treatments-xml/data/03/98/6E/03986E15FFD54B140EB54EA4FDD8FA89.xml
2024-06-21 12:22:17 +02:00

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<document id="27B5706C5967A2B8D5B9963EA742FB1E" ID-DOI="10.26879/832" ID-ISSN="1094-8074" ID-Zenodo-Dep="10961470" ID-ZooBank="152941A8-052F-4134-9E98-ABC4CAC8F01B" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1712855045557" checkinUser="felipe" docAuthor="Alekseev, Vitalii I. &amp; Tomaszewska, Wioletta" docDate="2018" docId="03986E15FFD54B140EB54EA4FDD8FA89" docLanguage="en" docName="PalaeontolElectron.21.1.6A.pdf" docOrigin="Palaeontologia Electronica (6 A) 21 (1)" docSource="http://dx.doi.org/10.26879/832" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.3:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="3" docTitle="Trochoideus resinatissimus Alekseev &amp; Tomaszewska 2018, sp. nov." docType="treatment" docUuid="2456F93D-89AA-4850-8452-99A33CF5E6B4" docUuidSource="ZooBank" docVersion="3" lastPageNumber="17" masterDocId="FFA1166DFFDB4B040D174A35FFF2FFD6" masterDocTitle="New handsome fungus beetles (Coleoptera: Coccinelloidea: Anamorphidae, Endomychidae) from European amber of the Upper Eocene" masterLastPageNumber="23" masterPageNumber="1" pageNumber="15" updateTime="1713983817784" updateUser="carolina" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="4F686DC6B16846B70F2ADB726F3A099D">New handsome fungus beetles (Coleoptera: Coccinelloidea: Anamorphidae, Endomychidae) from European amber of the Upper Eocene</mods:title>
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<mods:namePart id="8CE186D4095FBB45C35AE98D2C3F933C">Alekseev, Vitalii I.</mods:namePart>
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<mods:namePart id="7D210EC1126D03DA72E07F105534BB12">Tomaszewska, Wioletta</mods:namePart>
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<mods:title id="4EB1D1BE2B9F0152F9666FB9DADC7341">Palaeontologia Electronica</mods:title>
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<mods:part id="1F1F35DAA1E49C19521D06CF67943D06">
<mods:date id="79701C8952C3CAD5E7A319D23DB51293">2018</mods:date>
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<mods:title id="155C88C1F0CB05F644880FBE0CB79F8E">6 A</mods:title>
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<mods:number id="C9B4D78FDAC5C92D76155CC55B2043C2">2018-12-31</mods:number>
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<mods:number id="2796A8455BBBE3CFE55AF4295B537E4E">21</mods:number>
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<mods:number id="23C5B75C02D1896A71BC120D8CD8FB7C">1</mods:number>
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<subSubSection id="C32B8C88FFD54B0A0EB54EA4FAB5FB7C" box="[930,1351,1169,1194]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8B8EDF03FFD54B0A0EB54EA4FAB5FB7C" blockId="14.[930,1351,1169,1226]" box="[930,1351,1169,1194]" pageId="14" pageNumber="15">
<heading id="D0C6686FFFD54B0A0EB54EA4FAB5FB7C" box="[930,1351,1169,1194]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="15" reason="7">
<taxonomicName id="4C31A480FFD54B0A0EB54EA4FB11FB7C" authority="Alekseev &amp; Tomaszewska, 2018" authorityName="Alekseev &amp; Tomaszewska" authorityYear="2018" box="[930,1251,1169,1194]" class="Insecta" family="Endomychidae" genus="Trochoideus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="resinatissimus" status="sp. nov.">
<emphasis id="B9450311FFD54B0A0EB54EA4FB11FB7C" box="[930,1251,1169,1194]" italics="true" pageId="14" pageNumber="15">Trochoideus resinatissimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276BE6AFFD54B0A09FC4EA4FAB5FB7C" box="[1259,1351,1169,1194]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32B8C88FFD54B0A09174E84FA8CFB39" pageId="14" pageNumber="15" type="description">
<paragraph id="8B8EDF03FFD54B0A09174E84FB1AFB1C" blockId="14.[930,1351,1169,1226]" box="[1024,1256,1201,1226]" pageId="14" pageNumber="15">
<figureCitation id="130AC386FFD54B0A09174E84FB78FB1C" box="[1024,1162,1201,1226]" captionStart="FIGURE 5" captionStartId="12.[194,285,1662,1685]" captionTargetBox="[195,1438,164,1644]" captionTargetId="figure-13@12.[195,1438,164,1644]" captionTargetPageId="12" captionText="FIGURE 5. Endomychids from Baltic and Bitterfeld ambers. 1-3, Laima andreei gen. et sp. nov. (No. 1222-5 [CCHH], holotype), habitus in lateral (1), dorsal (2), and ventral (3) views. 4-5, Trochoideus koenigsbergicus sp. nov. (No. 1506-1 [CCHH], holotype), habitus in dorsal view (4), and antenna in lateral view (5). 6-7, Trochoideus resinatissimus sp. nov., antenna (6) and maxillary palp (7) in lateral views." figureDoi="http://doi.org/10.5281/zenodo.10961481" httpUri="https://zenodo.org/record/10961481/files/figure.png" pageId="14" pageNumber="15">Figures 5.6</figureCitation>
-7, 6.1-2
</paragraph>
<paragraph id="8B8EDF03FFD54B0A0E5F4EEEFA8CFB39" blockId="14.[840,1406,1243,1263]" box="[840,1406,1243,1263]" pageId="14" pageNumber="15">
<emphasis id="B9450311FFD54B0A0E5F4EEEFA8CFB39" bold="true" box="[840,1406,1243,1263]" pageId="14" pageNumber="15">
zoobank.org/
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</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32B8C88FFD54B0A0E5F4F34FB94F98C" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8B8EDF03FFD54B0A0E5F4F34FB97FA2C" blockId="14.[840,1441,1280,1882]" pageId="14" pageNumber="15">
<emphasis id="B9450311FFD54B0A0E5F4F34FBF3FACC" bold="true" box="[840,1025,1281,1306]" pageId="14" pageNumber="15">Type material.</emphasis>
<typeStatus id="548A61A1FFD54B0A09074F34FB8BFACC" box="[1040,1145,1281,1306]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
No. 53-2 [
<collectionCode id="ED2047C6FFD54B0A081D4F35FAACFACC" box="[1290,1374,1280,1306]" pageId="14" pageNumber="15">CCHH</collectionCode>
], sex unknown. The beetle is included in a polished piece of transparent amber with an orange shade. The amber was not subjected to any additional fixation. Measurements of the amber piece are
<quantity id="4CC972E6FFD54B0A08954FB4FC86FA6C" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="mm" value="20.0">20 mm</quantity>
x
<quantity id="4CC972E6FFD54B0A0E864F94FC10FA6C" box="[913,994,1441,1466]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="15" unit="mm" value="17.0">17 mm</quantity>
x
<quantity id="4CC972E6FFD54B0A0EE84F94FBB4FA6C" box="[1023,1094,1441,1466]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="15" unit="mm" value="5.0">5 mm</quantity>
. The amber piece was possibly subjected to thermal and high-pressure processing in an autoclave.
</paragraph>
<paragraph id="8B8EDF03FFD54B0A0E5F4C34FAFCF9CC" blockId="14.[840,1441,1280,1882]" box="[840,1294,1537,1562]" pageId="14" pageNumber="15">
<emphasis id="B9450311FFD54B0A0E5F4C34FC29F9CC" bold="true" box="[840,987,1537,1562]" pageId="14" pageNumber="15">
<typeStatus id="548A61A1FFD54B0A0E5F4C34FC74F9CC" box="[840,902,1537,1562]" pageId="14" pageNumber="15">Type</typeStatus>
strata.
</emphasis>
Bitterfeld amber. Eocene.
</paragraph>
<paragraph id="8B8EDF03FFD54B0A0E5F4C14FB94F98C" blockId="14.[840,1441,1280,1882]" pageId="14" pageNumber="15">
<materialsCitation id="3B59D55EFFD54B0A0E5F4C14FB90F98C" country="Germany" location="Goitzsche" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Sachsen-Anhalt" typeStatus="holotype">
<emphasis id="B9450311FFD54B0A0E5F4C14FC07F9EC" bold="true" box="[840,1013,1569,1594]" pageId="14" pageNumber="15">
<typeStatus id="548A61A1FFD54B0A0E5F4C14FC76F9EC" box="[840,900,1569,1594]" pageId="14" pageNumber="15">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3269F93FFD54B0A09134C15FB86F9EC" box="[1028,1140,1568,1594]" name="Germany" pageId="14" pageNumber="15">Germany</collectingCountry>
,
<collectingRegion id="49F511E1FFD54B0A09914C15FAB5F9EC" box="[1158,1351,1568,1594]" country="Germany" name="Sachsen-Anhalt" pageId="14" pageNumber="15">Sachsen-Anhalt</collectingRegion>
,
<location id="8EEE89D8FFD54B0A084E4C15FC72F98C" LSID="urn:lsid:plazi:treatment:03986E15FFD54B140EB54EA4FDD8FA89:8EEE89D8FFD54B0A084E4C15FC72F98C" country="Germany" name="Goitzsche" pageId="14" pageNumber="15" stateProvince="Sachsen-Anhalt">Goitzsche</location>
(Bitterfeld-Wolfen)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B8C88FFD54B0A0E5F4C54FA7CF94C" pageId="14" pageNumber="15" type="etymology">
<paragraph id="8B8EDF03FFD54B0A0E5F4C54FA7CF94C" blockId="14.[840,1441,1280,1882]" pageId="14" pageNumber="15">
<emphasis id="B9450311FFD54B0A0E5F4C54FC24F9AC" bold="true" box="[840,982,1633,1658]" pageId="14" pageNumber="15">Etymology.</emphasis>
The epithet of the new species is Latin adjective
<emphasis id="B9450311FFD54B0A0EAE4CB4FBC5F94C" box="[953,1079,1665,1690]" italics="true" pageId="14" pageNumber="15">“resinatus”</emphasis>
(resinous) in the superlative.
</paragraph>
</subSubSection>
<subSubSection id="C32B8C88FFD54B0A0E5F4C94FC7AF88C" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8B8EDF03FFD54B0A0E5F4C94FC7AF88C" blockId="14.[840,1441,1280,1882]" pageId="14" pageNumber="15">
<emphasis id="B9450311FFD54B0A0E5F4C94FC3CF96C" bold="true" box="[840,974,1697,1722]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
<taxonomicName id="4C31A480FFD54B0A0EFE4C94FACFF96C" authorityName="Alekseev &amp; Tomaszewska" authorityYear="2018" box="[1001,1341,1697,1722]" class="Insecta" family="Endomychidae" genus="Trochoideus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="resinatissimus">
<emphasis id="B9450311FFD54B0A0EFE4C94FACFF96C" box="[1001,1341,1697,1722]" italics="true" pageId="14" pageNumber="15">Trochoideus resinatissimus</emphasis>
</taxonomicName>
differs from congeneric and probably coeval species described above from Baltic amber, based on antennal structure (strongly transverse antennomere 4 and egg-shaped antennomere 5) and larger body.
</paragraph>
</subSubSection>
<caption id="DF4E8F8BFFD44B0B0DD74CD5FF16F8BB" ID-DOI="http://doi.org/10.5281/zenodo.10961483" ID-Zenodo-Dep="10961483" httpUri="https://zenodo.org/record/10961483/files/figure.png" pageId="15" pageNumber="16" startId="15.[192,283,1760,1783]" targetBox="[221,1410,168,1727]" targetPageId="15" targetType="figure">
<paragraph id="8B8EDF03FFD44B0B0DD74CD5FF16F8BB" blockId="15.[192,1432,1760,1901]" pageId="15" pageNumber="16">
<emphasis id="B9450311FFD44B0B0DD74CD5FEC9F92E" bold="true" box="[192,315,1760,1784]" pageId="15" pageNumber="16">FIGURE 6.</emphasis>
Endomychids from Bitterfeld and Baltic ambers.
<emphasis id="B9450311FFD44B0B0E724CD4FC7CF92E" bold="true" box="[869,910,1761,1784]" pageId="15" pageNumber="16">1-2,</emphasis>
<taxonomicName id="4C31A480FFD44B0B0E8D4CD4FB32F92E" authority="Alekseev &amp; Tomaszewska, 2018" authorityName="Alekseev &amp; Tomaszewska" authorityYear="2018" box="[922,1216,1761,1784]" class="Insecta" family="Endomychidae" genus="Trochoideus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="resinatissimus" status="sp. nov.">
<emphasis id="B9450311FFD44B0B0E8D4CD4FB32F92E" box="[922,1216,1761,1784]" italics="true" pageId="15" pageNumber="16">Trochoideus resinatissimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276BE6AFFD44B0B09DB4CD4FAD6F92E" box="[1228,1316,1761,1784]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
(No. 53-2 [CCHH], holotype), habitus in dorsal view (
<emphasis id="B9450311FFD44B0B0F904CCBFD66F8C3" bold="true" box="[647,660,1790,1813]" pageId="15" pageNumber="16">1</emphasis>
), and forebody in ventral view (
<emphasis id="B9450311FFD44B0B0EF04CCBFC06F8C3" bold="true" box="[999,1012,1790,1813]" pageId="15" pageNumber="16">2</emphasis>
).
<emphasis id="B9450311FFD44B0B091D4CCBFBC1F8C3" bold="true" box="[1034,1075,1790,1813]" pageId="15" pageNumber="16">3-5,</emphasis>
<taxonomicName id="4C31A480FFD44B0B092C4CCBFE93F8E4" authority="Shockley and Alekseev, 2014" authorityName="Shockley and Alekseev" authorityYear="2014" class="Insecta" family="Endomychidae" genus="Glesirhanis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bercioi">
<emphasis id="B9450311FFD44B0B092C4CCBFAF3F8C3" box="[1083,1281,1790,1813]" italics="true" pageId="15" pageNumber="16">Glesirhanis bercioi</emphasis>
<bibRefCitation id="EFA0A2F2FFD44B0B081F4CCBFE93F8E4" author="Shockley, F. W. &amp; Alekseev, V. I." pageId="15" pageNumber="16" refId="ref13126" refString="Shockley, F. W. and Alekseev, V. I. 2014. Glesirhanis bercioi, a new genus and species from Baltic" type="book" year="2014">Shockley and Alekseev, 2014</bibRefCitation>
</taxonomicName>
(No. 1219-2 [CCHH]), habitus in dorsal (
<emphasis id="B9450311FFD44B0B0E064D2EFCECF8E4" bold="true" box="[785,798,1819,1842]" pageId="15" pageNumber="16">3</emphasis>
), ventro-lateral (
<emphasis id="B9450311FFD44B0B0ED94D2EFC29F8E4" bold="true" box="[974,987,1819,1842]" pageId="15" pageNumber="16">4</emphasis>
), and lateral (
<emphasis id="B9450311FFD44B0B09794D2EFB89F8E4" bold="true" box="[1134,1147,1819,1842]" pageId="15" pageNumber="16">5</emphasis>
) views.
<emphasis id="B9450311FFD44B0B09C44D2EFB0EF8E4" bold="true" box="[1235,1276,1819,1842]" pageId="15" pageNumber="16">6-7,</emphasis>
<taxonomicName id="4C31A480FFD44B0B08144D2EFDC6F886" authority="Motschulsky, 1856" authorityName="Motschulsky" authorityYear="1856" class="Insecta" family="Endomychidae" genus="Phymaphoroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="B9450311FFD44B0B08144D2EFE94F886" italics="true" pageId="15" pageNumber="16">Phymaphoroides antennatus</emphasis>
<bibRefCitation id="EFA0A2F2FFD44B0B0C794D0CFDC6F886" author="Motschulsky, V. I." box="[366,564,1849,1872]" pageId="15" pageNumber="16" pagination="21 - 38" refId="ref12434" refString="Motschulsky, V. I. 1856. Voyages. Lettres de M. de Motschulsky a M. Menetries. No. 3. New York le 15 Juillet 1654. Etudes Entomologiques, 5: 21 - 38." type="journal article" year="1856">Motschulsky, 1856</bibRefCitation>
</taxonomicName>
(No. 1219-3 [CCHH]), close-up of antenna (
<emphasis id="B9450311FFD44B0B09024D0CFBD0F886" bold="true" box="[1045,1058,1849,1872]" pageId="15" pageNumber="16">6</emphasis>
), and habitus in dorso-lateral view (
<emphasis id="B9450311FFD44B0B0DDF4D63FF27F8BB" bold="true" box="[200,213,1878,1901]" pageId="15" pageNumber="16">7</emphasis>
).
</paragraph>
</caption>
<subSubSection id="C32B8C88FFCB4B140DD74A93FCE1FC89" pageId="16" pageNumber="17" type="description">
<paragraph id="8B8EDF03FFCB4B140DD74A93FF06FEC9" blockId="16.[192,793,166,1375]" pageId="16" pageNumber="17">
<emphasis id="B9450311FFCB4B140DD74A93FEA8FF69" bold="true" box="[192,346,166,191]" pageId="16" pageNumber="17">Description.</emphasis>
Body elongate, convex, uniformly dark. Total body length
<quantity id="4CC972E6FFCB4B140CF74AF3FDB2FF09" box="[480,576,198,223]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="16" pageNumber="17" unit="mm" value="4.1">4.1 mm</quantity>
, elytral length
<quantity id="4CC972E6FFCB4B140FE44AF3FF02FF29" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="16" pageNumber="17" unit="mm" value="2.7">2.7 mm</quantity>
, width (at widest point in middle of elytra) 2.0 mm.
</paragraph>
<paragraph id="8B8EDF03FFCB4B140DED4B13FCE1FC89" blockId="16.[192,793,166,1375]" pageId="16" pageNumber="17">Compound eyes prominent and coarsely facetted. Antennae short, extending to middle of pronotum; composed of five antennomeres, claviform. Antennomere length proportions follow formula: 3.2: 1.2: 1.4: 1.0: 8.0. Penultimate antennomere strongly transverse (three times as wide as long). Terminal antennomere egg-shaped, rounded apically, 2.0 times as long as wide. Terminal labial palpomere almost spherical. Maxillary palp with terminal palpomere as long as penultimate one, oblong and acuminate. Anterior angles of pronotum widely-rounded. Metathorax coarsely and densely punctate. Distance between metacoxae wider than distance between mesocoxae. Elytra laterally with fine, irregular, slightly longitudinally oval punctures. Apex of elytra with distinct and fine parasutural stria. Legs with tarsal formula 4-4-4. Abdomen with six ventrites, and ventrite 1 longest.</paragraph>
</subSubSection>
<subSubSection id="C32B8C88FFCB4B140DD74953FDD8FA89" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8B8EDF03FFCB4B140DD74953FEAEFC09" blockId="16.[192,793,166,1375]" pageId="16" pageNumber="17">
<emphasis id="B9450311FFCB4B140DD74953FECAFCA9" bold="true" box="[192,312,870,895]" pageId="16" pageNumber="17">Remarks.</emphasis>
The specimen is dorsally covered with dense milky cover, so the characters of upper surface are not visible (except lateral and apical parts of the elytra).
</paragraph>
<paragraph id="8B8EDF03FFCB4B140DD749D3FDD8FA89" blockId="16.[192,793,166,1375]" pageId="16" pageNumber="17">
<emphasis id="B9450311FFCB4B140DD749D3FEF0FC29" bold="true" box="[192,258,998,1023]" pageId="16" pageNumber="17">Note.</emphasis>
According to the note given under the description of
<taxonomicName id="4C31A480FFCB4B140CB54E33FCEAFBC9" authorityName="Alekseev &amp; Tomaszewska" authorityYear="2018" box="[418,792,1030,1055]" class="Insecta" family="Endomychidae" genus="Trochoideus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="koenigsbergicus">
<emphasis id="B9450311FFCB4B140CB54E33FCEAFBC9" box="[418,792,1030,1055]" italics="true" pageId="16" pageNumber="17">Trochoideus koenigsbergicus</emphasis>
</taxonomicName>
(above) from Baltic amber, the new species from Bitterfeld amber could theoretically be the male of the same, widely distributed Eocene species. The thickened antennomere 4 of
<taxonomicName id="4C31A480FFCB4B140F0D4EB3FD10FB49" authorityName="Alekseev &amp; Tomaszewska" authorityYear="2018" box="[538,738,1158,1183]" class="Insecta" family="Endomychidae" genus="Trochoideus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="resinatissimus">
<emphasis id="B9450311FFCB4B140F0D4EB3FD10FB49" box="[538,738,1158,1183]" italics="true" pageId="16" pageNumber="17">T. resinatissimus</emphasis>
</taxonomicName>
and different form of other antennomeres could be interpreted as sexual characters only. However, the question about reliable specific differences between specimens from Baltic and Bitterfeld amber would require further study of additional fossil material from both deposits.
</paragraph>
</subSubSection>
</treatment>
</document>