210 lines
24 KiB
XML
210 lines
24 KiB
XML
<document id="C3373EF8B3776360BD17AFFA60D0B5EE" ID-CLB-Dataset="24276" ID-DOI="10.11646/zootaxa.4728.3.4" ID-GBIF-Dataset="a8e9f6a6-8e63-4edc-b479-57383f972b74" ID-ISSN="1175-5326" ID-Zenodo-Dep="3625852" ID-ZooBank="30F4E480-BCE1-43CE-91FE-EF59BD3103C4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1579787967588" checkinUser="plazi" docAuthor="Deschodt, Christian M., Sole, Catherine L. & Scholtz, Clarke H." docDate="2020" docId="FD0887F3A642136DDACEFAF5FE29A21A" docLanguage="en" docName="zootaxa.4728.3.4.pdf" docOrigin="Zootaxa 4728 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Upsa centennial Deschodt, Sole & Scholtz 2020, new species" docType="treatment" docUuid="95FFD29C-BFA4-44D6-A1FA-A039BC362372" docUuidSource="ZooBank" docVersion="6" lastPageNumber="361" masterDocId="0131FF8BA6401369DA59FFF6FFC8A305" masterDocTitle="Upsa centennial, a new genus and species (Coleoptera: Scarabaeidae: Scarabaeinae: Endroedyolini) from the Southern Mistbelt forest in South Africa celebrating 100 years of entomology at the University of Pretoria" masterLastPageNumber="362" masterPageNumber="357" pageNumber="359" updateTime="1698778638551" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="703D426736AA86B1D1A24AFDC51AA344">
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<mods:title id="18DFA1A5AE201008679579F7F341AA29">Upsa centennial, a new genus and species (Coleoptera: Scarabaeidae: Scarabaeinae: Endroedyolini) from the Southern Mistbelt forest in South Africa celebrating 100 years of entomology at the University of Pretoria</mods:title>
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<mods:name id="B21330B908DAF14DFE9F512DB2E812F7" type="personal">
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<mods:role id="17E9359AF9B236E3E6817B8FE50672DF">
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<mods:roleTerm id="DF24BC2A89F3D3565AE6F538B38B4938">Author</mods:roleTerm>
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<mods:namePart id="4416D9B70E71EFC91C6622F4386CC3C1">Deschodt, Christian M.</mods:namePart>
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<mods:roleTerm id="7FA299109260B9F6CEA1F29658BBD512">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="004F64C973CBA867AE39F2BF821ABC1F">Sole, Catherine L.</mods:namePart>
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</mods:name>
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<mods:name id="2E07DB3FBCEF9EBE3937A7D6B9617B9F" type="personal">
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<mods:roleTerm id="777C76D16AA9A6598BB65591C0E2EBEC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9DFB3C5114E71495E0EF13A406957F54">Scholtz, Clarke H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A8336CB7D23DC3EB937B5F50DF84F894">text</mods:typeOfResource>
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<mods:title id="F4E0B27A58BEF9072ABD1160060CD322">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F8E25092AF46B8D85FD28948ADABA983">
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<mods:date id="113A0246D8A33DA6219DE7CC607D3BAC">2020</mods:date>
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<mods:detail id="B70C1DBE5B1629E346D377ADD5F4DE68" type="pubDate">
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<mods:number id="706F90746B6EA80FEC2A4A91841B52C3">2020-01-23</mods:number>
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<mods:detail id="C105032A46CC682FC51CC85BDEC37195" type="volume">
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<mods:number id="9ED8882605031B00392F8C1E65964190">4728</mods:number>
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<mods:detail id="1B488F88880CA815E12BF5B7C22B2520" type="issue">
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<mods:classification id="A2BF28A5B01B18165E6D97E48FE1133E">journal article</mods:classification>
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<mods:identifier id="762D6DBE1EF02140A4DA227EAF7292E7" type="CLB-Dataset">24276</mods:identifier>
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<mods:identifier id="A2803225B2C79F899A60A279E077F386" type="DOI">10.11646/zootaxa.4728.3.4</mods:identifier>
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</mods:mods>
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<treatment id="FD0887F3A642136DDACEFAF5FE29A21A" ID-GBIF-Taxon="161838034" LSID="urn:lsid:zoobank.org:act:95FFD29C-BFA4-44D6-A1FA-A039BC362372" httpUri="http://treatment.plazi.org/id/FD0887F3A642136DDACEFAF5FE29A21A" lastPageId="4" lastPageNumber="361" pageId="2" pageNumber="359">
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<subSubSection id="3DBB656EA642136BDACEFAF5FCF2A618" box="[151,826,1282,1309]" pageId="2" pageNumber="359" type="nomenclature">
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<paragraph id="751E36E5A642136BDACEFAF5FCF2A618" blockId="2.[151,826,1282,1345]" box="[151,826,1282,1309]" pageId="2" pageNumber="359">
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<heading id="2E568189A642136BDACEFAF5FCF2A618" bold="true" box="[151,826,1282,1309]" fontSize="11" level="1" pageId="2" pageNumber="359" reason="1">
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<emphasis id="47D5EAF7A642136BDACEFAF5FCF2A618" bold="true" box="[151,826,1282,1309]" pageId="2" pageNumber="359">
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<taxonomicName id="B2A14D66A642136BDACEFAF5FD56A618" ID-CoL="7DM72" authority="Deschodt, Sole & Scholtz" authorityName="Deschodt, Sole & Scholtz" authorityYear="2020" box="[151,670,1282,1309]" class="Insecta" family="Scarabaeidae" genus="Upsa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="359" phylum="Arthropoda" rank="species" species="centennial" status="sp. nov.">
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<emphasis id="47D5EAF7A642136BDACEFAF5FE93A618" bold="true" box="[151,347,1282,1309]" italics="true" pageId="2" pageNumber="359">Upsa centennial</emphasis>
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Deschodt, Sole & Scholtz
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</taxonomicName>
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,
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<taxonomicNameLabel id="5CE6578CA642136BD8F3FAF5FCF2A618" box="[682,826,1283,1309]" pageId="2" pageNumber="359" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="3DBB656EA642136BDACEFAD1FE41A644" box="[151,393,1319,1345]" pageId="2" pageNumber="359" type="description">
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<paragraph id="751E36E5A642136BDACEFAD1FE41A644" blockId="2.[151,826,1282,1345]" box="[151,393,1319,1345]" pageId="2" pageNumber="359">
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(
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<figureCitation id="ED9A2A60A642136BDAC6FAD1FF25A644" box="[159,237,1319,1345]" captionStart="FIGURE 1" captionStartId="2.[151,250,1104,1129]" captionTargetBox="[144,1444,181,1078]" captionTargetId="figure@2.[144,1444,181,1078]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of all the known Endroedyolini species with the type locality of Upsa centennial Deschodt, Sole & Scholtz, new species in Alexandria forest. All the forests in South Africa (Mucina & Rutherford 2006) are marked in green on the main map and major cities indicated with black circles." figureDoi="http://doi.org/10.5281/zenodo.3625854" httpUri="https://zenodo.org/record/3625854/files/figure.png" pageId="2" pageNumber="359">Figs. 1</figureCitation>
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,
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<figureCitation id="ED9A2A60A642136BDAA2FAD1FE88A644" box="[251,320,1319,1345]" captionStart="FIGURE 2" captionStartId="3.[151,250,1911,1936]" captionTargetBox="[168,1421,530,1887]" captionTargetId="figure@3.[151,1436,517,1892]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Upsa centennial Deschodt, Sole & Scholtz, new species (A, C, E–G) and Outenikwanus tomentosus Scholtz & Howden, 1987 (B, D). A, B, habitus, dorsal view; C, D, line diagram of the meso- and metasternum; E, aedeagus, dorsal view; F, pygidium; G, labels. Scale bar = 1.0 mm for 2A–D and 0.5 mm for 2E, F." figureDoi="http://doi.org/10.5281/zenodo.3625856" httpUri="https://zenodo.org/record/3625856/files/figure.png" pageId="2" pageNumber="359">2A, C</figureCitation>
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, E–G)
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</paragraph>
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</subSubSection>
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<subSubSection id="3DBB656EA642136BDACEFA99FEF1A6F0" pageId="2" pageNumber="359" type="materials_examined">
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<paragraph id="751E36E5A642136BDACEFA99FB67A6A8" blockId="2.[151,1437,1390,2029]" pageId="2" pageNumber="359">
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<emphasis id="47D5EAF7A642136BDACEFA99FE8DA68C" bold="true" box="[151,325,1390,1417]" pageId="2" pageNumber="359">Type material.</emphasis>
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<materialsCitation id="C5C93CB8A642136BDB15FA99FB67A6A8" ID-GBIF-Occurrence="2558265180" collectingDate="1987-05-12" collectionCode="TMSA" collectorName="S. Afr & Endrody-Younga" location="Alexandria For. St." pageId="2" pageNumber="359" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
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<typeStatus id="AA1A8847A642136BDB15FA99FE7CA68C" box="[332,436,1391,1417]" pageId="2" pageNumber="359" type="holotype">Holotype</typeStatus>
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<specimenCount id="63A7FD6CA642136BDBE5FA99FE1CA68C" box="[444,468,1391,1417]" pageId="2" pageNumber="359" type="male">♂</specimenCount>
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: “
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<collectorName id="D8545333A642136BDBB3FA99FDE0A68C" box="[490,552,1391,1417]" pageId="2" pageNumber="359">S.Afr</collectorName>
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;SE.
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<collectingRegion id="B765F807A642136BD80AFA99FD1CA68C" box="[595,724,1391,1417]" country="South Africa" name="Western Cape" pageId="2" pageNumber="359">Cape Prov.</collectingRegion>
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<location id="707E603EA642136BD8BFFA99FC7DA68C" LSID="urn:lsid:plazi:treatment:FD0887F3A642136DDACEFAF5FE29A21A:707E603EA642136BD8BFFA99FC7DA68C" box="[742,949,1391,1417]" name="Alexandria For. St." pageId="2" pageNumber="359" stateProvince="Western Cape">Alexandria For.St.</location>
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| 33[°].43[ʹ] S–26[°].23[ʹ] E ||
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<date id="011F1025A642136BDF53FA98FAB6A68C" box="[1290,1406,1390,1417]" pageId="2" pageNumber="359" value="1987-05-12">
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<collectingDate id="115BE9CDA642136BDF53FA98FAB6A68C" box="[1290,1406,1390,1417]" pageId="2" pageNumber="359" value="1987-05-12">5.12.1987</collectingDate>
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</date>
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;E- Y:2553 | groundtraps, 2 days | leg.
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<collectorName id="D8545333A642136BD845FA65FD14A6A8" box="[540,732,1427,1453]" pageId="2" pageNumber="359">Endrödy-Younga</collectorName>
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|| groundtrap with | faeces bait” (
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<collectionCode id="13B0AE20A642136BDE17FA65FB6AA6A8" box="[1102,1186,1427,1453]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="2" pageNumber="359">TMSA</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="751E36E5A642136BDA9EFA41FEF1A6F0" blockId="2.[151,1437,1390,2029]" pageId="2" pageNumber="359">Additional notes from Endrödy-Younga’s field diary: “old indig.[enous] for.[est] on sand. Very dry cond.[itions].”</paragraph>
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</subSubSection>
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<subSubSection id="3DBB656EA642136ADA9EFA08FA82A3FF" lastPageId="3" lastPageNumber="360" pageId="2" pageNumber="359" type="description">
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<paragraph id="751E36E5A642136BDA9EFA08FEAFA53B" blockId="2.[151,1437,1390,2029]" pageId="2" pageNumber="359">
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<emphasis id="47D5EAF7A642136BDA9EFA08FDAAA51C" bold="true" box="[199,610,1534,1561]" pageId="2" pageNumber="359">
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Description of the
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<typeStatus id="AA1A8847A642136BDBF3FA09FDDAA51C" box="[426,530,1535,1561]" pageId="2" pageNumber="359" type="holotype">holotype</typeStatus>
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, male.
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</emphasis>
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Body: 2.3 (maximum length with head extended) x
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<quantity id="B2599B00A642136BDE99FA09FAD1A51C" box="[1216,1305,1535,1561]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="2" pageNumber="359" unit="mm" value="1.7">1.7 mm</quantity>
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(maximum width over elytra).
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</paragraph>
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<paragraph id="751E36E5A642136BDA9EF9B1FAF8A5AC" blockId="2.[151,1437,1390,2029]" pageId="2" pageNumber="359">
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<emphasis id="47D5EAF7A642136BDA9EF9B1FECFA564" bold="true" box="[199,263,1607,1633]" pageId="2" pageNumber="359">Head</emphasis>
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. Clypeus bidentate, with genae slightly protruding anteriorly to form two small denticles next to each denticle; teeth separated by almost two times their length; surface punctate with punctures close together; genal suture straight forming obtuse angle with frontoclypeal suture; frontoclypeal suture medially incomplete.
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</paragraph>
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<paragraph id="751E36E5A642136BDA9EF945FCDFA5F4" blockId="2.[151,1437,1390,2029]" pageId="2" pageNumber="359">
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<emphasis id="47D5EAF7A642136BDA9EF945FE88A5C8" bold="true" box="[199,320,1715,1741]" pageId="2" pageNumber="359">Pronotum</emphasis>
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. Surface covered with small punctures, punctures sometimes bearing setae, close together; setae light brown, recurved; lateral edges somewhat explanate.
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</paragraph>
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<paragraph id="751E36E5A642136BDA9EF90DFE59A458" blockId="2.[151,1437,1390,2029]" pageId="2" pageNumber="359">
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<emphasis id="47D5EAF7A642136BDA9EF90DFEDBA410" bold="true" box="[199,275,1787,1813]" pageId="2" pageNumber="359">Elytra</emphasis>
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. Striae with punctures separated by about two puncture widths, punctures slightly larger than width of striae; interstriae with single rows of setiferous punctures with short recurved setae, punctures separated by up to eight puncture widths.
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</paragraph>
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<paragraph id="751E36E5A642136BDA9EF891FB4DA4CC" blockId="2.[151,1437,1390,2029]" pageId="2" pageNumber="359">
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<emphasis id="47D5EAF7A642136BDA9EF891FE10A484" bold="true" box="[199,472,1895,1921]" pageId="2" pageNumber="359">Meso- and metasterna</emphasis>
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. Meso- and metasterna fused, mesometasternal suture evenly curved; mesosternum punctate, medial area with fewer punctures, mesosternum about eight puncture widths wide medially; metasternum with slightly smaller punctures that are separated by about two puncture widths (
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<figureCitation id="ED9A2A60A642136BDE46F859FBB0A4CC" box="[1055,1144,1967,1993]" captionStart="FIGURE 2" captionStartId="3.[151,250,1911,1936]" captionTargetBox="[168,1421,530,1887]" captionTargetId="figure@3.[151,1436,517,1892]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Upsa centennial Deschodt, Sole & Scholtz, new species (A, C, E–G) and Outenikwanus tomentosus Scholtz & Howden, 1987 (B, D). A, B, habitus, dorsal view; C, D, line diagram of the meso- and metasternum; E, aedeagus, dorsal view; F, pygidium; G, labels. Scale bar = 1.0 mm for 2A–D and 0.5 mm for 2E, F." figureDoi="http://doi.org/10.5281/zenodo.3625856" httpUri="https://zenodo.org/record/3625856/files/figure.png" pageId="2" pageNumber="359">Fig. 2C</figureCitation>
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).
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</paragraph>
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<paragraph id="751E36E5A642136BDA9EF824FB87A4E8" blockId="2.[151,1437,1390,2029]" box="[199,1103,2002,2029]" pageId="2" pageNumber="359">
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<emphasis id="47D5EAF7A642136BDA9EF824FEF2A4E8" bold="true" box="[199,314,2002,2029]" pageId="2" pageNumber="359">Pygidium</emphasis>
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||
. Indistinctly punctate with small closely arranged punctures (
|
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<figureCitation id="ED9A2A60A642136BD9B4F825FB8AA4E8" box="[1005,1090,2003,2029]" captionStart="FIGURE 2" captionStartId="3.[151,250,1911,1936]" captionTargetBox="[168,1421,530,1887]" captionTargetId="figure@3.[151,1436,517,1892]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Upsa centennial Deschodt, Sole & Scholtz, new species (A, C, E–G) and Outenikwanus tomentosus Scholtz & Howden, 1987 (B, D). A, B, habitus, dorsal view; C, D, line diagram of the meso- and metasternum; E, aedeagus, dorsal view; F, pygidium; G, labels. Scale bar = 1.0 mm for 2A–D and 0.5 mm for 2E, F." figureDoi="http://doi.org/10.5281/zenodo.3625856" httpUri="https://zenodo.org/record/3625856/files/figure.png" pageId="2" pageNumber="359">Fig. 2F</figureCitation>
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).
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</paragraph>
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<paragraph id="751E36E5A643136ADA9EFF6EFAE4A3D2" blockId="3.[151,1436,152,502]" pageId="3" pageNumber="360">
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<emphasis id="47D5EAF7A643136ADA9EFF6EFEFDA3B6" bold="true" box="[199,309,152,179]" pageId="3" pageNumber="360">Foretibia</emphasis>
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||
. Narrow at base and widening to two times the basal width at the first denticle; outside margin crenulate in basal half with three denticles in distal half; end of foretibia blunt with a downward pointing spur.
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</paragraph>
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<paragraph id="751E36E5A643136ADA9EFF16FA82A3FF" blockId="3.[151,1436,152,502]" box="[199,1354,224,251]" pageId="3" pageNumber="360">
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<emphasis id="47D5EAF7A643136ADA9EFF16FEF2A3FE" bold="true" box="[199,314,224,251]" pageId="3" pageNumber="360">Aedeagus</emphasis>
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. Parameres symmetrical with terminal area flattened, similar to other Endroedyolini species.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="3DBB656EA643136ADA9EFEF3FB23A21B" box="[199,1259,260,287]" pageId="3" pageNumber="360" type="etymology">
|
||
<paragraph id="751E36E5A643136ADA9EFEF3FB23A21B" blockId="3.[151,1436,152,502]" box="[199,1259,260,287]" pageId="3" pageNumber="360">
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<emphasis id="47D5EAF7A643136ADA9EFEF3FE86A21A" bold="true" box="[199,334,261,287]" pageId="3" pageNumber="360">Etymology.</emphasis>
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This new species celebrates a century of entomology at the University of Pretoria.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="3DBB656EA643136DDA9EFEDFFE29A21A" lastPageId="4" lastPageNumber="361" pageId="3" pageNumber="360" type="discussion">
|
||
<paragraph id="751E36E5A643136ADA9EFEDFFED2A2F3" blockId="3.[151,1436,152,502]" pageId="3" pageNumber="360">
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||
<emphasis id="47D5EAF7A643136ADA9EFEDFFEFCA246" bold="true" box="[199,308,297,323]" pageId="3" pageNumber="360">Remarks</emphasis>
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. The new genus occurs within the Southern Mistbelt group that consists of three different forest
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<typeStatus id="AA1A8847A643136ADF38FEDFFA54A246" box="[1377,1436,297,323]" pageId="3" pageNumber="360">types</typeStatus>
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as defined by
|
||
<bibRefCitation id="11304B14A643136ADB62FEBBFD96A262" author="Von Maltitz, G. & Mucina, L. & Geldenhuys, C. J. & Lawes, M. & Eeley, H. & Adie, H. & Vink, D. & Fleming, G. & Bailey, C." box="[315,606,332,359]" pageId="3" pageNumber="360" refId="ref4092" refString="Von Maltitz, G., Mucina, L., Geldenhuys, C. J., Lawes, M., Eeley, H., Adie, H., Vink, D., Fleming, G. & Bailey, C. (2003) Classification system for South African Indigenous Forests: An objective classification for the Department of Water Affairs and Forestry. Unpublished Environmentek Report ENV-P-C 2003 - 017, CSIR, Pretoria, 284 pp." type="book" year="2003">
|
||
Von Maltitz
|
||
<emphasis id="47D5EAF7A643136ADB95FEBBFDC8A263" box="[460,512,332,358]" italics="true" pageId="3" pageNumber="360">et al</emphasis>
|
||
. (2003)
|
||
</bibRefCitation>
|
||
. One of these forest
|
||
<typeStatus id="AA1A8847A643136AD908FEBBFC44A262" box="[849,908,333,359]" pageId="3" pageNumber="360">types</typeStatus>
|
||
comprises the Albany Coastal Mistbelt forest consisting of five isolated forest patches. These patches often cover undulating coastal plains and include Alexandria Forest (
|
||
<bibRefCitation id="11304B14A643136ADB79FE63FDE6A2AA" author="Von Maltitz, G. & Mucina, L. & Geldenhuys, C. J. & Lawes, M. & Eeley, H. & Adie, H. & Vink, D. & Fleming, G. & Bailey, C." box="[288,558,404,431]" pageId="3" pageNumber="360" refId="ref4092" refString="Von Maltitz, G., Mucina, L., Geldenhuys, C. J., Lawes, M., Eeley, H., Adie, H., Vink, D., Fleming, G. & Bailey, C. (2003) Classification system for South African Indigenous Forests: An objective classification for the Department of Water Affairs and Forestry. Unpublished Environmentek Report ENV-P-C 2003 - 017, CSIR, Pretoria, 284 pp." type="book" year="2003">
|
||
Von Maltitz
|
||
<emphasis id="47D5EAF7A643136ADBE9FE63FE2AA2AB" box="[432,482,404,430]" italics="true" pageId="3" pageNumber="360">et al</emphasis>
|
||
. 2003
|
||
</bibRefCitation>
|
||
). Although it receives less rain than most of the other forest
|
||
<typeStatus id="AA1A8847A643136ADE85FE63FADFA2AA" box="[1244,1303,405,431]" pageId="3" pageNumber="360">types</typeStatus>
|
||
(
|
||
<bibRefCitation id="11304B14A643136ADF7EFE62FE9DA2D6" author="Mucina, L. & Rutherford, M. C." pageId="3" pageNumber="360" refId="ref3948" refString="Mucina, L. & Rutherford, M. C. (Eds.) (2006) The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina & Rutherford 2006</bibRefCitation>
|
||
), rain can be expected any time of the year. This forest is protected within the Addo National Park boundaries.
|
||
</paragraph>
|
||
<caption id="21DE666DA643136ADACEF881FBD3A4D2" ID-DOI="http://doi.org/10.5281/zenodo.3625856" ID-Zenodo-Dep="3625856" httpUri="https://zenodo.org/record/3625856/files/figure.png" pageId="3" pageNumber="360" startId="3.[151,250,1911,1936]" targetBox="[168,1421,530,1887]" targetPageId="3">
|
||
<paragraph id="751E36E5A643136ADACEF881FBD3A4D2" blockId="3.[151,1436,1911,2008]" pageId="3" pageNumber="360">
|
||
<emphasis id="47D5EAF7A643136ADACEF881FEDCA495" bold="true" box="[151,276,1911,1936]" pageId="3" pageNumber="360">FIGURE 2.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="B2A14D66A643136ADBC4F88FFC9DA495" authority="Deschodt, Sole & Scholtz" authorityName="Deschodt, Sole & Scholtz" authorityYear="2020" box="[413,853,1911,1936]" class="Insecta" family="Scarabaeidae" genus="Upsa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="centennial" status="sp. nov.">
|
||
<emphasis id="47D5EAF7A643136ADBC4F88FFD8BA495" box="[413,579,1912,1936]" italics="true" pageId="3" pageNumber="360">Upsa centennial</emphasis>
|
||
Deschodt, Sole & Scholtz
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="5CE6578CA643136AD939F88EFC12A495" box="[864,986,1911,1936]" pageId="3" pageNumber="360" rank="species">new species</taxonomicNameLabel>
|
||
(A, C, E–G) and
|
||
<taxonomicName id="B2A14D66A643136ADECBF88EFE55A4B1" authority="Scholtz & Howden, 1987" authorityName="Scholtz & Howden" authorityYear="1987" class="Insecta" family="Scarabaeidae" genus="Outenikwanus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="360" phylum="Arthropoda" rank="species" species="tomentosus">
|
||
<emphasis id="47D5EAF7A643136ADECBF88EFA54A495" box="[1170,1436,1912,1936]" italics="true" pageId="3" pageNumber="360">Outenikwanus tomentosus</emphasis>
|
||
<bibRefCitation id="11304B14A643136ADACEF86DFE55A4B1" author="Scholtz, C. H. & Howden, H. F." box="[151,413,1947,1972]" pageId="3" pageNumber="360" pagination="75 - 119" refId="ref4042" refString="Scholtz, C. H. & Howden, H. F. (1987) A revision of the southern African Ca [n] thonina (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of the Entomological Society of Southern Africa, 50, 75 - 119." type="journal article" year="1987">Scholtz & Howden, 1987</bibRefCitation>
|
||
</taxonomicName>
|
||
(B, D). A, B, habitus, dorsal view; C, D, line diagram of the meso- and metasternum; E, aedeagus, dorsal view; F, pygidium; G, labels. Scale bar = 1.0 mm for 2A–D and 0.5 mm for 2E, F.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="751E36E5A644136DDA9EFF6FFE29A21A" blockId="4.[151,1437,152,287]" pageId="4" pageNumber="361">
|
||
The main forest patch of Alexandria forest is
|
||
<quantity id="B2599B00A644136DD880FF6FFCEAA3B6" box="[729,802,153,179]" metricMagnitude="4" metricUnit="m" metricValue="7.8" pageId="4" pageNumber="361" unit="km" value="78.0">78 km</quantity>
|
||
<superScript id="82D49BADA644136DD97BFF6EFCE2A3A3" attach="left" box="[802,810,152,166]" fontSize="6" pageId="4" pageNumber="361">2</superScript>
|
||
(
|
||
<bibRefCitation id="11304B14A644136DD962FF6FFBB2A3B1" author="Mucina, L. & Rutherford, M. C." box="[827,1146,153,180]" pageId="4" pageNumber="361" refId="ref3948" refString="Mucina, L. & Rutherford, M. C. (Eds.) (2006) The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina & Rutherford 2006</bibRefCitation>
|
||
) and there are four other forest patches of the same
|
||
<typeStatus id="AA1A8847A644136DDB9CFF4BFE3DA3D2" box="[453,501,189,215]" pageId="4" pageNumber="361">type</typeStatus>
|
||
that this species may possibly inhabit. On the basis of current knowledge this new species should be assessed as endangered B2ac(iv)—see
|
||
<bibRefCitation id="11304B14A644136DD979FF17FABBA3FE" author="IUCN Standards & Petitions Subcommittee" box="[800,1395,225,251]" pageId="4" pageNumber="361" refId="ref3832" refString="IUCN Standards and Petitions Subcommittee (2014) Guidelines for Using the IUCN Red List Categories and Criteria. Version 11. Prepared by the Standards and Petitions Subcommittee. Available from: http: // www. iucnredlist. org / documents / RedList- Guidelines. pdf. (accessed 27 September 2019)" type="book" year="2014">IUCN Standards and Petitions Subcommittee (2014)</bibRefCitation>
|
||
for an explanation of the criteria.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |