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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.454.7269" ID-GBIF-Dataset="1d5a0bdd-db86-4f99-bbf0-d05a7b50deda" ID-PMC="PMC4258740" ID-Pensoft-Pub="1313-2970-454-69" ID-PubMed="25493067" ID-ZBK="9A22E524BD38402FABDC506BC238CF94" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-454-69" ModsDocOrigin="ZooKeys 454" ModsDocTitle="Duan Karst of Guangxi: a kingdom of the cavernicolous genus Dongodytes Deuve (Coleoptera, Carabidae, Trechinae)" checkinTime="1451245117888" checkinUser="pensoft" docAuthor="Tian, Mingyi, Yin, Haomin &amp; Huang, Sunbin" docDate="2014" docId="F0A01DD5E2EFC8AA0B764584E1BB97F6" docLanguage="en" docName="ZooKeys 454: 69-107" docOrigin="ZooKeys 454" docSource="http://dx.doi.org/10.3897/zookeys.454.7269" docTitle="Dongodytes Deuve" docType="treatment" docVersion="4" lastPageNumber="74" masterDocId="3E36DC35FFE8FFDB8B7DD75B2E35D00D" masterDocTitle="Du'an Karst of Guangxi: a kingdom of the cavernicolous genus Dongodytes Deuve (Coleoptera, Carabidae, Trechinae)" masterLastPageNumber="107" masterPageNumber="69" pageNumber="71" updateTime="1668159606652" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Du'an Karst of Guangxi: a kingdom of the cavernicolous genus Dongodytes Deuve (Coleoptera, Carabidae, Trechinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tian, Mingyi</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yin, Haomin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Huang, Sunbin</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>454</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>69</mods:start>
<mods:end>107</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.454.7269</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.454.7269</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-454-69</mods:identifier>
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<treatment ID-GBIF-Taxon="152056355" LSID="urn:lsid:plazi:treatment:F0A01DD5E2EFC8AA0B764584E1BB97F6" httpUri="http://treatment.plazi.org/id/F0A01DD5E2EFC8AA0B764584E1BB97F6" lastPageId="5" lastPageNumber="74" pageId="2" pageNumber="71">
<subSubSection pageId="2" pageNumber="71" type="multiple">
<paragraph pageId="2" pageNumber="71">Taxon classification Animalia Coleoptera Carabidae</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="71" type="nomenclature">
<paragraph pageId="2" pageNumber="71">
Genus
<taxonomicName authority="Deuve" class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="2" pageNumber="71" phylum="Arthropoda" rank="genus">Dongodytes Deuve</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="71" type="reference_group">
<paragraph pageId="2" pageNumber="71">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="2" pageNumber="71" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
Deuve, 1993: 292
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="71" type="type species">
<paragraph pageId="2" pageNumber="71">Type species.</paragraph>
<paragraph pageId="2" pageNumber="71">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes fowleri" order="Coleoptera" pageId="2" pageNumber="71" phylum="Arthropoda" rank="species" species="fowleri">Dongodytes fowleri</taxonomicName>
Deuve, 1993: 292
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="71" type="type locality">
<paragraph pageId="2" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="2" pageNumber="71">
Jiabao Dong, in Bama County, northwestern Guangxi (
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
1998).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="72" pageId="2" pageNumber="71" type="diagnostic characters of the genus">
<paragraph pageId="2" pageNumber="71">Diagnostic characters of the genus.</paragraph>
<paragraph lastPageId="3" lastPageNumber="72" pageId="2" pageNumber="71">
Members of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">
<pageBreakToken pageId="3" pageNumber="72" start="start">Dongodytes</pageBreakToken>
</taxonomicName>
share the following combined characters: elytra remarkably elongate though much wider than prothorax and without shoulders; eyes completely effaced and depigmented; body especially head and prothorax strongly elongate; frontal furrows very short, mandibles long and slender, right mandible bidentate, palps and antennae very elongate and thin; propleura strongly tumid, visible from above; pronotum much longer than wide; elytra depressed medially on humeral parts, anterior and preapical dorsal pores present or not, humeral set of marginal umbilicate pores not aggregated, 1st pore widely located from 2nd and 3rd which are close to each other, 1st to 3rd more or less adjoining marginal gutter, 4th distinctly dorsal wards located and far from marginal gutter; 5th and 6th of middle set are close to each other; legs thin and very long, tarsi slender; protarsi not distinctly modified in male.
</paragraph>
<paragraph pageId="3" pageNumber="72">
Male genitalia are strongly diversified in
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
and could be important character states for phylogenetic analysis of the genus. Three types of the median lobe can be recognized. Type I, species of the subgenus
<taxonomicName lsidName="(Dongodytes)" pageId="3" pageNumber="72" rank="subGenus" subGenus="Dongodytes">Dongodytes</taxonomicName>
(s. str.) which are known only from
<normalizedToken originalValue="Duan">Du'an</normalizedToken>
Karst, the median lobe is very short and stout, basal orifice very large, but with a very small, sometimes indistinct, sagittal aileron, and the parameres are broader (Figs 33-38); Type II, species of the subgenus
<taxonomicName lsidName="(Dongodytodes)" pageId="3" pageNumber="72" rank="subGenus" subGenus="Dongodytodes">Dongodytodes</taxonomicName>
, which are all known from
<normalizedToken originalValue="Duan">Du'an</normalizedToken>
Karst, the median lobe is very elongate and thin, basal orifice comparatively small, but with a large sagittal aileron, and slender parameres (Figs 39-46); Type III, species of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
(s. str.) which are all known from other counties rather than
<normalizedToken originalValue="Duan">Du'an</normalizedToken>
: the median lobe is intermediately shaped between the two patterns described above, moderately elongate, rather stout, and with a large sagittal aileron; in particular, the median lobe distinctly curved at apex (
<bibRefCitation author="Deuve, T" journalOrPublisher="Bulletin de la Societe entomologique de France" pageId="18" pageNumber="87" pagination="291 - 296" title="Description de Dongodytesfowleri n. gen., sp. n., Coleoptere troglobie des karsts du Guangxi, Chine (Adephaga: Trechidae)." volume="98" year="1993">Deuve 1993</bibRefCitation>
: Fig. 2;
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
1998: Figs 1-4).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="72" type="systematic position of dongodytes">
<paragraph pageId="3" pageNumber="72">
Systematic position of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
.
</paragraph>
<paragraph pageId="3" pageNumber="72">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
is one of the most modified troglobiomorphic genera of trechines in the world. To determine the taxonomic position of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
within the tribe
<taxonomicName lsidName="" pageId="3" pageNumber="72" rank="tribe" tribe="Trechini">Trechini</taxonomicName>
remains a challenge.
<bibRefCitation author="Vigna Taglianti, A" journalOrPublisher="International Journal of Speleology" pageId="18" pageNumber="87" pagination="33 - 41" title="A new genus and species of troglobitic Trechinae (Coleoptera, Carabidae) from southern China." url="10.5038/1827-806X.25.1.3" volume="25" year="1997">Vigna Taglianti (1997)</bibRefCitation>
and
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
(1998) suggested it may be allied to the European
<taxonomicName class="Insecta" family="Carabidae" genus="Aphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphaenops" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Aphaenops</taxonomicName>
series, contrary to
<bibRefCitation author="Deuve, T" journalOrPublisher="Bulletin de la Societe entomologique de France" pageId="18" pageNumber="87" pagination="291 - 296" title="Description de Dongodytesfowleri n. gen., sp. n., Coleoptere troglobie des karsts du Guangxi, Chine (Adephaga: Trechidae)." volume="98" year="1993">Deuve (1993)</bibRefCitation>
who compared
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
with
<taxonomicName class="Insecta" family="Carabidae" genus="Sinaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinaphaenops" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Sinaphaenops</taxonomicName>
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
&amp; Wang, 1991. Because recent study based on molecular phylogenetic analysis have clearly demonstrated that the
<taxonomicName class="Insecta" family="Carabidae" genus="Aphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Aphaenops" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Aphaenops</taxonomicName>
series in the widest and traditional sense are restricted to the Pyrenean lineage (
<bibRefCitation author="Faille, A" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="18" pageNumber="87" pagination="97 - 106" title="A molecular phylogeny shows the single origin of the Pyrenean subterranean Trechini ground beetles (Coleoptera: Carabidae)." url="10.1016/j.ympev.2009.10.008" volume="54" year="2010">Faille et al. 2010</bibRefCitation>
,
<bibRefCitation author="Faille, A" journalOrPublisher="BMC Evolutionary Biology" pageId="18" pageNumber="87" pagination="1 - 16" title="A molecular phylogeny of Alpine subterranean Trechini (Coleoptera: Carabidae)." volume="13" year="2013">2013</bibRefCitation>
), we agree with
<normalizedToken originalValue="Deuves">Deuve's</normalizedToken>
opinion.
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
seems to be allied to its Chinese congeners such as
<taxonomicName class="Insecta" family="Carabidae" genus="Sinaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinaphaenops" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Sinaphaenops</taxonomicName>
and
<taxonomicName class="Insecta" family="Carabidae" genus="Pilosaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilosaphaenops" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Pilosaphaenops</taxonomicName>
Deuve &amp; Tian, 2008. For example, by having very short frontal furrows, very elongate genae and slender neck constriction, the head structures of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
are more or less similar to those of
<taxonomicName class="Insecta" family="Carabidae" genus="Sinaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinaphaenops" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Sinaphaenops</taxonomicName>
and
<taxonomicName class="Insecta" family="Carabidae" genus="Pilosaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilosaphaenops" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Pilosaphaenops</taxonomicName>
. Furthermore, all of the above genera have similar prothorax although latero-marginal setae are always absent in
<taxonomicName class="Insecta" family="Carabidae" genus="Sinaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinaphaenops" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Sinaphaenops</taxonomicName>
and
<taxonomicName class="Insecta" family="Carabidae" genus="Pilosaphaenops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilosaphaenops" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Pilosaphaenops</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="72" type="geographical distribution">
<paragraph pageId="3" pageNumber="72">Geographical distribution.</paragraph>
<paragraph pageId="3" pageNumber="72">
Endemic to northwestern Guangxi (Fig. 73). Members of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
are recorded from several counties of Hechi Prefecture. However, only a single species is known in each of Bama, Fengshan,
<normalizedToken originalValue="Tiane">Tian'e</normalizedToken>
and Dahua Counties, respectively. On the other hand, majority of the species (eight) are distributed in
<normalizedToken originalValue="Duan">Du'an</normalizedToken>
County. Thus, it is clear that, from the present knowledge, all species of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
are distributed in a very limited area of the river Hongshui He drainages in northwestern Guangxi. The river acts as a natural barrier for dispersal of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="3" pageNumber="72" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
at only a specific level. The nominate subgenus
<taxonomicName lsidName="(Dongodytes)" pageId="3" pageNumber="72" rank="subGenus" subGenus="Dongodytes">Dongodytes</taxonomicName>
(s. str.) covers a larger distribution range than the subgenus
<taxonomicName lsidName="(Dongodytodes)" pageId="3" pageNumber="72" rank="subGenus" subGenus="Dongodytodes">Dongodytodes</taxonomicName>
which is restricted to
<normalizedToken originalValue="Duan">Du'an</normalizedToken>
Karst in the eastern or northern bank of Hongshui He.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="73" type="taxonomic treatment">
<paragraph pageId="4" pageNumber="73">
<pageBreakToken pageId="4" pageNumber="73" start="start">Taxonomic</pageBreakToken>
treatment.
</paragraph>
<paragraph pageId="4" pageNumber="73">
Species of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
are divided into two subgenera,
<taxonomicName lsidName="(Dongodytes)" pageId="4" pageNumber="73" rank="subGenus" subGenus="Dongodytes">Dongodytes</taxonomicName>
(s. str.) Deuve and
<taxonomicName lsidName="(Dongodytodes)" pageId="4" pageNumber="73" rank="subGenus" subGenus="Dongodytodes">Dongodytodes</taxonomicName>
Tian. Both subgenera can be separated each other by shape of head, length of antennae, body pubescent or not, and chaetotaxal pattern on head, pronotum, elytra, and abdominal ventrite VII of male (
<bibRefCitation author="Tian, MY" journalOrPublisher="Subterranean Biology" pageId="18" pageNumber="87" pagination="57 - 64" title="A new subgenus and two new species of the troglobitic genus Dongodytes Deuve from Guangxi, China (Coleoptera, Carabidae)." url="10.3897/subtbiol.8.1232" volume="8" year="2011">Tian 2011</bibRefCitation>
). See the following key for details.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="74" pageId="4" pageNumber="73" type="key to subgenera and species of dongodytes deuve">
<paragraph pageId="4" pageNumber="73">
Key to subgenera and species of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
Deuve
</paragraph>
<paragraph lastPageId="5" lastPageNumber="74" pageId="4" pageNumber="73">
<table lastPageId="5" lastPageNumber="74" pageId="4" pageNumber="73">
<tr pageId="4" pageNumber="73">
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">
8101619
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes elongatus" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" species="elongatus">Dongodytes elongatus</taxonomicName>
rd625
<taxonomicName lsidName="(Dongodytes)" pageId="4" pageNumber="73" rank="subGenus" subGenus="Dongodytes">Dongodytes</taxonomicName>
</td>
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">2</td>
</tr>
<tr pageId="4" pageNumber="73">
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">
11152024rd726
<taxonomicName lsidName="(Dongodytodes)" pageId="4" pageNumber="73" rank="subGenus" subGenus="Dongodytodes">Dongodytodes</taxonomicName>
</td>
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">8</td>
</tr>
<tr pageId="4" pageNumber="73">
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes giraffa" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="giraffa">Dongodytes (s. str.) giraffa</taxonomicName>
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
</td>
</tr>
<tr pageId="4" pageNumber="73">
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">3</td>
</tr>
<tr pageId="4" pageNumber="73">
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">4</td>
</tr>
<tr pageId="4" pageNumber="73">
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">5</td>
</tr>
<tr pageId="4" pageNumber="73">
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">rd</td>
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes elongatus" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="elongatus">Dongodytes (s. str.) elongatus</taxonomicName>
sp. n.
</td>
</tr>
<tr pageId="4" pageNumber="73">
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">rd</td>
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes lani" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="lani">Dongodytes (s. str.) lani</taxonomicName>
sp. n.
</td>
</tr>
<tr pageId="4" pageNumber="73">
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">6</td>
</tr>
<tr pageId="4" pageNumber="73">
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">7</td>
</tr>
<tr pageId="4" pageNumber="73">
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes fowleri" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="fowleri">Dongodytes (s. str.) fowleri</taxonomicName>
Deuve
</td>
</tr>
<tr pageId="4" pageNumber="73">
<td colspan="1" pageId="4" pageNumber="73" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes grandis" order="Coleoptera" pageId="4" pageNumber="73" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="grandis">Dongodytes (s. str.) grandis</taxonomicName>
<normalizedToken originalValue="Uéno">Ueno</normalizedToken>
</td>
</tr>
<tr pageId="5" pageNumber="74">
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">
<pageBreakToken pageId="5" pageNumber="74" start="start">1627</pageBreakToken>
</td>
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes baxian" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="baxian">Dongodytes (s. str.) baxian</taxonomicName>
Tian
</td>
</tr>
<tr pageId="5" pageNumber="74">
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">1930</td>
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes troglodytes" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="troglodytes">Dongodytes (s. str.) troglodytes</taxonomicName>
sp. n.
</td>
</tr>
<tr pageId="5" pageNumber="74">
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">9</td>
</tr>
<tr pageId="5" pageNumber="74">
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">10</td>
</tr>
<tr pageId="5" pageNumber="74">
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">2111</td>
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) jinzhuensis" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="jinzhuensis" subGenus="Dongodytodes">Dongodytes (Dongodytodes) jinzhuensis</taxonomicName>
sp. n.
</td>
</tr>
<tr pageId="5" pageNumber="74">
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">2212</td>
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) inexpectatus" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="inexpectatus" subGenus="Dongodytodes">Dongodytes (Dongodytodes) inexpectatus</taxonomicName>
sp. n.
</td>
</tr>
<tr pageId="5" pageNumber="74">
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">2031</td>
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) deharvengi" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="deharvengi" subGenus="Dongodytodes">Dongodytes (Dongodytodes) deharvengi</taxonomicName>
Tian
</td>
</tr>
<tr pageId="5" pageNumber="74">
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">11</td>
</tr>
<tr pageId="5" pageNumber="74">
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">13144344</td>
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) brevipenis" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="brevipenis" subGenus="Dongodytodes">Dongodytes (Dongodytodes) brevipenis</taxonomicName>
sp. n.
</td>
</tr>
<tr pageId="5" pageNumber="74">
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">134546</td>
<td colspan="1" pageId="5" pageNumber="74" rowspan="1">
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) yaophilus" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="yaophilus" subGenus="Dongodytodes">Dongodytes (Dongodytodes) yaophilus</taxonomicName>
sp. n.
</td>
</tr>
</table>
</paragraph>
<caption pageId="5" pageNumber="74">
<paragraph pageId="5" pageNumber="74">
Figures 6-7. Elytral chaetotaxal patterns in
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
. 6
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes lani" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="lani">Dongodytes (s. str.) lani</taxonomicName>
sp. n. 7
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) jinzhuensis" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="jinzhuensis" subGenus="Dongodytodes">Dongodytes (Dongodytodes) jinzhuensis</taxonomicName>
sp. n.
</paragraph>
</caption>
<caption pageId="5" pageNumber="74">
<paragraph pageId="5" pageNumber="74">
Figures 8-11. Habitus of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
species. 8
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes elongatus" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="elongatus">Dongodytes (s. str.) elongatus</taxonomicName>
sp. n. female, paratype 9
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes troglodytes" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="troglodytes">Dongodytes (s. str.) troglodytes</taxonomicName>
sp. n. male, holotype 10
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes lani" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="lani">Dongodytes (s. str.) lani</taxonomicName>
sp. n. female, paratype 11
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) jinzhuensis" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="jinzhuensis" subGenus="Dongodytodes">Dongodytes (Dongodytodes) jinzhuensis</taxonomicName>
sp. n. female, paratype, from Jinzhu Dong I. Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption pageId="5" pageNumber="74">
<paragraph pageId="5" pageNumber="74">
Figures 12-15. Habitus of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
species. 12
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) inexpectatus" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="inexpectatus" subGenus="Dongodytodes">Dongodytes (Dongodytodes) inexpectatus</taxonomicName>
sp. n. male, holotype 13
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) brevipenis" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="brevipenis" subGenus="Dongodytodes">Dongodytes (Dongodytodes) brevipenis</taxonomicName>
sp. n. male, paratype, from Longwan: Nongqu Dong 14 ibid, from Chengjinag: Diaomao Dong 15
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) yaophilus" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="yaophilus" subGenus="Dongodytodes">Dongodytes (Dongodytodes) yaophilus</taxonomicName>
sp. n. female, paratype. Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption pageId="5" pageNumber="74">
<paragraph pageId="5" pageNumber="74">
Figures 16-24. Head of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
species. 16
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes baxian" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="baxian">Dongodytes (s. str.) baxian</taxonomicName>
M, male, holotype 17
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes elongatus" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="elongatus">Dongodytes (s. str.) elongatus</taxonomicName>
sp. n. male, holotype 18
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes lani" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="lani">Dongodytes (s. str.) lani</taxonomicName>
sp. n. male, holotype 19
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes troglodytes" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="troglodytes">Dongodytes (s. str.) troglodytes</taxonomicName>
sp. n. male, holotype 20
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) deharvengi" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="deharvengi" subGenus="Dongodytodes">Dongodytes (Dongodytodes) deharvengi</taxonomicName>
, male, holotype 21
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) jinzhuensis" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="jinzhuensis" subGenus="Dongodytodes">Dongodytes (Dongodytodes) jinzhuensis</taxonomicName>
sp. n. female, paratype 22
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) inexpectatus" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="inexpectatus" subGenus="Dongodytodes">Dongodytes (Dongodytodes) inexpectatus</taxonomicName>
sp. n. male, holotype 23
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) brevipenis" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="brevipenis" subGenus="Dongodytodes">Dongodytes (Dongodytodes) brevipenis</taxonomicName>
sp. n. male, holotype 24
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) yaophilus" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="yaophilus" subGenus="Dongodytodes">Dongodytes (Dongodytodes) yaophilus</taxonomicName>
sp. n. male, holotype.
</paragraph>
</caption>
<caption pageId="5" pageNumber="74">
<paragraph pageId="5" pageNumber="74">
Figures 25-26. Abdominal ventrite VII of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
in male. 25
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes lani" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="lani">Dongodytes (s. str.) lani</taxonomicName>
sp. n., paratype 26
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) jinzhuensis" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="jinzhuensis" subGenus="Dongodytodes">Dongodytes (Dongodytodes) jinzhuensis</taxonomicName>
sp. n., paratype.
</paragraph>
</caption>
<caption pageId="5" pageNumber="74">
<paragraph pageId="5" pageNumber="74">
Figures 27-32. Basal parts of prothorax and elytra of subgenus
<taxonomicName lsidName="(Dongodytes)" pageId="5" pageNumber="74" rank="subGenus" subGenus="Dongodytes">Dongodytes</taxonomicName>
(s. str.). 27
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes baxian" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="baxian">Dongodytes (s. str.) baxian</taxonomicName>
, male, holotype 28
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes elongatus" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="elongatus">Dongodytes (s. str.) elongatus</taxonomicName>
sp. n., male, holotype 29
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes lani" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="lani">Dongodytes (s. str.) lani</taxonomicName>
sp. n., female, paratype 30
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes troglodytes" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="troglodytes">Dongodytes (s. str.) troglodytes</taxonomicName>
sp. n., male, holotype 31
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) deharvengi" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="deharvengi" subGenus="Dongodytodes">Dongodytes (Dongodytodes) deharvengi</taxonomicName>
Tian, male, paratype 32
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) jinzhuensis" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="jinzhuensis" subGenus="Dongodytodes">Dongodytes (Dongodytodes) jinzhuensis</taxonomicName>
sp. n., female, paratype.
</paragraph>
</caption>
<caption pageId="5" pageNumber="74">
<paragraph pageId="5" pageNumber="74">
Figures 33-38. Male genitalia of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="genus">Dongodytes</taxonomicName>
(s. str.) species (median lobe and parameres in lateral view, apical part of median lobe in dorsal view). 33-34
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes elongatus" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="elongatus">Dongodytes (s. str.) elongatus</taxonomicName>
sp. n. 35-36
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes lani" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="lani">Dongodytes (s. str.) lani</taxonomicName>
sp. n. 37-38
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes troglodytes" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="troglodytes">Dongodytes (s. str.) troglodytes</taxonomicName>
sp. n.
</paragraph>
</caption>
<caption pageId="5" pageNumber="74">
<paragraph pageId="5" pageNumber="74">
Figures 39-46. Male genitalia of
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes)" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="subGenus" subGenus="Dongodytodes">Dongodytes (Dongodytodes)</taxonomicName>
species (median lobe and parameres in lateral view, apical part of median lobe in dorsal view). 39-40
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) jinzhuensis" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="jinzhuensis" subGenus="Dongodytodes">Dongodytes (Dongodytodes) jinzhuensis</taxonomicName>
sp. n. 41-42
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) inexpectatus" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="inexpectatus" subGenus="Dongodytodes">Dongodytes (Dongodytodes) inexpectatus</taxonomicName>
sp. n. 43-44
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) brevipenis" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="brevipenis" subGenus="Dongodytodes">Dongodytes (Dongodytodes) brevipenis</taxonomicName>
sp. n. 45-46
<taxonomicName class="Insecta" family="Carabidae" genus="Dongodytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dongodytes (Dongodytodes) yaophilus" order="Coleoptera" pageId="5" pageNumber="74" phylum="Arthropoda" rank="species" species="yaophilus" subGenus="Dongodytodes">Dongodytes (Dongodytodes) yaophilus</taxonomicName>
sp. n.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>