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<document id="F077B170339EE57902A2A7E10E965F91" ID-DOI="10.5281/zenodo.158325" ID-GBIF-Dataset="d4b9c969-40fe-4848-9bf8-e1aae8cf182d" ID-ISSN="1175­5326" ID-Zenodo-Dep="158325" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461027947031" checkinUser="plazi" docAuthor="Waichert, C. &amp; Azevedo, C. O." docDate="2004" docId="FD7C87A2D52CFFF1AA6437811A10C9FE" docLanguage="en" docName="zt00661.pdf" docOrigin="Zootaxa 661" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Pseudisobrachium triacutum Waichert &amp; Azevedo, 2004, sp. nov." docType="treatment" docVersion="9" lastPageNumber="6" masterDocId="0145FFDAD528FFF4AB6C363D1916C878" masterDocTitle="Fourteen new species of Pseudisobrachium (Hymenoptera, Bethylidae) from Atlantic rain forest of Espírito Santo, Brazil" masterLastPageNumber="22" masterPageNumber="1" pageNumber="5" updateTime="1698572738771" updateUser="plazi">
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<mods:title id="9F38FB2A0B4235C361885D47493A967E">Fourteen new species of Pseudisobrachium (Hymenoptera, Bethylidae) from Atlantic rain forest of Espírito Santo, Brazil</mods:title>
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</mods:role>
<mods:namePart id="977B2A46CB5201800AB2BF2FA31B2FAD">Waichert, C.</mods:namePart>
</mods:name>
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<mods:namePart id="30FA4C3DF310C7B2A887DE5A15632A4D">Azevedo, C. O.</mods:namePart>
</mods:name>
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<mods:date id="57DDACE2DB1ECB3DC8EF61F413EBAF98">2004</mods:date>
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<mods:number id="7AAF01CEA64D5CC4E4E82CADB40F7E87">661</mods:number>
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<mods:classification id="2C310F3F2286881F14B4976B31967D71">journal article</mods:classification>
<mods:identifier id="47A851F5DD0FBD7BCD36F0D19775D5D5" type="DOI">10.5281/zenodo.158325</mods:identifier>
<mods:identifier id="D4E862CA69A66499099FC82DF62E26A4" type="GBIF-Dataset">d4b9c969-40fe-4848-9bf8-e1aae8cf182d</mods:identifier>
<mods:identifier id="B19D3C8775F25A240DEE065EE87CABD4" type="ISSN">1175­5326</mods:identifier>
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<treatment id="FD7C87A2D52CFFF1AA6437811A10C9FE" ID-DOI="http://doi.org/10.5281/zenodo.6269220" ID-GBIF-Taxon="119607008" ID-Zenodo-Dep="6269220" LSID="urn:lsid:plazi:treatment:FD7C87A2D52CFFF1AA6437811A10C9FE" httpUri="http://treatment.plazi.org/id/FD7C87A2D52CFFF1AA6437811A10C9FE" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<subSubSection id="3DCF653FD52CFFF0AA6437811A75C9AD" box="[264,867,443,470]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="756A36B4D52CFFF0AA6437811A75C9AD" blockId="4.[264,867,443,470]" box="[264,867,443,470]" pageId="4" pageNumber="5">
<heading id="2E2281D8D52CFFF0AA6437811A75C9AD" bold="true" box="[264,867,443,470]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="47A1EAA6D52CFFF0AA6437811A75C9AD" bold="true" box="[264,867,443,470]" pageId="4" pageNumber="5">
<taxonomicName id="B2D54D37D52CFFF0AA6437811B71C9AE" box="[264,615,444,470]" class="Insecta" family="Bethylidae" genus="Pseudisobrachium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="triacutum" status="sp. nov.">
<emphasis id="47A1EAA6D52CFFF0AA6437811B71C9AE" bold="true" box="[264,615,444,470]" italics="true" pageId="4" pageNumber="5">Pseudisobrachium triacutum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5C9257DDD52CFFF0A90237861BDAC9AD" box="[622,716,443,469]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="EDEE2A31D52CFFF0A9B037861A4DC9AD" box="[732,859,443,469]" captionStart="FIGURES 1 – 11. 1 – 5" captionStartId="3.[264,383,1728,1752]" captionTargetBox="[336,1260,315,1715]" captionTargetId="figure@3.[329,1292,293,1718]" captionTargetPageId="3" captionText="FIGURES 1 – 11. 1 – 5, Pseudisobrachium ventriosum sp. nov. 1, head, dorsal view; 2, mandible, frontal view; 3, genitalia, ventral view; 4, aedeagus, lateral view; 5, paramere, lateral view; 6 – 11, Pseudisobrachium triacutum sp. nov. 6, head, dorsal view; 7, mandible, frontal view; 8, genitalia, ventral view; 9, aedeagus, lateral view; 10, ventral arm of paramere, lateral view; 11, dorsal arm of paramere, lateral view. (Scale bars = 250 μm for head and mandible, = 130 μm for genitalia)." httpUri="https://zenodo.org/record/158326/files/figure.png" pageId="4" pageNumber="5">Figs. 6–11</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3DCF653FD52CFFF0AA64343B1A01CDE9" pageId="4" pageNumber="5" type="description">
<paragraph id="756A36B4D52CFFF0AA64343B1B92CAE1" blockId="4.[264,1325,518,1905]" pageId="4" pageNumber="5">
<emphasis id="47A1EAA6D52CFFF0AA64343B18A8CA58" bold="true" box="[264,446,518,544]" pageId="4" pageNumber="5">Description.—</emphasis>
Male. Body length
<quantity id="B22D9B51D52CFFF0A9F3343A1A1DCA59" box="[671,779,519,545]" metricMagnitude="-3" metricUnit="m" metricValue="2.46" pageId="4" pageNumber="5" unit="mm" value="2.46">2.46 mm</quantity>
; LFW
<quantity id="B22D9B51D52CFFF0A830343A1ADECA59" box="[860,968,519,545]" metricMagnitude="-3" metricUnit="m" metricValue="1.68" pageId="4" pageNumber="5" unit="mm" value="1.68">1.68 mm</quantity>
. Color: head and thorax black, metasoma dark castaneous; mandible light castaneous; antenna dark castaneous, the three first segments light castaneous; legs light castaneous; wings subhyaline, veins dark castaneous. Pubescence conspicuous.
</paragraph>
<paragraph id="756A36B4D52CFFF0AA54349B18D1CBA1" blockId="4.[264,1325,518,1905]" pageId="4" pageNumber="5">
<emphasis id="47A1EAA6D52CFFF0AA54349B186DCAB8" bold="true" box="[312,379,678,704]" pageId="4" pageNumber="5">Head</emphasis>
(
<figureCitation id="EDEE2A31D52CFFF0AAE7349A18C3CAB9" box="[395,469,679,705]" captionStart="FIGURES 1 – 11. 1 – 5" captionStartId="3.[264,383,1728,1752]" captionTargetBox="[336,1260,315,1715]" captionTargetId="figure@3.[329,1292,293,1718]" captionTargetPageId="3" captionText="FIGURES 1 – 11. 1 – 5, Pseudisobrachium ventriosum sp. nov. 1, head, dorsal view; 2, mandible, frontal view; 3, genitalia, ventral view; 4, aedeagus, lateral view; 5, paramere, lateral view; 6 – 11, Pseudisobrachium triacutum sp. nov. 6, head, dorsal view; 7, mandible, frontal view; 8, genitalia, ventral view; 9, aedeagus, lateral view; 10, ventral arm of paramere, lateral view; 11, dorsal arm of paramere, lateral view. (Scale bars = 250 μm for head and mandible, = 130 μm for genitalia)." httpUri="https://zenodo.org/record/158326/files/figure.png" pageId="4" pageNumber="5">Fig. 6</figureCitation>
): mandible with 5 sharp teeth (
<figureCitation id="EDEE2A31D52CFFF0A82A349A1A99CAB9" box="[838,911,679,705]" captionStart="FIGURES 1 – 11. 1 – 5" captionStartId="3.[264,383,1728,1752]" captionTargetBox="[336,1260,315,1715]" captionTargetId="figure@3.[329,1292,293,1718]" captionTargetPageId="3" captionText="FIGURES 1 – 11. 1 – 5, Pseudisobrachium ventriosum sp. nov. 1, head, dorsal view; 2, mandible, frontal view; 3, genitalia, ventral view; 4, aedeagus, lateral view; 5, paramere, lateral view; 6 – 11, Pseudisobrachium triacutum sp. nov. 6, head, dorsal view; 7, mandible, frontal view; 8, genitalia, ventral view; 9, aedeagus, lateral view; 10, ventral arm of paramere, lateral view; 11, dorsal arm of paramere, lateral view. (Scale bars = 250 μm for head and mandible, = 130 μm for genitalia)." httpUri="https://zenodo.org/record/158326/files/figure.png" pageId="4" pageNumber="5">Fig. 7</figureCitation>
). Clypeus with trapezoidal median lobe, strongly projected, apical margin emarginate and angled, median carina bifurcate apically. First four antennal segments in a ratio of 26:8:18:12, segment III 2.33 X longer than thick, segment XI 2.25 X; pubescence of flagellum subappressed, with some erect setae. Frons coriaceous, with regular punctures, shallow and sparse. WH 0.8 X LH, WF 0.56 X WH, WF 1.07 X HE;
<collectingCountry id="0DC27624D52CFFF0A90935521BB3CBF1" box="[613,677,879,905]" name="Iceland" pageId="4" pageNumber="5">DAO</collectingCountry>
0.28 X WF; ocellar triangle compact, with acute frons angle; OOL 0.87 X WOT. Vertex slightly convex, corners rounded. VOL 1.0 X HE. Temples subparallel.
</paragraph>
<paragraph id="756A36B4D52CFFF0AA5435DB1854CD61" blockId="4.[264,1325,518,1905]" pageId="4" pageNumber="5">
<emphasis id="47A1EAA6D52CFFF0AA5435DB18AACC78" bold="true" box="[312,444,998,1024]" pageId="4" pageNumber="5">Mesosoma</emphasis>
: pronotum and mesoscutum coriaceous, punctures similar to those of frons, but smaller; notauli narrow and short, occupying only the superior third of mesoscutum, parapsidal furrow narrow and complete; scutellar disc with same texture as mesoscutum. Propodeal disc 1.07 X as long as wide, 1.68 X as long as high, areolate anteriorly and transversally striate posteriorly, median carina incomplete, lateral and posterior carinae present, declivity strigulate, lateral coriaceous. Mesopleuron coriaceous with elongate coriaceous callus. Fore wings with discoidal vein spectral, not interstitial with median vein.
</paragraph>
<paragraph id="756A36B4D52CFFF0AA54331B1A01CDE9" blockId="4.[264,1325,518,1905]" pageId="4" pageNumber="5">
<emphasis id="47A1EAA6D52CFFF0AA54331B18BACD38" bold="true" box="[312,428,1318,1344]" pageId="4" pageNumber="5">Genitalia</emphasis>
(
<figureCitation id="EDEE2A31D52CFFF0AAD2331A1B54CD39" box="[446,578,1319,1345]" captionStart="FIGURES 1 – 11. 1 – 5" captionStartId="3.[264,383,1728,1752]" captionTargetBox="[336,1260,315,1715]" captionTargetId="figure@3.[329,1292,293,1718]" captionTargetPageId="3" captionText="FIGURES 1 – 11. 1 – 5, Pseudisobrachium ventriosum sp. nov. 1, head, dorsal view; 2, mandible, frontal view; 3, genitalia, ventral view; 4, aedeagus, lateral view; 5, paramere, lateral view; 6 – 11, Pseudisobrachium triacutum sp. nov. 6, head, dorsal view; 7, mandible, frontal view; 8, genitalia, ventral view; 9, aedeagus, lateral view; 10, ventral arm of paramere, lateral view; 11, dorsal arm of paramere, lateral view. (Scale bars = 250 μm for head and mandible, = 130 μm for genitalia)." httpUri="https://zenodo.org/record/158326/files/figure.png" pageId="4" pageNumber="5">Figs. 8–11</figureCitation>
): paramere evenly wide; ventral arm 2.8 X wider than dorsal arm, vannus with 5 inclined grooves; aedeagus narrow, only slightly wider medially, apex emarginate medially, truncate in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="3DCF653FD52CFFF0AA5433A31A4ACF59" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="756A36B4D52CFFF0AA5433A31D7BCEF9" blockId="4.[264,1325,518,1905]" pageId="4" pageNumber="5">
<emphasis id="47A1EAA6D52CFFF0AA5433A31B58CDC0" bold="true" box="[312,590,1438,1464]" pageId="4" pageNumber="5">Material examined.—</emphasis>
<typeStatus id="AA6E8816D52CFFF0A92233A21BFBCDC1" box="[590,749,1439,1465]" pageId="4" pageNumber="5" type="holotype">HOLOTYPE</typeStatus>
: male,
<collectingCountry id="0DC27624D52CFFF0A82633A21AA1CDC1" box="[842,951,1439,1465]" name="Brazil" pageId="4" pageNumber="5">BRAZIL</collectingCountry>
,
<emphasis id="47A1EAA6D52CFFF0A8A433A21D60CDC0" box="[968,1142,1439,1464]" italics="true" pageId="4" pageNumber="5">Espírito Santo</emphasis>
, Santa Teresa, Estação Biológica de
<collectingCountry id="0DC27624D52CFFF0A97C33FA1BB7CD99" box="[528,673,1479,1505]" name="Saint Lucia" pageId="4" pageNumber="5">Santa Lúcia</collectingCountry>
,
<date id="016B1074D52CFFF0A9DC33FA1A54CD99" box="[688,834,1479,1505]" pageId="4" pageNumber="5" value="2001-07-23">23.VII.2001</date>
, sweeping, C.O. Azevedo &amp; R. Kawada col. (
<collectionCode id="13C4AE71D52CFFF0AA2A33D21882CE71" box="[326,404,1519,1545]" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="4" pageNumber="5">UFES</collectionCode>
).
<typeStatus id="AA6E8816D52CFFF0AAC733D21B44CE71" box="[427,594,1519,1545]" pageId="4" pageNumber="5" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="0DC27624D52CFFF0A93333D21BDACE71" box="[607,716,1519,1545]" name="Brazil" pageId="4" pageNumber="5">BRAZIL</collectingCountry>
,
<emphasis id="47A1EAA6D52CFFF0A9B733D21A91CE70" box="[731,903,1519,1544]" italics="true" pageId="4" pageNumber="5">Espírito Santo</emphasis>
,
<specimenCount id="63D3FD3DD52CFFF0A8F533D21D0ECE71" box="[921,1048,1519,1545]" pageId="4" pageNumber="5" type="male">131 males</specimenCount>
, Santa Teresa, Estação Biológica de
<collectingCountry id="0DC27624D52CFFF0AAC4302A1B2ECE49" box="[424,568,1559,1585]" name="Saint Lucia" pageId="4" pageNumber="5">Santa Lúcia</collectingCountry>
,
<date id="016B1074D52CFFF0A92A302A1A03CE49" box="[582,789,1559,1585]" pageId="4" pageNumber="5" value="2001-01-30" valueMax="2001-12-10" valueMin="2001-01-30">30.I–10.XII.2001</date>
, sweeping, C.O. Azevedo &amp; R. Kawada col. (
<collectionCode id="13C4AE71D52CFFF0AA7C30021848CE21" box="[272,350,1599,1625]" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="4" pageNumber="5">UFES</collectionCode>
);
<specimenCount id="63D3FD3DD52CFFF0AA14300218CCCE21" box="[376,474,1599,1625]" pageId="4" pageNumber="5" type="male">5 males</specimenCount>
, Cariacica, Reserva Biológica de Reserva Biológica de Duas Bocas,
<date id="016B1074D52CFFF0AA64305A1886CEF9" box="[264,400,1639,1665]" pageId="4" pageNumber="5" value="1996-09-24">24.IX.1996</date>
–
<date id="016B1074D52CFFF0AACD305A1B0CCEF9" box="[417,538,1639,1665]" pageId="4" pageNumber="5" value="1997-02-25">25.II.1997</date>
, sweeping, C.O. Azevedo &amp; H. S. Sá col. (
<collectionCode id="13C4AE71D52CFFF0AF7F305A1D77CEF9" box="[1043,1121,1639,1665]" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="4" pageNumber="5">UFES</collectionCode>
).
</paragraph>
<paragraph id="756A36B4D52CFFF0AA5430B31A4ACF59" blockId="4.[264,1325,518,1905]" pageId="4" pageNumber="5">
<emphasis id="47A1EAA6D52CFFF0AA5430B318CCCED0" bold="true" box="[312,474,1678,1704]" pageId="4" pageNumber="5">Variations.—</emphasis>
Specimens from Reserva Biológica de Duas Bocas with pronotum somewhat light castaneous. Some specimens with body more robust; head larger or narrower; notauli almost absent. Propodeal disc completely areolate, but cells smaller than the specimens from Estação Biológica de
<collectingCountry id="0DC27624D52CFFF0A9A5313A1A4ECF59" box="[713,856,1799,1825]" name="Saint Lucia" pageId="4" pageNumber="5">Santa Lúcia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="3DCF653FD52CFFF0AA5431131889CF09" pageId="4" pageNumber="5" type="discussion">
<paragraph id="756A36B4D52CFFF0AA5431131889CF09" blockId="4.[264,1325,518,1905]" pageId="4" pageNumber="5">
<emphasis id="47A1EAA6D52CFFF0AA54311318F6CF30" bold="true" box="[312,480,1838,1864]" pageId="4" pageNumber="5">Discussion.—</emphasis>
This species has the genitalia slender and more elongate than is typical of the genus.
</paragraph>
</subSubSection>
<subSubSection id="3DCF653FD52DFFF1AA543726187FC926" pageId="5" pageNumber="6" type="etymology">
<paragraph id="756A36B4D52DFFF1AA543726187FC926" blockId="5.[264,1322,283,390]" pageId="5" pageNumber="6">
<emphasis id="47A1EAA6D52DFFF1AA54372618F4C94D" bold="true" box="[312,482,283,309]" pageId="5" pageNumber="6">Etymology.—</emphasis>
The specific epithet refers the apical margin of the median lobe of clypeus.
</paragraph>
</subSubSection>
<subSubSection id="3DCF653FD52DFFF1AA5437561A10C9FE" box="[312,774,363,390]" pageId="5" pageNumber="6" type="distribution">
<paragraph id="756A36B4D52DFFF1AA5437561A10C9FE" blockId="5.[264,1322,283,390]" box="[312,774,363,390]" pageId="5" pageNumber="6">
<emphasis id="47A1EAA6D52DFFF1AA54375618E0C9FD" bold="true" box="[312,502,363,389]" pageId="5" pageNumber="6">Distribution.—</emphasis>
<collectingCountry id="0DC27624D52DFFF1AA9937511B2BC9FE" box="[501,573,364,390]" name="Brazil" pageId="5" pageNumber="6">Brazil</collectingCountry>
(Espírito Santo).
</paragraph>
</subSubSection>
</treatment>
</document>
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