treatments-xml/data/03/87/BB/0387BB49FFA03D01FD3ACB22FE19FA1B.xml
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<document id="8B9C2DF9E13631630B5A86945E24DB8A" ID-DOI="10.5252/g2011n3a3" ID-ISSN="1638-9395" ID-Zenodo-Dep="5381380" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630292643081" checkinUser="marcus" docAuthor="Geraads, Denis, Spassov, Nikolaï, Hristova, Latinka, Markov, Georgi N. &amp; Tzankov, Tzanko" docDate="2011" docId="0387BB49FFA03D01FD3ACB22FE19FA1B" docLanguage="en" docName="Geodiversitas.2011.33.3.451-484.pdf" docOrigin="Geodiversitas 33 (3)" docSource="http://dx.doi.org/10.5252/g2011n3a3" docStyle="DocumentStyle:DF8BFA78045876A77BB343B212438C9A.3:Geodiversitas.2011-2012.journal_article.type1" docStyleId="DF8BFA78045876A77BB343B212438C9A" docStyleName="Geodiversitas.2011-2012.journal_article.type1" docStyleVersion="3" docTitle="Prostrepsiceros rotundicornis" docType="treatment" docVersion="3" lastPageNumber="476" masterDocId="FFBEC331FFB83D18FF8ACF6DFF9CFFBC" masterDocTitle="Upper Miocene mammals from Strumyani, South-Western Bulgaria" masterLastPageNumber="484" masterPageNumber="451" pageNumber="475" updateTime="1699217341029" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="958AF8BA1D5BEC2A497DE7C3D9774F40">Upper Miocene mammals from Strumyani, South-Western Bulgaria</mods:title>
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<mods:namePart id="0BD0A9FEA00E928C4E4A6E5E96CC8D5E">Geraads, Denis</mods:namePart>
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<mods:namePart id="D0E55BDDBD4B49FD752D8E9270FFADC9">Spassov, Nikolaï</mods:namePart>
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<mods:namePart id="8ED33BADA9E3C284AB89407F354B0D12">Hristova, Latinka</mods:namePart>
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<mods:namePart id="0DFB6DEF1A34F4AC07750AECEB927669">Markov, Georgi N.</mods:namePart>
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<paragraph id="8B910A5FFFA03D00FD3ACB22FB32FBD6" blockId="24.[688,1198,1102,1130]" box="[688,1198,1102,1130]" pageId="24" pageNumber="475">
<heading id="D0D9BD33FFA03D00FD3ACB22FB32FBD6" box="[688,1198,1102,1130]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="475" reason="2">
<taxonomicName id="4C2E71DCFFA03D00FD3ACB22FB32FBD6" authority="(Weithofer, 1888)" baseAuthorityName="Weithofer" baseAuthorityYear="1888" box="[688,1198,1102,1130]" class="Mammalia" family="Bovidae" genus="Prostrepsiceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="rotundicornis">
<emphasis id="B95AD64DFFA03D00FD3ACB22FC78FBD5" bold="true" box="[688,996,1102,1129]" italics="true" pageId="24" pageNumber="475">Prostrepsiceros rotundicornis</emphasis>
(
<bibRefCitation id="EFBF77AEFFA03D00FC78CB22FB3AFBD6" author="WEITHOFER K. A." box="[1010,1190,1103,1130]" pageId="24" pageNumber="475" pagination="225 - 292" refId="ref22756" refString="WEITHOFER K. A. 1888. - Beitrage zur Kenntnis der Fauna von Pikermi bei Athen. Beitrage zur Palaontologie Osterreich-Ungarns und des Orients 6: 225 - 292." type="journal article" year="1888">Weithofer, 1888</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B910A5FFFA03D00FD2FCBE3FCBCF999" blockId="24.[675,1212,1166,1700]" pageId="24" pageNumber="475">
The best preserved specimen is a frontlet with most of the right horn-core and the base of the left one, FM-2739 (
<figureCitation id="131516DAFFA03D00FC96CBA3FCEFFB55" box="[796,883,1230,1257]" captionStart="FIG" captionStartId="22.[113,124,1612,1629]" captionTargetBox="[146,1176,221,1589]" captionTargetId="figure-17@22.[113,1209,209,1608]" captionTargetPageId="22" captionText="FIG. 13. — Fossil mammals from the Miocene of Strumyani (SW Bulgaria): A-C, F, G, Palaeoreas lindermayeri Wagner, 1848; A, skull FM-2738, front view; B, mandible FM-2744; C, mandible FM-2783; F, frontlet FM-2740; G, frontlet FM-2780; D, I, J, Prostrepsiceros rotundicornis (Weithofer, 1888); D, mandible FM-2331; I, frontlet FM-2779; J, frontlet FM-2739; E, Palaeoryx sp., tooth row p2-m2 FM-2251; H, Tragoportax sp., skull FM-2094. Scale bar: A, G-J, 20 cm; B-E, 5 cm." figureDoi="http://doi.org/10.5281/zenodo.5381398" httpUri="https://zenodo.org/record/5381398/files/figure.png" pageId="24" pageNumber="475">Fig. 13J</figureCitation>
; measurements:
<tableCitation id="C6AC3FE4FFA03D00FBA9CBA3FBE9FB55" box="[1059,1141,1230,1257]" captionStart="TABLE" captionStartId="20.[183,194,220,237]" captionText="TABLE 4. — Measurements (in mm) of bovid frontlets. 1, PIUW; 2, NHM; 3, Kostopoulos 2005; 4, Kostopoulos 2006." pageId="24" pageNumber="475">Table 4</tableCitation>
). FM- 2765 and 2766 are horn-cores in bad condition. This taxon is also represented by a frontlet from Str-1, FM-2779 (
<figureCitation id="131516DAFFA03D00FCD5CA40FC2AFAF4" box="[863,950,1325,1352]" captionStart="FIG" captionStartId="22.[113,124,1612,1629]" captionTargetBox="[146,1176,221,1589]" captionTargetId="figure-17@22.[113,1209,209,1608]" captionTargetPageId="22" captionText="FIG. 13. — Fossil mammals from the Miocene of Strumyani (SW Bulgaria): A-C, F, G, Palaeoreas lindermayeri Wagner, 1848; A, skull FM-2738, front view; B, mandible FM-2744; C, mandible FM-2783; F, frontlet FM-2740; G, frontlet FM-2780; D, I, J, Prostrepsiceros rotundicornis (Weithofer, 1888); D, mandible FM-2331; I, frontlet FM-2779; J, frontlet FM-2739; E, Palaeoryx sp., tooth row p2-m2 FM-2251; H, Tragoportax sp., skull FM-2094. Scale bar: A, G-J, 20 cm; B-E, 5 cm." figureDoi="http://doi.org/10.5281/zenodo.5381398" httpUri="https://zenodo.org/record/5381398/files/figure.png" pageId="24" pageNumber="475">Fig. 13I</figureCitation>
). The neurocranium was certainly little inclined on the face, as the frontal is little bent. The supra-orbital foramina open at the lateral sides of large depressions that nearly meet at a mid-frontal suture; the latter is slightly raised and open, as is also the fronto-parietal one. There is a large post-cornual fossa. The orbital rims are prominent.
</paragraph>
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<subSubSection id="C33459D4FFA03D01FD31C947FE19FA1B" lastPageId="25" lastPageNumber="476" pageId="24" pageNumber="475" type="description">
<paragraph id="8B910A5FFFA03D01FD31C947FED8FE8E" blockId="24.[675,1212,1166,1700]" lastBlockId="25.[111,648,216,1447]" lastPageId="25" lastPageNumber="476" pageId="24" pageNumber="475">The pedicles are extremely short, as the horncores rise directly from the frontal bone. They are inserted rather uprightly, diverge from the very base, and the divergence increases upwards before decreasing in the second half. They are strongly spiralled but lack keels, although grooves run along their lateral surface.</paragraph>
<paragraph id="8B910A5FFFA13D01FF02CE5AFEFDFAB4" blockId="25.[111,648,216,1447]" pageId="25" pageNumber="476">
These frontlets obviously belong to a group of
<taxonomicName id="4C2E71DCFFA13D01FFF9CE3AFE14FECE" authority="Major, 1891" authorityName="Major" authorityYear="1891" box="[115,392,343,370]" class="Mammalia" family="Bovidae" genus="Prostrepsiceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="25" pageNumber="476" phylum="Chordata" rank="genus">
<emphasis id="B95AD64DFFA13D01FFF9CE3AFE9DFECC" box="[115,257,343,368]" italics="true" pageId="25" pageNumber="476">Prostrepsiceros</emphasis>
Major, 1891
</taxonomicName>
whose horn-cores virtually lack keels, and which includes
<taxonomicName id="4C2E71DCFFA13D01FE6FCE1AFD1AFE2C" baseAuthorityName="Weithofer" baseAuthorityYear="1888" box="[485,646,374,400]" class="Mammalia" family="Bovidae" genus="Prostrepsiceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="rotundicornis">
<emphasis id="B95AD64DFFA13D01FE6FCE1AFD1AFE2C" box="[485,646,374,400]" italics="true" pageId="25" pageNumber="476">P. rotundicornis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2E71DCFFA13D01FF14CEFAFE0EFE0D" authority="(Andree, 1826)" baseAuthorityName="Andree" baseAuthorityYear="1826" box="[158,402,406,433]" class="Mammalia" family="Bovidae" genus="Prostrepsiceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="fraasi">
<emphasis id="B95AD64DFFA13D01FF14CEFAFF77FE0C" box="[158,235,406,432]" italics="true" pageId="25" pageNumber="476">P. fraasi</emphasis>
(Andree, 1826)
</taxonomicName>
. The former species was defined at Pikermi (
<bibRefCitation id="EFBF77AEFFA13D01FEB7CEDBFE74FE6D" author="WEITHOFER K. A." box="[317,488,438,465]" pageId="25" pageNumber="476" pagination="225 - 292" refId="ref22756" refString="WEITHOFER K. A. 1888. - Beitrage zur Kenntnis der Fauna von Pikermi bei Athen. Beitrage zur Palaontologie Osterreich-Ungarns und des Orients 6: 225 - 292." type="journal article" year="1888">Weithofer 1888</bibRefCitation>
;
<bibRefCitation id="EFBF77AEFFA13D01FE7ACEDBFDE2FE6D" author="GAUDRY A." box="[496,638,438,465]" pageId="25" pageNumber="476" refId="ref19232" refString="GAUDRY A. 1862 - 1867. - Animaux fossiles et geologie de l'Attique. Savy, Paris, 476 p." type="book" year="1862">Gaudry 1862</bibRefCitation>
- 1867;
<bibRefCitation id="EFBF77AEFFA13D01FF3BCEB8FE5CFE4C" author="PILGRIM G. &amp; HOPWOOD A. T." box="[177,448,469,496]" pageId="25" pageNumber="476" refId="ref21581" refString="PILGRIM G. &amp; HOPWOOD A. T. 1928 - Catalogue of the Pontian Bovidae of Europe. British Museum (Natural History), London, 106 p." type="book" year="1928">Pilgrim &amp; Hopwood1928</bibRefCitation>
), the latter at Samos (
<bibRefCitation id="EFBF77AEFFA13D01FFFCCE98FE9DFDAC" author="ANDREE J." box="[118,257,501,528]" pageId="25" pageNumber="476" pagination="135 - 175" refId="ref18631" refString="ANDREE J. 1926. - Neue Cavicornier aus dem Pliocan von Samos. Palaeontographica 67: 135 - 175." type="journal article" year="1926">Andree 1926</bibRefCitation>
). The boundary between these species, and their occurrences in other localities have long been debated (
<bibRefCitation id="EFBF77AEFFA13D01FE9ACD59FE3CFDF3" author="GENTRY A. W." box="[272,416,564,591]" pageId="25" pageNumber="476" pagination="231 - 296" refId="ref19254" refString="GENTRY A. W. 1971. - The earliest goats and other antelopes from the Samos Hipparion fauna. Bulletin of the British Museum (Natural History) Geology 20: 231 - 296." type="journal article" year="1971">Gentry 1971</bibRefCitation>
;
<bibRefCitation id="EFBF77AEFFA13D01FE27CD59FDCEFDF3" author="BOUVRAIN G." box="[429,594,564,591]" pageId="25" pageNumber="476" pagination="113 - 124" refId="ref18947" refString="BOUVRAIN G. 1982. - Revision du genre Prostrepsiceros Major 1891 (Mammalia, Bovidae). Palaontologische Zeitschrift 56: 113 - 124." type="journal article" year="1982">Bouvrain 1982</bibRefCitation>
) but recent discoveries and taxonomic revisions tend to support the distinction (
<bibRefCitation id="EFBF77AEFFA13D01FEE4CD19FDA8FD33" author="KOSTOPOULOS D. S." box="[366,564,628,655]" pageId="25" pageNumber="476" pagination="747 - 791" refId="ref20360" refString="KOSTOPOULOS D. S. 2005. - The Bovidae (Mammalia, Artiodactyla) from the late Miocene of AkkasdaGI, Turkey. Geodiversitas 27 (4): 747 - 791." type="journal article" year="2005">Kostopoulos 2005</bibRefCitation>
, 2006).
<taxonomicName id="4C2E71DCFFA13D01FFFACDF9FEDDFD11" baseAuthorityName="Andree" baseAuthorityYear="1826" box="[112,321,659,685]" class="Mammalia" family="Bovidae" genus="Prostrepsiceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="fraasi">
<emphasis id="B95AD64DFFA13D01FFFACDF9FEDDFD11" box="[112,321,659,685]" italics="true" pageId="25" pageNumber="476">Prostrepsiceros fraasi</emphasis>
</taxonomicName>
is larger, has a more strongly angled face on the neurocranium, less salient orbital rims, shallower post-cornual fossa, and more divergent horn-cores. It occurs at Samos, Maragha, and perhaps Perivolaki (although
<bibRefCitation id="EFBF77AEFFA13D01FE7DCC7FFF50FCF1" author="KOSTOPOULOS D. S." pageId="25" pageNumber="476" pagination="879 - 891" refId="ref20467" refString="KOSTOPOULOS D. S. 2009 b. - Contribution to the systematics and phylogeny of Prostrepsiceros vallesiensis Bouvrain, 1982 (Mammalia, Bovidae). Geodiversitas 31 (4): 879 - 891." type="journal article" year="2009">Kostopoulos [2009b]</bibRefCitation>
now believes that the Perivolaki form is closer to
<taxonomicName id="4C2E71DCFFA13D01FF5ACC3FFF20FC30" authority="(Rodler &amp; Weithofer, 1890))" baseAuthorityName="Rodler &amp; Weithofer" baseAuthorityYear="1890" class="Mammalia" family="Bovidae" genus="Prostrepsiceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="houtumschindleri">
<emphasis id="B95AD64DFFA13D01FF5ACC3FFE07FCD7" box="[208,411,849,875]" italics="true" pageId="25" pageNumber="476">P. houtumschindleri</emphasis>
(Rodler &amp; Weithofer, 1890))
</taxonomicName>
, while
<taxonomicName id="4C2E71DCFFA13D01FE8CCC1FFE38FC37" baseAuthorityName="Weithofer" baseAuthorityYear="1888" box="[262,420,881,907]" class="Mammalia" family="Bovidae" genus="Prostrepsiceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="rotundicornis">
<emphasis id="B95AD64DFFA13D01FE8CCC1FFE38FC37" box="[262,420,881,907]" italics="true" pageId="25" pageNumber="476">P. rotundicornis</emphasis>
</taxonomicName>
is present at Pikermi, Halmyropotamos, AkkasdaĞI, Ravin des Zouaves 5 (
<bibRefCitation id="EFBF77AEFFA13D01FFFDCCDDFE98FC77" author="GENTRY A. W." box="[119,260,944,971]" pageId="25" pageNumber="476" pagination="231 - 296" refId="ref19254" refString="GENTRY A. W. 1971. - The earliest goats and other antelopes from the Samos Hipparion fauna. Bulletin of the British Museum (Natural History) Geology 20: 231 - 296." type="journal article" year="1971">Gentry 1971</bibRefCitation>
;
<bibRefCitation id="EFBF77AEFFA13D01FE84CCDDFE2CFC77" author="BOUVRAIN G." box="[270,432,944,971]" pageId="25" pageNumber="476" pagination="113 - 124" refId="ref18947" refString="BOUVRAIN G. 1982. - Revision du genre Prostrepsiceros Major 1891 (Mammalia, Bovidae). Palaontologische Zeitschrift 56: 113 - 124." type="journal article" year="1982">Bouvrain 1982</bibRefCitation>
;
<bibRefCitation id="EFBF77AEFFA13D01FE30CCDDFD1CFC77" author="KOSTOPOULOS D. S." box="[442,640,944,971]" pageId="25" pageNumber="476" pagination="747 - 791" refId="ref20360" refString="KOSTOPOULOS D. S. 2005. - The Bovidae (Mammalia, Artiodactyla) from the late Miocene of AkkasdaGI, Turkey. Geodiversitas 27 (4): 747 - 791." type="journal article" year="2005">Kostopoulos 2005</bibRefCitation>
, 2006) and at Vozarci in the
<collectingCountry id="F3394ACFFFA13D01FE2ACCBDFF0FFBB7" name="Macedonia" pageId="25" pageNumber="476">Republic of Macedonia</collectingCountry>
(FYROM). Further finds are needed before the issue can be settled, because it may look hard to accept the occurrence of two contemporaneous and partly sympatric closely related species. If the distinction between these two species is accepted, the characters of the Str-2 frontlet clearly align it with
<taxonomicName id="4C2E71DCFFA13D01FF23CBC2FED6FB74" baseAuthorityName="Weithofer" baseAuthorityYear="1888" box="[169,330,1198,1224]" class="Mammalia" family="Bovidae" genus="Prostrepsiceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="rotundicornis">
<emphasis id="B95AD64DFFA13D01FF23CBC2FED6FB74" box="[169,330,1198,1224]" italics="true" pageId="25" pageNumber="476">P. rotundicornis</emphasis>
</taxonomicName>
. Both species are best known from sites similar in age to Pikermi, although some records may be earlier.
</paragraph>
<paragraph id="8B910A5FFFA13D01FF02CA60FE19FA1B" blockId="25.[111,648,216,1447]" pageId="25" pageNumber="476">
Two mandibles, FM-2332 and 2331 (
<figureCitation id="131516DAFFA13D01FD9CCA60FDE4FA94" box="[534,632,1293,1320]" captionStart="FIG" captionStartId="22.[113,124,1612,1629]" captionTargetBox="[146,1176,221,1589]" captionTargetId="figure-17@22.[113,1209,209,1608]" captionTargetPageId="22" captionText="FIG. 13. — Fossil mammals from the Miocene of Strumyani (SW Bulgaria): A-C, F, G, Palaeoreas lindermayeri Wagner, 1848; A, skull FM-2738, front view; B, mandible FM-2744; C, mandible FM-2783; F, frontlet FM-2740; G, frontlet FM-2780; D, I, J, Prostrepsiceros rotundicornis (Weithofer, 1888); D, mandible FM-2331; I, frontlet FM-2779; J, frontlet FM-2739; E, Palaeoryx sp., tooth row p2-m2 FM-2251; H, Tragoportax sp., skull FM-2094. Scale bar: A, G-J, 20 cm; B-E, 5 cm." figureDoi="http://doi.org/10.5281/zenodo.5381398" httpUri="https://zenodo.org/record/5381398/files/figure.png" pageId="25" pageNumber="476">Fig. 13D</figureCitation>
), differ from those assigned above to
<taxonomicName id="4C2E71DCFFA13D01FE70CA40FDFFFAFB" authorityName="Gaudry" authorityYear="1861" box="[506,611,1325,1351]" class="Mammalia" family="Bovidae" genus="Palaeoreas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="25" pageNumber="476" phylum="Chordata" rank="genus">
<emphasis id="B95AD64DFFA13D01FE70CA40FDFFFAFB" box="[506,611,1325,1351]" italics="true" pageId="25" pageNumber="476">Palaeoreas</emphasis>
</taxonomicName>
by their longer premolar row (
<tableCitation id="C6AC3FE4FFA13D01FE11CA21FE73FADB" box="[411,495,1356,1383]" captionStart="TABLE" captionStartId="23.[217,226,219,236]" captionText="TABLE 5. — Measurements (in mm) of bovid lower tooth-rows from the Miocene of Strumyani (SW Bulgaria)." pageId="25" pageNumber="476">Table 5</tableCitation>
), but they are otherwise little different, and their assignment to
<taxonomicName id="4C2E71DCFFA13D01FFFBCAE1FE9DFA19" authorityName="Major" authorityYear="1891" box="[113,257,1420,1445]" class="Mammalia" family="Bovidae" genus="Prostrepsiceros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="25" pageNumber="476" phylum="Chordata" rank="genus">
<emphasis id="B95AD64DFFA13D01FFFBCAE1FE9DFA19" box="[113,257,1420,1445]" italics="true" pageId="25" pageNumber="476">Prostrepsiceros</emphasis>
</taxonomicName>
is tentative.
</paragraph>
</subSubSection>
</treatment>
</document>