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<document id="8489FB5765340FC2CCE4D6448ED66B1F" ID-DOI="10.5281/zenodo.193859" ID-GBIF-Dataset="2395cee2-3f03-43e9-b86f-e8b4d2d5816b" ID-ISSN="1175-5326" ID-Zenodo-Dep="193859" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460174789656" checkinUser="plazi" docAuthor="Welton, Luke J., Siler, Cameron D., Diesmos, Arvin C. &amp; Brown, Rafe M." docDate="2010" docId="038EF74D3D557E33FF59F8C9B7A220E5" docLanguage="en" docName="zt02390p068.pdf" docOrigin="Zootaxa 2390" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cyrtodactylus jambangan Welton, Siler, Diesmos &amp; Brown, 2010, sp. nov." docType="treatment" docVersion="10" lastPageNumber="63" masterDocId="FFB78F353D507E3CFFCEFFE4B4462019" masterDocTitle="Phylogeny-based species delimitation of southern Philippines bent-toed geckos and a new species of Cyrtodactylus (Squamata: Gekkonidae) from western Mindanao and the Sulu Archipelago" masterLastPageNumber="68" masterPageNumber="49" pageNumber="54" updateTime="1698236077467" updateUser="plazi">
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<mods:title id="587C7DB96118BFD7800ABCDC6ED4BBA2">Phylogeny-based species delimitation of southern Philippines bent-toed geckos and a new species of Cyrtodactylus (Squamata: Gekkonidae) from western Mindanao and the Sulu Archipelago</mods:title>
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<mods:namePart id="5CB4EA1F613ABF324EF67C6379B14035">Welton, Luke J.</mods:namePart>
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<mods:namePart id="95A8F3C43050F30C1652B4D801093B23">Siler, Cameron D.</mods:namePart>
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<mods:namePart id="EBB9F3CA0F6706536BEE0A4D39C329D3">Diesmos, Arvin C.</mods:namePart>
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<mods:namePart id="7599908ACA59F079623A2C6F05C3D7D0">Brown, Rafe M.</mods:namePart>
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<mods:date id="7935327820134AA8363525F5ED95CBB0">2010</mods:date>
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<treatment id="038EF74D3D557E33FF59F8C9B7A220E5" ID-DOI="http://doi.org/10.5281/zenodo.5627392" ID-GBIF-Taxon="119383519" ID-Zenodo-Dep="5627392" LSID="urn:lsid:plazi:treatment:038EF74D3D557E33FF59F8C9B7A220E5" httpUri="http://treatment.plazi.org/id/038EF74D3D557E33FF59F8C9B7A220E5" lastPageId="15" lastPageNumber="63" pageId="5" pageNumber="54">
<subSubSection id="C33D15D03D557E39FF59F8C9B5422773" pageId="5" pageNumber="54" type="nomenclature">
<paragraph id="8B98465B3D557E39FF59F8C9B674275E" blockId="5.[151,562,1837,1898]" box="[151,562,1837,1863]" pageId="5" pageNumber="54">
<heading id="D0D0F1373D557E39FF59F8C9B674275E" bold="true" box="[151,562,1837,1863]" fontSize="11" level="1" pageId="5" pageNumber="54" reason="1">
<emphasis id="B9539A493D557E39FF59F8C9B674275E" bold="true" box="[151,562,1837,1863]" pageId="5" pageNumber="54">
<taxonomicName id="4C273DD83D557E39FF59F8C9B58B275E" box="[151,461,1837,1863]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="54" phylum="Chordata" rank="species" species="jambangan" status="sp. nov.">
<emphasis id="B9539A493D557E39FF59F8C9B58B275E" bold="true" box="[151,461,1837,1863]" italics="true" pageId="5" pageNumber="54">Cyrtodactylus jambangan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26027323D557E39FE1AF8C9B674275E" box="[468,562,1837,1863]" pageId="5" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B98465B3D557E39FF59F8B4B5422773" blockId="5.[151,562,1837,1898]" box="[151,260,1872,1898]" pageId="5" pageNumber="54">
<figureCitation id="131C5ADE3D557E39FF59F8B4B5422773" box="[151,260,1872,1898]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="9.[151,255,1109,1133]" captionStartId-1="10.[151,258,1163,1187]" captionStartId-2="12.[151,255,1863,1887]" captionTargetBox-0="[409,1177,197,1086]" captionTargetBox-1="[417,1170,276,1117]" captionTargetBox-2="[154,1436,212,1822]" captionTargetId-0="figure@9.[409,1177,197,1086]" captionTargetId-1="figure@10.[409,1177,269,1142]" captionTargetId-2="figure@12.[151,1436,197,1837]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="12" captionText-0="FIGURE 3. Dorsal aspect of typical sized specimens of Cyrtodactylus annulatus (left: KU 309364; Camiguin Sur Isl.) and C. jambangan, n. sp., (right: holotype PNM 9593 specimen from Pasonanca, Zamboanga City Province)." captionText-1="FIGURE 4. Precloacal regions of (A) Cyrtodactylus jambangan, n. sp., and (B) C. annulatus (from Taylor 1922), showing the four-pour configuration typical of C. jambangan and the normal condition of six pores in C. annulatus. Note the presence of separated lateral patches of enlarged scales on either side of the precloacal pores (anterior precloacals undifferentiated or only slightly differentiated in C. jambangan) versus a continuous arc of greatly enlarged precloacals in C. annulatus." captionText-2="FIGURE 5. Photographs of Cyrtodactylus jambangan, n. sp., in life: typical muted, marbled color pattern of (A) adults (holotype PNM 9593) and (B) more strikingly patterned contrasting coloration of juveniles (KU 314812)." httpUri-0="https://zenodo.org/record/193862/files/figure.png" httpUri-1="https://zenodo.org/record/193863/files/figure.png" httpUri-2="https://zenodo.org/record/193864/files/figure.png" pageId="5" pageNumber="54">Figs. 35</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D15D03D557E3AFF59F871B58E2125" lastPageId="6" lastPageNumber="55" pageId="5" pageNumber="54" type="materials_examined">
<paragraph id="8B98465B3D557E39FF59F871B1DA27ED" blockId="5.[151,1436,1941,2036]" pageId="5" pageNumber="54">
<emphasis id="B9539A493D557E39FF59F871B55627B6" bold="true" box="[151,272,1941,1967]" pageId="5" pageNumber="54">
<typeStatus id="549CF8F93D557E39FF59F871B54D27B6" box="[151,267,1941,1967]" pageId="5" pageNumber="54" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="F33006CB3D557E39FED1F871B51927B6" box="[287,351,1941,1967]" name="Panama" pageId="5" pageNumber="54">
<collectionCode id="ED36DE9E3D557E39FED1F871B51927B6" box="[287,351,1941,1967]" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="5" pageNumber="54">PNM</collectionCode>
</collectingCountry>
9593 (formerly KU 314810, RMB Field No. 10,005), adult male, collected on the trunk of a sapling (20:0023:00) in Pasonanca Natural Park, Barangay Baluno, Zamboanga City Province, Mindanao Island,
<collectingCountry id="F33006CB3D557E39FF21F83EB53327ED" box="[239,373,2010,2036]" name="Philippines" pageId="5" pageNumber="54">Philippines</collectingCountry>
(N: 07º 1.0554, E: 122º 1.731, WGS-84;
<quantity id="4CDFEBBE3D557E39FCA4F83EB7F027ED" box="[874,950,2010,2036]" metricMagnitude="2" metricUnit="m" metricValue="7.58" pageId="5" pageNumber="54" unit="m" value="758.0">758 m</quantity>
above sea level), on
<date id="FF99609B3D557E39FB7AF83EB14B27ED" box="[1204,1293,2010,2036]" pageId="5" pageNumber="54" value="1912-07">12 July</date>
, 2008 by L.
</paragraph>
<paragraph id="8B98465B3D567E3AFF59FF7CB05320AB" blockId="6.[151,1437,152,2015]" box="[151,1045,152,178]" pageId="6" pageNumber="55">Welton, R. Brown, C. Siler, J. Fernandez, M. and V. Yngente, and J. Phenix.</paragraph>
<paragraph id="8B98465B3D567E3AFF09FF5EB58E2125" blockId="6.[151,1437,152,2015]" pageId="6" pageNumber="55">
<emphasis id="B9539A493D567E3AFF09FF5EB5C520CD" bold="true" box="[199,387,186,212]" pageId="6" pageNumber="55">Paratopotypes.</emphasis>
Four males (KU 314813, 314815, 314825, and 314828), fourteen females (KU 314808, 314811, 31481618, 314820, 313482224, 31482627, and 31482931), and five juveniles (KU 314809, 314812, 314814, 314819, and 314821) collected on the trunks of trees (18:0023:00),
<date id="FF99609B3D567E3AFB43FEE4B1122103" box="[1165,1364,256,282]" pageId="6" pageNumber="55" value="2008-07-12" valueMax="2008-07-18" valueMin="2008-07-12">1218 July 2008</date>
, other data identical to
<typeStatus id="549CF8F93D567E3AFE97FEC6B5852125" box="[345,451,290,316]" pageId="6" pageNumber="55" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D15D03D567E3AFF09FEA1B12324FD" pageId="6" pageNumber="55" type="description">
<paragraph id="8B98465B3D567E3AFF09FEA1B12324FD" blockId="6.[151,1437,152,2015]" pageId="6" pageNumber="55">
<emphasis id="B9539A493D567E3AFF09FEA1B5092146" bold="true" box="[199,335,325,351]" pageId="6" pageNumber="55">
<typeStatus id="549CF8F93D567E3AFF09FEA1B50E2146" box="[199,328,325,351]" pageId="6" pageNumber="55" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Four males (KU 31479394 and 31479697), two females (KU 314795 and 314798), and three juveniles (KU 314792, and 31483334), collected
<date id="FF99609B3D567E3AFC89FE8CB79C219B" box="[839,986,360,386]" pageId="6" pageNumber="55" value="2008-07-07">7 July 2008</date>
, in Pasonanca Natural Park, Tumaga River (N: 6º 35.174, E: 122º
<quantity id="4CDFEBBE3D567E3AFE29FE6EB62921BD" box="[487,623,394,420]" metricMagnitude="3" metricUnit="m" metricValue="2.42694" pageId="6" pageNumber="55" unit="m" value="2426.94">2.426, 94m</quantity>
, WGS-84), by L. Welton, R. Brown, C. Siler, J. Fernandez, M. and V. Yngente, and J. Phenix; one male (KU 314835), one female (KU 314806), and one juvenile (KU 314799), collected
<date id="FF99609B3D567E3AFEC7FE34B5D121F3" box="[265,407,464,490]" pageId="6" pageNumber="55" value="2008-07-08">8 July 2008</date>
, in Pasonanca Natural Park, Tumaga River (N: 6º 58.624, E: 122º
<quantity id="4CDFEBBE3D567E3AFB62FE34B10221F3" box="[1196,1348,464,490]" metricMagnitude="3" metricUnit="m" metricValue="3.0431039999999996" pageId="6" pageNumber="55" unit="m" value="3043.104">3.043, 104m</quantity>
, WGS- 84), by the same collectors; two males (KU 314807 and 314836) collected
<date id="FF99609B3D567E3AFBFEFE16B0B62215" box="[1072,1264,498,524]" pageId="6" pageNumber="55" value="2008-07-09" valueMax="2008-07-11" valueMin="2008-07-09">911 July 2008</date>
, in Pasonanca Natural Park, Tumaga River (N: 6º 59.534, E: 122º
<quantity id="4CDFEBBE3D567E3AFD36FDF1B7C62236" box="[760,896,533,559]" metricMagnitude="3" metricUnit="m" metricValue="3.62694" pageId="6" pageNumber="55" unit="m" value="3626.94">3.626, 94m</quantity>
, WGS-84), by the same collectors; two males (KU 314803 and 314805), one female (KU 314804), and three juveniles (KU 31480002), collected
<date id="FF99609B3D567E3AFA9DFDDCB490226D" pageId="6" pageNumber="55" value="2008-07-09">9 July 2008</date>
, Pasonanca Natural Park, Tumaga River (N: 6º 58.624, E: 122º
<quantity id="4CDFEBBE3D567E3AFC10FDBEB03C226D" box="[990,1146,602,628]" metricMagnitude="3" metricUnit="m" metricValue="4.043104" pageId="6" pageNumber="55" unit="m" value="4043.104">4.043, 104m</quantity>
, WGS-84), by the same collectors; two males (KU 314781 and 314784), one female (KU 314780), and four juveniles (KU 31477879 and 31478283), collected
<date id="FF99609B3D567E3AFE3CFD44B69E22A3" box="[498,728,672,698]" pageId="6" pageNumber="55" value="2008-04-20" valueMax="2008-04-22" valueMin="2008-04-20">2022 April 2008</date>
, in Pasonanca Natural Park, Barangay Pasonanca, Sitio Canucutan (N: 6º 59.534, E: 122º
<quantity id="4CDFEBBE3D567E3AFDF0FD26B69A22C5" box="[574,732,706,732]" metricMagnitude="3" metricUnit="m" metricValue="3.6261039999999993" pageId="6" pageNumber="55" unit="m" value="3626.104">3.626, 104m</quantity>
, WGS-84), by R. Brown and A. Diesmos; one male (KU 319654), one female (KU 319655), and one juvenile (KU 319652), collected
<date id="FF99609B3D567E3AFBF6FD01B0B422E6" box="[1080,1266,741,767]" pageId="6" pageNumber="55" value="2009-04-03" valueMax="2009-04-05" valueMin="2009-04-03">35 April 2009</date>
, in Pasonanca Natural Park, Barangay La Paz, Sitio Nancy (N: 7º 5.099, E: 122º
<quantity id="4CDFEBBE3D567E3AFC69FCECB00A233B" box="[935,1100,776,802]" metricMagnitude="3" metricUnit="m" metricValue="1.620113" pageId="6" pageNumber="55" unit="m" value="1620.113">1.620, 1130m</quantity>
, WGS-84), by R. Brown, C. Infante, and A. Diesmos; two males (KU 319656 and 319657), collected
<date id="FF99609B3D567E3AFBE4FCCEB0AA235D" box="[1066,1260,810,836]" pageId="6" pageNumber="55" value="2009-04-06" valueMax="2009-04-09" valueMin="2009-04-06">69 April 2009</date>
, in Pasonanca Natural Park, Barangay Tulosa, Sitio Santa Clara, Cabo Negros outpost (N: 7º 6.479, E: 122º
<quantity id="4CDFEBBE3D567E3AFB30FCA9B1D1237E" box="[1278,1431,845,871]" metricMagnitude="3" metricUnit="m" metricValue="7.139619999999999" pageId="6" pageNumber="55" unit="m" value="7139.62">7.139, 620m</quantity>
, WGS-84), by the same collectors; three males (KU 31966062), and one female (KU 319658), collected
<date id="FF99609B3D567E3AFAB1FC94B55A23B5" pageId="6" pageNumber="55" value="2009-04-11">11 April 2009</date>
, in Pasonanca Natural Park, Tumaga River (N: 6º 35.174, E: 122º
<quantity id="4CDFEBBE3D567E3AFBF6FC76B08523B5" box="[1080,1219,914,940]" metricMagnitude="3" metricUnit="m" metricValue="2.42694" pageId="6" pageNumber="55" unit="m" value="2426.94">2.426, 94m</quantity>
, WGS-84), by the same collectors; twelve males (CAS 60195, 60197, 60199, 60203205, 60208, 60212, 6021718, 60221, and 60453),
<specimenCount id="9D218DD23D567E3AFF32FC3CB5C623EB" box="[252,384,984,1010]" pageId="6" pageNumber="55" type="female">15 females</specimenCount>
(CAS 60196, 60200201, 60207, 6020910, 6021316, 60220, 6045455, 60458, and 60460), and six juveniles (CAS 6022224, 6045657, and 60459) collected at Abung-Abung and Port
<collectingCountry id="F33006CB3D567E3AFF59FBF9B4B0242E" box="[151,246,1053,1079]" name="Netherlands" pageId="6" pageNumber="55">Holland</collectingCountry>
on Basilan Island,
<date id="FF99609B3D567E3AFE12FBF9B6C6242E" box="[476,640,1053,1079]" pageId="6" pageNumber="55" value="1925-10-05">525 October</date>
, 1921, by E. H. Taylor; two males (CAS 6062223), three females (CAS 6061921), and one juvenile (CAS 60624) collected on New Govenen Island,
<date id="FF99609B3D567E3AFB4AFBA4B16E2443" box="[1156,1320,1088,1114]" pageId="6" pageNumber="55" value="1925-10-05">525 October</date>
, 1920, by E. H. Taylor; one male (CAS 60669), four females (CAS 6067072 and 60887), and two juveniles (CAS 60886 and 60888) collected on Bud Daho Mt. on Jolo Island,
<date id="FF99609B3D567E3AFC59FB61B09C2486" box="[919,1242,1157,1183]" pageId="6" pageNumber="55">25 October17 November</date>
, 1920, by E. H. Taylor; two males (CAS 62020 and 62022), three females (CAS 62017, 62019, and 62021), and one juvenile (CAS 62018) collected in Zamboanga del Sur Province,
<date id="FF99609B3D567E3AFCE0FB2EB01224FD" box="[814,1108,1226,1252]" pageId="6" pageNumber="55">23 September6 October</date>
, 1920, by E. H. Taylor.
</paragraph>
</subSubSection>
<subSubSection id="C33D15D03D567E35FF09FB09B0E3265C" lastPageId="9" lastPageNumber="58" pageId="6" pageNumber="55" type="diagnosis">
<paragraph id="8B98465B3D567E3AFF09FB09B06A2626" blockId="6.[151,1437,152,2015]" pageId="6" pageNumber="55">
<emphasis id="B9539A493D567E3AFF09FB09B50E251E" bold="true" box="[199,328,1261,1287]" pageId="6" pageNumber="55">Diagnosis.</emphasis>
The new species differs from most of Asias 115
<taxonomicName id="4C273DD83D567E3AFC50FB09B003251F" box="[926,1093,1261,1286]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="B9539A493D567E3AFC50FB09B003251F" box="[926,1093,1261,1286]" italics="true" pageId="6" pageNumber="55">Cyrtodactylus</emphasis>
</taxonomicName>
species by the same suite of character states that has facilitated the recognition and distinction of
<taxonomicName id="4C273DD83D567E3AFC1CFAF4B02C2530" box="[978,1130,1296,1321]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D567E3AFC1CFAF4B02C2530" box="[978,1130,1296,1321]" italics="true" pageId="6" pageNumber="55">C. annulatus</emphasis>
</taxonomicName>
: (1) small body size; (2) light gray body, with dark transverse bands; (3) presence of a weak precloacal groove; (4) absence of enlarged femoral scales and pores; (5) moderately spinose tuberculation; (6) presence of forelimb tuberculation; (7) and absence of enlarged, median subcaudal scales (Taylor 1922;
<bibRefCitation id="EFB63BAA3D567E3AFCBAFA9CB0C5258B" author="Brown" box="[884,1155,1400,1426]" pageId="6" pageNumber="66" refString="Brown, W. C. &amp; Alcala, A. C. (1978) Philippine Lizards of the Family Gekkonidae. Silliman University Press, Dumaguete City, pp. 1 - 146." type="book chapter" year="1978">Brown &amp; Alcala 1978</bibRefCitation>
;
<bibRefCitation id="EFB63BAA3D567E3AFB5EFA9CB49325AD" author="Manthey" pageId="6" pageNumber="67" refString="Manthey, U. &amp; Grossman, W. (1997) Amphibien und Reptilien sudostasiens. Natur undTier-Verlag, Berlin, Germany, 512 pp." type="book" year="1997">Manthey &amp; Grossman 1997</bibRefCitation>
; Grismer 2005;
<bibRefCitation id="EFB63BAA3D567E3AFE5DFA7EB63425AD" author="Welton" box="[403,626,1434,1460]" pageId="6" pageNumber="67" refString="Welton, L. J., Siler, C. D., Diesmos, A. C. &amp; Brown, R. M. (2009) A new bent-toed Gekko (Genus Cyrtodactylus) from southern Palawan Island, Philippines and clarification of the taxonomic status of C. annulatus. Herpetologica, 65: 328 - 343." type="journal article" year="2009">
Welton
<emphasis id="B9539A493D567E3AFE21FA7FB66C25AD" box="[495,554,1435,1460]" italics="true" pageId="6" pageNumber="55">et al.</emphasis>
2009
</bibRefCitation>
). Because the new species differs from non-Philippine congeners by characters that have distinguished
<taxonomicName id="4C273DD83D567E3AFDF7FA59B69225CF" box="[569,724,1469,1494]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D567E3AFDF7FA59B69225CF" box="[569,724,1469,1494]" italics="true" pageId="6" pageNumber="55">C. annulatus</emphasis>
</taxonomicName>
(and have never been challenged;
<bibRefCitation id="EFB63BAA3D567E3AFBB4FA59B15825CE" author="Taylor" box="[1146,1310,1469,1495]" pageId="6" pageNumber="67" refString="Taylor, E. H. (1922 a) The lizards of the Philippine Islands. Philippine Bureau of Science, Manila, Philippines, 269 pp." type="book" year="1922" yearSuffix="a">Taylor 1922a</bibRefCitation>
;
<bibRefCitation id="EFB63BAA3D567E3AFAE5FA59B56E25E3" author="Brown" pageId="6" pageNumber="66" refString="Brown, W. C. &amp; Alcala, A. C. (1978) Philippine Lizards of the Family Gekkonidae. Silliman University Press, Dumaguete City, pp. 1 - 146." type="book chapter" year="1978">Brown &amp; Alcala 1978</bibRefCitation>
), we do not provide exhaustive comparisons to all those species (see
<bibRefCitation id="EFB63BAA3D567E3AFB9FFA04B16A25E3" author="Welton" box="[1105,1324,1504,1530]" pageId="6" pageNumber="67" refString="Welton, L. J., Siler, C. D., Diesmos, A. C. &amp; Brown, R. M. (2009) A new bent-toed Gekko (Genus Cyrtodactylus) from southern Palawan Island, Philippines and clarification of the taxonomic status of C. annulatus. Herpetologica, 65: 328 - 343." type="journal article" year="2009">
Welton
<emphasis id="B9539A493D567E3AFB62FA04B0A125E0" box="[1196,1255,1504,1529]" italics="true" pageId="6" pageNumber="55">et al.</emphasis>
2009
</bibRefCitation>
). Instead we focus on the comparisons relevant for the recognition of the new species: the closely related taxa within the
<taxonomicName id="4C273DD83D567E3AFF0CF9C1B5112627" box="[194,343,1573,1598]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D567E3AFF0CF9C1B5112627" box="[194,343,1573,1598]" italics="true" pageId="6" pageNumber="55">C. annulatus</emphasis>
</taxonomicName>
complex and other
<taxonomicName id="4C273DD83D567E3AFD8EF9C1B6A22627" box="[576,740,1573,1598]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="B9539A493D567E3AFD8EF9C1B6A22627" box="[576,740,1573,1598]" italics="true" pageId="6" pageNumber="55">Cyrtodactylus</emphasis>
</taxonomicName>
endemic to the
<collectingCountry id="F33006CB3D567E3AFC6EF9C1B06E2626" box="[928,1064,1573,1599]" name="Philippines" pageId="6" pageNumber="55">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8B98465B3D567E3AFF09F9ACB5C427C6" blockId="6.[151,1437,152,2015]" pageId="6" pageNumber="55">
C
<emphasis id="B9539A493D567E3AFF14F9ACB5B12678" box="[218,503,1608,1633]" italics="true" pageId="6" pageNumber="55">
yrtodactylus
<taxonomicName id="4C273DD83D567E3AFEBDF9ACB5B12678" box="[371,503,1608,1633]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="jambangan">jambangan</taxonomicName>
</emphasis>
differs from all other species of
<collectingCountry id="F33006CB3D567E3AFCB7F9ACB7B7267B" box="[889,1009,1608,1634]" name="Philippines" pageId="6" pageNumber="55">Philippine</collectingCountry>
<taxonomicName id="4C273DD83D567E3AFC34F9ACB0D82678" box="[1018,1182,1608,1633]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="55" phylum="Chordata" rank="genus">
<emphasis id="B9539A493D567E3AFC34F9ACB0D82678" box="[1018,1182,1608,1633]" italics="true" pageId="6" pageNumber="55">Cyrtodactylus</emphasis>
</taxonomicName>
(i.e.,
<emphasis id="B9539A493D567E3AFB2DF9ACB65B269D" italics="true" pageId="6" pageNumber="55">
<taxonomicName id="4C273DD83D567E3AFB2DF9ACB1D12678" box="[1251,1431,1608,1633]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="agusanensis">C. agusanensis</taxonomicName>
,
<taxonomicName id="4C273DD83D567E3AFF59F98FB51C269D" box="[151,346,1643,1668]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="philippinicus">C. philippinicus</taxonomicName>
,
<taxonomicName id="4C273DD83D567E3AFEA8F98FB65F269D" box="[358,537,1643,1668]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="redimiculus">C. redimiculus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C273DD83D567E3AFD97F98FB756269D" box="[601,784,1643,1668]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="55" phylum="Chordata" rank="species" species="tautbatorum">
<emphasis id="B9539A493D567E3AFD97F98FB756269D" box="[601,784,1643,1668]" italics="true" pageId="6" pageNumber="55">C. tautbatorum</emphasis>
</taxonomicName>
; Table 2) by the following combination of characters: (1) small body size (2) dorsum darkly marbled lavender and brown or brown with indistinct wavy lavender blotches; (3) presence of bright yellow superciliaries, canthal stripe, and dorsal tubercles; (4) low dorsal scale counts (as measured by midbody dorsal and paravertebral scales; Table 2); (5) moderate numbers of midbody dorsal tubercle rows (Table 2); (6) absence of femoral pores in both sexes; (7) presence of precloacal pores in males and similarly enlarged, dimpled, scales in females, arranged in an inverted “V”-shaped configuration; (8) moderately depressed precloacal groove; (9) scales anterior to precloacal region undifferentiated; (10) lamellae under Finger
<date id="FF99609B3D567E3AFE53F8B9B648276E" box="[413,526,1885,1911]" pageId="6" pageNumber="55" value="1917-03-22">III 1722</date>
; (11) lamellae under Toe
<date id="FF99609B3D567E3AFCFBF8B9B7E2276E" box="[821,932,1885,1911]" pageId="6" pageNumber="55" value="1920-04-24">IV 2024</date>
(12) supralabials to beneath eye 811; (13) infralabials to beneath the eye 68; (14) absence of stripes connecting the lateral margins of transverse bands on trunk; and (15) midbody ventral scales 4863. Comparisons with additional Southeast Asian species are provided in Table 3.
</paragraph>
<caption id="DF5816D33D597E35FF59FBB1B1622494" httpUri="https://zenodo.org/record/193862/files/figure.png" pageId="9" pageNumber="58" targetBox="[409,1177,197,1086]" targetPageId="9">
<paragraph id="8B98465B3D597E35FF59FBB1B1622494" blockId="9.[151,1435,1109,1165]" pageId="9" pageNumber="58">
<emphasis id="B9539A493D597E35FF59FBB1B55F2474" bold="true" box="[151,281,1109,1133]" pageId="9" pageNumber="58">FIGURE 3.</emphasis>
Dorsal aspect of typical sized specimens of
<taxonomicName id="4C273DD83D597E35FCCEFBB2B0402474" box="[768,1030,1110,1133]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D597E35FCCEFBB2B0402474" box="[768,1030,1110,1133]" italics="true" pageId="9" pageNumber="58">Cyrtodactylus annulatus</emphasis>
</taxonomicName>
(left: KU 309364; Camiguin Sur Isl.) and
<taxonomicName id="4C273DD83D597E35FF0AFB92B51D2494" box="[196,347,1142,1165]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="jambangan" status="sp. nov.">
<emphasis id="B9539A493D597E35FF0AFB92B51D2494" box="[196,347,1142,1165]" italics="true" pageId="9" pageNumber="58">C. jambangan</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26027323D597E35FEA6FB92B5E52494" box="[360,419,1142,1165]" pageId="9" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
, (right: holotype PNM 9593 specimen from Pasonanca, Zamboanga City Province).
</paragraph>
</caption>
<paragraph id="8B98465B3D597E35FF09FB52B0E3265C" blockId="9.[151,1437,1206,2016]" pageId="9" pageNumber="58">
The critical comparison for the recognition of the new species is the diagnosis of the Zamboanga and Sulu populations from true
<taxonomicName id="4C273DD83D597E35FE7FFB38B61724EC" box="[433,593,1244,1269]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D597E35FE7FFB38B61724EC" box="[433,593,1244,1269]" italics="true" pageId="9" pageNumber="58">C. annulatus</emphasis>
</taxonomicName>
(
<typeStatus id="549CF8F93D597E35FDAAFB3FB6DA24EC" box="[612,668,1243,1269]" pageId="9" pageNumber="58">type</typeStatus>
locality from the Municipality of Bunawan, Agusan Del Sur Province, Mindanao Island), and newly designated
<taxonomicName id="4C273DD83D597E35FD39FAE5B7C82503" box="[759,910,1281,1306]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D597E35FD39FAE5B7C82503" box="[759,910,1281,1306]" italics="true" pageId="9" pageNumber="58">C. annulatus</emphasis>
</taxonomicName>
<typeStatus id="549CF8F93D597E35FC58FAE5B7B52502" box="[918,1011,1281,1307]" pageId="9" pageNumber="58" type="neotype">neotype</typeStatus>
locality (
<bibRefCitation id="EFB63BAA3D597E35FBADFAE5B1072502" author="Welton" box="[1123,1345,1281,1307]" pageId="9" pageNumber="67" refString="Welton, L. J., Siler, C. D., Diesmos, A. C. &amp; Brown, R. M. (2009) A new bent-toed Gekko (Genus Cyrtodactylus) from southern Palawan Island, Philippines and clarification of the taxonomic status of C. annulatus. Herpetologica, 65: 328 - 343." type="journal article" year="2009">
Welton
<emphasis id="B9539A493D597E35FB0FFAE5B0B32503" box="[1217,1269,1281,1306]" italics="true" pageId="9" pageNumber="58">et al</emphasis>
. 2009
</bibRefCitation>
) of Mt. Hilonghilong, Diwata Mountains, Agusan Del Norte Province, Mindanao Island. Although morphologically similar to
<taxonomicName id="4C273DD83D597E35FED3FAA8B585257C" box="[285,451,1356,1381]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D597E35FED3FAA8B585257C" box="[285,451,1356,1381]" italics="true" pageId="9" pageNumber="58">C. annulatus</emphasis>
</taxonomicName>
from throughout its range,
<taxonomicName id="4C273DD83D597E35FCE3FAA8B7A5257C" box="[813,995,1356,1381]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="jambangan">
<emphasis id="B9539A493D597E35FCE3FAA8B7A5257C" box="[813,995,1356,1381]" italics="true" pageId="9" pageNumber="58">C. jambangan</emphasis>
</taxonomicName>
has bright yellow canthal stripes, superciliaries, dorsal tubercles, and moderate-sized dorsolateral body tubercles, fewer precloacal pores, undifferentiated precloacals, and strongly spinose (versus moderate, domed) dorsal tuberculation. The new species is separated from
<taxonomicName id="4C273DD83D597E35FE07FA58B62625CC" box="[457,608,1468,1493]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D597E35FE07FA58B62625CC" box="[457,608,1468,1493]" italics="true" pageId="9" pageNumber="58">C. annulatus</emphasis>
</taxonomicName>
by 7.99.5% (mean = 8.6%) uncorrected sequence divergence (
<tableCitation id="C6A573E03D597E35FA96FA5FB4E125E2" captionStart="TABLE 4" captionStartId="11.[151,239,263,287]" captionTargetBox="[161,1410,450,2013]" captionTargetPageId="11" captionText="TABLE 4. Summary of univariate morphological variation among mensural and meristic characters in the type series of Cyrtodactylus jambangan, n. sp., from Pasonanca Natural Park, Zamboanga Peninsula, western Mindanao Island, and Cyrtodactylus annulatus from eastern Mindanao Island (including Bunawan, the C. annulatus type locality). Labial scale counts were counted to below the center of the eye and bilaterally symmetrical characters are separated into left (l) and right (r) sides of the body. See Materials and Methods for character abbreviations." httpUri="http://table.plazi.org/id/DF5816D33D5B7E37FF59FEE3B0732186" pageId="9" pageNumber="58" tableUuid="DF5816D33D5B7E37FF59FEE3B0732186">Table 4</tableCitation>
) in the ND2 protein-coding mitochondrial gene region (see Table 3 for complete univariate morphometric data for
<taxonomicName id="4C273DD83D597E35FF37F9E2B5C92606" box="[249,399,1542,1567]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D597E35FF37F9E2B5C92606" box="[249,399,1542,1567]" italics="true" pageId="9" pageNumber="58">C. annulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C273DD83D597E35FE06F9E2B62B2606" box="[456,621,1542,1567]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="58" phylum="Chordata" rank="species" species="jambangan">
<emphasis id="B9539A493D597E35FE06F9E2B62B2606" box="[456,621,1542,1567]" italics="true" pageId="9" pageNumber="58">C. jambangan</emphasis>
</taxonomicName>
). The new species differs from all remaining
<collectingCountry id="F33006CB3D597E35FB40F9E2B1412639" box="[1166,1287,1542,1568]" name="Philippines" pageId="9" pageNumber="58">Philippine</collectingCountry>
species by a variety of morphological and color pattern characters (Table 2;
<bibRefCitation id="EFB63BAA3D597E35FCB3F9CFB0DE265C" author="Brown" box="[893,1176,1579,1605]" pageId="9" pageNumber="66" refString="Brown, W. C. &amp; Alcala, A. C. (1978) Philippine Lizards of the Family Gekkonidae. Silliman University Press, Dumaguete City, pp. 1 - 146." type="book chapter" year="1978">Brown and Alcala 1978</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33D15D03D597E32FF09F9B4B78726D8" lastPageId="14" lastPageNumber="63" pageId="9" pageNumber="58" type="description">
<paragraph id="8B98465B3D597E35FF09F9B4B7B026C2" blockId="9.[151,1437,1206,2016]" pageId="9" pageNumber="58">
<emphasis id="B9539A493D597E35FF09F9B4B5B02673" bold="true" box="[199,502,1616,1642]" pageId="9" pageNumber="58">
Description of
<typeStatus id="549CF8F93D597E35FE4CF9B4B5B72673" box="[386,497,1616,1642]" pageId="9" pageNumber="58" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male, snoutvent length 67.7 mm; head moderately long, distinct from neck, 30.3% SVL; head width 64.7% and height 41.7% head length; head triangular in dorsal aspect; lores concave; snout elongate, 41.7% head length, anterior tip rounded; eye diameter 20.6% head length, 76.3% eyeear distance; auricular opening ovoid, longest axis 9.5% head length.
</paragraph>
<paragraph id="8B98465B3D597E36FF09F902B02A20CE" blockId="9.[151,1437,1206,2016]" lastBlockId="10.[151,1437,152,215]" lastPageId="10" lastPageNumber="59" pageId="9" pageNumber="58">Dorsal head scalation heterogenous, scales small and granular, with tubercle density sparse medially, tubercles increasing in size and density posteriorly and laterally; superciliaries increasing in size (laterally elongated) anteriorly, with largest scales at anterodorsal margin of orbit; rostral taller than wide, divided dorsally by inverted “Y” shaped crease; rostral bordered by large, anterior internasal and smaller, paired posterior internasals, supranasals, and first supralabials; internarial distance 2.2 mm; nostrils bordered by supranasals, rostral, 2 postnasals, and first supralabials; supralabials rectangular, decreasing in size posteriorly (left/right): 9/10 to midpoint of eye, 12/12 total; infralabials 7/7 to midpoint of eye, 11/10 total; supralabials bordered dorsally by secondary, slightly differentiated row of scales, extending to anterior margin of orbit; infralabials bordered ventrally by similar row, extending to anterior margin of orbit.</paragraph>
<caption id="DF5816D33D5A7E36FF59FB6FB586253B" httpUri="https://zenodo.org/record/193863/files/figure.png" pageId="10" pageNumber="59" targetBox="[417,1170,276,1117]" targetPageId="10">
<paragraph id="8B98465B3D5A7E36FF59FB6FB586253B" blockId="10.[151,1436,1163,1314]" pageId="10" pageNumber="59">
<emphasis id="B9539A493D5A7E36FF59FB6FB55824BA" bold="true" box="[151,286,1163,1187]" pageId="10" pageNumber="59">FIGURE 4.</emphasis>
Precloacal regions of (
<emphasis id="B9539A493D5A7E36FDE5FB6FB67824BA" bold="true" box="[555,574,1163,1187]" pageId="10" pageNumber="59">A</emphasis>
)
<taxonomicName id="4C273DD83D5A7E36FD9EFB6FB73624BB" box="[592,880,1163,1186]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="jambangan" status="sp. nov.">
<emphasis id="B9539A493D5A7E36FD9EFB6FB73624BB" box="[592,880,1163,1186]" italics="true" pageId="10" pageNumber="59">Cyrtodactylus jambangan</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26027323D5A7E36FCB1FB6FB7F824BB" box="[895,958,1163,1186]" pageId="10" pageNumber="59" rank="species">n. sp.</taxonomicNameLabel>
, and (
<emphasis id="B9539A493D5A7E36FBCAFB6FB05024BA" bold="true" box="[1028,1046,1163,1187]" pageId="10" pageNumber="59">B</emphasis>
)
<taxonomicName id="4C273DD83D5A7E36FBE9FB6FB0F024BB" box="[1063,1206,1163,1186]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D5A7E36FBE9FB6FB0F024BB" box="[1063,1206,1163,1186]" italics="true" pageId="10" pageNumber="59">C. annulatus</emphasis>
</taxonomicName>
(from Taylor 1922), showing the four-pour configuration typical of
<taxonomicName id="4C273DD83D5A7E36FD6FFB4FB77B24DB" box="[673,829,1195,1218]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="jambangan">
<emphasis id="B9539A493D5A7E36FD6FFB4FB77B24DB" box="[673,829,1195,1218]" italics="true" pageId="10" pageNumber="59">C. jambangan</emphasis>
</taxonomicName>
and the normal condition of six pores in
<taxonomicName id="4C273DD83D5A7E36FAC6FB4FB1D324DB" box="[1288,1429,1195,1218]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D5A7E36FAC6FB4FB1D324DB" box="[1288,1429,1195,1218]" italics="true" pageId="10" pageNumber="59">C. annulatus</emphasis>
</taxonomicName>
. Note the presence of separated lateral patches of enlarged scales on either side of the precloacal pores (anterior precloacals undifferentiated or only slightly differentiated in
<taxonomicName id="4C273DD83D5A7E36FCE8FB0FB7F8251B" box="[806,958,1259,1282]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="jambangan">
<emphasis id="B9539A493D5A7E36FCE8FB0FB7F8251B" box="[806,958,1259,1282]" italics="true" pageId="10" pageNumber="59">C. jambangan</emphasis>
</taxonomicName>
) versus a continuous arc of greatly enlarged precloacals in
<taxonomicName id="4C273DD83D5A7E36FEFFFAEFB5FF253B" box="[305,441,1291,1314]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="59" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D5A7E36FEFFFAEFB5FF253B" box="[305,441,1291,1314]" italics="true" pageId="10" pageNumber="59">C. annulatus</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B98465B3D5A7E36FF09FAAFB67625CC" blockId="10.[151,1437,1355,2016]" pageId="10" pageNumber="59">Ventral head scalation heterogenous; mental bordered by first infralabials, and paired postmentals, with triangular projection posteriorly; 7 differentiated gular scales posterior to postmentals; first infralabial in contact with postmental and single differentiated gular; remaining gulars heterogenous, small and granular, increasing in size in nuchal region.</paragraph>
<paragraph id="8B98465B3D5A7E36FF09FA05B02D273C" blockId="10.[151,1437,1355,2016]" pageId="10" pageNumber="59">Body elongate, axillagroin distance 44.5% snoutvent length, lacking distinct ventrolateral folds; dorsal scalation granular, heterogenous, with semi-regular rows of rounded, posteriorly-oriented tubercles, increasing in protuberance (from convex to conical) and size posterior to hind limb insertions; tubercles in 17 longitudinal midbody and 27 paravertebral rows; paravertebral scales 164; transverse midbody dorsal and ventral scales 91 and 54 respectively, between lateral tubercle rows; ventral scales imbricate, slightly larger than dorsals and increasing in size medially; differentiated precloacals 4, arranged in an inverted “V” configuration pierced with enlarged pores, surrounding shallow and longitudinally short precloacal groove; two patches of enlarged scales anterolateral to precloacals (14 left, 13 right); two rows of slightly enlarged scales anterior to precloacals; three slightly enlarged scales posterior to precloacals.</paragraph>
<paragraph id="8B98465B3D5A7E37FF09F8D5B15E20CE" blockId="10.[151,1437,1355,2016]" lastBlockId="11.[151,1436,152,215]" lastPageId="11" lastPageNumber="60" pageId="10" pageNumber="59">Forelimbs slender, forearm and upper arm 14.3% and 12.4% snoutvent length respectively; scalation on dorsal surfaces of forelimbs heterogenous, scales larger than ventrals; tubercles absent on upper arm, sparse on forearm, less spinose than dorsal trunk tubercles; ventral scalation homogenous, lacking tubercles; fingers well developed; Fingers I and III 37.4% and 62.2% forearm length, respectively; lamellae enlarged and slightly raised, lamellae proximal to inflection larger than those distal to inflection, slight increase in size from inflection to claw; finger number followed by subdigital lamellae (in parentheses): I (11), II (14), III (20), IV (18), V (15); all fingers clawed; claws well developed, sheathed by single dorsal and ventral scale.</paragraph>
<caption id="DF5816D33D5B7E37FF59FEE3B0732186" ID-Table-UUID="DF5816D33D5B7E37FF59FEE3B0732186" httpUri="http://table.plazi.org/id/DF5816D33D5B7E37FF59FEE3B0732186" pageId="11" pageNumber="60" targetBox="[161,1410,450,2013]" targetIsTable="true" targetPageId="11">
<paragraph id="8B98465B3D5B7E37FF59FEE3B0732186" blockId="11.[151,1436,263,415]" pageId="11" pageNumber="60">
<emphasis id="B9539A493D5B7E37FF59FEE3B54F2106" bold="true" box="[151,265,263,287]" pageId="11" pageNumber="60">TABLE 4.</emphasis>
Summary of univariate morphological variation among mensural and meristic characters in the type series of
<taxonomicName id="4C273DD83D5B7E37FF59FECCB5F62126" box="[151,432,296,319]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="60" phylum="Chordata" rank="species" species="jambangan" status="sp. nov.">
<emphasis id="B9539A493D5B7E37FF59FECCB5F62126" box="[151,432,296,319]" italics="true" pageId="11" pageNumber="60">Cyrtodactylus jambangan</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26027323D5B7E37FE70FECCB5BD2126" box="[446,507,296,319]" pageId="11" pageNumber="60" rank="species">n. sp.</taxonomicNameLabel>
, from Pasonanca Natural Park, Zamboanga Peninsula, western Mindanao Island, and
<taxonomicName id="4C273DD83D5B7E37FF59FEACB5E32146" box="[151,421,328,351]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="60" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D5B7E37FF59FEACB5E32146" box="[151,421,328,351]" italics="true" pageId="11" pageNumber="60">Cyrtodactylus annulatus</emphasis>
</taxonomicName>
from eastern Mindanao Island (including Bunawan, the
<taxonomicName id="4C273DD83D5B7E37FBD6FEACB0E32146" box="[1048,1189,328,351]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="60" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D5B7E37FBD6FEACB0E32146" box="[1048,1189,328,351]" italics="true" pageId="11" pageNumber="60">C. annulatus</emphasis>
</taxonomicName>
type locality). Labial scale counts were counted to below the center of the eye and bilaterally symmetrical characters are separated into left (l) and right (r) sides of the body. See Materials and Methods for character abbreviations.
</paragraph>
</caption>
<paragraph id="8B98465B3D5B7E37FE61FE26B17427C4" pageId="11" pageNumber="60">
<table id="F927B4FB3D5B81C3FF6FFE26B1C427C4" box="[161,1410,450,2013]" gridcols="5" gridrows="35" pageId="11" pageNumber="60">
<tr id="351744193D5B81C3FF6FFE26B1C421CE" box="[161,1410,450,471]" gridrow="0" pageId="11" pageNumber="60" rowspan-0="1">
<th id="76C62D653D5B81C3FE61FE26B6C321CE" box="[431,645,450,471]" gridcol="1" gridrow="0" pageId="11" pageNumber="60">
<taxonomicName id="4C273DD83D5B7E37FE61FE26B65D21CE" box="[431,539,450,471]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="60" phylum="Chordata" rank="species" species="jambangan">
<emphasis id="B9539A493D5B7E37FE61FE26B65D21CE" box="[431,539,450,471]" italics="true" pageId="11" pageNumber="60">jambangan</emphasis>
</taxonomicName>
</th>
<th id="76C62D653D5B81C3FD7FFE26B7CE21CE" box="[689,904,450,471]" gridcol="2" gridrow="0" pageId="11" pageNumber="60">
<taxonomicName id="4C273DD83D5B7E37FD7FFE26B75B21CE" box="[689,797,450,471]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="60" phylum="Chordata" rank="species" species="jambangan">
<emphasis id="B9539A493D5B7E37FD7FFE26B75B21CE" box="[689,797,450,471]" italics="true" pageId="11" pageNumber="60">jambangan</emphasis>
</taxonomicName>
</th>
<th id="76C62D653D5B81C3FC62FE26B0C221CE" box="[940,1156,450,471]" gridcol="3" gridrow="0" pageId="11" pageNumber="60">
<taxonomicName id="4C273DD83D5B7E37FC62FE26B04C21CE" box="[940,1034,450,471]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="60" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D5B7E37FC62FE26B04C21CE" box="[940,1034,450,471]" italics="true" pageId="11" pageNumber="60">annulatus</emphasis>
</taxonomicName>
</th>
<th id="76C62D653D5B81C3FB64FE26B1C421CE" box="[1194,1410,450,471]" gridcol="4" gridrow="0" pageId="11" pageNumber="60">
<taxonomicName id="4C273DD83D5B7E37FB64FE26B14F21CE" box="[1194,1289,450,471]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="60" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D5B7E37FB64FE26B14F21CE" box="[1194,1289,450,471]" italics="true" pageId="11" pageNumber="60">annulatus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="351744193D5B81C3FF6FFE0BB1C4221D" box="[161,1410,495,516]" gridrow="1" pageId="11" pageNumber="60" rowspan-0="1">
<td id="76C62D653D5B81C3FE61FE0BB6C3221D" box="[431,645,495,516]" gridcol="1" gridrow="1" pageId="11" pageNumber="60">Male (N=10)</td>
<td id="76C62D653D5B81C3FD7FFE0BB7CE221D" box="[689,904,495,516]" gridcol="2" gridrow="1" pageId="11" pageNumber="60">Female (N=10)</td>
<td id="76C62D653D5B81C3FC62FE0BB0C2221D" box="[940,1156,495,516]" gridcol="3" gridrow="1" pageId="11" pageNumber="60">Male (N=4)</td>
<td id="76C62D653D5B81C3FB64FE0BB1C4221D" box="[1194,1410,495,516]" gridcol="4" gridrow="1" pageId="11" pageNumber="60">Female (N=6)</td>
</tr>
<tr id="351744193D5B81C3FF6FFDF9B1C4222B" box="[161,1410,541,562]" gridrow="2" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFDF9B5CF222B" box="[161,393,541,562]" gridcol="0" gridrow="2" pageId="11" pageNumber="60">Snout-vent length</th>
<td id="76C62D653D5B81C3FE61FDF9B6C3222B" box="[431,645,541,562]" gridcol="1" gridrow="2" pageId="11" pageNumber="60">60.472.3 (67.1 ± 3.8)</td>
<td id="76C62D653D5B81C3FD7FFDF9B7CE222B" box="[689,904,541,562]" gridcol="2" gridrow="2" pageId="11" pageNumber="60">61.281.5 (71.1 ± 5.9)</td>
<td id="76C62D653D5B81C3FC62FDF9B0C2222B" box="[940,1156,541,562]" gridcol="3" gridrow="2" pageId="11" pageNumber="60">68.879.4 (74.1 ± 5.6)</td>
<td id="76C62D653D5B81C3FB64FDF9B1C4222B" box="[1194,1410,541,562]" gridcol="4" gridrow="2" pageId="11" pageNumber="60">59.380.5 (67.3 ± 8.3)</td>
</tr>
<tr id="351744193D5B81C3FF6FFDAEB1C42246" box="[161,1410,586,607]" gridrow="3" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFDAEB5CF2246" box="[161,393,586,607]" gridcol="0" gridrow="3" pageId="11" pageNumber="60">Lamellae of Finger 3</th>
<td id="76C62D653D5B81C3FE61FDAEB6C32246" box="[431,645,586,607]" gridcol="1" gridrow="3" pageId="11" pageNumber="60">1720 (18 ± 0.9)</td>
<td id="76C62D653D5B81C3FD7FFDAEB7CE2246" box="[689,904,586,607]" gridcol="2" gridrow="3" pageId="11" pageNumber="60">1622 (18 ± 1.7)</td>
<td id="76C62D653D5B81C3FC62FDAEB0C22246" box="[940,1156,586,607]" gridcol="3" gridrow="3" pageId="11" pageNumber="60">1721 (19 ± 1.0)</td>
<td id="76C62D653D5B81C3FB64FDAEB1C42246" box="[1194,1410,586,607]" gridcol="4" gridrow="3" pageId="11" pageNumber="60">1518 (17 ± 1.2)</td>
</tr>
<tr id="351744193D5B81C3FF6FFD93B1C42295" box="[161,1410,631,652]" gridrow="4" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFD93B5CF2295" box="[161,393,631,652]" gridcol="0" gridrow="4" pageId="11" pageNumber="60">Lamellae of Toe 4</th>
<td id="76C62D653D5B81C3FE61FD93B6C32295" box="[431,645,631,652]" gridcol="1" gridrow="4" pageId="11" pageNumber="60">2023 (21 ± 1.0)</td>
<td id="76C62D653D5B81C3FD7FFD93B7CE2295" box="[689,904,631,652]" gridcol="2" gridrow="4" pageId="11" pageNumber="60">2024 (22 ± 1.3)</td>
<td id="76C62D653D5B81C3FC62FD93B0C22295" box="[940,1156,631,652]" gridcol="3" gridrow="4" pageId="11" pageNumber="60">2022 (21 ± 1.0)</td>
<td id="76C62D653D5B81C3FB64FD93B1C42295" box="[1194,1410,631,652]" gridcol="4" gridrow="4" pageId="11" pageNumber="60">1721 (19 ± 1.5)</td>
</tr>
<tr id="351744193D5B81C3FF6FFD41B1C422A3" box="[161,1410,677,698]" gridrow="5" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFD41B5CF22A3" box="[161,393,677,698]" gridcol="0" gridrow="5" pageId="11" pageNumber="60">Finger 1 length</th>
<td id="76C62D653D5B81C3FE61FD41B6C322A3" box="[431,645,677,698]" gridcol="1" gridrow="5" pageId="11" pageNumber="60">2.73.7 (3.3 ± 0.4)</td>
<td id="76C62D653D5B81C3FD7FFD41B7CE22A3" box="[689,904,677,698]" gridcol="2" gridrow="5" pageId="11" pageNumber="60">2.64.1 (3.4 ± 0.4)</td>
<td id="76C62D653D5B81C3FC62FD41B0C222A3" box="[940,1156,677,698]" gridcol="3" gridrow="5" pageId="11" pageNumber="60">3.24.1 (3.8 ± 0.4)</td>
<td id="76C62D653D5B81C3FB64FD41B1C422A3" box="[1194,1410,677,698]" gridcol="4" gridrow="5" pageId="11" pageNumber="60">2.64.0 (3.2 ± 0.6)</td>
</tr>
<tr id="351744193D5B81C3FF6FFD36B1C422FE" box="[161,1410,722,743]" gridrow="6" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFD36B5CF22FE" box="[161,393,722,743]" gridcol="0" gridrow="6" pageId="11" pageNumber="60">Finger 3 length</th>
<td id="76C62D653D5B81C3FE61FD36B6C322FE" box="[431,645,722,743]" gridcol="1" gridrow="6" pageId="11" pageNumber="60">4.36.0 (5.4 ± 0.5)</td>
<td id="76C62D653D5B81C3FD7FFD36B7CE22FE" box="[689,904,722,743]" gridcol="2" gridrow="6" pageId="11" pageNumber="60">4.47.0 (5.5 ± 0.6)</td>
<td id="76C62D653D5B81C3FC62FD36B0C222FE" box="[940,1156,722,743]" gridcol="3" gridrow="6" pageId="11" pageNumber="60">4.06.2 (5.3 ± 1.1)</td>
<td id="76C62D653D5B81C3FB64FD36B1C422FE" box="[1194,1410,722,743]" gridcol="4" gridrow="6" pageId="11" pageNumber="60">4.05.9 (5.0 ± 0.8)</td>
</tr>
<tr id="351744193D5B81C3FF6FFD1BB1C4230D" box="[161,1410,767,788]" gridrow="7" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFD1BB5CF230D" box="[161,393,767,788]" gridcol="0" gridrow="7" pageId="11" pageNumber="60">Toe 1 length</th>
<td id="76C62D653D5B81C3FE61FD1BB6C3230D" box="[431,645,767,788]" gridcol="1" gridrow="7" pageId="11" pageNumber="60">2.93.7 (3.2 ± 0.3)</td>
<td id="76C62D653D5B81C3FD7FFD1BB7CE230D" box="[689,904,767,788]" gridcol="2" gridrow="7" pageId="11" pageNumber="60">2.53.9 (3.4 ± 0.4)</td>
<td id="76C62D653D5B81C3FC62FD1BB0C2230D" box="[940,1156,767,788]" gridcol="3" gridrow="7" pageId="11" pageNumber="60">3.03.9 (3.4 ± 0.4)</td>
<td id="76C62D653D5B81C3FB64FD1BB1C4230D" box="[1194,1410,767,788]" gridcol="4" gridrow="7" pageId="11" pageNumber="60">2.03.8 (3.2 ± 0.7)</td>
</tr>
<tr id="351744193D5B81C3FF6FFCC9B1C4235B" box="[161,1410,813,834]" gridrow="8" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFCC9B5CF235B" box="[161,393,813,834]" gridcol="0" gridrow="8" pageId="11" pageNumber="60">Toe 4 length</th>
<td id="76C62D653D5B81C3FE61FCC9B6C3235B" box="[431,645,813,834]" gridcol="1" gridrow="8" pageId="11" pageNumber="60">6.89.0 (7.8 ± 0.7)</td>
<td id="76C62D653D5B81C3FD7FFCC9B7CE235B" box="[689,904,813,834]" gridcol="2" gridrow="8" pageId="11" pageNumber="60">6.910.3 (8.6 ± 1.1)</td>
<td id="76C62D653D5B81C3FC62FCC9B0C2235B" box="[940,1156,813,834]" gridcol="3" gridrow="8" pageId="11" pageNumber="60">6.69.7 (8.1 ± 1.4)</td>
<td id="76C62D653D5B81C3FB64FCC9B1C4235B" box="[1194,1410,813,834]" gridcol="4" gridrow="8" pageId="11" pageNumber="60">5.58.7 (7.4 ± 1.2)</td>
</tr>
<tr id="351744193D5B81C3FF6FFCBEB1C42376" box="[161,1410,858,879]" gridrow="9" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFCBEB5CF2376" box="[161,393,858,879]" gridcol="0" gridrow="9" pageId="11" pageNumber="60">Supralabials (l)</th>
<td id="76C62D653D5B81C3FE61FCBEB6C32376" box="[431,645,858,879]" gridcol="1" gridrow="9" pageId="11" pageNumber="60">811 (9 ± 1.0)</td>
<td id="76C62D653D5B81C3FD7FFCBEB7CE2376" box="[689,904,858,879]" gridcol="2" gridrow="9" pageId="11" pageNumber="60">810 (9 ± 0.7)</td>
<td id="76C62D653D5B81C3FC62FCBEB0C22376" box="[940,1156,858,879]" gridcol="3" gridrow="9" pageId="11" pageNumber="60">810 (9 ± 1.0)</td>
<td id="76C62D653D5B81C3FB64FCBEB1C42376" box="[1194,1410,858,879]" gridcol="4" gridrow="9" pageId="11" pageNumber="60">710 (9 ± 1.2)</td>
</tr>
<tr id="351744193D5B81C3FF6FFC6CB1C42384" box="[161,1410,904,925]" gridrow="10" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFC6CB5CF2384" box="[161,393,904,925]" gridcol="0" gridrow="10" pageId="11" pageNumber="60">Supralabials (r)</th>
<td id="76C62D653D5B81C3FE61FC6CB6C32384" box="[431,645,904,925]" gridcol="1" gridrow="10" pageId="11" pageNumber="60">810 (9 ± 0.6)</td>
<td id="76C62D653D5B81C3FD7FFC6CB7CE2384" box="[689,904,904,925]" gridcol="2" gridrow="10" pageId="11" pageNumber="60">810 (9 ± 0.6)</td>
<td id="76C62D653D5B81C3FC62FC6CB0C22384" box="[940,1156,904,925]" gridcol="3" gridrow="10" pageId="11" pageNumber="60">810 (9 ± 1.0)</td>
<td id="76C62D653D5B81C3FB64FC6CB1C42384" box="[1194,1410,904,925]" gridcol="4" gridrow="10" pageId="11" pageNumber="60">710 (9 ± 0.8)</td>
</tr>
<tr id="351744193D5B81C3FF6FFC51B1C423D3" box="[161,1410,949,970]" gridrow="11" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFC51B5CF23D3" box="[161,393,949,970]" gridcol="0" gridrow="11" pageId="11" pageNumber="60">Infralabials (l)</th>
<td id="76C62D653D5B81C3FE61FC51B6C323D3" box="[431,645,949,970]" gridcol="1" gridrow="11" pageId="11" pageNumber="60">68 (7 ± 0.5)</td>
<td id="76C62D653D5B81C3FD7FFC51B7CE23D3" box="[689,904,949,970]" gridcol="2" gridrow="11" pageId="11" pageNumber="60">78 (7 ± 0.3)</td>
<td id="76C62D653D5B81C3FC62FC51B0C223D3" box="[940,1156,949,970]" gridcol="3" gridrow="11" pageId="11" pageNumber="60">78 (7 ± 1.0)</td>
<td id="76C62D653D5B81C3FB64FC51B1C423D3" box="[1194,1410,949,970]" gridcol="4" gridrow="11" pageId="11" pageNumber="60">78 (8 ± 0.5)</td>
</tr>
<tr id="351744193D5B81C3FF6FFC06B1C423EE" box="[161,1410,994,1015]" gridrow="12" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFC06B5CF23EE" box="[161,393,994,1015]" gridcol="0" gridrow="12" pageId="11" pageNumber="60">Infralabials (r)</th>
<td id="76C62D653D5B81C3FE61FC06B6C323EE" box="[431,645,994,1015]" gridcol="1" gridrow="12" pageId="11" pageNumber="60">68 (7 ± 0.6)</td>
<td id="76C62D653D5B81C3FD7FFC06B7CE23EE" box="[689,904,994,1015]" gridcol="2" gridrow="12" pageId="11" pageNumber="60">68 (7 ± 0.5)</td>
<td id="76C62D653D5B81C3FC62FC06B0C223EE" box="[940,1156,994,1015]" gridcol="3" gridrow="12" pageId="11" pageNumber="60">78 (7 ± 1.0)</td>
<td id="76C62D653D5B81C3FB64FC06B1C423EE" box="[1194,1410,994,1015]" gridcol="4" gridrow="12" pageId="11" pageNumber="60">78 (7 ± 0.5)</td>
</tr>
<tr id="351744193D5B81C3FF6FFBF4B1C4243C" box="[161,1410,1040,1061]" gridrow="13" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFBF4B5CF243C" box="[161,393,1040,1061]" gridcol="0" gridrow="13" pageId="11" pageNumber="60">Precloacals (l)</th>
<td id="76C62D653D5B81C3FE61FBF4B6C3243C" box="[431,645,1040,1061]" gridcol="1" gridrow="13" pageId="11" pageNumber="60">2 or 3 (2.2 ± 0.4)</td>
<td id="76C62D653D5B81C3FD7FFBF4B7CE243C" box="[689,904,1040,1061]" gridcol="2" gridrow="13" pageId="11" pageNumber="60">N/A</td>
<td id="76C62D653D5B81C3FC62FBF4B0C2243C" box="[940,1156,1040,1061]" gridcol="3" gridrow="13" pageId="11" pageNumber="60">2 or 3</td>
<td id="76C62D653D5B81C3FB64FBF4B1C4243C" box="[1194,1410,1040,1061]" gridcol="4" gridrow="13" pageId="11" pageNumber="60">N/A</td>
</tr>
<tr id="351744193D5B81C3FF6FFBD9B1C4244B" box="[161,1410,1085,1106]" gridrow="14" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFBD9B5CF244B" box="[161,393,1085,1106]" gridcol="0" gridrow="14" pageId="11" pageNumber="60">Precloacals (r)</th>
<td id="76C62D653D5B81C3FE61FBD9B6C3244B" box="[431,645,1085,1106]" gridcol="1" gridrow="14" pageId="11" pageNumber="60">02 (1.2 ± 0.6)</td>
<td id="76C62D653D5B81C3FD7FFBD9B7CE244B" box="[689,904,1085,1106]" gridcol="2" gridrow="14" pageId="11" pageNumber="60">N/A</td>
<td id="76C62D653D5B81C3FC62FBD9B0C2244B" box="[940,1156,1085,1106]" gridcol="3" gridrow="14" pageId="11" pageNumber="60">3</td>
<td id="76C62D653D5B81C3FB64FBD9B1C4244B" box="[1194,1410,1085,1106]" gridcol="4" gridrow="14" pageId="11" pageNumber="60">N/A</td>
</tr>
<tr id="351744193D5B81C3FF6FFB8EB1C42466" box="[161,1410,1130,1151]" gridrow="15" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFB8EB5CF2466" box="[161,393,1130,1151]" gridcol="0" gridrow="15" pageId="11" pageNumber="60">Total precloacals</th>
<td id="76C62D653D5B81C3FE61FB8EB6C32466" box="[431,645,1130,1151]" gridcol="1" gridrow="15" pageId="11" pageNumber="60">25 (3.4 ± 0.8)</td>
<td id="76C62D653D5B81C3FD7FFB8EB7CE2466" box="[689,904,1130,1151]" gridcol="2" gridrow="15" pageId="11" pageNumber="60">N/A</td>
<td id="76C62D653D5B81C3FC62FB8EB0C22466" box="[940,1156,1130,1151]" gridcol="3" gridrow="15" pageId="11" pageNumber="60">5 or 6</td>
<td id="76C62D653D5B81C3FB64FB8EB1C42466" box="[1194,1410,1130,1151]" gridcol="4" gridrow="15" pageId="11" pageNumber="60">N/A</td>
</tr>
<tr id="351744193D5B81C3FF6FFB7CB1C424B4" box="[161,1410,1176,1197]" gridrow="16" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFB7CB5CF24B4" box="[161,393,1176,1197]" gridcol="0" gridrow="16" pageId="11" pageNumber="60">Midbody dorsals</th>
<td id="76C62D653D5B81C3FE61FB7CB6C324B4" box="[431,645,1176,1197]" gridcol="1" gridrow="16" pageId="11" pageNumber="60">7591 (87 ± 4.6)</td>
<td id="76C62D653D5B81C3FD7FFB7CB7CE24B4" box="[689,904,1176,1197]" gridcol="2" gridrow="16" pageId="11" pageNumber="60">8696 (93 ± 3.1)</td>
<td id="76C62D653D5B81C3FC62FB7CB0C224B4" box="[940,1156,1176,1197]" gridcol="3" gridrow="16" pageId="11" pageNumber="60">8795 (91 ± 3.7)</td>
<td id="76C62D653D5B81C3FB64FB7CB1C424B4" box="[1194,1410,1176,1197]" gridcol="4" gridrow="16" pageId="11" pageNumber="60">8191 (85 ± 4.0)</td>
</tr>
<tr id="351744193D5B81C3FF6FFB21B1C424C3" box="[161,1410,1221,1242]" gridrow="17" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFB21B5CF24C3" box="[161,393,1221,1242]" gridcol="0" gridrow="17" pageId="11" pageNumber="60">Midbody tubercles</th>
<td id="76C62D653D5B81C3FE61FB21B6C324C3" box="[431,645,1221,1242]" gridcol="1" gridrow="17" pageId="11" pageNumber="60">1719 (18 ± 0.6)</td>
<td id="76C62D653D5B81C3FD7FFB21B7CE24C3" box="[689,904,1221,1242]" gridcol="2" gridrow="17" pageId="11" pageNumber="60">1618 (17 ± 0.9)</td>
<td id="76C62D653D5B81C3FC62FB21B0C224C3" box="[940,1156,1221,1242]" gridcol="3" gridrow="17" pageId="11" pageNumber="60">1416 (15.5 ± 1.3)</td>
<td id="76C62D653D5B81C3FB64FB21B1C424C3" box="[1194,1410,1221,1242]" gridcol="4" gridrow="17" pageId="11" pageNumber="60">1416 (15 ± 1.0)</td>
</tr>
<tr id="351744193D5B81C3FF6FFB17B1C42511" box="[161,1410,1267,1288]" gridrow="18" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFB17B5CF2511" box="[161,393,1267,1288]" gridcol="0" gridrow="18" pageId="11" pageNumber="60">Midbody ventrals</th>
<td id="76C62D653D5B81C3FE61FB17B6C32511" box="[431,645,1267,1288]" gridcol="1" gridrow="18" pageId="11" pageNumber="60">4859 (53 ± 3.6)</td>
<td id="76C62D653D5B81C3FD7FFB17B7CE2511" box="[689,904,1267,1288]" gridcol="2" gridrow="18" pageId="11" pageNumber="60">5063 (57 ± 4.6)</td>
<td id="76C62D653D5B81C3FC62FB17B0C22511" box="[940,1156,1267,1288]" gridcol="3" gridrow="18" pageId="11" pageNumber="60">4954 (51 ± 2.5)</td>
<td id="76C62D653D5B81C3FB64FB17B1C42511" box="[1194,1410,1267,1288]" gridcol="4" gridrow="18" pageId="11" pageNumber="60">5060 (53 ± 3.7)</td>
</tr>
<tr id="351744193D5B81C3FF6FFAC4B1C4252C" box="[161,1410,1312,1333]" gridrow="19" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFAC4B5CF252C" box="[161,393,1312,1333]" gridcol="0" gridrow="19" pageId="11" pageNumber="60">Paravertebrals</th>
<td id="76C62D653D5B81C3FE61FAC4B6C3252C" box="[431,645,1312,1333]" gridcol="1" gridrow="19" pageId="11" pageNumber="60">144174 (161 ± 8.4)</td>
<td id="76C62D653D5B81C3FD7FFAC4B7CE252C" box="[689,904,1312,1333]" gridcol="2" gridrow="19" pageId="11" pageNumber="60">150170 (163 ± 6.2)</td>
<td id="76C62D653D5B81C3FC62FAC4B0C2252C" box="[940,1156,1312,1333]" gridcol="3" gridrow="19" pageId="11" pageNumber="60">124166 (152 ± 18.9)</td>
<td id="76C62D653D5B81C3FB64FAC4B1C4252C" box="[1194,1410,1312,1333]" gridcol="4" gridrow="19" pageId="11" pageNumber="60">137163 (151 ± 12.5)</td>
</tr>
<tr id="351744193D5B81C3FF6FFAA9B1C4257B" box="[161,1410,1357,1378]" gridrow="20" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFAA9B5CF257B" box="[161,393,1357,1378]" gridcol="0" gridrow="20" pageId="11" pageNumber="60">Paravertebral tubercles</th>
<td id="76C62D653D5B81C3FE61FAA9B6C3257B" box="[431,645,1357,1378]" gridcol="1" gridrow="20" pageId="11" pageNumber="60">2530 (27 ± 1.8)</td>
<td id="76C62D653D5B81C3FD7FFAA9B7CE257B" box="[689,904,1357,1378]" gridcol="2" gridrow="20" pageId="11" pageNumber="60">2331 (27 ± 2.7)</td>
<td id="76C62D653D5B81C3FC62FAA9B0C2257B" box="[940,1156,1357,1378]" gridcol="3" gridrow="20" pageId="11" pageNumber="60">1727 (24 ± 4.7)</td>
<td id="76C62D653D5B81C3FB64FAA9B1C4257B" box="[1194,1410,1357,1378]" gridcol="4" gridrow="20" pageId="11" pageNumber="60">2128 (24 ± 2.4)</td>
</tr>
<tr id="351744193D5B81C3FF6FFA9FB1C42589" box="[161,1410,1403,1424]" gridrow="21" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFA9FB5CF2589" box="[161,393,1403,1424]" gridcol="0" gridrow="21" pageId="11" pageNumber="60">Eue-narial distance</th>
<td id="76C62D653D5B81C3FE61FA9FB6C32589" box="[431,645,1403,1424]" gridcol="1" gridrow="21" pageId="11" pageNumber="60">5.97.4 (6.5 ± 0.5)</td>
<td id="76C62D653D5B81C3FD7FFA9FB7CE2589" box="[689,904,1403,1424]" gridcol="2" gridrow="21" pageId="11" pageNumber="60">6.08.1 (7.0 ± 0.6)</td>
<td id="76C62D653D5B81C3FC62FA9FB0C22589" box="[940,1156,1403,1424]" gridcol="3" gridrow="21" pageId="11" pageNumber="60">5.47.7 (6.7 ± 1.1)</td>
<td id="76C62D653D5B81C3FB64FA9FB1C42589" box="[1194,1410,1403,1424]" gridcol="4" gridrow="21" pageId="11" pageNumber="60">4.67.5 (6.0 ± 1.1)</td>
</tr>
<tr id="351744193D5B81C3FF6FFA4CB1C425A4" box="[161,1410,1448,1469]" gridrow="22" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFA4CB5CF25A4" box="[161,393,1448,1469]" gridcol="0" gridrow="22" pageId="11" pageNumber="60">Eye-tympanum distance</th>
<td id="76C62D653D5B81C3FE61FA4CB6C325A4" box="[431,645,1448,1469]" gridcol="1" gridrow="22" pageId="11" pageNumber="60">5.36.5 (5.6 ± 0.4)</td>
<td id="76C62D653D5B81C3FD7FFA4CB7CE25A4" box="[689,904,1448,1469]" gridcol="2" gridrow="22" pageId="11" pageNumber="60">5.87.7 (6.4 ± 0.6)</td>
<td id="76C62D653D5B81C3FC62FA4CB0C225A4" box="[940,1156,1448,1469]" gridcol="3" gridrow="22" pageId="11" pageNumber="60">5.46.6 (6.0 ± 0.5)</td>
<td id="76C62D653D5B81C3FB64FA4CB1C425A4" box="[1194,1410,1448,1469]" gridcol="4" gridrow="22" pageId="11" pageNumber="60">5.06.4 (5.7 ± 0.6)</td>
</tr>
<tr id="351744193D5B81C3FF6FFA31B1C425F3" box="[161,1410,1493,1514]" gridrow="23" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FFA31B5CF25F3" box="[161,393,1493,1514]" gridcol="0" gridrow="23" pageId="11" pageNumber="60">Interorbital distance</th>
<td id="76C62D653D5B81C3FE61FA31B6C325F3" box="[431,645,1493,1514]" gridcol="1" gridrow="23" pageId="11" pageNumber="60">2.23.0 (2.6 ± 0.2)</td>
<td id="76C62D653D5B81C3FD7FFA31B7CE25F3" box="[689,904,1493,1514]" gridcol="2" gridrow="23" pageId="11" pageNumber="60">2.43.7 (3.0 ± 0.4)</td>
<td id="76C62D653D5B81C3FC62FA31B0C225F3" box="[940,1156,1493,1514]" gridcol="3" gridrow="23" pageId="11" pageNumber="60">2.93.9 (3.3 ± 0.5)</td>
<td id="76C62D653D5B81C3FB64FA31B1C425F3" box="[1194,1410,1493,1514]" gridcol="4" gridrow="23" pageId="11" pageNumber="60">2.63.4 (2.9 ± 0.3)</td>
</tr>
<tr id="351744193D5B81C3FF6FF9E7B1C42601" box="[161,1410,1539,1560]" gridrow="24" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FF9E7B5CF2601" box="[161,393,1539,1560]" gridcol="0" gridrow="24" pageId="11" pageNumber="60">Snout length</th>
<td id="76C62D653D5B81C3FE61F9E7B6C32601" box="[431,645,1539,1560]" gridcol="1" gridrow="24" pageId="11" pageNumber="60">6.38.7 (7.8 ± 0.8)</td>
<td id="76C62D653D5B81C3FD7FF9E7B7CE2601" box="[689,904,1539,1560]" gridcol="2" gridrow="24" pageId="11" pageNumber="60">7.19.5 (8.4 ± 0.7)</td>
<td id="76C62D653D5B81C3FC62F9E7B0C22601" box="[940,1156,1539,1560]" gridcol="3" gridrow="24" pageId="11" pageNumber="60">6.59.3 (8.2 ± 1.3)</td>
<td id="76C62D653D5B81C3FB64F9E7B1C42601" box="[1194,1410,1539,1560]" gridcol="4" gridrow="24" pageId="11" pageNumber="60">5.58.9 (7.0 ± 1.4)</td>
</tr>
<tr id="351744193D5B81C3FF6FF9D4B1C4265C" box="[161,1410,1584,1605]" gridrow="25" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FF9D4B5CF265C" box="[161,393,1584,1605]" gridcol="0" gridrow="25" pageId="11" pageNumber="60">Internarial distance</th>
<td id="76C62D653D5B81C3FE61F9D4B6C3265C" box="[431,645,1584,1605]" gridcol="1" gridrow="25" pageId="11" pageNumber="60">1.92.5 (2.2 ± 0.2)</td>
<td id="76C62D653D5B81C3FD7FF9D4B7CE265C" box="[689,904,1584,1605]" gridcol="2" gridrow="25" pageId="11" pageNumber="60">2.03.0 (2.5 ± 0.3)</td>
<td id="76C62D653D5B81C3FC62F9D4B0C2265C" box="[940,1156,1584,1605]" gridcol="3" gridrow="25" pageId="11" pageNumber="60">2.22.5 (2.4 ± 0.1)</td>
<td id="76C62D653D5B81C3FB64F9D4B1C4265C" box="[1194,1410,1584,1605]" gridcol="4" gridrow="25" pageId="11" pageNumber="60">2.12.6 (2.3 ± 0.2)</td>
</tr>
<tr id="351744193D5B81C3FF6FF9B9B1C4266B" box="[161,1410,1629,1650]" gridrow="26" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FF9B9B5CF266B" box="[161,393,1629,1650]" gridcol="0" gridrow="26" pageId="11" pageNumber="60">Eye diameter</th>
<td id="76C62D653D5B81C3FE61F9B9B6C3266B" box="[431,645,1629,1650]" gridcol="1" gridrow="26" pageId="11" pageNumber="60">3.84.6 (4.2 ± 0.3)</td>
<td id="76C62D653D5B81C3FD7FF9B9B7CE266B" box="[689,904,1629,1650]" gridcol="2" gridrow="26" pageId="11" pageNumber="60">3.75.1 (4.5 ± 0.5)</td>
<td id="76C62D653D5B81C3FC62F9B9B0C2266B" box="[940,1156,1629,1650]" gridcol="3" gridrow="26" pageId="11" pageNumber="60">4.34.8 (4.6 ± 0.2)</td>
<td id="76C62D653D5B81C3FB64F9B9B1C4266B" box="[1194,1410,1629,1650]" gridcol="4" gridrow="26" pageId="11" pageNumber="60">3.65.2 (4.3 ± 0.6)</td>
</tr>
<tr id="351744193D5B81C3FF6FF96FB1C426B9" box="[161,1410,1675,1696]" gridrow="27" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FF96FB5CF26B9" box="[161,393,1675,1696]" gridcol="0" gridrow="27" pageId="11" pageNumber="60">Upper arm length</th>
<td id="76C62D653D5B81C3FE61F96FB6C326B9" box="[431,645,1675,1696]" gridcol="1" gridrow="27" pageId="11" pageNumber="60">6.89.9 (7.8 ± 0.9)</td>
<td id="76C62D653D5B81C3FD7FF96FB7CE26B9" box="[689,904,1675,1696]" gridcol="2" gridrow="27" pageId="11" pageNumber="60">6.910.7 (9.0 ± 1.0)</td>
<td id="76C62D653D5B81C3FC62F96FB0C226B9" box="[940,1156,1675,1696]" gridcol="3" gridrow="27" pageId="11" pageNumber="60">6.29.6 (7.9 ± 1.5)</td>
<td id="76C62D653D5B81C3FB64F96FB1C426B9" box="[1194,1410,1675,1696]" gridcol="4" gridrow="27" pageId="11" pageNumber="60">5.110.4 (7.4 ± 2.2)</td>
</tr>
<tr id="351744193D5B81C3FF6FF95CB1C426D4" box="[161,1410,1720,1741]" gridrow="28" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FF95CB5CF26D4" box="[161,393,1720,1741]" gridcol="0" gridrow="28" pageId="11" pageNumber="60">Forearm length</th>
<td id="76C62D653D5B81C3FE61F95CB6C326D4" box="[431,645,1720,1741]" gridcol="1" gridrow="28" pageId="11" pageNumber="60">8.19.8 (9.0 ± 0.6)</td>
<td id="76C62D653D5B81C3FD7FF95CB7CE26D4" box="[689,904,1720,1741]" gridcol="2" gridrow="28" pageId="11" pageNumber="60">8.210.8 (9.6 ± 0.8)</td>
<td id="76C62D653D5B81C3FC62F95CB0C226D4" box="[940,1156,1720,1741]" gridcol="3" gridrow="28" pageId="11" pageNumber="60">8.611.1 (10.1 ± 1.2)</td>
<td id="76C62D653D5B81C3FB64F95CB1C426D4" box="[1194,1410,1720,1741]" gridcol="4" gridrow="28" pageId="11" pageNumber="60">7.611.5 (9.4 ± 1.4)</td>
</tr>
<tr id="351744193D5B81C3FF6FF902B1C426E2" box="[161,1410,1766,1787]" gridrow="29" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FF902B5CF26E2" box="[161,393,1766,1787]" gridcol="0" gridrow="29" pageId="11" pageNumber="60">Femur length</th>
<td id="76C62D653D5B81C3FE61F902B6C326E2" box="[431,645,1766,1787]" gridcol="1" gridrow="29" pageId="11" pageNumber="60">10.412.8 (11.7 ± 0.9)</td>
<td id="76C62D653D5B81C3FD7FF902B7CE26E2" box="[689,904,1766,1787]" gridcol="2" gridrow="29" pageId="11" pageNumber="60">9.514.9 (12.4 ± 1.4)</td>
<td id="76C62D653D5B81C3FC62F902B0C226E2" box="[940,1156,1766,1787]" gridcol="3" gridrow="29" pageId="11" pageNumber="60">11.413.3 (12.6 ± 0.9)</td>
<td id="76C62D653D5B81C3FB64F902B1C426E2" box="[1194,1410,1766,1787]" gridcol="4" gridrow="29" pageId="11" pageNumber="60">9.013.6 (11.6 ± 1.7)</td>
</tr>
<tr id="351744193D5B81C3FF6FF8F7B1C42731" box="[161,1410,1811,1832]" gridrow="30" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FF8F7B5CF2731" box="[161,393,1811,1832]" gridcol="0" gridrow="30" pageId="11" pageNumber="60">Tibia length</th>
<td id="76C62D653D5B81C3FE61F8F7B6C32731" box="[431,645,1811,1832]" gridcol="1" gridrow="30" pageId="11" pageNumber="60">9.411.1 (10.3 ± 0.5)</td>
<td id="76C62D653D5B81C3FD7FF8F7B7CE2731" box="[689,904,1811,1832]" gridcol="2" gridrow="30" pageId="11" pageNumber="60">8.713.6 (10.9 ± 1.2)</td>
<td id="76C62D653D5B81C3FC62F8F7B0C22731" box="[940,1156,1811,1832]" gridcol="3" gridrow="30" pageId="11" pageNumber="60">10.213.0 (11.4 ± 1.3)</td>
<td id="76C62D653D5B81C3FB64F8F7B1C42731" box="[1194,1410,1811,1832]" gridcol="4" gridrow="30" pageId="11" pageNumber="60">9.513.1 (10.7 ± 1.4)</td>
</tr>
<tr id="351744193D5B81C3FF6FF8A4B1C4274C" box="[161,1410,1856,1877]" gridrow="31" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FF8A4B5CF274C" box="[161,393,1856,1877]" gridcol="0" gridrow="31" pageId="11" pageNumber="60">Axilla-groin distance</th>
<td id="76C62D653D5B81C3FE61F8A4B6C3274C" box="[431,645,1856,1877]" gridcol="1" gridrow="31" pageId="11" pageNumber="60">26.332.8 (29.3 ± 2.3)</td>
<td id="76C62D653D5B81C3FD7FF8A4B7CE274C" box="[689,904,1856,1877]" gridcol="2" gridrow="31" pageId="11" pageNumber="60">27.734.6 (32.0 ± 2.1)</td>
<td id="76C62D653D5B81C3FC62F8A4B0C2274C" box="[940,1156,1856,1877]" gridcol="3" gridrow="31" pageId="11" pageNumber="60">26.331.2 (31.6 ± 4.1)</td>
<td id="76C62D653D5B81C3FB64F8A4B1C4274C" box="[1194,1410,1856,1877]" gridcol="4" gridrow="31" pageId="11" pageNumber="60">26.036.8 (29.9 ± 4.5)</td>
</tr>
<tr id="351744193D5B81C3FF6FF88AB1C4279A" box="[161,1410,1902,1923]" gridrow="32" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FF88AB5CF279A" box="[161,393,1902,1923]" gridcol="0" gridrow="32" pageId="11" pageNumber="60">Transverse bands</th>
<td id="76C62D653D5B81C3FE61F88AB6C3279A" box="[431,645,1902,1923]" gridcol="1" gridrow="32" pageId="11" pageNumber="60">35 (3 ± 0.7)</td>
<td id="76C62D653D5B81C3FD7FF88AB7CE279A" box="[689,904,1902,1923]" gridcol="2" gridrow="32" pageId="11" pageNumber="60">34 (3 ± 0.3)</td>
<td id="76C62D653D5B81C3FC62F88AB0C2279A" box="[940,1156,1902,1923]" gridcol="3" gridrow="32" pageId="11" pageNumber="60">3</td>
<td id="76C62D653D5B81C3FB64F88AB1C4279A" box="[1194,1410,1902,1923]" gridcol="4" gridrow="32" pageId="11" pageNumber="60">34 (3 ± 0.4)</td>
</tr>
<tr id="351744193D5B81C3FF6FF87FB1C427A9" box="[161,1410,1947,1968]" gridrow="33" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FF87FB5CF27A9" box="[161,393,1947,1968]" gridcol="0" gridrow="33" pageId="11" pageNumber="60">Postcloacal tubercles</th>
<td id="76C62D653D5B81C3FE61F87FB6C327A9" box="[431,645,1947,1968]" gridcol="1" gridrow="33" pageId="11" pageNumber="60">25 (4 ± 0.9)</td>
<td id="76C62D653D5B81C3FD7FF87FB7CE27A9" box="[689,904,1947,1968]" gridcol="2" gridrow="33" pageId="11" pageNumber="60">24 (4 ± 0.7)</td>
<td id="76C62D653D5B81C3FC62F87FB0C227A9" box="[940,1156,1947,1968]" gridcol="3" gridrow="33" pageId="11" pageNumber="60">24 (3 ± 1.2)</td>
<td id="76C62D653D5B81C3FB64F87FB1C427A9" box="[1194,1410,1947,1968]" gridcol="4" gridrow="33" pageId="11" pageNumber="60">24 (3 ± 1.0)</td>
</tr>
<tr id="351744193D5B81C3FF6FF82CB1C427C4" box="[161,1410,1992,2013]" gridrow="34" pageId="11" pageNumber="60">
<th id="76C62D653D5B81C3FF6FF82CB5CF27C4" box="[161,393,1992,2013]" gridcol="0" gridrow="34" pageId="11" pageNumber="60">Postementals</th>
<td id="76C62D653D5B81C3FE61F82CB6C327C4" box="[431,645,1992,2013]" gridcol="1" gridrow="34" pageId="11" pageNumber="60">69 (7 ± 1.0)</td>
<td id="76C62D653D5B81C3FD7FF82CB7CE27C4" box="[689,904,1992,2013]" gridcol="2" gridrow="34" pageId="11" pageNumber="60">69 (8 ± 1.2)</td>
<td id="76C62D653D5B81C3FC62F82CB0C227C4" box="[940,1156,1992,2013]" gridcol="3" gridrow="34" pageId="11" pageNumber="60">810 (9 ± 1.0)</td>
<td id="76C62D653D5B81C3FB64F82CB1C427C4" box="[1194,1410,1992,2013]" gridcol="4" gridrow="34" pageId="11" pageNumber="60">611 (8 ± 1.9)</td>
</tr>
</table>
</paragraph>
<caption id="DF5816D33D5C7E30FF59F8A3B0B82766" httpUri="https://zenodo.org/record/193864/files/figure.png" pageId="12" pageNumber="61" targetBox="[154,1436,212,1822]" targetPageId="12">
<paragraph id="8B98465B3D5C7E30FF59F8A3B0B82766" blockId="12.[151,1436,1863,1920]" pageId="12" pageNumber="61">
<emphasis id="B9539A493D5C7E30FF59F8A3B55F2746" bold="true" box="[151,281,1863,1887]" pageId="12" pageNumber="61">FIGURE 5.</emphasis>
Photographs of
<taxonomicName id="4C273DD83D5C7E30FE1EF8A3B6A32747" box="[464,741,1863,1886]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="61" phylum="Chordata" rank="species" species="jambangan" status="sp. nov.">
<emphasis id="B9539A493D5C7E30FE1EF8A3B6A32747" box="[464,741,1863,1886]" italics="true" pageId="12" pageNumber="61">Cyrtodactylus jambangan</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26027323D5C7E30FD3CF8A3B7682747" box="[754,814,1863,1886]" pageId="12" pageNumber="61" rank="species">n. sp.</taxonomicNameLabel>
, in life: typical muted, marbled color pattern of (
<emphasis id="B9539A493D5C7E30FAF7F8A3B10A2746" bold="true" box="[1337,1356,1863,1887]" pageId="12" pageNumber="61">A</emphasis>
) adults (holotype PNM 9593) and (
<emphasis id="B9539A493D5C7E30FE71F88CB5972799" bold="true" box="[447,465,1896,1920]" pageId="12" pageNumber="61">B</emphasis>
) more strikingly patterned contrasting coloration of juveniles (KU 314812).
</paragraph>
</caption>
<paragraph id="8B98465B3D5C7E31FF09F84CB0B1218B" blockId="12.[151,1436,1960,2023]" lastBlockId="13.[151,1437,152,2007]" lastPageId="13" lastPageNumber="62" pageId="12" pageNumber="61">
<collectingCountry id="F33006CB3D5C7E30FF09F84CB54427DB" box="[199,258,1960,1986]" name="India" pageId="12" pageNumber="61">Hind</collectingCountry>
limbs relatively robust, limb diameter at insertion twice that of forelimb; femur long, 16.8% snout vent length; dorsal hindlimb scalation heterogenous, scales increasing in size distally, with tubercles regularly distributed, increasing in size and becoming more spinose distally; ventral scalation heterogenous, increasing in size distally, lacking tubercles; right limb with two conspicuous patches dorsally, lacking scales or tubercles (possibly resulting from injury); enlarged femoral scales (and pores) absent; toes well developed, relatively longer than those on manus; Toes I and IV 31.2% and 85.4% tibia length, respectively; lamellae enlarged and slightly raised, lamellae proximal to inflection larger than those distal to inflection, slight increase in size from inflection to claw; toe number followed by subdigital lamellae (in parentheses): I (12), II (16), III (20), IV (23), V (22); all toes clawed; claws well developed, sheathed by single dorsal and ventral scale.
</paragraph>
<paragraph id="8B98465B3D5D7E31FF09FE79B78D226B" blockId="13.[151,1437,152,2007]" pageId="13" pageNumber="62">Tail original, length 78.2 mm, width and height 5.6 and 4.8 mm at base respectively, tapering abruptly posterior to hemipenal bulge, then gradually to terminus; 7 annuli discernable dorsally through lateral margins, posterior to which whorls cannot be distinguished, owing to absence of differentiated scales associated with annuli; tubercles smaller and less spinose than those of trunk; enlarged medial subcaudal scales absent; 1 enlarged post-cloacal tubercle present on each side of cloaca; anterior margin of hemipenal bulge with paired, laterally expanded, postcloacal glandular openings.</paragraph>
<paragraph id="8B98465B3D5D7E31FF09FD99B7CA2335" blockId="13.[151,1437,152,2007]" pageId="13" pageNumber="62">
<emphasis id="B9539A493D5D7E31FF09FD99B6D5228E" bold="true" box="[199,659,637,663]" pageId="13" pageNumber="62">
Measurements of
<typeStatus id="549CF8F93D5D7E31FE62FD99B65C228E" box="[428,538,637,663]" pageId="13" pageNumber="62" type="holotype">holotype</typeStatus>
(in mm).
</emphasis>
Snoutvent length 67.7; lamellae under Finger
<date id="FF99609B3D5D7E31FB2FFD99B16F228E" box="[1249,1321,637,663]" pageId="13" pageNumber="62" value="1920-03">III 20</date>
; lamellae under Toe
<date id="FF99609B3D5D7E31FED7FD46B52422A5" box="[281,354,674,700]" pageId="13" pageNumber="62" value="1923-04">IV 23</date>
; Finger I length 3.6; Finger III length 6.0; Toe I length 3.3; Toe IV length 9.0; eye-narial distance 6.5; eyeear distance 5.6; interorbital distance 2.6; snout length 8.6; internarial distance 2.2; eye diameter 4.2; upper arm length 8.4; upper arm length 9.7; femur length 11.4; tibia length 10.5; hand length 9.1; foot length 10.9; axillagroin distance 10.1; tail length 84.6.
</paragraph>
<paragraph id="8B98465B3D5D7E31FF09FCD3B08024BB" blockId="13.[151,1437,152,2007]" pageId="13" pageNumber="62">
<emphasis id="B9539A493D5D7E31FF09FCD3B6ED2348" bold="true" box="[199,683,823,849]" pageId="13" pageNumber="62">
Coloration of
<typeStatus id="549CF8F93D5D7E31FEB8FCD3B5A72348" box="[374,481,823,849]" pageId="13" pageNumber="62" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
Dorsal ground coloration (of head, neck, body, limbs, and tail) dark brown, with irregularly shaped dark gray to lavender transverse dorsal bands, varying from irregular dark gray blotches anteriorly, to a single medial blotch followed posteriorly by a “V-shaped” marking in the nuchal region, to three conspicuous, “butterfly-shaped” patterns interspersed with three irregularly shaped transverse blotches throughout the trunk, and becoming less-defined laterally; supralabials with four conspicuous light tan to white blotches (left and right) irregularly distributed between the second and twelfth scales; light gray band bordered by dark brown extends from posterior margin of eye to auricular opening; trunk tubercles primarily light tan to cream; limbs mottled, with dark gray and dark brown, lacking distinct bands; digits with light tan to cream blotches at joints; tail more conspicuously banded than trunk, with dark brown ground coloration fading to white at the terminus of tail, overlain with 10 dark gray to black bands.
</paragraph>
<paragraph id="8B98465B3D5D7E31FF09FB49B7D4250B" blockId="13.[151,1437,152,2007]" pageId="13" pageNumber="62">Ventral portions of head, trunk, and limbs tan, darkening at margins of ventrolateral tubercle row; hands and feet tan, fingers and toes slightly darker; subcaudal coloration cream with increased dark speckling through anterior third; seven discernable white bands posteriorly.</paragraph>
<paragraph id="8B98465B3D5D7E31FF09FAF9B5B427AB" blockId="13.[151,1437,152,2007]" pageId="13" pageNumber="62">
<emphasis id="B9539A493D5D7E31FF09FAF9B5D5252E" bold="true" box="[199,403,1309,1335]" pageId="13" pageNumber="62">Color Variation.</emphasis>
Our sample of
<specimenCount id="9D218DD23D5D7E31FD9EFAF9B6FD252E" box="[592,699,1309,1335]" pageId="13" pageNumber="62" type="male">36 males</specimenCount>
,
<specimenCount id="9D218DD23D5D7E31FD09FAF9B70F252E" box="[711,841,1309,1335]" pageId="13" pageNumber="62" type="female">52 females</specimenCount>
, and
<specimenCount id="9D218DD23D5D7E31FC49FAF9B053252E" box="[903,1045,1309,1335]" pageId="13" pageNumber="62" type="juvenile">26 juveniles</specimenCount>
exhibits minimal color variation. Four adult males (KU 314794, 314796, 314813, and 314825) and four adult females (KU 314804, 314823 24, and 314831) are darker, with dorsal bands through the axilla-groin region of higher contrast to the lighter ground color. One adult male (KU 314815) and two adult females (KU 314822, and 314827) are lighter, with distinct dorsal banding, and light brown to light gray ground color overlaid by medium to dark gray band. One adult male (KU 314805) and one adult female (KU 314795) lack a defined “V-shaped” blotch spanning the posterior dorsum of the head and nuchal region. Three adult males (KU 314794, 314805, and 314835) lack the conspicuous blotch medially in the nuchal region. All other specimens have color and patterning consistent with the
<typeStatus id="549CF8F93D5D7E31FE46F9ACB5BE267B" box="[392,504,1608,1634]" pageId="13" pageNumber="62" type="holotype">holotype</typeStatus>
. Additionally, nine adult males (KU 314781, 31478485, 314789, 314791, 314797, 314803, 314812, and 314836),
<specimenCount id="9D218DD23D5D7E31FD48F989B6AC269E" box="[646,746,1645,1671]" pageId="13" pageNumber="62" type="adult">19 adult</specimenCount>
females (KU 31477880, 31478788, 314790, 314792, 314798, 31480809, 31481619, 31482021, 314829, 314832, and 314834), and
<specimenCount id="9D218DD23D5D7E31FBAFF976B0B626B5" box="[1121,1264,1682,1708]" pageId="13" pageNumber="62" type="juvenile">10 juveniles</specimenCount>
(KU 314782 83, 314883, 314799802, 314807, 314786, and 314814) show similar trends in variation, with one adult male (KU 314797), two adult females (KU 314816 and 314832), and two juveniles (KU 314814 and 314833) with a darker overall coloration and more highly defined dorsal banding. Four adult males (KU 314781, 314784, 314789, and 314803) and
<specimenCount id="9D218DD23D5D7E31FE15F8CCB679275B" box="[475,575,1832,1858]" pageId="13" pageNumber="62" type="adult">13 adult</specimenCount>
females (KU 314734, 31477980, 314790, 314792, 314798, 314809, 31481721, and 314829) have a light gray ground color with dark gray, well-defined dorsal banding. Regenerated tails have highly variable dark speckling overlaying a medium gray ground color, to a completely dark brown dorsal coloration.
</paragraph>
<paragraph id="8B98465B3D5D7E32FF09F859B54021AE" blockId="13.[151,1437,152,2007]" lastBlockId="14.[151,1436,152,738]" lastPageId="14" lastPageNumber="63" pageId="13" pageNumber="62">Generally, a greater level of definition in dorsal banding is present in female and juveniles; these specimens have greater contrast in dorsal banding; a few individuals nearly lack dorsal bands all together (KU 314799, 314801, and 314802). Ventral coloration is less variable except on the tail. Original tails are similarly colored with speckling increasing from the anterior margin of the tail to the terminus, with the tail tip being nearly black. Dorsal, light-colored bands extend through venter, becoming lighter in color and more defined towards terminus of tail. One adult male (KU 314825) has speckling at the base and the tip of the tail, a cream ground color throughout, and lacks dark patterning in between. All others exhibit increasing amounts of dark speckling over a light to medium gray ground color from the anterior margin of the tail to the terminus.</paragraph>
<paragraph id="8B98465B3D5E7E32FF09FE26B168223E" blockId="14.[151,1436,152,738]" pageId="14" pageNumber="63">
<emphasis id="B9539A493D5E7E32FF09FE26B5B921C5" bold="true" box="[199,511,450,476]" pageId="14" pageNumber="63">Morphological variation.</emphasis>
Summaries of univariate morphological variation in the series are presented in Table 3. Our series shows minimal morphological variation between males and females. However, females are generally larger and more robust (the largest specimen, KU 314831, is female with SVL = 81.5).
</paragraph>
<paragraph id="8B98465B3D5E7E32FF09FDD6B6DB22FB" blockId="14.[151,1436,152,738]" pageId="14" pageNumber="63">
<emphasis id="B9539A493D5E7E32FF09FDD6B5252255" bold="true" box="[199,355,562,588]" pageId="14" pageNumber="63">Color in life.</emphasis>
<taxonomicName id="4C273DD83D5E7E32FEBFFDD6B6E62252" box="[369,672,562,587]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="63" phylum="Chordata" rank="species" species="jambangan">
<emphasis id="B9539A493D5E7E32FEBFFDD6B6E62252" box="[369,672,562,587]" italics="true" pageId="14" pageNumber="63">Cyrtodactylus jambangan</emphasis>
</taxonomicName>
has light yellow to gold dorsal tubercles through the axillagroin region, a yellow canthal stripe, bright yellow superciliaries, and a variable dorsal banding pattern. All three characters are in contrast to observations of true
<taxonomicName id="4C273DD83D5E7E32FD35FD99B7DC228F" box="[763,922,637,662]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="63" phylum="Chordata" rank="species" species="annulatus">
<emphasis id="B9539A493D5E7E32FD35FD99B7DC228F" box="[763,922,637,662]" italics="true" pageId="14" pageNumber="63">C. annulatus</emphasis>
</taxonomicName>
, which has regular, distinct, and brightly contrasting dorsal bands through the axilla-groin region, more pattern variation, and also lacks bright yellow canthal stripe and superciliaries (see
<figureCitation id="131C5ADE3D5E7E32FD89FD2CB6CB22FB" box="[583,653,712,738]" captionStart="FIGURE 5" captionStartId="12.[151,255,1863,1887]" captionTargetBox="[154,1436,212,1822]" captionTargetId="figure@12.[151,1436,197,1837]" captionTargetPageId="12" captionText="FIGURE 5. Photographs of Cyrtodactylus jambangan, n. sp., in life: typical muted, marbled color pattern of (A) adults (holotype PNM 9593) and (B) more strikingly patterned contrasting coloration of juveniles (KU 314812)." httpUri="https://zenodo.org/record/193864/files/figure.png" pageId="14" pageNumber="63">Fig. 5</figureCitation>
).
</paragraph>
<caption id="DF5816D33D5E7E32FF59FA13B73B2637" httpUri="https://zenodo.org/record/193865/files/figure.png" pageId="14" pageNumber="63" targetBox="[217,1369,791,1504]" targetPageId="14">
<paragraph id="8B98465B3D5E7E32FF59FA13B73B2637" blockId="14.[151,1436,1527,1582]" pageId="14" pageNumber="63">
<emphasis id="B9539A493D5E7E32FF59FA13B55F2616" bold="true" box="[151,281,1527,1551]" pageId="14" pageNumber="63">FIGURE 6.</emphasis>
Preferred riparian habitat characteristics of
<taxonomicName id="4C273DD83D5E7E32FD37FA13B7D72617" box="[761,913,1527,1550]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="63" phylum="Chordata" rank="species" species="jambangan" status="sp. nov.">
<emphasis id="B9539A493D5E7E32FD37FA13B7D72617" box="[761,913,1527,1550]" italics="true" pageId="14" pageNumber="63">C. jambangan</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26027323D5E7E32FC50FA13B79F2617" box="[926,985,1527,1550]" pageId="14" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
, along the Tumaga River, 90 m above sea level, Pasonanca Natural Park, Zamboanga City Province, Mindanao.
</paragraph>
</caption>
<paragraph id="8B98465B3D5E7E32FF09F9B8B78726D8" blockId="14.[151,1437,1628,2028]" pageId="14" pageNumber="63">
<emphasis id="B9539A493D5E7E32FF09F9B8B674266F" bold="true" box="[199,562,1628,1654]" pageId="14" pageNumber="63">Ecology and natural history.</emphasis>
The new species is found at low and mid-elevations in riparian habitats (gallery forests) along streams. We encountered
<taxonomicName id="4C273DD83D5E7E32FD38F966B7E52682" box="[758,931,1666,1691]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="63" phylum="Chordata" rank="species" species="jambangan">
<emphasis id="B9539A493D5E7E32FD38F966B7E52682" box="[758,931,1666,1691]" italics="true" pageId="14" pageNumber="63">C. jambangan</emphasis>
</taxonomicName>
specimens on trunks of trees, rocks, and overhanging exposed root masses on the banks of rivers and streams.
</paragraph>
</subSubSection>
<subSubSection id="C33D15D03D5E7E32FF09F928B6BF2765" pageId="14" pageNumber="63" type="distribution">
<paragraph id="8B98465B3D5E7E32FF09F928B6BF2765" blockId="14.[151,1437,1628,2028]" pageId="14" pageNumber="63">
<emphasis id="B9539A493D5E7E32FF09F928B53526FE" bold="true" box="[199,371,1740,1767]" pageId="14" pageNumber="63">Distribution.</emphasis>
The distribution of the new species includes Pasonanca Natural Park, Zamboanga Peninsula, extreme western Mindanao Island, and adjacent Sulu Archipelago islands to the southwest (
<figureCitation id="131C5ADE3D5E7E32FA88F916B1CC2715" box="[1350,1418,1778,1804]" captionStart="FIGURE 1" captionStartId="4.[151,255,718,742]" captionTargetBox="[226,1337,192,701]" captionTargetId="figure@4.[217,1347,183,710]" captionTargetPageId="4" captionText="FIGURE 1. The potential distribution of Cyrtodactylus annulatus in the southern and central Philippines (left, darkly shaded islands), known C. annulatus localities on Mindanao Isl. (right, black circles; square = Bunawan, type locality), the distribution of C. tautbatorum from Palawan (Welton et al. 2009; left, diamond, southern Palawan) and know localities for C. jambangan, n. sp. (the type locality indicated with a star; additional localities substantiated with specimens encircled with dashed line). The question mark adjacent to Tawi-tawi Island indicates uncertainty with respect to the taxonomic status of the Cyrtodactylus of the southwestern Sulu Archipelago. The occurrence of Cyrtodactylus from these islands was reported by Taylor (1922; as C. annulatus), but specimens from southern Sulu Archipelago islands currently are not available in collections. The single dark spot north of Mt. Malingdang at the base of the Zamboanga peninsula is the westernmost locality for true C. annulatus." httpUri="https://zenodo.org/record/193860/files/figure.png" pageId="14" pageNumber="63">Fig. 1</figureCitation>
). Other localities in the Sulu Archipelago where
<bibRefCitation id="EFB63BAA3D5E7E32FD01F8F3B7C42728" author="Taylor" box="[719,898,1815,1841]" pageId="14" pageNumber="67" refString="Taylor, E. H. (1922 a) The lizards of the Philippine Islands. Philippine Bureau of Science, Manila, Philippines, 269 pp." type="book" year="1922" yearSuffix="a">Taylor (1922a)</bibRefCitation>
observed this species include Basilan, Great Santa Cruz, Teipono, Tamuk, Cancuman, Dipolod, Bitinan, Jolo, Tulian, Tawitawi, Papahag, Bongao, Buban Islands of the Tapiantana and Taipan Island Groups.
</paragraph>
</subSubSection>
<subSubSection id="C33D15D03D5E7E33FF09F863B7A220E5" lastPageId="15" lastPageNumber="64" pageId="14" pageNumber="63" type="etymology">
<paragraph id="8B98465B3D5E7E33FF09F863B7A220E5" blockId="14.[151,1437,1628,2028]" lastBlockId="15.[151,1437,152,252]" lastPageId="15" lastPageNumber="64" pageId="14" pageNumber="63">
<emphasis id="B9539A493D5E7E32FF09F863B51B27B8" bold="true" box="[199,349,1927,1953]" pageId="14" pageNumber="63">Etymology.</emphasis>
The specific epithet is derived from the term
<taxonomicName id="4C273DD83D5E7E32FC78F86CB00E27B8" box="[950,1096,1928,1953]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="68" phylum="Chordata" rank="species" species="jambangan">
<emphasis id="B9539A493D5E7E32FC78F86CB00E27B8" box="[950,1096,1928,1953]" italics="true" pageId="14" pageNumber="63">Jambangan</emphasis>
</taxonomicName>
—the ancient name for the Zamboanga City area. Sulu and Zamboanga folklore suggest that the name was bestowed upon the Zamboanga City area by the immigrant Subanons (“People of the River”) who arrived in western Mindanao and the Sulu archipelago in approximately 1200 A.D., after traveling by boat through the Sulu Archipelago, from what is now
<collectingCountry id="F33006CB3D5F7E33FE4EFF59B64620CE" box="[384,512,189,215]" name="Indonesia" pageId="15" pageNumber="64">Indonesia</collectingCountry>
.
<taxonomicName id="4C273DD83D5F7E33FDDBFF59B6E120CF" box="[533,679,189,214]" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="68" phylum="Chordata" rank="species" species="jambangan">
<emphasis id="B9539A493D5F7E33FDDBFF59B6E120CF" box="[533,679,189,214]" italics="true" pageId="15" pageNumber="64">Jambangan</emphasis>
</taxonomicName>
means “The Land of Flowers,” a name presumably used in reference to the natural beauty of the area surrounding Zamboanga City.
</paragraph>
</subSubSection>
</treatment>
</document>