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<document ID-CLB-Dataset="269602" ID-DOI="http://dx.doi.org/10.3897/dez.70.107425" ID-GBIF-Dataset="360868d3-9133-466c-bfae-e8de48028017" ID-Pensoft-Pub="1860-1324-2-311" ID-Pensoft-UUID="65D0B2390EBE553382C9727ECC9CBF8F" ID-ZooBank="8D3E277C424C440B8FE878085239C2A2" ModsDocID="1860-1324-70-2-311" checkinTime="1695893248727" checkinUser="pensoft" docAuthor="Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek &amp; Schmidt, Joachim" docDate="2023" docId="F06932799AAE5261B94FFF356556DA17" docLanguage="en" docName="DeutEntomolZeit 70(2): 311-335" docOrigin="Deutsche Entomologische Zeitschrift 70 (2)" docPubDate="2023-09-27" docSource="http://dx.doi.org/10.3897/dez.70.107425" docTitle="Trechus (Abunetrechus) habeshanicus Quéinnec &amp; Ollivier" docType="treatment" docVersion="2" id="65D0B2390EBE553382C9727ECC9CBF8F" lastPageNumber="311" masterDocId="65D0B2390EBE553382C9727ECC9CBF8F" masterDocTitle="Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini)" masterLastPageNumber="335" masterPageNumber="311" pageNumber="311" updateTime="1695893870965" updateUser="ExternalLinkService">
<mods:mods id="E0BA9898FA79820511986FF8B8220B89" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="255383C982A0FC2E07DF53EF06C7CEA1">Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini)</mods:title>
</mods:titleInfo>
<mods:name id="08FFF5DA1878B7EE302E0846359B05BD" type="personal">
<mods:role id="9D65AE98315074916B8C5A69BFF8F443">
<mods:roleTerm id="FCECA1590997242BA35B56EFE7F3D816">Author</mods:roleTerm>
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<mods:namePart id="59F56B72DED918A4DCF0C65B78E183B7">Faille, Arnaud</mods:namePart>
<mods:nameIdentifier id="178F2A6C2E4C8C05C624F4408652E3F9" type="ORCID">https://orcid.org/0000-0003-3274-5915</mods:nameIdentifier>
<mods:affiliation id="95C24EDAB1C0E001A9B2FE76684601F2">Department of Entomology, Stuttgart State Museum of Natural History, Stuttgart, Germany</mods:affiliation>
<mods:nameIdentifier id="9359A50594B05180A95AFF3D177C008E" type="email">arnaud.faille@smns-bw.de</mods:nameIdentifier>
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<mods:namePart id="D6F85413D375BDB88E657B9D532C3725">Hofmann, Sylvia</mods:namePart>
<mods:affiliation id="0F4551AD04AE4F8D3B9638E5B1315A91">Leibniz Institute for the Analysis of Biodiversity Change, Museum Koenig, 53113 Bonn, Germany</mods:affiliation>
</mods:name>
<mods:name id="07DB4453F06BB6D0335F95A0D359FAE0" type="personal">
<mods:role id="F11F6279061E77FD1EA46BF399114FF4">
<mods:roleTerm id="96A2ED78A9F737605DB64F344C2D954B">Author</mods:roleTerm>
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<mods:namePart id="72586838149F14AAEC1196E55DF6AE35">Merene, Yeshitla</mods:namePart>
<mods:nameIdentifier id="73D000B1BDD369237DA98DD9CE3F7A95" type="ORCID">https://orcid.org/0000-0003-2998-822X</mods:nameIdentifier>
<mods:affiliation id="B62EAF3CB231651ED8993E29F20B574E">Faculty of Biology, Philipps University Marburg, Marburg, Germany &amp; Department of Zoological Sciences, Addis Ababa University, Addis Ababa, Ethiopia &amp; Amhara Agricultural Research Institute, Bahir Dar, Ethiopia</mods:affiliation>
</mods:name>
<mods:name id="51B233FB1114506044320BC2AFA6C0D2" type="personal">
<mods:role id="6FF527C11344209C44A6F0BF6E2DB172">
<mods:roleTerm id="CC4BCB4467D6B032C0DAB7CAE6EE205C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D3D31C8468881EDF1E18023CBF20F97D">Hauth, David</mods:namePart>
<mods:affiliation id="AB2BFFAFA36ECCFFD3AA40BE50D2019B">Faculty of Biology, Philipps University Marburg, Marburg, Germany</mods:affiliation>
</mods:name>
<mods:name id="79E6F5ED7BC1F19176A3069A52145D62" type="personal">
<mods:role id="7E20F1E23020022A444548FD19BEDD4C">
<mods:roleTerm id="94D356C7689D0372BFE12CE5195CC384">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="15D8146D8B595FF10F20FF6CC40F743B">Opgenoorth, Lars</mods:namePart>
<mods:affiliation id="1CAB1F917A9312325DFBD5B7827B640B">Faculty of Biology, Philipps University Marburg, Marburg, Germany</mods:affiliation>
</mods:name>
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<mods:role id="CF42256378BD180481FE028736659926">
<mods:roleTerm id="396FBD4E51CF335FC00473FAEAEED9AA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="26E0A965E8C1F9E37CCD7AD0BD138B46">Woldehawariat, Yitbarek</mods:namePart>
<mods:nameIdentifier id="C2E3550210ED35A46839BD9918FC73C6" type="ORCID">https://orcid.org/0000-0002-0918-8906</mods:nameIdentifier>
<mods:affiliation id="A85DDDDB11B58F89FBC1C40A5D4C9E4C">Department of Zoological Sciences, Addis Ababa University, Addis Ababa, Ethiopia</mods:affiliation>
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<mods:roleTerm id="9F1FC6878ACDC53D5CDD622A06A3C9C0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F171FC0F4B887E82A0CC50B5AC5F1D74">Schmidt, Joachim</mods:namePart>
<mods:affiliation id="C097A3F478296AE2869FDF973B5CDFBB">General and Systematic Zoology, University of Rostock, Rostock, Germany</mods:affiliation>
<mods:nameIdentifier id="D1DC2F63C027D8956C32985680F08273" type="email">agonumschmidt@hotmail.com</mods:nameIdentifier>
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<mods:title id="9AD6A9081CE136BF1562C2B6EAE4C973">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="D14AAD5AF102EE63DDBF5E8ECA5DC617">2023</mods:date>
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<mods:number id="5967E24DCD5AE5E27D49DEAF100A7FF5">2023-09-27</mods:number>
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<mods:number id="08E1580D88F9C8383978CCCD6C773525">70</mods:number>
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<mods:start id="DAD7CB283AD06DED791A57061FC0612B">311</mods:start>
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<mods:url id="78D5E88EF8E672F1F65E46FC8E9E9F19">http://dx.doi.org/10.3897/dez.70.107425</mods:url>
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<mods:classification id="D0FF3CC57BFF0BA77FE36F1F61046CBC">journal article</mods:classification>
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<treatment id="F06932799AAE5261B94FFF356556DA17" LSID="urn:lsid:plazi:treatment:F06932799AAE5261B94FFF356556DA17" httpUri="http://treatment.plazi.org/id/F06932799AAE5261B94FFF356556DA17" lastPageNumber="311" pageId="0" pageNumber="311">
<subSubSection id="25D1B06E232585608EF4B86DB97AD47A" pageId="0" pageNumber="311" type="nomenclature">
<paragraph id="F88492FEF2C2488918DB6BC7393ED311" pageId="0" pageNumber="311">
<taxonomicName id="C1A11BD92123A5C888ACA487AF1BD614" LSID="F0693279-9AAE-5261-B94F-FF356556DA17" authority="Queinnec &amp; Ollivier" authorityName="Quéinnec &amp; Ollivier" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus (Abunetrechus) habeshanicus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="species" species="habeshanicus" subGenus="Abunetrechus">
Trechus (Abunetrechus) habeshanicus
<normalizedToken id="95140E34852718A90C43BDDEC5460E15" originalValue="Quéinnec">Queinnec</normalizedToken>
&amp; Ollivier
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="CC520C47D8E537A791F1C44D0A5487A3" pageId="0" pageNumber="311" type="description">
<paragraph id="CD6EF1F865AD422AB568DC79ED5A6C32" pageId="0" pageNumber="311">
<figureCitation id="6DA29DCE0FECA9F75A0CDEB397EA3C2A" captionStart="Figures 21–23" captionStartId="F8" captionText="Figures 21 - 23. Trechus subgenus Abunetrechus nov., dorsal aspect of body (21) and head (22), and anterior part of left elytron (23). 21, 22. T. habeshaicus Queinnec &amp; Ollivier; 23. T. bipartitus Raffray. The small white circles in Fig. 22 mark the insertion points of the clypeal setae; the arrow in Fig. 23 points to the insertion of the anterior elytral discal seta." figureDoi="10.3897/dez.70.107425.figures21-23" httpUri="https://binary.pensoft.net/fig/913118" pageId="0" pageNumber="311">Figs 21</figureCitation>
<figureCitation id="315B676C5D515D600BE9C5A69139EE03" captionStart="Figures 21–23" captionStartId="F8" captionText="Figures 21 - 23. Trechus subgenus Abunetrechus nov., dorsal aspect of body (21) and head (22), and anterior part of left elytron (23). 21, 22. T. habeshaicus Queinnec &amp; Ollivier; 23. T. bipartitus Raffray. The small white circles in Fig. 22 mark the insertion points of the clypeal setae; the arrow in Fig. 23 points to the insertion of the anterior elytral discal seta." figureDoi="10.3897/dez.70.107425.figures21-23" httpUri="https://binary.pensoft.net/fig/913118" pageId="0" pageNumber="311">, 22</figureCitation>
<figureCitation id="9D93C74C7AE614787CBA494366ED088C" captionStart="Figures 24–29" captionStartId="F9" captionText="Figures 24 - 29. Trechus subgenus Abunetrechus nov., aedeagus in left lateral aspect (24, 26, 28) and dorsal aspect (25, 27, 29). 24, 25. T. bipartitus Raffray; 26, 27. T. lalibelae Queinnec &amp; Ollivier; 28, 29. T. habeshaicus Queinnec &amp; Ollivier." figureDoi="10.3897/dez.70.107425.figures24-29" httpUri="https://binary.pensoft.net/fig/913119" pageId="0" pageNumber="311">, 28</figureCitation>
<figureCitation id="E5C7A2E9499F3DD2C0D0B244E59256E3" captionStart="Figures 24–29" captionStartId="F9" captionText="Figures 24 - 29. Trechus subgenus Abunetrechus nov., aedeagus in left lateral aspect (24, 26, 28) and dorsal aspect (25, 27, 29). 24, 25. T. bipartitus Raffray; 26, 27. T. lalibelae Queinnec &amp; Ollivier; 28, 29. T. habeshaicus Queinnec &amp; Ollivier." figureDoi="10.3897/dez.70.107425.figures24-29" httpUri="https://binary.pensoft.net/fig/913119" pageId="0" pageNumber="311">, 29</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6C445B1704E951B6CA9E16E3690ED369" pageId="0" pageNumber="311" type="reference_group">
<paragraph id="F0FBB3A99941A0C574B86D381AC62C35" pageId="0" pageNumber="311">
<taxonomicName id="05C06701DE37BAACC58F1581A22E42AE" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus (Abunetrechus) habeshanicus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="species" species="habeshanicus" subGenus="Abunetrechus">Trechus (Abunetrechus) habeshanicus</taxonomicName>
:
<bibRefCitation id="410F54089AE4FEEB5F7BD776463C2350" DOI="https://doi.org/10.12976/jib/2021.27.1.1" author="Queinnec, E" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="311" pagination="1 - 70" refId="B43" refString="Queinnec, E, Ollivier, E, Reeb, C, 2021. Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae). Journal of Insect Biodiversity 27 (1): 1 - 70, DOI: https://doi.org/10.12976/jib/2021.27.1.1" title="Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae)." url="https://doi.org/10.12976/jib/2021.27.1.1" volume="27" year="2021">
<normalizedToken id="386F069AFF1514A20A054FF2B57BF720" originalValue="Quéinnec">Queinnec</normalizedToken>
et al. 2021
</bibRefCitation>
: 23; locus typicus: Guassa Plateau
<geoCoordinate id="0D45A3087221E234C6A509B70EDE2D47" degrees="10" direction="north" minutes="17" orientation="latitude" precision="15" seconds="19" value="10.288611">10°17'19&quot;N</geoCoordinate>
,
<geoCoordinate id="EE7AAB37E62F7AF860B7DF3354DDD90D" degrees="39" direction="east" minutes="48" orientation="longitude" precision="15" seconds="13" value="39.80361">39°48'13&quot;E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="A650FF4AF680FEFA54CFDC455B060EAF" pageId="0" pageNumber="311" type="type material">
<paragraph id="E39B6EC7F21103C4DAAB06C387F5CA5C" pageId="0" pageNumber="311">Type material.</paragraph>
<paragraph id="9E8B0FBBABD4AF9FE1E23E335A309A00" pageId="0" pageNumber="311">
Not studied. Identification is based on the original description, including habitus and male genital figures of the type specimens (
<bibRefCitation id="D98C587AF47FAD69F8689B610D3BE15B" DOI="https://doi.org/10.12976/jib/2021.27.1.1" author="Queinnec, E" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="311" pagination="1 - 70" refId="B43" refString="Queinnec, E, Ollivier, E, Reeb, C, 2021. Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae). Journal of Insect Biodiversity 27 (1): 1 - 70, DOI: https://doi.org/10.12976/jib/2021.27.1.1" title="Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae)." url="https://doi.org/10.12976/jib/2021.27.1.1" volume="27" year="2021">
<normalizedToken id="F0E0B532E0A5F218A5393812B62FFFF6" originalValue="Quéinnec">Queinnec</normalizedToken>
et al. 2021
</bibRefCitation>
), as well as on comprehensive material collected at the type locality (see below).
</paragraph>
</subSubSection>
<subSubSection id="6F21EE094ABB00A61F312E47A2C641ED" pageId="0" pageNumber="311" type="materials_examined">
<paragraph id="A60FEF316EBEC67359B547BFA1443580" pageId="0" pageNumber="311">Additional material.</paragraph>
<paragraph id="B0FA66119BEDC6D48495C7397B2709AB" pageId="0" pageNumber="311">
<materialsCitation id="B2B74283A2E31E4309F68A7D0E9F9AED" collectingDate="2022-05-18" collectingDateMax="2022-05-20" collectingDateMin="2022-05-18" collectorName="J. Schmidt, Yeshitla M." country="Ethiopia" elevation="3330" latitude="10.434167" location="northern Guassa Plateau" longLatPrecision="21" longitude="39.787777" specimenCount="1" stateProvince="Amhara">
98 exx. (males, females),
<collectingCountry id="02255B5613493F60CAD14AE37DDB3585" name="Ethiopia">Ethiopia</collectingCountry>
,
<collectingRegion id="8E28D1E702740CF5F9C540F941F26C60" country="Ethiopia" name="Amara">Amhara</collectingRegion>
,
<location id="77EA4CBFF14B80A3076A0B620915BA36" LSID="urn:lsid:plazi:treatment:F06932799AAE5261B94FFF356556DA17:77EA4CBFF14B80A3076A0B620915BA36" country="Ethiopia" latitude="10.434167" longLatPrecision="21" longitude="39.787777" name="northern Guassa Plateau" stateProvince="Amhara">northern Guassa Plateau</location>
, near Guassa Comm. Lodge
<quantity id="5B36D05DC624267905B1EC2620DF4A34" metricMagnitude="3" metricUnit="m" metricValue="3.3299999999999996" unit="m" value="3330.0">
<elevation id="446025733DFD16BA9B5CAA8201AE304D" metricMagnitude="3" metricUnit="m" metricValue="3.3299999999999996" unit="m" value="3330.0">3330 m</elevation>
</quantity>
,
<geoCoordinate id="DFFA82F7FD1D7A09517DBF3260A23420" degrees="10" direction="north" minutes="17" orientation="latitude" precision="15" seconds="17" value="10.288056">10°17'17&quot;N</geoCoordinate>
,
<geoCoordinate id="06E66FEF279D2730A8E1D6A8128C773A" degrees="39" direction="east" minutes="47" orientation="longitude" precision="15" seconds="54" value="39.798332">39°47'54&quot;E</geoCoordinate>
,
<collectingDate id="BA3DF4A4DDADEFE5A6A17A45BED4DD37" value="2022-05-18">18.V.2022</collectingDate>
, leg.
<collectorName id="03FBB9DE8983B3DB10FCF9FEAD87AB8C">J. Schmidt</collectorName>
,
<collectorName id="66747C911A10563EFA680272804822E6">Yeshitla M.</collectorName>
(CAF, CSCHM, NHMAA); 360 exx. (males, females), ditto, &quot;Aste wuha&quot;
<quantity id="DE5CA6F81945941BAF5634F5DFBDB6D0" metricMagnitude="3" metricUnit="m" metricValue="3.4" unit="m" value="3400.0">
<elevation id="94C14E291E6FB679AF110AC5CE5F785A" metricMagnitude="3" metricUnit="m" metricValue="3.4" unit="m" value="3400.0">3400 m</elevation>
</quantity>
,
<geoCoordinate id="02546DDBD51B6BFC1A40A0595F54D33B" degrees="10" direction="north" minutes="24" orientation="latitude" precision="925" value="10.4">10°24'N</geoCoordinate>
,
<geoCoordinate id="191EDA8BF893920878D79BD06AF57EC2" degrees="39" direction="east" minutes="48" orientation="longitude" precision="925" value="39.8">39°48'E</geoCoordinate>
,
<collectingDate id="B1C4F064232940DB7E50F1C8A6AE7B23" value="2022-05-19">19.V.2022</collectingDate>
, leg.
<collectorName id="A8D269644C1025EE216634E76AB55C95">J. Schmidt</collectorName>
,
<collectorName id="4B0F88EE2DB751259B587A735AA22206">Yeshitla M.</collectorName>
(CAF, CSCHM, NHMAA); 11 exx. (males, females), ditto, &quot;Yegana Natural Forest&quot;, river valley,
<quantity id="BFCD849C7187F24905FDC78F1B225EC8" metricMagnitude="3" metricUnit="m" metricValue="3.125" unit="m" value="3125.0">
<elevation id="2C153E805569FC8EC8494D133BC18336" metricMagnitude="3" metricUnit="m" metricValue="3.125" unit="m" value="3125.0">3125 m</elevation>
</quantity>
,
<geoCoordinate id="FBE06F84CF5508F31B82C3561E64C2E5" degrees="10" direction="north" minutes="26" orientation="latitude" precision="15" seconds="03" value="10.434167">10°26'03&quot;N</geoCoordinate>
,
<geoCoordinate id="B64F887AE4BE7A0E2992323E8AC75C4C" degrees="39" direction="east" minutes="47" orientation="longitude" precision="15" seconds="16" value="39.787777">39°47'16&quot;E</geoCoordinate>
,
<collectingDate id="77940C8C412D973F70AF68F797AA2666" value="2022-05-20">20.V.2022</collectingDate>
, leg.
<collectorName id="5B2516B03533ADBAC84D1C80179BDC5E">J. Schmidt</collectorName>
,
<collectorName id="FD905D84AEFB0A0CBE0E9867A37E67B6">Yeshitla M.</collectorName>
(CSCHM)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2127731CAA73C08E8CBB5A19AC41720F" pageId="0" pageNumber="311" type="description">
<paragraph id="BEC03FF6982A00E36A9CE9ABE1EA7A7C" pageId="0" pageNumber="311">Additions to the species description.</paragraph>
<paragraph id="1390A5AF4E0B13CA9E3DDDA14A597F09" pageId="0" pageNumber="311">
Mature species with elytra blackish brown, and with suture and first interval reddish brown lightened in most specimens (
<bibRefCitation id="5408EFDDDB415632A9ABA2A9097F33CA" DOI="https://doi.org/10.12976/jib/2021.27.1.1" author="Queinnec, E" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="311" pagination="1 - 70" refId="B43" refString="Queinnec, E, Ollivier, E, Reeb, C, 2021. Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae). Journal of Insect Biodiversity 27 (1): 1 - 70, DOI: https://doi.org/10.12976/jib/2021.27.1.1" title="Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae)." url="https://doi.org/10.12976/jib/2021.27.1.1" volume="27" year="2021">
<normalizedToken id="09781BC524DD7EC28C77006DED141EE7" originalValue="Quéinnec">Queinnec</normalizedToken>
et al. 2021
</bibRefCitation>
: &quot;suture distinctly darkened&quot;; this information is probably based on confusion with
<taxonomicName id="ABCA8B5151E131715D3F77D3A307607B" family="Carabidae" kingdom="Animalia" lsidName="T. bipartitus" order="Coleoptera" pageId="0" pageNumber="311" rank="species" species="bipartitus">
<emphasis id="6D299191245D821776C05D15AB983A31" italics="true" pageId="0" pageNumber="311">T. bipartitus</emphasis>
</taxonomicName>
). Body length 4.1-5.0 mm (
<bibRefCitation id="14B3B24372B452555D3AC2C24C3B1BCB" DOI="https://doi.org/10.12976/jib/2021.27.1.1" author="Queinnec, E" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="311" pagination="1 - 70" refId="B43" refString="Queinnec, E, Ollivier, E, Reeb, C, 2021. Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae). Journal of Insect Biodiversity 27 (1): 1 - 70, DOI: https://doi.org/10.12976/jib/2021.27.1.1" title="Insights on Ethiopian montane ground beetle biodiversity: Taxonomic study of afro-alpine and sub-alpine Trechini (Coleoptera: Carabidae: Trechinae)." url="https://doi.org/10.12976/jib/2021.27.1.1" volume="27" year="2021">
<normalizedToken id="226ABFAD2CC959B91437C25E26AD4FF2" originalValue="Quéinnec">Queinnec</normalizedToken>
et al. 2021
</bibRefCitation>
: 4.1-4.3 mm). Length of aedeagus 0.70-0.80 mm. PW/PL = 1.28-1.38; n = 20. PW/TL = 1.08-1.13; n = 20. EL/AL = 3.00-3.33; n = 10.
</paragraph>
</subSubSection>
<subSubSection id="C48547513521E7221A7D77A8E56B5A84" pageId="0" pageNumber="311" type="differential diagnosis">
<paragraph id="946EDE8CB4339091EA174C9FC50E83FF" pageId="0" pageNumber="311">Differential diagnosis.</paragraph>
<paragraph id="9966432A73F742748E8F40392198E7C0" pageId="0" pageNumber="311">
See Key to species of the subgenus
<taxonomicName id="B2579CAB5DE863181CD490AB5527AFBE" authorityName="Schmidt &amp; Faille" authorityYear="2023" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus (Abunetrechus)" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="subGenus" subGenus="Abunetrechus">Trechus Abunetrechus</taxonomicName>
, below.
</paragraph>
</subSubSection>
<subSubSection id="E0A2F81EC1A910C5B13AF99CBF71354D" pageId="0" pageNumber="311" type="distribution">
<paragraph id="81D5986606306B71B5395EF365441B87" pageId="0" pageNumber="311">Distribution.</paragraph>
<paragraph id="5BABD9F80F37461FD41DB385EE127384" pageId="0" pageNumber="311">Endemic to the Guassa Plateau of the northern Ethiopian Highlands.</paragraph>
</subSubSection>
<subSubSection id="513768E461CE2C7982DC3E7F2EC2F836" pageId="0" pageNumber="311" type="habitat">
<paragraph id="B67375F7816E98680BFFAB95F3F1EFD9" pageId="0" pageNumber="311">Habitat.</paragraph>
<paragraph id="D364FFB810DDD2FF670050F65C5B5FAF" pageId="0" pageNumber="311">
Specimens of
<taxonomicName id="F481FDF6C51860119AE6EDC877577327" family="Carabidae" kingdom="Animalia" lsidName="T. habeshanicus" order="Coleoptera" pageId="0" pageNumber="311" rank="species" species="habeshanicus">
<emphasis id="7EDF329100FE52CCCCB68307B64ADBE1" italics="true" pageId="0" pageNumber="311">T. habeshanicus</emphasis>
</taxonomicName>
were found syntopic with
<taxonomicName id="647BA163E07F3173F5F3BB0C07AE064A" family="Carabidae" kingdom="Animalia" lsidName="T. guassaensis" order="Coleoptera" pageId="0" pageNumber="311" rank="species" species="guassaensis">
<emphasis id="C96DCA37E461E37EC7AE59EF45270179" italics="true" pageId="0" pageNumber="311">T. guassaensis</emphasis>
</taxonomicName>
(
<normalizedToken id="5BC8A095C9B98B4E6DF9FEB1356F40BC" originalValue="Quéinnec">Queinnec</normalizedToken>
&amp; Ollivier) and two hitherto undescribed
<taxonomicName id="0F38491BEF5751BE4113A28CACDE588A" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="genus">
<emphasis id="A8A176DF13FD6F3A20EE06D9D1990220" italics="true" pageId="0" pageNumber="311">Trechus</emphasis>
</taxonomicName>
species in humus and rotten plant material along brooks in the afromontane zone.
</paragraph>
<caption id="48B7F9760FF6E6C87EC96253BE28F08F" doi="10.3897/dez.70.107425.figures17-20" httpUri="https://binary.pensoft.net/fig/913117" pageId="0" pageNumber="311" start="Figures 17–20" startId="F7">
<paragraph id="8A98DA6245A11343885128AD5F2324CB" pageId="0" pageNumber="311">
<emphasis id="AD23E15BAA32081E3B4E42DAF3AA4A4C" bold="true" pageId="0" pageNumber="311">Figures 17-20.</emphasis>
<taxonomicName id="73C82C71E8CBCB7325715660DE09FFFB" authority="Abunetrechus" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus (Abunetrechus) subsp. subgenus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="subGenus" subGenus="Abunetrechus">Trechus subgenus Abunetrechus</taxonomicName>
nov., dorsal aspect of body (
<emphasis id="E1F9BDF05D4A301C4F90169DCC1090B4" bold="true" pageId="0" pageNumber="311">17, 19</emphasis>
), head and pronotum (
<emphasis id="C2690155D6E82353B407AC27D40A0831" bold="true" pageId="0" pageNumber="311">18, 20</emphasis>
).
<emphasis id="6B66407DEDEC9BB011AA2B240E07835A" bold="true" pageId="0" pageNumber="311">17, 18.</emphasis>
<taxonomicName id="A36231813984CF009EEB9EF56D67715E" family="Carabidae" kingdom="Animalia" lsidName="T. bipartitus" order="Coleoptera" pageId="0" pageNumber="311" rank="species" species="bipartitus">
<emphasis id="17AF2F03B45D020A52AC4386B9690000" italics="true" pageId="0" pageNumber="311">T. bipartitus</emphasis>
</taxonomicName>
Raffray;
<emphasis id="A6000C819A91FD9D9C8B3BCD1493F3A6" bold="true" pageId="0" pageNumber="311">19, 20.</emphasis>
<taxonomicName id="DEB2E5F8B48E71F2A2FB178C88A071D4" family="Carabidae" kingdom="Animalia" lsidName="T. lalibelae" order="Coleoptera" pageId="0" pageNumber="311" rank="species" species="lalibelae">
<emphasis id="E05109F3604F4B3E54438F000C7E1324" italics="true" pageId="0" pageNumber="311">T. lalibelae</emphasis>
</taxonomicName>
<normalizedToken id="F6A87EC9A6CFFA2DA61C9B5A2768A50B" originalValue="Quéinnec">Queinnec</normalizedToken>
&amp; Ollivier. The small white circles in Figs
<figureCitation id="6BD2495893216DA8B660802034C3B619" captionStart="Figures 17–20" captionStartId="F7" captionText="Figures 17 - 20. Trechus subgenus Abunetrechus nov., dorsal aspect of body (17, 19), head and pronotum (18, 20). 17, 18. T. bipartitus Raffray; 19, 20. T. lalibelae Queinnec &amp; Ollivier. The small white circles in Figs 18 and 20 mark the insertion points of the clypeal setae." figureDoi="10.3897/dez.70.107425.figures17-20" httpUri="https://binary.pensoft.net/fig/913117" pageId="0" pageNumber="311">18</figureCitation>
and
<figureCitation id="F558CDB7E5EB98BBE9F43C45647070E1" captionStart="Figures 17–20" captionStartId="F7" captionText="Figures 17 - 20. Trechus subgenus Abunetrechus nov., dorsal aspect of body (17, 19), head and pronotum (18, 20). 17, 18. T. bipartitus Raffray; 19, 20. T. lalibelae Queinnec &amp; Ollivier. The small white circles in Figs 18 and 20 mark the insertion points of the clypeal setae." figureDoi="10.3897/dez.70.107425.figures17-20" httpUri="https://binary.pensoft.net/fig/913117" pageId="0" pageNumber="311">20</figureCitation>
mark the insertion points of the clypeal setae.
</paragraph>
</caption>
<caption id="1B0FD78E16DAD3E1E2B7FB2C78346860" doi="10.3897/dez.70.107425.figures21-23" httpUri="https://binary.pensoft.net/fig/913118" pageId="0" pageNumber="311" start="Figures 21–23" startId="F8">
<paragraph id="73FA7A79CA28922E7E28403ADF02DD92" pageId="0" pageNumber="311">
<emphasis id="B4B70E3FB32938D5C9BDF4E4AB5B6C69" bold="true" pageId="0" pageNumber="311">Figures 21-23.</emphasis>
<taxonomicName id="0A7C09AFE5CB977C3372F385C1502CBE" authority="Abunetrechus" class="Insecta" family="Carabidae" genus="Trechus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trechus (Abunetrechus) subsp. subgenus" order="Coleoptera" pageId="0" pageNumber="311" phylum="Arthropoda" rank="subGenus" subGenus="Abunetrechus">Trechus subgenus Abunetrechus</taxonomicName>
nov., dorsal aspect of body (
<emphasis id="2BDCE49A56FE2612A272B7B8619BEAEC" bold="true" pageId="0" pageNumber="311">21</emphasis>
) and head (
<emphasis id="290B51281DE0712416C52954DA529501" bold="true" pageId="0" pageNumber="311">22</emphasis>
), and anterior part of left elytron (
<emphasis id="30EA266A670E6D2D3FCDE2E63176DFEE" bold="true" pageId="0" pageNumber="311">23</emphasis>
).
<emphasis id="8609DFE7F7CB041B638B8CAA5001907F" bold="true" pageId="0" pageNumber="311">21, 22.</emphasis>
<taxonomicName id="6C9EA7D07B6EE6B29D0562F3767A5F75" family="Carabidae" kingdom="Animalia" lsidName="T. habeshaicus" order="Coleoptera" pageId="0" pageNumber="311" rank="species" species="habeshaicus">
<emphasis id="89D91974FE51EB729DE96F78154FACDE" italics="true" pageId="0" pageNumber="311">T. habeshaicus</emphasis>
</taxonomicName>
<normalizedToken id="D49457ED12DE8469649C6656DA1384AB" originalValue="Quéinnec">Queinnec</normalizedToken>
&amp; Ollivier;
<emphasis id="06C351F66BE474F5CAAB12305F020A6C" bold="true" pageId="0" pageNumber="311">23.</emphasis>
<taxonomicName id="107BE469659C3F14512A2DB5E72CCD45" family="Carabidae" kingdom="Animalia" lsidName="T. bipartitus" order="Coleoptera" pageId="0" pageNumber="311" rank="species" species="bipartitus">
<emphasis id="882854FA4F3C71626B8678F4947A6F85" italics="true" pageId="0" pageNumber="311">T. bipartitus</emphasis>
</taxonomicName>
Raffray. The small white circles in Fig.
<figureCitation id="E5DFD9DC58EAB8F2FDC6F6D2BE3A962C" captionStart="Figures 21–23" captionStartId="F8" captionText="Figures 21 - 23. Trechus subgenus Abunetrechus nov., dorsal aspect of body (21) and head (22), and anterior part of left elytron (23). 21, 22. T. habeshaicus Queinnec &amp; Ollivier; 23. T. bipartitus Raffray. The small white circles in Fig. 22 mark the insertion points of the clypeal setae; the arrow in Fig. 23 points to the insertion of the anterior elytral discal seta." figureDoi="10.3897/dez.70.107425.figures21-23" httpUri="https://binary.pensoft.net/fig/913118" pageId="0" pageNumber="311">22</figureCitation>
mark the insertion points of the clypeal setae; the arrow in Fig.
<figureCitation id="70699C56E8B8C06455D365AC7255E6B7" captionStart="Figures 21–23" captionStartId="F8" captionText="Figures 21 - 23. Trechus subgenus Abunetrechus nov., dorsal aspect of body (21) and head (22), and anterior part of left elytron (23). 21, 22. T. habeshaicus Queinnec &amp; Ollivier; 23. T. bipartitus Raffray. The small white circles in Fig. 22 mark the insertion points of the clypeal setae; the arrow in Fig. 23 points to the insertion of the anterior elytral discal seta." figureDoi="10.3897/dez.70.107425.figures21-23" httpUri="https://binary.pensoft.net/fig/913118" pageId="0" pageNumber="311">23</figureCitation>
points to the insertion of the anterior elytral discal seta.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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