188 lines
21 KiB
XML
188 lines
21 KiB
XML
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<mods:title id="37F15F7918972BD7E0B856850CDDB340">New Black Fungus Gnats (Diptera, Sciaridae) of North America. Part IV. Genera Eugnoriste Coquillett and Keilbachia Mohrig</mods:title>
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<mods:title id="742C46E75714805CA441EF749B4697AA">Zootaxa</mods:title>
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<treatment id="039944034F1E6D58388AF800FA4656FB" ID-DOI="http://doi.org/10.5281/zenodo.6003900" ID-GBIF-Taxon="134017001" ID-Zenodo-Dep="6003900" LSID="urn:lsid:plazi:treatment:039944034F1E6D58388AF800FA4656FB" httpUri="http://treatment.plazi.org/id/039944034F1E6D58388AF800FA4656FB" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54">
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<subSubSection id="C32AA69E4F1E6D5B388AF800FD5953BB" box="[151,643,1939,1965]" pageId="1" pageNumber="54" type="nomenclature">
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<emphasis id="B94429074F1E6D5B388AF800FD5953BB" bold="true" box="[151,643,1939,1965]" pageId="1" pageNumber="54">
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Key to the species of
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<taxonomicName id="4C308E964F1E6D5B3981F800FDF853BB" ID-CoL="635DQ" ID-ENA="1895447" box="[412,546,1939,1965]" class="Insecta" family="Sciaridae" genus="Eugnoriste" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="54" phylum="Arthropoda" rank="genus">
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<emphasis id="B94429074F1E6D5B3981F800FDF853BB" bold="true" box="[412,546,1939,1965]" italics="true" pageId="1" pageNumber="54">Eugnoriste</emphasis>
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</taxonomicName>
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(males)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32AA69E4F1E6D58388AF849FA4656FB" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54" type="key">
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<keyStep id="30C4EDB04F1E6D58388AF849FA4654DC" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54">
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<paragraph id="8B8FF5154F1E6D5B388AF849FA4253F9" blockId="1.[151,1432,2010,2031]" box="[151,1432,2010,2031]" pageId="1" pageNumber="54">
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<keyLead id="30C156204F1E6D5B388AF849FA4253F9" box="[151,1432,2010,2031]" pageId="1" pageNumber="54">1 Mouth part very long in both sexes, at least as long as the head high; gonostylus with long and dense hairs on the outer side..</keyLead>
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</paragraph>
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<paragraph id="8B8FF5154F1D6D5838FEFF04FA4654DC" blockId="2.[151,1437,151,750]" pageId="2" pageNumber="55">
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.................................................................................................... 2
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<keyLead id="30C156204F1D6D58388AFF26FA4654DC" box="[151,1436,181,202]" pageId="2" pageNumber="55">- Mouth part long in both sexes, but shorter than the head high; gonostylus with short hairs on the on outer side............ 3</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30C4EDB04F1D6D58388AFF41FA465557" pageId="2" pageNumber="55">
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<paragraph id="8B8FF5154F1D6D58388AFF41FA415512" blockId="2.[151,1437,151,750]" pageId="2" pageNumber="55">
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<keyLead id="30C156204F1D6D58388AFF41FA415512" pageId="2" pageNumber="55">
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2 Mouth part in both sexes twice as long as the head high; haired areas on distal tergites not distinctly darkened............................................................................... ..
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<emphasis id="B94429074F1D6D583C15FF7CFBED5513" bold="true" box="[1032,1079,239,261]" italics="true" pageId="2" pageNumber="55">Eug.</emphasis>
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<taxonomicName id="4C308E964F1D6D583C20FF7CFA945512" ID-CoL="3C888" authority="Coquillett, 1896" authorityName="CoquIllett. The" authorityYear="1896" box="[1085,1358,239,261]" class="Insecta" family="Sciaridae" genus="Eugnoriste" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="occidentalis">
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<emphasis id="B94429074F1D6D583C20FF7CFB685513" bold="true" box="[1085,1202,239,261]" italics="true" pageId="2" pageNumber="55">occidentalis</emphasis>
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Coquillett,1896
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</taxonomicName>
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(
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<figureCitation id="130BE9904F1D6D583D46FF7CFA4F5512" box="[1371,1429,239,260]" captionStart="FIGURE 6" captionStartId="9.[151,250,1971,1993]" captionTargetBox="[173,1400,877,1926]" captionTargetId="figure@9.[151,1436,873,1949]" captionTargetPageId="9" captionText="FIGURE 6. Eugnoriste occidentalis Coquillett, 1896. A. Male. Part of hypopygium (speCimen from California), ventral vieW. B. Female. Head (after Menzel & Mohrig 2000, Fig. 573)." httpUri="https://zenodo.org/record/888609/files/figure.png" pageId="2" pageNumber="55">Fig. 6</figureCitation>
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)
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</keyLead>
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</paragraph>
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<paragraph id="8B8FF5154F1D6D58388AFE9CFA465557" blockId="2.[151,1437,151,750]" pageId="2" pageNumber="55">
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<keyLead id="30C156204F1D6D58388AFE9CFA465557" pageId="2" pageNumber="55">
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- Mouth part in both sexes as long as the head high; haired area of the 7th tergite black................................................................................................... ..
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<emphasis id="B94429074F1D6D583C1AFEBFFBEC5554" bold="true" box="[1031,1078,300,322]" italics="true" pageId="2" pageNumber="55">Eug.</emphasis>
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<taxonomicName id="4C308E964F1D6D583C21FEBFFA945557" ID-CoL="3C887" authority="Coquillett, 1904" authorityName="Coquillett" authorityYear="1904" box="[1084,1358,300,322]" class="Insecta" family="Sciaridae" genus="Eugnoriste" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="brevirostris">
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<emphasis id="B94429074F1D6D583C21FEBFFB715554" bold="true" box="[1084,1195,300,322]" italics="true" pageId="2" pageNumber="55">brevirostris</emphasis>
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<bibRefCitation id="EFA188E44F1D6D583CAFFEBFFA945557" author="Coquillett" box="[1202,1358,300,321]" pageId="2" pageNumber="55" refString="Coquillett, D. W. (1904) NeW North AmeriCan Diptera. Proceedings of the Entomological Society of Washington, 6 (3), 166 - 192." type="journal article" year="1904">Coquillett, 1904</bibRefCitation>
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</taxonomicName>
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(
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<figureCitation id="130BE9904F1D6D583D46FEBFFA4E5557" box="[1371,1428,300,321]" captionStart="FIGURE 3" captionStartId="5.[151,250,1969,1991]" captionTargetBox="[157,1428,221,1930]" captionTargetId="figure@5.[151,1436,195,1948]" captionTargetPageId="5" captionText="FIGURE 3. Eugnoriste brevirostris Coquillett, 1904. Male. Holotype. A. Part of hypopygium, ventral vieW. B. Gonostylus (draWing, 200 X). C. Head. D. Antennal flagellomeres 3 – 4. E. Abdominal tergites 3 – 6. F. CerCus (draWing, 200 X)." httpUri="https://zenodo.org/record/888603/files/figure.png" pageId="2" pageNumber="55">Fig. 3</figureCitation>
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)
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30C4EDB04F1D6D58388AFEDFFA41558A" pageId="2" pageNumber="55">
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<paragraph id="8B8FF5154F1D6D58388AFEDFFA465569" blockId="2.[151,1437,151,750]" pageId="2" pageNumber="55">
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<keyLead id="30C156204F1D6D58388AFEDFFA465569" pageId="2" pageNumber="55">
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3 Gonostylus slender, tapered towards the apex; ventral base of the hypopygium bare; 9th tergite bilobate, length of lobes equals one third of the length of the gonocoxites...............................................
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<emphasis id="B94429074F1D6D583CBFFEFAFA945569" bold="true" box="[1186,1358,361,383]" pageId="2" pageNumber="55">
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||
<emphasis id="B94429074F1D6D583CBFFEFAFAC85569" bold="true" box="[1186,1298,361,383]" italics="true" pageId="2" pageNumber="55">
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Eug.
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<taxonomicName id="4C308E964F1D6D583CCAFEFAFAC85569" ID-CoL="9HYK8" authority="Kauschke, 2017" authorityName="Kauschke" authorityYear="2017" box="[1239,1298,361,383]" class="Insecta" family="Sciaridae" genus="Eugnoriste" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="florea" status="sp. nov.">florea</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A277947C4F1D6D583D05FEFAFA945569" box="[1304,1358,361,383]" pageId="2" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="130BE9904F1D6D583D46FEF9FA4F5569" box="[1371,1429,362,383]" captionStart="FIGURE 4" captionStartId="7.[151,250,1600,1622]" captionTargetBox="[188,1396,217,1557]" captionTargetId="figure@7.[151,1436,195,1576]" captionTargetPageId="7" captionText="FIGURE 4. Eugnoriste florea sp. n. Male. Holotype. A. Part of hypopygium, ventral vieW. B. Head and thoraX. C. 4 th flagellomere. D. Wing. E. ApeX of fore tibia. F. Abdominal tergite 6, dorsal vieW." httpUri="https://zenodo.org/record/888605/files/figure.png" pageId="2" pageNumber="55">Fig. 4</figureCitation>
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)
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</keyLead>
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</paragraph>
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<paragraph id="8B8FF5154F1D6D58388AFE14FA41558A" blockId="2.[151,1437,151,750]" box="[151,1435,391,412]" pageId="2" pageNumber="55">
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<keyLead id="30C156204F1D6D58388AFE14FA41558A" box="[151,1435,391,412]" pageId="2" pageNumber="55">- Gonostylus broad, not taper towards the apex, apically more or less rounded....................................... 4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30C4EDB04F1D6D58388AFE35FA415600" pageId="2" pageNumber="55">
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<paragraph id="8B8FF5154F1D6D58388AFE35FA4155CF" blockId="2.[151,1437,151,750]" pageId="2" pageNumber="55">
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<keyLead id="30C156204F1D6D58388AFE35FA4155CF" pageId="2" pageNumber="55">
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4 Gonocoxites with a patch of 5–6 bristles in the middle of the inner side; ventral base of the hypopygium bare; 5th–7th abdominal tergites with distinct bristle patches; mouth part shorter than the head high...........
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<emphasis id="B94429074F1D6D583C29FE57FAE955CF" bold="true" box="[1076,1331,451,474]" pageId="2" pageNumber="55">
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<emphasis id="B94429074F1D6D583C29FE57FB2D55CC" bold="true" box="[1076,1271,452,474]" italics="true" pageId="2" pageNumber="55">
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Eug.
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<taxonomicName id="4C308E964F1D6D583C77FE57FB2D55CC" ID-CoL="9HW5Y" authority="Kauschke, 2017" authorityName="Kauschke" authorityYear="2017" box="[1130,1271,452,474]" class="Insecta" family="Sciaridae" genus="Eugnoriste" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="brachycostalis" status="sp. nov.">brachycostalis</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A277947C4F1D6D583CE1FE50FAE955CF" box="[1276,1331,451,473]" pageId="2" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
(
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<figureCitation id="130BE9904F1D6D583D22FE57FAA655CF" box="[1343,1404,452,473]" captionStart="FIGURE 1" captionStartId="3.[151,250,1991,2013]" captionTargetBox="[232,1345,213,1927]" captionTargetId="figure@3.[211,1364,195,1951]" captionTargetPageId="3" captionText="FIGURE 1. Eugnoriste brachycostalis sp. n. Male. Holotype. A. Gonostylus, ventral vieW. B. Ventral base of hypopygium." httpUri="https://zenodo.org/record/888599/files/figure.png" pageId="2" pageNumber="55">Figs 1</figureCitation>
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,
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<figureCitation id="130BE9904F1D6D583D95FE57FA4F55CF" box="[1416,1429,452,473]" captionStart="FIGURE 2" captionStartId="4.[151,250,1713,1735]" captionTargetBox="[155,1413,174,1681]" captionTargetId="figure@4.[151,1436,173,1713]" captionTargetPageId="4" captionText="FIGURE 2. Eugnoriste brachycostalis sp. n. Male. Holotype. A. Head and thoraX. B. Abdominal tergites 3 – 8. C. Antennal flagellomeres 3 – 4. D. ApeX of fore tibia. E. C / W-seCtion of the Wing apeX (draWing, 80 X)." httpUri="https://zenodo.org/record/888601/files/figure.png" pageId="2" pageNumber="55">2</figureCitation>
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)
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</keyLead>
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</paragraph>
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<paragraph id="8B8FF5154F1D6D58388AFE77FA415600" blockId="2.[151,1437,151,750]" pageId="2" pageNumber="55">
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||
<keyLead id="30C156204F1D6D58388AFE77FA415600" pageId="2" pageNumber="55">- Gonocoxites without distinct bristles in the middle of the inner side; ventral base of hypopygium with short hairs, or the 9th sternite bilobated...................................................................................... 5</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30C4EDB04F1D6D58388AFDB2FA415686" pageId="2" pageNumber="55">
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||
<paragraph id="8B8FF5154F1D6D58388AFDB2FA465645" blockId="2.[151,1437,151,750]" pageId="2" pageNumber="55">
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<keyLead id="30C156204F1D6D58388AFDB2FA465645" pageId="2" pageNumber="55">
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5 6th and 7th tergites with dense hairs on the distal borders; gonostylus is wider in the apical third than in the middle, concaved in the apical third only...............................................................
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<emphasis id="B94429074F1D6D583C8BFDADFA945642" bold="true" box="[1174,1358,574,596]" pageId="2" pageNumber="55">
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||
<emphasis id="B94429074F1D6D583C8BFDADFACB5642" bold="true" box="[1174,1297,574,596]" italics="true" pageId="2" pageNumber="55">
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Eug.
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<taxonomicName id="4C308E964F1D6D583CD6FDADFACB5642" ID-CoL="9HS8J" authority="Kauschke, 2017" authorityName="Kauschke" authorityYear="2017" box="[1227,1297,574,596]" class="Insecta" family="Sciaridae" genus="Eugnoriste" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="hirsuta" status="sp. nov.">hirsuta</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A277947C4F1D6D583D05FDADFA945642" box="[1304,1358,574,596]" pageId="2" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="130BE9904F1D6D583D46FDADFA4E5645" box="[1371,1428,574,595]" captionStart="FIGURE 5" captionStartId="8.[151,250,1909,1931]" captionTargetBox="[166,1423,633,1871]" captionTargetId="figure@8.[151,1436,630,1887]" captionTargetPageId="8" captionText="FIGURE 5. Eugnoriste hirsuta sp. n. Male. Holotype. A. Part of hypopygium, ventral vieW. B. Head With basal flagellomeres. C. Abdominal tergites 5 – 8." httpUri="https://zenodo.org/record/888607/files/figure.png" pageId="2" pageNumber="55">Fig. 5</figureCitation>
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)
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</keyLead>
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</paragraph>
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<paragraph id="8B8FF5154F1D6D58388AFDCDFA415686" blockId="2.[151,1437,151,750]" pageId="2" pageNumber="55">
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<keyLead id="30C156204F1D6D58388AFDCDFA415686" pageId="2" pageNumber="55">- 7th tergite with dense hairs on the distal border only; gonostylus wider in the middle than in the apical third, concaved in the middle or the basal half................................................................................. 6</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="30C4EDB04F1D6D58388AFD08FA4656FB" pageId="2" pageNumber="55">
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<paragraph id="8B8FF5154F1D6D58388AFD08FA4656A6" blockId="2.[151,1437,151,750]" box="[151,1436,664,689]" pageId="2" pageNumber="55">
|
||
<keyLead id="30C156204F1D6D58388AFD08FA4656A6" box="[151,1436,664,689]" pageId="2" pageNumber="55">
|
||
6 Gonostylus with two apical spines; 9th sternite shortly bilobated............
|
||
<emphasis id="B94429074F1D6D583BC8FD08FBDF56A7" bold="true" box="[981,1029,667,689]" italics="true" pageId="2" pageNumber="55">Eug.</emphasis>
|
||
<taxonomicName id="4C308E964F1D6D583C16FD08FA9456A6" ID-CoL="3C88B" authority="Mohrig, 2003" authorityName="Mohrig" authorityYear="2003" box="[1035,1358,667,689]" class="Insecta" family="Sciaridae" genus="Eugnoriste" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="villosoabdominalis">
|
||
<emphasis id="B94429074F1D6D583C16FD08FB1E56A7" bold="true" box="[1035,1220,667,689]" italics="true" pageId="2" pageNumber="55">villosoabdominalis</emphasis>
|
||
<bibRefCitation id="EFA188E44F1D6D583CD7FD08FA9456A6" author="Mohrig" box="[1226,1358,667,688]" pageId="2" pageNumber="55" refString="Mohrig, W. (2003) BlaCk fungus gnats of Central AmeriCa. Part I. (Diptera, SCiaridae). Beitrage zur Entomologie, 53 (1), 1 - 69." type="journal article" year="2003">Mohrig, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130BE9904F1D6D583D46FD08FA4E56A6" box="[1371,1428,667,688]" captionStart="FIGURE 8" captionStartId="12.[151,250,1865,1887]" captionTargetBox="[175,1397,458,1825]" captionTargetId="figure@12.[173,1409,433,1842]" captionTargetPageId="12" captionText="FIGURE 8. Eugnoriste villosoabdominalis Mohrig, 2003. Male. A. Part of hypopygium, ventral vieW. B. Gonostylus, ventral vieW (draWing, 400 X). C. Head. D. 4 th flagellomere. E. Abdominal segments 5 – 6. F. ApeX of fore tibia (draWing, 200 X). G. Tegmen (draWing, 200 X)." httpUri="https://zenodo.org/record/888613/files/figure.png" pageId="2" pageNumber="55">Fig. 8</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8B8FF5154F1D6D58388AFD28FA4656FB" blockId="2.[151,1437,151,750]" pageId="2" pageNumber="55">
|
||
<keyLead id="30C156204F1D6D58388AFD28FA4656FB" pageId="2" pageNumber="55">
|
||
- Gonostylus with two spines near to the middle; 9th sternite bilobated, with long lobes (half as long as gonocoxites)........................................................................................ ..
|
||
<emphasis id="B94429074F1D6D583C8AFD4BFA9456F8" bold="true" box="[1175,1358,728,750]" pageId="2" pageNumber="55">
|
||
<emphasis id="B94429074F1D6D583C8AFD4BFACB56F8" bold="true" box="[1175,1297,728,750]" italics="true" pageId="2" pageNumber="55">
|
||
Eug.
|
||
<taxonomicName id="4C308E964F1D6D583CD1FD4BFACB56F8" ID-CoL="9HY3R" authority="Kauschke, 2017" authorityName="Kauschke" authorityYear="2017" box="[1228,1297,728,750]" class="Insecta" family="Sciaridae" genus="Eugnoriste" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="55" phylum="Arthropoda" rank="species" species="ptilosis" status="sp. nov.">ptilosis</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A277947C4F1D6D583D05FD4BFA9456F8" box="[1304,1358,728,750]" pageId="2" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="130BE9904F1D6D583D46FD4BFA4F56FB" box="[1371,1429,728,749]" captionStart="FIGURE 7" captionStartId="11.[151,250,1881,1903]" captionTargetBox="[153,1435,207,1843]" captionTargetId="figure@11.[151,1436,195,1859]" captionTargetPageId="11" captionText="FIGURE 7. Eugnoriste ptilosis sp. n. Male. Holotype. A. Hypopygium, ventral vieW. B. Gonostylus, ventral vieW (draWing, 400 X). C. ApeX of gonostylus With Whiplash-hair and the tWo subapiCal spines. D. 4 th flagellomere. E. Head. F. Abdominal tergites 3 – 6. G. C / W - indeX of the Wing apeX (draWing, 80 X)." httpUri="https://zenodo.org/record/888611/files/figure.png" pageId="2" pageNumber="55">Fig. 7</figureCitation>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |