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<document id="BFECEE43E54ACF5265E63ED3179A4801" ID-CLB-Dataset="24141" ID-DOI="10.11646/zootaxa.4731.3.4" ID-GBIF-Dataset="a9228bfa-3781-4c60-b06c-af3e12a20234" ID-ISSN="1175-5326" ID-Zenodo-Dep="3653608" ID-ZooBank="61966E54-BC89-4301-A8FA-671FA803597E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581059455841" checkinUser="plazi" docAuthor="Scioli, Justin A. &amp; Anker, Arthur" docDate="2020" docId="039987FC96744D7BFF4D99FEFB0DAE61" docLanguage="en" docName="zootaxa.4731.3.4.pdf" docOrigin="Zootaxa 4731 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alpheus gallicus Scioli &amp; Anker 2020, n. sp." docType="treatment" docVersion="8" lastPageNumber="355" masterDocId="FFA0FF8496764D73FFDA9B4BFFD4AB52" masterDocTitle="Description of Alpheus gallicus, a new deep-water snapping shrimp from Galicia Bank, northeastern Atlantic (Malacostraca, Decapoda, Alpheidae)" masterLastPageNumber="358" masterPageNumber="347" pageNumber="349" updateTime="1698780902693" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DB3AD7B52255442B41E5AFF1F4A2ED1B">Description of Alpheus gallicus, a new deep-water snapping shrimp from Galicia Bank, northeastern Atlantic (Malacostraca, Decapoda, Alpheidae)</mods:title>
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<mods:namePart id="98C7B111BF2A66E3BF30F950AD831A51">Scioli, Justin A.</mods:namePart>
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<mods:namePart id="E158456DD73FA9F4122277D54A0E9D8E">Anker, Arthur</mods:namePart>
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<paragraph id="8B8F36EA96744D71FF4D99FEFE7EA99D" blockId="2.[151,426,693,755]" box="[151,426,693,720]" pageId="2" pageNumber="349">
<heading id="D0C7818696744D71FF4D99FEFE7EA99D" bold="true" box="[151,426,693,720]" fontSize="11" level="1" pageId="2" pageNumber="349" reason="1">
<emphasis id="B944EAF896744D71FF4D99FEFE7EA99D" bold="true" box="[151,426,693,720]" pageId="2" pageNumber="349">
<taxonomicName id="4C304D6996744D71FF4D99FEFEB5A982" ID-CoL="86B5H" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[151,353,693,720]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="349" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896744D71FF4D99FEFEB5A982" bold="true" box="[151,353,693,720]" italics="true" pageId="2" pageNumber="349">Alpheus gallicus</emphasis>
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<taxonomicNameLabel id="A277578396744D71FEB299FEFE7EA99D" box="[360,426,693,719]" pageId="2" pageNumber="349" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8B8F36EA96744D71FF4D9992FEC6A9A1" blockId="2.[151,426,693,755]" box="[151,274,729,755]" pageId="2" pageNumber="349">
(
<figureCitation id="130B2A6F96744D71FF459992FEDEA9A1" box="[159,266,729,755]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1909,1934]" captionStartId-1="4.[151,250,1614,1639]" captionStartId-2="5.[151,250,1899,1924]" captionStartId-3="6.[151,250,1182,1207]" captionTargetBox-0="[187,1400,361,1856]" captionTargetBox-1="[187,1390,209,1543]" captionTargetBox-2="[210,1376,398,1801]" captionTargetBox-3="[151,1436,181,1156]" captionTargetId-0="figure@3.[160,1427,342,1881]" captionTargetId-1="figure@4.[164,1423,181,1589]" captionTargetId-2="figure@5.[164,1423,345,1874]" captionTargetId-3="figure@6.[151,1436,181,1156]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Alpheus gallicus n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, frontal region, dorsal; b, same, lateral; c, telson, dorsal; d, ventromesial carina of first article of antennular peduncle; e, third maxilliped, lateral; f, second pleopod, detail of appendix masculina and appendix interna, anterior (lateral); g, uropod, dorsal." captionText-1="FIGURE 2. Alpheus gallicus, n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, major cheliped, lateral; b, same, mesial; c, same, fingers; opened, lateral; d, same, dactylus, opened, lateral; e, minor cheliped, lateral; f, same, mesial." captionText-2="FIGURE 3. Alpheus gallicus, n. sp., paratype male (CL 9.6 mm), ICM 002619 (ad), and holotype male (CL 11.5 mm) (eg), OUMNH.ZC.2017-01-0061, both from Galicia Bank, NE Atlantic Ocean off western Spain: a, second pereiopod, lateral; b, third pereiopod, lateral; c, same, propodus and dactylus, lateral; d, same, detail of dactylus, lateral; e, fifth pereiopod, lateral; f, same, propodus and dactylus, lateral; g, same, dactylus, lateral." captionText-3="FIGURE 4. Alpheus gallicus n. sp., unspecified specimen, from Galicia Bank, NE Atlantic Ocean off western Spain: colouration of living individual, lateral. Scale below in cm. Photograph courtesy of Dr. Antonio Puzón." figureDoi-0="http://doi.org/10.5281/zenodo.3653610" figureDoi-1="http://doi.org/10.5281/zenodo.3653612" figureDoi-2="http://doi.org/10.5281/zenodo.3653614" figureDoi-3="http://doi.org/10.5281/zenodo.3653618" httpUri-0="https://zenodo.org/record/3653610/files/figure.png" httpUri-1="https://zenodo.org/record/3653612/files/figure.png" httpUri-2="https://zenodo.org/record/3653614/files/figure.png" httpUri-3="https://zenodo.org/record/3653618/files/figure.png" pageId="2" pageNumber="349">Figs. 14</figureCitation>
)
</paragraph>
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<paragraph id="8B8F36EA96744D71FF4D9856FB10A864" blockId="2.[151,1349,797,852]" box="[151,1220,797,822]" pageId="2" pageNumber="349">
<treatmentCitationGroup id="AB2011C496744D71FF4D9856FB10A864" box="[151,1220,797,822]" pageId="2" pageNumber="349">
<treatmentCitation id="0A9110FB96744D71FF4D9856FD5AA864" author="Cartes, J. E. &amp; Papiol, V. &amp; Frutos, I. &amp; Macpherson, E. &amp; Gonzalez-Pola, C. &amp; Punzon, A. &amp; Valeiras, X. &amp; Serrano, A." box="[151,654,797,822]" page="168" pageId="2" pageNumber="349" year="2014">
<taxonomicName id="4C304D6996744D71FF4D9856FD5AA864" ID-CoL="84W8H" authority="- Cartes et al. 2014: 168" authorityName="Cartes" authorityPageNumber="168" authorityYear="2014" box="[151,654,797,822]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="349" phylum="Arthropoda" rank="species" species="macrocheles">
<emphasis id="B944EAF896744D71FF4D9856FF3DA867" box="[151,233,797,821]" italics="true" pageId="2" pageNumber="349">Alpheus</emphasis>
cf.
<emphasis id="B944EAF896744D71FED59856FE5AA867" box="[271,398,797,821]" italics="true" pageId="2" pageNumber="349">macrocheles</emphasis>
<bibRefCitation id="EFA14B1B96744D71FE729856FD5AA864" author="Cartes, J. E. &amp; Papiol, V. &amp; Frutos, I. &amp; Macpherson, E. &amp; Gonzalez-Pola, C. &amp; Punzon, A. &amp; Valeiras, X. &amp; Serrano, A." box="[424,654,797,822]" pageId="2" pageNumber="349" pagination="165 - 178" refId="ref6562" refString="Cartes, J. E., Papiol, V., Frutos, I., Macpherson, E., Gonzalez-Pola, C., Punzon, A., Valeiras, X. &amp; Serrano, A. (2014) Distribution and biogeographic trends of decapod assemblages from Galicia Bank (NE Atlantic) at depths between 700 and 1800 m, with connexions to regional water masses. Deep Sea Research II, 106, 165 - 178. https: // doi. org / 10.1016 / j. dsr 2.2013.09.034" type="journal article" year="2014">
Cartes
<emphasis id="B944EAF896744D71FE359855FDF6A867" box="[495,546,797,821]" italics="true" pageId="2" pageNumber="349">et al.</emphasis>
2014: 168
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<tableCitation id="C6B2035196744D71FD419856FD32A867" box="[667,742,797,822]" captionStart="TABLE 1" captionStartId="1.[151,244,1232,1259]" captionTargetBox="[163,1415,1382,1769]" captionText="TABLE 1. Kimura 2-Parameter (= K2P) distances of the COI gene between Alpheus gallicus n. sp. and morphologically similar (putatively related) species of the A. macrocheles species group. COI sequences of three species (A. lentiginosus Anker &amp; Nizinski, 2011, A. platydactylus Coutière 1898 and A. romensky Burukovsky, 1990) are presently unavailable. See text for further details" httpUri="http://table.plazi.org/id/DF4F666296774D72FF4D9F9BFD68AE05" pageId="2" pageNumber="349" tableUuid="DF4F666296774D72FF4D9F9BFD68AE05">Table 1</tableCitation>
) [not
<taxonomicName id="4C304D6996744D71FCFC9856FB6DA864" ID-CoL="84W8H" authority="(Hailstone, 1835)" baseAuthorityName="Hailstone" baseAuthorityYear="1835" box="[806,1209,797,822]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="349" phylum="Arthropoda" rank="species" species="macrocheles">
<emphasis id="B944EAF896744D71FCFC9856FC29A867" box="[806,1021,797,821]" italics="true" pageId="2" pageNumber="349">Alpheus macrocheles</emphasis>
(
<bibRefCitation id="EFA14B1B96744D71FBD09856FB65A864" author="Hailstone, S." box="[1034,1201,797,822]" pageId="2" pageNumber="349" pagination="394 - 395" refId="ref7628" refString="Hailstone, S. (1835) The species of crustaceous animals discovered and described by Mr. Hailstone, and illustrated and annotated upon by Mr. Westwood. The Magazine of Natural History and Journal of Zoology, Botany, Mineralogy, Geology, and Meteorology, 8, 394 - 395." type="journal article" year="1835">Hailstone, 1835</bibRefCitation>
)
</taxonomicName>
].
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</paragraph>
<paragraph id="8B8F36EA96744D71FF4D9877FA91A806" blockId="2.[151,1349,797,852]" box="[151,1349,827,852]" pageId="2" pageNumber="349">
(?)
<emphasis id="B944EAF896744D71FF609877FE7CA806" box="[186,424,828,852]" italics="true" pageId="2" pageNumber="349">
<taxonomicName id="4C304D6996744D71FF609877FED8A806" ID-CoL="84LB5" authorityName="Fabricius" authorityYear="1798" box="[186,268,828,852]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="349" phylum="Arthropoda" rank="genus">Alpheus</taxonomicName>
macrocheles—
</emphasis>
<bibRefCitation id="EFA14B1B96744D71FE7D9877FD44A806" author="Garcia Raso, J. E." box="[423,656,827,852]" pageId="2" pageNumber="349" pagination="730 - 752" refId="ref7386" refString="Garcia Raso, J. E. (1996) Crustacea Decapoda (excl. Sergestidae) from Ibero-Moroccan waters. Results of Balgim- 84 expedition. Bulletin of Marine Science, 58, 730 - 752." type="journal article" year="1996">García Raso 1996: 735</bibRefCitation>
[not
<taxonomicName id="4C304D6996744D71FD1C9877FB83A806" ID-CoL="84W8H" authority="(Hailstone, 1835)" baseAuthorityName="Hailstone" baseAuthorityYear="1835" box="[710,1111,827,852]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="349" phylum="Arthropoda" rank="species" species="macrocheles">
<emphasis id="B944EAF896744D71FD1C9877FC48A806" box="[710,924,828,852]" italics="true" pageId="2" pageNumber="349">Alpheus macrocheles</emphasis>
(
<bibRefCitation id="EFA14B1B96744D71FC709870FB9BA806" author="Hailstone, S." box="[938,1103,827,852]" pageId="2" pageNumber="349" pagination="394 - 395" refId="ref7628" refString="Hailstone, S. (1835) The species of crustaceous animals discovered and described by Mr. Hailstone, and illustrated and annotated upon by Mr. Westwood. The Magazine of Natural History and Journal of Zoology, Botany, Mineralogy, Geology, and Meteorology, 8, 394 - 395." type="journal article" year="1835">Hailstone, 1835</bibRefCitation>
)
</taxonomicName>
, see discussion below].
</paragraph>
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<emphasis id="B944EAF896744D71FF4D98C9FEEBA8CE" bold="true" box="[151,319,897,924]" pageId="2" pageNumber="349">Type material</emphasis>
.
<materialsCitation id="3B583CB796744D71FE9498C9FA4CA892" ID-GBIF-Occurrence="2564241280" collectingDate="2011-08-09" collectionCode="OUMNH, ZC, INDE-" collectorName="J. E. Cartes" country="Spain" elevation="768" latitude="42.736816" location="Bank" longLatPrecision="1" longitude="-11.772533" pageId="2" pageNumber="349" specimenCode="MARES 0711, V11" specimenCount="1" stateProvince="Galicia" typeStatus="holotype">
<typeStatus id="548B884896744D71FE9498C9FE6FA8CE" box="[334,443,898,924]" pageId="2" pageNumber="349" type="holotype">Holotype</typeStatus>
: male (cl
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),
<collectionCode id="ED21AE2F96744D71FD7F98C9FCC3A8CE" box="[677,791,898,924]" pageId="2" pageNumber="349">OUMNH</collectionCode>
.
<collectionCode id="ED21AE2F96744D71FCC198C9FC95A8CE" box="[795,833,898,924]" country="Austria" name="Zoological Collection, University of Vienna" pageId="2" pageNumber="349" type="University or college">ZC</collectionCode>
.2017-01-061,
<collectingCountry id="F327767A96744D71FC3A98C9FBF7A8CE" box="[992,1059,898,924]" name="Spain" pageId="2" pageNumber="349">Spain</collectingCountry>
,
<collectingRegion id="49F4F80896744D71FBF598C9FB54A8CE" box="[1071,1152,898,924]" country="Spain" name="Galicia" pageId="2" pageNumber="349">Galicia</collectingRegion>
<location id="8EEF603196744D71FB5298C9FB12A8CE" LSID="urn:lsid:plazi:treatment:039987FC96744D7BFF4D99FEFB0DAE61:8EEF603196744D71FB5298C9FB12A8CE" box="[1160,1222,898,924]" country="Spain" latitude="42.736816" longLatPrecision="1" longitude="-11.772533" name="Bank" pageId="2" pageNumber="349" stateProvince="Galicia">Bank</location>
, expedition
<collectionCode id="ED21AE2F96744D71FA8A98C8FA48A8CF" box="[1360,1436,899,925]" pageId="2" pageNumber="349">INDE-</collectionCode>
<specimenCode id="DB969E9196744D71FF4D98EDFEECA892" box="[151,312,934,960]" collectionCode="MARES" pageId="2" pageNumber="349">MARES 0711</specimenCode>
, sta.
<specimenCode id="DB969E9196744D71FEB798EDFE4AA892" box="[365,414,934,960]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="349" type="Museum">V11</specimenCode>
,
<geoCoordinate id="EE04502D96744D71FE7398EDFDECA892" box="[425,568,934,960]" degrees="42" direction="north" minutes="44.209" orientation="latitude" pageId="2" pageNumber="349" precision="1" value="42.736816">42º44.209N</geoCoordinate>
<geoCoordinate id="EE04502D96744D71FDE598EEFD07A892" box="[575,723,933,960]" degrees="11" direction="west" minutes="46.352" orientation="longitude" pageId="2" pageNumber="349" precision="1" value="-11.772533">11º46.352W</geoCoordinate>
, depth:
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<elevation id="001DD1D996744D71FCF098EDFCA6A893" box="[810,882,934,961]" metricMagnitude="2" metricUnit="m" metricValue="7.68" pageId="2" pageNumber="349" unit="m" value="768.0">768 m</elevation>
</quantity>
, beam trawl, leg.
<collectorName id="26C5533C96744D71FBEE98ECFB78A892" box="[1076,1196,934,961]" pageId="2" pageNumber="349">J.E. Cartes</collectorName>
<emphasis id="B944EAF896744D71FB6B98ECFB36A892" box="[1201,1250,934,960]" italics="true" pageId="2" pageNumber="349">et al</emphasis>
.,
<date id="FF8E102A96744D71FB2F98EDFA4CA892" box="[1269,1432,934,960]" pageId="2" pageNumber="349" value="2011-08-09">
<collectingDate id="EFCAE9C296744D71FB2F98EDFA4CA892" box="[1269,1432,934,960]" pageId="2" pageNumber="349" value="2011-08-09">9 August 2011</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B583CB796744D71FF4D9880FAD0A8B6" ID-GBIF-Occurrence="2564241279" box="[151,1284,970,997]" collectingDate="2011-08-09" collectionCode="OUMNH, ZC" collectorName="J. E. Cartes" country="Spain" elevation="768" latitude="42.736816" location="Bank" longLatPrecision="1" longitude="-11.772533" pageId="2" pageNumber="349" specimenCount="1" specimenCount-female="1" stateProvince="Galicia" typeStatus="paratype">
<typeStatus id="548B884896744D71FF4D9880FEDCA8B7" box="[151,264,971,997]" pageId="2" pageNumber="349" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D36FD6396744D71FECF9881FEADA8B6" box="[277,377,970,996]" pageId="2" pageNumber="349" type="female">1 female</specimenCount>
(CL
<quantity id="4CC89B0F96744D71FE629881FDDAA8B7" box="[440,526,970,997]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="349" unit="mm" value="13.0">13 mm</quantity>
),
<collectionCode id="ED21AE2F96744D71FDF99881FD40A8B6" box="[547,660,970,996]" pageId="2" pageNumber="349">OUMNH</collectionCode>
.
<collectionCode id="ED21AE2F96744D71FD429881FD6AA8B6" box="[664,702,970,996]" country="Austria" name="Zoological Collection, University of Vienna" pageId="2" pageNumber="349" type="University or college">ZC</collectionCode>
.2017-01-062, same collection data as for holotype
</materialsCitation>
;
<materialsCitation id="3B583CB796744D71FACB9881FC26AF5A" ID-GBIF-Occurrence="2564241277" collectingDate="2011-08-09" collectionCode="OUMNH, ZC" collectorName="J. E. Cartes" country="Spain" elevation="768" latitude="42.736816" location="Bank" longLatPrecision="1" longitude="-11.772533" pageId="2" pageNumber="349" specimenCount="1" specimenCount-female="1" stateProvince="Galicia" typeStatus="paratype">
<specimenCount id="9D36FD6396744D71FACB9881FAA1A8B6" box="[1297,1397,970,996]" pageId="2" pageNumber="349" type="female">1 female</specimenCount>
(cl
<quantity id="4CC89B0F96744D71FF4D98A6FED8AF5B" box="[151,268,1005,1033]" metricMagnitude="-2" metricUnit="m" metricValue="1.125" pageId="2" pageNumber="349" unit="mm" value="11.25">11.25 mm</quantity>
),
<collectionCode id="ED21AE2F96744D71FEC598A5FE45AF5A" box="[287,401,1006,1032]" pageId="2" pageNumber="349">OUMNH</collectionCode>
.
<collectionCode id="ED21AE2F96744D71FE4F98A5FE6FAF5A" box="[405,443,1006,1032]" country="Austria" name="Zoological Collection, University of Vienna" pageId="2" pageNumber="349" type="University or college">ZC</collectionCode>
.2017-01-063, same collection data as for holotype
</materialsCitation>
;
<materialsCitation id="3B583CB796744D71FC2798A5FDEFAF7E" ID-GBIF-Occurrence="2564241278" collectingDate="2011-08-09" collectionCode="ICM" collectorName="J. E. Cartes" country="Spain" elevation="768" latitude="42.736816" location="Bank" longLatPrecision="1" longitude="-11.772533" pageId="2" pageNumber="349" specimenCode="ICM 002618" specimenCount="1" specimenCount-female="1" stateProvince="Galicia" typeStatus="paratype">
<specimenCount id="9D36FD6396744D71FC2798A5FB8BAF5A" box="[1021,1119,1006,1032]" pageId="2" pageNumber="349" type="female">1 female</specimenCount>
(cl
<quantity id="4CC89B0F96744D71FB5198A5FB26AF5B" box="[1163,1266,1006,1033]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="2" pageNumber="349" unit="mm" value="11.9">11.9 mm</quantity>
),
<specimenCode id="DB969E9196744D71FADF98A5FA4CAF5A" box="[1285,1432,1006,1032]" collectionCode="ICM" country="Spain" httpUri="http://grbio.org/cool/x81t-7j82" name="Instituto de Ciencias del Mar" pageId="2" pageNumber="349">ICM 002618</specimenCode>
, same collection data as for holotype
</materialsCitation>
;
<materialsCitation id="3B583CB796744D71FD929F59FE81AF26" ID-GBIF-Occurrence="2564241281" collectingDate="2011-07-30" collectionCode="INDE-" collectorName="J. E. Cartes" country="Spain" elevation="785" latitude="42.649532" location="Bank" longLatPrecision="1" longitude="-11.8182" pageId="2" pageNumber="349" specimenCode="ICM 002619, MARES 0711" specimenCount="1" specimenCount-male="1" stateProvince="Galicia" typeStatus="paratype">
<specimenCount id="9D36FD6396744D71FD929F59FD41AF7E" box="[584,661,1042,1068]" pageId="2" pageNumber="349" type="male">1 male</specimenCount>
(cl
<quantity id="4CC89B0F96744D71FD1C9F59FCF6AF7F" box="[710,802,1042,1069]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="2" pageNumber="349" unit="mm" value="9.6">9.6 mm</quantity>
),
<specimenCode id="DB969E9196744D71FCE29F59FC19AF7E" box="[824,973,1042,1068]" collectionCode="ICM" country="Spain" httpUri="http://grbio.org/cool/x81t-7j82" name="Instituto de Ciencias del Mar" pageId="2" pageNumber="349">ICM 002619</specimenCode>
,
<collectingCountry id="F327767A96744D71FC019F59FBCAAF7E" box="[987,1054,1042,1068]" name="Spain" pageId="2" pageNumber="349">Spain</collectingCountry>
,
<collectingRegion id="49F4F80896744D71FBF19F59FBA8AF7E" box="[1067,1148,1042,1068]" country="Spain" name="Galicia" pageId="2" pageNumber="349">Galicia</collectingRegion>
<location id="8EEF603196744D71FB5F9F59FB17AF7E" LSID="urn:lsid:plazi:treatment:039987FC96744D7BFF4D99FEFB0DAE61:8EEF603196744D71FB5F9F59FB17AF7E" box="[1157,1219,1042,1068]" country="Spain" latitude="42.649532" longLatPrecision="1" longitude="-11.8182" name="Bank" pageId="2" pageNumber="349" stateProvince="Galicia">Bank</location>
, expedition
<collectionCode id="ED21AE2F96744D71FA8A9F58FA48AF7F" box="[1360,1436,1043,1069]" pageId="2" pageNumber="349">INDE-</collectionCode>
<specimenCode id="DB969E9196744D71FF4D9F7DFEEDAF02" box="[151,313,1078,1104]" collectionCode="MARES" pageId="2" pageNumber="349">MARES 0711</specimenCode>
, sta. GOC3,
<geoCoordinate id="EE04502D96744D71FE119F7DFD8EAF02" box="[459,602,1078,1104]" degrees="42" direction="north" minutes="38.972" orientation="latitude" pageId="2" pageNumber="349" precision="1" value="42.649532">42º38.972N</geoCoordinate>
<geoCoordinate id="EE04502D96744D71FDBB9F7DFD21AF02" box="[609,757,1078,1104]" degrees="11" direction="west" minutes="49.092" orientation="longitude" pageId="2" pageNumber="349" precision="1" value="-11.8182">11º49.092W</geoCoordinate>
, depth:
<quantity id="4CC89B0F96744D71FC959F7EFC4DAF03" box="[847,921,1077,1105]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="2" pageNumber="349" unit="m" value="785.0">
<elevation id="001DD1D996744D71FC959F7EFC4DAF03" box="[847,921,1077,1105]" metricMagnitude="2" metricUnit="m" metricValue="7.85" pageId="2" pageNumber="349" unit="m" value="785.0">785 m</elevation>
</quantity>
, otter trawl / 2-doors trawl, leg.
<collectorName id="26C5533C96744D71FADD9F7CFA55AF02" box="[1287,1409,1078,1105]" pageId="2" pageNumber="349">J.E. Cartes</collectorName>
<emphasis id="B944EAF896744D71FA529F7CFF79AF26" italics="true" pageId="2" pageNumber="349">et al</emphasis>
.,
<date id="FF8E102A96744D71FF189F11FE81AF26" box="[194,341,1114,1140]" pageId="2" pageNumber="349" value="2011-07-30">
<collectingDate id="EFCAE9C296744D71FF189F11FE81AF26" box="[194,341,1114,1140]" pageId="2" pageNumber="349" value="2011-07-30">30 July 2011</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A656196744D71FF1D9F36FC2EADAE" pageId="2" pageNumber="349" type="diagnosis">
<paragraph id="8B8F36EA96744D71FF1D9F36FC2EADAE" blockId="2.[151,1437,897,2005]" pageId="2" pageNumber="349">
<emphasis id="B944EAF896744D71FF1D9F36FEEEAFCA" bold="true" box="[199,314,1149,1176]" pageId="2" pageNumber="349">Diagnosis</emphasis>
. Species of
<taxonomicName id="4C304D6996744D71FE169F35FD68AFCA" baseAuthorityName="Hailstone" baseAuthorityYear="1835" box="[460,700,1150,1176]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="349" phylum="Arthropoda" rank="species" species="macrocheles">
<emphasis id="B944EAF896744D71FE169F35FD68AFCA" box="[460,700,1150,1176]" italics="true" pageId="2" pageNumber="349">Alpheus macrocheles</emphasis>
</taxonomicName>
species group. Frontal margin of carapace with well-developed rostrum and orbital teeth; rostrum distally acute, about 1.3 times as long as wide at base, tip not reaching halflength of first article of antennular peduncle; orbital teeth narrow and acute; stylocerite of antennule not reaching distal margin of first article or antennular peduncle; second antennular article slender, four times as long as wide; scaphocerite with shallowly concave lateral margin, stout distolateral tooth, reaching far beyond distal margin of narrow blade; third maxilliped not especially modified; major cheliped much larger and stouter than minor cheliped; major cheliped ischium with small spiniform seta on ventromesial margin; major cheliped merus with several small spiniform setae on ventromesial margin and acute distomesial tooth; major chela with distinct inferior plaque and superior crests separated by superior and inferior grooves, as well as with superior and inferior transverse grooves; superior and inferior crests ending distally in acute, anteriorly directed teeth; distal portion of major cheliped pollex twisted laterally, cutting edge bearing small tooth and notch; major chela dactylus twisted laterally, dorsally convex; plunger reduced to small proximal protuberance; minor cheliped ischium bearing small stout spiniform seta on ventral margin; ventromesial margin of minor cheliped merus without spiniform setae, with acute distomesial tooth; minor cheliped dactylus compressed, twisted laterally, with high dorsal crest; length ratio of subdivisions of second pereiopod carpus approximately equal to 9: 5: 3: 3: 4; meri of third and fourth pereiopods without distoventral tooth, ventral margin without spiniform setae; dactyli of third to fifth pereiopods inconspicuously biunguiculate; male second pleopod with appendix masculina slightly overreaching appendix interna; uropod with distolateral spiniform seta flanked laterally by acute tooth, mesially by obtuse projection.
</paragraph>
</subSubSection>
<subSubSection id="C32A656196744D74FF1D9C4EFF26A869" lastPageId="7" lastPageNumber="354" pageId="2" pageNumber="349" type="description">
<paragraph id="8B8F36EA96744D71FF1D9C4EFC7FAC86" blockId="2.[151,1437,897,2005]" pageId="2" pageNumber="349">
<emphasis id="B944EAF896744D71FF1D9C4EFE85AC72" bold="true" box="[199,337,1797,1824]" pageId="2" pageNumber="349">Description</emphasis>
. Carapace glabrous, not setose; frontal margin with well-developed rostrum and orbital teeth; rostrum subtriangular, distally acute, about 1.3 times as long as wide at base, without setae, tip not reaching half-length of first article of antennular peduncle; rostral carina not distinct; orbital teeth narrow, distally acute, reaching halflength of rostrum, directed anteriorly and slightly mesially; frontal margin between rostrum and orbital teeth nearly straight at oblique angle; orbital hoods moderately swollen, completely enclosing eyes (
<figureCitation id="130B2A6F96744D71FBAA9CDDFB0AACE3" box="[1136,1246,1942,1969]" captionStart="FIGURE 1" captionStartId="3.[151,250,1909,1934]" captionTargetBox="[187,1400,361,1856]" captionTargetId="figure@3.[160,1427,342,1881]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus gallicus n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, frontal region, dorsal; b, same, lateral; c, telson, dorsal; d, ventromesial carina of first article of antennular peduncle; e, third maxilliped, lateral; f, second pleopod, detail of appendix masculina and appendix interna, anterior (lateral); g, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3653610" httpUri="https://zenodo.org/record/3653610/files/figure.png" pageId="2" pageNumber="349">Fig. 1a, b</figureCitation>
). Pterygostomial angle rounded, not protruding anteriorly (
<figureCitation id="130B2A6F96744D71FDB39CF1FD6EAC87" box="[617,698,1978,2005]" captionStart="FIGURE 1" captionStartId="3.[151,250,1909,1934]" captionTargetBox="[187,1400,361,1856]" captionTargetId="figure@3.[160,1427,342,1881]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus gallicus n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, frontal region, dorsal; b, same, lateral; c, telson, dorsal; d, ventromesial carina of first article of antennular peduncle; e, third maxilliped, lateral; f, second pleopod, detail of appendix masculina and appendix interna, anterior (lateral); g, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3653610" httpUri="https://zenodo.org/record/3653610/files/figure.png" pageId="2" pageNumber="349">Fig. 1b</figureCitation>
); cardiac notch deep.
</paragraph>
<paragraph id="8B8F36EA96754D70FF1D9BD2FC42AA11" blockId="3.[151,1437,153,323]" pageId="3" pageNumber="350">
Telson about twice as long as basal width; lateral margin slightly constricted posterior to half-length, tapering distally; dorsal surface smooth, bearing two pairs of stout cuspidate setae; anterior pair inserted slightly anterior to telson mid-length; posterior pair inserted at about 0.7 telson length; posterior margin slightly convex, distolateral angles each bearing one pair of spiniform setae, mesial about four times longer than lateral; margin between spiniform setae without spinules (
<figureCitation id="130B2A6F96754D70FE019A62FDFFAA11" box="[475,555,297,323]" captionStart="FIGURE 1" captionStartId="3.[151,250,1909,1934]" captionTargetBox="[187,1400,361,1856]" captionTargetId="figure@3.[160,1427,342,1881]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus gallicus n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, frontal region, dorsal; b, same, lateral; c, telson, dorsal; d, ventromesial carina of first article of antennular peduncle; e, third maxilliped, lateral; f, second pleopod, detail of appendix masculina and appendix interna, anterior (lateral); g, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3653610" httpUri="https://zenodo.org/record/3653610/files/figure.png" pageId="3" pageNumber="350">Fig. 1c</figureCitation>
); anal tubercles well developed.
</paragraph>
<caption id="DF4F666296754D70FF4D9C3EFE7CACA8" ID-DOI="http://doi.org/10.5281/zenodo.3653610" ID-Zenodo-Dep="3653610" httpUri="https://zenodo.org/record/3653610/files/figure.png" pageId="3" pageNumber="350" startId="3.[151,250,1909,1934]" targetBox="[187,1400,361,1856]" targetPageId="3">
<paragraph id="8B8F36EA96754D70FF4D9C3EFE7CACA8" blockId="3.[151,1437,1909,2042]" pageId="3" pageNumber="350">
<emphasis id="B944EAF896754D70FF4D9C3EFEC6ACDC" bold="true" box="[151,274,1909,1934]" pageId="3" pageNumber="350">FIGURE 1.</emphasis>
<taxonomicName id="4C304D6996754D70FEC29C3DFE6BACDC" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[280,447,1910,1934]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896754D70FEC29C3DFE6BACDC" box="[280,447,1910,1934]" italics="true" pageId="3" pageNumber="350">Alpheus gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF896754D70FE1E9C3DFE28ACDC" bold="true" box="[452,508,1910,1934]" pageId="3" pageNumber="350">
<taxonomicNameLabel id="A277578396754D70FE1E9C3DFE28ACDC" box="[452,508,1910,1934]" pageId="3" pageNumber="350" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, frontal region, dorsal; b, same, lateral; c, telson, dorsal; d, ventromesial carina of first article of antennular peduncle; e, third maxilliped, lateral; f, second pleopod, detail of appendix masculina and appendix interna, anterior (lateral); g, uropod, dorsal.
</paragraph>
</caption>
<caption id="DF4F666296724D77FF4D9D05FD37ADFD" ID-DOI="http://doi.org/10.5281/zenodo.3653612" ID-Zenodo-Dep="3653612" httpUri="https://zenodo.org/record/3653612/files/figure.png" pageId="4" pageNumber="351" startId="4.[151,250,1614,1639]" targetBox="[187,1390,209,1543]" targetPageId="4">
<paragraph id="8B8F36EA96724D77FF4D9D05FD37ADFD" blockId="4.[151,1436,1614,1711]" pageId="4" pageNumber="351">
<emphasis id="B944EAF896724D77FF4D9D05FEC1AD35" bold="true" box="[151,277,1614,1639]" pageId="4" pageNumber="351">FIGURE 2.</emphasis>
<emphasis id="B944EAF896724D77FEC79D04FE19AD35" box="[285,461,1615,1639]" italics="true" pageId="4" pageNumber="351">
<taxonomicName id="4C304D6996724D77FEC79D04FE1DAD35" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[285,457,1615,1639]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">Alpheus gallicus</taxonomicName>
,
</emphasis>
<emphasis id="B944EAF896724D77FE0F9D04FDC4AD35" bold="true" box="[469,528,1615,1639]" pageId="4" pageNumber="351">
<taxonomicNameLabel id="A277578396724D77FE0F9D04FDC4AD35" box="[469,528,1615,1639]" pageId="4" pageNumber="351" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, major cheliped, lateral; b, same, mesial; c, same, fingers; opened, lateral; d, same, dactylus, opened, lateral; e, minor cheliped, lateral; f, same, mesial.
</paragraph>
</caption>
<paragraph id="8B8F36EA96724D77FF1D9D96FEA0ACA6" blockId="4.[151,1437,1757,2036]" pageId="4" pageNumber="351">
Antennular peduncle slender; stylocerite well developed, with acute tip, not reaching distal margin of first article of antennular peduncle (
<figureCitation id="130B2A6F96724D77FE3E9C4AFD81AC4E" box="[484,597,1793,1820]" captionStart="FIGURE 1" captionStartId="3.[151,250,1909,1934]" captionTargetBox="[187,1400,361,1856]" captionTargetId="figure@3.[160,1427,342,1881]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus gallicus n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, frontal region, dorsal; b, same, lateral; c, telson, dorsal; d, ventromesial carina of first article of antennular peduncle; e, third maxilliped, lateral; f, second pleopod, detail of appendix masculina and appendix interna, anterior (lateral); g, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3653610" httpUri="https://zenodo.org/record/3653610/files/figure.png" pageId="4" pageNumber="351">Fig. 1a, b</figureCitation>
); ventromesial carina of first article subtriangular, with minute, anteriorly directed tooth (
<figureCitation id="130B2A6F96724D77FEE59C6EFE5BAC12" box="[319,399,1829,1856]" captionStart="FIGURE 1" captionStartId="3.[151,250,1909,1934]" captionTargetBox="[187,1400,361,1856]" captionTargetId="figure@3.[160,1427,342,1881]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus gallicus n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, frontal region, dorsal; b, same, lateral; c, telson, dorsal; d, ventromesial carina of first article of antennular peduncle; e, third maxilliped, lateral; f, second pleopod, detail of appendix masculina and appendix interna, anterior (lateral); g, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3653610" httpUri="https://zenodo.org/record/3653610/files/figure.png" pageId="4" pageNumber="351">Fig. 1d</figureCitation>
); second article about twice as long as visible portion of first, four times as long as wide; third article shortest, one-fourth length of second (
<figureCitation id="130B2A6F96724D77FD489C02FCD5AC36" box="[658,769,1865,1892]" captionStart="FIGURE 1" captionStartId="3.[151,250,1909,1934]" captionTargetBox="[187,1400,361,1856]" captionTargetId="figure@3.[160,1427,342,1881]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus gallicus n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, frontal region, dorsal; b, same, lateral; c, telson, dorsal; d, ventromesial carina of first article of antennular peduncle; e, third maxilliped, lateral; f, second pleopod, detail of appendix masculina and appendix interna, anterior (lateral); g, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3653610" httpUri="https://zenodo.org/record/3653610/files/figure.png" pageId="4" pageNumber="351">Fig. 1a, b</figureCitation>
); lateral flagellum with aesthetascs starting from about 15
<superScript id="7C459BA296724D77FA4A9C03FA49AC04" attach="right" box="[1424,1437,1864,1878]" fontSize="6" pageId="4" pageNumber="351">th</superScript>
subdivision. Antenna with basicerite bearing short, acute distoventral tooth; carpocerite slender, about eight times as long as broad, overreaching both antennular peduncle and scaphocerite; scaphocerite with shallowly concave lateral margin; distolateral tooth stout, reaching far beyond distal margin of blade; blade fairly narrow, gradually tapering distally (
<figureCitation id="130B2A6F96724D77FF239C92FEB3ACA6" box="[249,359,2009,2036]" captionStart="FIGURE 1" captionStartId="3.[151,250,1909,1934]" captionTargetBox="[187,1400,361,1856]" captionTargetId="figure@3.[160,1427,342,1881]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus gallicus n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, frontal region, dorsal; b, same, lateral; c, telson, dorsal; d, ventromesial carina of first article of antennular peduncle; e, third maxilliped, lateral; f, second pleopod, detail of appendix masculina and appendix interna, anterior (lateral); g, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3653610" httpUri="https://zenodo.org/record/3653610/files/figure.png" pageId="4" pageNumber="351">Fig. 1a, b</figureCitation>
).
</paragraph>
<paragraph id="8B8F36EA96734D76FF1D9BD2FB28AA11" blockId="5.[151,1436,153,323]" pageId="5" pageNumber="352">
Mouthparts typical for genus in external view. Third maxilliped slender, not especially modified; coxa with lateral plate not strongly produced, distal margin sparsely setose; exopod not overreaching penultimate article; antepenultimate article gradually flattening ventrally, ventral margin setose; penultimate article subcylindrical, slightly broadening distally, roughly 4.5 as long as maximal width; ultimate article long, slender, about 11 times as long as maximal width, unarmed distally, with transverse rows of serrulate setae on mesial surface (
<figureCitation id="130B2A6F96734D76FB469A62FB38AA11" box="[1180,1260,297,323]" captionStart="FIGURE 1" captionStartId="3.[151,250,1909,1934]" captionTargetBox="[187,1400,361,1856]" captionTargetId="figure@3.[160,1427,342,1881]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus gallicus n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, frontal region, dorsal; b, same, lateral; c, telson, dorsal; d, ventromesial carina of first article of antennular peduncle; e, third maxilliped, lateral; f, second pleopod, detail of appendix masculina and appendix interna, anterior (lateral); g, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3653610" httpUri="https://zenodo.org/record/3653610/files/figure.png" pageId="5" pageNumber="352">Fig. 1e</figureCitation>
).
</paragraph>
<caption id="DF4F666296734D76FF4D9C20FD02ACA2" ID-DOI="http://doi.org/10.5281/zenodo.3653614" ID-Zenodo-Dep="3653614" httpUri="https://zenodo.org/record/3653614/files/figure.png" pageId="5" pageNumber="352" startId="5.[151,250,1899,1924]" targetBox="[210,1376,398,1801]" targetPageId="5">
<paragraph id="8B8F36EA96734D76FF4D9C20FD02ACA2" blockId="5.[151,1436,1899,2032]" pageId="5" pageNumber="352">
<emphasis id="B944EAF896734D76FF4D9C20FEC7ACD6" bold="true" box="[151,275,1899,1924]" pageId="5" pageNumber="352">FIGURE 3.</emphasis>
<emphasis id="B944EAF896734D76FEC09C20FE1DACD1" box="[282,457,1899,1923]" italics="true" pageId="5" pageNumber="352">
<taxonomicName id="4C304D6996734D76FEC09C20FE11ACD1" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[282,453,1899,1923]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">Alpheus gallicus</taxonomicName>
,
</emphasis>
<emphasis id="B944EAF896734D76FE159C20FDDDACD1" bold="true" box="[463,521,1899,1923]" pageId="5" pageNumber="352">
<taxonomicNameLabel id="A277578396734D76FE159C20FDDDACD1" box="[463,521,1899,1923]" pageId="5" pageNumber="352" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, paratype male (CL 9.6 mm), ICM 002619 (ad), and holotype male (CL 11.5 mm) (eg), OUMNH.ZC.2017-01-0061, both from Galicia Bank, NE Atlantic Ocean off western Spain: a, second pereiopod, lateral; b, third pereiopod, lateral; c, same, propodus and dactylus, lateral; d, same, detail of dactylus, lateral; e, fifth pereiopod, lateral; f, same, propodus and dactylus, lateral; g, same, dactylus, lateral.
</paragraph>
</caption>
<caption id="DF4F666296704D75FF4D9FD5FBB4AF89" ID-DOI="http://doi.org/10.5281/zenodo.3653618" ID-Zenodo-Dep="3653618" httpUri="https://zenodo.org/record/3653618/files/figure.png" pageId="6" pageNumber="353" startId="6.[151,250,1182,1207]" targetBox="[151,1436,181,1156]" targetPageId="6">
<paragraph id="8B8F36EA96704D75FF4D9FD5FBB4AF89" blockId="6.[151,1436,1182,1243]" pageId="6" pageNumber="353">
<emphasis id="B944EAF896704D75FF4D9FD5FED9AFE5" bold="true" box="[151,269,1182,1207]" pageId="6" pageNumber="353">FIGURE 4</emphasis>
.
<taxonomicName id="4C304D6996704D75FEC09FD5FE16AFE4" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[282,450,1182,1206]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="353" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896704D75FEC09FD5FE16AFE4" box="[282,450,1182,1206]" italics="true" pageId="6" pageNumber="353">Alpheus gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF896704D75FE139FD4FDD6AFE5" bold="true" box="[457,514,1183,1207]" pageId="6" pageNumber="353">
<taxonomicNameLabel id="A277578396704D75FE139FD4FDD6AFE5" box="[457,514,1183,1207]" pageId="6" pageNumber="353" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, unspecified specimen, from Galicia Bank, NE Atlantic Ocean off western Spain: colouration of living individual, lateral. Scale below in cm. Photograph courtesy of Dr. Antonio Puzón.
</paragraph>
</caption>
<paragraph id="8B8F36EA96704D75FF1D9E42FCE1ADE2" blockId="6.[151,1437,1289,2035]" pageId="6" pageNumber="353">
Major cheliped noticeably larger and stouter and somewhat longer than minor cheliped; ischium short, bearing small spiniform seta on ventromesial margin; merus broad, relatively short, subtriangular in cross-section, about 2.5 times as long as maximal width; ventrolateral margin unarmed; ventromesial margin bearing seven or so small spiniform setae and ending in sharp distal tooth; distodorsal margin bluntly protruding; carpus short, cup-shaped; chela somewhat compressed, twisted laterally; plaque crest deviating from dorsal margin in proximal third of chela, continuing as elevation on lateral surface; superior crest lamellar, dorsally rounded, ending distally in acute tooth; inferior crest distinct, ending distally as sharp acute tooth; superior groove deep, proximally extending beyond superior transverse groove; inferior groove originating shallowly near linea impressa, gradually deepening distally; superior transverse groove deep, with overhanging proximal shoulder; inferior depression deep, sharply delimited proximally; pollex short, distal portion twisted laterally, cutting edge bearing small tooth followed by triangular notch; dactylus twisted laterally, dorsally convex, flattened; tip with deep notch bearing tufts of setae; plunger greatly reduced to small proximal protuberance (
<figureCitation id="130B2A6F96704D75FD609DDEFD28ADE2" box="[698,764,1685,1712]" captionStart="FIGURE 2" captionStartId="4.[151,250,1614,1639]" captionTargetBox="[187,1390,209,1543]" captionTargetId="figure@4.[164,1423,181,1589]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus gallicus, n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, major cheliped, lateral; b, same, mesial; c, same, fingers; opened, lateral; d, same, dactylus, opened, lateral; e, minor cheliped, lateral; f, same, mesial." figureDoi="http://doi.org/10.5281/zenodo.3653612" httpUri="https://zenodo.org/record/3653612/files/figure.png" pageId="6" pageNumber="353">Fig. 2</figureCitation>
ad).
</paragraph>
<paragraph id="8B8F36EA96704D75FF1D9DF2FACFACDA" blockId="6.[151,1437,1289,2035]" pageId="6" pageNumber="353">
Minor cheliped with ischium short, bearing small, stout spiniform seta on ventral margin; merus short, broad, subtriangular in cross-section, about 2.5 times longer than maximal width; ventrolateral margin smooth; ventromesial margin without spiniform setae, with acute distomesial tooth; distodorsal margin not protruding; carpus short, cup-shaped; chela strongly compressed, twisted laterally; crests and grooves on lateral face of chela similar to those of major chela but less distinct; fingers subequal to palm in length; pollex with ventral margin distally curving, cutting edge excavated; dactylus compressed, lamellar, twisted laterally, with high dorsal crest (
<figureCitation id="130B2A6F96704D75FB7E9C26FADAACDA" box="[1188,1294,1901,1928]" captionStart="FIGURE 2" captionStartId="4.[151,250,1614,1639]" captionTargetBox="[187,1390,209,1543]" captionTargetId="figure@4.[164,1423,181,1589]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus gallicus, n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, major cheliped, lateral; b, same, mesial; c, same, fingers; opened, lateral; d, same, dactylus, opened, lateral; e, minor cheliped, lateral; f, same, mesial." figureDoi="http://doi.org/10.5281/zenodo.3653612" httpUri="https://zenodo.org/record/3653612/files/figure.png" pageId="6" pageNumber="353">Fig. 2e, f</figureCitation>
).
</paragraph>
<paragraph id="8B8F36EA96704D75FF1D9CDAFF2BACA1" blockId="6.[151,1437,1289,2035]" pageId="6" pageNumber="353">
Second pereiopod long, slender; ischium slightly longer than merus; carpus with five subdivisions with length ratio approximately equal to 9: 5: 3: 3: 4; chela simple, fingers slightly longer than palm, with distal tufts of setae (
<figureCitation id="130B2A6F96704D75FF7A9C92FF25ACA1" box="[160,241,2009,2035]" captionStart="FIGURE 3" captionStartId="5.[151,250,1899,1924]" captionTargetBox="[210,1376,398,1801]" captionTargetId="figure@5.[164,1423,345,1874]" captionTargetPageId="5" captionText="FIGURE 3. Alpheus gallicus, n. sp., paratype male (CL 9.6 mm), ICM 002619 (ad), and holotype male (CL 11.5 mm) (eg), OUMNH.ZC.2017-01-0061, both from Galicia Bank, NE Atlantic Ocean off western Spain: a, second pereiopod, lateral; b, third pereiopod, lateral; c, same, propodus and dactylus, lateral; d, same, detail of dactylus, lateral; e, fifth pereiopod, lateral; f, same, propodus and dactylus, lateral; g, same, dactylus, lateral." figureDoi="http://doi.org/10.5281/zenodo.3653614" httpUri="https://zenodo.org/record/3653614/files/figure.png" pageId="6" pageNumber="353">Fig. 3a</figureCitation>
).
</paragraph>
<paragraph id="8B8F36EA96714D74FF1D9BD2FD2AAAA5" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="354">
Third pereiopod long, slender; ischium with one cuspidate seta on ventrolateral surface; merus compressed, about 6.5 times as long as maximal width; ventral margin without spiniform setae and without distal tooth; carpus about half-length of merus, slenderer than latter, distal portion of dorsal margin obtusely projecting; propodus slen- der, subequal in length to carpus, armed with twelve or so slender spiniform setae on ventral margin, in addition to two distoventral spiniform setae flanking base of dactylus; dactylus inconspicuously biunguiculate, with proximal unguis much smaller and shorter than distal one, extensor margin bearing small distal notch (
<figureCitation id="130B2A6F96714D74FB139A06FAD9AA3A" box="[1225,1293,333,360]" captionStart="FIGURE 3" captionStartId="5.[151,250,1899,1924]" captionTargetBox="[210,1376,398,1801]" captionTargetId="figure@5.[164,1423,345,1874]" captionTargetPageId="5" captionText="FIGURE 3. Alpheus gallicus, n. sp., paratype male (CL 9.6 mm), ICM 002619 (ad), and holotype male (CL 11.5 mm) (eg), OUMNH.ZC.2017-01-0061, both from Galicia Bank, NE Atlantic Ocean off western Spain: a, second pereiopod, lateral; b, third pereiopod, lateral; c, same, propodus and dactylus, lateral; d, same, detail of dactylus, lateral; e, fifth pereiopod, lateral; f, same, propodus and dactylus, lateral; g, same, dactylus, lateral." figureDoi="http://doi.org/10.5281/zenodo.3653614" httpUri="https://zenodo.org/record/3653614/files/figure.png" pageId="7" pageNumber="354">Fig. 3</figureCitation>
bd). Fourth pereiopod generally similar to third in armature and proportions of articles, somewhat slenderer. Fifth pereiopod slenderest; ischium bearing small cuspidate seta on ventrolateral surface; merus and carpus subequal in length, unarmed; propodus bearing 10 or so spiniform setae on ventral margin and setal brush on distolateral surface; dactylus generally similar to that of third pereiopod (
<figureCitation id="130B2A6F96714D74FD589A96FD10AAA5" box="[642,708,477,503]" captionStart="FIGURE 3" captionStartId="5.[151,250,1899,1924]" captionTargetBox="[210,1376,398,1801]" captionTargetId="figure@5.[164,1423,345,1874]" captionTargetPageId="5" captionText="FIGURE 3. Alpheus gallicus, n. sp., paratype male (CL 9.6 mm), ICM 002619 (ad), and holotype male (CL 11.5 mm) (eg), OUMNH.ZC.2017-01-0061, both from Galicia Bank, NE Atlantic Ocean off western Spain: a, second pereiopod, lateral; b, third pereiopod, lateral; c, same, propodus and dactylus, lateral; d, same, detail of dactylus, lateral; e, fifth pereiopod, lateral; f, same, propodus and dactylus, lateral; g, same, dactylus, lateral." figureDoi="http://doi.org/10.5281/zenodo.3653614" httpUri="https://zenodo.org/record/3653614/files/figure.png" pageId="7" pageNumber="354">Fig. 3</figureCitation>
eg).
</paragraph>
<paragraph id="8B8F36EA96714D74FF1D994AFEEEA936" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="354">
Second pleopod with lateral margin of protopod unarmed, bearing long, fine setae; in males appendix masculina slightly overreaching appendix interna, but not overreaching distal margin of endopod, with numerous long distal setae (
<figureCitation id="130B2A6F96714D74FF049902FEF8A936" box="[222,300,585,612]" captionStart="FIGURE 1" captionStartId="3.[151,250,1909,1934]" captionTargetBox="[187,1400,361,1856]" captionTargetId="figure@3.[160,1427,342,1881]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus gallicus n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, frontal region, dorsal; b, same, lateral; c, telson, dorsal; d, ventromesial carina of first article of antennular peduncle; e, third maxilliped, lateral; f, second pleopod, detail of appendix masculina and appendix interna, anterior (lateral); g, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3653610" httpUri="https://zenodo.org/record/3653610/files/figure.png" pageId="7" pageNumber="354">Fig. 1f</figureCitation>
).
</paragraph>
<paragraph id="8B8F36EA96714D74FF1D9926FCA0A99D" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="354">
Uropod with protopod bearing large acute distolateral tooth; exopod with sinuous transverse suture (diaeresis); distolateral spiniform seta flanked by acute distolateral tooth laterally and broad obtuse projection mesially; endopod bearing row of minute spinules along distal margin (
<figureCitation id="130B2A6F96714D74FCC999FEFCB0A99D" box="[787,868,693,719]" captionStart="FIGURE 1" captionStartId="3.[151,250,1909,1934]" captionTargetBox="[187,1400,361,1856]" captionTargetId="figure@3.[160,1427,342,1881]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus gallicus n. sp., holotype male (CL 11.5 mm), OUMNH.ZC.2017-01-0061, from Galicia Bank, NE Atlantic Ocean off western Spain: a, frontal region, dorsal; b, same, lateral; c, telson, dorsal; d, ventromesial carina of first article of antennular peduncle; e, third maxilliped, lateral; f, second pleopod, detail of appendix masculina and appendix interna, anterior (lateral); g, uropod, dorsal." figureDoi="http://doi.org/10.5281/zenodo.3653610" httpUri="https://zenodo.org/record/3653610/files/figure.png" pageId="7" pageNumber="354">
Fig.
<quantity id="4CC89B0F96714D74FC9D99FEFCB0A99D" box="[839,868,693,719]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="7" pageNumber="354" unit="g" value="1.0">1g</quantity>
</figureCitation>
).
</paragraph>
<paragraph id="8B8F36EA96714D74FF1D9993FF26A869" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="354">
<emphasis id="B944EAF896714D74FF1D9993FE8CA9A1" bold="true" box="[199,344,728,755]" pageId="7" pageNumber="354">Colouration</emphasis>
. Body uniformly reddish, with reddish chromatophores distributed throughout the body (carapace, pleon, telson); major and minor chelae orange-reddish; antennules, antennae and walking legs pale orange-reddish (
<figureCitation id="130B2A6F96714D74FF7A986AFF31A869" box="[160,229,801,827]" captionStart="FIGURE 4" captionStartId="6.[151,250,1182,1207]" captionTargetBox="[151,1436,181,1156]" captionTargetId="figure@6.[151,1436,181,1156]" captionTargetPageId="6" captionText="FIGURE 4. Alpheus gallicus n. sp., unspecified specimen, from Galicia Bank, NE Atlantic Ocean off western Spain: colouration of living individual, lateral. Scale below in cm. Photograph courtesy of Dr. Antonio Puzón." figureDoi="http://doi.org/10.5281/zenodo.3653618" httpUri="https://zenodo.org/record/3653618/files/figure.png" pageId="7" pageNumber="354">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32A656196714D74FF1D980EFEAAA8F5" pageId="7" pageNumber="354" type="etymology">
<paragraph id="8B8F36EA96714D74FF1D980EFEAAA8F5" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="354">
<emphasis id="B944EAF896714D74FF1D980EFE9CA80D" bold="true" box="[199,328,837,863]" pageId="7" pageNumber="354">Etymology</emphasis>
. The new species name refers to the
<typeStatus id="548B884896714D74FCDA980EFCE4A80D" box="[768,816,837,863]" pageId="7" pageNumber="354">type</typeStatus>
locality,
<collectingRegion id="49F4F80896714D74FC46980EFC39A80D" box="[924,1005,837,863]" country="Spain" name="Galicia" pageId="7" pageNumber="354">Galicia</collectingRegion>
Bank, situated in the offshore waters of
<collectingRegion id="49F4F80896714D74FF6C9822FEDEA8D1" box="[182,266,873,899]" country="Spain" name="Galicia" pageId="7" pageNumber="354">Galicia</collectingRegion>
, an autonomous community of
<collectingCountry id="F327767A96714D74FDB49822FD7AA8D1" box="[622,686,873,899]" name="Spain" pageId="7" pageNumber="354">Spain</collectingCountry>
(
<taxonomicName id="4C304D6996714D74FD659822FCCCA8D1" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[703,792,873,899]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="gallicus">
<emphasis id="B944EAF896714D74FD659822FCCCA8D1" box="[703,792,873,899]" italics="true" pageId="7" pageNumber="354">gallicus</emphasis>
</taxonomicName>
—Latin adjective for Galician, with a masculine ending); used as an adjective.
</paragraph>
</subSubSection>
<subSubSection id="C32A656196714D74FF1D98FAFD62A8BD" pageId="7" pageNumber="354" type="biology_ecology">
<paragraph id="8B8F36EA96714D74FF1D98FAFD62A8BD" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="354">
<emphasis id="B944EAF896714D74FF1D98FAFEF2A899" bold="true" box="[199,294,945,971]" pageId="7" pageNumber="354">Ecology</emphasis>
. All specimens were trawled at considerable depths ranging from
<quantity id="4CC89B0F96714D74FBD898FAFB40A899" box="[1026,1172,944,971]" metricMagnitude="2" metricUnit="m" metricValue="7.765" metricValueMax="7.85" metricValueMin="7.68" pageId="7" pageNumber="354" unit="m" value="776.5" valueMax="785.0" valueMin="768.0">768 to 785 m</quantity>
(
<typeStatus id="548B884896714D74FB7B98FAFB07A899" box="[1185,1235,945,971]" pageId="7" pageNumber="354">type</typeStatus>
series), and down to
<quantity id="4CC89B0F96714D74FF6E989EFF2FA8BD" box="[180,251,981,1007]" metricMagnitude="2" metricUnit="m" metricValue="9.09" pageId="7" pageNumber="354" unit="m" value="909.0">909 m</quantity>
(other material, see
<bibRefCitation id="EFA14B1B96714D74FE3A989EFD7FA8BD" author="Cartes, J. E. &amp; Papiol, V. &amp; Frutos, I. &amp; Macpherson, E. &amp; Gonzalez-Pola, C. &amp; Punzon, A. &amp; Valeiras, X. &amp; Serrano, A." box="[480,683,981,1007]" pageId="7" pageNumber="354" pagination="165 - 178" refId="ref6562" refString="Cartes, J. E., Papiol, V., Frutos, I., Macpherson, E., Gonzalez-Pola, C., Punzon, A., Valeiras, X. &amp; Serrano, A. (2014) Distribution and biogeographic trends of decapod assemblages from Galicia Bank (NE Atlantic) at depths between 700 and 1800 m, with connexions to regional water masses. Deep Sea Research II, 106, 165 - 178. https: // doi. org / 10.1016 / j. dsr 2.2013.09.034" type="journal article" year="2014">
Cartes
<emphasis id="B944EAF896714D74FDF5989DFDB4A8BD" box="[559,608,981,1007]" italics="true" pageId="7" pageNumber="354">et al</emphasis>
. 2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32A656196714D7BFF1D98B2FB0DAE61" lastPageId="8" lastPageNumber="355" pageId="7" pageNumber="354" type="discussion">
<paragraph id="8B8F36EA96714D74FF1D98B2FE9FAE29" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="354">
<emphasis id="B944EAF896714D74FF1D98B2FEE0AF41" bold="true" box="[199,308,1017,1043]" pageId="7" pageNumber="354">Remarks</emphasis>
. The distinct pattern of crests and grooves on the chelipeds clearly place
<taxonomicName id="4C304D6996714D74FB4998B2FA84AF41" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[1171,1360,1017,1043]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896714D74FB4998B2FA84AF41" box="[1171,1360,1017,1043]" italics="true" pageId="7" pageNumber="354">Alpheus gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF896714D74FA8098B2FA48AF41" bold="true" box="[1370,1436,1017,1043]" pageId="7" pageNumber="354">
<taxonomicNameLabel id="A277578396714D74FA8098B2FA48AF41" box="[1370,1436,1017,1043]" pageId="7" pageNumber="354" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
within the
<taxonomicName id="4C304D6996714D74FECA9F56FE2AAF65" baseAuthorityName="Hailstone" baseAuthorityYear="1835" box="[272,510,1053,1079]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="macrocheles">
<emphasis id="B944EAF896714D74FECA9F56FE2AAF65" box="[272,510,1053,1079]" italics="true" pageId="7" pageNumber="354">Alpheus macrocheles</emphasis>
</taxonomicName>
species group
<emphasis id="B944EAF896714D74FD729F55FD32AF65" box="[680,742,1054,1079]" italics="true" pageId="7" pageNumber="354">sensu</emphasis>
<bibRefCitation id="EFA14B1B96714D74FD349F56FC71AF65" author="Coutiere, H." box="[750,933,1052,1079]" pageId="7" pageNumber="354" pagination="852 - 921" refId="ref6971" refString="Coutiere, H. (1905) Les Alpheidae. In: Gardiner, J. S. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes. Being the account of the work carried on and of the collections made by an expedition during the years 1899 and 1900. University Press, Cambridge, pp. 852 - 921, pls. 70 - 87." type="book chapter" year="1905">Coutière (1905)</bibRefCitation>
. Within this species group,
<taxonomicName id="4C304D6996714D74FB069F56FA81AF65" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[1244,1365,1053,1079]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896714D74FB069F56FA81AF65" box="[1244,1365,1053,1079]" italics="true" pageId="7" pageNumber="354">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF896714D74FA869F56FA48AF65" bold="true" box="[1372,1436,1053,1079]" pageId="7" pageNumber="354">
<taxonomicNameLabel id="A277578396714D74FA869F56FA48AF65" box="[1372,1436,1053,1079]" pageId="7" pageNumber="354" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is morphologically closest to three deep-water species, viz.
<taxonomicName id="4C304D6996714D74FCED9F0AFC0AAF09" authorityName="Anker &amp; Nizinski" authorityYear="2011" box="[823,990,1089,1115]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="lentiginosus">
<emphasis id="B944EAF896714D74FCED9F0AFC0AAF09" box="[823,990,1089,1115]" italics="true" pageId="7" pageNumber="354">A. lentiginosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C304D6996714D74FC369F0AFBA2AF09" authorityName="Burukovsky" authorityYear="1990" box="[1004,1142,1089,1115]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="romensky">
<emphasis id="B944EAF896714D74FC369F0AFBA2AF09" box="[1004,1142,1089,1115]" italics="true" pageId="7" pageNumber="354">A. romensky</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C304D6996714D74FB6F9F0AFABEAF09" authorityName="Coutiere" authorityYear="1897" box="[1205,1386,1089,1115]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="platydactylus">
<emphasis id="B944EAF896714D74FB6F9F0AFABEAF09" box="[1205,1386,1089,1115]" italics="true" pageId="7" pageNumber="354">A. platydactylus</emphasis>
</taxonomicName>
, but also to the shallow-water
<taxonomicName id="4C304D6996714D74FE669F2EFDBCAF2D" baseAuthorityName="Hailstone" baseAuthorityYear="1835" box="[444,616,1125,1151]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="macrocheles">
<emphasis id="B944EAF896714D74FE669F2EFDBCAF2D" box="[444,616,1125,1151]" italics="true" pageId="7" pageNumber="354">A. macrocheles</emphasis>
</taxonomicName>
. These five taxa are indeed similar in the shape of the frontal margin of the carapace and also many features of the major and minor chelipeds, as well as the walking legs (third to fifth pereiopods). The most important morphological characters uniting these taxa are: (1) the well-developed rostrum and orbital teeth; (2) the major chela palm with well-developed crests and deep grooves, including a superior transverse groove; (3) the major chela dactylus with a greatly reduced plunger; (4) the major chela pollex not or only feebly reduced; (5) the meri of the third and fourth pereiopods without a sharp distoventral tooth; (6) the second article of the antennular peduncle slender (length to width ratio&gt;3); and (7) the dorsal margin of the cheliped meri without spiniform setae.
</paragraph>
<paragraph id="8B8F36EA96714D74FF1D9ECEFBF8AD59" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="354">
<taxonomicName id="4C304D6996714D74FF1D9ECEFE64AECD" authorityName="Anker &amp; Nizinski" authorityYear="2011" box="[199,432,1413,1439]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="lentiginosus">
<emphasis id="B944EAF896714D74FF1D9ECEFE64AECD" box="[199,432,1413,1439]" italics="true" pageId="7" pageNumber="354">Alpheus lentiginosus</emphasis>
</taxonomicName>
, presently known only from its
<typeStatus id="548B884896714D74FCC69ECEFC98AECD" box="[796,844,1413,1439]" pageId="7" pageNumber="354">type</typeStatus>
locality in the Gulf of
<collectingCountry id="F327767A96714D74FB889ECEFB7EAECD" box="[1106,1194,1413,1439]" name="Mexico" pageId="7" pageNumber="354">Mexico</collectingCountry>
, can be distinguished from
<taxonomicName id="4C304D6996714D74FF0F9EE2FE9AAE91" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[213,334,1449,1475]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896714D74FF0F9EE2FE9AAE91" box="[213,334,1449,1475]" italics="true" pageId="7" pageNumber="354">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF896714D74FE8F9EE2FE41AE91" bold="true" box="[341,405,1449,1475]" pageId="7" pageNumber="354">
<taxonomicNameLabel id="A277578396714D74FE8F9EE2FE41AE91" box="[341,405,1449,1475]" pageId="7" pageNumber="354" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the absence of a notch the extensor margin of the dactyli of the third to fifth pereiopods (present in
<taxonomicName id="4C304D6996714D74FEC39E86FE47AEB5" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[281,403,1485,1511]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896714D74FEC39E86FE47AEB5" box="[281,403,1485,1511]" italics="true" pageId="7" pageNumber="354">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF896714D74FE419E86FE0FAEB5" bold="true" box="[411,475,1485,1511]" pageId="7" pageNumber="354">
<taxonomicNameLabel id="A277578396714D74FE419E86FE0FAEB5" box="[411,475,1485,1511]" pageId="7" pageNumber="354" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
), and by the number of spiniform setae on the ventral margin of the cheliped ischia (three in
<emphasis id="B944EAF896714D74FF269EBAFE1CAD59" box="[252,456,1521,1547]" italics="true" pageId="7" pageNumber="354">
<taxonomicName id="4C304D6996714D74FF269EBAFE76AD59" authorityName="Anker &amp; Nizinski" authorityYear="2011" box="[252,418,1521,1547]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="lentiginosus">A. lentiginosus</taxonomicName>
vs.
</emphasis>
one in
<taxonomicName id="4C304D6996714D74FDC19EBAFD47AD59" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[539,659,1521,1547]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896714D74FDC19EBAFD47AD59" box="[539,659,1521,1547]" italics="true" pageId="7" pageNumber="354">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF896714D74FD409EBAFD0DAD59" bold="true" box="[666,729,1521,1547]" pageId="7" pageNumber="354">
<taxonomicNameLabel id="A277578396714D74FD409EBAFD0DAD59" box="[666,729,1521,1547]" pageId="7" pageNumber="354" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
) (cf.
<bibRefCitation id="EFA14B1B96714D74FCCF9EBAFBF5AD59" author="Anker, A. &amp; Nizinski, M." box="[789,1057,1521,1547]" pageId="7" pageNumber="354" pagination="49 - 56" refId="ref6326" refString="Anker, A. &amp; Nizinski, M. (2011) Description of a new deep-water species of Alpheus Fabricius, 1798 from the Gulf of Mexico (Crustacea, Decapoda, Alpheidae). Zootaxa, 2925 (1), 49 - 56. https: // doi. org / 10.11646 / zootaxa. 2925.1.5" type="journal article" year="2011">Anker &amp; Nizinski 2011</bibRefCitation>
).
</paragraph>
<paragraph id="8B8F36EA96714D74FF1D9D5EFD98ADB1" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="354">
<taxonomicName id="4C304D6996714D74FF1D9D5EFE47AD7D" authorityName="Burukovsky" authorityYear="1990" box="[199,403,1557,1583]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="romensky">
<emphasis id="B944EAF896714D74FF1D9D5EFE47AD7D" box="[199,403,1557,1583]" italics="true" pageId="7" pageNumber="354">Alpheus romensky</emphasis>
</taxonomicName>
, was described from a single incomplete specimen collected from the Salas y Gomez Ridge in the Central Pacific. The frontal margin and major cheliped of this species are similar to those of
<taxonomicName id="4C304D6996714D74FB009D72FA85AD01" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[1242,1361,1593,1619]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896714D74FB009D72FA85AD01" box="[1242,1361,1593,1619]" italics="true" pageId="7" pageNumber="354">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF896714D74FA8D9D72FA41AD01" bold="true" box="[1367,1429,1593,1619]" pageId="7" pageNumber="354">
<taxonomicNameLabel id="A277578396714D74FA8D9D72FA41AD01" box="[1367,1429,1593,1619]" pageId="7" pageNumber="354" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, but it can be easily distinguished from the new species by the shape of the minor chela dactylus (slender, mesially somewhat concave in
<taxonomicName id="4C304D6996714D74FE4A9DCAFDCDADC9" authorityName="Burukovsky" authorityYear="1990" box="[400,537,1665,1691]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="romensky">
<emphasis id="B944EAF896714D74FE4A9DCAFDCDADC9" box="[400,537,1665,1691]" italics="true" pageId="7" pageNumber="354">A. romensky</emphasis>
</taxonomicName>
vs. greatly expanded dorsally, distinctly convex in
<taxonomicName id="4C304D6996714D74FB839DCAFB06ADC9" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[1113,1234,1665,1691]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896714D74FB839DCAFB06ADC9" box="[1113,1234,1665,1691]" italics="true" pageId="7" pageNumber="354">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF896714D74FB029DCAFAC3ADC9" bold="true" box="[1240,1303,1665,1691]" pageId="7" pageNumber="354">
<taxonomicNameLabel id="A277578396714D74FB029DCAFAC3ADC9" box="[1240,1303,1665,1691]" pageId="7" pageNumber="354" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
), as well as the development of the superior transverse groove of the major chela (shallow in
<emphasis id="B944EAF896714D74FBF49DEEFB0AADED" box="[1070,1246,1701,1727]" italics="true" pageId="7" pageNumber="354">
<taxonomicName id="4C304D6996714D74FBF49DEEFB6CADED" authorityName="Burukovsky" authorityYear="1990" box="[1070,1208,1701,1727]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="romensky">A. romensky</taxonomicName>
vs.
</emphasis>
fairly deep in
<taxonomicName id="4C304D6996714D74FA5E9DEEFF24ADB1" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896714D74FA5E9DEEFF24ADB1" italics="true" pageId="7" pageNumber="354">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF896714D74FF2D9D82FEE2ADB1" bold="true" box="[247,310,1737,1763]" pageId="7" pageNumber="354">
<taxonomicNameLabel id="A277578396714D74FF2D9D82FEE2ADB1" box="[247,310,1737,1763]" pageId="7" pageNumber="354" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
) (cf.
<bibRefCitation id="EFA14B1B96714D74FEA99D82FDEBADB1" author="Burukovsky, R. N." box="[371,575,1737,1763]" pageId="7" pageNumber="354" pagination="187 - 217" refId="ref6528" refString="Burukovsky, R. N. (1990) Shrimps from the Sala-y-Gomez and Nazca Ridges [in Russian]. Trudy Instituta Okeanologii Akademii Nauk SSSR, 124, 187 - 217." type="journal article" year="1990">Burukovsky 1990</bibRefCitation>
).
</paragraph>
<paragraph id="8B8F36EA96714D74FF1D9DA6FCDDACE9" blockId="7.[151,1437,153,1979]" pageId="7" pageNumber="354">
<taxonomicName id="4C304D6996714D74FF1D9DA6FE6FAC55" authorityName="Coutiere" authorityYear="1897" box="[199,443,1773,1799]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="platydactylus">
<emphasis id="B944EAF896714D74FF1D9DA6FE6FAC55" box="[199,443,1773,1799]" italics="true" pageId="7" pageNumber="354">Alpheus platydactylus</emphasis>
</taxonomicName>
is a deep-water eastern Atlantic species found at depths between
<quantity id="4CC89B0F96714D74FB5A9DA7FB63AC55" box="[1152,1207,1772,1799]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="7" pageNumber="354" unit="m" value="55.0">55 m</quantity>
and
<quantity id="4CC89B0F96714D74FB339DA6FAFAAC55" box="[1257,1326,1773,1799]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="354" unit="m" value="600.0">600 m</quantity>
(
<bibRefCitation id="EFA14B1B96714D74FAE09DA6FF06AC79" author="Coutiere, H." pageId="7" pageNumber="354" pagination="303 - 306" refId="ref6813" refString="Coutiere, H. (1897) Note sur quelques Alphees nouveaux. Bulletin du Museum d'Histoire Naturelle, 3 (7), 303 - 306. https: // doi. org / 10.5962 / bhl. part. 19261" type="journal article" year="1897">Coutière 1897</bibRefCitation>
;
<bibRefCitation id="EFA14B1B96714D74FF049C5AFE3EAC7E" author="Crosnier, A. &amp; Forest, J." box="[222,490,1809,1836]" pageId="7" pageNumber="354" pagination="199 - 314" refId="ref7093" refString="Crosnier, A. &amp; Forest, J. (1966) Crustaces decapodes: Alpheidae. In: Campagnes de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome et Annobon (1956), et Campagne aux Iles du Cap Vert (1959). Part 19. Resultats Scientifiques des Campagnes de la Calypso 7 (27). Annales de l'Institut Oceanographique de Monaco, 44, 199 - 314." type="journal article" year="1966">Crosnier &amp; Forest 1966</bibRefCitation>
). Based on both previously published descriptions and the presently examined material (see comparative material),
<taxonomicName id="4C304D6996714D74FE249C7EFD67AC1D" authorityName="Coutiere" authorityYear="1897" box="[510,691,1845,1871]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="platydactylus">
<emphasis id="B944EAF896714D74FE249C7EFD67AC1D" box="[510,691,1845,1871]" italics="true" pageId="7" pageNumber="354">A. platydactylus</emphasis>
</taxonomicName>
is distinguishable from
<taxonomicName id="4C304D6996714D74FC199C7EFBE8AC1D" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[963,1084,1845,1871]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896714D74FC199C7EFBE8AC1D" box="[963,1084,1845,1871]" italics="true" pageId="7" pageNumber="354">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF896714D74FB999C7EFB56AC1D" bold="true" box="[1091,1154,1845,1871]" pageId="7" pageNumber="354">
<taxonomicNameLabel id="A277578396714D74FB999C7EFB56AC1D" box="[1091,1154,1845,1871]" pageId="7" pageNumber="354" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
mainly by the shape and setation of the minor chela. In
<taxonomicName id="4C304D6996714D74FE239C12FD7AAC21" authorityName="Coutiere" authorityYear="1897" box="[505,686,1881,1907]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="platydactylus">
<emphasis id="B944EAF896714D74FE239C12FD7AAC21" box="[505,686,1881,1907]" italics="true" pageId="7" pageNumber="354">A. platydactylus</emphasis>
</taxonomicName>
, the minor chela dactylus (cf.
<bibRefCitation id="EFA14B1B96714D74FBD49C12FAF4AC21" author="Crosnier, A. &amp; Forest, J." box="[1038,1312,1881,1907]" pageId="7" pageNumber="354" pagination="199 - 314" refId="ref7093" refString="Crosnier, A. &amp; Forest, J. (1966) Crustaces decapodes: Alpheidae. In: Campagnes de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome et Annobon (1956), et Campagne aux Iles du Cap Vert (1959). Part 19. Resultats Scientifiques des Campagnes de la Calypso 7 (27). Annales de l'Institut Oceanographique de Monaco, 44, 199 - 314." type="journal article" year="1966">Crosnier &amp; Forest 1966</bibRefCitation>
: fig.
<quantity id="4CC89B0F96714D74FA819C12FAACAC21" box="[1371,1400,1881,1907]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="7" pageNumber="354" unit="g" value="2.0">2g</quantity>
) is slender and not expanded dorsally (
<emphasis id="B944EAF896714D74FDF69C35FD9FACC5" box="[556,587,1918,1943]" italics="true" pageId="7" pageNumber="354">vs.</emphasis>
strongly expanded in
<taxonomicName id="4C304D6996714D74FC939C36FC16ACC5" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[841,962,1917,1943]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896714D74FC939C36FC16ACC5" box="[841,962,1917,1943]" italics="true" pageId="7" pageNumber="354">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF896714D74FC109C36FBDEACC5" bold="true" box="[970,1034,1917,1943]" pageId="7" pageNumber="354">
<taxonomicNameLabel id="A277578396714D74FC109C36FBDEACC5" box="[970,1034,1917,1943]" pageId="7" pageNumber="354" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
), and bears dense rows of elongate setae throughout its length (absent in
<taxonomicName id="4C304D6996714D74FDE19CEAFD67ACE9" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[571,691,1953,1979]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF896714D74FDE19CEAFD67ACE9" box="[571,691,1953,1979]" italics="true" pageId="7" pageNumber="354">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF896714D74FD609CEAFD2DACE9" bold="true" box="[698,761,1953,1979]" pageId="7" pageNumber="354">
<taxonomicNameLabel id="A277578396714D74FD609CEAFD2DACE9" box="[698,761,1953,1979]" pageId="7" pageNumber="354" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
<paragraph id="8B8F36EA967E4D7BFF1D9BD2FE05A96D" blockId="8.[151,1437,153,1331]" pageId="8" pageNumber="355">
<taxonomicName id="4C304D69967E4D7BFF1D9BD2FE60ABE1" baseAuthorityName="Hailstone" baseAuthorityYear="1835" box="[199,436,153,179]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="macrocheles">
<emphasis id="B944EAF8967E4D7BFF1D9BD2FE60ABE1" box="[199,436,153,179]" italics="true" pageId="8" pageNumber="355">Alpheus macrocheles</emphasis>
</taxonomicName>
is a shallow to moderately deep-water species (typically
<quantity id="4CC89B0F967E4D7BFBF59BD2FB47ABE1" box="[1071,1171,153,179]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="8" pageNumber="355" unit="m" value="100.0" valueMax="200.0" valueMin="0.0">0200 m</quantity>
, but see below) that has been reported from throughout the eastern Atlantic from
<collectingCountry id="F327767A967E4D7BFCCC9BF6FC7EAB85" box="[790,938,189,215]" name="United Kingdom" pageId="8" pageNumber="355">Great Britain</collectingCountry>
to
<collectingCountry id="F327767A967E4D7BFC149BF6FBCFAB85" box="[974,1051,189,215]" name="Gabon" pageId="8" pageNumber="355">Gabon</collectingCountry>
, including the Mediterranean Sea, as well as from
<collectingCountry id="F327767A967E4D7BFE949BAAFDC4ABA9" box="[334,528,225,251]" name="Ascension Island" pageId="8" pageNumber="355">Ascension Island</collectingCountry>
and
<collectingCountry id="F327767A967E4D7BFD919BAAFD15ABA9" box="[587,705,225,251]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="8" pageNumber="355">St. Helena</collectingCountry>
in the central Atlantic (
<bibRefCitation id="EFA14B1B967E4D7BFC089BAAFBACABA9" author="Holthuis, L. B." box="[978,1144,224,251]" pageId="8" pageNumber="355" pagination="7 - 187" refId="ref7683" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical west Africa. Atlantide Report Scientific Results of the Danish Expedition to the Coasts of Tropical West Africa 1945 - 1946, 2, 7 - 187." type="journal article" year="1951">Holthuis 1951</bibRefCitation>
;
<bibRefCitation id="EFA14B1B967E4D7BFB5C9BAAFA43ABA9" author="Crosnier, A. &amp; Forest, J." box="[1158,1431,225,251]" pageId="8" pageNumber="355" pagination="199 - 314" refId="ref7093" refString="Crosnier, A. &amp; Forest, J. (1966) Crustaces decapodes: Alpheidae. In: Campagnes de la Calypso dans le Golfe de Guinee et aux Iles Principe, Sao Tome et Annobon (1956), et Campagne aux Iles du Cap Vert (1959). Part 19. Resultats Scientifiques des Campagnes de la Calypso 7 (27). Annales de l'Institut Oceanographique de Monaco, 44, 199 - 314." type="journal article" year="1966">Crosnier &amp; Forest 1966</bibRefCitation>
;
<bibRefCitation id="EFA14B1B967E4D7BFF4D9A4EFEF4AA4D" author="Chace, F. A. Jr." box="[151,288,261,287]" pageId="8" pageNumber="355" pagination="622 - 662" refId="ref6659" refString="Chace, F. A. Jr. (1966) Decapod crustaceans from St. Helena Island, South Atlantic. Proceedings of the United States National Museum, 118, 622 - 662. https: // doi. org / 10.5479 / si. 00963801.118 - 3536.623" type="journal article" year="1966">Chace 1966</bibRefCitation>
; Manning &amp; Chance 1990,
<bibRefCitation id="EFA14B1B967E4D7BFD809A4EFC92AA4D" author="Soledade, G. O. &amp; Scioli, J. A. &amp; Terossi, M. &amp; Mantelatto, F. L. &amp; Almeida, A. O." box="[602,838,261,287]" pageId="8" pageNumber="355" pagination="1 - 22" refId="ref8121" refString="Soledade, G. O., Scioli, J. A., Terossi, M., Mantelatto, F. L. &amp; Almeida, A. O. (2019) Congruence between morphology and molecular data supports the distinction between the snapping shrimps Alpheus macrocheles and A. amblyonyx (Caridea, Alpheidae). Zootaxa, 4624 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 4624.1.1" type="journal article" year="2019">
Soledade
<emphasis id="B944EAF8967E4D7BFD129A4DFCD5AA4D" box="[712,769,261,287]" italics="true" pageId="8" pageNumber="355">et al.</emphasis>
2019
</bibRefCitation>
).
<taxonomicName id="4C304D69967E4D7BFC839A4EFB93AA4D" baseAuthorityName="Hailstone" baseAuthorityYear="1835" box="[857,1095,261,287]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="macrocheles">
<emphasis id="B944EAF8967E4D7BFC839A4EFB93AA4D" box="[857,1095,261,287]" italics="true" pageId="8" pageNumber="355">Alpheus macrocheles</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName id="4C304D69967E4D7BFA5E9A4EFF24AA11" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFA5E9A4EFF24AA11" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFF239A62FEEDAA11" bold="true" box="[249,313,297,323]" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFF239A62FEEDAA11" box="[249,313,297,323]" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the shape of the dactyli of the third to fifth pereiopods (simple in
<taxonomicName id="4C304D69967E4D7BFB809A62FAD3AA11" baseAuthorityName="Hailstone" baseAuthorityYear="1835" box="[1114,1287,297,323]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="macrocheles">
<emphasis id="B944EAF8967E4D7BFB809A62FAD3AA11" box="[1114,1287,297,323]" italics="true" pageId="8" pageNumber="355">A. macrocheles</emphasis>
</taxonomicName>
vs. minutely biunguiculate in
<taxonomicName id="4C304D69967E4D7BFE8D9A06FE05AA35" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[343,465,333,359]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFE8D9A06FE05AA35" box="[343,465,333,359]" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFE039A06FDCEAA35" bold="true" box="[473,538,333,359]" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFE039A06FDCEAA35" box="[473,538,333,359]" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
); the shorter and stouter antennular peduncles (length-with ratio of the second article
<quantity id="4CC89B0F967E4D7BFF3E9A3BFE94AAD9" box="[228,320,368,395]" metricMagnitude="-2" metricUnit="m" metricValue="6.985" metricValueMax="7.62" metricValueMin="6.35" pageId="8" pageNumber="355" unit="in" value="2.75" valueMax="3.0" valueMin="2.5">2.53 in</quantity>
<taxonomicName id="4C304D69967E4D7BFE9D9A3AFE3CAAD9" box="[327,488,369,395]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="macrochele">
<emphasis id="B944EAF8967E4D7BFE9D9A3AFE3CAAD9" box="[327,488,369,395]" italics="true" pageId="8" pageNumber="355">A. macrochele</emphasis>
</taxonomicName>
s vs. closer to
<quantity id="4CC89B0F967E4D7BFD5D9A3AFD66AAD9" box="[647,690,369,395]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="8" pageNumber="355" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C304D69967E4D7BFD629A3AFCE4AAD9" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[696,816,369,395]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFD629A3AFCE4AAD9" box="[696,816,369,395]" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFCED9A3AFCA2AAD9" bold="true" box="[823,886,369,395]" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFCED9A3AFCA2AAD9" box="[823,886,369,395]" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
); the length of the stylocerite (reaching or nearly reaching the distal margin of the first antennular article in
<taxonomicName id="4C304D69967E4D7BFCE99ADEFC34AAFD" baseAuthorityName="Hailstone" baseAuthorityYear="1835" box="[819,992,405,431]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="macrocheles">
<emphasis id="B944EAF8967E4D7BFCE99ADEFC34AAFD" box="[819,992,405,431]" italics="true" pageId="8" pageNumber="355">A. macrocheles</emphasis>
</taxonomicName>
vs. not reaching its distal margin in
<taxonomicName id="4C304D69967E4D7BFA5E9ADEFF24AA81" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFA5E9ADEFF24AA81" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFF2D9AF2FEE1AA81" bold="true" box="[247,309,441,467]" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFF2D9AF2FEE1AA81" box="[247,309,441,467]" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
); and the number of spiniform setae on the ventral surface of the major cheliped ischium (three in
<taxonomicName id="4C304D69967E4D7BFA5E9AF2FEF7AAA5" baseAuthorityName="Hailstone" baseAuthorityYear="1835" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="macrocheles">
<emphasis id="B944EAF8967E4D7BFA5E9AF2FEF7AAA5" italics="true" pageId="8" pageNumber="355">A. macrocheles</emphasis>
</taxonomicName>
vs. one in
<taxonomicName id="4C304D69967E4D7BFE469A96FDC0AAA5" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[412,532,477,503]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFE469A96FDC0AAA5" box="[412,532,477,503]" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFDC19A96FD8DAAA5" bold="true" box="[539,601,477,503]" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFDC19A96FD8DAAA5" box="[539,601,477,503]" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
). The two species also differ by the colouration of the major chela, with
<taxonomicName id="4C304D69967E4D7BFA5E9A96FEF7A949" baseAuthorityName="Hailstone" baseAuthorityYear="1835" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="macrocheles">
<emphasis id="B944EAF8967E4D7BFA5E9A96FEF7A949" italics="true" pageId="8" pageNumber="355">A. macrocheles</emphasis>
</taxonomicName>
displaying a transverse, more or less W-shaped blotch on the distomesial face of the palm (cf.
<bibRefCitation id="EFA14B1B967E4D7BFAE0994AFE92A96D" author="Anker, A. &amp; De Grave, S." pageId="8" pageNumber="355" pagination="1 - 15" refId="ref6264" refString="Anker, A. &amp; De Grave, S. (2012) Description of Alpheus cedrici sp. n., a strikingly coloured snapping shrimp (Crustacea, Decapoda, Alpheidae) from Ascension Island, central Atlantic Ocean. ZooKeys, 183, 1 - 15. https: // doi. org / 10.3897 / zookeys. 183.3073" type="journal article" year="2012">Anker &amp; De Grave 2012</bibRefCitation>
: fig. 4A, B).
</paragraph>
<paragraph id="8B8F36EA967E4D7BFF1D9902FE81AF41" blockId="8.[151,1437,153,1331]" pageId="8" pageNumber="355">
Several other members of the
<taxonomicName id="4C304D69967E4D7BFDC39902FD11A931" baseAuthorityName="Hailstone" baseAuthorityYear="1835" box="[537,709,585,611]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="macrocheles">
<emphasis id="B944EAF8967E4D7BFDC39902FD11A931" box="[537,709,585,611]" italics="true" pageId="8" pageNumber="355">A. macrocheles</emphasis>
</taxonomicName>
species group seem to be more distantly related to
<taxonomicName id="4C304D69967E4D7BFADC9902FAABA931" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[1286,1407,585,611]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFADC9902FAABA931" box="[1286,1407,585,611]" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFA5C9902FF6CA9D5" bold="true" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFA5C9902FF6CA9D5" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
based on their morphological characteristics.
<taxonomicName id="4C304D69967E4D7BFD609926FBBCA9D5" authority="Christoffersen, 1979" authorityName="Christoffersen" authorityYear="1979" box="[698,1128,621,647]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="puapeba">
<emphasis id="B944EAF8967E4D7BFD609926FCAFA9D5" box="[698,891,621,647]" italics="true" pageId="8" pageNumber="355">Alpheus puapeba</emphasis>
<bibRefCitation id="EFA14B1B967E4D7BFC599926FBBCA9D5" author="Christoffersen, M. L." box="[899,1128,621,647]" pageId="8" pageNumber="355" pagination="297 - 377" refId="ref6767" refString="Christoffersen, M. L. (1979) Campagne de la Calypso au large des cotes Atlantiques de l'Amerique du Sud (1961 - 1962). I. Decapod Crustacea: Alpheoidea. Annales de l'Institut Oceanographique, 55, 297 - 377." type="journal article" year="1979">Christoffersen, 1979</bibRefCitation>
</taxonomicName>
is a moderately deep-water species (
<quantity id="4CC89B0F967E4D7BFF2D99DBFEBDA9F9" box="[247,361,656,683]" metricMagnitude="2" metricUnit="m" metricValue="1.1" metricValueMax="1.75" metricValueMin="0.45" pageId="8" pageNumber="355" unit="m" value="110.0" valueMax="175.0" valueMin="45.0">45175 m</quantity>
) known from
<collectingCountry id="F327767A967E4D7BFDD299DAFD9BA9F9" box="[520,591,657,683]" name="Brazil" pageId="8" pageNumber="355">Brazil</collectingCountry>
,
<collectingCountry id="F327767A967E4D7BFD8099DAFD68A9F9" box="[602,700,657,683]" name="Uruguay" pageId="8" pageNumber="355">Uruguay</collectingCountry>
and
<collectingCountry id="F327767A967E4D7BFD2A99DAFC8BA9F9" box="[752,863,657,683]" name="Argentina" pageId="8" pageNumber="355">Argentina</collectingCountry>
(
<bibRefCitation id="EFA14B1B967E4D7BFCB799DAFB85A9F9" author="Christoffersen, M. L." box="[877,1105,657,683]" pageId="8" pageNumber="355" pagination="297 - 377" refId="ref6767" refString="Christoffersen, M. L. (1979) Campagne de la Calypso au large des cotes Atlantiques de l'Amerique du Sud (1961 - 1962). I. Decapod Crustacea: Alpheoidea. Annales de l'Institut Oceanographique, 55, 297 - 377." type="journal article" year="1979">Christoffersen 1979</bibRefCitation>
). It can be distinguished from
<taxonomicName id="4C304D69967E4D7BFF4D99FEFEC6A99D" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[151,274,693,719]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFF4D99FEFEC6A99D" box="[151,274,693,719]" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFEC199FEFE88A99D" bold="true" box="[283,348,693,719]" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFEC199FEFE88A99D" box="[283,348,693,719]" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the shape of the dactyli of the third to fifth pereiopods (simple in
<emphasis id="B944EAF8967E4D7BFB5C99FEFAFBA99D" box="[1158,1327,693,719]" italics="true" pageId="8" pageNumber="355">
<taxonomicName id="4C304D69967E4D7BFB5C99FEFAD3A99D" authorityName="Christoffersen" authorityYear="1979" box="[1158,1287,693,719]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="puapeba">A. puapeba</taxonomicName>
vs.
</emphasis>
minutely biunguiculate in
<taxonomicName id="4C304D69967E4D7BFE8E9992FE19A9A1" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[340,461,729,755]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFE8E9992FE19A9A1" box="[340,461,729,755]" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFE0E9992FDC0A9A1" bold="true" box="[468,532,729,755]" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFE0E9992FDC0A9A1" box="[468,532,729,755]" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
); the shape of the minor cheliped dactylus (not expanded in
<emphasis id="B944EAF8967E4D7BFB1B9992FAB3A9A1" box="[1217,1383,729,755]" italics="true" pageId="8" pageNumber="355">
<taxonomicName id="4C304D69967E4D7BFB1B9992FA94A9A1" authorityName="Christoffersen" authorityYear="1979" box="[1217,1344,729,755]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="puapeba">A. puapeba</taxonomicName>
vs.
</emphasis>
dorsally expanded in
<taxonomicName id="4C304D69967E4D7BFEBB99B6FE0CA845" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[353,472,765,791]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFEBB99B6FE0CA845" box="[353,472,765,791]" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFE0599B6FDC9A845" bold="true" box="[479,541,765,791]" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFE0599B6FDC9A845" box="[479,541,765,791]" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
); and the development of the inferior transverse groove of the major chela (with strongly overhanging shoulder in
<emphasis id="B944EAF8967E4D7BFDCC986AFD6AA869" box="[534,702,801,827]" italics="true" pageId="8" pageNumber="355">
<taxonomicName id="4C304D69967E4D7BFDCC986AFD43A869" authorityName="Christoffersen" authorityYear="1979" box="[534,663,801,827]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="puapeba">A. puapeba</taxonomicName>
vs.
</emphasis>
with very shallowly overhanging shoulder in
<taxonomicName id="4C304D69967E4D7BFB10986AFA90A869" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[1226,1348,801,827]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFB10986AFA90A869" box="[1226,1348,801,827]" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFA96986AFA58A869" bold="true" box="[1356,1420,801,827]" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFA96986AFA58A869" box="[1356,1420,801,827]" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
). The members of the Indo-West Pacific
<taxonomicName id="4C304D69967E4D7BFD93980EFD01A80D" authorityName="Coutiere" authorityYear="1905" box="[585,725,837,863]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="hailstonei">
<emphasis id="B944EAF8967E4D7BFD93980EFD01A80D" box="[585,725,837,863]" italics="true" pageId="8" pageNumber="355">A. hailstonei</emphasis>
</taxonomicName>
species complex, which includes the non-monophyletic
<taxonomicName id="4C304D69967E4D7BFA93980EFF0EA8D1" authorityName="Coutiere" authorityYear="1905" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="hailstonei">
<emphasis id="B944EAF8967E4D7BFA93980EFF0EA8D1" italics="true" pageId="8" pageNumber="355">A. hailstonei</emphasis>
</taxonomicName>
(Scioli
<emphasis id="B944EAF8967E4D7BFEE89821FEBEA8D1" box="[306,362,873,899]" italics="true" pageId="8" pageNumber="355">et al.</emphasis>
, in prep.),
<taxonomicName id="4C304D69967E4D7BFE399822FCE4A8D1" authority="Banner, 1953" authorityName="Banner" authorityYear="1953" box="[483,816,872,899]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="brachymerus">
<emphasis id="B944EAF8967E4D7BFE399822FD40A8D1" box="[483,660,873,899]" italics="true" pageId="8" pageNumber="355">A. brachymerus</emphasis>
<bibRefCitation id="EFA14B1B967E4D7BFD419822FCE4A8D1" author="Banner, A. H." box="[667,816,872,899]" pageId="8" pageNumber="355" pagination="3 - 147" refId="ref6385" refString="Banner, A. H. (1953) The Crangonidae, or snapping shrimp, of Hawaii. Pacific Science, 7, 3 - 147." type="journal article" year="1953">Banner, 1953</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C304D69967E4D7BFCBC9822FB7AA8D1" authority="Banner, 1959" authorityName="Banner" authorityYear="1959" box="[870,1198,872,899]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="lanceostylus">
<emphasis id="B944EAF8967E4D7BFCBC9822FBDAA8D1" box="[870,1038,873,899]" italics="true" pageId="8" pageNumber="355">A. lanceostylus</emphasis>
<bibRefCitation id="EFA14B1B967E4D7BFBCF9822FB7AA8D1" author="Banner, A. H." box="[1045,1198,872,899]" pageId="8" pageNumber="355" pagination="130 - 155" refId="ref6413" refString="Banner, A. H. (1959) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part IV. Various small collections from the Central Pacific area, including supplementary notes on alpheids from Hawaii. Pacific Science, 13, 130 - 155." type="journal article" year="1959">Banner, 1959</bibRefCitation>
</taxonomicName>
, can be distinguished from
<taxonomicName id="4C304D69967E4D7BFF0E98C6FE9FA8F5" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[212,331,909,935]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFF0E98C6FE9FA8F5" box="[212,331,909,935]" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFE8898C6FE44A8F5" bold="true" box="[338,400,909,935]" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFE8898C6FE44A8F5" box="[338,400,909,935]" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by the presence of spiniform setae on the dorsal margin of the cheliped meri (present in most members of this complex, absent in
<taxonomicName id="4C304D69967E4D7BFDF098FAFD76A899" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[554,674,945,971]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFDF098FAFD76A899" box="[554,674,945,971]" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFD7398FAFD33A899" bold="true" box="[681,743,945,971]" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFD7398FAFD33A899" box="[681,743,945,971]" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
), as well as by several features on the minor cheliped, such as the proportions of articles and the general shape of the minor chela (cf.
<bibRefCitation id="EFA14B1B967E4D7BFC62989EFB8FA8BD" author="Coutiere, H." box="[952,1115,980,1007]" pageId="8" pageNumber="355" pagination="852 - 921" refId="ref6971" refString="Coutiere, H. (1905) Les Alpheidae. In: Gardiner, J. S. (Ed.), The fauna and geography of the Maldive and Laccadive Archipelagoes. Being the account of the work carried on and of the collections made by an expedition during the years 1899 and 1900. University Press, Cambridge, pp. 852 - 921, pls. 70 - 87." type="book chapter" year="1905">Coutière 1905</bibRefCitation>
;
<bibRefCitation id="EFA14B1B967E4D7BFBBD989EFB2EA8BD" author="Banner, A. H." box="[1127,1274,980,1007]" pageId="8" pageNumber="355" pagination="3 - 147" refId="ref6385" refString="Banner, A. H. (1953) The Crangonidae, or snapping shrimp, of Hawaii. Pacific Science, 7, 3 - 147." type="journal article" year="1953">Banner 1953</bibRefCitation>
,
<bibRefCitation id="EFA14B1B967E4D7BFADF989FFA94A8BD" author="Banner, A. H." box="[1285,1344,980,1007]" pageId="8" pageNumber="355" pagination="130 - 155" refId="ref6413" refString="Banner, A. H. (1959) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part IV. Various small collections from the Central Pacific area, including supplementary notes on alpheids from Hawaii. Pacific Science, 13, 130 - 155." type="journal article" year="1959">1959</bibRefCitation>
;
<bibRefCitation id="EFA14B1B967E4D7BFA91989EFE9CAF41" author="Banner, D. M. &amp; Banner, A. H." pageId="8" pageNumber="355" pagination="1 - 357" refId="ref6464" refString="Banner, D. M. &amp; Banner, A. H. (1982) The alpheid shrimp of Australia. Part III: The remaining alpheids, principally the genus Alpheus and the family Ogyrididae. Records of the Australian Museum, 34, 1 - 357. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.434" type="journal article" year="1982">Banner &amp; Banner 1982</bibRefCitation>
).
</paragraph>
<paragraph id="8B8F36EA967E4D7BFF1D9F56FF01AE5D" blockId="8.[151,1437,153,1331]" pageId="8" pageNumber="355">
The poorly known
<taxonomicName id="4C304D69967E4D7BFE429F56FCC2AF65" authority="Guerin-Meneville, 1855" authorityName="Guerin-Meneville" authorityYear="1855" box="[408,790,1052,1079]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="candei">
<emphasis id="B944EAF8967E4D7BFE429F56FDD5AF65" box="[408,513,1053,1079]" italics="true" pageId="8" pageNumber="355">A. candei</emphasis>
Guérin-Méneville, 1855
</taxonomicName>
, originally described from
<collectingCountry id="F327767A967E4D7BFBE49F56FBA8AF65" box="[1086,1148,1053,1079]" name="Cuba" pageId="8" pageNumber="355">Cuba</collectingCountry>
, is a taxonomically highly problematic species that has not been collected since
<bibRefCitation id="EFA14B1B967E4D7BFD309F0AFC7AAF09" author="Coutiere, H." box="[746,942,1089,1115]" pageId="8" pageNumber="355" pagination="485 - 487" refId="ref7045" refString="Coutiere, H. (1910) The snapping shrimps (Alpheidae) of the Dry Tortugas, Florida. Proceedings of the United States National Museum, 37, 485 - 487. https: // doi. org / 10.5479 / si. 00963801.37 - 1716.485" type="journal article" year="1910">Coutières (1910)</bibRefCitation>
tentative identification of a single specimen from Dry
<collectingCountry id="F327767A967E4D7BFEDF9F2EFEBEAF2D" box="[261,362,1125,1151]" name="Haiti" pageId="8" pageNumber="355">Tortugas</collectingCountry>
, Florida.
<taxonomicName id="4C304D69967E4D7BFE089F2EFDA9AF2D" authorityName="Guerin-Meneville" authorityYear="1855" box="[466,637,1125,1151]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="candei">
<emphasis id="B944EAF8967E4D7BFE089F2EFDA9AF2D" box="[466,637,1125,1151]" italics="true" pageId="8" pageNumber="355">Alpheus candei</emphasis>
</taxonomicName>
, if confirmed to be valid, can be easily distinguished from
<taxonomicName id="4C304D69967E4D7BFAD39F2EFA54AF2D" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[1289,1408,1125,1151]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFAD39F2EFA54AF2D" box="[1289,1408,1125,1151]" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFA5C9F2EFF6CAFF1" bold="true" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFA5C9F2EFF6CAFF1" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by multiple characters of the frontal region (shape of the rostrum and orbital teeth, proportions of the antennular articles, much shorter scaphocerite blade, etc.) and minor cheliped (e.g. with the dactylus not expanded in
<taxonomicName id="4C304D69967E4D7BFAE99FE6FA49AF95" authorityName="Guerin-Meneville" authorityYear="1855" box="[1331,1437,1197,1223]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="candei">
<emphasis id="B944EAF8967E4D7BFAE99FE6FA49AF95" box="[1331,1437,1197,1223]" italics="true" pageId="8" pageNumber="355">A. candei</emphasis>
</taxonomicName>
vs. dorsally expanded in
<taxonomicName id="4C304D69967E4D7BFE6F9F9AFDFBAFB9" authority="Scioli &amp; Anker, 2020" authorityName="Scioli &amp; Anker" authorityYear="2020" box="[437,559,1233,1259]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="355" phylum="Arthropoda" rank="species" species="gallicus" status="sp. nov.">
<emphasis id="B944EAF8967E4D7BFE6F9F9AFDFBAFB9" box="[437,559,1233,1259]" italics="true" pageId="8" pageNumber="355">A. gallicus</emphasis>
</taxonomicName>
<emphasis id="B944EAF8967E4D7BFDED9F9AFDA3AFB9" bold="true" box="[567,631,1233,1259]" pageId="8" pageNumber="355">
<taxonomicNameLabel id="A2775783967E4D7BFDED9F9AFDA3AFB9" box="[567,631,1233,1259]" pageId="8" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
). In addition, it appears to be a shallow-water species inhabiting coral reefs.
</paragraph>
<paragraph id="8B8F36EA967E4D7BFF1D9E53FB0DAE61" blockId="8.[151,1437,153,1331]" box="[199,1241,1304,1331]" pageId="8" pageNumber="355">
<emphasis id="B944EAF8967E4D7BFF1D9E53FE0DAE61" bold="true" box="[199,473,1304,1331]" pageId="8" pageNumber="355">Genbank Accession No</emphasis>
. OUMNH.ZC.2017-01-063:
<accessionNumber id="9463AB09967E4D7BFCF89E53FC72AE61" box="[802,934,1304,1331]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN754223" pageId="8" pageNumber="355">MN754223</accessionNumber>
, ICM 002618:
<accessionNumber id="9463AB09967E4D7BFB8A9E53FB01AE61" box="[1104,1237,1304,1331]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN754224" pageId="8" pageNumber="355">MN754224</accessionNumber>
.
</paragraph>
</subSubSection>
</treatment>
</document>