C76E522" box="[1085,1141,332,356]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="3" pageNumber="59">UNS</collectionCode>
00459/ 
<collectionCode id="ED21AE2FFFF7517BFB34F66C1D0BE522" box="[1228,1288,332,356]" pageId="3" pageNumber="59">AMS</collectionCode>
R 176523 (
<specimenCount id="9D36FD63FFF7517BFA76F66C18CCE5C1" pageId="3" pageNumber="59" type="adult">3 adult</specimenCount>
females) collected at night in large rocky stream with no riparian vegetation (due to storm damage) in montane evergreen forest in Ngoc Linh Nature Reserve, Kon Tum Province, 
<collectingCountry id="F327767AFFF7517BFC35F6B41C2FE5EA" box="[973,1068,404,428]" name="Vietnam" pageId="3" pageNumber="59">Vietnam</collectingCountry>
(
<geoCoordinate id="EE04502DFFF7517BFBC3F6B41CBCE5EA" box="[1083,1215,404,428]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="5" value="15.0407">15.0407º N</geoCoordinate>
, 
<geoCoordinate id="EE04502DFFF7517BFB33F6B41D5BE5EA" box="[1227,1368,404,428]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="5" value="107.8165">107.8165º E</geoCoordinate>
, 
<quantity id="4CC89B0FFFF7517BFA9CF6B418ACE589" metricMagnitude="3" metricUnit="m" metricValue="1.494" pageId="3" pageNumber="59" unit="m" value="1494.0">1494 m</quantity>
, 
<figureCitation id="130B2A6FFFF7517BFF44F697191AE589" box="[188,281,439,463]" captionStart="FIGURE 1" captionStartId="2.[151,250,1343,1366]" captionTargetBox="[153,1433,197,1320]" captionTargetId="figure@2.[151,1436,195,1322]" captionTargetPageId="2" captionText="FIGURE 1. Type locality (black star) and collection localities (black circles) of Leptolalax firthi sp. nov." httpUri="https://zenodo.org/record/215102/files/figure.png" pageId="3" pageNumber="59">Figure 1</figureCitation>
) on 
<date id="FF8E102AFFF7517BFEA9F6971A00E589" box="[337,515,439,463]" pageId="3" pageNumber="59" value="2010-03-30">30 March 2010</date>
. 
<collectionCode id="ED21AE2FFFF7517BFDF6F6971A48E589" box="[526,587,439,463]" pageId="3" pageNumber="59">AMS</collectionCode>
R 173736 (adult female), collected at 21:05 h in 
<quantity id="4CC89B0FFFF7517BFB86F6971CA9E589" box="[1150,1194,439,463]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="59" unit="m" value="3.0">3 m</quantity>
wide, shallow, rocky stream in montane evergreen forest in Ngoc Linh Nature Reserve, Kon Tum Province, 
<collectingCountry id="F327767AFFF7517BFB66F6FC1CFEE5B2" box="[1182,1277,476,500]" name="Vietnam" pageId="3" pageNumber="59">Vietnam</collectingCountry>
(
<geoCoordinate id="EE04502DFFF7517BFAE8F6FC1D9BE5B2" box="[1296,1432,476,500]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="5" value="15.2359">15.2359º N</geoCoordinate>
, 
<geoCoordinate id="EE04502DFFF7517BFF6FF6DF1920E651" box="[151,291,511,535]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="5" value="107.7098">107.7098º E</geoCoordinate>
, 
<quantity id="4CC89B0FFFF7517BFEC8F6DF198BE651" box="[304,392,511,535]" metricMagnitude="3" metricUnit="m" metricValue="1.316" pageId="3" pageNumber="59" unit="m" value="1316.0">1316 m</quantity>
, 
<figureCitation id="130B2A6FFFF7517BFE6DF6DF19F1E651" box="[405,498,511,535]" captionStart="FIGURE 1" captionStartId="2.[151,250,1343,1366]" captionTargetBox="[153,1433,197,1320]" captionTargetId="figure@2.[151,1436,195,1322]" captionTargetPageId="2" captionText="FIGURE 1. Type locality (black star) and collection localities (black circles) of Leptolalax firthi sp. nov." httpUri="https://zenodo.org/record/215102/files/figure.png" pageId="3" pageNumber="59">Figure 1</figureCitation>
) on 
<date id="FF8E102AFFF7517BFDD1F6DF1ABCE651" box="[553,703,511,535]" pageId="3" pageNumber="59" value="2009-07-12">12 July 2009</date>
. 
<collectionCode id="ED21AE2FFFF7517BFD32F6DF1B04E651" box="[714,775,511,535]" pageId="3" pageNumber="59">AMS</collectionCode>
R 173774 (sub-adult female), collected in 
<quantity id="4CC89B0FFFF7517BFB15F6DF1D1BE651" box="[1261,1304,511,535]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="59" unit="m" value="3.0">3 m</quantity>
wide, shallow, rocky stream in montane evergreen forest in Ngoc Linh Nature Reserve, Kon Tum Province, 
<collectingCountry id="F327767AFFF7517BFAC5F5041D9FE67A" box="[1341,1436,548,572]" name="Vietnam" pageId="3" pageNumber="59">Vietnam</collectingCountry>
(
<geoCoordinate id="EE04502DFFF7517BFF66F5671926E619" box="[158,293,583,607]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="5" value="15.2359">15.2359º N</geoCoordinate>
, 
<geoCoordinate id="EE04502DFFF7517BFECCF56719C0E619" box="[308,451,583,607]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="5" value="107.7098">107.7098º E</geoCoordinate>
, 
<quantity id="4CC89B0FFFF7517BFE2BF5671A2EE619" box="[467,557,583,607]" metricMagnitude="3" metricUnit="m" metricValue="1.316" pageId="3" pageNumber="59" unit="m" value="1316.0">1316 m</quantity>
, 
<figureCitation id="130B2A6FFFF7517BFDC4F5671A9FE619" box="[572,668,583,607]" captionStart="FIGURE 1" captionStartId="2.[151,250,1343,1366]" captionTargetBox="[153,1433,197,1320]" captionTargetId="figure@2.[151,1436,195,1322]" captionTargetPageId="2" captionText="FIGURE 1. Type locality (black star) and collection localities (black circles) of Leptolalax firthi sp. nov." httpUri="https://zenodo.org/record/215102/files/figure.png" pageId="3" pageNumber="59">Figure 1</figureCitation>
) on 
<date id="FF8E102AFFF7517BFD20F5671B77E619" box="[728,884,583,607]" pageId="3" pageNumber="59" value="2009-07-13">13 July 2009</date>
. 
<collectionCode id="ED21AE2FFFF7517BFC79F5671BBDE619" box="[897,958,583,607]" pageId="3" pageNumber="59">AMS</collectionCode>
R 171722 (adult female), collected from medium gradient, narrow, rocky stream in montane evergreen forest in Song Thanh Nature Reserve, Quang Nam Province, 
<collectingCountry id="F327767AFFF7517BFEF1F5AF196BE6E1" box="[265,360,655,679]" name="Vietnam" pageId="3" pageNumber="59">Vietnam</collectingCountry>
(
<geoCoordinate id="EE04502DFFF7517BFE8FF5AF19F8E6E1" box="[375,507,655,679]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="5" value="15.3293">15.3293º N</geoCoordinate>
, 
<geoCoordinate id="EE04502DFFF7517BFDFFF5AF1A90E6E1" box="[519,659,655,679]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="5" value="107.7043">107.7043º E</geoCoordinate>
, 
<quantity id="4CC89B0FFFF7517BFD67F5AF1AEAE6E1" box="[671,745,655,679]" metricMagnitude="2" metricUnit="m" metricValue="8.629999999999999" pageId="3" pageNumber="59" unit="m" value="863.0">863 m</quantity>
, 
<figureCitation id="130B2A6FFFF7517BFD0DF5AF1B51E6E1" box="[757,850,655,679]" captionStart="FIGURE 1" captionStartId="2.[151,250,1343,1366]" captionTargetBox="[153,1433,197,1320]" captionTargetId="figure@2.[151,1436,195,1322]" captionTargetPageId="2" captionText="FIGURE 1. Type locality (black star) and collection localities (black circles) of Leptolalax firthi sp. nov." httpUri="https://zenodo.org/record/215102/files/figure.png" pageId="3" pageNumber="59">Figure 1</figureCitation>
) at 18:30 h on 
<date id="FF8E102AFFF7517BFBF8F5AF1C95E6E1" box="[1024,1174,655,679]" pageId="3" pageNumber="59" value="2007-07-26">26 July 2007</date>
. 
<collectionCode id="ED21AE2FFFF7517BFB58F5AF1CDEE6E1" box="[1184,1245,655,679]" pageId="3" pageNumber="59">AMS</collectionCode>
R 171714 (adult female), collected on leaf-litter 
<quantity id="4CC89B0FFFF7517BFDFAF5941A2CE68A" box="[514,559,692,716]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="59" unit="m" value="4.0">4 m</quantity>
from wide, low gradient, rocky stream in montane evergreen forest in Song Thanh Nature Reserve, Quang Nam Province, 
<collectingCountry id="F327767AFFF7517BFD5BF5F71B01E6A9" box="[675,770,727,751]" name="Vietnam" pageId="3" pageNumber="59">Vietnam</collectingCountry>
(
<geoCoordinate id="EE04502DFFF7517BFCE8F5F71B97E6A9" box="[784,916,727,751]" direction="north" orientation="latitude" pageId="3" pageNumber="59" precision="5" value="15.2681">15.2681º N</geoCoordinate>
, 
<geoCoordinate id="EE04502DFFF7517BFC67F5F71C28E6A9" box="[927,1067,727,751]" direction="east" orientation="longitude" pageId="3" pageNumber="59" precision="5" value="107.7546">107.7546º E</geoCoordinate>
, 
<quantity id="4CC89B0FFFF7517BFBC0F5F71C8DE6A9" box="[1080,1166,727,751]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="3" pageNumber="59" unit="m" value="1125.0">1125 m</quantity>
, 
<figureCitation id="130B2A6FFFF7517BFB62F5F71CF5E6A9" box="[1178,1270,727,751]" captionStart="FIGURE 1" captionStartId="2.[151,250,1343,1366]" captionTargetBox="[153,1433,197,1320]" captionTargetId="figure@2.[151,1436,195,1322]" captionTargetPageId="2" captionText="FIGURE 1. Type locality (black star) and collection localities (black circles) of Leptolalax firthi sp. nov." httpUri="https://zenodo.org/record/215102/files/figure.png" pageId="3" pageNumber="59">Figure 1</figureCitation>
) at 19:20 h on 
<date id="FF8E102AFFF7517BFF6FF5DC192FE752" box="[151,300,764,788]" pageId="3" pageNumber="59" value="2007-07-22">22 July 2007</date>
. Specimens from 
<date id="FF8E102AFFF7517BFE0DF5DC1A71E752" box="[501,626,764,788]" pageId="3" pageNumber="59" value="2009" valueMax="2010">2009–2010</date>
were collected by Jodi J. L. Rowley, Huy 
<collectingCountry id="F327767AFFF7517BFBA9F5DD1C83E752" box="[1105,1152,765,788]" name="Germany" pageId="3" pageNumber="59">Duc</collectingCountry>
Hoang, Duong Thi Thuy Le, Vinh Quang Dau. Specimens from 2007 were collected by Jodi J. L. Rowley, and Trung Tien Cao.
</paragraph>
</subSubSection>
<subSubSection id="C32A6561FFF7517BFF31F4641AE9E739" pageId="3" pageNumber="59" type="etymology">
<paragraph id="8B8F36EAFFF7517BFF31F4641AE9E739" blockId="3.[151,1437,151,895]" pageId="3" pageNumber="59">
<emphasis id="B944EAF8FFF7517BFF31F4641949E71A" bold="true" box="[201,330,836,860]" pageId="3" pageNumber="59">Etymology</emphasis>
: Specific epithet is a patronym honouring Denys Firth, for his support of amphibian biodiversity conservation and scientific capacity building in Asia.
</paragraph>
</subSubSection>
<caption id="DF4F6662FFF7517BFF6FF0A11A5AE3F1" httpUri="https://zenodo.org/record/215103/files/figure.png" pageId="3" pageNumber="59" targetBox="[151,1410,949,1900]" targetPageId="3">
<paragraph id="8B8F36EAFFF7517BFF6FF0A11A5AE3F1" blockId="3.[151,1436,1921,1975]" pageId="3" pageNumber="59">
<emphasis id="B944EAF8FFF7517BFF6FF0A11911E3D1" bold="true" box="[151,274,1921,1944]" pageId="3" pageNumber="59">FIGURE 2.</emphasis>
Adult male holotype (AMS R 176524) of 
<taxonomicName id="4C304D69FFF7517BFD46F0A31B08E3DE" box="[702,779,1922,1944]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="59" phylum="Chordata" rank="species" species="firthi" status="sp. nov.">
<emphasis id="B944EAF8FFF7517BFD46F0A31B08E3DE" box="[702,779,1922,1944]" italics="true" pageId="3" pageNumber="59">L. firthi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775783FFF7517BFCE8F0A21B63E3DE" box="[784,864,1922,1944]" pageId="3" pageNumber="59" rank="species">
<emphasis id="B944EAF8FFF7517BFCE8F0A21B58E3D1" bold="true" box="[784,859,1922,1943]" pageId="3" pageNumber="59">sp. nov</emphasis>
.
</taxonomicNameLabel>
in life. (A) 
<emphasis id="B944EAF8FFF7517BFC2FF0A31C17E3DE" box="[983,1044,1923,1944]" italics="true" pageId="3" pageNumber="59">in situ</emphasis>
, frontodorsolateral view; (B) dorsolateral view; (C) dorsal view; (D) ventral view.
</paragraph>
</caption>
<caption id="DF4F6662FFF0517CFF6FF38C1A9DE17B" httpUri="https://zenodo.org/record/215104/files/figure.png" pageId="4" pageNumber="60" targetBox="[154,1414,188,1171]" targetPageId="4">
<paragraph id="8B8F36EAFFF0517CFF6FF38C1A9DE17B" blockId="4.[151,1437,1196,1341]" pageId="4" pageNumber="60">
<emphasis id="B944EAF8FFF0517CFF6FF38C1910E084" bold="true" box="[151,275,1196,1219]" pageId="4" pageNumber="60">FIGURE 3.</emphasis>
(A) Adult male holotype of 
<taxonomicName id="4C304D69FFF0517CFDCEF38E1A87E085" box="[566,644,1197,1219]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="60" phylum="Chordata" rank="species" species="firthi" status="sp. nov.">
<emphasis id="B944EAF8FFF0517CFDCEF38E1A87E085" box="[566,644,1197,1219]" italics="true" pageId="4" pageNumber="60">L. firthi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775783FFF0517CFD73F38D1ADFE084" box="[651,732,1197,1218]" pageId="4" pageNumber="60" rank="species">
<emphasis id="B944EAF8FFF0517CFD73F38D1AD5E084" bold="true" box="[651,726,1197,1218]" pageId="4" pageNumber="60">sp. nov</emphasis>
.
</taxonomicNameLabel>
AMS R 176524 in preservative (i) dorsal view, (ii) ventral view, (iii) ventral view of left hand showing lateral dermal fringes on Finger II, and (iv) ventral view of left foot showing wide lateral dermal fringes on all toes. (B) Adult female paratype of 
<taxonomicName id="4C304D69FFF0517CFD53F3CB1AFBE146" box="[683,760,1258,1280]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="60" phylum="Chordata" rank="species" species="firthi" status="sp. nov.">
<emphasis id="B944EAF8FFF0517CFD53F3CB1AFBE146" box="[683,760,1258,1280]" italics="true" pageId="4" pageNumber="60">L. firthi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775783FFF0517CFD06F3CA1B4DE146" box="[766,846,1258,1280]" pageId="4" pageNumber="60" rank="species">
<emphasis id="B944EAF8FFF0517CFD06F3CA1B4BE0B9" bold="true" box="[766,840,1258,1279]" pageId="4" pageNumber="60">sp. nov</emphasis>
.
</taxonomicNameLabel>
AMS R 176515 in preservative (i) dorsal view, (ii) ventral view, (iii) ventral view of left hand with Finger II lacking lateral dermal fringes, and (iv) ventral view of left foot showing poorly-developed lateral dermal fringes on all toes.
</paragraph>
</caption>
<subSubSection id="C32A6561FFF0517CFF31F24A1D5BE2E4" pageId="4" pageNumber="60" type="diagnosis">
<paragraph id="8B8F36EAFFF0517CFF31F24A1D5BE2E4" blockId="4.[151,1437,1386,2022]" pageId="4" pageNumber="60">
<emphasis id="B944EAF8FFF0517CFF31F24A193FE1C5" bold="true" box="[201,316,1386,1411]" pageId="4" pageNumber="60">Diagnosis</emphasis>
: Assigned to the genus 
<taxonomicName id="4C304D69FFF0517CFDB1F24A1AC2E1C4" box="[585,705,1386,1410]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="60" phylum="Chordata" rank="genus">
<emphasis id="B944EAF8FFF0517CFDB1F24A1AC2E1C4" box="[585,705,1386,1410]" italics="true" pageId="4" pageNumber="60">Leptolalax</emphasis>
</taxonomicName>
on the basis of the following: small size, rounded finger tips, the presence of an elevated inner palmar tubercle not continuous to the thumb, presence of macroglands on body (including supra-axillary, pectoral, femoral and ventrolateral glands), vomerine teeth absent, tubercles on eyelids, anterior tip of snout with pale vertical bar (
<bibRefCitation id="EFA14B1BFFF0517CFD74F2F61B27E1A8" author="Dubois" box="[652,804,1494,1518]" pageId="4" pageNumber="60" refString="Dubois, A. (1983) Note preliminaire sur le genre Leptolalax Dubois, 1980 (Amphibiens, Anoures), avec diagnose d'une espece novelle du Vietnam. Alytes, 2, 147 - 153." type="journal article" year="1983">Dubois 1983</bibRefCitation>
; 
<bibRefCitation id="EFA14B1BFFF0517CFCCAF2F61C17E1A8" author="Lathrop" box="[818,1044,1494,1518]" pageId="4" pageNumber="60" refString="Lathrop, A., Murphy, R. W., Orlov, N. &amp; Ho, C. T. (1998) Two new species of Leptolalax (Anura: Megophryidae) from northern Vietnam. Amphibia-Reptilia, 19, 253 - 267." type="journal article" year="1998">
Lathrop 
<emphasis id="B944EAF8FFF0517CFC6CF2F71BCBE1A8" box="[916,968,1494,1518]" italics="true" pageId="4" pageNumber="60">et al</emphasis>
. 1998
</bibRefCitation>
; 
<bibRefCitation id="EFA14B1BFFF0517CFBDAF2F61D0CE1A8" author="Delorme" box="[1058,1295,1494,1518]" pageId="4" pageNumber="60" refString="Delorme, M., Dubois, A., Grosjean, S., &amp; Ohler, A. (2006) Une nouvelle ergotaxinomie des Megophryidae (Amphibia, Anura). Alytes, 24, 6 - 21." type="journal article" year="2006">
Delorme 
<emphasis id="B944EAF8FFF0517CFB76F2F71CC2E1A8" box="[1166,1217,1494,1518]" italics="true" pageId="4" pageNumber="60">et al</emphasis>
. 2006
</bibRefCitation>
). 
<taxonomicName id="4C304D69FFF0517CFADCF2F618CCE254" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="60" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFF0517CFADCF2F618CCE254" italics="true" pageId="4" pageNumber="60">Leptolalax firthi</emphasis>
</taxonomicName>
is distinguished from its congeners by a combination of (1) an absence of distinct dark brown/black dorsolateral markings, (2) toes with rudimentary webbing, (3) toes with wide lateral dermal fringes in males and weak or absent lateral dermal fringes in females, (4) most males with wide lateral dermal fringes on Finger II, (5) medium size for the genus (
<quantity id="4CC89B0FFFF0517CFE91F1461A0EE238" box="[361,525,1638,1662]" metricMagnitude="-2" metricUnit="m" metricValue="2.78" metricValueMax="2.92" metricValueMin="2.6399999999999997" pageId="4" pageNumber="60" unit="mm" value="27.799999999999997" valueMax="29.2" valueMin="26.4">26.4–29.2 mm</quantity>
in 
<specimenCount id="9D36FD63FFF0517CFDC8F1461A89E238" box="[560,650,1638,1662]" pageId="4" pageNumber="60" type="adult">21 adult</specimenCount>
males, 
<quantity id="4CC89B0FFFF0517CFD18F1461B81E238" box="[736,898,1638,1662]" metricMagnitude="-2" metricUnit="m" metricValue="3.13" metricValueMax="3.69" metricValueMin="2.57" pageId="4" pageNumber="60" unit="mm" value="31.299999999999997" valueMax="36.9" valueMin="25.7">25.7–36.9 mm</quantity>
in 
<specimenCount id="9D36FD63FFF0517CFC5DF1461C27E238" box="[933,1060,1638,1662]" pageId="4" pageNumber="60" type="female">14 females</specimenCount>
), (6) near immaculate white chest and belly, and (7) a call consisting of 2–5 notes with a dominant frequency of 5.4–6.6 kHz (at 18.3–21.2º C).
</paragraph>
</subSubSection>
<subSubSection id="C32A6561FFF05171FF31F18E1D03E3FE" lastPageId="9" lastPageNumber="65" pageId="4" pageNumber="60" type="description">
<paragraph id="8B8F36EAFFF0517DFF31F18E1B63E5EA" blockId="4.[151,1437,1386,2022]" lastBlockId="5.[151,1437,151,428]" lastPageId="5" lastPageNumber="61" pageId="4" pageNumber="60">
<emphasis id="B944EAF8FFF0517CFF31F18E19E1E280" bold="true" box="[201,482,1710,1735]" pageId="4" pageNumber="60">
Description of 
<typeStatus id="548B8848FFF0517CFE84F18E19E1E280" box="[380,482,1710,1734]" pageId="4" pageNumber="60" type="holotype">holotype</typeStatus>
</emphasis>
: Head slightly wider than long; snout rounded with slight point in dorsal view and rounded but relatively truncate in profile, projecting slightly beyond margin of the lower jaw; nostril closer to tip of snout than eye; canthus rostralis distinct, bluntly rounded; lores slightly concave; vertical pupil; eye diameter smaller than snout length; tympanum distinct, round, diameter smaller than that of the eye; tympanic rim elevated relative to skin of temporal region; vomerine teeth absent; pineal ocellus absent; vocal sac openings slit-like, located posteriolaterally on floor of mouth; tongue long, wide, with deep notch at posterior tip; supratympanic ridge distinct, running from eye towards axillary gland, with raised tubercles. Tips of fingers rounded, slightly swollen; relative finger lengths I &lt;II = IV &lt;III; nuptial pad absent; subarticular tubercles absent; a large, round inner palmar tubercle distinctly separated from small, laterally compressed outer palmar tubercle; no finger webbing, moderate lateral dermal fringes on Finger II only (
<figureCitation id="130B2A6FFFF1517DFCF0F7B71B64E4E9" box="[776,871,151,175]" captionStart="FIGURE 3" captionStartId="4.[151,250,1196,1219]" captionTargetBox="[154,1414,188,1171]" captionTargetId="figure@4.[151,1436,175,1175]" captionTargetPageId="4" captionText="FIGURE 3. (A) Adult male holotype of L. firthi sp. nov. AMS R 176524 in preservative (i) dorsal view, (ii) ventral view, (iii) ventral view of left hand showing lateral dermal fringes on Finger II, and (iv) ventral view of left foot showing wide lateral dermal fringes on all toes. (B) Adult female paratype of L. firthi sp. nov. AMS R 176515 in preservative (i) dorsal view, (ii) ventral view, (iii) ventral view of left hand with Finger II lacking lateral dermal fringes, and (iv) ventral view of left foot showing poorly-developed lateral dermal fringes on all toes." httpUri="https://zenodo.org/record/215104/files/figure.png" pageId="5" pageNumber="61">Figure 3</figureCitation>
Aiii). Tips of toes like fingers; relative toe length I &lt;II &lt;V &lt;III &lt;IV; subarticular tubercles absent, replaced by dermal ridges, distinct on toes II–V; large, oval inner metatarsal tubercle pronounced, outer metatarsal tubercle absent; basal webbing; wide lateral fringes (
<figureCitation id="130B2A6FFFF1517DFAA9F7FF18A7E55A" captionStart="FIGURE 3" captionStartId="4.[151,250,1196,1219]" captionTargetBox="[154,1414,188,1171]" captionTargetId="figure@4.[151,1436,175,1175]" captionTargetPageId="4" captionText="FIGURE 3. (A) Adult male holotype of L. firthi sp. nov. AMS R 176524 in preservative (i) dorsal view, (ii) ventral view, (iii) ventral view of left hand showing lateral dermal fringes on Finger II, and (iv) ventral view of left foot showing wide lateral dermal fringes on all toes. (B) Adult female paratype of L. firthi sp. nov. AMS R 176515 in preservative (i) dorsal view, (ii) ventral view, (iii) ventral view of left hand with Finger II lacking lateral dermal fringes, and (iv) ventral view of left foot showing poorly-developed lateral dermal fringes on all toes." httpUri="https://zenodo.org/record/215104/files/figure.png" pageId="5" pageNumber="61">Figure 3</figureCitation>
Aiv). Tibia 50% of snout-vent length; tibiotarsal articulation reaches anterior margin of eye. Skin on dorsum shagreened, with fine, round, relatively evenly scattered tubercles; ventral skin smooth; pectoral gland oval, 1.0 mm diameter, relatively indistinct in preservative; femoral gland oval, approximately 
<quantity id="4CC89B0FFFF1517DFBD0F66C1C7DE522" box="[1064,1150,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="61" unit="mm" value="1.2">1.2 mm</quantity>
diameter, on posteroventral surface of thigh, closer to knee than to vent; supra-axillary gland raised, 
<quantity id="4CC89B0FFFF1517DFC08F64F1C45E5C1" box="[1008,1094,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="61" unit="mm" value="1.1">1.1 mm</quantity>
diameter. Ventrolateral glands present, dorsolaterally compressed, forming an incomplete line.
</paragraph>
<caption id="DF4F6662FFF1517DFF6FF0BF1989E3B4" httpUri="https://zenodo.org/record/215105/files/figure.png" pageId="5" pageNumber="61" targetBox="[152,1436,470,1928]" targetPageId="5">
<paragraph id="8B8F36EAFFF1517DFF6FF0BF1989E3B4" blockId="5.[151,1436,1951,2034]" pageId="5" pageNumber="61">
<emphasis id="B944EAF8FFF1517DFF6FF0BF1910E3F2" bold="true" box="[151,275,1951,1974]" pageId="5" pageNumber="61">FIGURE 4.</emphasis>
Paratypes of 
<taxonomicName id="4C304D69FFF1517DFE58F08019EDE3F3" box="[416,494,1951,1973]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="61" phylum="Chordata" rank="species" species="firthi" status="sp. nov.">
<emphasis id="B944EAF8FFF1517DFE58F08019EDE3F3" box="[416,494,1951,1973]" italics="true" pageId="5" pageNumber="61">L. firthi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775783FFF1517DFE0DF0801A45E3F3" box="[501,582,1952,1973]" pageId="5" pageNumber="61" rank="species">
<emphasis id="B944EAF8FFF1517DFE0DF0801A43E3F3" bold="true" box="[501,576,1952,1973]" pageId="5" pageNumber="61">sp. nov</emphasis>
.
</taxonomicNameLabel>
in life. (A) adult male AMS R 176506, (B) adult female AMS R 176502, (C) adult female AMS R 176515, (D) adult female UNS 00458/AMS R 176519, (E) adult female AMS R 173736, and (F) sub-adult female AMS R 173774.
</paragraph>
</caption>
<paragraph id="8B8F36EAFFF2517EFF31F7B71CE1E5EA" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="62">
<emphasis id="B944EAF8FFF2517EFF31F7B71A0DE4F6" bold="true" box="[201,526,151,176]" pageId="6" pageNumber="62">
Colour of 
<typeStatus id="548B8848FFF2517EFEB3F7B819B1E4F6" box="[331,434,152,176]" pageId="6" pageNumber="62" type="holotype">holotype</typeStatus>
in life:
</emphasis>
Dorsal surface brown with no distinct darker markings, only faint transverse brownish grey bars on the dorsal surface of the thighs, tibia, tarsus, lower arms, fingers and toes; tiny white flecks scattered on back (
<figureCitation id="130B2A6FFFF2517EFE91F7FF19C8E4B1" box="[361,459,223,247]" captionStart="FIGURE 2" captionStartId="3.[151,250,1921,1944]" captionTargetBox="[151,1410,949,1900]" captionTargetId="figure@3.[151,1436,949,1901]" captionTargetPageId="3" captionText="FIGURE 2. Adult male holotype (AMS R 176524) of L. firthi sp. nov. in life. (A) in situ, frontodorsolateral view; (B) dorsolateral view; (C) dorsal view; (D) ventral view." httpUri="https://zenodo.org/record/215103/files/figure.png" pageId="6" pageNumber="62">Figure 2</figureCitation>
). The dorsolateral surfaces of body, fingers, toes and elbow to upper arm are pale copper. Ventral surfaces of chest and belly opaque creamy white; throat transparent pink with brown dusting along anterior margin. Ventral surface of arms transparent pink with brown dusting along lateral margins; ventral surface of thighs, tibia, and tarsus brownish pink with faint cream flecking. Supra-axillary gland pale copper; pectoral glands opaque cream; femoral glands pale copper, lined with dark brown. Iris bright gold, slightly coppery orange in upper half; minute, black reticulations throughout. Iris periphery lined with black. Sclera white.
</paragraph>
<paragraph id="8B8F36EAFFF2517EFF31F6971B9CE651" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="62">
<emphasis id="B944EAF8FFF2517EFF31F6971A74E596" bold="true" box="[201,631,439,464]" pageId="6" pageNumber="62">
Colour of 
<typeStatus id="548B8848FFF2517EFEB0F69819ACE596" box="[328,431,440,464]" pageId="6" pageNumber="62" type="holotype">holotype</typeStatus>
in preservative:
</emphasis>
Dorsum dark brown with slightly paler limbs. Banding on limbs and white flecks on dorsum are more pronounced (
<figureCitation id="130B2A6FFFF2517EFD50F6FC1B0AE5B2" box="[680,777,476,500]" captionStart="FIGURE 3" captionStartId="4.[151,250,1196,1219]" captionTargetBox="[154,1414,188,1171]" captionTargetId="figure@4.[151,1436,175,1175]" captionTargetPageId="4" captionText="FIGURE 3. (A) Adult male holotype of L. firthi sp. nov. AMS R 176524 in preservative (i) dorsal view, (ii) ventral view, (iii) ventral view of left hand showing lateral dermal fringes on Finger II, and (iv) ventral view of left foot showing wide lateral dermal fringes on all toes. (B) Adult female paratype of L. firthi sp. nov. AMS R 176515 in preservative (i) dorsal view, (ii) ventral view, (iii) ventral view of left hand with Finger II lacking lateral dermal fringes, and (iv) ventral view of left foot showing poorly-developed lateral dermal fringes on all toes." httpUri="https://zenodo.org/record/215104/files/figure.png" pageId="6" pageNumber="62">Figure 3</figureCitation>
A). Ventral surface of chest, belly, throat, interior portions of arms and thighs are creamy white. Macrogrands are creamy white.
</paragraph>
<paragraph id="8B8F36EAFFF2517EFF31F5041A6FE626" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="62">
<emphasis id="B944EAF8FFF2517EFF31F504197BE67A" bold="true" box="[201,376,548,572]" pageId="6" pageNumber="62">Measurements</emphasis>
: 
<typeStatus id="548B8848FFF2517EFE7FF50419F7E67A" box="[391,500,548,572]" pageId="6" pageNumber="62" type="holotype">Holotype</typeStatus>
: SVL 28.9, HDL 9.6, HDW 11.0, SNT 4.3, EYE 3.6, IOD 3.5, TMP 1.6, TEY 1.4, TIB 14.5, ML 8.4, PL 13.8, weight 
<quantity id="4CC89B0FFFF2517EFDDEF5671A6FE626" box="[550,620,583,608]" metricMagnitude="-3" metricUnit="kg" metricValue="1.95" pageId="6" pageNumber="62" unit="g" value="1.95">1.95 g</quantity>
</paragraph>
<caption id="DF4F6662FFF2517EFF6FF2811BEEE217" httpUri="https://zenodo.org/record/215106/files/figure.png" pageId="6" pageNumber="62" targetBox="[154,1432,667,1417]" targetPageId="6">
<paragraph id="8B8F36EAFFF2517EFF6FF2811BEEE217" blockId="6.[151,1436,1441,1617]" pageId="6" pageNumber="62">
<emphasis id="B944EAF8FFF2517EFF6FF2811910E1F1" bold="true" box="[151,275,1441,1464]" pageId="6" pageNumber="62">FIGURE 5.</emphasis>
(A) Advertisement call of 
<taxonomicName id="4C304D69FFF2517EFDDDF2821AC8E1FE" box="[549,715,1442,1464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="62" phylum="Chordata" rank="species" species="firthi" status="sp. nov.">
<emphasis id="B944EAF8FFF2517EFDDDF2821AC8E1FE" box="[549,715,1442,1464]" italics="true" pageId="6" pageNumber="62">Leptolalax firthi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775783FFF2517EFD29F2821B21E1F1" box="[721,802,1442,1463]" pageId="6" pageNumber="62" rank="species">
<emphasis id="B944EAF8FFF2517EFD29F2821B21E1F1" bold="true" box="[721,802,1442,1463]" pageId="6" pageNumber="62">sp. nov.</emphasis>
</taxonomicNameLabel>
holotype AMS R 176524 (i) 30 s waveform of relative amplitude (Rel. amp.) over time for 23 calls, (ii) waveform and (iii) corresponding spectrogram of single representative call containing two notes, expanded from section shown in i, and (iv) power spectrum (relative amplitude vs. frequency) of first note; all recorded at an ambient air temperature of 20.2º C. (B) Representative single advertisement calls of (i) paratype AMS R 176522 recorded at an ambient air temperature of 18.8º C, (ii) paratype AMS R 176520 recorded at an ambient air temperature of 19.3º C, and (iii) paratype AMS R 176513 recorded at ambient air temperature of 21.2º C.
</paragraph>
</caption>
<paragraph id="8B8F36EAFFF2517FFF31F1581911E55A" blockId="6.[151,1437,1656,2005]" lastBlockId="7.[151,1436,151,391]" lastPageId="7" lastPageNumber="63" pageId="6" pageNumber="62">
<emphasis id="B944EAF8FFF2517EFF31F1581934E2D7" bold="true" box="[201,311,1656,1681]" pageId="6" pageNumber="62">Variation</emphasis>
: Measurements of the 
<typeStatus id="548B8848FFF2517EFDC3F15A1A68E2D4" box="[571,619,1658,1682]" pageId="6" pageNumber="62">type</typeStatus>
series are shown in 
<tableCitation id="C6B20351FFF2517EFCA9F1591BA7E2D7" box="[849,932,1657,1681]" captionStart="TABLE 1" captionStartId="7.[151,235,434,455]" captionText="TABLE 1. Measurements (mm) of Leptolalax firthi sp. nov. Abbreviations defined in text. S. D. = standard deviation. Measurements Males Females" pageId="6" pageNumber="62">Table 1</tableCitation>
and representative photographs of 
<typeStatus id="548B8848FFF2517EFAC9F15A1D9FE2D4" box="[1329,1436,1658,1682]" pageId="6" pageNumber="62" type="paratype">paratypes</typeStatus>
(in life) are shown in 
<figureCitation id="130B2A6FFFF2517EFE62F1BD19FDE2F3" box="[410,510,1693,1717]" captionStart="FIGURE 4" captionStartId="5.[151,250,1951,1974]" captionTargetBox="[152,1436,470,1928]" captionTargetId="figure@5.[151,1436,457,1930]" captionTargetPageId="5" captionText="FIGURE 4. Paratypes of L. firthi sp. nov. in life. (A) adult male AMS R 176506, (B) adult female AMS R 176502, (C) adult female AMS R 176515, (D) adult female UNS 00458 / AMS R 176519, (E) adult female AMS R 173736, and (F) sub-adult female AMS R 173774." httpUri="https://zenodo.org/record/215105/files/figure.png" pageId="6" pageNumber="62">Figure 4</figureCitation>
. Specimens vary in the extent of white speckling on the dorsum; male AMS R 176518 and to a lesser extent male AMS R 176517 and females AMS R 176515 and UNS 00459/AMS R 176523 are highly speckled; two males (AMS R 176504 and UNS 00453/AMS R 176509), and three females (AMS R 171714, AMS R 173774, and AMS R 171722) lack white specks on the dorsum entirely. In most specimens, iris colour was relatively uniformly gold or coppery gold, with minute black reticulations throughout; in some specimens (eg. UNS 00456/AMS R 176514, UNS 00458/AMS R 176519 [
<figureCitation id="130B2A6FFFF2517EFC21F0711C42E32F" box="[985,1089,1873,1897]" captionStart="FIGURE 4" captionStartId="5.[151,250,1951,1974]" captionTargetBox="[152,1436,470,1928]" captionTargetId="figure@5.[151,1436,457,1930]" captionTargetPageId="5" captionText="FIGURE 4. Paratypes of L. firthi sp. nov. in life. (A) adult male AMS R 176506, (B) adult female AMS R 176502, (C) adult female AMS R 176515, (D) adult female UNS 00458 / AMS R 176519, (E) adult female AMS R 173736, and (F) sub-adult female AMS R 173774." httpUri="https://zenodo.org/record/215105/files/figure.png" pageId="6" pageNumber="62">Figure 4</figureCitation>
D], AMS R 176521, AMS R 176524), the upper third to half of the iris was slightly orange or reddish orange in life. A faint silvery grey horizontal band along the middle of the iris was also indistinctly present in some specimens in life (eg. AMS R 176515, UNS 00458/AMS R 176519). Females AMS R 171722 and AMS R 173736 have more obvious limb banding and darker brown dorsal botches although patterning is still indistinct. Males vary in the extent of the lateral dermal fringes on Finger II, with AMS R 176522, NCSM 78995 and NCSM 78997 lacking distinct lateral dermal fringes on Finger II (two of which were observed calling, suggesting the degree of fringing may not reflect reproductive condition).
</paragraph>
<paragraph id="8B8F36EAFFF3517FFF31F60718E3E5C1" blockId="7.[151,1436,151,391]" pageId="7" pageNumber="63">
Females are generally larger and are heavier in life (all measured males were &lt;
<quantity id="4CC89B0FFFF3517FFBA0F6071C7FE506" box="[1112,1148,295,320]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="7" pageNumber="63" unit="g" value="2.0">2 g</quantity>
and females were&gt; 
<quantity id="4CC89B0FFFF3517FFA9FF6071D8DE579" box="[1383,1422,295,319]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="7" pageNumber="63" unit="g" value="2.0">2 g</quantity>
). Females have, at most, weak lateral dermal fringes on the toes and all lack lateral dermal fringes on Finger II (Figure 3).
</paragraph>
<caption id="DF4F6662FFF3517FFF6FF6921C0EE5BF" pageId="7" pageNumber="63">
<paragraph id="8B8F36EAFFF3517FFF6FF6921C0EE5BF" blockId="7.[151,1349,434,1160]" pageId="7" pageNumber="63">
<emphasis id="B944EAF8FFF3517FFF6FF69218FDE58E" bold="true" box="[151,254,434,456]" pageId="7" pageNumber="63">TABLE 1</emphasis>
. Measurements (mm) of 
<taxonomicName id="4C304D69FFF3517FFE06F6921AA7E58E" box="[510,676,434,456]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="63" phylum="Chordata" rank="species" species="firthi" status="sp. nov.">
<emphasis id="B944EAF8FFF3517FFE06F6921AA7E58E" box="[510,676,434,456]" italics="true" pageId="7" pageNumber="63">Leptolalax firthi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775783FFF3517FFD52F6931AF9E58E" box="[682,762,435,456]" pageId="7" pageNumber="63" rank="species">
<emphasis id="B944EAF8FFF3517FFD52F6931AF9E58E" bold="true" box="[682,762,435,456]" pageId="7" pageNumber="63">sp. nov.</emphasis>
</taxonomicNameLabel>
Abbreviations defined in text. S. D. = standard deviation. Measurements Males Females
</paragraph>
</caption>
<paragraph id="8B8F36EAFFF3517FFE75F52E1C68E6F6" blockId="7.[151,1349,434,1160]" pageId="7" pageNumber="63">
<emphasis id="B944EAF8FFF3517FFE75F52E1CE2E665" box="[397,1249,525,547]" italics="true" pageId="7" pageNumber="63">Range; Mean ± S. D. (N=21) Range; Mean ± S. D. (N=14)</emphasis>
SVL 26.4–29.2; 27.8 ± 0.9 25.7–36.9; 33.1 ± 2.6 HDL 9.6–12.5; 11.1 ± 0.6 11.9–14.6; 13.6 ± 1.0 HDW 10.4–11.8; 11.1 ± 0.3 10.3–14.2; 13.0 ± 1.0 SNT 3.5–4.6; 4.2 ± 0.3 4.0–5.3; 4.9 ± 0.3
</paragraph>
<paragraph id="8B8F36EAFFF3517FFF61F59E1C68E695" blockId="7.[151,1349,434,1160]" box="[153,1131,701,723]" pageId="7" pageNumber="63">EYE 3.3–4.1; 3.7 ± 0.2 3.0–4.1; 3.6 ± 0.5</paragraph>
<paragraph id="8B8F36EAFFF3517FFF61F5C11C68E6B0" blockId="7.[151,1349,434,1160]" box="[153,1131,737,759]" pageId="7" pageNumber="63">IOD 2.8–3.7; 3.2 ± 0.2 3.0–4.1; 3.6 ± 0.3</paragraph>
<paragraph id="8B8F36EAFFF3517FFF61F4251C6FE75F" blockId="7.[151,1349,434,1160]" box="[153,1132,772,794]" pageId="7" pageNumber="63">TMP 1.54–2.2; 1.7 ± 0.2 1.4–2.4; 2.1 ± 0.2</paragraph>
<paragraph id="8B8F36EAFFF3517FFF61F4081C68E77B" blockId="7.[151,1349,434,1160]" box="[153,1131,807,829]" pageId="7" pageNumber="63">TEY 0.8–1.4; 1.2 ± 0.2 1.1–3.3; 1.6 ± 0.5</paragraph>
<paragraph id="8B8F36EAFFF3517FFF61F46C1C68E7C2" blockId="7.[151,1349,434,1160]" pageId="7" pageNumber="63">TIB 12.9–14.5; 13.7 ± 0.5 13.7–17.2; 16.3 ± 0.9 ML 7.0–9.0; 7.8 ± 0.5 7.1–9.6; 8.9 ± 0.6</paragraph>
<paragraph id="8B8F36EAFFF3517FFF61F4B21C91E7E1" blockId="7.[151,1349,434,1160]" box="[153,1170,913,935]" pageId="7" pageNumber="63">PL 11.0–13.8; 13.0 ± 0.7 11.7–16.0; 14.6 ± 0.6</paragraph>
<paragraph id="8B8F36EAFFF3517FFE75F49C1C5BE7B2" blockId="7.[151,1349,434,1160]" pageId="7" pageNumber="63">
<emphasis id="B944EAF8FFF3517FFE75F49C1CABE797" box="[397,1192,955,977]" italics="true" pageId="7" pageNumber="63">Range; Median (N=21) Range; Median (N=14)</emphasis>
HDL:HDW 0.87–1.11; 1.02 0.91–1.09; 1.05
</paragraph>
<paragraph id="8B8F36EAFFF3517FFF61F3221C5BE051" blockId="7.[151,1349,434,1160]" box="[153,1112,1025,1047]" pageId="7" pageNumber="63">HDL:SVL 0.33–0.43; 0.40 0.35–0.44; 0.41</paragraph>
<paragraph id="8B8F36EAFFF3517FFF61F3051C5BE07C" blockId="7.[151,1349,434,1160]" box="[153,1112,1061,1083]" pageId="7" pageNumber="63">TIB:SVL 0.47–0.51; 0.50 0.46–0.53; 0.49</paragraph>
<paragraph id="8B8F36EAFFF3517FFE75F36F1C68E0C1" blockId="7.[151,1349,434,1160]" pageId="7" pageNumber="63">
<emphasis id="B944EAF8FFF3517FFE75F36F1CE4E022" box="[397,1255,1102,1124]" italics="true" pageId="7" pageNumber="63">Range; Mean ± S. D. (N=19) Range; Mean ± S. D. (N=12)</emphasis>
Weight in life (g) 1.5–1.9; 1.8 ± 0.1 2.2–3.4; 2.9 ± 0.6
</paragraph>
<paragraph id="8B8F36EAFFF3517FFF31F3E11AC4E1F7" blockId="7.[151,1437,1217,1458]" pageId="7" pageNumber="63">
<emphasis id="B944EAF8FFF3517FFF31F3E119AAE09C" bold="true" box="[201,425,1217,1242]" pageId="7" pageNumber="63">Advertisement call</emphasis>
: Call descriptions are based on the calls of the 
<typeStatus id="548B8848FFF3517FFC41F3E11C1DE09F" box="[953,1054,1217,1241]" pageId="7" pageNumber="63" type="holotype">holotype</typeStatus>
, recorded at 20.2º C ambient temperature. Calls were an average of 24 ms in duration and consisted of two notes (
<tableCitation id="C6B20351FFF3517FFBCAF3C51C89E0BB" box="[1074,1162,1253,1277]" captionStart="TABLE 2" captionStartId="7.[151,235,1505,1526]" captionTargetBox="[151,1436,1656,2026]" captionTargetPageId="7" captionText="TABLE 2. Measurements of advertisement call parameters for Leptolalax firthi sp. nov. Parameter values are given as means (and ranges)." httpUri="http://table.plazi.org/id/DF4F6662FFF3517FFF6FF2C1191FE250" pageId="7" pageNumber="63" tableUuid="DF4F6662FFF3517FFF6FF2C1191FE250">Table 2</tableCitation>
, 
<figureCitation id="130B2A6FFFF3517FFB60F3C51CF4E0BB" box="[1176,1271,1253,1277]" captionStart="FIGURE 5" captionStartId="6.[151,250,1441,1464]" captionTargetBox="[154,1432,667,1417]" captionTargetId="figure@6.[151,1436,661,1421]" captionTargetPageId="6" captionText="FIGURE 5. (A) Advertisement call of Leptolalax firthi sp. nov. holotype AMS R 176524 (i) 30 s waveform of relative amplitude (Rel. amp.) over time for 23 calls, (ii) waveform and (iii) corresponding spectrogram of single representative call containing two notes, expanded from section shown in i, and (iv) power spectrum (relative amplitude vs. frequency) of first note; all recorded at an ambient air temperature of 20.2 º C. (B) Representative single advertisement calls of (i) paratype AMS R 176522 recorded at an ambient air temperature of 18.8 º C, (ii) paratype AMS R 176520 recorded at an ambient air temperature of 19.3 º C, and (iii) paratype AMS R 176513 recorded at ambient air temperature of 21.2 º C." httpUri="https://zenodo.org/record/215106/files/figure.png" pageId="7" pageNumber="63">Figure 5</figureCitation>
A). Calls were highly amplitude modulated, with amplitude peaking at the start of the call and the start of each note. Notes generally contained two, rather indistinct pulses, with less distinct pulses also present (
<figureCitation id="130B2A6FFFF3517FFBC9F20D1C92E103" box="[1073,1169,1325,1349]" captionStart="FIGURE 5" captionStartId="6.[151,250,1441,1464]" captionTargetBox="[154,1432,667,1417]" captionTargetId="figure@6.[151,1436,661,1421]" captionTargetPageId="6" captionText="FIGURE 5. (A) Advertisement call of Leptolalax firthi sp. nov. holotype AMS R 176524 (i) 30 s waveform of relative amplitude (Rel. amp.) over time for 23 calls, (ii) waveform and (iii) corresponding spectrogram of single representative call containing two notes, expanded from section shown in i, and (iv) power spectrum (relative amplitude vs. frequency) of first note; all recorded at an ambient air temperature of 20.2 º C. (B) Representative single advertisement calls of (i) paratype AMS R 176522 recorded at an ambient air temperature of 18.8 º C, (ii) paratype AMS R 176520 recorded at an ambient air temperature of 19.3 º C, and (iii) paratype AMS R 176513 recorded at ambient air temperature of 21.2 º C." httpUri="https://zenodo.org/record/215106/files/figure.png" pageId="7" pageNumber="63">Figure 5</figureCitation>
Aii). The dominant frequency was 5.6–6.4 kHz, and harmonics were present at approximately 12, 18, 24 and 31 kHz. A fundamental frequency was present at approximately 0.2 kHz. Calls were repeated at a rate of approximately 0.9 calls per second, and had an average intercall interval of 
<geoCoordinate id="EE04502DFFF3517FFD72F2B91AC1E1F7" box="[650,706,1433,1457]" direction="south" orientation="latitude" pageId="7" pageNumber="63" precision="5555" value="-1.1">1.1 s</geoCoordinate>
.
</paragraph>
<caption id="DF4F6662FFF3517FFF6FF2C1191FE250" ID-Table-UUID="DF4F6662FFF3517FFF6FF2C1191FE250" httpUri="http://table.plazi.org/id/DF4F6662FFF3517FFF6FF2C1191FE250" pageId="7" pageNumber="63" targetBox="[151,1436,1656,2026]" targetIsTable="true" targetPageId="7">
<paragraph id="8B8F36EAFFF3517FFF6FF2C1191FE250" blockId="7.[151,1436,1505,1638]" pageId="7" pageNumber="63">
<emphasis id="B944EAF8FFF3517FFF6FF2C11906E1B0" bold="true" box="[151,261,1505,1526]" pageId="7" pageNumber="63">TABLE 2.</emphasis>
Measurements of advertisement call parameters for 
<taxonomicName id="4C304D69FFF3517FFCE6F2C11BC6E1B1" box="[798,965,1505,1527]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="63" phylum="Chordata" rank="species" species="firthi" status="sp. nov.">
<emphasis id="B944EAF8FFF3517FFCE6F2C11BC6E1B1" box="[798,965,1505,1527]" italics="true" pageId="7" pageNumber="63">Leptolalax firthi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775783FFF3517FFC34F2C21C1FE1B1" box="[972,1052,1506,1527]" pageId="7" pageNumber="63" rank="species">
<emphasis id="B944EAF8FFF3517FFC34F2C21C1FE1B1" bold="true" box="[972,1052,1506,1527]" pageId="7" pageNumber="63">sp. nov.</emphasis>
</taxonomicNameLabel>
Parameter values are given as means (and ranges).
</paragraph>
</caption>
<paragraph id="8B8F36EAFFF3517FFE2DF10F1D52E203" blockId="7.[151,1436,1505,1638]" box="[469,1361,1583,1605]" pageId="7" pageNumber="63">AMS R AMS R AMS R NCSM AMS R AMS R</paragraph>
<paragraph id="8B8F36EAFFF35170FE2DF1701AB1E5C1" blockId="7.[151,1436,1505,1638]" lastBlockId="8.[151,1436,151,572]" lastPageId="8" lastPageNumber="64" pageId="7" pageNumber="63">
176524* 176504 176513 78996 176520 176522 In the calls of the six individuals recorded, call repetition rate, intercall interval, the number of notes per call, the number and distinctiveness of pulses within notes and dominant frequency varied both within and among individuals (
<tableCitation id="C6B20351FFFC5170FF0EF7FF194DE4B1" box="[246,334,223,247]" captionStart="TABLE 2" captionStartId="7.[151,235,1505,1526]" captionTargetBox="[151,1436,1656,2026]" captionTargetPageId="7" captionText="TABLE 2. Measurements of advertisement call parameters for Leptolalax firthi sp. nov. Parameter values are given as means (and ranges)." httpUri="http://table.plazi.org/id/DF4F6662FFF3517FFF6FF2C1191FE250" pageId="8" pageNumber="64" tableUuid="DF4F6662FFF3517FFF6FF2C1191FE250">Table 2</tableCitation>
, 
<figureCitation id="130B2A6FFFFC5170FEA2F7FF19B8E4B1" box="[346,443,223,247]" captionStart="FIGURE 5" captionStartId="6.[151,250,1441,1464]" captionTargetBox="[154,1432,667,1417]" captionTargetId="figure@6.[151,1436,661,1421]" captionTargetPageId="6" captionText="FIGURE 5. (A) Advertisement call of Leptolalax firthi sp. nov. holotype AMS R 176524 (i) 30 s waveform of relative amplitude (Rel. amp.) over time for 23 calls, (ii) waveform and (iii) corresponding spectrogram of single representative call containing two notes, expanded from section shown in i, and (iv) power spectrum (relative amplitude vs. frequency) of first note; all recorded at an ambient air temperature of 20.2 º C. (B) Representative single advertisement calls of (i) paratype AMS R 176522 recorded at an ambient air temperature of 18.8 º C, (ii) paratype AMS R 176520 recorded at an ambient air temperature of 19.3 º C, and (iii) paratype AMS R 176513 recorded at ambient air temperature of 21.2 º C." httpUri="https://zenodo.org/record/215106/files/figure.png" pageId="8" pageNumber="64">Figure 5</figureCitation>
). Most noteworthy, while the number of notes per call was invariably two in the recording of the 
<typeStatus id="548B8848FFFC5170FEFFF624196FE55A" box="[263,364,260,284]" pageId="8" pageNumber="64" type="holotype">holotype</typeStatus>
, the number of notes per call in the recordings of 
<typeStatus id="548B8848FFFC5170FC69F6251BFFE55B" box="[913,1020,261,285]" pageId="8" pageNumber="64" type="paratype">paratypes</typeStatus>
ranged from 2–5, and was invariably three for two 
<typeStatus id="548B8848FFFC5170FED7F6081999E506" box="[303,410,296,320]" pageId="8" pageNumber="64" type="paratype">paratypes</typeStatus>
(
<tableCitation id="C6B20351FFFC5170FE50F60719FCE579" box="[424,511,295,319]" captionStart="TABLE 2" captionStartId="7.[151,235,1505,1526]" captionTargetBox="[151,1436,1656,2026]" captionTargetPageId="7" captionText="TABLE 2. Measurements of advertisement call parameters for Leptolalax firthi sp. nov. Parameter values are given as means (and ranges)." httpUri="http://table.plazi.org/id/DF4F6662FFF3517FFF6FF2C1191FE250" pageId="8" pageNumber="64" tableUuid="DF4F6662FFF3517FFF6FF2C1191FE250">Table 2</tableCitation>
). Dominant frequency also varied within and between individuals (over &lt;3 ºC difference in ambient temperature), from 5.4–6.6 kHz. To the human ear, the advertisement call of 
<taxonomicName id="4C304D69FFFC5170FB28F66D1D25E522" box="[1232,1318,332,356]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFC5170FB28F66D1D25E522" box="[1232,1318,332,356]" italics="true" pageId="8" pageNumber="64">L. firthi</emphasis>
</taxonomicName>
is a rapid, high-pitched chirping, similar to an orthopteran.
</paragraph>
<paragraph id="8B8F36EAFFF3517FFF5DF15818FFE3AC" pageId="7" pageNumber="63">
<table id="F930C44AFFF3AE87FF6FF1581D9FE3AC" box="[151,1436,1656,2026]" gridcols="7" gridrows="7" pageId="7" pageNumber="63">
<tr id="350034A8FFF3AE87FF6FF1581D9FE2C8" box="[151,1436,1656,1678]" gridrow="0" pageId="7" pageNumber="63">
<th id="76D15DD4FFF3AE87FF6FF15819BAE2C8" box="[151,441,1656,1678]" gridcol="0" gridrow="0" pageId="7" pageNumber="63">Number of calls measured</th>
<th id="76D15DD4FFF3AE87FE2DF1581A5BE2C8" box="[469,600,1656,1678]" gridcol="1" gridrow="0" pageId="7" pageNumber="63">10</th>
<th id="76D15DD4FFF3AE87FD8EF1581AFBE2C8" box="[630,760,1656,1678]" gridcol="2" gridrow="0" pageId="7" pageNumber="63">10</th>
<th id="76D15DD4FFF3AE87FCE0F1581B99E2C8" box="[792,922,1656,1678]" gridcol="3" gridrow="0" pageId="7" pageNumber="63">10</th>
<th id="76D15DD4FFF3AE87FC40F1581C38E2C8" box="[952,1083,1656,1678]" gridcol="4" gridrow="0" pageId="7" pageNumber="63">10</th>
<th id="76D15DD4FFF3AE87FBA1F1581CCEE2C8" box="[1113,1229,1656,1678]" gridcol="5" gridrow="0" pageId="7" pageNumber="63">10</th>
<th id="76D15DD4FFF3AE87FAFBF1581D9FE2C8" box="[1283,1436,1656,1678]" gridcol="6" gridrow="0" pageId="7" pageNumber="63">10</th>
</tr>
<tr id="350034A8FFF3AE87FF6FF1801D9FE2F0" box="[151,1436,1696,1718]" gridrow="1" pageId="7" pageNumber="63">
<th id="76D15DD4FFF3AE87FF6FF18019BAE2F0" box="[151,441,1696,1718]" gridcol="0" gridrow="1" pageId="7" pageNumber="63">Number of notes</th>
<td id="76D15DD4FFF3AE87FE2DF1801A5BE2F0" box="[469,600,1696,1718]" gridcol="1" gridrow="1" pageId="7" pageNumber="63">20</td>
<td id="76D15DD4FFF3AE87FD8EF1801AFBE2F0" box="[630,760,1696,1718]" gridcol="2" gridrow="1" pageId="7" pageNumber="63">28</td>
<td id="76D15DD4FFF3AE87FCE0F1801B99E2F0" box="[792,922,1696,1718]" gridcol="3" gridrow="1" pageId="7" pageNumber="63">30</td>
<td id="76D15DD4FFF3AE87FC40F1801C38E2F0" box="[952,1083,1696,1718]" gridcol="4" gridrow="1" pageId="7" pageNumber="63">45</td>
<td id="76D15DD4FFF3AE87FBA1F1801CCEE2F0" box="[1113,1229,1696,1718]" gridcol="5" gridrow="1" pageId="7" pageNumber="63">30</td>
<td id="76D15DD4FFF3AE87FAFBF1801D9FE2F0" box="[1283,1436,1696,1718]" gridcol="6" gridrow="1" pageId="7" pageNumber="63">34</td>
</tr>
<tr id="350034A8FFF3AE87FF6FF1E71D9FE299" box="[151,1436,1735,1759]" gridrow="2" pageId="7" pageNumber="63">
<th id="76D15DD4FFF3AE87FF6FF1E719BAE299" box="[151,441,1735,1759]" gridcol="0" gridrow="2" pageId="7" pageNumber="63">Call duration (ms)</th>
<td id="76D15DD4FFF3AE87FE2DF1E71A5BE299" box="[469,600,1735,1759]" gridcol="1" gridrow="2" pageId="7" pageNumber="63">21 (18 24)</td>
<td id="76D15DD4FFF3AE87FD8EF1E71AFBE299" box="[630,760,1735,1759]" gridcol="2" gridrow="2" pageId="7" pageNumber="63">46 (40 51)</td>
<td id="76D15DD4FFF3AE87FCE0F1E71B99E299" box="[792,922,1735,1759]" gridcol="3" gridrow="2" pageId="7" pageNumber="63">41 (39 42)</td>
<td id="76D15DD4FFF3AE87FC40F1E71C38E299" box="[952,1083,1735,1759]" gridcol="4" gridrow="2" pageId="7" pageNumber="63">35 (32 36)</td>
<td id="76D15DD4FFF3AE87FBA1F1E71CCEE299" box="[1113,1229,1735,1759]" gridcol="5" gridrow="2" pageId="7" pageNumber="63">34 (31 37)</td>
<td id="76D15DD4FFF3AE87FAFBF1E71D9FE299" box="[1283,1436,1735,1759]" gridcol="6" gridrow="2" pageId="7" pageNumber="63">38 (35 42)</td>
</tr>
<tr id="350034A8FFF3AE87FF6FF1D01D9FE328" box="[151,1436,1776,1902]" gridrow="3" pageId="7" pageNumber="63">
<th id="76D15DD4FFF3AE87FF6FF1D019BAE328" box="[151,441,1776,1902]" gridcol="0" gridrow="3" pageId="7" pageNumber="63">Call repetition rate (calls/s) Intercall interval (ms) Notes/call</th>
<td id="76D15DD4FFF3AE87FE2DF1D01A5BE328" box="[469,600,1776,1902]" gridcol="1" gridrow="3" pageId="7" pageNumber="63">0.86 1140 (1098 1221) 2</td>
<td id="76D15DD4FFF3AE87FD8EF1D01AFBE328" box="[630,760,1776,1902]" gridcol="2" gridrow="3" pageId="7" pageNumber="63">0.72 1346 (1075 1625) 2.8 (2 3)</td>
<td id="76D15DD4FFF3AE87FCE0F1D01B99E328" box="[792,922,1776,1902]" gridcol="3" gridrow="3" pageId="7" pageNumber="63">0.97 920 (327 1184) 3</td>
<td id="76D15DD4FFF3AE87FC40F1D01C38E328" box="[952,1083,1776,1902]" gridcol="4" gridrow="3" pageId="7" pageNumber="63">0.58 1674 (1261 2285) 4.5 (4 5)</td>
<td id="76D15DD4FFF3AE87FBA1F1D01CCEE328" box="[1113,1229,1776,1902]" gridcol="5" gridrow="3" pageId="7" pageNumber="63">0.68 1431 (1187 1716) 3</td>
<td id="76D15DD4FFF3AE87FAFBF1D01D9FE328" box="[1283,1436,1776,1902]" gridcol="6" gridrow="3" pageId="7" pageNumber="63">0.63 1545 (1401 2034) 3.4 (3 4)</td>
</tr>
<tr id="350034A8FFF3AE87FF6FF05F1D9FE3D0" box="[151,1436,1919,1942]" gridrow="4" pageId="7" pageNumber="63">
<th id="76D15DD4FFF3AE87FF6FF05F19BAE3D0" box="[151,441,1919,1942]" gridcol="0" gridrow="4" pageId="7" pageNumber="63">Dominant frequency (kHz)</th>
<td id="76D15DD4FFF3AE87FE2DF05F1A5BE3D0" box="[469,600,1919,1942]" gridcol="1" gridrow="4" pageId="7" pageNumber="63">5.8 (5.6 6.4)</td>
<td id="76D15DD4FFF3AE87FD8EF05F1AFBE3D0" box="[630,760,1919,1942]" gridcol="2" gridrow="4" pageId="7" pageNumber="63">6.2 (5.8 6.6)</td>
<td id="76D15DD4FFF3AE87FCE0F05F1B99E3D0" box="[792,922,1919,1942]" gridcol="3" gridrow="4" pageId="7" pageNumber="63">6.3 (6.2 6.6)</td>
<td id="76D15DD4FFF3AE87FC40F05F1C38E3D0" box="[952,1083,1919,1942]" gridcol="4" gridrow="4" pageId="7" pageNumber="63">5.7 (5.4 5.8)</td>
<td id="76D15DD4FFF3AE87FBA1F05F1CCEE3D0" box="[1113,1229,1919,1942]" gridcol="5" gridrow="4" pageId="7" pageNumber="63">6.2</td>
<td id="76D15DD4FFF3AE87FAFBF05F1D9FE3D0" box="[1283,1436,1919,1942]" gridcol="6" gridrow="4" pageId="7" pageNumber="63">5.8</td>
</tr>
<tr id="350034A8FFF3AE87FF6FF0891D9FE3F9" box="[151,1436,1961,1983]" gridrow="5" pageId="7" pageNumber="63">
<th id="76D15DD4FFF3AE87FF6FF08919BAE3F9" box="[151,441,1961,1983]" gridcol="0" gridrow="5" pageId="7" pageNumber="63">Temperature (°C)</th>
<td id="76D15DD4FFF3AE87FE2DF0891A5BE3F9" box="[469,600,1961,1983]" gridcol="1" gridrow="5" pageId="7" pageNumber="63">20.2</td>
<td id="76D15DD4FFF3AE87FD8EF0891AFBE3F9" box="[630,760,1961,1983]" gridcol="2" gridrow="5" pageId="7" pageNumber="63">20.4</td>
<td id="76D15DD4FFF3AE87FCE0F0891B99E3F9" box="[792,922,1961,1983]" gridcol="3" gridrow="5" pageId="7" pageNumber="63">21.2</td>
<td id="76D15DD4FFF3AE87FC40F0891C38E3F9" box="[952,1083,1961,1983]" gridcol="4" gridrow="5" pageId="7" pageNumber="63">18.3</td>
<td id="76D15DD4FFF3AE87FBA1F0891CCEE3F9" box="[1113,1229,1961,1983]" gridcol="5" gridrow="5" pageId="7" pageNumber="63">19.3</td>
<td id="76D15DD4FFF3AE87FAFBF0891D9FE3F9" box="[1283,1436,1961,1983]" gridcol="6" gridrow="5" pageId="7" pageNumber="63">18.8</td>
</tr>
<tr id="350034A8FFF3AE87FF6FF0F41D9FE3AC" box="[151,1436,2004,2026]" gridrow="6" pageId="7" pageNumber="63" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1">
<th id="76D15DD4FFF3AE87FF6FF0F419BAE3AC" box="[151,441,2004,2026]" gridcol="0" gridrow="6" pageId="7" pageNumber="63">*holotype</th>
</tr>
</table>
</paragraph>
<paragraph id="8B8F36EAFFFC5170FF31F6B31A23E67A" blockId="8.[151,1436,151,572]" pageId="8" pageNumber="64">
<emphasis id="B944EAF8FFFC5170FF31F6B319C5E5EA" bold="true" box="[201,454,403,428]" pageId="8" pageNumber="64">Sequence divergence</emphasis>
: Uncorrected sequence divergences between 
<taxonomicName id="4C304D69FFFC5170FC26F6B51C3AE5EA" box="[990,1081,404,428]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFC5170FC26F6B51C3AE5EA" box="[990,1081,404,428]" italics="true" pageId="8" pageNumber="64">L. firthi</emphasis>
</taxonomicName>
16S rRNA sequences and all homologous sequences available on GenBank (sequences assigned to 16 species; Appendix II) were&gt;10%. This degree of pairwise divergence in the 16S rRNA gene is greater than that usually representing differentiation at the species level in frogs (
<bibRefCitation id="EFA14B1BFFFC5170FE6DF6DF1A6EE651" author="Vences" box="[405,621,511,536]" pageId="8" pageNumber="64" refString="Vences, M., M. Thomas, R. M. Bonett &amp; Vieites D. R. (2005) Deciphering amphibian diversity through DNA barcoding: chances and challenges, Philosophical Transactions of the Royal Society London, Ser. B, 360, 1859 - 1868." type="journal article" year="2005">
Vences 
<emphasis id="B944EAF8FFFC5170FE16F5211A2BE65E" box="[494,552,512,536]" italics="true" pageId="8" pageNumber="64">et al.</emphasis>
2005
</bibRefCitation>
). Intraspecific variation in this gene fragment for 
<taxonomicName id="4C304D69FFFC5170FB59F5211CFBE65E" box="[1185,1272,512,536]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFC5170FB59F5211CFBE65E" box="[1185,1272,512,536]" italics="true" pageId="8" pageNumber="64">L. firthi</emphasis>
</taxonomicName>
sampled from sites up to 
<quantity id="4CC89B0FFFFC5170FEE8F5041955E67A" box="[272,342,548,572]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="64" unit="km" value="27.0">27 km</quantity>
apart was &lt;0.8%.
</paragraph>
<caption id="DF4F6662FFFC5170FF6FF1321B35E220" httpUri="https://zenodo.org/record/215107/files/figure.png" pageId="8" pageNumber="64" targetBox="[151,1436,625,1532]" targetPageId="8">
<paragraph id="8B8F36EAFFFC5170FF6FF1321B35E220" blockId="8.[151,1436,1554,1638]" pageId="8" pageNumber="64">
<emphasis id="B944EAF8FFFC5170FF6FF1321917E26E" bold="true" box="[151,276,1554,1577]" pageId="8" pageNumber="64">FIGURE 6.</emphasis>
Habitat of 
<taxonomicName id="4C304D69FFFC5170FE75F1331A36E26F" box="[397,565,1555,1577]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="firthi" status="sp. nov.">
<emphasis id="B944EAF8FFFC5170FE75F1331A36E26F" box="[397,565,1555,1577]" italics="true" pageId="8" pageNumber="64">Leptolalax firthi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775783FFFC5170FDC5F1331A8CE26E" box="[573,655,1555,1576]" pageId="8" pageNumber="64" rank="species">
<emphasis id="B944EAF8FFFC5170FDC5F1331A8CE26E" bold="true" box="[573,655,1555,1576]" pageId="8" pageNumber="64">sp. nov.</emphasis>
</taxonomicNameLabel>
(A) Habitat at ~900 m elevation in Song Thanh Proposed Nature Reserve, Quang Nam Province. (B) Slightly storm-damaged habitat at ~1700 m elevation, and (C–D) heavily storm-damaged habitat at ~1300–1500 m in Ngoc Linh Nature Reserve, Kon Tum Province.
</paragraph>
</caption>
<paragraph id="8B8F36EAFFFC5170FF31F1AE1D8FE383" blockId="8.[151,1436,1677,1990]" pageId="8" pageNumber="64">
<emphasis id="B944EAF8FFFC5170FF31F1AE1932E2E0" bold="true" box="[201,305,1678,1702]" pageId="8" pageNumber="64">Ecology:</emphasis>
All specimens of the new species were found in montane evergreen forest between ~ 
<quantity id="4CC89B0FFFFC5170FAF4F1AD1D9FE2E3" box="[1292,1436,1677,1701]" metricMagnitude="3" metricUnit="m" metricValue="1.29" metricValueMax="1.72" metricValueMin="0.86" pageId="8" pageNumber="64" unit="m" value="1290.0" valueMax="1720.0" valueMin="860.0">860–1720 m</quantity>
elevation (
<figureCitation id="130B2A6FFFFC5170FEE8F1911975E28F" box="[272,374,1713,1737]" captionStart="FIGURE 6" captionStartId="8.[151,250,1554,1577]" captionTargetBox="[151,1436,625,1532]" captionTargetId="figure@8.[151,1436,625,1533]" captionTargetPageId="8" captionText="FIGURE 6. Habitat of Leptolalax firthi sp. nov. (A) Habitat at ~ 900 m elevation in Song Thanh Proposed Nature Reserve, Quang Nam Province. (B) Slightly storm-damaged habitat at ~ 1700 m elevation, and (C – D) heavily storm-damaged habitat at ~ 1300 – 1500 m in Ngoc Linh Nature Reserve, Kon Tum Province." httpUri="https://zenodo.org/record/215107/files/figure.png" pageId="8" pageNumber="64">Figure 6</figureCitation>
). During wet-season surveys (
<date id="FF8E102AFFFC5170FD22F1911B4DE28F" box="[730,846,1713,1737]" pageId="8" pageNumber="64" value="2007-07">July 2007</date>
and 2009), only females of the new species were observed, generally from within the forest, away from streams. During these surveys, females were rarely encountered (4 individuals over 19 survey nights), and no males were ever observed or heard calling. In contrast, hundreds of males were observed and heard calling at the end of the dry season in March/ 
<date id="FF8E102AFFFC5170FBC3F03D1CB8E373" box="[1083,1211,1821,1845]" pageId="8" pageNumber="64" value="2010-04">April 2010</date>
. Females were also observed at this time, but far less abundant (and/or detectable) than males. Males were observed calling on the banks of medium sized (~ 
<quantity id="4CC89B0FFFFC5170FE41F0451A03E33B" box="[441,512,1893,1917]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="8" pageNumber="64" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">1–5 m</quantity>
wide), rocky streams, most of which had been highly disturbed by recent flooding (riparian vegetation was largely absent, with bare rocks for many meters at either side of most streams). The new species occurs in syntopy with at least three other species of 
<taxonomicName id="4C304D69FFFC5170FCC7F08E1BB4E380" box="[831,951,1966,1990]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="64" phylum="Chordata" rank="genus">
<emphasis id="B944EAF8FFFC5170FCC7F08E1BB4E380" box="[831,951,1966,1990]" italics="true" pageId="8" pageNumber="64">Leptolalax</emphasis>
</taxonomicName>
: 
<emphasis id="B944EAF8FFFC5170FC3EF08F1CC6E380" box="[966,1221,1966,1990]" italics="true" pageId="8" pageNumber="64">
<taxonomicName id="4C304D69FFFC5170FC3EF08F1C45E380" box="[966,1094,1966,1990]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="applebyi">L. applebyi</taxonomicName>
, 
<taxonomicName id="4C304D69FFFC5170FBA9F08F1CC6E380" box="[1105,1221,1967,1990]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="croceus">L. croceus</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C304D69FFFC5170FB04F08F1D86E380" box="[1276,1413,1966,1990]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="64" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B944EAF8FFFC5170FB04F08F1D86E380" box="[1276,1413,1966,1990]" italics="true" pageId="8" pageNumber="64">L. tuberosus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8F36EAFFFD5171FF31F7B71C31E522" blockId="9.[151,1437,151,1976]" pageId="9" pageNumber="65">
<emphasis id="B944EAF8FFFD5171FF31F7B719BAE4F6" bold="true" box="[201,441,151,176]" pageId="9" pageNumber="65">Conservation status</emphasis>
: 
<taxonomicName id="4C304D69FFFD5171FE33F7B81A86E4F6" box="[459,645,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FE33F7B81A86E4F6" box="[459,645,152,176]" italics="true" pageId="9" pageNumber="65">Leptolalax firthi</emphasis>
</taxonomicName>
is known from a number of sites in Song Thanh Nature Reserve in Quang Nam Province, and the adjoining Ngoc Linh Nature Reserve in Kon Tum Province. The farthest distance between two collection localities is approximately 
<quantity id="4CC89B0FFFFD5171FD22F7FF1B25E4B1" box="[730,806,223,247]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="65" unit="km" value="35.0">35 km</quantity>
. The actual distribution of the new species is unknown but probably extends to adjacent forested areas in the Kon Tum Plateau, possibly including Dong Am Pham National Biodiversity Conservation Area in 
<collectingCountry id="F327767AFFFD5171FD74F6081AC5E579" box="[652,710,296,319]" name="Laos" pageId="9" pageNumber="65">Laos</collectingCountry>
. Given the available information, we suggest the species should be considered Data Deficient following IUCN’s Red List categories (
<bibRefCitation id="EFA14B1BFFFD5171FC64F66C1C26E522" author="IUCN" box="[924,1061,332,356]" pageId="9" pageNumber="65" refString="IUCN. (2001) IUCN Red List Categories and Criteria: Version 3.1. IUCN Species Survival Commission. IUCN, Gland, Switzerland and Cambridge, UK. ii + 30 pp." type="book" year="2001">IUCN 2001</bibRefCitation>
).
</paragraph>
<paragraph id="8B8F36EAFFFD5171FF31F64F1BDDE771" blockId="9.[151,1437,151,1976]" pageId="9" pageNumber="65">
<emphasis id="B944EAF8FFFD5171FF31F64F196BE5CE" bold="true" box="[201,360,367,392]" pageId="9" pageNumber="65">Comparisons</emphasis>
: In lacking distinct dorsolateral markings including blackish spots on the flank and dark canthal and/or temporal streaks, 
<taxonomicName id="4C304D69FFFD5171FE53F6B51A02E5EA" box="[427,513,404,428]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FE53F6B51A02E5EA" box="[427,513,404,428]" italics="true" pageId="9" pageNumber="65">L. firthi</emphasis>
</taxonomicName>
is distinguished from all but 
<taxonomicName id="4C304D69FFFD5171FCB1F6B51BACE5EA" box="[841,943,405,428]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="aereus">
<emphasis id="B944EAF8FFFD5171FCB1F6B51BACE5EA" box="[841,943,405,428]" italics="true" pageId="9" pageNumber="65">L. aereus</emphasis>
</taxonomicName>
, 
<emphasis id="B944EAF8FFFD5171FC46F6B51CA5E5EA" box="[958,1190,405,428]" italics="true" pageId="9" pageNumber="65">
<taxonomicName id="4C304D69FFFD5171FC46F6B51C24E5EA" box="[958,1063,405,428]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FBCAF6B51CA5E5EA" box="[1074,1190,405,428]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="croceus">L. croceus</taxonomicName>
</emphasis>
, 
<taxonomicName id="4C304D69FFFD5171FB4CF6B51CF5E5EA" box="[1204,1270,405,428]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="eos">
<emphasis id="B944EAF8FFFD5171FB4CF6B51CF5E5EA" box="[1204,1270,405,428]" italics="true" pageId="9" pageNumber="65">L. eos</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C304D69FFFD5171FAD5F6B518B8E596" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B944EAF8FFFD5171FAD5F6B518B8E596" italics="true" pageId="9" pageNumber="65">L. tuberosus</emphasis>
</taxonomicName>
(
<emphasis id="B944EAF8FFFD5171FF33F6991D14E65E" italics="true" pageId="9" pageNumber="65">
<taxonomicName id="4C304D69FFFD5171FF33F6991934E596" box="[203,311,440,464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="alpinis">L. alpinis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FEBAF69919C1E596" box="[322,450,440,464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="applebyi">L. applebyi</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FE35F6991A71E596" box="[461,626,440,464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="bidoupensis">L. bidoupensis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FD86F6991AFBE596" box="[638,760,440,464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="bourreti">L. bourreti</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FCFBF6991B6BE596" box="[771,872,440,464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="dringi">L. dringi</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FC8BF6991C05E596" box="[883,1030,440,464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="fulignosus">L. fulignosus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FBE9F6991C85E596" box="[1041,1158,440,464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FB6AF6991D03E596" box="[1170,1280,440,464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="hamidi">L. hamidi</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FAF2F6991D94E596" box="[1290,1431,440,464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="heteropus">L. heteropus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FF6FF6FD193FE5B2" box="[151,316,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="kajangensis">L. kajangensis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FEB1F6FD199DE5B2" box="[329,414,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FE53F6FD1A40E5B2" box="[427,579,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="khasiorum">L. khasiorum</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FDB7F6FD1ACEE5B2" box="[591,717,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="lateralis">L. lateralis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FD21F6FD1B23E5B2" box="[729,800,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="liui">L. liui</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FCD4F6FD1BA1E5B2" box="[812,930,477,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="maurus">L. maurus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FC56F6FD1C69E5B2" box="[942,1130,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="melanoleucus">L. melanoleucus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FB8EF6FD1CEFE5B2" box="[1142,1260,476,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="melicus">L. melicus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FB01F6FD1D7AE5B2" box="[1273,1401,477,500]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="minimus">L. minimus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FA7EF6FD1925E65E" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="nahangensis">L. nahangensis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FEC9F521197AE65E" box="[305,377,513,536]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="nyx">L. nyx</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FE7CF5211A23E65E" box="[388,544,512,536]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="oshanensis">L. oshanensis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FDD4F5211AD9E65E" box="[556,730,512,536]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FD1EF5211B4AE65E" box="[742,841,513,536]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="pictus">L. pictus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FCADF5211BD4E65E" box="[853,983,512,536]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="pluvialis">L. pluvialis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FC1AF5211C3FE65E" box="[994,1084,512,536]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="solus">L. solus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FBB0F5211CA6E65E" box="[1096,1189,513,536]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="sungi">L. sungi</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FB49F5211D14E65E" box="[1201,1303,512,536]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="tamdil">L. tamdil</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C304D69FFFD5171FAB6F521191DE67A" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="B944EAF8FFFD5171FAB6F521191DE67A" italics="true" pageId="9" pageNumber="65">L. ventripunctatus</emphasis>
</taxonomicName>
all have blackish spots on the flank, and dark canthal and/or dark temporal streaks). From 
<emphasis id="B944EAF8FFFD5171FAE9F50518CCE626" italics="true" pageId="9" pageNumber="65">
<taxonomicName id="4C304D69FFFD5171FAE9F5051D79E67A" box="[1297,1402,549,572]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="aereus">L. aereus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FA7DF50518CCE626" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">L. firthi</taxonomicName>
</emphasis>
can be distinguished molecularly, by the male advertisement call and by the degree of lateral fringing on the toes (see below); from 
<emphasis id="B944EAF8FFFD5171FE6EF54D1A5CE6C2" box="[406,607,620,644]" italics="true" pageId="9" pageNumber="65">
<taxonomicName id="4C304D69FFFD5171FE6EF54D19FCE6C2" box="[406,511,621,644]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FDF1F54D1A5CE6C2" box="[521,607,620,644]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">L. firthi</taxonomicName>
</emphasis>
can be distinguished molecularly, by the degree of lateral fringing on toes, and by the male advertisement call (see below); from 
<taxonomicName id="4C304D69FFFD5171FD17F5B11B61E6EE" box="[751,866,657,680]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="croceus">
<emphasis id="B944EAF8FFFD5171FD17F5B11B61E6EE" box="[751,866,657,680]" italics="true" pageId="9" pageNumber="65">L. croceus</emphasis>
</taxonomicName>
, the new species can be distinguished molecularly, by ventral coloration, degree of lateral fringing on toes, visibility of the tympanum, and by the male advertisement call (see below); from 
<emphasis id="B944EAF8FFFD5171FE5CF5F91A55E6B6" box="[420,598,728,752]" italics="true" pageId="9" pageNumber="65">
<taxonomicName id="4C304D69FFFD5171FE5CF5F919EEE6B6" box="[420,493,729,752]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="eos">L. eos</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FE04F5F91A55E6B6" box="[508,598,728,752]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">L. firthi</taxonomicName>
</emphasis>
can be distinguished molecularly and by size (see below); and from 
<taxonomicName id="4C304D69FFFD5171FA7DF5F91900E752" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B944EAF8FFFD5171FA7DF5F91900E752" italics="true" pageId="9" pageNumber="65">L. tuberosus</emphasis>
</taxonomicName>
, the new species can be distinguished molecularly, by ventral pattern, degree of lateral fringing on toes, visibility of the tympanum, and by the male advertisement call (see below)
</paragraph>
<paragraph id="8B8F36EAFFFD5171FF31F4651D36E78F" blockId="9.[151,1437,151,1976]" pageId="9" pageNumber="65">
In having toes with broad lateral fringing in males, 
<taxonomicName id="4C304D69FFFD5171FCEAF4651B6AE71A" box="[786,873,836,860]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FCEAF4651B6AE71A" box="[786,873,836,860]" italics="true" pageId="9" pageNumber="65">L. firthi</emphasis>
</taxonomicName>
is further distinguished from all but 
<emphasis id="B944EAF8FFFD5171FAF4F46518BFE7C6" italics="true" pageId="9" pageNumber="65">
<taxonomicName id="4C304D69FFFD5171FAF4F4651D7EE71A" box="[1292,1405,836,860]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="alpinus">L. alpinus</taxonomicName>
<taxonomicName id="4C304D69FFFD5171FA7DF46518BFE7C6" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="eos">L. eos</taxonomicName>
</emphasis>
, and 
<taxonomicName id="4C304D69FFFD5171FF03F449193DE7C6" box="[251,318,872,896]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="liui">
<emphasis id="B944EAF8FFFD5171FF03F449193DE7C6" box="[251,318,872,896]" italics="true" pageId="9" pageNumber="65">L. liui</emphasis>
</taxonomicName>
(all other male congeners lack fringing or have only weak fringing on their toes. In addition, topotypic adult male 
<taxonomicName id="4C304D69FFFD5171FEABF4AD1995E7E2" box="[339,406,908,932]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="liui">
<emphasis id="B944EAF8FFFD5171FEABF4AD1995E7E2" box="[339,406,908,932]" italics="true" pageId="9" pageNumber="65">L. liui</emphasis>
</taxonomicName>
examined [N = 3] have considerably less developed fringing compared to 
<taxonomicName id="4C304D69FFFD5171FB26F4AD1D37E7E2" box="[1246,1332,908,932]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FB26F4AD1D37E7E2" box="[1246,1332,908,932]" italics="true" pageId="9" pageNumber="65">L. firthi</emphasis>
</taxonomicName>
). In having males with wide lateral dermal fringes on Finger II only, 
<taxonomicName id="4C304D69FFFD5171FCB9F4911B94E78E" box="[833,919,944,968]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FCB9F4911B94E78E" box="[833,919,944,968]" italics="true" pageId="9" pageNumber="65">L. firthi</emphasis>
</taxonomicName>
appears to differ from all congeners.
</paragraph>
<paragraph id="8B8F36EAFFFD5171FF31F4F41C0AE0E6" blockId="9.[151,1437,151,1976]" pageId="9" pageNumber="65">
<taxonomicName id="4C304D69FFFD5171FF31F4F41983E7AA" box="[201,384,980,1004]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FF31F4F41983E7AA" box="[201,384,980,1004]" italics="true" pageId="9" pageNumber="65">Leptolalax firthi</emphasis>
</taxonomicName>
is a medium-sized species of 
<taxonomicName id="4C304D69FFFD5171FD2DF4F41B4EE7AA" box="[725,845,980,1004]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="genus">
<emphasis id="B944EAF8FFFD5171FD2DF4F41B4EE7AA" box="[725,845,980,1004]" italics="true" pageId="9" pageNumber="65">Leptolalax</emphasis>
</taxonomicName>
(
<quantity id="4CC89B0FFFFD5171FCA4F4F41C01E7AA" box="[860,1026,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="2.78" metricValueMax="2.92" metricValueMin="2.6399999999999997" pageId="9" pageNumber="65" unit="mm" value="27.799999999999997" valueMax="29.2" valueMin="26.4">26.4–29.2 mm</quantity>
in 
<specimenCount id="9D36FD63FFFD5171FBDFF4F41C80E7AA" box="[1063,1155,980,1004]" pageId="9" pageNumber="65" type="adult">21 adult</specimenCount>
males, 
<quantity id="4CC89B0FFFFD5171FB23F4F41D7CE7AA" box="[1243,1407,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="3.13" metricValueMax="3.69" metricValueMin="2.57" pageId="9" pageNumber="65" unit="mm" value="31.299999999999997" valueMax="36.9" valueMin="25.7">25.7–36.9 mm</quantity>
in 
<specimenCount id="9D36FD63FFFD5171FF6FF4D7191AE049" box="[151,281,1015,1039]" pageId="9" pageNumber="65" type="female">14 females</specimenCount>
), and can be distinguished on the basis of size from the smaller 
<taxonomicName id="4C304D69FFFD5171FBF6F4D91C8CE056" box="[1038,1167,1016,1040]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B944EAF8FFFD5171FBF6F4D91C8CE056" box="[1038,1167,1016,1040]" italics="true" pageId="9" pageNumber="65">L. applebyi</emphasis>
</taxonomicName>
(males 
<quantity id="4CC89B0FFFFD5171FB17F4D71D9BE056" box="[1263,1432,1015,1040]" metricMagnitude="-2" metricUnit="m" metricValue="2.095" metricValueMax="2.23" metricValueMin="1.96" pageId="9" pageNumber="65" unit="mm" value="20.95" valueMax="22.3" valueMin="19.6">19.6–22.3 mm</quantity>
, females 
<quantity id="4CC89B0FFFFD5171FF0EF33C199DE072" box="[246,414,1052,1076]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" metricValueMax="2.59" metricValueMin="2.17" pageId="9" pageNumber="65" unit="mm" value="23.799999999999997" valueMax="25.9" valueMin="21.7">21.7–25.9 mm</quantity>
), 
<taxonomicName id="4C304D69FFFD5171FE4AF33D1A05E072" box="[434,518,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="kecil">
<emphasis id="B944EAF8FFFD5171FE4AF33D1A05E072" box="[434,518,1052,1076]" italics="true" pageId="9" pageNumber="65">L. kecil</emphasis>
</taxonomicName>
(males 
<quantity id="4CC89B0FFFFD5171FD99F33C1B04E072" box="[609,775,1052,1076]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" metricValueMax="2.05" metricValueMin="1.93" pageId="9" pageNumber="65" unit="mm" value="19.9" valueMax="20.5" valueMin="19.3">19.3–20.5 mm</quantity>
, female 
<quantity id="4CC89B0FFFFD5171FC90F33C1BBFE072" box="[872,956,1052,1076]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="9" pageNumber="65" unit="mm" value="25.0">25 mm</quantity>
), 
<taxonomicName id="4C304D69FFFD5171FC28F33D1C47E072" box="[976,1092,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B944EAF8FFFD5171FC28F33D1C47E072" box="[976,1092,1052,1076]" italics="true" pageId="9" pageNumber="65">L. melicus</emphasis>
</taxonomicName>
(males 
<quantity id="4CC89B0FFFFD5171FB58F33C1D46E072" box="[1184,1349,1052,1076]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" metricValueMax="2.27" metricValueMin="1.95" pageId="9" pageNumber="65" unit="mm" value="21.1" valueMax="22.7" valueMin="19.5">19.5–22.7 mm</quantity>
) and 
<taxonomicName id="4C304D69FFFD5171FA7DF33D18FBE01E" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="B944EAF8FFFD5171FA7DF33D18FBE01E" italics="true" pageId="9" pageNumber="65">L. pluvialis</emphasis>
</taxonomicName>
(males 
<quantity id="4CC89B0FFFFD5171FEB7F36019F5E01E" box="[335,502,1088,1112]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" metricValueMax="2.23" metricValueMin="2.13" pageId="9" pageNumber="65" unit="mm" value="21.8" valueMax="22.3" valueMin="21.3">21.3–22.3 mm</quantity>
), and the larger 
<taxonomicName id="4C304D69FFFD5171FD50F3611B23E01E" box="[680,800,1088,1112]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="bourreti">
<emphasis id="B944EAF8FFFD5171FD50F3611B23E01E" box="[680,800,1088,1112]" italics="true" pageId="9" pageNumber="65">L. bourreti</emphasis>
</taxonomicName>
(male 
<quantity id="4CC89B0FFFFD5171FC94F3601BD7E01E" box="[876,980,1088,1112]" metricMagnitude="-2" metricUnit="m" metricValue="3.62" pageId="9" pageNumber="65" unit="mm" value="36.2">36.2 mm</quantity>
), 
<taxonomicName id="4C304D69FFFD5171FC1FF3611C29E01E" box="[999,1066,1089,1112]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="eos">
<emphasis id="B944EAF8FFFD5171FC1FF3611C29E01E" box="[999,1066,1089,1112]" italics="true" pageId="9" pageNumber="65">L. eos</emphasis>
</taxonomicName>
(males 
<quantity id="4CC89B0FFFFD5171FB79F3601D2BE01E" box="[1153,1320,1088,1112]" metricMagnitude="-2" metricUnit="m" metricValue="3.39" metricValueMax="3.47" metricValueMin="3.31" pageId="9" pageNumber="65" unit="mm" value="33.9" valueMax="34.7" valueMin="33.1">33.1–34.7 mm</quantity>
), 
<taxonomicName id="4C304D69FFFD5171FAC2F36118B1E03A" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B944EAF8FFFD5171FAC2F36118B1E03A" italics="true" pageId="9" pageNumber="65">L. gracilis</emphasis>
</taxonomicName>
(males 
<quantity id="4CC89B0FFFFD5171FEF4F3441989E03A" box="[268,394,1124,1148]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.6" metricValueMin="3.0" pageId="9" pageNumber="65" unit="mm" value="33.0" valueMax="36.0" valueMin="30.0">30–36 mm</quantity>
), 
<taxonomicName id="4C304D69FFFD5171FE66F3451A42E03A" box="[414,577,1124,1148]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="kajangensis">
<emphasis id="B944EAF8FFFD5171FE66F3451A42E03A" box="[414,577,1124,1148]" italics="true" pageId="9" pageNumber="65">L. kajangensis</emphasis>
</taxonomicName>
(males 
<quantity id="4CC89B0FFFFD5171FD64F3441B19E03A" box="[668,794,1124,1148]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="9" pageNumber="65" unit="mm" value="34.5" valueMax="35.0" valueMin="34.0">34–35 mm</quantity>
), 
<taxonomicName id="4C304D69FFFD5171FCD6F3451BDAE03A" box="[814,985,1124,1148]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="nahangensis">
<emphasis id="B944EAF8FFFD5171FCD6F3451BDAE03A" box="[814,985,1124,1148]" italics="true" pageId="9" pageNumber="65">L. nahangensis</emphasis>
</taxonomicName>
(male 
<quantity id="4CC89B0FFFFD5171FBD1F3441C91E03A" box="[1065,1170,1124,1148]" metricMagnitude="-2" metricUnit="m" metricValue="4.08" pageId="9" pageNumber="65" unit="mm" value="40.8">40.8 mm</quantity>
), 
<taxonomicName id="4C304D69FFFD5171FB5EF3451D01E03A" box="[1190,1282,1125,1148]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="sungi">
<emphasis id="B944EAF8FFFD5171FB5EF3451D01E03A" box="[1190,1282,1125,1148]" italics="true" pageId="9" pageNumber="65">L. sungi</emphasis>
</taxonomicName>
(males 
<quantity id="4CC89B0FFFFD5171FAA5F34418FEE0E6" metricMagnitude="-2" metricUnit="m" metricValue="5.05" metricValueMax="5.27" metricValueMin="4.83" pageId="9" pageNumber="65" unit="mm" value="50.5" valueMax="52.7" valueMin="48.3">48.3– 52.7 mm</quantity>
, females 56.7–58.9), and 
<taxonomicName id="4C304D69FFFD5171FDE5F3A91A80E0E6" box="[541,643,1160,1184]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="tamdil">
<emphasis id="B944EAF8FFFD5171FDE5F3A91A80E0E6" box="[541,643,1160,1184]" italics="true" pageId="9" pageNumber="65">L. tamdil</emphasis>
</taxonomicName>
(male 
<quantity id="4CC89B0FFFFD5171FD28F3A81B35E0E6" box="[720,822,1160,1184]" metricMagnitude="-2" metricUnit="m" metricValue="3.2299999999999995" pageId="9" pageNumber="65" unit="mm" value="32.3">32.3 mm</quantity>
, female 
<quantity id="4CC89B0FFFFD5171FC6DF3A81BFEE0E6" box="[917,1021,1160,1184]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="9" pageNumber="65" unit="mm" value="31.8">31.8 mm</quantity>
).
</paragraph>
<paragraph id="8B8F36EAFFFD5171FF31F38D1C61E112" blockId="9.[151,1437,151,1976]" pageId="9" pageNumber="65">
In having an immaculate white chest and belly with only slight brown specking at the margins, 
<taxonomicName id="4C304D69FFFD5171FB0FF38D1D4FE082" box="[1271,1356,1196,1220]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FB0FF38D1D4FE082" box="[1271,1356,1196,1220]" italics="true" pageId="9" pageNumber="65">L. firthi</emphasis>
</taxonomicName>
can be distinguished from 
<emphasis id="B944EAF8FFFD5171FE97F3F11900E176" italics="true" pageId="9" pageNumber="65">
<taxonomicName id="4C304D69FFFD5171FE97F3F119D8E0AE" box="[367,475,1232,1256]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="alpinis">L. alpinis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FE1EF3F11A65E0AE" box="[486,614,1232,1256]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="applebyi">L. applebyi</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FD89F3F11B14E0AE" box="[625,791,1232,1256]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="bidoupensis">L. bidoupensis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FCD9F3F11B94E0AE" box="[801,919,1233,1256]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="croceus">L. croceus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FC5AF3F11C05E0AE" box="[930,1030,1232,1256]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="dringi">L. dringi</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FBE9F3F11CA7E0AE" box="[1041,1188,1232,1256]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="fulignosus">L. fulignosus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FB57F3F11D26E0AE" box="[1199,1317,1232,1256]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FAD7F3F118C2E14A" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="heteropus">L. heteropus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FF37F3D51974E14A" box="[207,375,1268,1292]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="kajangensis">L. kajangensis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FE7EF3D519DDE14A" box="[390,478,1268,1292]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FE15F3D51A61E14A" box="[493,610,1269,1292]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="maurus">L. maurus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FD8CF3D51B32E14A" box="[628,817,1268,1292]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="melanoleucus">L. melanoleucus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FCB8F3D51BBAE14A" box="[832,953,1268,1292]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="melicus">L. melicus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FC30F3D51C7BE14A" box="[968,1144,1268,1292]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="nahangensis">L. nahangensis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FB7FF3D51D08E14A" box="[1159,1291,1268,1292]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="pluvialis">L. pluvialis</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FAE1F3D51D75E14A" box="[1305,1398,1268,1292]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="solus">L. solus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FA7EF3D51900E176" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="tuberosus">L. tuberosus</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C304D69FFFD5171FEC4F2391A0AE176" box="[316,521,1305,1328]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="B944EAF8FFFD5171FEC4F2391A0AE176" box="[316,521,1305,1328]" italics="true" pageId="9" pageNumber="65">L. ventripunctatus</emphasis>
</taxonomicName>
, all of which have dark or otherwise maculate chests and/or bellies. In having a visible tympanum, 
<taxonomicName id="4C304D69FFFD5171FE96F21D19C7E112" box="[366,452,1340,1364]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FE96F21D19C7E112" box="[366,452,1340,1364]" italics="true" pageId="9" pageNumber="65">L. firthi</emphasis>
</taxonomicName>
is differentiated from 
<emphasis id="B944EAF8FFFD5171FD47F21D1B98E112" box="[703,923,1341,1364]" italics="true" pageId="9" pageNumber="65">
<taxonomicName id="4C304D69FFFD5171FD47F21D1B36E112" box="[703,821,1341,1364]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="croceus">L. croceus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FCB8F21D1B98E112" box="[832,923,1341,1364]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="sungi">L. sungi</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C304D69FFFD5171FC29F21D1C58E112" box="[977,1115,1340,1364]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B944EAF8FFFD5171FC29F21D1C58E112" box="[977,1115,1340,1364]" italics="true" pageId="9" pageNumber="65">L. tuberosus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8F36EAFFFD5171FF31F27F18F6E329" blockId="9.[151,1437,151,1976]" pageId="9" pageNumber="65">
The advertisement call of 
<taxonomicName id="4C304D69FFFD5171FE0BF2411A49E13E" box="[499,586,1376,1400]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FE0BF2411A49E13E" box="[499,586,1376,1400]" italics="true" pageId="9" pageNumber="65">L. firthi</emphasis>
</taxonomicName>
is unique among congeners with known calls. In lacking strong frequency modulation, the call of 
<taxonomicName id="4C304D69FFFD5171FE61F2A519EDE1DA" box="[409,494,1412,1436]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FE61F2A519EDE1DA" box="[409,494,1412,1436]" italics="true" pageId="9" pageNumber="65">L. firthi</emphasis>
</taxonomicName>
differs from 
<taxonomicName id="4C304D69FFFD5171FD79F2A51AE1E1DA" box="[641,738,1412,1436]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="dringi">
<emphasis id="B944EAF8FFFD5171FD79F2A51AE1E1DA" box="[641,738,1412,1436]" italics="true" pageId="9" pageNumber="65">L. dringi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C304D69FFFD5171FCEFF2A51B81E1DA" box="[791,898,1412,1436]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="hamidi">
<emphasis id="B944EAF8FFFD5171FCEFF2A51B81E1DA" box="[791,898,1412,1436]" italics="true" pageId="9" pageNumber="65">L. hamidi</emphasis>
</taxonomicName>
. In having a relatively high dominant frequency (5.4–6.6 kHz; 18.3–21.2 ºC) the call of 
<taxonomicName id="4C304D69FFFD5171FDABF2891AABE186" box="[595,680,1448,1472]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FDABF2891AABE186" box="[595,680,1448,1472]" italics="true" pageId="9" pageNumber="65">L. firthi</emphasis>
</taxonomicName>
can be readily distinguished from that of 
<taxonomicName id="4C304D69FFFD5171FB85F2891CF8E186" box="[1149,1275,1448,1472]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B944EAF8FFFD5171FB85F2891CF8E186" box="[1149,1275,1448,1472]" italics="true" pageId="9" pageNumber="65">L. applebyi</emphasis>
</taxonomicName>
(4.0–4.3 kHz, 21.5 ºC), 
<taxonomicName id="4C304D69FFFD5171FEF9F2ED19A6E1A2" box="[257,421,1484,1508]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B944EAF8FFFD5171FEF9F2ED19A6E1A2" box="[257,421,1484,1508]" italics="true" pageId="9" pageNumber="65">L. bidoupensis</emphasis>
</taxonomicName>
(1.9–3.8 kHz, 19.9–20.0 ºC), 
<taxonomicName id="4C304D69FFFD5171FD05F2ED1B72E1A2" box="[765,881,1485,1508]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="croceus">
<emphasis id="B944EAF8FFFD5171FD05F2ED1B72E1A2" box="[765,881,1485,1508]" italics="true" pageId="9" pageNumber="65">L. croceus</emphasis>
</taxonomicName>
(2.6–3.0 kHz, 21.6–25.1 ºC), 
<taxonomicName id="4C304D69FFFD5171FB3AF2ED1D58E1A2" box="[1218,1371,1484,1508]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="B944EAF8FFFD5171FB3AF2ED1D58E1A2" box="[1218,1371,1484,1508]" italics="true" pageId="9" pageNumber="65">L. fuliginosus</emphasis>
</taxonomicName>
(2.3– 2.4 kHz, 19.3–19.6 ºC), 
<taxonomicName id="4C304D69FFFD5171FE5FF2D11A19E24E" box="[423,538,1520,1544]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B944EAF8FFFD5171FE5FF2D11A19E24E" box="[423,538,1520,1544]" italics="true" pageId="9" pageNumber="65">L. gracilis</emphasis>
</taxonomicName>
(2.5–2.7 kHz, 20 ºC), 
<taxonomicName id="4C304D69FFFD5171FCE0F2D11BA1E24E" box="[792,930,1520,1544]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="B944EAF8FFFD5171FCE0F2D11BA1E24E" box="[792,930,1520,1544]" italics="true" pageId="9" pageNumber="65">L. heteropus</emphasis>
</taxonomicName>
(2.8 kHz, 21 ºC), 
<taxonomicName id="4C304D69FFFD5171FB97F2D11CC2E24E" box="[1135,1217,1520,1544]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="kecil">
<emphasis id="B944EAF8FFFD5171FB97F2D11CC2E24E" box="[1135,1217,1520,1544]" italics="true" pageId="9" pageNumber="65">L. kecil</emphasis>
</taxonomicName>
(3.2 kHz, 21.4 ºC), 
<taxonomicName id="4C304D69FFFD5171FF6FF135194DE26A" box="[151,334,1556,1580]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="melanoleucus">
<emphasis id="B944EAF8FFFD5171FF6FF135194DE26A" box="[151,334,1556,1580]" italics="true" pageId="9" pageNumber="65">L. melanoleucus</emphasis>
</taxonomicName>
(3.1–3.2 kHz, 23.9 ºC), 
<taxonomicName id="4C304D69FFFD5171FDA5F1351AD3E26A" box="[605,720,1556,1580]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B944EAF8FFFD5171FDA5F1351AD3E26A" box="[605,720,1556,1580]" italics="true" pageId="9" pageNumber="65">L. melicus</emphasis>
</taxonomicName>
(2.6–4.0 kHz, 26.1–26.2 ºC), 
<taxonomicName id="4C304D69FFFD5171FBE7F1351CB4E26A" box="[1055,1207,1556,1580]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="B944EAF8FFFD5171FBE7F1351CB4E26A" box="[1055,1207,1556,1580]" italics="true" pageId="9" pageNumber="65">L. oshanensis</emphasis>
</taxonomicName>
(4.4–4.6 kHz, 14ºC; recorded from c. 
<quantity id="4CC89B0FFFFD5171FEAEF118199EE216" box="[342,413,1592,1616]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="65" unit="km" value="40.0">40 km</quantity>
from 
<typeStatus id="548B8848FFFD5171FE18F1191A13E217" box="[480,528,1593,1617]" pageId="9" pageNumber="65">type</typeStatus>
locality of 
<taxonomicName id="4C304D69FFFD5171FD77F1191B24E216" box="[655,807,1592,1616]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="B944EAF8FFFD5171FD77F1191B24E216" box="[655,807,1592,1616]" italics="true" pageId="9" pageNumber="65">L. oshanensis</emphasis>
</taxonomicName>
), 
<taxonomicName id="4C304D69FFFD5171FCC6F1191B96E216" box="[830,917,1592,1616]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="solus">
<emphasis id="B944EAF8FFFD5171FCC6F1191B96E216" box="[830,917,1592,1616]" italics="true" pageId="9" pageNumber="65">L. solus</emphasis>
</taxonomicName>
(3.1–3.2 kHz, 24.2–24.3 ºC), and 
<taxonomicName id="4C304D69FFFD5171FAEBF1191D9FE216" box="[1299,1436,1592,1616]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B944EAF8FFFD5171FAEBF1191D9FE216" box="[1299,1436,1592,1616]" italics="true" pageId="9" pageNumber="65">L. tuberosus</emphasis>
</taxonomicName>
(2.6– 2.8 kHz, 22.5–24.5 ºC). The call of 
<taxonomicName id="4C304D69FFFD5171FD90F17D1ABDE232" box="[616,702,1628,1652]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FD90F17D1ABDE232" box="[616,702,1628,1652]" italics="true" pageId="9" pageNumber="65">L. firthi</emphasis>
</taxonomicName>
also appears to be lower in frequency compared to that attributed to 
<taxonomicName id="4C304D69FFFD5171FF4DF1A1191AE2DE" box="[181,281,1664,1688]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="dringi">
<emphasis id="B944EAF8FFFD5171FF4DF1A1191AE2DE" box="[181,281,1664,1688]" italics="true" pageId="9" pageNumber="65">L. dringi</emphasis>
</taxonomicName>
(7.6–8.1 kHz, 24.3 ºC). Although frequency can vary with temperature, differences among species of the scale reported here are very unlikely to be attributed to temperature differences. Congeners with known calls of a similar dominant frequency are 
<emphasis id="B944EAF8FFFD5171FDF6F1E91C5BE2A6" box="[526,1112,1736,1760]" italics="true" pageId="9" pageNumber="65">
<taxonomicName id="4C304D69FFFD5171FDF6F1E91A7BE2A6" box="[526,632,1737,1760]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="aereus">L. aereus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FD7BF1E91AF6E2A6" box="[643,757,1736,1760]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="alpinus">L. alpinus</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FCF8F1E91B69E2A6" box="[768,874,1737,1760]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FC8DF1E91BE0E2A6" box="[885,995,1736,1760]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="hamidi">L. hamidi</taxonomicName>
, 
<taxonomicName id="4C304D69FFFD5171FC15F1E91C5BE2A6" box="[1005,1112,1737,1760]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="maurus">L maurus</taxonomicName>
</emphasis>
and 
<taxonomicName id="4C304D69FFFD5171FB75F1E91CEDE2A6" box="[1165,1262,1737,1760]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B944EAF8FFFD5171FB75F1E91CEDE2A6" box="[1165,1262,1737,1760]" italics="true" pageId="9" pageNumber="65">L. pictus</emphasis>
</taxonomicName>
. The advertisement call of 
<taxonomicName id="4C304D69FFFD5171FEDEF1CD197EE342" box="[294,381,1772,1796]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FEDEF1CD197EE342" box="[294,381,1772,1796]" italics="true" pageId="9" pageNumber="65">L. firthi</emphasis>
</taxonomicName>
has a lower call repetition rate (0.58–0.97 calls/s) compared to 
<taxonomicName id="4C304D69FFFD5171FBA9F1CD1CB9E342" box="[1105,1210,1773,1796]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="aereus">
<emphasis id="B944EAF8FFFD5171FBA9F1CD1CB9E342" box="[1105,1210,1773,1796]" italics="true" pageId="9" pageNumber="65">L. aereus</emphasis>
</taxonomicName>
(2.4–8.0 calls/s), 
<taxonomicName id="4C304D69FFFD5171FA7DF1CD18E3E36E" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="arayai">
<emphasis id="B944EAF8FFFD5171FA7DF1CD18E3E36E" italics="true" pageId="9" pageNumber="65">L. arayai</emphasis>
</taxonomicName>
(9.0–9.3 calls/s), 
<taxonomicName id="4C304D69FFFD5171FE53F0311A14E36E" box="[427,535,1808,1832]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="hamidi">
<emphasis id="B944EAF8FFFD5171FE53F0311A14E36E" box="[427,535,1808,1832]" italics="true" pageId="9" pageNumber="65">L. hamidi</emphasis>
</taxonomicName>
(9.0–9.3 calls/s), and 
<taxonomicName id="4C304D69FFFD5171FCEAF0311B77E36E" box="[786,884,1809,1832]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B944EAF8FFFD5171FCEAF0311B77E36E" box="[786,884,1809,1832]" italics="true" pageId="9" pageNumber="65">L. pictus</emphasis>
</taxonomicName>
(11–13 calls/s), has a lower number of notes per call (2–5) compared to 
<taxonomicName id="4C304D69FFFD5171FE58F0151A11E30A" box="[416,530,1844,1868]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="alpinus">
<emphasis id="B944EAF8FFFD5171FE58F0151A11E30A" box="[416,530,1844,1868]" italics="true" pageId="9" pageNumber="65">L. alpinus</emphasis>
</taxonomicName>
(9.45 2.73), and has a lower call duration (18–51 ms) compared to 
<taxonomicName id="4C304D69FFFD5171FAD1F0151D9FE30A" box="[1321,1436,1845,1868]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="maurus">
<emphasis id="B944EAF8FFFD5171FAD1F0151D9FE30A" box="[1321,1436,1845,1868]" italics="true" pageId="9" pageNumber="65">L. maurus</emphasis>
</taxonomicName>
(85 ms).
</paragraph>
<paragraph id="8B8F36EAFFFD5171FF31F05D1D03E3FE" blockId="9.[151,1437,151,1976]" pageId="9" pageNumber="65">
In addition, uncorrected sequence divergences between 
<taxonomicName id="4C304D69FFFD5171FCA9F05D1BA8E3D2" box="[849,939,1916,1940]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="65" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B944EAF8FFFD5171FCA9F05D1BA8E3D2" box="[849,939,1916,1940]" italics="true" pageId="9" pageNumber="65">L. firthi</emphasis>
</taxonomicName>
16S rRNA sequences and all homologous sequences available on GenBank (from individuals assigned to 16 species; Appendix II) were&gt;10%.
</paragraph>
</subSubSection>
</treatment>
</document>