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<document id="0B55A741F9126CB8A6CB7928F4EFD9FF" ID-CLB-Dataset="8744" ID-DOI="10.11646/zootaxa.4838.2.1" ID-GBIF-Dataset="761fcec7-c79d-4e80-bb2b-23cc9e78e0b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403783" ID-ZooBank="0AD43D95-7508-466C-980F-CBEF38CA1198" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609352386978" checkinUser="plazi" docAuthor="Gompel, Nicolas" docDate="2020" docId="039187D8FFD1BB09FF1880B9FAEFFE76" docLanguage="en" docName="zootaxa.4838.2.1.pdf" docOrigin="Zootaxa 4838 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zarcosia sumatrensis Gompel 2020, n. sp." docType="treatment" docUuid="0AF41C62-6C5E-40C6-8FB6-0F5B93D10D04" docUuidSource="ZooBank" docVersion="8" lastPageNumber="175" masterDocId="FFA8FFA0FFC6BB11FF8F844AFF9DFFFF" masterDocTitle="Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species" masterLastPageNumber="178" masterPageNumber="151" pageNumber="174" updateTime="1698850145988" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B2ACFD8516A29B970A88FCE1BB562890">Synopsis of the Asian species in the genus Zarcosia Collado &amp; Alonso-Zarazaga 1996 with new combinations and descriptions of fourteen new species</mods:title>
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<mods:namePart id="64B5D8053C5B1CAC94DFD53131661B22">Gompel, Nicolas</mods:namePart>
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<mods:date id="A059FADA460AAC9FB95C6A2881963497">2020</mods:date>
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<mods:number id="55A00520482C706C136F9272FBA22BF4">2020-08-26</mods:number>
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<mods:identifier id="72E22EFE354B3148DCD599B640AEDF4A" type="DOI">10.11646/zootaxa.4838.2.1</mods:identifier>
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<treatment id="039187D8FFD1BB09FF1880B9FAEFFE76" ID-DOI="http://doi.org/10.5281/zenodo.4404275" ID-GBIF-Taxon="177012511" ID-Zenodo-Dep="4404275" LSID="urn:lsid:zoobank.org:act:0AF41C62-6C5E-40C6-8FB6-0F5B93D10D04" httpUri="http://treatment.plazi.org/id/039187D8FFD1BB09FF1880B9FAEFFE76" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174">
<subSubSection id="C3226545FFD1BB06FF1880B9FE7EFAF2" box="[151,483,1267,1293]" pageId="23" pageNumber="174" type="nomenclature">
<paragraph id="8B8736CEFFD1BB06FF1880B9FE7EFAF2" blockId="23.[151,483,1267,1330]" box="[151,483,1267,1293]" pageId="23" pageNumber="174">
<heading id="D0CF81A2FFD1BB06FF1880B9FE7EFAF2" bold="true" box="[151,483,1267,1293]" fontSize="11" level="1" pageId="23" pageNumber="174" reason="1">
<emphasis id="B94CEADCFFD1BB06FF1880B9FE7EFAF2" bold="true" box="[151,483,1267,1293]" pageId="23" pageNumber="174">
<taxonomicName id="4C384D4DFFD1BB06FF1880B9FE04FAF2" authority="Gompel, 2020" authorityName="Gompel" authorityYear="2020" box="[151,409,1267,1293]" class="Insecta" family="Aderidae" genus="Zarcosia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="sumatrensis" status="sp. nov.">
<emphasis id="B94CEADCFFD1BB06FF1880B9FE04FAF2" bold="true" box="[151,409,1267,1293]" italics="true" pageId="23" pageNumber="174">Zarcosia sumatrensis</emphasis>
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<taxonomicNameLabel id="A27F57A7FFD1BB06FE2E80B9FE7EFAF2" box="[417,483,1267,1293]" pageId="23" pageNumber="174" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C3226545FFD1BB06FF18815DFE53FACD" box="[151,462,1302,1330]" pageId="23" pageNumber="174" type="description">
<paragraph id="8B8736CEFFD1BB06FF18815DFE53FACD" blockId="23.[151,483,1267,1330]" box="[151,462,1302,1330]" pageId="23" pageNumber="174">
(
<figureCitation id="13032A4BFFD1BB06FF10815DFE96FACE" box="[159,267,1303,1329]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="23" pageNumber="174">Figures 1</figureCitation>
;
<figureCitation id="13032A4BFFD1BB06FE97815DFED1FACE" box="[280,332,1303,1329]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="23" pageNumber="174">
<quantity id="4CC09B2BFFD1BB06FE97815DFEABFACE" box="[280,310,1303,1329]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="23" pageNumber="174" unit="g" value="3.0">3g</quantity>
,h
</figureCitation>
;
<figureCitation id="13032A4BFFD1BB06FED7815DFEE5FACE" box="[344,376,1303,1329]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="23" pageNumber="174">4p</figureCitation>
;
<figureCitation id="13032A4BFFD1BB06FE0C815CFE3FFACE" box="[387,418,1302,1329]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="23" pageNumber="174">5o</figureCitation>
;
<figureCitation id="13032A4BFFD1BB06FE21815DFE5BFACD" box="[430,454,1303,1330]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="23" pageNumber="174">6f</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFD1BB06FF188115FCFCFA86" box="[151,865,1375,1401]" pageId="23" pageNumber="174" type="etymology">
<paragraph id="8B8736CEFFD1BB06FF188115FCFCFA86" blockId="23.[151,1437,1375,1942]" box="[151,865,1375,1401]" pageId="23" pageNumber="174">
<emphasis id="B94CEADCFFD1BB06FF188115FE83FA86" bold="true" box="[151,286,1375,1401]" pageId="23" pageNumber="174">Etymology.</emphasis>
The species’ name refers to its geographical origin.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFD1BB06FF4881C9FB42FA62" box="[199,1247,1410,1437]" pageId="23" pageNumber="174" type="materials_examined">
<paragraph id="8B8736CEFFD1BB06FF4881C9FB42FA62" blockId="23.[151,1437,1375,1942]" box="[199,1247,1410,1437]" pageId="23" pageNumber="174">
<materialsCitation id="3B503C93FFD1BB06FF4881C9FB46FA62" ID-GBIF-Occurrence="3020160374" box="[199,1243,1410,1437]" country="Indonesia" latitude="4.233333" location="Goea Limau Moangkaer" longLatPrecision="1306" longitude="98.166664" pageId="23" pageNumber="174" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<emphasis id="B94CEADCFFD1BB06FF4881C9FEFBFA63" bold="true" box="[199,358,1410,1437]" pageId="23" pageNumber="174">
<typeStatus id="5483886CFFD1BB06FF4881C9FE9FFA62" box="[199,258,1411,1437]" pageId="23" pageNumber="174">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32F765EFFD1BB06FEE381C9FDF5FA62" box="[364,616,1411,1437]" name="Indonesia" pageId="23" pageNumber="174">Republic of Indonesia</collectingCountry>
:
<collectingRegion id="49FCF82CFFD1BB06FDFD81C9FD4FFA62" box="[626,722,1411,1437]" country="Indonesia" name="Sumatera Utara" pageId="23" pageNumber="174">Sumatra</collectingRegion>
,
<location id="8EE76015FFD1BB06FD5181C9FC6DFA62" LSID="urn:lsid:plazi:treatment:039187D8FFD1BB09FF1880B9FAEFFE76:8EE76015FFD1BB06FD5181C9FC6DFA62" box="[734,1008,1411,1437]" country="Indonesia" latitude="4.233333" longLatPrecision="1306" longitude="98.166664" name="Goea Limau Moangkaer" pageId="23" pageNumber="174" stateProvince="Sumatera Utara">Goea Limau Moangkaer</location>
[
<geoCoordinate id="EE0C5009FFD1BB06FC7181C9FBF8FA62" box="[1022,1125,1411,1437]" degrees="04" direction="north" minutes="14" orientation="latitude" pageId="23" pageNumber="174" precision="925" value="4.233333">04°14’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFD1BB06FBE381C9FB49FA62" box="[1132,1236,1410,1437]" degrees="98" direction="east" minutes="10" orientation="longitude" pageId="23" pageNumber="174" precision="925" value="98.166664">98°10’ E</geoCoordinate>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFD1BB06FF4881ECFCD4FA3E" box="[199,841,1446,1473]" pageId="23" pageNumber="174" type="distribution">
<paragraph id="8B8736CEFFD1BB06FF4881ECFCD4FA3E" blockId="23.[151,1437,1375,1942]" box="[199,841,1446,1473]" pageId="23" pageNumber="174">
<emphasis id="B94CEADCFFD1BB06FF4881ECFEFCFA3F" bold="true" box="[199,353,1446,1472]" pageId="23" pageNumber="174">Distribution.</emphasis>
<collectingCountry id="F32F765EFFD1BB06FEE881EDFDFEFA3E" box="[359,611,1447,1473]" name="Indonesia" pageId="23" pageNumber="174">Republic of Indonesia</collectingCountry>
:
<collectingRegion id="49FCF82CFFD1BB06FDE181EDFD56FA3E" box="[622,715,1447,1473]" country="Indonesia" name="Sumatera Utara" pageId="23" pageNumber="174">Sumatra</collectingRegion>
(
<figureCitation id="13032A4BFFD1BB06FD5681EDFCA5FA3E" box="[729,824,1447,1473]" captionStart="FIGURE 1" captionStartId="3.[151,250,1474,1498]" captionTargetBox="[158,1432,565,1446]" captionTargetId="figure-172@3.[151,1437,561,1451]" captionTargetPageId="3" captionText="FIGURE 1. Distribution of Zarcosia species across South Asia. New taxa are indicated in boldface. For Z. lemairei n. sp., the broader geographical range is highlighted with a red area with a dotted line." figureDoi="http://doi.org/10.5281/zenodo.4403787" httpUri="https://zenodo.org/record/4403787/files/figure.png" pageId="23" pageNumber="174">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFD1BB09FF488180FB47FF2A" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174" type="description">
<paragraph id="8B8736CEFFD1BB06FF488180FB51F9F6" blockId="23.[151,1437,1375,1942]" pageId="23" pageNumber="174">
<emphasis id="B94CEADCFFD1BB06FF488180FEC4FA1B" bold="true" box="[199,345,1482,1508]" pageId="23" pageNumber="174">Description.</emphasis>
(
<figureCitation id="13032A4BFFD1BB06FEE78181FE72FA19" box="[360,495,1483,1510]" captionStart="FIGURE 3" captionStartId="20.[151,250,188,212]" captionTargetId="figure-15@19.[244,1343,179,2016]" captionTargetPageId="19" captionText="FIGURE 3. Habitus of new Zarcosia species. a, Z. lemairei n. sp., ♂ holotype, from Laos. b, Z. lemairei n. sp., ♀ paratype, from Laos. c, Z. ilonae n. sp., ♂ holotype, from Thailand. d, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. e, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). f, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). g, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). h, Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra). i, Z. barclayi n. sp., ♂ holotype, from Malaysia. j, Z. barclayi n. sp., ♀ paratype, from Malaysia. k, Z. schawalleri n. sp., ♂ holotype, from Malaysia. l, Z. schawalleri n. sp., ♀ paratype, from Malaysia. m, Z. bedosae n. sp., ♂ holotype, from Laos. n, Z. sinuata n. sp., ♂ holotype, from Thailand. o, Z. sinuata n. sp., ♀ paratype, from Thailand. p, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. q, Z. dentatifemur (Pic, 1912), ♂ holotype, from Sri Lanka, after remounting (same specimen as in Fig. 2o). r, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. s, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. s, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4453388" httpUri="https://zenodo.org/record/4453388/files/figure.png" pageId="23" pageNumber="174">
Figure
<quantity id="4CC09B2BFFD1BB06FE348181FE44FA19" box="[443,473,1483,1510]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="23" pageNumber="174" unit="g" value="3.0">3g</quantity>
,h
</figureCitation>
). Body length.
<specimenCount id="9D3EFD47FFD1BB06FD128181FD2FFA1A" box="[669,690,1483,1509]" pageId="23" pageNumber="174" type="male">♂</specimenCount>
<typeStatus id="5483886CFFD1BB06FD358181FCBDFA1A" box="[698,800,1483,1509]" pageId="23" pageNumber="174" type="holotype">holotype</typeStatus>
: 3.00 mm;
<specimenCount id="9D3EFD47FFD1BB06FC2F8181FC2AFA1A" box="[928,951,1483,1509]" pageId="23" pageNumber="174" type="female">♀</specimenCount>
: 2.65±
<quantity id="4CC09B2BFFD1BB06FB8B8180FBF7FA1A" box="[1028,1130,1482,1509]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="23" pageNumber="174" unit="mm" value="0.19">0.19 mm</quantity>
(n=7). Dark brown, darker anteriorly, gracile, covered with very short, pulverulent pubescence. Appendages orange-brown.
</paragraph>
<paragraph id="8B8736CEFFD1BB06FF488259FA06F9AE" blockId="23.[151,1437,1375,1942]" pageId="23" pageNumber="174">
<emphasis id="B94CEADCFFD1BB06FF488259FE99F9D2" box="[199,260,1555,1581]" italics="true" pageId="23" pageNumber="174">Head</emphasis>
. Small, dark, barely wider than pronotum. Palpi orange, last article securiform. Integument, finely and densely punctuated, lightly rugose. Occiput lined with a very fine fringe of dense and short silver hairs posteriorly.
</paragraph>
<paragraph id="8B8736CEFFD1BB06FF488211FE3DF966" blockId="23.[151,1437,1375,1942]" pageId="23" pageNumber="174">
<emphasis id="B94CEADCFFD1BB06FF488211FEACF98A" box="[199,305,1627,1653]" italics="true" pageId="23" pageNumber="174">Antennae</emphasis>
. A third longer than elytra in
<specimenCount id="9D3EFD47FFD1BB06FDFC8211FD17F98A" box="[627,650,1627,1653]" pageId="23" pageNumber="174" type="male">♂</specimenCount>
, as long as elytra in
<specimenCount id="9D3EFD47FFD1BB06FCE78211FCE0F98A" box="[872,893,1627,1653]" pageId="23" pageNumber="174" type="female">♀</specimenCount>
, orange-brown, regularly filiform with antenommeres 3–10 cylindrical.
</paragraph>
<paragraph id="8B8736CEFFD1BB06FF4882E9FB2FF91E" blockId="23.[151,1437,1375,1942]" pageId="23" pageNumber="174">
<emphasis id="B94CEADCFFD1BB06FF4882E9FEABF943" box="[199,310,1699,1724]" italics="true" pageId="23" pageNumber="174">Pronotum</emphasis>
(
<figureCitation id="13032A4BFFD1BB06FECB82E9FE2DF941" box="[324,432,1699,1726]" captionStart="FIGURE 6" captionStartId="26.[151,250,1631,1655]" captionTargetBox="[158,1431,530,1605]" captionTargetId="figure-126@26.[151,1436,525,1607]" captionTargetPageId="26" captionText="FIGURE 6. Thoraces of selected Zarcosia species in dorsal view. a, Z. lemairei n. sp., ♂ holotype, from Laos (left) and Z. lemairei n. sp., ♀ paratype, from Laos (right). b, Z. ilonae n. sp., ♂ holotype, from Thailand. c, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam. d, Z. weigeli n. sp., ♂ holotype, from China (Yunnan). e, Z. deharvengi n. sp., ♂ holotype, from Republic of Indonesia (Sumatra). f, Z. sumatrensis n. sp., ♂ holotype, from Republic of Indonesia (Sumatra) (left) and Z. sumatrensis n. sp., ♀ paratype, from Republic of Indonesia (Sumatra) (right). g, Z. barclayi n. sp., ♂ holotype, from Malaysia (left) and Z. barclayi n. sp., ♀ paratype, from Malaysia (right). h, Z. schawalleri n. sp., ♂ holotype, from Malaysia (left) and Z. schawalleri n. sp., ♀ paratype, from Malaysia (right). i, Z. bedosae n. sp., ♂ holotype, from Laos. j, Z. sinuata n. sp., ♂ holotype, from Thailand (left) and Z. sinuata n. sp., ♀ paratype, from Thailand (right). k, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka. l, Z. intermedia n. sp., ♂ paratype, from Sri Lanka. m, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka. n, Z. spinifemur n. sp., ♂ holotype, from Sri Lanka. o, Z. brunnea n. sp., ♂ paratype, from Sri Lanka. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403795" httpUri="https://zenodo.org/record/4403795/files/figure.png" pageId="23" pageNumber="174">Figure 6f</figureCitation>
). Round and convex, slightly elongated in
<specimenCount id="9D3EFD47FFD1BB06FC0582E9FC3CF942" box="[906,929,1699,1725]" pageId="23" pageNumber="174" type="male">♂</specimenCount>
, as long as wide in
<specimenCount id="9D3EFD47FFD1BB06FBF682E9FB12F942" box="[1145,1167,1699,1725]" pageId="23" pageNumber="174" type="female">♀</specimenCount>
. Sides regularly curved. Surface mat, finely and densely punctuated, covered with dense, pruinose double pubescence.
</paragraph>
<paragraph id="8B8736CEFFD1BB06FF4882A1FC12F8B2" blockId="23.[151,1437,1375,1942]" pageId="23" pageNumber="174">
<emphasis id="B94CEADCFFD1BB06FF4882A1FE90F8FA" box="[199,269,1771,1797]" italics="true" pageId="23" pageNumber="174">Elytra</emphasis>
. Length:
<specimenCount id="9D3EFD47FFD1BB06FEF482A1FE0DF8FA" box="[379,400,1771,1797]" pageId="23" pageNumber="174" type="male">♂</specimenCount>
<typeStatus id="5483886CFFD1BB06FE1882A1FE60F8FA" box="[407,509,1771,1797]" pageId="23" pageNumber="174" type="holotype">holotype</typeStatus>
: 2.00 mm;
<specimenCount id="9D3EFD47FFD1BB06FDF282A1FD09F8FA" box="[637,660,1771,1797]" pageId="23" pageNumber="174" type="female">♀</specimenCount>
: 1.95±
<quantity id="4CC09B2BFFD1BB06FD6F82A0FCDBF8FA" box="[736,838,1770,1797]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="174" unit="mm" value="0.1">0.10 mm</quantity>
(n=7). Punctation fine and shallow, dense (space between points as wide as the points themselves). Disc depressed. Pubescence dense and short, whitish, comprised of short hairs borne from the points and interspersed pulverulent hairs.
</paragraph>
<paragraph id="8B8736CEFFD1BB06FF48831DFB3DF869" blockId="23.[151,1437,1375,1942]" pageId="23" pageNumber="174">
<emphasis id="B94CEADCFFD1BB06FF48831DFF61F88F" box="[199,252,1879,1904]" italics="true" pageId="23" pageNumber="174">Legs</emphasis>
. Particularly elongated. Foretibia thickened in a smooth tooth on their inner side (
<figureCitation id="13032A4BFFD1BB06FB0E831DFB6CF88E" box="[1153,1265,1878,1905]" captionStart="FIGURE 5" captionStartId="25.[151,250,1602,1626]" captionTargetBox="[170,1408,188,1563]" captionTargetId="figure-15@25.[169,1419,181,1578]" captionTargetPageId="25" captionText="FIGURE 5. Morphological details of selected Zarcosia species. a–d, Z. lemairei n. sp., ♂ holotype, from Laos: head in lateral (a), frontal (b) and dorsal (c) views; mesothoracic (left) and metathoracic (right) tarsae (d). e–g, Z. palliditarsis (Pic, 1912), ♂, from Sri Lanka: head in lateral (e), frontal (f) and dorsal (g) views. h, Z. srilankaensis n. sp., ♂ paratype, from Sri Lanka: tuft of pubescence at elytral apex. i, Z. weigeli n. sp., ♂ holotype, from China (Yunnan): elytral pubescence on disc. j, Z. gerstmeieri n. sp., ♂ holotype, from Vietnam: elytral pubescence on disc. k–t: male forelegs of Z. ilonae n. sp., holotype, from Thailand (k); Z. deharvengi n. sp., holotype, from Republic of Indonesia (Sumatra) (l); Z. schawalleri n. sp., holotype, from Malaysia (m); Z. sinuata n. sp., holotype, from Thailand (n); Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra) (o); Z. srilankaensis n. sp., paratype, from Sri Lanka (p); Z. intermedia n. sp., paratype, from Sri Lanka (q); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (r); Z. brunnea n. sp., paratype, from Sri Lanka (s); Z. weigeli n. sp., holotype, from China (Yunnan) (t). u–x: male mesothoracic legs of Z. cephalica (Pic, 1901) from Republic of Indonesia (u); Z. gerstmeieri n. sp., holotype, from Vietnam (v); Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka (w); Z. spinifemur n. sp., holotype, from Sri Lanka (x). Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4403791" httpUri="https://zenodo.org/record/4403791/files/figure.png" pageId="23" pageNumber="174">Figure 5o</figureCitation>
). Mesothoracic legs not observed (a single male devoid of mesothoracic legs was present in the
<typeStatus id="5483886CFFD1BB06FB968331FBD4F86A" box="[1049,1097,1915,1941]" pageId="23" pageNumber="174">type</typeStatus>
series).
</paragraph>
<paragraph id="8B8736CEFFDEBB09FF4884DDFB47FF2A" blockId="24.[151,1437,151,393]" pageId="24" pageNumber="175">
<emphasis id="B94CEADCFFDEBB09FF4884DDFEA9FF4E" box="[199,308,151,177]" italics="true" pageId="24" pageNumber="175">Aedeagus</emphasis>
(
<figureCitation id="13032A4BFFDEBB09FEC984DDFE21FF4E" box="[326,444,151,177]" captionStart="FIGURE 4" captionStartId="22.[151,250,186,210]" captionTargetId="figure-15@21.[275,1312,181,1998]" captionTargetPageId="21" captionText="FIGURE 4. Aedeagus of selected Zarcosia species. a, Z. barclayi n. sp., holotype, from Malaysia, lateral and dorsal views. b, Z. bedosae n. sp., holotype, from Laos, lateral and dorsal views. c, Z. bipartita (Pic, 1901) from India, lateral view. d, Z. cephalica (Pic, 1901) from Republic of Indonesia, lateral view. e, Z. deharvengi n. sp., holotype, from Laos, lateral and dorsal views. f, Z. gerstmeieri n. sp., holotype, from Vietnam, lateral and dorsal views. g, Z. ilonae n. sp., holotype, from Thailand, lateral and dorsal views. h, Z. kempi (Blair, 1924), holotype, from India, lateral and dorsal views. i, Z. lemairei n. sp., holotype, from Laos, lateral and dorsal views. j, Z. obscuricornis (Pic, 1927) from Vietnam, lateral view. k, Z. palliditarsis (Pic, 1912) from Sri Lanka, lateral view. l, Z. picina (Fairmaire, 1893), lectotype, from Vietnam, lateral view. m, Z. rubrobasalis (Pic, 1921) (slight uncertainty on the identity of this specimen) from Singapore, lateral view. n, Z. schawalleri n. sp., holotype, from Malaysia, lateral and dorsal views. o, Z. sinuata n. sp., holotype, from Thailand, lateral and dorsal views. p, Z. sumatrensis n. sp., holotype, from Republic of Indonesia (Sumatra), lateral and dorsal views. q, Z. troglodytes (Champion, 1916), holotype, from Malaysia, lateral view. r, Z. weigeli n. sp., holotype, from China (Yunnan), lateral and dorsal views. s, Z. spinifemur n. sp., holotype, from Sri Lanka, lateral and dorsal views. t, Z. dentatifemur (Pic, 1912), holotype, from Sri Lanka, lateral and dorsal views. u, Z. srilankaensis n. sp., paratype, from Sri Lanka, lateral and dorsal views. v, Z. intermedia n. sp., paratype, from Sri Lanka, lateral and dorsal views. w, Z. brunnea n. sp., paratype, from Sri Lanka, lateral and dorsal views. Scale bars: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4453392" httpUri="https://zenodo.org/record/4453392/files/figure.png" pageId="24" pageNumber="175">Figure 4p</figureCitation>
). Sides subparallel in dorsal view, apex tapered in apical fifth, ending with a button. Strongly bent in lateral view, apex with a weak upward deflection. Appendages visible near apex.
</paragraph>
</subSubSection>
<subSubSection id="C3226545FFDEBB09FF488495FAEFFE76" pageId="24" pageNumber="175" type="materials_examined">
<paragraph id="8B8736CEFFDEBB09FF488495FAA3FE9A" blockId="24.[151,1437,151,393]" pageId="24" pageNumber="175">
<emphasis id="B94CEADCFFDEBB09FF488495FEEAFF07" bold="true" box="[199,375,222,249]" pageId="24" pageNumber="175">Type material.</emphasis>
<materialsCitation id="3B503C93FFDEBB09FEF08495FD32FEE2" ID-GBIF-Occurrence="3020160366" collectingDate="1932-03-20" collectionCode="BMNH" collectorName="J. C. van der Meer Mohr" country="Indonesia" latitude="4.233333" location="Goea Limau Moangkaer" longLatPrecision="1306" longitude="98.166664" pageId="24" pageNumber="175" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<typeStatus id="5483886CFFDEBB09FEF08495FE7AFF06" box="[383,487,223,249]" pageId="24" pageNumber="175" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D3EFD47FFDEBB09FE768495FD8DFF06" box="[505,528,223,249]" pageId="24" pageNumber="175" type="male">♂</specimenCount>
,
<collectionCode id="ED29AE0BFFDEBB09FD918495FDEAFF06" box="[542,631,223,249]" pageId="24" pageNumber="175">BMNH</collectionCode>
).
<emphasis id="B94CEADCFFDEBB09FD048494FC09FF07" bold="true" box="[651,916,222,248]" pageId="24" pageNumber="175">
<collectingCountry id="F32F765EFFDEBB09FD048494FC09FF07" box="[651,916,222,248]" name="Indonesia" pageId="24" pageNumber="175">Republic of Indonesia</collectingCountry>
</emphasis>
:
<collectingRegion id="49FCF82CFFDEBB09FC2B8495FB99FF06" box="[932,1028,223,249]" country="Indonesia" name="Sumatera Utara" pageId="24" pageNumber="175">Sumatra</collectingRegion>
,
<location id="8EE76015FFDEBB09FB9E8495FAB5FF06" LSID="urn:lsid:plazi:treatment:039187D8FFD1BB09FF1880B9FAEFFE76:8EE76015FFDEBB09FB9E8495FAB5FF06" box="[1041,1320,223,249]" country="Indonesia" latitude="4.233333" longLatPrecision="1306" longitude="98.166664" name="Goea Limau Moangkaer" pageId="24" pageNumber="175" stateProvince="Sumatera Utara">Goea Limau Moangkaer</location>
,
<geoCoordinate id="EE0C5009FFDEBB09FABA8495FA01FF06" box="[1333,1436,223,249]" degrees="04" direction="north" minutes="14" orientation="latitude" pageId="24" pageNumber="175" precision="925" value="4.233333">04°14’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFDEBB09FF188549FF64FEE2" box="[151,249,259,285]" degrees="98" direction="east" minutes="10" orientation="longitude" pageId="24" pageNumber="175" precision="925" value="98.166664">98°10’ E</geoCoordinate>
(
<collectorName id="26CD5318FFDEBB09FE858549FD83FEE1" box="[266,542,259,286]" pageId="24" pageNumber="175">J. C. van der Meer Mohr</collectorName>
)
<date id="FF86100EFFDEBB09FDA28549FD32FEE2" box="[557,687,259,285]" pageId="24" pageNumber="175" value="1932-03-20">
<collectingDate id="EFC2E9E6FFDEBB09FDA28549FD32FEE2" box="[557,687,259,285]" pageId="24" pageNumber="175" value="1932-03-20">20.III.1932</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B503C93FFDEBB09FD368549FBE9FEBE" ID-GBIF-Occurrence="3020160459" collectingDate="1932-03-20" collectionCode="BMNH" collectorName="J. C. van der Meer Mohr" country="Indonesia" latitude="4.233333" location="Goea Limau Moangkaer" longLatPrecision="1306" longitude="98.166664" pageId="24" pageNumber="175" specimenCount="3" specimenCount-female="2" stateProvince="Sumatera Utara" typeStatus="paratype">
<typeStatus id="5483886CFFDEBB09FD368549FCB7FEE2" box="[697,810,259,285]" pageId="24" pageNumber="175" type="paratype">Paratypes</typeStatus>
.
<emphasis id="B94CEADCFFDEBB09FCBC8548FBA5FEE3" bold="true" box="[819,1080,258,284]" pageId="24" pageNumber="175">
<collectingCountry id="F32F765EFFDEBB09FCBC8548FBA5FEE3" box="[819,1080,258,284]" name="Indonesia" pageId="24" pageNumber="175">Republic of Indonesia</collectingCountry>
</emphasis>
:
<collectingRegion id="49FCF82CFFDEBB09FBC88549FB3AFEE2" box="[1095,1191,259,285]" country="Indonesia" name="Sumatera Utara" pageId="24" pageNumber="175">Sumatra</collectingRegion>
,
<location id="8EE76015FFDEBB09FB3D8549FF54FEBE" LSID="urn:lsid:plazi:treatment:039187D8FFD1BB09FF1880B9FAEFFE76:8EE76015FFDEBB09FB3D8549FF54FEBE" country="Indonesia" latitude="4.233333" longLatPrecision="1306" longitude="98.166664" name="Goea Limau Moangkaer" pageId="24" pageNumber="175" stateProvince="Sumatera Utara">Goea Limau Moangkaer</location>
,
<geoCoordinate id="EE0C5009FFDEBB09FF5B856DFEA7FEBE" box="[212,314,295,321]" degrees="04" direction="north" minutes="14" orientation="latitude" pageId="24" pageNumber="175" precision="925" value="4.233333">04°14’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFDEBB09FECE856DFE39FEBE" box="[321,420,295,321]" degrees="98" direction="east" minutes="10" orientation="longitude" pageId="24" pageNumber="175" precision="925" value="98.166664">98°10’ E</geoCoordinate>
(
<collectorName id="26CD5318FFDEBB09FE3A856DFD56FEBD" box="[437,715,295,322]" pageId="24" pageNumber="175">J. C. van der Meer Mohr</collectorName>
)
<date id="FF86100EFFDEBB09FD55856DFCC5FEBE" box="[730,856,295,321]" pageId="24" pageNumber="175" value="1932-03-20">
<collectingDate id="EFC2E9E6FFDEBB09FD55856DFCC5FEBE" box="[730,856,295,321]" pageId="24" pageNumber="175" value="1932-03-20">20.III.1932</collectingDate>
</date>
(
<specimenCount id="9D3EFD47FFDEBB09FCE8856DFC0FFEBE" box="[871,914,295,321]" pageId="24" pageNumber="175" type="female">2 ♀</specimenCount>
<typeStatus id="5483886CFFDEBB09FC16856DFB9AFEBE" box="[921,1031,295,321]" pageId="24" pageNumber="175" type="paratype">paratypes</typeStatus>
,
<collectionCode id="ED29AE0BFFDEBB09FB9C856DFBF1FEBE" box="[1043,1132,295,321]" pageId="24" pageNumber="175">BMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B503C93FFDEBB09FBF0856DFAA7FE9A" ID-GBIF-Occurrence="3020160310" collectingDate="1934-10-05" collectionCode="BMNH, NGPC" collectorName="Goea Lau Rakit I &amp; J. C. van der Meer Mohr" country="Indonesia" latitude="2.3333333" location="Sumatra" longLatPrecision="1307" longitude="98.666664" pageId="24" pageNumber="175" specimenCount="8" specimenCount-female="8" stateProvince="Sumatera Utara" typeStatus="paratype">
<collectingRegion id="49FCF82CFFDEBB09FBF0856DFB42FEBE" box="[1151,1247,295,321]" country="Indonesia" name="Sumatera Utara" pageId="24" pageNumber="175">Sumatra</collectingRegion>
,
<collectorName id="26CD5318FFDEBB09FB64856DFF3FFE9A" pageId="24" pageNumber="175">Goea Lau Rakit I</collectorName>
,
<geoCoordinate id="EE0C5009FFDEBB09FF208501FE89FE9A" box="[175,276,331,357]" degrees="02" direction="north" minutes="20" orientation="latitude" pageId="24" pageNumber="175" precision="925" value="2.3333333">02°20’ N</geoCoordinate>
<geoCoordinate id="EE0C5009FFDEBB09FE948501FEE3FE9A" box="[283,382,331,357]" degrees="98" direction="east" minutes="40" orientation="longitude" pageId="24" pageNumber="175" precision="925" value="98.666664">98°40’ E</geoCoordinate>
(
<collectorName id="26CD5318FFDEBB09FE018501FD39FE99" box="[398,676,331,358]" pageId="24" pageNumber="175">J. C. van der Meer Mohr</collectorName>
)
<date id="FF86100EFFDEBB09FD3D8500FC86FE9A" box="[690,795,330,357]" pageId="24" pageNumber="175" value="1934-10-05">
<collectingDate id="EFC2E9E6FFDEBB09FD3D8500FC86FE9A" box="[690,795,330,357]" pageId="24" pageNumber="175" value="1934-10-05">5.X.1934</collectingDate>
</date>
(
<specimenCount id="9D3EFD47FFDEBB09FCA58501FCC8FE9A" box="[810,853,331,357]" pageId="24" pageNumber="175" type="female">6 ♀</specimenCount>
<typeStatus id="5483886CFFDEBB09FCD38501FC57FE9A" box="[860,970,331,357]" pageId="24" pageNumber="175" type="paratype">paratypes</typeStatus>
,
<collectionCode id="ED29AE0BFFDEBB09FC5A8501FBB0FE9A" box="[981,1069,331,357]" pageId="24" pageNumber="175">BMNH</collectionCode>
;
<specimenCount id="9D3EFD47FFDEBB09FBB78501FBFFFE9A" box="[1080,1122,331,357]" pageId="24" pageNumber="175" type="female">2 ♀</specimenCount>
<typeStatus id="5483886CFFDEBB09FBE68501FB4AFE9A" box="[1129,1239,331,357]" pageId="24" pageNumber="175" type="paratype">paratypes</typeStatus>
,
<collectionCode id="ED29AE0BFFDEBB09FB6D8501FAAFFE9A" box="[1250,1330,331,357]" pageId="24" pageNumber="175">NGPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8736CEFFDEBB09FF488525FAEFFE76" blockId="24.[151,1437,151,393]" box="[199,1394,367,393]" pageId="24" pageNumber="175">
<emphasis id="B94CEADCFFDEBB09FF488525FF62FE76" bold="true" box="[199,255,367,393]" pageId="24" pageNumber="175">Note</emphasis>
. The extremely thin and elongated appendages of species likely represent adaptations to life in caves.
</paragraph>
</subSubSection>
</treatment>
</document>
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