666 lines
74 KiB
XML
666 lines
74 KiB
XML
<document id="C08ADBF9E20660BA7F3396A1E838D324" ID="10.11646/zootaxa.4126.1.3" ID-CLB-Dataset="38804" ID-DOI="10.11646/zootaxa.4126.1.3" ID-GBIF-Dataset="5419f9c9-7f5b-469e-ac7e-fc34a0c9d54a" ID-ISSN="1175-5326" ID-Zenodo-Dep="262880" ID-ZooBank="B3A0D94A-3CEB-4B5F-99A2-23B595FA4219" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1466059820839" checkinUser="plazi" docAuthor="Barjadze, Shalva, Baquero, Enrique, Soto-Adames, Felipe N., Giordano, Rosanna & Jordana, Rafael" docDate="2016" docId="039987FCFFC9FF9BFF14398D6DF08B95" docLanguage="en" docName="zootaxa.4126.1.3.pdf" docOrigin="Zootaxa 4126 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Plutomurus revazi Barjadze, Baquero, Soto-Adames, Giordano & Jordana, 2016, sp. nov." docType="treatment" docUuid="602A1B34-57E6-41C8-93C8-831BACCCCD99" docUuidSource="ZooBank" docVersion="12" lastPageNumber="84" masterDocId="FFA0FF84FFCDFF9CFF833D0669018C63" masterDocTitle="New diagnosis for species of Plutomurus Yosii (Collembola, Tomoceridae), with descriptions of two new species from Georgian caves" masterLastPageNumber="96" masterPageNumber="77" pageNumber="81" updateTime="1698663001192" updateUser="ExternalLinkService">
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<mods:title id="C6DE1704F894A29D00A8BFD78D598DC9">New diagnosis for species of Plutomurus Yosii (Collembola, Tomoceridae), with descriptions of two new species from Georgian caves</mods:title>
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<mods:namePart id="6D66DED65FC299DAF5F06F9BD3511785">Barjadze, Shalva</mods:namePart>
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<mods:namePart id="42DDA67B47A5A5EC8E5A63B1C593A27C">Baquero, Enrique</mods:namePart>
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<mods:namePart id="1E74BDB562A0EC837554FF53897B3963">Soto-Adames, Felipe N.</mods:namePart>
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<mods:title id="11681A56BF4E8BE51577DDF48B80BFD8">Zootaxa</mods:title>
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<treatment id="039987FCFFC9FF9BFF14398D6DF08B95" ID-DOI="http://doi.org/10.5281/zenodo.5625761" ID-GBIF-Taxon="120828680" ID-Zenodo-Dep="5625761" LSID="urn:lsid:zoobank.org:act:602A1B34-57E6-41C8-93C8-831BACCCCD99" httpUri="http://treatment.plazi.org/id/039987FCFFC9FF9BFF14398D6DF08B95" lastPageId="7" lastPageNumber="84" pageId="4" pageNumber="81">
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<subSubSection id="C32A6561FFC9FF98FF14398D686688A4" pageId="4" pageNumber="81" type="nomenclature">
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<paragraph id="8B8F36EAFFC9FF98FF14398D68D888C5" blockId="4.[151,473,1163,1223]" box="[151,473,1163,1190]" pageId="4" pageNumber="81">
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<heading id="D0C78186FFC9FF98FF14398D68D888C5" bold="true" box="[151,473,1163,1190]" fontSize="11" level="1" pageId="4" pageNumber="81" reason="1">
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<taxonomicName id="4C304D69FFC9FF98FF14398D687788C6" box="[151,374,1163,1189]" class="Entognatha" family="Tomoceridae" genus="Plutomurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="81" phylum="Arthropoda" rank="species" species="revazi" status="sp. nov.">
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<emphasis id="B944EAF8FFC9FF98FF14398D687788C6" bold="true" box="[151,374,1163,1189]" italics="true" pageId="4" pageNumber="81">Plutomurus revazi</emphasis>
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<taxonomicNameLabel id="A2775783FFC9FF98FEFF398A68D888C5" box="[380,473,1164,1190]" pageId="4" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B8F36EAFFC9FF98FF1439A8686688A4" blockId="4.[151,473,1163,1223]" box="[151,359,1198,1223]" pageId="4" pageNumber="81">
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<figureCitation id="130B2A6FFFC9FF98FF1439A8680888A4" box="[151,265,1198,1223]" captionStart-0="FIGURES 5 – 6" captionStart-1="FIGURES 7 – 14" captionStart-2="FIGURES 15 – 17" captionStart-3="FIGURES 18 – 21" captionStart-4="FIGURES 22 – 24" captionStartId-0="5.[151,264,1231,1254]" captionStartId-1="6.[151,264,1868,1891]" captionStartId-2="8.[151,264,953,976]" captionStartId-3="8.[151,264,1839,1862]" captionStartId-4="9.[151,264,1911,1934]" captionTargetBox-0="[465,1121,301,1210]" captionTargetBox-1="[180,1411,203,1844]" captionTargetBox-2="[248,1321,223,912]" captionTargetBox-3="[249,1319,1050,1821]" captionTargetBox-4="[299,1282,207,1875]" captionTargetId-0="figure@5.[465,1121,301,1210]" captionTargetId-1="figure@6.[170,1424,193,1850]" captionTargetId-2="figure@8.[244,1333,195,933]" captionTargetId-3="figure@8.[249,1320,1050,1822]" captionTargetId-4="figure@9.[286,1297,193,1889]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="8" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="FIGURES 5 – 6. Plutomurus revazi sp. nov. 5, habitus (light microscopy); 6, habitus (SEM) (bar: 0.4 mm)." captionText-1="FIGURES 7 – 14. Plutomurus revazi sp. nov. 7, labrum (bar: 0.04 mm); 8, labial palp (bar: 0.02 mm); 9, outer lobe of maxillary palp (bar: 0.02 mm); 10, trochanteral and femoral organs (bar: 0.06 mm); 11, leg III tibiotarsus, showing the location of the spine-like chaetae (bar: 0.06 mm) (some ordinary chaetae are illustrated for comparison); 12 – 14, claw and Emp. I – III respectively (bar: 0.05 mm) (a, basal tooth to internal lamellae; b, first unpaired tooth, the more developed, c and d, additional unpaired teeth; The arrows point each tooth)." captionText-2="FIGURES 15 – 17. Plutomurus revazi sp. nov. 15, mucro (bar: 0.03 mm) (The arrows show the presence of the two medial teeth); 16, spines on dens (bar: 0.05 mm); 17, tenaculum (bar: 0.03 mm)." captionText-3="FIGURES 18 – 21. Plutomurus revazi sp. nov. 18, SEM microphotograph of claw I and Emp. I (bar: 0.03 mm); 18 ’, detail of internal lamellae on claw I, showing teeth: a, basal to internal lamellae; b, first unpaired tooth, the more developed (broken), c and d, additional unpaired teeth, not always presents); 19, detail of Emp. I, showing the two internal ridges with three denticles on each (bar: 0.01 mm); 20, Abd. VI, ventral, view, and male genital plate (bar: 0.06 mm); 21, detail of male genital plate (bar: 0.02 mm)." captionText-4="FIGURES 22 – 24. Plutomurus revazi sp. nov., dorsal chaetotaxy: 22, head; 23, Th. II – III; 24, Abd. I – VI (dash-lined hollow circles represent eyes; large black circles represent macrochaetae; white circles represent mesochaetae or small macrochaetae; small black circles represent microchaetae; squares with a black dot represent bothriothrix; triangles are microsensillae; circles with a slash represent pseudopores)." httpUri-0="https://zenodo.org/record/262883/files/figure.png" httpUri-1="https://zenodo.org/record/262884/files/figure.png" httpUri-2="https://zenodo.org/record/262885/files/figure.png" httpUri-3="https://zenodo.org/record/262886/files/figure.png" httpUri-4="https://zenodo.org/record/262887/files/figure.png" pageId="4" pageNumber="81">Figs 5–24</figureCitation>
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,
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<tableCitation id="C6B20351FFC9FF98FE9739A8686688A4" box="[276,359,1198,1223]" captionStart="TABLE 2" captionStartId="7.[151,239,152,175]" captionText="TABLE 2. Plutomurus revazi sp. nov. Variability of number of teeth on claw, on each lamella of the Emp. and number of dental spines on both sides of each specimen. (Claw I – III and Emp. I – III represent I – III leg’s claw and Emp. respectively; b – with claw is basal tooth of claw; the Arabic numbers for dens mean the number of small spines; dental large spines are in Roman, bold and in Italics for basal and apical parts of dens)." pageId="4" pageNumber="81">Table 2</tableCitation>
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locality
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.
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, Imereti region, Tskaltubo district, Kumistavi village, Sataplia-Tskaltubo karst massif, Satsurblia Cave,
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,
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,
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.
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alt.
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, female in slide: dark zone,
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(code CAU-SATSURBLIA 20141130 01).
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(same data as
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): three females in slide, dark zone,
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(
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(
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(
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(
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20150124 0 2, 0 5, 0 6, 11, 12); one specimen mounted on SEM stub (
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20141130 13). Deposited material:
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and
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0 4, 0 6 and 0
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; remaining
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|
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.
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFC9FF98FF443B106DE78AF8" blockId="4.[151,1437,1270,2015]" pageId="4" pageNumber="81">
|
||
<emphasis id="B944EAF8FFC9FF98FF443B1068FD8A4C" bold="true" box="[199,508,1558,1583]" pageId="4" pageNumber="81">Complementary material.</emphasis>
|
||
Six specimens in slide (codes
|
||
<collectionCode id="ED21AE2FFFC9FF98FCD73B116A8E8A4C" LSID="urn:lsid:biocol.org:col:33074" box="[852,911,1559,1583]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="4" pageNumber="81">CAU</collectionCode>
|
||
<collectionCode id="ED21AE2FFFC9FF98FC153B116D588A4C" box="[918,1113,1559,1583]" pageId="4" pageNumber="81">PROMETHEUS-</collectionCode>
|
||
20141130 0 1 to 06), and two specimens mounted on SEM stub (
|
||
<collectionCode id="ED21AE2FFFC9FF98FDA13B3D6B5E8A30" LSID="urn:lsid:biocol.org:col:33074" box="[546,607,1595,1619]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="4" pageNumber="81">CAU</collectionCode>
|
||
<collectionCode id="ED21AE2FFFC9FF98FDE43B3D6A238A30" box="[615,802,1595,1619]" pageId="4" pageNumber="81">PROMETHEUS</collectionCode>
|
||
20141130 0 7 and 08), from
|
||
<collectingCountry id="F327767AFFC9FF98FBEB3B3D6DE88A30" box="[1128,1257,1595,1619]" name="Georgia" pageId="4" pageNumber="81">GEORGIA</collectingCountry>
|
||
, Imereti region, Tskaltubo district, Kumistavi village, Sataplia-Tskaltubo karst massif, Prometheus Cave,
|
||
<geoCoordinate id="EE04502DFFC9FF98FB5E3B586C998A15" box="[1245,1432,1630,1655]" direction="north" orientation="latitude" pageId="4" pageNumber="81" precision="1" value="42.377235">42°22'38.05" N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE04502DFFC9FF98FF143B8468378AF9" box="[151,310,1666,1691]" direction="east" orientation="longitude" pageId="4" pageNumber="81" precision="1" value="42.600945">42°36'3.40" E</geoCoordinate>
|
||
,
|
||
<specimenCount id="9D36FD63FFC9FF98FEC13B8468918AF9" box="[322,400,1666,1691]" pageId="4" pageNumber="81" type="generic">
|
||
<quantity id="4CC89B0FFFC9FF98FEC13B84688D8AF9" box="[322,396,1666,1691]" metricMagnitude="2" metricUnit="m" metricValue="1.75" pageId="4" pageNumber="81" unit="m" value="175.0">175 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
alt., dark zone,
|
||
<date id="FF8E102AFFC9FF98FDC03B846BBE8AF8" box="[579,703,1666,1691]" pageId="4" pageNumber="81" value="2014-11-30">30.xi.2014</date>
|
||
. Specimens 0 1 and 0
|
||
<quantity id="4CC89B0FFFC9FF98FC2A3B856AD48AF8" box="[937,981,1666,1691]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="4" pageNumber="81" unit="in" value="4.0">4 in</quantity>
|
||
<collectionCode id="ED21AE2FFFC9FF98FC583B856D318AF8" box="[987,1072,1667,1691]" httpUri="http://grbio.org/cool/s63m-4f6f" name="Universidad de Navarra, Museum of Zoology" pageId="4" pageNumber="81">MZNA</collectionCode>
|
||
, the rest in
|
||
<collectionCode id="ED21AE2FFFC9FF98FB323B856DE38AF8" box="[1201,1250,1667,1691]" httpUri="http://grbio.org/cool/jgjv-qdg3" name="Indiana State University" pageId="4" pageNumber="81">ISU</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFC9FF98FF443BA068938A80" blockId="4.[151,1437,1270,2015]" pageId="4" pageNumber="81">
|
||
The new species was previously mentioned from Prometheus and Satsurblia caves as “
|
||
<taxonomicName id="4C304D69FFC9FF98FB4B3BA06C4A8ADC" box="[1224,1355,1702,1727]" class="Entognatha" family="Tomoceridae" genus="Plutomurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="4" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944EAF8FFC9FF98FB4B3BA06C4A8ADC" box="[1224,1355,1702,1727]" italics="true" pageId="4" pageNumber="81">Plutomurus</emphasis>
|
||
</taxonomicName>
|
||
sp. 2” (Barjadze
|
||
<emphasis id="B944EAF8FFC9FF98FE883BCA683D8A80" box="[267,316,1738,1763]" italics="true" pageId="4" pageNumber="81">et al</emphasis>
|
||
. 2015).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A6561FFC9FF9BFF443BE869EE8AB6" lastPageId="7" lastPageNumber="84" pageId="4" pageNumber="81" type="description">
|
||
<paragraph id="8B8F36EAFFC9FF98FF443BE86D488B10" blockId="4.[151,1437,1270,2015]" pageId="4" pageNumber="81">
|
||
<emphasis id="B944EAF8FFC9FF98FF443BE868588B64" bold="true" box="[199,345,1774,1799]" pageId="4" pageNumber="81">Description.</emphasis>
|
||
Body length up to
|
||
<quantity id="4CC89B0FFFC9FF98FDD13BE86BA98B65" box="[594,680,1774,1799]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="81" unit="mm" value="5.6">5.6 mm</quantity>
|
||
excluding antennae (
|
||
<typeStatus id="548B8848FFC9FF98FC283BE86D0E8B64" box="[939,1039,1774,1799]" pageId="4" pageNumber="81" type="holotype">holotype</typeStatus>
|
||
<quantity id="4CC89B0FFFC9FF98FB9E3BE86D758B64" box="[1053,1140,1774,1799]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="4" pageNumber="81" unit="mm" value="3.6">3.6 mm</quantity>
|
||
) (Satsurblia Cave, mean
|
||
<quantity id="4CC89B0FFFC9FF98FF143A1469FC8B49" box="[151,253,1810,1835]" metricMagnitude="-3" metricUnit="m" metricValue="4.56" pageId="4" pageNumber="81" unit="mm" value="4.56">4.56 mm</quantity>
|
||
, n = 10; Prometheus Cave, mean
|
||
<quantity id="4CC89B0FFFC9FF98FDFA3A156BDE8B49" box="[633,735,1811,1835]" metricMagnitude="-3" metricUnit="m" metricValue="3.42" pageId="4" pageNumber="81" unit="mm" value="3.42">3.42 mm</quantity>
|
||
, n = 6). Body cuticular, colour brownish-grey, with patches of darker pigmentation over the whole body except on antennae, legs and dens; scales covering antennal I and II segments, head, body and legs; colour and scale patterns, and habitus as in
|
||
<figureCitation id="130B2A6FFFC9FF98FC633A5C6D448B10" box="[992,1093,1882,1907]" captionStart="FIGURES 5 – 6" captionStartId="5.[151,264,1231,1254]" captionTargetBox="[465,1121,301,1210]" captionTargetId="figure@5.[465,1121,301,1210]" captionTargetPageId="5" captionText="FIGURES 5 – 6. Plutomurus revazi sp. nov. 5, habitus (light microscopy); 6, habitus (SEM) (bar: 0.4 mm)." httpUri="https://zenodo.org/record/262883/files/figure.png" pageId="4" pageNumber="81">Figs 5–6</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFC9FF99FF443A786A748C9B" blockId="4.[151,1437,1270,2015]" lastBlockId="5.[151,1436,151,248]" lastPageId="5" lastPageNumber="82" pageId="4" pageNumber="81">
|
||
<emphasis id="B944EAF8FFC9FF98FF443A78680A8BF4" box="[199,267,1918,1943]" italics="true" pageId="4" pageNumber="81">Head.</emphasis>
|
||
Eye patch small, eyes not visible under light microscope, while in SEM exterior faded 3-4 (5) cornea present. Ratio of body length to antennae length up to 1.31 (
|
||
<typeStatus id="548B8848FFC9FF98FCD73AA46AB98BD8" box="[852,952,1954,1979]" pageId="4" pageNumber="81" type="holotype">holotype</typeStatus>
|
||
1.02) (Satsurblia Cave, mean 1.18, n = 6; Prometheus Cave, mean 0.99, n = 6). Prelabral chaetae 3 + 3; labrum with 5, 5, 4 chaetae arising from tubercles and with four curved hooks on distal part of labrum (
|
||
<figureCitation id="130B2A6FFFC8FF99FC8A3D916A4C8CD3" box="[777,845,151,176]" captionStart="FIGURES 7 – 14" captionStartId="6.[151,264,1868,1891]" captionTargetBox="[180,1411,203,1844]" captionTargetId="figure@6.[170,1424,193,1850]" captionTargetPageId="6" captionText="FIGURES 7 – 14. Plutomurus revazi sp. nov. 7, labrum (bar: 0.04 mm); 8, labial palp (bar: 0.02 mm); 9, outer lobe of maxillary palp (bar: 0.02 mm); 10, trochanteral and femoral organs (bar: 0.06 mm); 11, leg III tibiotarsus, showing the location of the spine-like chaetae (bar: 0.06 mm) (some ordinary chaetae are illustrated for comparison); 12 – 14, claw and Emp. I – III respectively (bar: 0.05 mm) (a, basal tooth to internal lamellae; b, first unpaired tooth, the more developed, c and d, additional unpaired teeth; The arrows point each tooth)." httpUri="https://zenodo.org/record/262884/files/figure.png" pageId="5" pageNumber="82">Fig. 7</figureCitation>
|
||
). Maxillary palp trifurcate, chaeta on distal papilla longer than basal papilla; sublobal plate with four sublobals, all shorter than basal chaeta (
|
||
<figureCitation id="130B2A6FFFC8FF99FBFC3DBA6DC28CB7" box="[1151,1219,188,213]" captionStart="FIGURES 7 – 14" captionStartId="6.[151,264,1868,1891]" captionTargetBox="[180,1411,203,1844]" captionTargetId="figure@6.[170,1424,193,1850]" captionTargetPageId="6" captionText="FIGURES 7 – 14. Plutomurus revazi sp. nov. 7, labrum (bar: 0.04 mm); 8, labial palp (bar: 0.02 mm); 9, outer lobe of maxillary palp (bar: 0.02 mm); 10, trochanteral and femoral organs (bar: 0.06 mm); 11, leg III tibiotarsus, showing the location of the spine-like chaetae (bar: 0.06 mm) (some ordinary chaetae are illustrated for comparison); 12 – 14, claw and Emp. I – III respectively (bar: 0.05 mm) (a, basal tooth to internal lamellae; b, first unpaired tooth, the more developed, c and d, additional unpaired teeth; The arrows point each tooth)." httpUri="https://zenodo.org/record/262884/files/figure.png" pageId="5" pageNumber="82">Fig. 9</figureCitation>
|
||
). Labial papillae as in
|
||
<figureCitation id="130B2A6FFFC8FF99FF373DD969F68C9B" box="[180,247,223,248]" captionStart="FIGURES 7 – 14" captionStartId="6.[151,264,1868,1891]" captionTargetBox="[180,1411,203,1844]" captionTargetId="figure@6.[170,1424,193,1850]" captionTargetPageId="6" captionText="FIGURES 7 – 14. Plutomurus revazi sp. nov. 7, labrum (bar: 0.04 mm); 8, labial palp (bar: 0.02 mm); 9, outer lobe of maxillary palp (bar: 0.02 mm); 10, trochanteral and femoral organs (bar: 0.06 mm); 11, leg III tibiotarsus, showing the location of the spine-like chaetae (bar: 0.06 mm) (some ordinary chaetae are illustrated for comparison); 12 – 14, claw and Emp. I – III respectively (bar: 0.05 mm) (a, basal tooth to internal lamellae; b, first unpaired tooth, the more developed, c and d, additional unpaired teeth; The arrows point each tooth)." httpUri="https://zenodo.org/record/262884/files/figure.png" pageId="5" pageNumber="82">Fig. 8</figureCitation>
|
||
; proximal labial chaetae up to 16; labial chaetae smooth.
|
||
</paragraph>
|
||
<caption id="DF4F6662FFC8FF99FF1439C96DD18886" box="[151,1232,1231,1254]" httpUri="https://zenodo.org/record/262883/files/figure.png" pageId="5" pageNumber="82" targetBox="[465,1121,301,1210]" targetPageId="5">
|
||
<paragraph id="8B8F36EAFFC8FF99FF1439C96DD18886" blockId="5.[151,1232,1231,1254]" box="[151,1232,1231,1254]" pageId="5" pageNumber="82">
|
||
<emphasis id="B944EAF8FFC8FF99FF1439C9683B8887" bold="true" box="[151,314,1231,1254]" pageId="5" pageNumber="82">FIGURES 5–6.</emphasis>
|
||
<taxonomicName id="4C304D69FFC8FF99FEC339C968FB8885" box="[320,506,1231,1254]" class="Entognatha" family="Tomoceridae" genus="Plutomurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="5" pageNumber="82" phylum="Arthropoda" rank="species" species="revazi" status="sp. nov.">
|
||
<emphasis id="B944EAF8FFC8FF99FEC339C968FB8885" box="[320,506,1231,1254]" italics="true" pageId="5" pageNumber="82">Plutomurus revazi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2775783FFC8FF99FD8339D66B518886" box="[512,592,1232,1253]" pageId="5" pageNumber="82" rank="species">
|
||
<emphasis id="B944EAF8FFC8FF99FD8339D66B518886" bold="true" box="[512,592,1232,1253]" pageId="5" pageNumber="82">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
5, habitus (light microscopy); 6, habitus (SEM) (bar: 0.4 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8F36EAFFC8FF99FF4438146DB88A9C" blockId="5.[151,1437,1298,1941]" pageId="5" pageNumber="82">
|
||
<emphasis id="B944EAF8FFC8FF99FF44381468078948" box="[199,262,1298,1323]" italics="true" pageId="5" pageNumber="82">Body.</emphasis>
|
||
Third legs with well developed trochanteral and femoral organs, with about 17 and 40 chaetae respectively, a few chaetae longer than others (
|
||
<figureCitation id="130B2A6FFFC8FF99FD4A38306A1D892C" box="[713,796,1334,1359]" captionStart="FIGURES 7 – 14" captionStartId="6.[151,264,1868,1891]" captionTargetBox="[180,1411,203,1844]" captionTargetId="figure@6.[170,1424,193,1850]" captionTargetPageId="6" captionText="FIGURES 7 – 14. Plutomurus revazi sp. nov. 7, labrum (bar: 0.04 mm); 8, labial palp (bar: 0.02 mm); 9, outer lobe of maxillary palp (bar: 0.02 mm); 10, trochanteral and femoral organs (bar: 0.06 mm); 11, leg III tibiotarsus, showing the location of the spine-like chaetae (bar: 0.06 mm) (some ordinary chaetae are illustrated for comparison); 12 – 14, claw and Emp. I – III respectively (bar: 0.05 mm) (a, basal tooth to internal lamellae; b, first unpaired tooth, the more developed, c and d, additional unpaired teeth; The arrows point each tooth)." httpUri="https://zenodo.org/record/262884/files/figure.png" pageId="5" pageNumber="82">Fig. 10</figureCitation>
|
||
). Ventral tube with 65 distal, 49 + 49 anterior and 54 posterior chaetae in one specimen. Legs scaled to tibiotarsus. Tibiotarsus III with two ventral spine-like chaetae (
|
||
<figureCitation id="130B2A6FFFC8FF99FF1C387869F089F4" box="[159,241,1406,1431]" captionStart="FIGURES 7 – 14" captionStartId="6.[151,264,1868,1891]" captionTargetBox="[180,1411,203,1844]" captionTargetId="figure@6.[170,1424,193,1850]" captionTargetPageId="6" captionText="FIGURES 7 – 14. Plutomurus revazi sp. nov. 7, labrum (bar: 0.04 mm); 8, labial palp (bar: 0.02 mm); 9, outer lobe of maxillary palp (bar: 0.02 mm); 10, trochanteral and femoral organs (bar: 0.06 mm); 11, leg III tibiotarsus, showing the location of the spine-like chaetae (bar: 0.06 mm) (some ordinary chaetae are illustrated for comparison); 12 – 14, claw and Emp. I – III respectively (bar: 0.05 mm) (a, basal tooth to internal lamellae; b, first unpaired tooth, the more developed, c and d, additional unpaired teeth; The arrows point each tooth)." httpUri="https://zenodo.org/record/262884/files/figure.png" pageId="5" pageNumber="82">Fig. 11</figureCitation>
|
||
), such chaetae absent on tibiotarsi I to II (formula 002). Claw on all legs with characteristic small, unpaired tooth between the two lamellae (
|
||
<figureCitation id="130B2A6FFFC8FF99FD8638A46B8789D8" box="[517,646,1442,1467]" captionStart="FIGURES 7 – 14" captionStartId="6.[151,264,1868,1891]" captionTargetBox="[180,1411,203,1844]" captionTargetId="figure@6.[170,1424,193,1850]" captionTargetPageId="6" captionText="FIGURES 7 – 14. Plutomurus revazi sp. nov. 7, labrum (bar: 0.04 mm); 8, labial palp (bar: 0.02 mm); 9, outer lobe of maxillary palp (bar: 0.02 mm); 10, trochanteral and femoral organs (bar: 0.06 mm); 11, leg III tibiotarsus, showing the location of the spine-like chaetae (bar: 0.06 mm) (some ordinary chaetae are illustrated for comparison); 12 – 14, claw and Emp. I – III respectively (bar: 0.05 mm) (a, basal tooth to internal lamellae; b, first unpaired tooth, the more developed, c and d, additional unpaired teeth; The arrows point each tooth)." httpUri="https://zenodo.org/record/262884/files/figure.png" pageId="5" pageNumber="82">Figs 12–14</figureCitation>
|
||
,
|
||
<figureCitation id="130B2A6FFFC8FF99FD1138A56BB089D8" box="[658,689,1443,1467]" captionStart="FIGURES 18 – 21" captionStartId="8.[151,264,1839,1862]" captionTargetBox="[249,1319,1050,1821]" captionTargetId="figure@8.[249,1320,1050,1822]" captionTargetPageId="8" captionText="FIGURES 18 – 21. Plutomurus revazi sp. nov. 18, SEM microphotograph of claw I and Emp. I (bar: 0.03 mm); 18 ’, detail of internal lamellae on claw I, showing teeth: a, basal to internal lamellae; b, first unpaired tooth, the more developed (broken), c and d, additional unpaired teeth, not always presents); 19, detail of Emp. I, showing the two internal ridges with three denticles on each (bar: 0.01 mm); 20, Abd. VI, ventral, view, and male genital plate (bar: 0.06 mm); 21, detail of male genital plate (bar: 0.02 mm)." httpUri="https://zenodo.org/record/262886/files/figure.png" pageId="5" pageNumber="82">18</figureCitation>
|
||
); number of internal unpaired teeth varying from 2 to 5, (
|
||
<tableCitation id="C6B20351FFC8FF99FAB438A46C8F89D8" box="[1335,1422,1442,1467]" captionStart="TABLE 2" captionStartId="7.[151,239,152,175]" captionText="TABLE 2. Plutomurus revazi sp. nov. Variability of number of teeth on claw, on each lamella of the Emp. and number of dental spines on both sides of each specimen. (Claw I – III and Emp. I – III represent I – III leg’s claw and Emp. respectively; b – with claw is basal tooth of claw; the Arabic numbers for dens mean the number of small spines; dental large spines are in Roman, bold and in Italics for basal and apical parts of dens)." pageId="5" pageNumber="82">Table 2</tableCitation>
|
||
), with a minute basal tooth at 20–40% of the claw length from the base, at base of the two internal lamellae; three unpaired teeth usually present, second and third teeth occasionally absent, formed by the fusion of both lamellae; distal tooth formed by terminal fusion of lamellae (
|
||
<figureCitation id="130B2A6FFFC8FF99FD673B086A388A44" box="[740,825,1550,1575]" captionStart="FIGURES 18 – 21" captionStartId="8.[151,264,1839,1862]" captionTargetBox="[249,1319,1050,1821]" captionTargetId="figure@8.[249,1320,1050,1822]" captionTargetPageId="8" captionText="FIGURES 18 – 21. Plutomurus revazi sp. nov. 18, SEM microphotograph of claw I and Emp. I (bar: 0.03 mm); 18 ’, detail of internal lamellae on claw I, showing teeth: a, basal to internal lamellae; b, first unpaired tooth, the more developed (broken), c and d, additional unpaired teeth, not always presents); 19, detail of Emp. I, showing the two internal ridges with three denticles on each (bar: 0.01 mm); 20, Abd. VI, ventral, view, and male genital plate (bar: 0.06 mm); 21, detail of male genital plate (bar: 0.02 mm)." httpUri="https://zenodo.org/record/262886/files/figure.png" pageId="5" pageNumber="82">Fig. 18</figureCitation>
|
||
’); Emp. blade-like, usually with two internal serrated lamellae bearing 0–8 teeth starting near distal part and extending to base (
|
||
<figureCitation id="130B2A6FFFC8FF99FC513B346D528A28" box="[978,1107,1586,1611]" captionStart="FIGURES 7 – 14" captionStartId="6.[151,264,1868,1891]" captionTargetBox="[180,1411,203,1844]" captionTargetId="figure@6.[170,1424,193,1850]" captionTargetPageId="6" captionText="FIGURES 7 – 14. Plutomurus revazi sp. nov. 7, labrum (bar: 0.04 mm); 8, labial palp (bar: 0.02 mm); 9, outer lobe of maxillary palp (bar: 0.02 mm); 10, trochanteral and femoral organs (bar: 0.06 mm); 11, leg III tibiotarsus, showing the location of the spine-like chaetae (bar: 0.06 mm) (some ordinary chaetae are illustrated for comparison); 12 – 14, claw and Emp. I – III respectively (bar: 0.05 mm) (a, basal tooth to internal lamellae; b, first unpaired tooth, the more developed, c and d, additional unpaired teeth; The arrows point each tooth)." httpUri="https://zenodo.org/record/262884/files/figure.png" pageId="5" pageNumber="82">Figs 12–14</figureCitation>
|
||
,
|
||
<figureCitation id="130B2A6FFFC8FF99FBDC3B356DA68A28" box="[1119,1191,1587,1611]" captionStart="FIGURES 18 – 21" captionStartId="8.[151,264,1839,1862]" captionTargetBox="[249,1319,1050,1821]" captionTargetId="figure@8.[249,1320,1050,1822]" captionTargetPageId="8" captionText="FIGURES 18 – 21. Plutomurus revazi sp. nov. 18, SEM microphotograph of claw I and Emp. I (bar: 0.03 mm); 18 ’, detail of internal lamellae on claw I, showing teeth: a, basal to internal lamellae; b, first unpaired tooth, the more developed (broken), c and d, additional unpaired teeth, not always presents); 19, detail of Emp. I, showing the two internal ridges with three denticles on each (bar: 0.01 mm); 20, Abd. VI, ventral, view, and male genital plate (bar: 0.06 mm); 21, detail of male genital plate (bar: 0.02 mm)." httpUri="https://zenodo.org/record/262886/files/figure.png" pageId="5" pageNumber="82">18–19</figureCitation>
|
||
;
|
||
<tableCitation id="C6B20351FFC8FF99FB373B346C088A28" box="[1204,1289,1586,1611]" captionStart="TABLE 2" captionStartId="7.[151,239,152,175]" captionText="TABLE 2. Plutomurus revazi sp. nov. Variability of number of teeth on claw, on each lamella of the Emp. and number of dental spines on both sides of each specimen. (Claw I – III and Emp. I – III represent I – III leg’s claw and Emp. respectively; b – with claw is basal tooth of claw; the Arabic numbers for dens mean the number of small spines; dental large spines are in Roman, bold and in Italics for basal and apical parts of dens)." pageId="5" pageNumber="82">Table 2</tableCitation>
|
||
). Tenent hair long, clavate. Ratio of claw III: Emp. III: tenent hair 1.8: 1.1: 1 (
|
||
<typeStatus id="548B8848FFC8FF99FC213B506D0B8A0C" box="[930,1034,1622,1647]" pageId="5" pageNumber="82" type="holotype">holotype</typeStatus>
|
||
); claw with a thin pseudonychium, 0.76 times length of inner edge of claw and two basal and lateral teeth. Tenaculum with 4 + 4 teeth and one chaeta on corpus (
|
||
<figureCitation id="130B2A6FFFC8FF99FE9A3B98686D8AD4" box="[281,364,1694,1719]" captionStart="FIGURES 15 – 17" captionStartId="8.[151,264,953,976]" captionTargetBox="[248,1321,223,912]" captionTargetId="figure@8.[244,1333,195,933]" captionTargetPageId="8" captionText="FIGURES 15 – 17. Plutomurus revazi sp. nov. 15, mucro (bar: 0.03 mm) (The arrows show the presence of the two medial teeth); 16, spines on dens (bar: 0.05 mm); 17, tenaculum (bar: 0.03 mm)." httpUri="https://zenodo.org/record/262885/files/figure.png" pageId="5" pageNumber="82">Fig. 17</figureCitation>
|
||
). Ratio of manubrium: dens: mucro 5.7: 7.5: 1 (n = 8, including
|
||
<typeStatus id="548B8848FFC8FF99FBE43B986DCF8AD4" box="[1127,1230,1694,1719]" pageId="5" pageNumber="82" type="holotype">holotype</typeStatus>
|
||
). Outer margin of dens with four thick, spine-like chaetae; dental formula very variable (
|
||
<tableCitation id="C6B20351FFC8FF99FC253BC46AFD8AB8" box="[934,1020,1730,1755]" captionStart="TABLE 2" captionStartId="7.[151,239,152,175]" captionText="TABLE 2. Plutomurus revazi sp. nov. Variability of number of teeth on claw, on each lamella of the Emp. and number of dental spines on both sides of each specimen. (Claw I – III and Emp. I – III represent I – III leg’s claw and Emp. respectively; b – with claw is basal tooth of claw; the Arabic numbers for dens mean the number of small spines; dental large spines are in Roman, bold and in Italics for basal and apical parts of dens)." pageId="5" pageNumber="82">Table 2</tableCitation>
|
||
) (
|
||
<figureCitation id="130B2A6FFFC8FF99FB903BC46D678AB8" box="[1043,1126,1730,1755]" captionStart="FIGURES 15 – 17" captionStartId="8.[151,264,953,976]" captionTargetBox="[248,1321,223,912]" captionTargetId="figure@8.[244,1333,195,933]" captionTargetPageId="8" captionText="FIGURES 15 – 17. Plutomurus revazi sp. nov. 15, mucro (bar: 0.03 mm) (The arrows show the presence of the two medial teeth); 16, spines on dens (bar: 0.05 mm); 17, tenaculum (bar: 0.03 mm)." httpUri="https://zenodo.org/record/262885/files/figure.png" pageId="5" pageNumber="82">Fig. 16</figureCitation>
|
||
). Mucro with 2 basal, fused teeth, 0–2 intermediate denticles, and 2 distal teeth (
|
||
<figureCitation id="130B2A6FFFC8FF99FD5C3BE06A338A9C" box="[735,818,1766,1791]" captionStart="FIGURES 15 – 17" captionStartId="8.[151,264,953,976]" captionTargetBox="[248,1321,223,912]" captionTargetId="figure@8.[244,1333,195,933]" captionTargetPageId="8" captionText="FIGURES 15 – 17. Plutomurus revazi sp. nov. 15, mucro (bar: 0.03 mm) (The arrows show the presence of the two medial teeth); 16, spines on dens (bar: 0.05 mm); 17, tenaculum (bar: 0.03 mm)." httpUri="https://zenodo.org/record/262885/files/figure.png" pageId="5" pageNumber="82">Fig. 15</figureCitation>
|
||
). Male genital plate in
|
||
<figureCitation id="130B2A6FFFC8FF99FBB63BE06DB78A9C" box="[1077,1206,1766,1791]" captionStart="FIGURES 18 – 21" captionStartId="8.[151,264,1839,1862]" captionTargetBox="[249,1319,1050,1821]" captionTargetId="figure@8.[249,1320,1050,1822]" captionTargetPageId="8" captionText="FIGURES 18 – 21. Plutomurus revazi sp. nov. 18, SEM microphotograph of claw I and Emp. I (bar: 0.03 mm); 18 ’, detail of internal lamellae on claw I, showing teeth: a, basal to internal lamellae; b, first unpaired tooth, the more developed (broken), c and d, additional unpaired teeth, not always presents); 19, detail of Emp. I, showing the two internal ridges with three denticles on each (bar: 0.01 mm); 20, Abd. VI, ventral, view, and male genital plate (bar: 0.06 mm); 21, detail of male genital plate (bar: 0.02 mm)." httpUri="https://zenodo.org/record/262886/files/figure.png" pageId="5" pageNumber="82">Figs 20–21</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFC8FF99FF443A0C6B7E8BF7" blockId="5.[151,1437,1298,1941]" pageId="5" pageNumber="82">
|
||
<emphasis id="B944EAF8FFC8FF99FF443A0C68548B40" bold="true" box="[199,341,1802,1827]" pageId="5" pageNumber="82">Chaetotaxy.</emphasis>
|
||
Dorsally Mac on each side: Th. II–III with 5 (m3, m4, m5, p3, p4) and 2 (p3, p5) Mac, respectively; Abd. I–V with 3 (m2, m3, m4) 3 (m2, m3, m4) 4 (m2, m6, p1, p6) 3 (A6, C6, E3) and 3 (p2, p3, p4) Mac, respectively. Bothriotrichal formula 0/2–1/0–0–1–2–0. Bothriothrix bordered anteriorly by 1–3 small chaetae. Macrochaetotaxy and Bothriotrichal pattern as in
|
||
<figureCitation id="130B2A6FFFC8FF99FE793A7A6B7A8BF7" box="[506,635,1916,1941]" captionStart="FIGURES 22 – 24" captionStartId="9.[151,264,1911,1934]" captionTargetBox="[299,1282,207,1875]" captionTargetId="figure@9.[286,1297,193,1889]" captionTargetPageId="9" captionText="FIGURES 22 – 24. Plutomurus revazi sp. nov., dorsal chaetotaxy: 22, head; 23, Th. II – III; 24, Abd. I – VI (dash-lined hollow circles represent eyes; large black circles represent macrochaetae; white circles represent mesochaetae or small macrochaetae; small black circles represent microchaetae; squares with a black dot represent bothriothrix; triangles are microsensillae; circles with a slash represent pseudopores)." httpUri="https://zenodo.org/record/262887/files/figure.png" pageId="5" pageNumber="82">Figs 22–24</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF4F6662FFCBFF9AFF143A4A6B618BBD" httpUri="https://zenodo.org/record/262884/files/figure.png" pageId="6" pageNumber="83" targetBox="[180,1411,203,1844]" targetPageId="6">
|
||
<paragraph id="8B8F36EAFFCBFF9AFF143A4A6B618BBD" blockId="6.[151,1436,1868,2014]" pageId="6" pageNumber="83">
|
||
<emphasis id="B944EAF8FFCBFF9AFF143A4A68468B01" bold="true" box="[151,327,1868,1891]" pageId="6" pageNumber="83">FIGURES 7–14.</emphasis>
|
||
<taxonomicName id="4C304D69FFCBFF9AFECF3A4B6B048B00" box="[332,517,1869,1891]" class="Entognatha" family="Tomoceridae" genus="Plutomurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="6" pageNumber="83" phylum="Arthropoda" rank="species" species="revazi" status="sp. nov.">
|
||
<emphasis id="B944EAF8FFCBFF9AFECF3A4B6B048B00" box="[332,517,1869,1891]" italics="true" pageId="6" pageNumber="83">Plutomurus revazi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2775783FFCBFF9AFD883A486B5A8B00" box="[523,603,1870,1891]" pageId="6" pageNumber="83" rank="species">
|
||
<emphasis id="B944EAF8FFCBFF9AFD883A486B5A8B00" bold="true" box="[523,603,1870,1891]" pageId="6" pageNumber="83">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
7, labrum (bar: 0.04 mm); 8, labial palp (bar: 0.02 mm); 9, outer lobe of maxillary palp (bar: 0.02 mm); 10, trochanteral and femoral organs (bar: 0.06 mm); 11, leg III tibiotarsus, showing the location of the spine-like chaetae (bar: 0.06 mm) (some ordinary chaetae are illustrated for comparison); 12–14, claw and Emp. I–III respectively (bar: 0.05 mm) (a, basal tooth to internal lamellae; b, first unpaired tooth, the more developed, c and d, additional unpaired teeth; The arrows point each tooth).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F6662FFCAFF9BFF143D9E6AEF8D6C" pageId="7" pageNumber="84">
|
||
<paragraph id="8B8F36EAFFCAFF9BFF143D9E6AEF8D6C" blockId="7.[151,1437,151,271]" pageId="7" pageNumber="84">
|
||
<emphasis id="B944EAF8FFCAFF9BFF143D9E68088CCC" bold="true" box="[151,265,152,175]" pageId="7" pageNumber="84">TABLE 2.</emphasis>
|
||
<taxonomicName id="4C304D69FFCAFF9BFE9B3D9168DD8CCC" box="[280,476,151,175]" class="Entognatha" family="Tomoceridae" genus="Plutomurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="revazi" status="sp. nov.">
|
||
<emphasis id="B944EAF8FFCAFF9BFE9B3D9168DD8CCC" box="[280,476,151,175]" italics="true" pageId="7" pageNumber="84">Plutomurus revazi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2775783FFCAFF9BFE673D9E6B3B8CCC" box="[484,570,152,175]" pageId="7" pageNumber="84" rank="species">
|
||
<emphasis id="B944EAF8FFCAFF9BFE673D9E6B3B8CCC" bold="true" box="[484,570,152,175]" pageId="7" pageNumber="84">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Variability of number of teeth on claw, on each lamella of the Emp. and number of dental spines on both sides of each specimen. (Claw I
|
||
<emphasis id="B944EAF8FFCAFF9BFCA03DBE6A318CAC" bold="true" box="[803,816,184,207]" pageId="7" pageNumber="84">–</emphasis>
|
||
III and Emp. I
|
||
<emphasis id="B944EAF8FFCAFF9BFC553DBE6AE28CAC" bold="true" box="[982,995,184,207]" pageId="7" pageNumber="84">–</emphasis>
|
||
III represent I
|
||
<emphasis id="B944EAF8FFCAFF9BFB033DBE6D8C8CAC" bold="true" box="[1152,1165,184,207]" pageId="7" pageNumber="84">–</emphasis>
|
||
III leg’s claw and Emp. respectively; b–with claw is basal tooth of claw; the Arabic numbers for dens mean the number of small spines; dental large spines are in Roman, bold and in Italics for basal and apical parts of dens).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C3C2C6C478D23" blockId="7.[159,1407,298,1476]" box="[159,1350,298,320]" pageId="7" pageNumber="84">Specimen Claw I Emp. I Claw II Emp. II Claw III Emp. III dens dens apical</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C3C4F6D9E8D3C" blockId="7.[159,1407,298,1476]" box="[159,1183,329,351]" pageId="7" pageNumber="84">number basal</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C3C746C398DD2" blockId="7.[159,1407,298,1476]" pageId="7" pageNumber="84">
|
||
<typeStatus id="548B8848FFCAFF9BFF1C3C7469FC8DEB" box="[159,253,370,392]" pageId="7" pageNumber="84" type="holotype">Holotype</typeStatus>
|
||
3+1b 3+3 2+1b 4+3 3+1b 5+4
|
||
<date id="FF8E102AFFCAFF9BFBE83C756C398DEB" box="[1131,1336,370,392]" pageId="7" pageNumber="84">
|
||
7
|
||
<emphasis id="B944EAF8FFCAFF9BFBFD3C756D9D8DEB" bold="true" box="[1150,1180,371,392]" italics="true" pageId="7" pageNumber="84">III</emphasis>
|
||
7
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B3C756DF38DEB" bold="true" box="[1256,1266,371,392]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFA883C756C148DEB" bold="true" box="[1291,1301,371,392]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD3C756C398DEB" bold="true" box="[1326,1336,371,392]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
3+1b 5+? 3+1b 4+3 3+1b 4+0 10
|
||
<emphasis id="B944EAF8FFCAFF9BFB083C9A6D9E8DD2" bold="true" box="[1163,1183,412,433]" italics="true" pageId="7" pageNumber="84">II</emphasis>
|
||
<date id="FF8E102AFFCAFF9BFB563C9A6C398DD2" box="[1237,1336,412,434]" pageId="7" pageNumber="84">
|
||
4
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B3C9A6DF38DD2" bold="true" box="[1256,1266,412,433]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFA883C9A6C148DD2" bold="true" box="[1291,1301,412,433]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD3C9A6C398DD2" bold="true" box="[1326,1336,412,433]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C3CC06C5D8E67" blockId="7.[159,1407,298,1476]" pageId="7" pageNumber="84">
|
||
<typeStatus id="548B8848FFCAFF9BFF1C3CC069F98DB8" box="[159,248,454,475]" pageId="7" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
1 3+1b 2+0 3+1b - 3+1b 5+
|
||
<date id="FF8E102AFFCAFF9BFB813CC36D9E8DB9" box="[1026,1183,453,475]" pageId="7" pageNumber="84" value="1923-02-03">
|
||
3 23
|
||
<emphasis id="B944EAF8FFCAFF9BFB083CC36D9E8DB9" bold="true" box="[1163,1183,453,474]" italics="true" pageId="7" pageNumber="84">II</emphasis>
|
||
</date>
|
||
<date id="FF8E102AFFCAFF9BFB563CC36C5D8DB9" box="[1237,1372,453,475]" pageId="7" pageNumber="84">
|
||
4
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B3CC36DF38DB9" bold="true" box="[1256,1266,453,474]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
7
|
||
<emphasis id="B944EAF8FFCAFF9BFA883CC36C148DB9" bold="true" box="[1291,1301,453,474]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD3CC36C398DB9" bold="true" box="[1326,1336,453,474]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
1
|
||
<emphasis id="B944EAF8FFCAFF9BFAD13CC36C5D8DB9" bold="true" box="[1362,1372,453,474]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
0
|
||
<date id="FF8E102AFFCAFF9BFAF63CC3685B8E67" pageId="7" pageNumber="84">
|
||
<emphasis id="B944EAF8FFCAFF9BFAF63CC36C7E8DB9" bold="true" box="[1397,1407,453,474]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
</date>
|
||
+1b 5+2 3+1b 7+5 - -
|
||
<date id="FF8E102AFFCAFF9BFBE83CE86C5D8E67" box="[1131,1372,494,516]" pageId="7" pageNumber="84">
|
||
9
|
||
<emphasis id="B944EAF8FFCAFF9BFBFD3CE96D938E67" bold="true" box="[1150,1170,495,516]" italics="true" pageId="7" pageNumber="84">II</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B3CE96DF38E67" bold="true" box="[1256,1266,495,516]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
5
|
||
<emphasis id="B944EAF8FFCAFF9BFA883CE96C148E67" bold="true" box="[1291,1301,495,516]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD3CE96C398E67" bold="true" box="[1326,1336,495,516]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFAD13CE96C5D8E67" bold="true" box="[1362,1372,495,516]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C3F1F6C398E4E" blockId="7.[159,1407,298,1476]" box="[159,1336,536,558]" pageId="7" pageNumber="84">
|
||
<typeStatus id="548B8848FFCAFF9BFF1C3F1F69F98E4D" box="[159,248,537,558]" pageId="7" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
2 3+1b 4+3 2+1b 4+5 3+1b 6+
|
||
<date id="FF8E102AFFCAFF9BFB823F1E6D948E4E" box="[1025,1173,536,558]" pageId="7" pageNumber="84" value="2014-01-06">
|
||
6 14
|
||
<emphasis id="B944EAF8FFCAFF9BFB083F1E6D948E4E" bold="true" box="[1163,1173,536,557]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
<date id="FF8E102AFFCAFF9BFB563F1E6C398E4E" box="[1237,1336,536,558]" pageId="7" pageNumber="84">
|
||
5
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B3F1E6DF38E4E" bold="true" box="[1256,1266,536,557]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
5
|
||
<emphasis id="B944EAF8FFCAFF9BFA883F1E6C148E4E" bold="true" box="[1291,1301,536,557]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
4
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD3F1E6C398E4E" bold="true" box="[1326,1336,536,557]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFECD3F476C5D8E34" blockId="7.[159,1407,298,1476]" box="[334,1372,577,599]" pageId="7" pageNumber="84">
|
||
2+1b 4+? 2+1b 4+? 3+1b - 17
|
||
<emphasis id="B944EAF8FFCAFF9BFB083F446DA98E34" bold="true" box="[1163,1192,578,599]" italics="true" pageId="7" pageNumber="84">III</emphasis>
|
||
<date id="FF8E102AFFCAFF9BFB563F476C5D8E34" box="[1237,1372,577,599]" pageId="7" pageNumber="84">
|
||
4
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B3F446DF38E34" bold="true" box="[1256,1266,578,599]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
5
|
||
<emphasis id="B944EAF8FFCAFF9BFA883F446C148E34" bold="true" box="[1291,1301,578,599]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
1
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD3F446C398E34" bold="true" box="[1326,1336,578,599]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
4
|
||
<emphasis id="B944EAF8FFCAFF9BFAD13F446C5D8E34" bold="true" box="[1362,1372,578,599]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C3F6D6C5D8EE3" blockId="7.[159,1407,298,1476]" box="[159,1372,618,640]" pageId="7" pageNumber="84">
|
||
<typeStatus id="548B8848FFCAFF9BFF1C3F6D69F98EE3" box="[159,248,619,640]" pageId="7" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
3 3+1b 4+5 3+1b 3+1 3+1b 6+
|
||
<date id="FF8E102AFFCAFF9BFB813F6C6DA98EE3" box="[1026,1192,618,640]" pageId="7" pageNumber="84" value="2015-03-03">
|
||
3 15
|
||
<emphasis id="B944EAF8FFCAFF9BFB083F6D6DA98EE3" bold="true" box="[1163,1192,619,640]" italics="true" pageId="7" pageNumber="84">III</emphasis>
|
||
</date>
|
||
<date id="FF8E102AFFCAFF9BFB563F6C6C5D8EE3" box="[1237,1372,618,640]" pageId="7" pageNumber="84">
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B3F6D6DF38EE3" bold="true" box="[1256,1266,619,640]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFA883F6D6C148EE3" bold="true" box="[1291,1301,619,640]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD3F6D6C398EE3" bold="true" box="[1326,1336,619,640]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
4
|
||
<emphasis id="B944EAF8FFCAFF9BFAD13F6D6C5D8EE3" bold="true" box="[1362,1372,619,640]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFECD3F926DDC8EC9" blockId="7.[159,1407,298,1476]" box="[334,1245,660,682]" pageId="7" pageNumber="84">3+1b 2+3 - 4+4 3+1b - - -</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C3FB86DDC8EB0" blockId="7.[159,1407,298,1476]" box="[159,1245,701,723]" pageId="7" pageNumber="84">
|
||
<typeStatus id="548B8848FFCAFF9BFF1C3FB869F98EB0" box="[159,248,702,723]" pageId="7" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
4 3+1b 4+3 4+1b 8+3 2+1b 1+0 - -
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFECD3FE06DDC8E9F" blockId="7.[159,1407,298,1476]" box="[334,1245,742,764]" pageId="7" pageNumber="84">3+1b 5+6 - - - - - -</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C3E176C398F46" blockId="7.[159,1407,298,1476]" box="[159,1336,784,806]" pageId="7" pageNumber="84">
|
||
<typeStatus id="548B8848FFCAFF9BFF1C3E1769F98F45" box="[159,248,785,806]" pageId="7" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
5 3+1b 2+0 3+1b - 3+1b 4+
|
||
<date id="FF8E102AFFCAFF9BFB813E166D9E8F46" box="[1026,1183,784,806]" pageId="7" pageNumber="84" value="2014-02-05">
|
||
5 14
|
||
<emphasis id="B944EAF8FFCAFF9BFB083E166D9E8F46" bold="true" box="[1163,1183,784,805]" italics="true" pageId="7" pageNumber="84">II</emphasis>
|
||
</date>
|
||
<date id="FF8E102AFFCAFF9BFB563E166C398F46" box="[1237,1336,784,806]" pageId="7" pageNumber="84">
|
||
6
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B3E166DF38F46" bold="true" box="[1256,1266,784,805]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
4
|
||
<emphasis id="B944EAF8FFCAFF9BFA883E166C148F46" bold="true" box="[1291,1301,784,805]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD3E166C398F46" bold="true" box="[1326,1336,784,805]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFECD3E3F6C5D8F2C" blockId="7.[159,1407,298,1476]" box="[334,1372,825,847]" pageId="7" pageNumber="84">
|
||
1+1b 3+? 3+1b 3+3 3+1b 4+
|
||
<date id="FF8E102AFFCAFF9BFB813E3F6DA98F2C" box="[1026,1192,825,847]" pageId="7" pageNumber="84" value="2012-03-05">
|
||
5 12
|
||
<emphasis id="B944EAF8FFCAFF9BFB083E3C6DA98F2C" bold="true" box="[1163,1192,826,847]" italics="true" pageId="7" pageNumber="84">III</emphasis>
|
||
</date>
|
||
<date id="FF8E102AFFCAFF9BFB563E3F6C5D8F2C" box="[1237,1372,825,847]" pageId="7" pageNumber="84">
|
||
5
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B3E3C6DF38F2C" bold="true" box="[1256,1266,826,847]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFA883E3C6C148F2C" bold="true" box="[1291,1301,826,847]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD3E3C6C398F2C" bold="true" box="[1326,1336,826,847]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFAD13E3C6C5D8F2C" bold="true" box="[1362,1372,826,847]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C3E656C148F1B" blockId="7.[159,1407,298,1476]" box="[159,1301,866,888]" pageId="7" pageNumber="84">
|
||
<typeStatus id="548B8848FFCAFF9BFF1C3E6569F98F1B" box="[159,248,867,888]" pageId="7" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
6 2+1b 3+4 3+1b 1+1 3+1b 3+4
|
||
<date id="FF8E102AFFCAFF9BFBE83E646C148F1B" box="[1131,1301,866,888]" pageId="7" pageNumber="84">
|
||
9
|
||
<emphasis id="B944EAF8FFCAFF9BFBFD3E656D938F1B" bold="true" box="[1150,1170,867,888]" italics="true" pageId="7" pageNumber="84">II</emphasis>
|
||
9
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B3E656DF38F1B" bold="true" box="[1256,1266,867,888]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
5
|
||
<emphasis id="B944EAF8FFCAFF9BFA883E656C148F1B" bold="true" box="[1291,1301,867,888]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFECD3E8A6C398FC2" blockId="7.[159,1407,298,1476]" box="[334,1336,908,930]" pageId="7" pageNumber="84">
|
||
1+1b 2+4 - - 3+1b 4+
|
||
<date id="FF8E102AFFCAFF9BFB823E8A6DA48FC2" box="[1025,1189,908,930]" pageId="7" pageNumber="84" value="2012-04-04">
|
||
4 12
|
||
<emphasis id="B944EAF8FFCAFF9BFB083E8A6DA48FC2" bold="true" box="[1163,1189,908,929]" italics="true" pageId="7" pageNumber="84">IV</emphasis>
|
||
</date>
|
||
<date id="FF8E102AFFCAFF9BFB563E8A6C398FC2" box="[1237,1336,908,930]" pageId="7" pageNumber="84">
|
||
5
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B3E8A6DF38FC2" bold="true" box="[1256,1266,908,929]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFA883E8A6C148FC2" bold="true" box="[1291,1301,908,929]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD3E8A6C398FC2" bold="true" box="[1326,1336,908,929]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C3EB06C398FA9" blockId="7.[159,1407,298,1476]" box="[159,1336,949,971]" pageId="7" pageNumber="84">
|
||
<typeStatus id="548B8848FFCAFF9BFF1C3EB069F98FA8" box="[159,248,950,971]" pageId="7" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
7 1+1b - 2+1b 4+5 1+1b 7+
|
||
<date id="FF8E102AFFCAFF9BFB823EB06D9F8FA9" box="[1025,1182,949,971]" pageId="7" pageNumber="84" value="2011-02-07">
|
||
7 11
|
||
<emphasis id="B944EAF8FFCAFF9BFB093EB36D9F8FA9" bold="true" box="[1162,1182,949,970]" italics="true" pageId="7" pageNumber="84">II</emphasis>
|
||
</date>
|
||
<date id="FF8E102AFFCAFF9BFB563EB36C398FA9" box="[1237,1336,949,971]" pageId="7" pageNumber="84">
|
||
6
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B3EB36DF38FA9" bold="true" box="[1256,1266,949,970]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
4
|
||
<emphasis id="B944EAF8FFCAFF9BFA883EB36C148FA9" bold="true" box="[1291,1301,949,970]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD3EB36C398FA9" bold="true" box="[1326,1336,949,970]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFECD3ED86C5D8F97" blockId="7.[159,1407,298,1476]" box="[334,1372,990,1012]" pageId="7" pageNumber="84">
|
||
2+1b 4+3 1+1b 7+3 2+1b 3+
|
||
<date id="FF8E102AFFCAFF9BFB833ED86D9E8F97" box="[1024,1183,990,1012]" pageId="7" pageNumber="84" value="2012-02-04">
|
||
4 12
|
||
<emphasis id="B944EAF8FFCAFF9BFB083ED96D9E8F97" bold="true" box="[1163,1183,991,1012]" italics="true" pageId="7" pageNumber="84">II</emphasis>
|
||
</date>
|
||
<date id="FF8E102AFFCAFF9BFB563ED86C5D8F97" box="[1237,1372,990,1012]" pageId="7" pageNumber="84">
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B3ED96DF38F97" bold="true" box="[1256,1266,991,1012]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
6
|
||
<emphasis id="B944EAF8FFCAFF9BFA883ED96C148F97" bold="true" box="[1291,1301,991,1012]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD3ED96C398F97" bold="true" box="[1326,1336,991,1012]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFAD13ED96C5D8F97" bold="true" box="[1362,1372,991,1012]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C390F6C39887E" blockId="7.[159,1407,298,1476]" box="[159,1336,1032,1054]" pageId="7" pageNumber="84">
|
||
<typeStatus id="548B8848FFCAFF9BFF1C390F69F9887D" box="[159,248,1033,1054]" pageId="7" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
8 2+1b 3+2 - - 1+1b 6+? 12
|
||
<emphasis id="B944EAF8FFCAFF9BFB08390E6D9E887E" bold="true" box="[1163,1183,1032,1053]" italics="true" pageId="7" pageNumber="84">II</emphasis>
|
||
<date id="FF8E102AFFCAFF9BFB56390E6C39887E" box="[1237,1336,1032,1054]" pageId="7" pageNumber="84">
|
||
9
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B390E6DF3887E" bold="true" box="[1256,1266,1032,1053]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFA88390E6C14887E" bold="true" box="[1291,1301,1032,1053]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD390E6C39887E" bold="true" box="[1326,1336,1032,1053]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFECD39376C5D8824" blockId="7.[159,1407,298,1476]" box="[334,1372,1073,1095]" pageId="7" pageNumber="84">
|
||
2+1b 3+2 - - - -
|
||
<date id="FF8E102AFFCAFF9BFBE839376C5D8824" box="[1131,1372,1073,1095]" pageId="7" pageNumber="84">
|
||
9
|
||
<emphasis id="B944EAF8FFCAFF9BFBFD39346D938824" bold="true" box="[1150,1170,1074,1095]" italics="true" pageId="7" pageNumber="84">II</emphasis>
|
||
5
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B39346DF38824" bold="true" box="[1256,1266,1074,1095]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFA8839346C148824" bold="true" box="[1291,1301,1074,1095]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD39346C398824" bold="true" box="[1326,1336,1074,1095]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFAD139346C5D8824" bold="true" box="[1362,1372,1074,1095]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C395D6C3988FA" blockId="7.[159,1407,298,1476]" pageId="7" pageNumber="84">
|
||
<typeStatus id="548B8848FFCAFF9BFF1C395D69F98813" box="[159,248,1115,1136]" pageId="7" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
9 1+1b 3+4 1+1b 4+4? 1+1b 6+
|
||
<date id="FF8E102AFFCAFF9BFB81395C6D9F8813" box="[1026,1182,1114,1136]" pageId="7" pageNumber="84" value="2011-02-05">
|
||
5 11
|
||
<emphasis id="B944EAF8FFCAFF9BFB09395D6D9F8813" bold="true" box="[1162,1182,1115,1136]" italics="true" pageId="7" pageNumber="84">II</emphasis>
|
||
</date>
|
||
<date id="FF8E102AFFCAFF9BFB56395C6C398813" box="[1237,1336,1114,1136]" pageId="7" pageNumber="84">
|
||
8
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B395D6DF38813" bold="true" box="[1256,1266,1115,1136]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
4
|
||
<emphasis id="B944EAF8FFCAFF9BFA88395D6C148813" bold="true" box="[1291,1301,1115,1136]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD395D6C398813" bold="true" box="[1326,1336,1115,1136]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
1+1b 4+? 1+1b 3+? 1+1b 6+
|
||
<date id="FF8E102AFFCAFF9BFB8239826D9E88FA" box="[1025,1183,1156,1178]" pageId="7" pageNumber="84" value="2015-02-04">
|
||
4 15
|
||
<emphasis id="B944EAF8FFCAFF9BFB0839826D9E88FA" bold="true" box="[1163,1183,1156,1177]" italics="true" pageId="7" pageNumber="84">II</emphasis>
|
||
</date>
|
||
<date id="FF8E102AFFCAFF9BFB5639826C3988FA" box="[1237,1336,1156,1178]" pageId="7" pageNumber="84">
|
||
8
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B39826DF388FA" bold="true" box="[1256,1266,1156,1177]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFA8839826C1488FA" bold="true" box="[1291,1301,1156,1177]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
4
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD39826C3988FA" bold="true" box="[1326,1336,1156,1177]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C39A86DDC88A0" blockId="7.[159,1407,298,1476]" box="[159,1245,1197,1219]" pageId="7" pageNumber="84">
|
||
<typeStatus id="548B8848FFCAFF9BFF1C39A869F988A0" box="[159,248,1198,1219]" pageId="7" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
10 3+1b 4+3 4+1b 4+4 3+1b 6+0 - -
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFECD39D06DDC888F" blockId="7.[159,1407,298,1476]" box="[334,1245,1238,1260]" pageId="7" pageNumber="84">3+1b 2+2 4+1b 2+2 3+1b - - -</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C38076C5D8976" blockId="7.[159,1407,298,1476]" box="[159,1372,1280,1302]" pageId="7" pageNumber="84">
|
||
<typeStatus id="548B8848FFCAFF9BFF1C380769F98975" box="[159,248,1281,1302]" pageId="7" pageNumber="84" type="paratype">Paratype</typeStatus>
|
||
11 3+1b 2+0 2+1b 4+? - - 17
|
||
<emphasis id="B944EAF8FFCAFF9BFB0838066D9E8976" bold="true" box="[1163,1183,1280,1301]" italics="true" pageId="7" pageNumber="84">II</emphasis>
|
||
<date id="FF8E102AFFCAFF9BFB5638066C5D8976" box="[1237,1372,1280,1302]" pageId="7" pageNumber="84">
|
||
2
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B38066DF38976" bold="true" box="[1256,1266,1280,1301]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
5
|
||
<emphasis id="B944EAF8FFCAFF9BFA8838066C148976" bold="true" box="[1291,1301,1280,1301]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD38066C398976" bold="true" box="[1326,1336,1280,1301]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFAD138066C5D8976" bold="true" box="[1362,1372,1280,1301]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFECD382F6C39895C" blockId="7.[159,1407,298,1476]" box="[334,1336,1321,1343]" pageId="7" pageNumber="84">
|
||
3+1b 5+4 2+1b 2+1 - - 17
|
||
<emphasis id="B944EAF8FFCAFF9BFB08382C6D9E895C" bold="true" box="[1163,1183,1322,1343]" italics="true" pageId="7" pageNumber="84">II</emphasis>
|
||
<date id="FF8E102AFFCAFF9BFB56382F6C39895C" box="[1237,1336,1321,1343]" pageId="7" pageNumber="84">
|
||
6
|
||
<emphasis id="B944EAF8FFCAFF9BFB6B382C6DF3895C" bold="true" box="[1256,1266,1322,1343]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
3
|
||
<emphasis id="B944EAF8FFCAFF9BFA88382C6C14895C" bold="true" box="[1291,1301,1322,1343]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
4
|
||
<emphasis id="B944EAF8FFCAFF9BFAAD382C6C39895C" bold="true" box="[1326,1336,1322,1343]" italics="true" pageId="7" pageNumber="84">I</emphasis>
|
||
</date>
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF1C38556D9F89C5" blockId="7.[159,1407,298,1476]" pageId="7" pageNumber="84">
|
||
Range 1–3+ 1b 2–5+0–6 1–4+1b 1–8+1–5 1–3+1b 1–7+0–
|
||
<date id="FF8E102AFFCAFF9BFBB738546D41890B" box="[1076,1088,1362,1384]" pageId="7" pageNumber="84" value="1923-07-07">7</date>
|
||
7–23 9–15 small small 1– 2–5 big spines 4 big
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFBE838A86DAB89A7" blockId="7.[159,1407,298,1476]" box="[1131,1194,1454,1476]" pageId="7" pageNumber="84">spines</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF443B0F69EE8AB6" blockId="7.[151,1437,1545,2038]" pageId="7" pageNumber="84">
|
||
<emphasis id="B944EAF8FFCAFF9BFF443B0F68B98A41" bold="true" box="[199,440,1545,1570]" pageId="7" pageNumber="84">Variable characters.</emphasis>
|
||
The number of inner teeth on claw is variable. In the
|
||
<specimenCount id="9D36FD63FFCAFF9BFB8E3B0C6DA58A41" box="[1037,1188,1545,1570]" pageId="7" pageNumber="84" type="generic">12 specimens</specimenCount>
|
||
examined the number of inner claw teeth vary from 2 to 5; the number of teeth on inner two lamellae of the Emp. vary from 0 to 8. The number of dental spines is also variable. The number of small basal dental spines ranges from 7 to 23. In
|
||
<typeStatus id="548B8848FFCAFF9BFABF3B546C9D8A09" box="[1340,1436,1618,1642]" pageId="7" pageNumber="84" type="paratype">paratype</typeStatus>
|
||
1 the dental spine formula on one dens is 23 II /
|
||
<date id="FF8E102AFFCAFF9BFD3B3B706A4D8AEE" box="[696,844,1654,1678]" pageId="7" pageNumber="84">4 I 7 I 2 I 1 I</date>
|
||
0 I, whereas on the other it is
|
||
<date id="FF8E102AFFCAFF9BFB223B706DCD8AEE" box="[1185,1228,1654,1678]" pageId="7" pageNumber="84">9 II</date>
|
||
/
|
||
<date id="FF8E102AFFCAFF9BFB613B706C7C8AEE" box="[1250,1405,1653,1678]" pageId="7" pageNumber="84">3 I 5 I 2 I 2 I.</date>
|
||
In most specimens the mucro lacks intermediate denticles, but in some specimens 1 or 2 intermediate denticles are present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A6561FFCAFF9BFF443BE76D348BE9" pageId="7" pageNumber="84" type="discussion">
|
||
<paragraph id="8B8F36EAFFCAFF9BFF443BE76D348BE9" blockId="7.[151,1437,1545,2038]" pageId="7" pageNumber="84">
|
||
<emphasis id="B944EAF8FFCAFF9BFF443BE768448A99" bold="true" box="[199,325,1761,1786]" pageId="7" pageNumber="84">Discussion</emphasis>
|
||
. The new species is most similar to the North American
|
||
<taxonomicName id="4C304D69FFCAFF9BFC4A3BE46C308A99" authority="Christiansen, 1980" authorityName="Christiansen" authorityYear="1980" box="[969,1329,1761,1786]" class="Entognatha" family="Tomoceridae" genus="Plutomurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="B944EAF8FFCAFF9BFC4A3BE46D458A99" box="[969,1092,1761,1786]" italics="true" pageId="7" pageNumber="84">P. grahami</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA14B1BFFCAFF9BFBD03BE76C288A99" author="Christiansen" box="[1107,1321,1761,1786]" pageId="7" pageNumber="84" refString="Christiansen, K. (1980) A new nearctic species of the genus Tomocerus (Collembola: Entomobryidae). Proceedings of the Iowa Academy of Sciences, 87, 121 - 123." type="journal article" year="1980">Christiansen, 1980</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
based on the following characters: apparently blind under light microscopy, presence of 3 + 3 prelabral chaetae, and presence of clavate tenent hair. Both are cave-dwelling species. The new species differs from
|
||
<taxonomicName id="4C304D69FFCAFF9BFB2B3A2C6C238B21" box="[1192,1314,1833,1858]" class="Entognatha" family="Tomoceridae" genus="Plutomurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="grahami">
|
||
<emphasis id="B944EAF8FFCAFF9BFB2B3A2C6C238B21" box="[1192,1314,1833,1858]" italics="true" pageId="7" pageNumber="84">P. grahami</emphasis>
|
||
</taxonomicName>
|
||
by: (1) the formula of spine-like chaetae on the tibiotarsi: 0 0
|
||
<quantity id="4CC89B0FFFCAFF9BFD4F3A486BF98B05" box="[716,760,1869,1894]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="84" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName id="4C304D69FFCAFF9BFC833A486A608B06" box="[768,865,1870,1893]" class="Entognatha" family="Tomoceridae" genus="Plutomurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="revazi" status="sp. nov.">
|
||
<emphasis id="B944EAF8FFCAFF9BFC833A486A608B06" box="[768,865,1870,1893]" italics="true" pageId="7" pageNumber="84">P. revazi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2775783FFCAFF9BFCE93A4B6AC48B05" box="[874,965,1869,1894]" pageId="7" pageNumber="84" rank="species">
|
||
<emphasis id="B944EAF8FFCAFF9BFCE93A4B6AC48B05" bold="true" box="[874,965,1869,1894]" pageId="7" pageNumber="84">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, while it is 0 0
|
||
<quantity id="4CC89B0FFFCAFF9BFBEC3A486D998B05" box="[1135,1176,1869,1894]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="84" unit="in" value="1.0">1 in</quantity>
|
||
<emphasis id="B944EAF8FFCAFF9BFB233A486C278B05" box="[1184,1318,1869,1894]" italics="true" pageId="7" pageNumber="84">
|
||
<taxonomicName id="4C304D69FFCAFF9BFB233A486C1F8B05" box="[1184,1310,1869,1894]" class="Entognatha" family="Tomoceridae" genus="Plutomurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="grahami">P. grahami</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
(2) dorsal macrochaetotaxy: 52/
|
||
<quantity id="4CC89B0FFFCAFF9BFE0A3A7768ED8BE9" box="[393,492,1905,1930]" metricMagnitude="2" metricUnit="m" metricValue="8.491982" pageId="7" pageNumber="84" unit="in" value="33433.0">33433 in</quantity>
|
||
the new species, while it is 42/
|
||
<quantity id="4CC89B0FFFCAFF9BFCC93A746AAC8BE9" box="[842,941,1905,1930]" metricMagnitude="2" metricUnit="m" metricValue="8.232902" pageId="7" pageNumber="84" unit="in" value="32413.0">32413 in</quantity>
|
||
<emphasis id="B944EAF8FFCAFF9BFC303A746D348BE9" box="[947,1077,1905,1930]" italics="true" pageId="7" pageNumber="84">
|
||
<taxonomicName id="4C304D69FFCAFF9BFC303A746D308BE9" box="[947,1073,1905,1930]" class="Entognatha" family="Tomoceridae" genus="Plutomurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="84" phylum="Arthropoda" rank="species" species="grahami">P. grahami</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A6561FFCAFF9BFF443A936DF08B95" pageId="7" pageNumber="84" type="etymology">
|
||
<paragraph id="8B8F36EAFFCAFF9BFF443A936BFA8BB1" blockId="7.[151,1437,1545,2038]" pageId="7" pageNumber="84">
|
||
<emphasis id="B944EAF8FFCAFF9BFF443A93684F8BCD" bold="true" box="[199,334,1941,1966]" pageId="7" pageNumber="84">Etymology.</emphasis>
|
||
The species is dedicated to Dr. Revaz Djanashvili, who, beginning in the 1960s, was the first Georgian biospeleologist and collembolan taxonomist.
|
||
</paragraph>
|
||
<paragraph id="8B8F36EAFFCAFF9BFF443ADB6DF08B95" blockId="7.[151,1437,1545,2038]" box="[199,1265,2013,2038]" pageId="7" pageNumber="84">
|
||
<emphasis id="B944EAF8FFCAFF9BFF443ADB682A8B95" bold="true" box="[199,299,2013,2038]" pageId="7" pageNumber="84">Ecology.</emphasis>
|
||
It is probably troglophilous species, because the body and eye patches are pigmented.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |