143 lines
17 KiB
XML
143 lines
17 KiB
XML
<document id="181C2EB6FD9EF061FABA64F50AFAB036" ID-CLB-Dataset="45678" ID-DOI="10.5281/zenodo.279694" ID-GBIF-Dataset="51ab7190-d4ae-4c04-9e20-05e2c15196a1" ID-ISSN="1175-5326" ID-Zenodo-Dep="279694" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460426162541" checkinUser="plazi" docAuthor="Arce-Pérez, Roberto, Shepard, William D. & Morón, Miguel A." docDate="2012" docId="038887BAFF8EFFD3FF692579FC703087" docLanguage="en" docName="zt03157p040.pdf" docOrigin="Zootaxa 3157" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Belicinus Arce-Perez and Shepard, gen. nov." docType="treatment" docVersion="9" lastPageNumber="32" masterDocId="FFB1FFC2FF8FFFD2FFFE261FFF8A3732" masterDocTitle="Belicinus Rhomboideus, a new genus and species of Psepheninae (Coleoptera: Byrrhoidea: Psephenidae) from Belize" masterLastPageNumber="40" masterPageNumber="31" pageNumber="32" updateTime="1698330708399" updateUser="ExternalLinkService">
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<mods:mods id="013CF21E1136603BA965271E56EB689D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FE5C98F95D00C6DA933B0CA16396A8C8">
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<mods:title id="E326D6087480FBABFE89798B76EEC65D">Belicinus Rhomboideus, a new genus and species of Psepheninae (Coleoptera: Byrrhoidea: Psephenidae) from Belize</mods:title>
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</mods:titleInfo>
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<mods:name id="CE4EFE1158E199EE3B5CA414C6C7B22B" type="personal">
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<mods:role id="2C8741CFF3C1C1CEDC46BBE6CC767ADD">
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<mods:roleTerm id="631E5BBA76A3ECE93143CDE00FB0F72F">Author</mods:roleTerm>
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<mods:namePart id="5A8D8BFF3FF195FE28BB51C3EE176F61">Arce-Pérez, Roberto</mods:namePart>
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</mods:name>
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<mods:name id="1298F53459B205947FD5CB834C5893F8" type="personal">
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<mods:role id="42DC6CA0416B2FB858A6AE292CD70E03">
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<mods:roleTerm id="CBE099226BC83150E1DEBA79DF337B6D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6607CC11E205A57EB90167CDDB35B81C">Shepard, William D.</mods:namePart>
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</mods:name>
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<mods:name id="8CFA57A46A2CBE13155E81D960E7EAD9" type="personal">
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<mods:roleTerm id="8C2841C4FCE66841CBDF9973B73F3CA9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AF3C1E2F9D7C139614D6ED691A460651">Morón, Miguel A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B2180B17BCDE3E73A4E6136C165A2B51">text</mods:typeOfResource>
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<mods:title id="2925D67E52E72D35E94C8EF3774394E9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CC12384572391C1CAF900E3E9D50136F">
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<mods:date id="DC4826FD7C17A7F440375322E19E640F">2012</mods:date>
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<mods:detail id="C8EEF2B83A8C241F650DA5097253F1EB" type="volume">
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<mods:number id="549EA6EEA7D00BB0DF4D4C12DDEE48F1">3157</mods:number>
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<mods:classification id="DA0961C70AE016A7F4D2916D36992F99">journal article</mods:classification>
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<mods:identifier id="1AA7EDAF778854B641A60EC883123F19" type="CLB-Dataset">45678</mods:identifier>
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<mods:identifier id="EEEF11EB64BDD5836244B8BF69041F77" type="DOI">10.5281/zenodo.279694</mods:identifier>
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<mods:identifier id="15B7FB780BAD246C9EA9C79F700EFA96" type="GBIF-Dataset">51ab7190-d4ae-4c04-9e20-05e2c15196a1</mods:identifier>
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<mods:identifier id="F7E36D40150E1EE0FB1D135B6FF2253E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C49B2BCED47CF7A32A994275750E28CB" type="Zenodo-Dep">279694</mods:identifier>
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<treatment id="038887BAFF8EFFD3FF692579FC703087" ID-DOI="http://doi.org/10.5281/zenodo.6169048" ID-GBIF-Taxon="119428669" ID-Zenodo-Dep="6169048" LSID="urn:lsid:plazi:treatment:038887BAFF8EFFD3FF692579FC703087" httpUri="http://treatment.plazi.org/id/038887BAFF8EFFD3FF692579FC703087" lastPageNumber="32" pageId="1" pageNumber="32">
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<subSubSection id="C33B6527FF8EFFD3FF692579FE083493" pageId="1" pageNumber="32" type="nomenclature">
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<paragraph id="8B9E36ACFF8EFFD3FF692579FD3D34B2" blockId="1.[151,695,870,929]" box="[151,695,870,896]" pageId="1" pageNumber="32">
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<heading id="D0D681C0FF8EFFD3FF692579FD3D34B2" bold="true" box="[151,695,870,896]" fontSize="11" level="1" pageId="1" pageNumber="32" reason="1">
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<emphasis id="B955EABEFF8EFFD3FF692579FD3D34B2" bold="true" box="[151,695,870,896]" pageId="1" pageNumber="32">
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<taxonomicName id="4C214D2FFF8EFFD3FF692579FDC834B2" authority="Arce-Perez and Shepard" authorityName="Arce-Perez and Shepard" box="[151,578,870,896]" class="Insecta" family="Psephenidae" genus="Belicinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B955EABEFF8EFFD3FF692579FE8334B2" bold="true" box="[151,265,870,896]" italics="true" pageId="1" pageNumber="32">Belicinus</emphasis>
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Arce-Pérez and Shepard
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</taxonomicName>
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<taxonomicNameLabel id="A26657C5FF8EFFD3FDB42579FD3D34B2" box="[586,695,870,896]" pageId="1" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9E36ACFF8EFFD3FF692596FE083493" blockId="1.[151,695,870,929]" box="[151,386,905,929]" pageId="1" pageNumber="32">
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(
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<figureCitation id="131A2A29FF8EFFD3FF612596FF673493" box="[159,237,905,929]" captionStart="FIGURES 1 6" captionStartId="3.[151,264,1661,1684]" captionTargetBox="[165,1424,181,1637]" captionTargetId="figure@3.[165,1425,181,1637]" captionTargetPageId="3" captionText="FIGURES 1 6. Belicinus rhomboideus sp. nov. 1. Head and pronotum, dosal view. 2. Head and prosternum, ventral view. 3. Labrum and maxillary palpus, ventral view. 4. Midline of pronotum with a fine carina in the distal half. 5 6. Protibiae slender, clearly arched toward the exterior, with tiny teeth on their internal margin." httpUri="https://zenodo.org/record/279695/files/figure.png" pageId="1" pageNumber="32">Figs. 1</figureCitation>
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, 2, 4, 5, 6, 9)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6527FF8EFFD3FF6925CEFF543247" pageId="1" pageNumber="32" type="description">
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<paragraph id="8B9E36ACFF8EFFD3FF6925CEFF543247" blockId="1.[151,1437,977,1973]" pageId="1" pageNumber="32">
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<emphasis id="B955EABEFF8EFFD3FF6925CEFEA334D8" bold="true" box="[151,297,977,1002]" pageId="1" pageNumber="32">Description.</emphasis>
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Head (
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<figureCitation id="131A2A29FF8EFFD3FE8225CEFE3434DB" box="[380,446,977,1001]" captionStart="FIGURES 1 6" captionStartId="3.[151,264,1661,1684]" captionTargetBox="[165,1424,181,1637]" captionTargetId="figure@3.[165,1425,181,1637]" captionTargetPageId="3" captionText="FIGURES 1 6. Belicinus rhomboideus sp. nov. 1. Head and pronotum, dosal view. 2. Head and prosternum, ventral view. 3. Labrum and maxillary palpus, ventral view. 4. Midline of pronotum with a fine carina in the distal half. 5 6. Protibiae slender, clearly arched toward the exterior, with tiny teeth on their internal margin." httpUri="https://zenodo.org/record/279695/files/figure.png" pageId="1" pageNumber="32">Fig. 1</figureCitation>
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) long, eyes spherical, very prominent; antennae short (e.g. hardly reaching the apex of scutellum), filiform and subserrated, 11 antennomeres, scape more robust and larger than remainder of antennomeres (globose); clypeus subtrapezoidal, declivous at an angle of less than 45 degrees from the plane of the frons, distal margin slightly lifted and bent backwards. Pronotum (
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<figureCitation id="131A2A29FF8EFFD3FCB22222FC113367" box="[844,923,1085,1109]" captionStart="FIGURES 1 6" captionStartId="3.[151,264,1661,1684]" captionTargetBox="[165,1424,181,1637]" captionTargetId="figure@3.[165,1425,181,1637]" captionTargetPageId="3" captionText="FIGURES 1 6. Belicinus rhomboideus sp. nov. 1. Head and pronotum, dosal view. 2. Head and prosternum, ventral view. 3. Labrum and maxillary palpus, ventral view. 4. Midline of pronotum with a fine carina in the distal half. 5 6. Protibiae slender, clearly arched toward the exterior, with tiny teeth on their internal margin." httpUri="https://zenodo.org/record/279695/files/figure.png" pageId="1" pageNumber="32">Figs. 1</figureCitation>
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, 4) trapezoidal, with a longitudinal glabrous line and a fine carina in the distal half, with posterior margin quite smooth, never crenulate or rugose; lateral margins rounded, unexpanded at posterior half. Legs with fore coxae and middle coxae not far apart (Fig. 2); fore femora robust subtriangular; tibiae (Figs. 5, 6) slender, clearly arched, with tiny teeth on their internal margin; basal four tarsomeres triangular (
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<figureCitation id="131A2A29FF8EFFD3FE3922D2FD8133D7" box="[455,523,1229,1253]" captionStart="FIGURES 7 12" captionStartId="4.[151,264,1932,1955]" captionTargetBox="[167,1410,193,1890]" captionTargetId="figure@4.[167,1411,193,1891]" captionTargetPageId="4" captionText="FIGURES 7 12. Belicinus rhomboideus sp. nov. 7. First four tarsomeres (dorsal and lateral view). 8. Abdominal ventrites. 9. Aedeagus on ventral view, showing phallobase, parameres and middle lobe (penis). 10. Aedeagus, dorsal view. 11. Aedeagus, close-up of penis, ventral view. 12. Aedeagus, dorsal view." httpUri="https://zenodo.org/record/279696/files/figure.png" pageId="1" pageNumber="32">Fig. 7</figureCitation>
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), flattened and widely emarginate at apex, fifth tarsomere (Fig. 5) longest, as long as preceding ones together, cylindrical; tarsal claws entire, never bifid or split, widely separate, widened at base and without teeth or accessory membranes. Male genitalia (Fig. 9) in ventral view subrectangular, trilobate; phallobase shorter than parameres; parameres shorter than median lobe; median lobe long and of expanded rhombic shape.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6527FF8EFFD3FF39239DFE59307B" pageId="1" pageNumber="32" type="discussion">
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<paragraph id="8B9E36ACFF8EFFD3FF39239DFE59307B" blockId="1.[151,1437,977,1973]" pageId="1" pageNumber="32">
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<emphasis id="B955EABEFF8EFFD3FF39239DFEB132A8" bold="true" box="[199,315,1410,1434]" pageId="1" pageNumber="32">Remarks.</emphasis>
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Other
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<taxonomicName id="4C214D2FFF8EFFD3FE75239EFD9D32AB" box="[395,535,1409,1433]" class="Insecta" family="Psephenidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="32" phylum="Arthropoda" rank="subFamily" subFamily="Psepheninae">Psepheninae</taxonomicName>
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genera can be distinguished from
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<taxonomicName id="4C214D2FFF8EFFD3FC5C239EFB8032AB" box="[930,1034,1409,1433]" class="Insecta" family="Psephenidae" genus="Belicinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B955EABEFF8EFFD3FC5C239EFB8032AB" box="[930,1034,1409,1433]" italics="true" pageId="1" pageNumber="32">Belicinus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26657C5FF8EFFD3FBEC239DFBF732A8" box="[1042,1149,1410,1434]" pageId="1" pageNumber="32" rank="genus">
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<emphasis id="B955EABEFF8EFFD3FBEC239DFBF732A8" bold="true" box="[1042,1149,1410,1434]" pageId="1" pageNumber="32">gen. nov.</emphasis>
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</taxonomicNameLabel>
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by the following characters:
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<taxonomicName id="4C214D2FFF8EFFD3FF3023BAFECC328F" box="[206,326,1445,1469]" class="Insecta" family="Psephenidae" genus="Psephenus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="B955EABEFF8EFFD3FF3023BAFECC328F" box="[206,326,1445,1469]" italics="true" pageId="1" pageNumber="32">Psephenus</emphasis>
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</taxonomicName>
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has the posterior half of the pronotum occasionally expanded; tibiae straight; four basal tarsomeres slender and subcylindrical, slightly emarginate at apex, with delicate, minute setae or papillae ventrally (Fig. 22); and the median lobe of the male genitalia is mostly membranous and subcylindric.
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<taxonomicName id="4C214D2FFF8EFFD3FBB723F2FB453137" box="[1097,1231,1517,1541]" class="Insecta" family="Psephenidae" genus="Psephenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="B955EABEFF8EFFD3FBB723F2FB453137" box="[1097,1231,1517,1541]" italics="true" pageId="1" pageNumber="32">Psephenops</emphasis>
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</taxonomicName>
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has the pronotum expanded laterally on posterior half; tibiae straight; tarsomeres 1 and 2, and occasionally 3, with wide ventral expansions that cover apical tarsomeres (Fig. 19); and the median lobe of the male genitalia is mostly membranous, subtriangular or finger-like. In
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<taxonomicName id="4C214D2FFF8EFFD3FE012046FDD43143" box="[511,606,1625,1649]" class="Insecta" family="Psephenidae" genus="Pheneps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="B955EABEFF8EFFD3FE012046FDD43143" box="[511,606,1625,1649]" italics="true" pageId="1" pageNumber="32">Pheneps</emphasis>
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</taxonomicName>
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, the body is more slender; the antennae are long approximately 0.75 times the length of the elytra; tibiae straight; tarsomeres 1 2 clearly stouter than any of the three apical tarsomeres (Fig. 15); and the phallobase is notably larger than the parameres.
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<taxonomicName id="4C214D2FFF8EFFD3FC7520BEFBBC318B" box="[907,1078,1697,1721]" class="Insecta" family="Psephenidae" genus="Psephenotarsis" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="B955EABEFF8EFFD3FC7520BEFBBC318B" box="[907,1078,1697,1721]" italics="true" pageId="1" pageNumber="32">Psephenotarsis</emphasis>
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</taxonomicName>
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has the antennae short, moniliform or filiform; pronotum expanded laterally in posterior half; tibiae straight; and the median lobe of the male genitalia is long, subcylindric and sclerotized, or finger-like and membranous.
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<taxonomicName id="4C214D2FFF8EFFD3FBE620F6FB453033" box="[1048,1231,1769,1793]" class="Insecta" family="Psephenidae" genus="Psephenopalpus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="32" phylum="Arthropoda" rank="genus">
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<emphasis id="B955EABEFF8EFFD3FBE620F6FB453033" box="[1048,1231,1769,1793]" italics="true" pageId="1" pageNumber="32">Psephenopalpus</emphasis>
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</taxonomicName>
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has the maxillary palpus extremely long, extending beyond sixth antennomere (Fig. 18); tibiae straight; and the median lobe of the male genitalia is finger-like.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6527FF8EFFD3FF39214AFDE0305F" box="[199,618,1877,1902]" pageId="1" pageNumber="32" type="materials_examined">
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<paragraph id="8B9E36ACFF8EFFD3FF39214AFDE0305F" blockId="1.[151,1437,977,1973]" box="[199,618,1877,1902]" pageId="1" pageNumber="32">
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<emphasis id="B955EABEFF8EFFD3FF39214AFEE8305C" bold="true" box="[199,354,1877,1902]" pageId="1" pageNumber="32">
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<typeStatus id="549A880EFF8EFFD3FF39214AFE88305F" box="[199,258,1877,1901]" pageId="1" pageNumber="32">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C214D2FFF8EFFD3FE97214AFDE0305F" box="[361,618,1877,1901]" class="Insecta" family="Psephenidae" genus="Belicinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="32" phylum="Arthropoda" rank="species" species="rhomboideus">
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<emphasis id="B955EABEFF8EFFD3FE97214AFDE0305F" box="[361,618,1877,1901]" italics="true" pageId="1" pageNumber="32">Belicinus rhomboideus</emphasis>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6527FF8EFFD3FF392166FE2530A3" box="[199,431,1913,1938]" pageId="1" pageNumber="32" type="distribution">
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<paragraph id="8B9E36ACFF8EFFD3FF392166FE2530A3" blockId="1.[151,1437,977,1973]" box="[199,431,1913,1938]" pageId="1" pageNumber="32">
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<emphasis id="B955EABEFF8EFFD3FF392166FEEA30A0" bold="true" box="[199,352,1913,1938]" pageId="1" pageNumber="32">Distribution.</emphasis>
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<collectingCountry id="F336763CFF8EFFD3FE992166FE2530A3" box="[359,431,1913,1937]" name="Belize" pageId="1" pageNumber="32">Belize</collectingCountry>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6527FF8EFFD3FF392182FC703087" box="[199,1018,1949,1973]" pageId="1" pageNumber="32" type="etymology">
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<paragraph id="8B9E36ACFF8EFFD3FF392182FC703087" blockId="1.[151,1437,977,1973]" box="[199,1018,1949,1973]" pageId="1" pageNumber="32">
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<emphasis id="B955EABEFF8EFFD3FF392182FEC43087" bold="true" box="[199,334,1949,1973]" pageId="1" pageNumber="32">Etymology.</emphasis>
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Named for the first record of the family in
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<collectingCountry id="F336763CFF8EFFD3FCCA2182FCF53087" box="[820,895,1949,1973]" name="Belize" pageId="1" pageNumber="32">Belize</collectingCountry>
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;
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<taxonomicName id="4C214D2FFF8EFFD3FC742182FC783087" box="[906,1010,1949,1973]" class="Insecta" family="Psephenidae" genus="Belicinus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B955EABEFF8EFFD3FC742182FC783087" box="[906,1010,1949,1973]" italics="true" pageId="1" pageNumber="32">Belicinus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |